1021 lines
158 KiB
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1021 lines
158 KiB
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<document id="65D692165769C701D3592B420C09912D" ID-DOI="10.11646/zootaxa.5541.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14241160" ID-ZooBank="4F812F60-9242-4F44-8E25-99381FD7E8B3" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1732835906282" checkinUser="plazi" docAuthor="Martinez-Soares, Pablo, Roux, Michel, Giusberti, Luca, Gatto, Roberto, Eléaume, Marc & Améziane, Nadia" docDate="2024" docId="2F1AC921FFDDFFD8F2F0FC18FBC25D28" docLanguage="en" docName="zootaxa.5541.4.1.pdf" docOrigin="Zootaxa 5541 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holopus fabianii Martinez-Soares, Roux, Giusberti, Gatto, Eléaume & Améziane, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="423" masterDocId="D323B159FFCDFFCEF267FFEBFFFE5C4D" masterDocTitle="New Eocene species of the crinoid genera Holopus and Cyathidium (Cyrtocrinida: Holopodidae) from north-eastern Italy" masterLastPageNumber="437" masterPageNumber="401" pageNumber="417" updateTime="1733513786105" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3E0472898EBC3425EBCB8031E6531AB6">New Eocene species of the crinoid genera Holopus and Cyathidium (Cyrtocrinida: Holopodidae) from north-eastern Italy</mods:title>
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<mods:namePart id="38FB431401D79EB9F13FBD93ECE219AC">Martinez-Soares, Pablo</mods:namePart>
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<mods:affiliation id="C294BD19BD05AEF8B055E193552A91CD">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national d’Histoire naturelle, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="269A3338D390E5BA9F022132B66C7A95">Roux, Michel</mods:namePart>
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<mods:affiliation id="41F7C53822709BA1E8EDE8762F3B39DE">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national d’Histoire naturelle, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<mods:nameIdentifier id="B62BB1FA57FE0C1623EFDB2AF7B58864" type="email">michel.roux51@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7534CF32DD634CA59D137BD8F4A168FB">Giusberti, Luca</mods:namePart>
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<mods:namePart id="9DFD3BB44B26F8EAEEF5BD474A09A2F6">Gatto, Roberto</mods:namePart>
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<mods:namePart id="80B2217F823F9C79363A4C6A3037961A">Eléaume, Marc</mods:namePart>
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<mods:affiliation id="72CB7F811DA27C4394BD6CAA66CCDC3E">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national d’Histoire naturelle, 57 rue Cuvier, 75005 Paris, France. & Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national d’Histoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
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<mods:namePart id="A240EC1317B800C228C2225B525F304B">Améziane, Nadia</mods:namePart>
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<mods:affiliation id="999F03DF250996289FD9ABD6CA62D72B">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national d’Histoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
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<mods:nameIdentifier id="343628006541FE079A1AB6666ACE408F" type="email">nadia.ameziane@mnhn.fr</mods:nameIdentifier>
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<mods:title id="BB6403D64B2A22A663153301F5E6AFA7">Zootaxa</mods:title>
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<mods:date id="B2B720BA18E0907D9529CEE4D4DBAB29">2024</mods:date>
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<taxonomicName id="60B303B4FFDDFFDEF2F0FC18FE9A5843" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[151,356,1011,1038]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="16" pageNumber="417" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
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<emphasis id="95C7A425FFDDFFDEF2F0FC18FE9A5843" bold="true" box="[151,356,1011,1038]" italics="true" pageId="16" pageNumber="417">Holopus fabianii</emphasis>
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<taxonomicNameLabel id="8EF4195EFFDDFFDEF30CFC18FE375840" box="[363,457,1011,1037]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="EFA92BBCFFDDFFDEF2F0FBFCFA995839" pageId="16" pageNumber="417" type="description">
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<paragraph id="A70C7837FFDDFFDEF2F0FBFCFDF9587C" blockId="16.[151,519,1011,1074]" box="[151,519,1047,1074]" pageId="16" pageNumber="417">
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<figureCitation id="3F8864B2FFDDFFDEF2F0FBFCFF1B587F" box="[151,229,1047,1074]" captionStart="FIGURE 7" captionStartId="11.[151,250,810,834]" captionTargetBox="[210,1376,188,783]" captionTargetId="figure-19@11.[204,1383,181,785]" captionTargetPageId="11" captionText="FIGURE 7. Main quantitative characters of aboral cup and IBrax in fossil Holopodidae. S: Cyathidium holopus from Seymour Island. Straight lines: values of H/D and H/Wd. Other values in mm." figureDoi="http://doi.org/10.5281/zenodo.14241174" httpUri="https://zenodo.org/record/14241174/files/figure.png" pageId="16" pageNumber="417">Figs. 7</figureCitation>
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,
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<figureCitation id="3F8864B2FFDDFFDEF296FBFCFED1587C" box="[241,303,1047,1073]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="16" pageNumber="417">8H–J</figureCitation>
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,
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<figureCitation id="3F8864B2FFDDFFDEF35CFBFCFE8B587C" box="[315,373,1047,1073]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="17.[151,250,798,822]" captionStartId-1="19.[151,250,1793,1817]" captionStartId-2="20.[151,250,1793,1817]" captionTargetBox-0="[202,1386,187,770]" captionTargetBox-1="[151,1436,181,1769]" captionTargetBox-2="[151,1436,181,1769]" captionTargetId-0="figure-19@17.[196,1392,181,774]" captionTargetId-1="figure-19@19.[151,1436,181,1769]" captionTargetId-2="figure-29@20.[151,1436,181,1769]" captionTargetPageId-0="17" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURE 9. Main quantitative characters of Holopodidae from the Eocene of north-eastern Italy. Cy: Cyathidium, Ho: Holopus, Ho.a.: juvenile of H. alidis (MNHN-IE-2014-2185). Straight lines: values of H/D and H/Wd. Other values in mm." captionText-1="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." captionText-2="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14241178" figureDoi-1="http://doi.org/10.5281/zenodo.14241180" figureDoi-2="http://doi.org/10.5281/zenodo.14241182" httpUri-0="https://zenodo.org/record/14241178/files/figure.png" httpUri-1="https://zenodo.org/record/14241180/files/figure.png" httpUri-2="https://zenodo.org/record/14241182/files/figure.png" pageId="16" pageNumber="417">9–11</figureCitation>
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;
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||
<tableCitation id="EA314D8CFFDDFFDEF3E7FBFCFDF9587C" box="[384,519,1047,1073]" captionStart-0="TABLE 7" captionStart-1="TABLE 8" captionStart-2="TABLE 9" captionStart-3="TABLE 10" captionStartId-0="21.[152,244,543,567]" captionStartId-1="21.[152,244,1163,1187]" captionStartId-2="22.[152,244,404,428]" captionStartId-3="22.[152,244,1573,1597]" captionTargetPageId-0="21" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="22" captionText-0="TABLE 7. Quantitative characters of the eight aboral cups in the type-series of Holopus fabianii sp. nov. Abbreviations: see Fig. 1A. *: estimated value (broken ossicle). D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33232; Paratype 1: MGP-PD 33233; Paratypes 2–7: MGP-PD 33234." captionText-1="TABLE 8. Quantitative characters of IBrax of Holopus fabianii sp. nov. belonging to figured paratypes. Abbreviations: see Fig. 1B. Wd used as growth index. Values in mm except ratios. Paratypes 8–19: MGP-PD 33235." captionText-2="TABLE 9. Main quantitative characters of IIBr from the proximal arm series (except keystone plates) of Holopus fabianii sp. nov. belonging to the type-series. Abbreviations: see Fig. 1C. Wp used as growth index. Values in mm except ratios. Paratypes 20–23: MGP-PD 33236; Paratypes 25–39: MGP-PD 33238." captionText-3="TABLE 10. Main quantitative characters of distal IIBr and keystone plates of Holopus fabianii sp. nov. belonging to the type-series.Abbreviations: see Fig. 1C. H for keystone plates: height of triangular external face. Wp used as growth index. Values in mm except ratios. Paratypes 24: MGP-PD 33236; Paratypes 40–45: MGP-PD 33238." httpUri-0="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFDF4FC835E32" httpUri-1="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFB60FB30588A" httpUri-2="http://table.plazi.org/id/F3CC28BFFFDBFFD8F2FFFE7FFC795DB8" httpUri-3="http://table.plazi.org/id/F3CC28BFFFDBFFD8F2FFF9CEFB735AC8" pageId="16" pageNumber="417" tableUuid-0="F3CC28BFFFD8FFDBF2FFFDF4FC835E32" tableUuid-1="F3CC28BFFFD8FFDBF2FFFB60FB30588A" tableUuid-2="F3CC28BFFFDBFFD8F2FFFE7FFC795DB8" tableUuid-3="F3CC28BFFFDBFFD8F2FFF9CEFB735AC8">Tables 7–10</tableCitation>
