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<document id="19177BEE6396007E368E691B961AE955" ID-CLB-Dataset="7325" ID-DOI="10.5852/ejt.2021.745.1311" ID-GBIF-Dataset="ac540bdb-8587-411a-8ab1-a2115525c618" ID-ISSN="2118-9773" ID-Zenodo-Dep="4690362" ID-ZooBank="6C85E664-6DE6-442A-9410-D94254E429F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618441012519" checkinUser="plazi" docAuthor="Yeshwanth, H. M. &amp; Konstantinov, Fedor V." docDate="2021" docId="BE301C0DF86CFF82FD098CE47438FB7D" docLanguage="en" docName="ejt-745_1-69.pdf" docOrigin="European Journal of Taxonomy 745" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Harpedona Distant 1904" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="42096475F864FF8BFF8A896B714FFF90" masterDocTitle="Review of the plant bug tribe Eccritotarsini (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka with description of two new genera and six new species" masterLastPageNumber="69" masterPageNumber="1" pageNumber="9" updateTime="1698936684705" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F2CFF73E799447400E8751357C3B9067">Review of the plant bug tribe Eccritotarsini (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka with description of two new genera and six new species</mods:title>
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<mods:name id="5B0A74F3329B528E6135941375B1A360" type="personal">
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<mods:roleTerm id="BBD64F06B860C3B5FB91E968CB0BE3E3">Author</mods:roleTerm>
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<mods:namePart id="0AF5E88A5122FFF26539519BECBEF626">Yeshwanth, H. M.</mods:namePart>
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<mods:affiliation id="99A943916DEAAC0B99B7A4FFE4757472">Department of Entomology, University of Agricultural Sciences (GKVK), Bangalore 560 065, India.</mods:affiliation>
<mods:nameIdentifier id="7CEACD0B898A620E5DE58BECC62E2B56" type="email">hmyeshwanth@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C04BA84213920AC0EC02A1113155C66A">Konstantinov, Fedor V.</mods:namePart>
<mods:nameIdentifier id="42EFF4C67C5EEA04FE4E54C75649DBBA" type="LSID">B5DF0683-A68F-4075-9B0C-171DFADD75E6</mods:nameIdentifier>
<mods:affiliation id="669B155F8FFA543074EB89D9A19F2FB4">Saint Petersburg State University, 7 / 9 Universitetskaya Emb., St. Petersburg 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="B80750C65FBAE7D91DF5068ABC196D31" type="email">f.konstantinov@spbu.ru</mods:nameIdentifier>
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<mods:title id="A5F725AAA9784EFC06B53E38092EDD95">European Journal of Taxonomy</mods:title>
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<mods:part id="87ABF4ADAD8FC2CBC68ECABFB59DBF97">
<mods:date id="8E685188CD3C5494BED8A31494D09370">2021</mods:date>
<mods:detail id="AAAEFCEB0D7A871B3F38D898B3982746" type="volume">
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<mods:classification id="FA5E325DC14280095D7CDA55249F9ED9">journal article</mods:classification>
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<mods:identifier id="FB5DC2D7D531978C8E17C2DC6FDCD296" type="DOI">10.5852/ejt.2021.745.1311</mods:identifier>
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<treatment id="BE301C0DF86CFF82FD098CE47438FB7D" ID-DOI="http://doi.org/10.5281/zenodo.4695073" ID-GBIF-Taxon="181011232" ID-Zenodo-Dep="4695073" LSID="urn:lsid:plazi:treatment:BE301C0DF86CFF82FD098CE47438FB7D" httpUri="http://treatment.plazi.org/id/BE301C0DF86CFF82FD098CE47438FB7D" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="7E83FE90F86CFF83FD098CE472FEFA3A" box="[643,945,1423,1450]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="3626AD1BF86CFF83FD098CE472FEFA3A" blockId="8.[563,1025,1423,1486]" box="[643,945,1423,1450]" pageId="8" pageNumber="9">
<heading id="6D6E1A77F86CFF83FD098CE472FEFA3A" box="[643,945,1423,1450]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="F199D698F86CFF83FD098CE472FEFA3A" ID-CoL="4TTY" ID-ENA="566903" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[643,945,1423,1450]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="04ED7109F86CFF83FD098CE47243FA3A" bold="true" box="[643,780,1423,1450]" italics="true" pageId="8" pageNumber="9">Harpedona</emphasis>
Distant, 1904
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<subSubSection id="7E83FE90F86CFF83FDB98CD8754EFA5E" box="[563,1025,1459,1486]" pageId="8" pageNumber="9" type="description">
<paragraph id="3626AD1BF86CFF83FDB98CD8754EFA5E" blockId="8.[563,1025,1423,1486]" box="[563,1025,1459,1486]" pageId="8" pageNumber="9">
<figureCitation id="AEA2B19EF86CFF83FDB98CD873FAFA5E" box="[563,693,1459,1486]" captionStart="Fig" captionStartId="43.[189,232,1894,1920]" captionTargetBox="[255,1348,414,1811]" captionTargetId="figure-12@43.[189,1398,376,1856]" captionTargetPageId="43" captionText="Fig. 2. Habitus of Ernestinus, Harpedona and Lopidolon spp. A. Ernestinus mimicus Distant, 1911, ♂, UASB. B. Ernestinus ramkeshariae Yasunaga &amp; Ishikawa, 2016, ♂, UASB. C. Harpedona sanguinipes Distant, 1904, ♂, UASB. D. Harpedona vittlaensis sp. nov, ♀, UASB. E–F. Lopidolon dandeliensis sp. nov., UASB. E. ♂. F. ♀." figureDoi="http://doi.org/10.5281/zenodo.4690366" httpUri="https://zenodo.org/record/4690366/files/figure.png" pageId="8" pageNumber="9">Figs 2C–D</figureCitation>