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</paragraph>
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<paragraph id="A70C7837FFDDFFDEF2F0FBB0FA995839" blockId="16.[151,1383,1115,1140]" box="[151,1383,1115,1140]" pageId="16" pageNumber="417">
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<emphasis id="95C7A425FFDDFFDEF2F0FBB0FEF0583E" bold="true" box="[151,270,1115,1139]" pageId="16" pageNumber="417">Synonymy.</emphasis>
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<taxonomicName id="60B303B4FFDDFFDEF373FBB7FCD65839" authority="Manni & Pacioni, 2021: 69 - 73" authorityName="Manni & Pacioni" authorityPageNumber="69 - 73" authorityYear="2021" box="[276,808,1116,1140]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="16" pageNumber="417" phylum="Echinodermata" rank="species" species="spileccense">
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<emphasis id="95C7A425FFDDFFDEF373FBB7FE1A5839" box="[276,484,1116,1140]" italics="true" pageId="16" pageNumber="417">Holopus spileccense</emphasis>
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<bibRefCitation id="C32205C6FFDDFFDEF38DFBB7FCD65839" author="Manni, R. & Pacioni, E." box="[490,808,1116,1140]" pageId="16" pageNumber="417" pagination="65 - 77" refId="ref22121" refString="Manni, R. & Pacioni, E. (2021) New records of the cyrtocrinid crinoids Cyathidium gastaldii (Michelin, 1851) and Holopus spileccense (Schluter, 1878) from Eocene strata in Veneto, north-east Italy. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen, 301 (1), 65 - 77. https: // doi. org / 10.1127 / njgpa / 2021 / 0997" type="journal article" year="2021">Manni & Pacioni, 2021: 69–73</bibRefCitation>
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</taxonomicName>
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, figs. 4–6;
|
||
<bibRefCitation id="C32205C6FFDDFFDEF1FFFBB7FB715839" author="Frisone, V. & Preto, N. & Pisera, A. & Agnini, C. & Giusberti, L. & Papazzoni, C. A. & De Angeli, A. & Beschin, C. & Mietto, P. & Quaggiotto, E. & Monaco, P. & Dominici, S. & Kiessling, W. & Luciani, V. & Roux, M. & Bosellini, F. R." box="[920,1167,1116,1140]" pageId="16" pageNumber="417" pagination="299 - 313" refId="ref21160" refString="Frisone, V., Preto, N., Pisera, A., Agnini, C., Giusberti, L., Papazzoni, C. A., De Angeli, A., Beschin, C., Mietto, P., Quaggiotto, E., Monaco, P., Dominici, S., Kiessling, W., Luciani, V., Roux, M. & Bosellini, F. R. (2020) A first glimpse on the taphonomy and sedimentary environment of the Eocene siliceous sponges from Chiampo, Lessini Mts, NE Italy. Bollettino della Societa Paleontologica Italiana, 59 (3), 299 - 313." type="journal article" year="2020">
|
||
Frisone
|
||
<emphasis id="95C7A425FFDDFFDEF18DFBB7FBE95839" box="[1002,1047,1116,1140]" italics="true" pageId="16" pageNumber="417">et al</emphasis>
|
||
., 2020: 306
|
||
</bibRefCitation>
|
||
(
|
||
<emphasis id="95C7A425FFDDFFDEF6F9FBB7FB3A583E" box="[1182,1220,1116,1139]" italics="true" pageId="16" pageNumber="417">non</emphasis>
|
||
<bibRefCitation id="C32205C6FFDDFFDEF6ADFBB7FAA25839" author="Schluter, C." box="[1226,1372,1116,1140]" pageId="16" pageNumber="417" pagination="28 - 66" refId="ref23579" refString="Schluter, C. (1878) Uber einige astylide Crinoiden. Zeitschrift der Deutschen Geologischen Gesellschaft, 30 (1), 28 - 66." type="journal article" year="1878">Schlüter 1878</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDDFFDEF2F0FB49FB6F5901" pageId="16" pageNumber="417" type="materials_examined">
|
||
<paragraph id="A70C7837FFDDFFDEF2F0FB49FC435965" blockId="16.[151,1437,1186,1968]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2F0FB49FEBA58F1" bold="true" box="[151,324,1186,1212]" pageId="16" pageNumber="417">Type material.</emphasis>
|
||
<materialsCitation id="17DB726AFFDDFFDEF32CFB49FC6558F1" box="[331,923,1186,1212]" collectionCode="H, I" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCount="8" specimenCount-egg="8" typeStatus="holotype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF32CFB49FE7658F1" box="[331,392,1186,1212]" count="8" pageId="16" pageNumber="417" type="egg">Eight</specimenCount>
|
||
aboral cups including the
|
||
<typeStatus id="7808C695FFDDFFDEF0CBFB49FCF058F1" box="[684,782,1186,1212]" pageId="16" pageNumber="417" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="3F8864B2FFDDFFDEF17AFB49FC7058F1" box="[797,910,1186,1212]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="16" pageNumber="417">
|
||
Fig. 8
|
||
<collectionCode id="C1A2E0F2FFDDFFDEF13AFB49FC8D58F1" box="[861,883,1186,1212]" pageId="16" pageNumber="417">H</collectionCode>
|
||
–
|
||
<collectionCode id="C1A2E0F2FFDDFFDEF1E3FB49FC7058F1" box="[900,910,1186,1212]" pageId="16" pageNumber="417">I</collectionCode>
|
||
</figureCitation>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="17DB726AFFDDFFDEF1C6FB49FA8B58F1" box="[929,1397,1186,1212]" collectionCode="J" country="Italy" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCode="paratype 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF1C6FB49FC3758F1" box="[929,969,1186,1212]" count="1" pageId="16" pageNumber="417" type="generic">one</specimenCount>
|
||
figured
|
||
<typeStatus id="7808C695FFDDFFDEF640FB49FB7958F1" box="[1063,1159,1186,1212]" pageId="16" pageNumber="417" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="F715D04CFFDDFFDEF6F2FB49FAF258F1" box="[1173,1292,1186,1212]" pageId="16" pageNumber="417">paratype 1</specimenCode>
|
||
,
|
||
<figureCitation id="3F8864B2FFDDFFDEF77EFB49FA9958F1" box="[1305,1383,1186,1212]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="16" pageNumber="417">Fig. 8J</figureCitation>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="17DB726AFFDDFFDEF71CFB49FDB058AD" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCode="paratypes 2-7" specimenCount="6" typeStatus="paratype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF71CFB49FA6258F1" box="[1403,1436,1186,1212]" count="6" pageId="16" pageNumber="417" type="generic">six</specimenCount>
|
||
non-figured
|
||
<typeStatus id="7808C695FFDDFFDEF344FB2DFE7058AD" box="[291,398,1222,1248]" pageId="16" pageNumber="417" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<specimenCode id="F715D04CFFDDFFDEF3F9FB2DFDBE58AD" box="[414,576,1222,1248]" pageId="16" pageNumber="417">paratypes 2–7</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="17DB726AFFDDFFDEF031FB2DFD965949" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCode="paratypes 8-19, paratypes 20-24" specimenCount="17" typeStatus="holotype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF031FB2DFD8C58AD" box="[598,626,1222,1248]" count="17" pageId="16" pageNumber="417" type="generic">17</specimenCount>
|
||
figured brachials including 12 IBrax (
|
||
<specimenCode id="F715D04CFFDDFFDEF644FB2DFB2C58AD" box="[1059,1234,1222,1248]" pageId="16" pageNumber="417">paratypes 8–19</specimenCode>
|
||
,
|
||
<figureCitation id="3F8864B2FFDDFFDEF6B8FB2DFA9A58AD" box="[1247,1380,1222,1248]" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="16" pageNumber="417">Fig. 10A–L</figureCitation>
|
||
) and five IIBr (
|
||
<specimenCode id="F715D04CFFDDFFDEF36FFB01FE385949" box="[264,454,1258,1284]" pageId="16" pageNumber="417">paratypes 20–24</specimenCode>
|
||
,
|
||
<figureCitation id="3F8864B2FFDDFFDEF3B5FB01FDA25949" box="[466,604,1258,1284]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="16" pageNumber="417">Fig. 11A–H</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="17DB726AFFDDFFDEF017FB01FBC45948" box="[624,1082,1258,1285]" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCode="paratypes 46-59" specimenCount="14" typeStatus="paratype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF017FB01FD725949" box="[624,652,1258,1284]" count="14" pageId="16" pageNumber="417" type="generic">14</specimenCount>
|
||
non-figured IBrax (
|
||
<specimenCode id="F715D04CFFDDFFDEF108FB01FBD05948" box="[879,1070,1258,1285]" pageId="16" pageNumber="417">paratypes 46–59</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="17DB726AFFDDFFDEF625FB01FC465965" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="16" pageNumber="417" specimenCode="paratypes 25-45" specimenCount="21" typeStatus="paratype">
|
||
<specimenCount id="B1B5B3BEFFDDFFDEF625FB01FBA05949" box="[1090,1118,1258,1284]" pageId="16" pageNumber="417">21</specimenCount>
|
||
non-figured IIBr (
|
||
<specimenCode id="F715D04CFFDDFFDEF748FB01FF205964" pageId="16" pageNumber="417">paratypes 25–45</specimenCode>
|
||
) including three keystone plates and three IIBr from distal series
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDDFFDEF2A0FAD9FB6F5901" blockId="16.[151,1437,1186,1968]" box="[199,1169,1330,1357]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2A0FAD9FE025901" bold="true" box="[199,508,1330,1356]" pageId="16" pageNumber="417">Other material examined.</emphasis>
|
||
17 IBrax, 58 IIBr of Wp<
|
||
<quantity id="604BD5D2FFDDFFDEF145FAD9FC845901" box="[802,890,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="417" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, 21 IIBr of Wp>
|
||
<quantity id="604BD5D2FFDDFFDEF652FAD9FB735901" box="[1077,1165,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="417" unit="mm" value="2.8">2.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDDFFDEF2A0FABDFCEE59D9" pageId="16" pageNumber="417" type="etymology">
|
||
<paragraph id="A70C7837FFDDFFDEF2A0FABDFCEE59D9" blockId="16.[151,1437,1186,1968]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2A0FABDFEB0593D" bold="true" box="[199,334,1366,1392]" pageId="16" pageNumber="417">Etymology.</emphasis>
|
||
Species dedicated to Ramiro Fabiani (1879–1954) for his contributions to stratigraphy and palaeontology of the Paleogene from north-eastern
|
||
<collectingCountry id="DFA438A7FFDDFFDEF0B0FA91FCF259D9" box="[727,780,1402,1428]" name="Italy" pageId="16" pageNumber="417">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDDFFDEF2A0FA75FF395B25" pageId="16" pageNumber="417" type="diagnosis">
|
||
<paragraph id="A70C7837FFDDFFDEF2A0FA75FF395B25" blockId="16.[151,1437,1186,1968]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2A0FA75FEBC59F5" bold="true" box="[199,322,1438,1464]" pageId="16" pageNumber="417">Diagnosis.</emphasis>
|
||
Cup with conspicuous constriction in the lower half; upper part of cup with rounded pentagonal cross-section; pronounced radial ribs and marked interradial ribs; adoral face without external interradial process; calycinal cavity unknown; estimated maximum cup size: H 21.0 mm, D
|
||
<quantity id="604BD5D2FFDDFFDEF1BCFA0DFBBA5A4D" box="[987,1092,1510,1536]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="16" pageNumber="417" unit="mm" value="14.8">14.8 mm</quantity>
|
||
; IBrax about as high as wide, with Y-shaped ridge, upper part of ridge (V-shape) irregularly doubled, usually with outer branches reaching angles of maximum brachial width (Wd) and inner branches corresponding to radial axis of brachial ramifications; small IBrax with only a few tubercles or none at all; large IBrax with several tubercles especially in the proximal half; external ornamentation of proximal IIBr consisting of X-shaped positioned tubercles, muscular synarthries with fulcral ridge shorter than maximum brachial width axis, adoral border forming an obtuse angle giving relatively narrow V-shape section; keystone plates with distal muscular synarthry roughly triangular and rounded, proximal synarthry symmetrical with fulcral ridge as long as Wp, external face smooth, shaped like an isosceles triangle, four conspicuous parallel symplexial crenulations on each side; distal IIBr series higher than broad, muscular synarthries with inconspicuous fulcral ridge, marked aboral ligament pit and axial nervous canal, conspicuous adoral muscle pits.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDDFFDEF2A0F899FB9D5BFD" pageId="16" pageNumber="417" type="materials_examined">
|
||
<paragraph id="A70C7837FFDDFFDEF2A0F899FD275BC1" blockId="16.[151,1437,1186,1968]" box="[199,729,1906,1932]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2A0F899FE915BC1" bold="true" box="[199,367,1906,1932]" pageId="16" pageNumber="417">Type stratum.</emphasis>
|
||
<geologicalTimeScale id="D593F89CFFDDFFDEF311F899FDD85BC1" box="[374,550,1906,1932]" pageId="16" pageNumber="417">Lower Lutetian</geologicalTimeScale>
|
||
, tuffite horizon.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDDFFDEF2A0F87DFB9D5BFD" blockId="16.[151,1437,1186,1968]" box="[199,1123,1942,1968]" pageId="16" pageNumber="417">
|
||
<emphasis id="95C7A425FFDDFFDEF2A0F87DFE985BFD" bold="true" box="[199,358,1942,1968]" pageId="16" pageNumber="417">Type locality.</emphasis>
|
||
<location id="A26C2EECFFDDFFDEF30BF87DFD7F5BFD" LSID="urn:lsid:plazi:treatment:2F1AC921FFDDFFD8F2F0FC18FBC25D28:A26C2EECFFDDFFDEF30BF87DFD7F5BFD" box="[364,641,1942,1968]" country="Italy" municipality="Chiampo Valley" name="Cengio dell'Orbo quarry" pageId="16" pageNumber="417">Cengio dell’Orbo quarry</location>
|
||
in the
|
||
<collectingMunicipality id="4768E24DFFDDFFDEF0A9F87DFC7A5BFD" box="[718,900,1942,1968]" pageId="16" pageNumber="417">Chiampo Valley</collectingMunicipality>
|
||
, north-eastern
|
||
<collectingCountry id="DFA438A7FFDDFFDEF64DF87DFBA15BFD" box="[1066,1119,1942,1968]" name="Italy" pageId="16" pageNumber="417">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F3CC28BFFFDCFFDFF2F0FCF5FAE55F17" ID-DOI="http://doi.org/10.5281/zenodo.14241178" ID-Zenodo-Dep="14241178" httpUri="https://zenodo.org/record/14241178/files/figure.png" pageId="17" pageNumber="418" startId="17.[151,250,798,822]" targetBox="[202,1386,187,770]" targetPageId="17" targetType="figure">
|
||
<paragraph id="A70C7837FFDCFFDFF2F0FCF5FAE55F17" blockId="17.[151,1436,798,858]" pageId="17" pageNumber="418">
|
||
<emphasis id="95C7A425FFDCFFDFF2F0FCF5FEEC5F7B" bold="true" box="[151,274,798,822]" pageId="17" pageNumber="418">FIGURE 9.</emphasis>
|
||
Main quantitative characters of
|
||
<taxonomicName id="60B303B4FFDCFFDFF030FCF5FD245F7B" authorityName="Zittel" authorityYear="1879" box="[599,730,798,822]" class="Crinoidea" family="Holopodidae" kingdom="Animalia" order="Cyrtocrinida" pageId="17" pageNumber="418" phylum="Echinodermata" rank="family">Holopodidae</taxonomicName>
|
||
from the Eocene of north-eastern Italy. Cy:
|
||
<taxonomicName id="60B303B4FFDCFFDFF6F5FCF5FAC65F7B" authority=", Ho" authorityName="Ho" box="[1170,1336,798,822]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="17" pageNumber="418" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="95C7A425FFDCFFDFF6F5FCF5FAF65F7B" box="[1170,1288,798,822]" italics="true" pageId="17" pageNumber="418">Cyathidium</emphasis>
|
||
, Ho
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="60B303B4FFDCFFDFF727FCF5FF3F5F17" authority=", Ho." authorityName="Ho." class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="17" pageNumber="418" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="95C7A425FFDCFFDFF727FCF5FA625F7B" box="[1344,1436,798,822]" italics="true" pageId="17" pageNumber="418">Holopus,</emphasis>
|
||
Ho.
|
||
</taxonomicName>
|
||
a.: juvenile of
|
||
<taxonomicName id="60B303B4FFDCFFDFF328FCA8FE585F17" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[335,422,834,858]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="17" pageNumber="418" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDCFFDFF328FCA8FE585F17" box="[335,422,834,858]" italics="true" pageId="17" pageNumber="418">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(MNHN-IE-2014-2185). Straight lines: values of H/D and H/Wd. Other values in mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EFA92BBCFFDCFFDCF2A0FC47FCEF5DB8" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418" type="description">
|
||
<paragraph id="A70C7837FFDCFFDFF2A0FC47FAE9591E" blockId="17.[151,1437,940,2011]" pageId="17" pageNumber="418">
|
||
<emphasis id="95C7A425FFDCFFDFF2A0FC47FE035F8B" bold="true" box="[199,509,940,967]" pageId="17" pageNumber="418">Description of type series.</emphasis>
|
||
Among the 8 aboral cups of the type series (quantitative characters listed in
|
||
<tableCitation id="EA314D8CFFDCFFDFF739FC47FF595FA6" captionStart="TABLE 7" captionStartId="21.[152,244,543,567]" captionTargetPageId="21" captionText="TABLE 7. Quantitative characters of the eight aboral cups in the type-series of Holopus fabianii sp. nov. Abbreviations: see Fig. 1A. *: estimated value (broken ossicle). D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33232; Paratype 1: MGP-PD 33233; Paratypes 2–7: MGP-PD 33234." httpUri="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFDF4FC835E32" pageId="17" pageNumber="418" tableUuid="F3CC28BFFFD8FFDBF2FFFDF4FC835E32">Table 7</tableCitation>
|
||
), the
|
||
<typeStatus id="7808C695FFDCFFDFF281FC3BFEB65FA7" box="[230,328,976,1002]" pageId="17" pageNumber="418" type="holotype">holotype</typeStatus>
|
||
was selected as the best preserved aboral cup (
|
||
<figureCitation id="3F8864B2FFDCFFDFF13EFC3BFC365FA6" box="[857,968,976,1003]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="17" pageNumber="418">Fig. 8H, I</figureCitation>
|
||
).