,
<figureCitation id="AEA2B19EF86CFF83FD4B8CD87248FA5E" box="[705,775,1459,1486]" captionStart="Fig" captionStartId="46.[189,232,1824,1850]" captionTargetBox="[263,1330,330,1736]" captionTargetId="figure-17@46.[189,1398,297,1786]" captionTargetPageId="46" captionText="Fig. 5. Dorsal view and label data of type specimens. A–C. Dioclerus bengalicus Stonedahl, 1988, ♀, holotype, BPBM.D–E. Dioclerus lutheri Stonedahl, 1988, ♂, holotype, AMNH_PBI 00338430, FMNH. F–G. Dioclerus praefectus Distant,1910, lectotype, AMNH_PBI 00340384, NHM. H–I. Ernestinus mimicus Distant,1911, ♂, syntype, AMNH_PBI 00085668, BMNH(E) 1633157, NHM. J–K. Harpedona marginata Distant, 1904, ♂, holotype, NHM. L–M. Harpedona sanguinipes Distant, 1909, ♂, lectotype, NHM." figureDoi="http://doi.org/10.5281/zenodo.4690372" httpUri="https://zenodo.org/record/4690372/files/figure.png" pageId="8" pageNumber="9">5J–M</figureCitation>
,
<figureCitation id="AEA2B19EF86CFF83FC998CD87211FA5D" box="[787,862,1459,1485]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="55.[189,232,1927,1953]" captionStartId-1="56.[189,232,1928,1954]" captionStartId-2="57.[189,232,1928,1954]" captionTargetBox-0="[310,1285,295,1867]" captionTargetBox-1="[209,1328,425,1859]" captionTargetBox-2="[302,1333,446,1858]" captionTargetId-0="figure-52@55.[227,1360,265,1889]" captionTargetId-1="figure-57@56.[189,1398,401,1890]" captionTargetId-2="figure-53@57.[189,1398,408,1890]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionText-0="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." captionText-1="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." captionText-2="Fig. 16. Male genitalia of Harpedona vittlaensis sp. nov., holotype, UASB. A. Genital capsule in lateral view. B–C. Left paramere in dorsal and lateral views. D–E. Right paramere in dorsal and lateral views. F. Aedeagus in lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4690392" figureDoi-1="http://doi.org/10.5281/zenodo.4690395" figureDoi-2="http://doi.org/10.5281/zenodo.4690397" httpUri-0="https://zenodo.org/record/4690392/files/figure.png" httpUri-1="https://zenodo.org/record/4690395/files/figure.png" httpUri-2="https://zenodo.org/record/4690397/files/figure.png" pageId="8" pageNumber="9">14–16</figureCitation>
,
<figureCitation id="AEA2B19EF86CFF83FCE18CD872EFFA5E" box="[875,928,1459,1486]" captionStart="Fig" captionStartId="66.[189,232,1893,1919]" captionTargetBox="[190,1392,292,1805]" captionTargetId="figure-72@66.[186,1401,286,1856]" captionTargetPageId="66" captionText="Fig. 25. Scent gland evaporatory area. A. Harpedona vittlaensis sp. nov., UASB. B. Lopidolon dandeliensis sp. nov., UASB. C. Mertila rubrocephala sp. nov., UASB. D. Stonedahlia mishmiensis gen. et sp. nov., UASB. E. Namyatovia castlerockensis gen. et sp. nov., UASB. F. Namyatovia sirsiensis gen. et sp. nov., UASB." figureDoi="http://doi.org/10.5281/zenodo.4690417" httpUri="https://zenodo.org/record/4690417/files/figure.png" pageId="8" pageNumber="9">25A</figureCitation>
,
<figureCitation id="AEA2B19EF86CFF83FC268CDF754EFA5E" box="[940,1025,1460,1486]" captionStart="Fig" captionStartId="68.[189,232,1893,1919]" captionTargetBox="[197,1389,264,1854]" captionTargetId="figure-72@68.[197,1389,264,1854]" captionTargetPageId="68" captionText="Fig. 27. Habitats and living specimens. A–B. Harpedona sanguinipes Distant, 1909 on Colocasia esculenta. A. Nymphs.B. Mating adults.C. Prodromus clypeatusDistant, 1904 on banana.D. Stonedahlia mishmiensis gen. et sp. nov. on Colocasia sp. E. Lopidolon dandeliensis sp. nov. on Diplocentrum recurvum. All specimens at UASB." figureDoi="http://doi.org/10.5281/zenodo.4690421" httpUri="https://zenodo.org/record/4690421/files/figure.png" pageId="8" pageNumber="9">27A–B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86CFF83FF378C9D73BEF9C3" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="3626AD1BF86CFF83FF378C9D7375F980" blockId="8.[189,570,1526,1552]" box="[189,570,1526,1552]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="16898A35F86CFF83FF378C9D7375F980" box="[189,570,1526,1552]" pageId="8" pageNumber="9">
<treatmentCitation id="B7388B0AF86CFF83FF378C9D7379F980" author="Distant W. L." box="[189,566,1526,1552]" page="418" pageId="8" pageNumber="9" year="1904">
<taxonomicName id="F199D698F86CFF83FF378C9D7379F980" ID-CoL="4TTY" authority="Distant, 1904 b: 418" authorityName="Distant" authorityPageNumber="418" authorityYear="1904" box="[189,566,1526,1552]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="04ED7109F86CFF83FF378C9D700CF980" box="[189,323,1526,1552]" italics="true" pageId="8" pageNumber="9">Harpedona</emphasis>
<bibRefCitation id="5208D0EAF86CFF83FEC08C9D7379F980" author="Distant W. L." box="[330,566,1526,1552]" pageId="8" pageNumber="9" refId="ref21440" refString="Distant W. L. 1904 b. Rhynchota. Vol. 2. In: Blanford W. T. (ed.) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London. Available from: https: // www. biodiversitylibrary. org / page / 12704190" type="url" year="1904">Distant, 1904b: 418</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3626AD1BF86CFF83FF378F5373BEF9C3" blockId="8.[189,753,1592,1619]" box="[189,753,1592,1619]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="16898A35F86CFF83FF378F5373BEF9C3" box="[189,753,1592,1619]" pageId="8" pageNumber="9">