|
||
<typeStatus id="7808C695FFDCFFDFF1B9FC3BFBB85FA7" box="[990,1094,976,1002]" pageId="17" pageNumber="418" type="holotype">Holotype</typeStatus>
|
||
medium sized (H
|
||
<quantity id="604BD5D2FFDCFFDFF771FC3AFA825FA6" box="[1302,1404,977,1003]" metricMagnitude="-3" metricUnit="m" metricValue="3.83" pageId="17" pageNumber="418" unit="mm" value="3.83">3.83 mm</quantity>
|
||
, D
|
||
<quantity id="604BD5D2FFDCFFDFF2F0FC1FFEFF5842" box="[151,257,1012,1039]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="17" pageNumber="418" unit="mm" value="2.75">2.75 mm</quantity>
|
||
), with a strong constriction (Db/Dcs 1.68) located in the aboral half and separating adoral two-thirds of cup from a broad basal part with flat attachment surface with a greater diameter than the adoral side (D/Db 0.88); adoral part with rounded pentagonal cross-section demarcated by five conspicuous radial ribs with intermediate ribs more pronounced on one side (
|
||
<figureCitation id="3F8864B2FFDCFFDFF055FB8BFD725836" box="[562,652,1120,1147]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="17" pageNumber="418">Fig. 8H</figureCitation>
|
||
) than on the other (
|
||
<figureCitation id="3F8864B2FFDCFFDFF112FB8BFC385836" box="[885,966,1120,1147]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="17" pageNumber="418">Fig. 8I</figureCitation>
|
||
); adoral face without external interradial process, calycinal cavity filled with recrystallized sediment making it impossible to identify any possible internal interradial processes, if present. Other aboral cups of type series are of variable sizes (H
|
||
<quantity id="604BD5D2FFDCFFDFF6E2FB42FABC588E" box="[1157,1346,1192,1219]" metricMagnitude="-3" metricUnit="m" metricValue="4.135" metricValueMax="6.55" metricValueMin="1.72" pageId="17" pageNumber="418" unit="mm" value="4.135" valueMax="6.55" valueMin="1.72">1.72 to 6.55 mm</quantity>
|
||
, D
|
||
<quantity id="604BD5D2FFDCFFDFF70CFB43FEE258AA" metricMagnitude="-3" metricUnit="m" metricValue="3.19" metricValueMax="4.62" metricValueMin="1.76" pageId="17" pageNumber="418" unit="mm" value="3.19" valueMax="4.62" valueMin="1.76">1.76 to 4.62 mm</quantity>
|
||
), usually height greater than adoral diameter (H/D 1.21 to 1.48) except in small
|
||
<typeStatus id="7808C695FFDCFFDFF6F9FB26FB0058AA" box="[1182,1278,1229,1255]" pageId="17" pageNumber="418" type="paratype">paratype</typeStatus>
|
||
7 (H/D 0.98), H/D increasing from nearly 1 to more than 1.4 with size (
|
||
<figureCitation id="3F8864B2FFDCFFDFF14EFB1BFC915946" box="[809,879,1264,1291]" captionStart="FIGURE 9" captionStartId="17.[151,250,798,822]" captionTargetBox="[202,1386,187,770]" captionTargetId="figure-19@17.[196,1392,181,774]" captionTargetPageId="17" captionText="FIGURE 9. Main quantitative characters of Holopodidae from the Eocene of north-eastern Italy. Cy: Cyathidium, Ho: Holopus, Ho.a.: juvenile of H. alidis (MNHN-IE-2014-2185). Straight lines: values of H/D and H/Wd. Other values in mm." figureDoi="http://doi.org/10.5281/zenodo.14241178" httpUri="https://zenodo.org/record/14241178/files/figure.png" pageId="17" pageNumber="418">Fig. 9</figureCitation>
|
||
), similar characters as the
|
||
<typeStatus id="7808C695FFDCFFDFF6F8FB1BFAFA5947" box="[1183,1284,1264,1290]" pageId="17" pageNumber="418" type="holotype">holotype</typeStatus>
|
||
, more or less conspicuous, attachment surface often smaller than adoral diameter and very irregular, sometimes appearing as a mold of a subcylindrical substrate (
|
||
<figureCitation id="3F8864B2FFDCFFDFF045FAD3FD8F591E" box="[546,625,1336,1363]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.A–C: Cyathidium chiampoensis sp. nov., A–B: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), D–E: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; F–G: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; H–J: aboral cups of Holopus fabianii sp. nov., H–I: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="17" pageNumber="418">Fig. 8J</figureCitation>
|
||
), cup with moderate curvature (H/h~1.3) only in
|
||
<typeStatus id="7808C695FFDCFFDFF6FDFAD2FB04591E" box="[1178,1274,1337,1363]" pageId="17" pageNumber="418" type="paratype">paratype</typeStatus>
|
||
3.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDCFFDFF2A0FAB6FD035AF7" blockId="17.[151,1437,940,2011]" pageId="17" pageNumber="418">
|
||
Type series of IBrax (
|
||
<typeStatus id="7808C695FFDCFFDFF3D5FAB6FDE1593A" box="[434,543,1373,1399]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
|
||
8 to 19) illustrate changes in form and ornamentation with size (
|
||
<figureCitation id="3F8864B2FFDCFFDFF687FAB7FACF593A" box="[1248,1329,1372,1399]" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="17" pageNumber="418">Fig. 10</figureCitation>
|
||
,
|
||
<tableCitation id="EA314D8CFFDCFFDFF75CFAB7FA70593A" box="[1339,1422,1372,1399]" captionStart="TABLE 8" captionStartId="21.[152,244,1163,1187]" captionTargetPageId="21" captionText="TABLE 8. Quantitative characters of IBrax of Holopus fabianii sp. nov. belonging to figured paratypes. Abbreviations: see Fig. 1B. Wd used as growth index. Values in mm except ratios. Paratypes 8–19: MGP-PD 33235." httpUri="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFB60FB30588A" pageId="17" pageNumber="418" tableUuid="F3CC28BFFFD8FFDBF2FFFB60FB30588A">Table 8</tableCitation>
|
||
). Smallest IBrax nearly as high as wide; external surface (
|
||
<figureCitation id="3F8864B2FFDCFFDFF162FA6BFC7759D6" box="[773,905,1408,1435]" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="17" pageNumber="418">Fig. 10A–E</figureCitation>
|
||
) with a more or less thick Y-shaped ridge, distal part of ridge (V-shape) irregularly doubled with usually outer branches reaching angles of maximum brachial width (Wp) and inner branches corresponding to radial axis of brachial ramifications; proximal external surface without tubercles on either side of axial ridge. Larger IBrax about as wide as high except
|
||
<typeStatus id="7808C695FFDCFFDFF675FA06FB835A4A" box="[1042,1149,1517,1543]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
|
||
10 and 18 which show H/ Wd 0.82 and 0.84, respectively; external ornamentation (
|
||
<figureCitation id="3F8864B2FFDCFFDFF17FF9FBFC6D5A66" box="[792,915,1552,1579]" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="17" pageNumber="418">Fig. 10I–L</figureCitation>
|
||
) showing Y-shaped ridge composed of a short lower segment and predominantly longer V-shaped segments corresponding to radial axis of brachial ramifications, 10 to 20 large tubercles developed irregularly over the entire external surface, sometimes transforming former ridges into rows of tubercles (
|
||
<figureCitation id="3F8864B2FFDCFFDFF385F997FD945ADA" box="[482,618,1660,1687]" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="17" pageNumber="418">Fig. 10K–L</figureCitation>
|
||
). Asymmetric
|
||
<typeStatus id="7808C695FFDCFFDFF16CF996FC955ADA" box="[779,875,1661,1687]" pageId="17" pageNumber="418" type="paratype">paratype</typeStatus>
|
||
17 possibly belongs to a lateral trivium plate (
|
||
<figureCitation id="3F8864B2FFDCFFDFF708F997FF3F5AF6" captionStart="FIGURE 10" captionStartId="19.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-19@19.[151,1436,181,1769]" captionTargetPageId="19" captionText="FIGURE 10. Variation in primibrachial ornamentation in Holopus fabianii sp. nov. from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="17" pageNumber="418">Fig. 10J</figureCitation>
|
||
). Aboral interradial process moderately developed.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDCFFDCF2A0F92EFB545CB4" blockId="17.[151,1437,940,2011]" lastBlockId="18.[151,1437,151,2013]" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418">
|
||
Type series of IIBr, proximal series to keystone plate (
|
||
<typeStatus id="7808C695FFDCFFDFF16CF92EFC865A92" box="[779,888,1733,1759]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
|
||
20 to23 and 25 to 39) with external ornamentation predominantly X-shaped, always with a more or less marked axial ridge and few tubercles (
|
||
<figureCitation id="3F8864B2FFDCFFDFF6E0F903FAF25B4E" box="[1159,1292,1768,1795]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="17" pageNumber="418">Fig. 11D–E</figureCitation>
|
||
). Ratios H/h, H/Wp and Wp/Wd independent of size (
|
||
<tableCitation id="EA314D8CFFDCFFDFF03EF8E7FD4E5B6A" box="[601,688,1804,1831]" captionStart="TABLE 9" captionStartId="22.[152,244,404,428]" captionTargetPageId="22" captionText="TABLE 9. Main quantitative characters of IIBr from the proximal arm series (except keystone plates) of Holopus fabianii sp. nov. belonging to the type-series. Abbreviations: see Fig. 1C. Wp used as growth index. Values in mm except ratios. Paratypes 20–23: MGP-PD 33236; Paratypes 25–39: MGP-PD 33238." httpUri="http://table.plazi.org/id/F3CC28BFFFDBFFD8F2FFFE7FFC795DB8" pageId="17" pageNumber="418" tableUuid="F3CC28BFFFDBFFD8F2FFFE7FFC795DB8">Table 9</tableCitation>
|
||
). Height asymmetry mostly due to oblique articulation with IBrax, 12 cases with H/h<1.3, 4 cases with 1.5<H/h<1.7 (
|
||
<figureCitation id="3F8864B2FFDCFFDFF0BDF8DBFCC35B06" box="[730,829,1840,1867]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="17" pageNumber="418">Fig. 11E</figureCitation>
|
||
) and 3 cases with 1.95<H/h<2.05 (
|
||
<figureCitation id="3F8864B2FFDCFFDFF6A8F8DBFAC95B06" box="[1231,1335,1840,1867]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="17" pageNumber="418">Fig. 11D</figureCitation>
|
||
). Lateral faces with 4 to 6 parallel symplexial crenulations, sometimes restricted to the highest one. Muscular synarthries with fulcral ridge shorter than maximum brachial width (W).