<taxonomicName id="F199D698F86CFF83FF378F53700CF9C2" ID-CoL="4TTY" authorityName="Distant" authorityYear="1904" box="[189,323,1592,1618]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="04ED7109F86CFF83FF378F53700CF9C2" box="[189,323,1592,1618]" italics="true" pageId="8" pageNumber="9">Harpedona</emphasis>
</taxonomicName>
–
<treatmentCitation id="B7388B0AF86CFF83FEEA8F53733FF9C2" author="Stonedahl G. M." box="[352,624,1592,1618]" page="16" pageId="8" pageNumber="9" year="1988">
<bibRefCitation id="5208D0EAF86CFF83FEEA8F53733FF9C2" author="Stonedahl G. M." box="[352,624,1592,1618]" pageId="8" pageNumber="9" pagination="1 - 99" refId="ref23015" refString="Stonedahl G. M. 1988. Revisions of Dioclerus, Harpedona, Mertila, Myiocapsus, Prodromus and Thaumastomiris (Heteroptera: Miridae, Bryocorinae: Eccritotarsini). Bulletin of the American Museum of Natural History 187: 1 - 99." type="journal article" year="1988">Stonedahl 1988: 16–32</bibRefCitation>
</treatmentCitation>
(revision).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86CFF83FF378F117293F954" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="3626AD1BF86CFF83FF378F117017F904" blockId="8.[189,344,1658,1684]" box="[189,344,1658,1684]" pageId="8" pageNumber="9">
<heading id="6D6E1A77F86CFF83FF378F117017F904" bold="true" box="[189,344,1658,1684]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="04ED7109F86CFF83FF378F117017F904" bold="true" box="[189,344,1658,1684]" pageId="8" pageNumber="9">
<typeStatus id="E92213B9F86CFF83FF378F1171B5F904" box="[189,250,1658,1684]" pageId="8" pageNumber="9">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="3626AD1BF86CFF83FF378FC27293F954" blockId="8.[189,988,1705,1732]" box="[189,988,1705,1732]" pageId="8" pageNumber="9">
<taxonomicName id="F199D698F86CFF83FF378FC27326F953" ID-CoL="6LDF8" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[189,617,1705,1731]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="04ED7109F86CFF83FF378FC2708CF953" box="[189,451,1705,1731]" italics="true" pageId="8" pageNumber="9">Harpedona marginata</emphasis>
Distant, 1904
</taxonomicName>
(by monotypy;
<bibRefCitation id="5208D0EAF86CFF83FCAF8FC27280F954" author="Distant W. L." box="[805,975,1705,1732]" pageId="8" pageNumber="9" refId="ref21440" refString="Distant W. L. 1904 b. Rhynchota. Vol. 2. In: Blanford W. T. (ed.) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London. Available from: https: // www. biodiversitylibrary. org / page / 12704190" type="url" year="1904">Distant 1904b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86CFF82FF378F8073F5FE45" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="3626AD1BF86CFF83FF378F807079F895" blockId="8.[189,310,1771,1797]" box="[189,310,1771,1797]" pageId="8" pageNumber="9">
<heading id="6D6E1A77F86CFF83FF378F807079F895" bold="true" box="[189,310,1771,1797]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="04ED7109F86CFF83FF378F807079F895" bold="true" box="[189,310,1771,1797]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="3626AD1BF86CFF82FF378E7073F5FE45" blockId="8.[188,1399,1819,2025]" lastBlockId="9.[188,1399,265,469]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Recognized by the following combination of characters: body narrow, parallel-sided, pronotum at base less than 1.5× as wide as head (
<figureCitation id="AEA2B19EF86CFF83FDA08E5473E1F8C9" box="[554,686,1855,1881]" captionStart="Fig" captionStartId="43.[189,232,1894,1920]" captionTargetBox="[255,1348,414,1811]" captionTargetId="figure-12@43.[189,1398,376,1856]" captionTargetPageId="43" captionText="Fig. 2. Habitus of Ernestinus, Harpedona and Lopidolon spp. A. Ernestinus mimicus Distant, 1911, ♂, UASB. B. Ernestinus ramkeshariae Yasunaga &amp; Ishikawa, 2016, ♂, UASB. C. Harpedona sanguinipes Distant, 1904, ♂, UASB. D. Harpedona vittlaensis sp. nov, ♀, UASB. E–F. Lopidolon dandeliensis sp. nov., UASB. E. ♂. F. ♀." figureDoi="http://doi.org/10.5281/zenodo.4690366" httpUri="https://zenodo.org/record/4690366/files/figure.png" pageId="8" pageNumber="9">Figs 2C–D</figureCitation>
,
<figureCitation id="AEA2B19EF86CFF83FD338E55724EF8C9" box="[697,769,1854,1881]" captionStart="Fig" captionStartId="46.[189,232,1824,1850]" captionTargetBox="[263,1330,330,1736]" captionTargetId="figure-17@46.[189,1398,297,1786]" captionTargetPageId="46" captionText="Fig. 5. Dorsal view and label data of type specimens. A–C. Dioclerus bengalicus Stonedahl, 1988, ♀, holotype, BPBM.D–E. Dioclerus lutheri Stonedahl, 1988, ♂, holotype, AMNH_PBI 00338430, FMNH. F–G. Dioclerus praefectus Distant,1910, lectotype, AMNH_PBI 00340384, NHM. H–I. Ernestinus mimicus Distant,1911, ♂, syntype, AMNH_PBI 00085668, BMNH(E) 1633157, NHM. J–K. Harpedona marginata Distant, 1904, ♂, holotype, NHM. L–M. Harpedona sanguinipes Distant, 1909, ♂, lectotype, NHM." figureDoi="http://doi.org/10.5281/zenodo.4690372" httpUri="https://zenodo.org/record/4690372/files/figure.png" pageId="8" pageNumber="9">5J–M</figureCitation>
); coloration of dorsum uniform, dirty yellow to black, sometimes with paler embolium, legs contrastingly whitish or reddish yellow, comparatively long and slender, apex of hind femur reaching or surpassing apex of abdomen; dorsum clothed with dense, short, adpressed, silvery simple setae; pronotal collar wide and large; calli medially confluent, dorsally and laterally separated by deep impressed lines giving appearance of three distinct sections of pronotum; genital capsule highly modified, with large, lobate process on right margin, and usually with bulbous, twisted and spinelike outgrowths on dorsal margin (