|
||
<specimenCount id="B1B5B3BEFFDCFFDFF173F892FC775BDE" box="[788,905,1913,1939]" count="2" pageId="17" pageNumber="418" type="generic">Two types</specimenCount>
|
||
of muscular synarthries: one symmetrical with equal fulcral ridge segments parallel to W (
|
||
<figureCitation id="3F8864B2FFDCFFDFF0EFF877FD0E5BFA" box="[648,752,1948,1975]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="17" pageNumber="418">Fig. 11A</figureCitation>
|
||
), another asymmetrical with unequal fulcral ridge segments oblique to W (
|
||
<figureCitation id="3F8864B2FFDCFFDFF35CF82BFE5F5B96" box="[315,417,1984,2011]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="17" pageNumber="418">Fig. 11B</figureCitation>
|
||
); most IIBr with one synarthry asymmetrical and the other symmetrical (IIBr1 or IIBr2?), other IIBr with two articular facets asymmetrical (IIBr1) or symmetrical (IIBr2 or 3?). Four IIBr nearly as high as wide and asymmetrical, assigned to IIBr1 of a bivium (
|
||
<figureCitation id="3F8864B2FFDFFFDCF091FF50FCA65C98" box="[758,856,187,213]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11E</figureCitation>
|
||
). Articulations with adoral border forming an obtuse angle, ambulacral groove shallow, giving the whole ossicle a relatively flat V-shaped section.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FEE8FE5E5DC4" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
Keystone plates of type series (
|
||
<typeStatus id="7808C695FFDFFFDCF05FFEE8FD5B5D50" box="[568,677,259,285]" pageId="18" pageNumber="419" type="paratype">paratypes</typeStatus>
|
||
24 and 40–42,
|
||
<tableCitation id="EA314D8CFFDFFFDCF105FEE8FC395D50" box="[866,967,259,285]" captionStart="TABLE 10" captionStartId="22.[152,244,1573,1597]" captionTargetPageId="22" captionText="TABLE 10. Main quantitative characters of distal IIBr and keystone plates of Holopus fabianii sp. nov. belonging to the type-series.Abbreviations: see Fig. 1C. H for keystone plates: height of triangular external face. Wp used as growth index. Values in mm except ratios. Paratypes 24: MGP-PD 33236; Paratypes 40–45: MGP-PD 33238." httpUri="http://table.plazi.org/id/F3CC28BFFFDBFFD8F2FFF9CEFB735AC8" pageId="18" pageNumber="419" tableUuid="F3CC28BFFFDBFFD8F2FFF9CEFB735AC8">Table 10</tableCitation>
|
||
) with distal muscular synarthry roughly triangular and rounded (
|
||
<figureCitation id="3F8864B2FFDFFFDCF3CCFECCFDEE5D0C" box="[427,528,295,321]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11F</figureCitation>
|
||
); proximal synarthry symmetrical, with fulcral ridge as long as Wp (
|
||
<figureCitation id="3F8864B2FFDFFFDCF741FECCFA715D0C" box="[1318,1423,295,321]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11H</figureCitation>
|
||
); external face smooth, shaped as an isosceles triangle (
|
||
<figureCitation id="3F8864B2FFDFFFDCF093FEA0FCA45D28" box="[756,858,331,357]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11G</figureCitation>
|
||
), four conspicuous parallel symplexial crenulations on each side (
|
||
<figureCitation id="3F8864B2FFDFFFDCF356FE84FE6A5DC4" box="[305,404,367,393]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FE78FCEF5DB8" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
A few brachials from the distal IIBr series (
|
||
<typeStatus id="7808C695FFDFFFDCF0CDFE78FCE95DE0" box="[682,791,403,429]" pageId="18" pageNumber="419" type="paratype">paratypes</typeStatus>
|
||
43–45,
|
||
<tableCitation id="EA314D8CFFDFFFDCF116FE78FC2C5DE0" box="[881,978,403,429]" captionStart="TABLE 10" captionStartId="22.[152,244,1573,1597]" captionTargetPageId="22" captionText="TABLE 10. Main quantitative characters of distal IIBr and keystone plates of Holopus fabianii sp. nov. belonging to the type-series.Abbreviations: see Fig. 1C. H for keystone plates: height of triangular external face. Wp used as growth index. Values in mm except ratios. Paratypes 24: MGP-PD 33236; Paratypes 40–45: MGP-PD 33238." httpUri="http://table.plazi.org/id/F3CC28BFFFDBFFD8F2FFF9CEFB735AC8" pageId="18" pageNumber="419" tableUuid="F3CC28BFFFDBFFD8F2FFF9CEFB735AC8">Table 10</tableCitation>
|
||
) higher than broad, muscular synarthries similar to the distal articulation of the keystone plates with inconspicuous fulcral ridge, marked aboral ligament pit and axial nervous canal, conspicuous adoral muscle pits.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDFFFDCF2A0FE15FA955F74" pageId="18" pageNumber="419" type="materials_examined">
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FE15FA955F74" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
<emphasis id="95C7A425FFDFFFDCF2A0FE15FBFE5E55" bold="true" box="[199,1024,510,537]" pageId="18" pageNumber="419">
|
||
Complementary data from material not included into the
|
||
<typeStatus id="7808C695FFDFFFDCF11BFE14FC515E54" box="[892,943,511,537]" pageId="18" pageNumber="419">type</typeStatus>
|
||
series.
|
||
</emphasis>
|
||
The additional material allows us to investigate in more detail the range of variation of quantitative characters (Supplementary
|
||
<tableCitation id="EA314D8CFFDFFFDCF6EEFDC8FAFC5E70" box="[1161,1282,547,573]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="1.[152,244,1155,1179]" captionStartId-1="2.[152,244,151,175]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="TABLE 1. Material of extant taxa examined." captionText-1="TABLE 2. Cenozoic Holopodidae examined. MGP-PD: Museum of Nature and Humankind of the University of Padua; MNHN: Muséum national d’Histoire naturelle, Paris. Spilecco Hill: near Bolca, lower Ypresian; Chiampo: Cengio dell’Orbo quarry, Chiampo Valley, lower Lutetian; Superga Hill: near Turin, Miocene (Serravallian); Man21: Manni & Pacioni, 2021." httpUri-0="http://table.plazi.org/id/F3CC28BFFFCCFFCFF2FFFB68FD8058D6" httpUri-1="http://table.plazi.org/id/F3CC28BFFFCFFFCCF2FFFF7CFECF5D51" pageId="18" pageNumber="419" tableUuid-0="F3CC28BFFFCCFFCFF2FFFB68FD8058D6" tableUuid-1="F3CC28BFFFCFFFCCF2FFFF7CFECF5D51">Tables 1–2</tableCitation>
|
||
) and ratios in the main
|
||
<typeStatus id="7808C695FFDFFFDCF299FDACFEC65E2C" box="[254,312,583,609]" pageId="18" pageNumber="419">types</typeStatus>
|
||
of brachials. All material examined including the types-series: 8 aboral cups, 43 IBrax, 46 IIBr of the proximal series (including 14 with Wp>
|
||
<quantity id="604BD5D2FFDFFFDCF039FD80FD485EC8" box="[606,694,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="419" unit="mm" value="2.8">2.8 mm</quantity>
|
||
) and 60 IIBr with Wp<
|
||
<quantity id="604BD5D2FFDFFFDCF1A2FD80FBE55EC8" box="[965,1051,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="419" unit="mm" value="2.8">2.8 mm</quantity>
|
||
(distal series plus a few keystone plates poorly preserved). Ratio of IBrax to aboral cups 5.4, very close to expected ratio of 5; ratio of IBrax to proximal IIBr 1.07, far from the expected ratio of 4 or 6 indicating a high IIBr deficit. Height asymmetry (H/h) of the proximal IIBr is independent of their size. In 50% of the proximal IIBr 1.1<H/h<1.3 (possibly IIBr2 or IIBr3), in 34.6% 1.4<H/ h<1.75 (possibly IIBr1 or IIBr2) and in 15.4% 1.75<H/h<2.3 (possibly IIBr1). Quantitative characters of IBrax attaining maximum values with H
|
||
<quantity id="604BD5D2FFDFFFDCF070FCF4FD825F74" box="[535,636,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="4.78" pageId="18" pageNumber="419" unit="mm" value="4.78">4.78 mm</quantity>
|
||
, Wd
|
||
<quantity id="604BD5D2FFDFFFDCF0D1FCF5FCE55F74" box="[694,795,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="18" pageNumber="419" unit="mm" value="5.3">5.30 mm</quantity>
|
||
, Wp
|
||
<quantity id="604BD5D2FFDFFFDCF133FCF5FC495F74" box="[852,951,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" pageId="18" pageNumber="419" unit="mm" value="4.15">4.15 mm</quantity>
|
||
and Wm
|
||
<quantity id="604BD5D2FFDFFFDCF644FCF5FB7B5F74" box="[1059,1157,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="18" pageNumber="419" unit="mm" value="3.25">3.25 mm</quantity>
|
||
(Appendix
|
||
<tableCitation id="EA314D8CFFDFFFDCF76EFCF4FAA45F74" box="[1289,1370,799,825]" captionStart="TABLE 1" captionStartId="1.[152,244,1155,1179]" captionTargetPageId="1" captionText="TABLE 1. Material of extant taxa examined." httpUri="http://table.plazi.org/id/F3CC28BFFFCCFFCFF2FFFB68FD8058D6" pageId="18" pageNumber="419" tableUuid="F3CC28BFFFCCFFCFF2FFFB68FD8058D6">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFA92BBCFFDFFFDCF2A0FCA8FB305B90" pageId="18" pageNumber="419" type="discussion">
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FCA8FA785878" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
<emphasis id="95C7A425FFDFFFDCF2A0FCA8FEC45F10" bold="true" box="[199,314,835,861]" pageId="18" pageNumber="419">Remarks.</emphasis>
|
||
The high number of isolated ossicles of
|
||
<taxonomicName id="60B303B4FFDFFFDCF160FCA8FC3D5F10" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[775,963,835,861]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF160FCA8FC3D5F10" box="[775,963,835,861]" italics="true" pageId="18" pageNumber="419">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF1ACFCA8FBDB5F10" bold="true" box="[971,1061,835,861]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF1ACFCA8FBDB5F10" box="[971,1061,835,861]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the
|
||
<typeStatus id="7808C695FFDFFFDCF6F2FCA8FB3B5F10" box="[1173,1221,835,861]" pageId="18" pageNumber="419">type</typeStatus>
|
||
locality provides a unique insight into intraspecific character variation in the genus
|
||
<taxonomicName id="60B303B4FFDFFFDCF114FC8CFC2F5FCC" box="[883,977,871,897]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="95C7A425FFDFFFDCF114FC8CFC2F5FCC" box="[883,977,871,897]" italics="true" pageId="18" pageNumber="419">Holopus</emphasis>
|
||
</taxonomicName>
|
||
. To date, no comparable data have been published for extant species. However, the absence of a crown preserved with connected brachials makes it impossible to establish the precise number of IIBr in the arm’s proximal series. The combination of H/h asymmetry, which appears to increase proximally in extant species, and the fact that 50% of the IIBr have a H/h close to 1 suggest that there might have been at least 2–3 IIBr preceding the keystone plate. Like in the extant species
|
||