<figureCitation id="AEA2B19EF86DFF82FC9E884772EAFED7" box="[788,933,300,327]" captionStart="Fig" captionStartId="55.[189,232,1927,1953]" captionTargetBox="[310,1285,295,1867]" captionTargetId="figure-52@55.[227,1360,265,1889]" captionTargetPageId="55" captionText="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690392" httpUri="https://zenodo.org/record/4690392/files/figure.png" pageId="9" pageNumber="10">Figs 14A–C</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FC3A88477548FED7" box="[944,1031,300,327]" captionStart="Fig" captionStartId="56.[189,232,1928,1954]" captionTargetBox="[209,1328,425,1859]" captionTargetId="figure-57@56.[189,1398,401,1890]" captionTargetPageId="56" captionText="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690395" httpUri="https://zenodo.org/record/4690395/files/figure.png" pageId="9" pageNumber="10">15A–B</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FB9888477507FED7" box="[1042,1096,300,327]" captionStart="Fig" captionStartId="57.[189,232,1928,1954]" captionTargetBox="[302,1333,446,1858]" captionTargetId="figure-53@57.[189,1398,408,1890]" captionTargetPageId="57" captionText="Fig. 16. Male genitalia of Harpedona vittlaensis sp. nov., holotype, UASB. A. Genital capsule in lateral view. B–C. Left paramere in dorsal and lateral views. D–E. Right paramere in dorsal and lateral views. F. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690397" httpUri="https://zenodo.org/record/4690397/files/figure.png" pageId="9" pageNumber="10">16A</figureCitation>
); parameres and aedeagus reduced, very small; left paramere with long and thin, twisted apical process (
<figureCitation id="AEA2B19EF86DFF82FBFF883B7446FEFA" box="[1141,1289,336,362]" captionStart="Fig" captionStartId="55.[189,232,1927,1953]" captionTargetBox="[310,1285,295,1867]" captionTargetId="figure-52@55.[227,1360,265,1889]" captionTargetPageId="55" captionText="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690392" httpUri="https://zenodo.org/record/4690392/files/figure.png" pageId="9" pageNumber="10">Figs 14D–E</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FA93883B743FFEFB" box="[1305,1392,336,363]" captionStart="Fig" captionStartId="56.[189,232,1928,1954]" captionTargetBox="[209,1328,425,1859]" captionTargetId="figure-57@56.[189,1398,401,1890]" captionTargetPageId="56" captionText="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690395" httpUri="https://zenodo.org/record/4690395/files/figure.png" pageId="9" pageNumber="10">15C–D</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FF378818705AFE1E" box="[189,277,371,398]" captionStart="Fig" captionStartId="57.[189,232,1928,1954]" captionTargetBox="[302,1333,446,1858]" captionTargetId="figure-53@57.[189,1398,408,1890]" captionTargetPageId="57" captionText="Fig. 16. Male genitalia of Harpedona vittlaensis sp. nov., holotype, UASB. A. Genital capsule in lateral view. B–C. Left paramere in dorsal and lateral views. D–E. Right paramere in dorsal and lateral views. F. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690397" httpUri="https://zenodo.org/record/4690397/files/figure.png" pageId="9" pageNumber="10">16B–C</figureCitation>
); right paramere simple, with tapering incurved apex (
<figureCitation id="AEA2B19EF86DFF82FC18881872B3FE1D" box="[914,1020,371,397]" captionStart="Fig" captionStartId="55.[189,232,1927,1953]" captionTargetBox="[310,1285,295,1867]" captionTargetId="figure-52@55.[227,1360,265,1889]" captionTargetPageId="55" captionText="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690392" httpUri="https://zenodo.org/record/4690392/files/figure.png" pageId="9" pageNumber="10">Figs 14F</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FB8288187518FE1E" box="[1032,1111,371,398]" captionStart="Fig" captionStartId="56.[189,232,1928,1954]" captionTargetBox="[209,1328,425,1859]" captionTargetId="figure-57@56.[189,1398,401,1890]" captionTargetPageId="56" captionText="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690395" httpUri="https://zenodo.org/record/4690395/files/figure.png" pageId="9" pageNumber="10">15E–F</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FBE9881875F4FE1E" box="[1123,1211,371,398]" captionStart="Fig" captionStartId="57.[189,232,1928,1954]" captionTargetBox="[302,1333,446,1858]" captionTargetId="figure-53@57.[189,1398,408,1890]" captionTargetPageId="57" captionText="Fig. 16. Male genitalia of Harpedona vittlaensis sp. nov., holotype, UASB. A. Genital capsule in lateral view. B–C. Left paramere in dorsal and lateral views. D–E. Right paramere in dorsal and lateral views. F. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690397" httpUri="https://zenodo.org/record/4690397/files/figure.png" pageId="9" pageNumber="10">16D–E</figureCitation>
); aedeagus with entirely membranous phallotheca, simple membranous endosoma and sclerotized, spine-like apical part of ductus seminis (
<figureCitation id="AEA2B19EF86DFF82FE1388D17345FE44" box="[409,522,442,468]" captionStart="Fig" captionStartId="55.[189,232,1927,1953]" captionTargetBox="[310,1285,295,1867]" captionTargetId="figure-52@55.[227,1360,265,1889]" captionTargetPageId="55" captionText="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690392" httpUri="https://zenodo.org/record/4690392/files/figure.png" pageId="9" pageNumber="10">Figs 14G</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FD9C88D17320FE45" box="[534,623,442,469]" captionStart="Fig" captionStartId="56.[189,232,1928,1954]" captionTargetBox="[209,1328,425,1859]" captionTargetId="figure-57@56.[189,1398,401,1890]" captionTargetPageId="56" captionText="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690395" httpUri="https://zenodo.org/record/4690395/files/figure.png" pageId="9" pageNumber="10">15G–H</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FDF188D173E2FE45" box="[635,685,442,469]" captionStart="Fig" captionStartId="57.[189,232,1928,1954]" captionTargetBox="[302,1333,446,1858]" captionTargetId="figure-53@57.[189,1398,408,1890]" captionTargetPageId="57" captionText="Fig. 16. Male genitalia of Harpedona vittlaensis sp. nov., holotype, UASB. A. Genital capsule in lateral view. B–C. Left paramere in dorsal and lateral views. D–E. Right paramere in dorsal and lateral views. F. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690397" httpUri="https://zenodo.org/record/4690397/files/figure.png" pageId="9" pageNumber="10">16F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86DFF82FF3788907218FDF8" pageId="9" pageNumber="10" type="distribution">
<paragraph id="3626AD1BF86DFF82FF3788907019FD85" blockId="9.[189,342,507,533]" box="[189,342,507,533]" pageId="9" pageNumber="10">
<heading id="6D6E1A77F86DFF82FF3788907019FD85" bold="true" box="[189,342,507,533]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="04ED7109F86DFF82FF3788907019FD85" bold="true" box="[189,342,507,533]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="3626AD1BF86DFF82FF378B407218FDF8" blockId="9.[186,1398,554,617]" pageId="9" pageNumber="10">
Known from
<collectingCountry id="4E8EED8BF86DFF82FEDC8B417086FDD4" box="[342,457,554,580]" name="Sri Lanka" pageId="9" pageNumber="10">Sri Lanka</collectingCountry>
, southwestern and northern
<taxonomicName id="F199D698F86DFF82FC838B41720AFDD4" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[777,837,554,580]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="37" phylum="Tracheophyta" rank="genus">India</taxonomicName>
to Fukien Prov. of
<collectingCountry id="4E8EED8BF86DFF82FB918B41752BFDD4" box="[1051,1124,554,580]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
,
<collectingCountry id="4E8EED8BF86DFF82FBE78B4175BBFDD4" box="[1133,1268,554,580]" name="Philippines" pageId="9" pageNumber="10">Philippines</collectingCountry>
,
<collectingCountry id="4E8EED8BF86DFF82FB778B41743DFDD4" box="[1277,1394,554,580]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
, New
<collectingCountry id="4E8EED8BF86DFF82FF738B25701EFDF8" box="[249,337,590,616]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
, and the
<collectingCountry id="4E8EED8BF86DFF82FE338B257331FDF8" box="[441,638,590,616]" name="Solomon Islands" pageId="9" pageNumber="10">Solomon Islands</collectingCountry>
(
<bibRefCitation id="5208D0EAF86DFF82FD078B257204FDF8" author="Stonedahl G. M." box="[653,843,590,616]" pageId="9" pageNumber="10" pagination="1 - 99" refId="ref23015" refString="Stonedahl G. M. 1988. Revisions of Dioclerus, Harpedona, Mertila, Myiocapsus, Prodromus and Thaumastomiris (Heteroptera: Miridae, Bryocorinae: Eccritotarsini). Bulletin of the American Museum of Natural History 187: 1 - 99." type="journal article" year="1988">Stonedahl 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86DFF82FF378BFB728EFC5B" pageId="9" pageNumber="10" type="discussion">
<paragraph id="3626AD1BF86DFF82FF378BFB7060FD3A" blockId="9.[189,303,656,682]" box="[189,303,656,682]" pageId="9" pageNumber="10">
<heading id="6D6E1A77F86DFF82FF378BFB7060FD3A" bold="true" box="[189,303,656,682]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="04ED7109F86DFF82FF378BFB7060FD3A" bold="true" box="[189,303,656,682]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="3626AD1BF86DFF82FF378BD5725EFC1A" blockId="9.[188,1401,702,906]" pageId="9" pageNumber="10">
This genus was revised by
<bibRefCitation id="5208D0EAF86DFF82FD8F8BD5739CFD49" author="Stonedahl G. M." box="[517,723,702,729]" pageId="9" pageNumber="10" pagination="1 - 99" refId="ref23015" refString="Stonedahl G. M. 1988. Revisions of Dioclerus, Harpedona, Mertila, Myiocapsus, Prodromus and Thaumastomiris (Heteroptera: Miridae, Bryocorinae: Eccritotarsini). Bulletin of the American Museum of Natural History 187: 1 - 99." type="journal article" year="1988">Stonedahl (1988)</bibRefCitation>
and, with addition of the recently described
<taxonomicName id="F199D698F86DFF82FB798BD573C8FD6C" authority="Yasunaga &amp; Ishikawa, 2016" authorityName="Yasunaga" authorityYear="2016" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stonedahli">
<emphasis id="04ED7109F86DFF82FB798BD57078FD6C" italics="true" pageId="9" pageNumber="10">Harpedona stonedahli</emphasis>