<taxonomicName id="60B303B4FFDFFFDCF6C8FC1CFAF3585C" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[1199,1293,1015,1041]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF6C8FC1CFAF3585C" box="[1199,1293,1015,1041]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
, the position of the keystone plate could have varied depending on the arm and the developmental stage (
|
||
<bibRefCitation id="C32205C6FFDFFFDCF6E8FBF0FA875878" author="Bourseau, J. - P. & Ameziane-Cominardi, N. & Avocat, R. & Roux, M." box="[1167,1401,1051,1077]" pageId="18" pageNumber="419" pagination="229 - 334" refId="ref20453" refString="Bourseau, J. - P., Ameziane-Cominardi, N., Avocat, R. & Roux, M. (1991) Echinodermata: Les Crinoides pedoncules de Nouvelle-Caledonie. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM. Vol. 8. Memoires du Museum National d'Histoire Naturelle, 151, pp. 229 - 334." type="journal article" year="1991">
|
||
Bourseau
|
||
<emphasis id="95C7A425FFDFFFDCF767FBF0FACE5878" box="[1280,1328,1051,1077]" italics="true" pageId="18" pageNumber="419">et al</emphasis>
|
||
. 1991
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FBD4FD3558EC" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
The recrystallized sediment within the calycinal cavity masks possible inner interradial processes, whose presence and characters would have been useful to describe and compare to extant species. These structures might have been absent as in
|
||
<taxonomicName id="60B303B4FFDFFFDCF3FEFB6CFDC558EC" authorityName="Manni & Pacioni" authorityYear="2021" box="[409,571,1159,1185]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="spileccense">
|
||
<emphasis id="95C7A425FFDFFFDCF3FEFB6CFDC558EC" box="[409,571,1159,1185]" italics="true" pageId="18" pageNumber="419">H. spileccense</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FB40FBA2598D" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
The maximum size of the aboral cup and IBrax of
|
||
<taxonomicName id="60B303B4FFDFFFDCF17CFB40FC695888" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[795,919,1195,1221]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF17CFB40FC695888" box="[795,919,1195,1221]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF1C6FB40FC005888" bold="true" box="[929,1022,1195,1221]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF1C6FB40FC005888" box="[929,1022,1195,1221]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be estimated by comparing the greatest width of the facets corresponding to the same articulation joining two adjacent ossicles. The proximal articulation of the largest IIBr (
|
||
<typeStatus id="7808C695FFDFFFDCF392FB18FDA95940" box="[501,599,1267,1293]" pageId="18" pageNumber="419" type="paratype">paratype</typeStatus>
|
||
20) is 1.6 times wider (Wp
|
||
<quantity id="604BD5D2FFDFFFDCF1F4FB19FC075940" box="[915,1017,1266,1293]" metricMagnitude="-3" metricUnit="m" metricValue="5.06" pageId="18" pageNumber="419" unit="mm" value="5.06">5.06 mm</quantity>
|
||
) than the largest distal articulation of IBrax (Wm
|
||
<quantity id="604BD5D2FFDFFFDCF346FAFDFE74597C" box="[289,394,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="18" pageNumber="419" unit="mm" value="3.25">3.25 mm</quantity>
|
||
). The largest proximal articulation of IBrax is 2 times wider (Wp
|
||
<quantity id="604BD5D2FFDFFFDCF61EFAFDFB21597C" box="[1145,1247,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" pageId="18" pageNumber="419" unit="mm" value="4.15">4.15 mm</quantity>
|
||
) than the largest distal articulation of the available aboral cups (
|
||
<typeStatus id="7808C695FFDFFFDCF0C2FAD0FCF95918" box="[677,775,1339,1365]" pageId="18" pageNumber="419" type="paratype">paratype</typeStatus>
|
||
2, Wr
|
||
<quantity id="604BD5D2FFDFFFDCF133FAD0FC425918" box="[852,956,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="18" pageNumber="419" unit="mm" value="2.03">2.03 mm</quantity>
|
||
). Thus, the estimated size of the individual to which
|
||
<typeStatus id="7808C695FFDFFFDCF29AFAB4FEA35934" box="[253,349,1375,1401]" pageId="18" pageNumber="419" type="paratype">paratype</typeStatus>
|
||
20 brachial belonged was 3.2 times larger than
|
||
<typeStatus id="7808C695FFDFFFDCF10DFAB4FC345934" box="[874,970,1375,1401]" pageId="18" pageNumber="419" type="paratype">paratype</typeStatus>
|
||
2. Therefore, the aboral cup of
|
||
<taxonomicName id="60B303B4FFDFFFDCF743FAB4FA625934" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[1316,1436,1375,1401]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF743FAB4FA625934" box="[1316,1436,1375,1401]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF2F0FA68FF0E59D0" bold="true" box="[151,240,1411,1437]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF2F0FA68FF0E59D0" box="[151,240,1411,1437]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
could possibly be as large as H 21.0 mm and D
|
||
<quantity id="604BD5D2FFDFFFDCF162FA68FC9459D0" box="[773,874,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="18" pageNumber="419" unit="mm" value="14.8">14.8 mm</quantity>
|
||
, a size intermediate between the known maximum aboral cup sizes of
|
||
<taxonomicName id="60B303B4FFDFFFDCF308FA4CFE2E598C" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[367,464,1447,1473]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF308FA4CFE2E598C" box="[367,464,1447,1473]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(H
|
||
<quantity id="604BD5D2FFDFFFDCF39CFA4CFDB2598C" box="[507,588,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="419" unit="mm" value="13.0">13 mm</quantity>
|
||
, D
|
||
<quantity id="604BD5D2FFDFFFDCF014FA4CFD4B598C" box="[627,693,1447,1473]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="419" unit="mm" value="9.0">9 mm</quantity>
|
||
) and
|
||
<taxonomicName id="60B303B4FFDFFFDCF094FA4CFCA4598D" authorityName="Orbigny" authorityYear="1837" box="[755,858,1447,1472]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="rangii">
|
||
<emphasis id="95C7A425FFDFFFDCF094FA4CFCA4598D" box="[755,858,1447,1472]" italics="true" pageId="18" pageNumber="419">H. rangii</emphasis>
|
||
</taxonomicName>
|
||
(H
|
||
<quantity id="604BD5D2FFDFFFDCF1E1FA4CFC29598C" box="[902,983,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="419" unit="mm" value="30.0">30 mm</quantity>
|
||
, D
|
||
<quantity id="604BD5D2FFDFFFDCF19AFA4DFBB3598C" box="[1021,1101,1446,1473]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="419" unit="mm" value="25.0">25 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDFFFDCF2A0FA20FB305B90" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="419">
|
||
<taxonomicName id="60B303B4FFDFFFDCF2A0FA20FE7E59A8" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[199,384,1483,1509]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF2A0FA20FE7E59A8" box="[199,384,1483,1509]" italics="true" pageId="18" pageNumber="419">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF3E4FA20FE2759A8" bold="true" box="[387,473,1483,1509]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF3E4FA20FE2759A8" box="[387,473,1483,1509]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
displays closer affinities with
|
||
<taxonomicName id="60B303B4FFDFFFDCF17BFA20FC8659A8" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[796,888,1483,1509]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF17BFA20FC8659A8" box="[796,888,1483,1509]" italics="true" pageId="18" pageNumber="419">H.alidis</emphasis>
|
||
</taxonomicName>
|
||
than with
|
||
<taxonomicName id="60B303B4FFDFFFDCF184FA20FBB859A9" authorityName="Orbigny" authorityYear="1837" box="[995,1094,1483,1508]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="rangii">
|
||
<emphasis id="95C7A425FFDFFFDCF184FA20FBB859A9" box="[995,1094,1483,1508]" italics="true" pageId="18" pageNumber="419">H.rangii</emphasis>
|
||
</taxonomicName>
|
||
,as both possess a differentiated keystone plate beyond IIBr2 separating proximal and distal arm series (
|
||
<figureCitation id="3F8864B2FFDFFFDCF1DCFA04FBC05A44" box="[955,1086,1518,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">Figs. 5C–H</figureCitation>
|
||
and
|
||
<figureCitation id="3F8864B2FFDFFFDCF612FA04FB045A44" box="[1141,1274,1519,1545]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11F–H</figureCitation>
|
||
). Both species also have an aboral cup with rather marked radial ribs. Yet,
|
||
<taxonomicName id="60B303B4FFDFFFDCF142F9F8FC7A5A60" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[805,900,1555,1581]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF142F9F8FC7A5A60" box="[805,900,1555,1581]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
differs in having a distinctly granulated external cup and brachials, including the keystone plates, with a predominantly tuberculated ornamentation unlike the nearly smooth keystone plates found for
|
||
<taxonomicName id="60B303B4FFDFFFDCF07BF9B0FD695A38" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[540,663,1627,1653]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF07BF9B0FD695A38" box="[540,663,1627,1653]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF0F8F9B0FD055A38" bold="true" box="[671,763,1627,1653]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF0F8F9B0FD055A38" box="[671,763,1627,1653]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF16BF9B0FC9B5A38" box="[780,869,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">Fig. 5C</figureCitation>
|
||
,
|
||
<figureCitation id="3F8864B2FFDFFFDCF115F9B0FC325A38" box="[882,972,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">
|
||
Fig. 5
|
||
<bibRefCitation id="C32205C6FFDFFFDCF1D2F9B1FC325A38" author="Barbieri, G. & De Zanche, V. & Sedea, R." box="[949,972,1626,1653]" pageId="18" pageNumber="419" pagination="5 - 12" refId="ref19830" refString="Barbieri, G., De Zanche, V. & Sedea, R. (1991) Vulcanismo Paleogenico ed evoluzione del semigraben Alpone-Agno. Rendiconti della Societa Geologica Italiana, 14, 5 - 12." type="journal article" year="1991">G</bibRefCitation>
|
||
</figureCitation>
|
||
, Bourseau
|
||
<emphasis id="95C7A425FFDFFFDCF62CF9B0FB805A38" box="[1099,1150,1627,1653]" italics="true" pageId="18" pageNumber="419">et al</emphasis>
|
||
. 1991, pl. 12). However, judging from our observation on
|
||
<taxonomicName id="60B303B4FFDFFFDCF072F994FD665AD4" box="[533,664,1663,1689]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="95C7A425FFDFFFDCF072F994FD665AD4" box="[533,664,1663,1689]" italics="true" pageId="18" pageNumber="419">Cyathidium</emphasis>
|
||
</taxonomicName>
|
||
, the relevance of the latter character might be called into question. The IBrax ornamentation with Y- or V-shaped ridges common in
|
||
<taxonomicName id="60B303B4FFDFFFDCF1E3F948FC015AF0" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[900,1023,1699,1725]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF1E3F948FC015AF0" box="[900,1023,1699,1725]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF66FF948FB9D5AF0" bold="true" box="[1032,1123,1699,1725]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF66FF948FB9D5AF0" box="[1032,1123,1699,1725]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is restricted to juveniles of
|
||
<taxonomicName id="60B303B4FFDFFFDCF2F0F92CFF045AAC" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[151,250,1735,1761]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF2F0F92CFF045AAC" box="[151,250,1735,1761]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF36CF92CFE7E5AAC" box="[267,384,1735,1761]" captionStart="FIGURE 4" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-29@6.[151,1436,181,1770]" captionTargetPageId="6" captionText="FIGURE 4. Holopus alidis. A: lateral distal view of paratype 4 without its arms (MNHN-IE-2013-10012); B: lateral distal view of a juvenile specimen with closed crown (MNHN-IE-2014-2185), C: close-up of IBrax of B; D: lateral oblique view of rolled arm attached to IBrax (arrow heads: keystone plates (IIBr4) of the two arms of the same ray) (MNHN-IE-2013-10012), E: pinnule socket on IIBr1 of Fig. 5B (adoral part at bottom with two small muscular pits); p: interradial process, r: thin radial ridge. Scale bars: 0.5 mm in E, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241168" httpUri="https://zenodo.org/record/14241168/files/figure.png" pageId="18" pageNumber="419">Fig 4B, C</figureCitation>
|
||
) and tends to disappear under the tuberculate ornamentation in older specimens. Moreover, the adoral side of the fulcral ridge of the proximal-most brachials often conspicuously crenulated in
|
||
<taxonomicName id="60B303B4FFDFFFDCF699F900FAA15B48" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[1278,1375,1771,1797]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF699F900FAA15B48" box="[1278,1375,1771,1797]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF708F900FF255B64" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">Fig. 5A–B</figureCitation>
|
||
) is not found in more distal synarthries (
|
||
<figureCitation id="3F8864B2FFDFFFDCF0C1F8E4FD035B64" box="[678,765,1806,1833]" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">Fig. 5E</figureCitation>
|
||
). This character appears to be absent in
|
||
<taxonomicName id="60B303B4FFDFFFDCF6A6F8E4FAC55B64" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[1217,1339,1807,1833]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF6A6F8E4FAC55B64" box="[1217,1339,1807,1833]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF725F8E4FA625B64" bold="true" box="[1346,1436,1807,1833]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF725F8E4FA625B64" box="[1346,1436,1807,1833]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF2F8F8D8FED75B00" box="[159,297,1843,1869]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11A–B</figureCitation>
|
||
). Furthermore, the IIBr fulcral ridges in
|
||
<taxonomicName id="60B303B4FFDFFFDCF091F8D8FCA95B00" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[758,855,1843,1869]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF091F8D8FCA95B00" box="[758,855,1843,1869]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF10FF8D8FC415B00" box="[872,959,1842,1869]" captionStart="FIGURE 5" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-19@7.[151,1436,181,1768]" captionTargetPageId="7" captionText="FIGURE 5. Holopus alidis (MNHN-IE-2013-10012). A: distal view of IBrax; B: adoral distal view of IIBr1; C–H: keystone plate (IIBr4) ending proximal brachial series (e.g., see arrow head in Fig. 4D), C: outer proximal side view (proximal muscular synarthial articulation to IIBr3 at left), D: close-up of lateral crenulation, E: proximal muscular synarthry, F: proximal adoral view, G: oblique distal lateral view, H: distal muscular synarthry (articulation to distal brachial series) at opposite side of external surface; arrow: adoral process, arrow heads: location of external triangular surface shown in C; d: distal tip of the keystone plate; *: pinnule insertion. Scale bars: 0.5 mm in D, 1.0 mm in all others." figureDoi="http://doi.org/10.5281/zenodo.14241170" httpUri="https://zenodo.org/record/14241170/files/figure.png" pageId="18" pageNumber="419">Fig. 5B</figureCitation>
|
||
) and
|
||
<taxonomicName id="60B303B4FFDFFFDCF667F8D8FB965B01" authorityName="Orbigny" authorityYear="1837" box="[1024,1128,1843,1868]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="rangii">
|
||
<emphasis id="95C7A425FFDFFFDCF667F8D8FB965B01" box="[1024,1128,1843,1868]" italics="true" pageId="18" pageNumber="419">H. rangii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF61EF8D8FB0F5B00" box="[1145,1265,1843,1869]" captionStart="FIGURE 6" captionStartId="10.[151,250,1884,1908]" captionTargetBox="[151,1436,273,1861]" captionTargetId="figure-77@10.[151,1436,273,1861]" captionTargetPageId="10" captionText="FIGURE 6. Holopus rangii (MNHN-IE-2013-11946). A–C: IBrax, A: oblique view of distal muscular synarthries, B: adoral view, C: close-up of fulcral ridge and outer ligament area (la) of proximal muscular synarthry, D–E: IIBr1, D: oblique distal view, E: close-up of pinnule socket, F: oblique aboral view of IIBr2 with proximal synarthrial facet; arrow: adoral interradial process, arrow head: pinnule socket. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241172" httpUri="https://zenodo.org/record/14241172/files/figure.png" pageId="18" pageNumber="419">Fig. 6D, F</figureCitation>
|
||
) are as long as the brachial is wide, which contrasts significantly with what is observed in
|
||
<taxonomicName id="60B303B4FFDFFFDCF182F8BCFBA05B3C" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[997,1118,1879,1905]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF182F8BCFBA05B3C" box="[997,1118,1879,1905]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF602F8BCFB415B3C" bold="true" box="[1125,1215,1879,1905]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF602F8BCFB415B3C" box="[1125,1215,1879,1905]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="3F8864B2FFDFFFDCF6A8F8BCFAA65B3C" box="[1231,1368,1879,1905]" captionStart="FIGURE 11" captionStartId="20.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure-29@20.[151,1436,181,1769]" captionTargetPageId="20" captionText="FIGURE 11. Secundibrachials of Holopus fabianii sp. nov. from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="18" pageNumber="419">Fig. 11A–C</figureCitation>
|
||
). This appears to be due to a lateral aboral displacement of the adoral surfaces of the synarthries, “opening” the articulation and thus giving the brachial a rather flat V-shape. The brachials of
|
||
<taxonomicName id="60B303B4FFDFFFDCF119F874FC205BF4" authorityName="Bourseau, Ameziane-Cominardi, Avocat & Roux" authorityYear="1991" box="[894,990,1951,1977]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="alidis">
|
||
<emphasis id="95C7A425FFDFFFDCF119F874FC205BF4" box="[894,990,1951,1977]" italics="true" pageId="18" pageNumber="419">H. alidis</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="60B303B4FFDFFFDCF67AF874FB7D5BF5" authorityName="Orbigny" authorityYear="1837" box="[1053,1155,1951,1976]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="rangii">
|
||
<emphasis id="95C7A425FFDFFFDCF67AF874FB7D5BF5" box="[1053,1155,1951,1976]" italics="true" pageId="18" pageNumber="419">H. rangii</emphasis>
|
||
</taxonomicName>
|
||
) are thus far thicker than those of
|
||
<taxonomicName id="60B303B4FFDFFFDCF29FF828FE8F5B90" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[248,369,1987,2013]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="18" pageNumber="419" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDFFFDCF29FF828FE8F5B90" box="[248,369,1987,2013]" italics="true" pageId="18" pageNumber="419">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDFFFDCF31FF828FE2F5B90" bold="true" box="[376,465,1987,2013]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel id="8EF4195EFFDFFFDCF31FF828FE2F5B90" box="[376,465,1987,2013]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and, in IBrax, exhibit a more pronounced distal interradial process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F3CC28BFFFDEFFDDF2F0F8EAFA8E5B2C" ID-DOI="http://doi.org/10.5281/zenodo.14241180" ID-Zenodo-Dep="14241180" httpUri="https://zenodo.org/record/14241180/files/figure.png" pageId="19" pageNumber="420" startId="19.[151,250,1793,1817]" targetBox="[151,1436,181,1769]" targetPageId="19" targetType="figure">
|
||
<paragraph id="A70C7837FFDEFFDDF2F0F8EAFA8E5B2C" blockId="19.[151,1436,1793,1890]" pageId="19" pageNumber="420">
|
||
<emphasis id="95C7A425FFDEFFDDF2F0F8EAFEDE5B54" bold="true" box="[151,288,1793,1817]" pageId="19" pageNumber="420">FIGURE 10.</emphasis>
|
||
Variation in primibrachial ornamentation in
|
||
<taxonomicName id="60B303B4FFDEFFDDF081F8EAFC6C5B54" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[742,914,1793,1817]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="19" pageNumber="420" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDEFFDDF081F8EAFC6C5B54" box="[742,914,1793,1817]" italics="true" pageId="19" pageNumber="420">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDEFFDDF1FFF8EAFC175B54" bold="true" box="[920,1001,1793,1817]" pageId="19" pageNumber="420">
|
||
<taxonomicNameLabel id="8EF4195EFFDEFFDDF1FFF8EAFC175B54" box="[920,1001,1793,1817]" pageId="19" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Chiampo (MGP-PD 33235). A–G, J: small specimens, A: paratype 8, B: paratype 9, C: paratype 10, D: paratype 11, E: paratype 12, F: paratype 13, G: paratype 14, J: paratype 15, H, I, K, L: large specimens, H: paratype 16, I: paratype 17, K: paratype 18, L: paratype 19. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F3CC28BFFFD9FFDAF2F0F8EAFD1A5B80" ID-DOI="http://doi.org/10.5281/zenodo.14241182" ID-Zenodo-Dep="14241182" httpUri="https://zenodo.org/record/14241182/files/figure.png" pageId="20" pageNumber="421" startId="20.[151,250,1793,1817]" targetBox="[151,1436,181,1769]" targetPageId="20" targetType="figure">