<bibRefCitation id="5208D0EAF86DFF82FEB78B8973C8FD6C" author="Yasunaga Y. &amp; Ishikawa T." box="[317,647,738,764]" pageId="9" pageNumber="10" pagination="411 - 469" refId="ref23196" refString="Yasunaga Y. &amp; Ishikawa T. 2016. Twelve new species of the Asian plant bug genus Ernestinus Distant (Heteroptera: Miridae: Bryocorinae: Eccritotarsini), with emphasis on unique biology and descriptions of three additional new eccritotarsine species. Insect Systematics &amp; Evolution 47 (5): 411 - 469. https: // doi. org / 10.1163 / 1876312 X- 47052149" type="journal article" year="2016">Yasunaga &amp; Ishikawa, 2016</bibRefCitation>
</taxonomicName>
, it currently comprises 13 species. Only
<taxonomicName id="F199D698F86DFF82FBD78B897010FC8F" authority="Distant, 1909" authorityName="Distant" authorityYear="1909" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanguinipes">
<emphasis id="04ED7109F86DFF82FBD78B89743AFD6C" box="[1117,1397,738,764]" italics="true" pageId="9" pageNumber="10">Harpedona sanguinipes</emphasis>
<bibRefCitation id="5208D0EAF86DFF82FF368A6E7010FC8F" author="Distant W. L." box="[188,351,773,799]" pageId="9" pageNumber="10" pagination="440 - 454" refId="ref21497" refString="Distant W. L. 1909. Descriptions of Oriental Capsidae. Annals and Magazine of Natural History 8 (4): 440 - 454. https: // doi. org / 10.1080 / 00222930908692696" type="journal article" year="1909">Distant, 1909</bibRefCitation>
</taxonomicName>
was known from
<taxonomicName id="F199D698F86DFF82FDC98A6E7330FC8F" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[579,639,773,799]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="37" phylum="Tracheophyta" rank="genus">India</taxonomicName>
prior to our study; the description of a new species from the
<collectingRegion id="F45D63F9F86DFF82FF378A427037FCD3" box="[189,376,809,835]" country="India" name="Karnataka" pageId="9" pageNumber="10">Karnataka state</collectingRegion>
of
<taxonomicName id="F199D698F86DFF82FE238A4270AAFCD3" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[425,485,809,835]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="37" phylum="Tracheophyta" rank="genus">India</taxonomicName>
is given below.
<taxonomicName id="F199D698F86DFF82FD338A427538FCD3" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[697,1143,809,835]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="04ED7109F86DFF82FD338A42728BFCD3" box="[697,964,809,835]" italics="true" pageId="9" pageNumber="10">Harpedona marginata</emphasis>
Distant, 1904
</taxonomicName>
, the most frequently collected and widely distributed species of the genus was originally described from
<collectingCountry id="4E8EED8BF86DFF82FB1D8A277445FCF6" box="[1175,1290,844,870]" name="Sri Lanka" pageId="9" pageNumber="10">Sri Lanka</collectingCountry>
and may likely be found in
<taxonomicName id="F199D698F86DFF82FE1E8A1B709FFC1A" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[404,464,880,906]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="37" phylum="Tracheophyta" rank="genus">India</taxonomicName>
with more sampling effort.
</paragraph>
<paragraph id="3626AD1BF86DFF82FF378ADA728EFC5B" blockId="9.[189,961,945,972]" box="[189,961,945,972]" pageId="9" pageNumber="10">
<heading id="6D6E1A77F86DFF82FF378ADA728EFC5B" bold="true" box="[189,961,945,972]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="04ED7109F86DFF82FF378ADA728EFC5B" bold="true" box="[189,961,945,972]" pageId="9" pageNumber="10">
Key to species of the genus
<taxonomicName id="F199D698F86DFF82FD988ADA73D4FC5C" authorityName="Distant" authorityYear="1904" box="[530,667,945,972]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="04ED7109F86DFF82FD988ADA73D4FC5C" bold="true" box="[530,667,945,972]" italics="true" pageId="9" pageNumber="10">Harpedona</emphasis>
</taxonomicName>
of
<taxonomicName id="F199D698F86DFF82FD488ADA724AFC5B" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[706,773,945,971]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="37" phylum="Tracheophyta" rank="genus">India</taxonomicName>
and
<collectingCountry id="4E8EED8BF86DFF82FCC88ADA728EFC5B" box="[834,961,945,971]" name="Sri Lanka" pageId="9" pageNumber="10">Sri Lanka</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7E83FE90F86DFF82FF378A8B7438FB7D" pageId="9" pageNumber="10" type="key">
<key id="4208FA2AF86DFF82FF378A8B7438FB7D" pageId="9" pageNumber="10">
<keyStep id="8D6DB5BEF86DFF82FF378A8B7438FB8E" pageId="9" pageNumber="10">
<paragraph id="3626AD1BF86DFF82FF378A8B7438FC6B" blockId="9.[189,1399,991,1054]" box="[189,1399,991,1019]" pageId="9" pageNumber="10">
<keyLead id="8D680E2EF86DFF82FF378A8B7438FC6B" box="[189,1399,991,1019]" pageId="9" pageNumber="10">
1. Pronotum dirty orange, with distinct punctures (
<figureCitation id="AEA2B19EF86DFF82FC948A8B7235FC6A" box="[798,890,992,1018]" captionStart="Fig" captionStartId="43.[189,232,1894,1920]" captionTargetBox="[255,1348,414,1811]" captionTargetId="figure-12@43.[189,1398,376,1856]" captionTargetPageId="43" captionText="Fig. 2. Habitus of Ernestinus, Harpedona and Lopidolon spp. A. Ernestinus mimicus Distant, 1911, ♂, UASB. B. Ernestinus ramkeshariae Yasunaga &amp; Ishikawa, 2016, ♂, UASB. C. Harpedona sanguinipes Distant, 1904, ♂, UASB. D. Harpedona vittlaensis sp. nov, ♀, UASB. E–F. Lopidolon dandeliensis sp. nov., UASB. E. ♂. F. ♀." figureDoi="http://doi.org/10.5281/zenodo.4690366" httpUri="https://zenodo.org/record/4690366/files/figure.png" pageId="9" pageNumber="10">Fig. 2D</figureCitation>
) ................................
<taxonomicName id="F199D698F86DFF82FBFD8A8B745AFC6A" authority="Yeshwanth &amp; Konstantinov, 2021" authorityName="Yeshwanth &amp; Konstantinov" authorityYear="2021" box="[1143,1301,991,1018]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vittlaensis" status="sp. nov.">
<emphasis id="04ED7109F86DFF82FBFD8A8B745AFC6A" bold="true" box="[1143,1301,991,1018]" italics="true" pageId="9" pageNumber="10">H. vittlaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FDECC72F86DFF82FA968A8A7438FC6B" box="[1308,1399,993,1019]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3626AD1BF86DFF82FF4B8D6F7438FB8E" blockId="9.[189,1399,991,1054]" box="[193,1399,1027,1054]" pageId="9" pageNumber="10">
<keyLead id="8D680E2EF86DFF82FF4B8D6F7438FB8E" box="[193,1399,1027,1054]" pageId="9" pageNumber="10">
<emphasis id="04ED7109F86DFF82FF4B8D6F719FFB8D" box="[193,208,1028,1053]" italics="true" pageId="9" pageNumber="10">–</emphasis>
Pronotum dark brown, smooth or very faintly punctate (
<figureCitation id="AEA2B19EF86DFF82FCF08D687295FB8E" box="[890,986,1027,1054]" captionStart="Fig" captionStartId="43.[189,232,1894,1920]" captionTargetBox="[255,1348,414,1811]" captionTargetId="figure-12@43.[189,1398,376,1856]" captionTargetPageId="43" captionText="Fig. 2. Habitus of Ernestinus, Harpedona and Lopidolon spp. A. Ernestinus mimicus Distant, 1911, ♂, UASB. B. Ernestinus ramkeshariae Yasunaga &amp; Ishikawa, 2016, ♂, UASB. C. Harpedona sanguinipes Distant, 1904, ♂, UASB. D. Harpedona vittlaensis sp. nov, ♀, UASB. E–F. Lopidolon dandeliensis sp. nov., UASB. E. ♂. F. ♀." figureDoi="http://doi.org/10.5281/zenodo.4690366" httpUri="https://zenodo.org/record/4690366/files/figure.png" pageId="9" pageNumber="10">Figs 2C</figureCitation>
,
<figureCitation id="AEA2B19EF86DFF82FC6D8D68756DFB8E" box="[999,1058,1027,1054]" captionStart="Fig" captionStartId="46.[189,232,1824,1850]" captionTargetBox="[263,1330,330,1736]" captionTargetId="figure-17@46.[189,1398,297,1786]" captionTargetPageId="46" captionText="Fig. 5. Dorsal view and label data of type specimens. A–C. Dioclerus bengalicus Stonedahl, 1988, ♀, holotype, BPBM.D–E. Dioclerus lutheri Stonedahl, 1988, ♂, holotype, AMNH_PBI 00338430, FMNH. F–G. Dioclerus praefectus Distant,1910, lectotype, AMNH_PBI 00340384, NHM. H–I. Ernestinus mimicus Distant,1911, ♂, syntype, AMNH_PBI 00085668, BMNH(E) 1633157, NHM. J–K. Harpedona marginata Distant, 1904, ♂, holotype, NHM. L–M. Harpedona sanguinipes Distant, 1909, ♂, lectotype, NHM." figureDoi="http://doi.org/10.5281/zenodo.4690372" httpUri="https://zenodo.org/record/4690372/files/figure.png" pageId="9" pageNumber="10">5J, L</figureCitation>
) ......................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8D6DB5BEF86DFF82FF378D2E7438FB7D" pageId="9" pageNumber="10">
<paragraph id="3626AD1BF86DFF82FF378D2E7438FB36" blockId="9.[189,1399,1093,1262]" pageId="9" pageNumber="10">
<keyLead id="8D680E2EF86DFF82FF378D2E7438FB36" pageId="9" pageNumber="10">
2. Frons in males with a distinct longitudinal sulcus (
<figureCitation id="AEA2B19EF86DFF82FCD98D2E72E5FBF0" box="[851,938,1093,1120]" captionStart="Fig" captionStartId="46.[189,232,1824,1850]" captionTargetBox="[263,1330,330,1736]" captionTargetId="figure-17@46.[189,1398,297,1786]" captionTargetPageId="46" captionText="Fig. 5. Dorsal view and label data of type specimens. A–C. Dioclerus bengalicus Stonedahl, 1988, ♀, holotype, BPBM.D–E. Dioclerus lutheri Stonedahl, 1988, ♂, holotype, AMNH_PBI 00338430, FMNH. F–G. Dioclerus praefectus Distant,1910, lectotype, AMNH_PBI 00340384, NHM. H–I. Ernestinus mimicus Distant,1911, ♂, syntype, AMNH_PBI 00085668, BMNH(E) 1633157, NHM. J–K. Harpedona marginata Distant, 1904, ♂, holotype, NHM. L–M. Harpedona sanguinipes Distant, 1909, ♂, lectotype, NHM." figureDoi="http://doi.org/10.5281/zenodo.4690372" httpUri="https://zenodo.org/record/4690372/files/figure.png" pageId="9" pageNumber="10">Fig. 5J</figureCitation>
). Labium at most reaching mesocoxa. Femora pale yellow. Phallotheca with a large sclerotized sac (
<figureCitation id="AEA2B19EF86DFF82FC378D037567FB13" box="[957,1064,1128,1155]" captionStart="Fig" captionStartId="55.[189,232,1927,1953]" captionTargetBox="[310,1285,295,1867]" captionTargetId="figure-52@55.[227,1360,265,1889]" captionTargetPageId="55" captionText="Fig. 14. Male genitalia of Harpedona marginata Distant, 1904, NHM. A–C. Genital capsule in ventral and lateral views. D–E. Right paramere in dorsal and lateral views. F. Right paramere in dorsal view. G. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690392" httpUri="https://zenodo.org/record/4690392/files/figure.png" pageId="9" pageNumber="10">Fig. 14G</figureCitation>
) .......................................... .................................................................................................................
<taxonomicName id="F199D698F86DFF82FBA48DE77438FB36" ID-CoL="6LDF8" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[1070,1399,1164,1190]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="04ED7109F86DFF82FBA48DE7759DFB36" bold="true" box="[1070,1234,1164,1190]" italics="true" pageId="9" pageNumber="10">H. marginata</emphasis>
Distant, 1904
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3626AD1BF86DFF82FF4B8DDB7438FB7D" blockId="9.[189,1399,1093,1262]" pageId="9" pageNumber="10">
<keyLead id="8D680E2EF86DFF82FF4B8DDB7438FB7D" pageId="9" pageNumber="10">
<emphasis id="04ED7109F86DFF82FF4B8DDB719FFB59" box="[193,208,1200,1225]" italics="true" pageId="9" pageNumber="10">–</emphasis>
Frons without sulcus. Labium surpassing metacoxae. Femora with a distinct reddish tinge (
<figureCitation id="AEA2B19EF86DFF82FA878DC47426FB5A" box="[1293,1385,1199,1226]" captionStart="Fig" captionStartId="43.[189,232,1894,1920]" captionTargetBox="[255,1348,414,1811]" captionTargetId="figure-12@43.[189,1398,376,1856]" captionTargetPageId="43" captionText="Fig. 2. Habitus of Ernestinus, Harpedona and Lopidolon spp. A. Ernestinus mimicus Distant, 1911, ♂, UASB. B. Ernestinus ramkeshariae Yasunaga &amp; Ishikawa, 2016, ♂, UASB. C. Harpedona sanguinipes Distant, 1904, ♂, UASB. D. Harpedona vittlaensis sp. nov, ♀, UASB. E–F. Lopidolon dandeliensis sp. nov., UASB. E. ♂. F. ♀." figureDoi="http://doi.org/10.5281/zenodo.4690366" httpUri="https://zenodo.org/record/4690366/files/figure.png" pageId="9" pageNumber="10">Fig. 2C</figureCitation>
). Phallotheca membranous, without a sclerotized sac (
<figureCitation id="AEA2B19EF86DFF82FCDB8DB872AFFB7E" box="[849,992,1235,1262]" captionStart="Fig" captionStartId="56.[189,232,1928,1954]" captionTargetBox="[209,1328,425,1859]" captionTargetId="figure-57@56.[189,1398,401,1890]" captionTargetPageId="56" captionText="Fig. 15. Male genitalia of Harpedona sanguinipes Distant, 1904, UASB. A–B. Genital capsule in dorsal and ventral views. C–D. Left paramere in lateral view. E–F. Right paramere in dorsal view. G–H. Aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4690395" httpUri="https://zenodo.org/record/4690395/files/figure.png" pageId="9" pageNumber="10">Fig. 15G–H</figureCitation>
) .....
<taxonomicName id="F199D698F86DFF82FB908DB87438FB7D" ID-CoL="6LDH6" authority="Distant, 1909" authorityName="Distant" authorityYear="1909" box="[1050,1399,1235,1261]" class="Insecta" family="Miridae" genus="Harpedona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanguinipes">
<emphasis id="04ED7109F86DFF82FB908DB8759EFB7D" bold="true" box="[1050,1233,1235,1261]" italics="true" pageId="9" pageNumber="10">H. sanguinipes</emphasis>
<bibRefCitation id="5208D0EAF86DFF82FB528DB87438FB7D" author="Distant W. L." box="[1240,1399,1235,1261]" pageId="9" pageNumber="10" pagination="440 - 454" refId="ref21497" refString="Distant W. L. 1909. Descriptions of Oriental Capsidae. Annals and Magazine of Natural History 8 (4): 440 - 454. https: // doi. org / 10.1080 / 00222930908692696" type="journal article" year="1909">Distant, 1909</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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