|
||
<paragraph id="A70C7837FFD9FFDAF2F0F8EAFD1A5B80" blockId="20.[151,1437,1793,1997]" pageId="20" pageNumber="421">
|
||
<emphasis id="95C7A425FFD9FFDAF2F0F8EAFEDF5B54" bold="true" box="[151,289,1793,1817]" pageId="20" pageNumber="421">FIGURE 11.</emphasis>
|
||
Secundibrachials of
|
||
<taxonomicName id="60B303B4FFD9FFDAF39BF8EAFD575B54" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[508,681,1793,1817]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="20" pageNumber="421" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD9FFDAF39BF8EAFD575B54" box="[508,681,1793,1817]" italics="true" pageId="20" pageNumber="421">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD9FFDAF0D5F8EAFCFA5B54" bold="true" box="[690,772,1793,1817]" pageId="20" pageNumber="421">
|
||
<taxonomicNameLabel id="8EF4195EFFD9FFDAF0D5F8EAFCFA5B54" box="[690,772,1793,1817]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Chiampo belonging to proximal series (MGP-PD 33236); A–E: IIBr from proximal part of proximal series, A: symmetrical muscular synarthry from a small specimen (paratype 20), B: asymmetrical muscular synarthry (paratype 21), C–D: proximal-most IIBr of a large specimen (paratype 22), C: oblique distal view, D: external face, E: IIBr of a bivium (paratype 23), F–H: keystone plate, distalmost IIBr of proximal series (paratype 24), F: oblique view of distal muscular synarthry, G: oblique view showing triangular external face and proximal muscular synarthry at left, H: proximal muscular synarthry. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EFA92BBCFFD8FFD8F2A0FF7CFBC25D28" lastPageId="22" lastPageNumber="423" pageId="21" pageNumber="422" type="description">
|
||
<paragraph id="A70C7837FFD8FFDBF2A0FF7CFE145DB8" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="422">
|
||
<taxonomicName id="60B303B4FFD8FFDBF2A0FF7CFE7E5CFC" authorityName="Roux, Martinez & Vizcaino" authorityYear="2021" box="[199,384,151,177]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="plaziati">
|
||
<emphasis id="95C7A425FFD8FFDBF2A0FF7CFE7E5CFC" box="[199,384,151,177]" italics="true" pageId="21" pageNumber="422">Holopus plaziati</emphasis>
|
||
</taxonomicName>
|
||
, known exclusively from isolated brachials (7 IBrax + 1 proximal IIBr), comes from the lower Eocene of the Corbières in southern
|
||
<collectingCountry id="DFA438A7FFD8FFDBF050FF50FD7D5C98" box="[567,643,187,213]" name="France" pageId="21" pageNumber="422">France</collectingCountry>
|
||
(
|
||
<bibRefCitation id="C32205C6FFD8FFDBF0F4FF50FCA95C98" author="Roux, M. & Martinez, A. & Vizcaino, D." box="[659,855,187,213]" pageId="21" pageNumber="422" pagination="201 - 242" refId="ref23311" refString="Roux, M., Martinez, A. & Vizcaino, D. (2021) A diverse crinoid fauna (Echinodermata, Crinoidea) from the Lower Eocene of the Gulf of Languedoc (Corbieres, Aude, southern France). Zootaxa, 4963 (2), 201 - 242. https: // doi. org / 10.11646 / zootaxa. 4963.2.1" type="journal article" year="2021">
|
||
Roux
|
||
<emphasis id="95C7A425FFD8FFDBF0BDFF50FCF25C98" box="[730,780,187,213]" italics="true" pageId="21" pageNumber="422">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
). Its IBrax differ from those of
|
||
<taxonomicName id="60B303B4FFD8FFDBF6D8FF50FAC75C98" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[1215,1337,187,213]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF6D8FF50FAC75C98" box="[1215,1337,187,213]" italics="true" pageId="21" pageNumber="422">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF725FF50FA625C98" bold="true" box="[1346,1436,187,213]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF725FF50FA625C98" box="[1346,1436,187,213]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="60B303B4FFD8FFDBF2A1FF34FE995CB4" authorityName="Manni & Pacioni" authorityYear="2021" box="[198,359,223,249]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="spileccense">
|
||
<emphasis id="95C7A425FFD8FFDBF2A1FF34FE995CB4" box="[198,359,223,249]" italics="true" pageId="21" pageNumber="422">H. spileccense</emphasis>
|
||
</taxonomicName>
|
||
in being generally smaller with a smaller H/Wd (
|
||
<figureCitation id="3F8864B2FFD8FFDBF1EEFF34FC325CB4" box="[905,972,223,249]" captionStart="FIGURE 7" captionStartId="11.[151,250,810,834]" captionTargetBox="[210,1376,188,783]" captionTargetId="figure-19@11.[204,1383,181,785]" captionTargetPageId="11" captionText="FIGURE 7. Main quantitative characters of aboral cup and IBrax in fossil Holopodidae. S: Cyathidium holopus from Seymour Island. Straight lines: values of H/D and H/Wd. Other values in mm." figureDoi="http://doi.org/10.5281/zenodo.14241174" httpUri="https://zenodo.org/record/14241174/files/figure.png" pageId="21" pageNumber="422">Fig. 7</figureCitation>
|
||
), and in having muscular synarthries with usually conspicuous crenulation on the adoral side of the fulcral ridge and sometimes on the adoral border of the muscular area. While
|
||
<typeStatus id="7808C695FFD8FFDBF3ECFECCFE1C5D0C" box="[395,482,295,321]" pageId="21" pageNumber="422" type="syntype">syntype</typeStatus>
|
||
2 of
|
||
<taxonomicName id="60B303B4FFD8FFDBF07CFECCFD6E5D0C" authorityName="Roux, Martinez & Vizcaino" authorityYear="2021" box="[539,656,295,321]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="plaziati">
|
||
<emphasis id="95C7A425FFD8FFDBF07CFECCFD6E5D0C" box="[539,656,295,321]" italics="true" pageId="21" pageNumber="422">H. plaziati</emphasis>
|
||
</taxonomicName>
|
||
appears similar to the IBrax of
|
||
<taxonomicName id="60B303B4FFD8FFDBF197FECCFB975D0C" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[1008,1129,295,321]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF197FECCFB975D0C" box="[1008,1129,295,321]" italics="true" pageId="21" pageNumber="422">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF617FECCFB375D0C" bold="true" box="[1136,1225,295,321]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF617FECCFB375D0C" box="[1136,1225,295,321]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, IBrax of
|
||
<typeStatus id="7808C695FFD8FFDBF75DFECCFA625D0C" box="[1338,1436,295,321]" pageId="21" pageNumber="422" type="syntype">syntypes</typeStatus>
|
||
3 and 7 have adoral articulation facets larger than in
|
||
<taxonomicName id="60B303B4FFD8FFDBF082FEA0FCA05D28" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[741,862,331,357]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF082FEA0FCA05D28" box="[741,862,331,357]" italics="true" pageId="21" pageNumber="422">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF101FEA0FC415D28" bold="true" box="[870,959,331,357]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF101FEA0FC415D28" box="[870,959,331,357]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
In
|
||
<taxonomicName id="60B303B4FFD8FFDBF182FEA0FBA55D28" authorityName="Roux, Martinez & Vizcaino" authorityYear="2021" box="[997,1115,331,357]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="plaziati">
|
||
<emphasis id="95C7A425FFD8FFDBF182FEA0FBA55D28" box="[997,1115,331,357]" italics="true" pageId="21" pageNumber="422">H. plaziati</emphasis>
|
||
</taxonomicName>
|
||
this increases the size of the distal interradial process and the general thickness of the brachial, resulting in an ambulacral groove that is more deeply carved into the ossicle. The conspicuous lateral symplexial crenulations in
|
||
<taxonomicName id="60B303B4FFD8FFDBF652FE78FB525DE0" authorityName="Roux, Martinez & Vizcaino" authorityYear="2021" box="[1077,1196,403,429]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="plaziati">
|
||
<emphasis id="95C7A425FFD8FFDBF652FE78FB525DE0" box="[1077,1196,403,429]" italics="true" pageId="21" pageNumber="422">H. plaziati</emphasis>
|
||
</taxonomicName>
|
||
are variable (parallel to undulated or branched). Its aboral surface displays irregular tuberculate ornamentation without formation of interradial or Y-shaped ridges.
|
||
</paragraph>
|
||
<caption id="F3CC28BFFFD8FFDBF2FFFDF4FC835E32" ID-Table-UUID="F3CC28BFFFD8FFDBF2FFFDF4FC835E32" httpUri="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFDF4FC835E32" pageId="21" pageNumber="422" startId="21.[152,244,543,567]" targetBox="[163,1398,655,1079]" targetIsTable="true" targetPageId="21" targetType="table">
|
||
<paragraph id="A70C7837FFD8FFDBF2FFFDF4FC835E32" blockId="21.[151,1437,543,640]" pageId="21" pageNumber="422">
|
||
<emphasis id="95C7A425FFD8FFDBF2FFFDF4FEF05E7A" bold="true" box="[152,270,543,567]" pageId="21" pageNumber="422">TABLE 7.</emphasis>
|
||
Quantitative characters of the eight aboral cups in the type-series of
|
||
<taxonomicName id="60B303B4FFD8FFDBF18DFDF4FB605E7A" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[1002,1182,543,567]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF18DFDF4FB605E7A" box="[1002,1182,543,567]" italics="true" pageId="21" pageNumber="422">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF6C3FDF4FB075E7A" bold="true" box="[1188,1273,543,567]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF6C3FDF4FB075E7A" box="[1188,1273,543,567]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations: see Fig. 1A. *: estimated value (broken ossicle). D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33232; Paratype 1: MGP-PD 33233; Paratypes 2–7: MGP-PD 33234.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A70C7837FFD8FFDBF2C4FD64FA9C587A" pageId="21" pageNumber="422">
|
||
<table id="D5B38A97FFD80031F2C4FD64FA88587A" box="[163,1398,655,1079]" gridcols="12" gridrows="9" pageId="21" pageNumber="422">
|
||
<tr id="19837A75FFD80031F2C4FD64FA885EEA" box="[163,1398,655,679]" gridrow="0" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FD64FEF05EEA" box="[163,270,655,679]" gridcol="0" gridrow="0" pageId="21" pageNumber="422">Specimen</th>
|
||
<th id="5A521309FFD80031F331FD64FE835EEA" box="[342,381,655,679]" gridcol="1" gridrow="0" pageId="21" pageNumber="422">D</th>
|
||
<th id="5A521309FFD80031F3D5FD64FE275EEA" box="[434,473,655,679]" gridcol="2" gridrow="0" pageId="21" pageNumber="422">H</th>
|
||
<th id="5A521309FFD80031F068FD64FDC85EEA" box="[527,566,655,679]" gridcol="3" gridrow="0" pageId="21" pageNumber="422">h</th>
|
||
<th id="5A521309FFD80031F00DFD64FD745EEA" box="[618,650,655,679]" gridcol="4" gridrow="0" pageId="21" pageNumber="422">ht</th>
|
||
<th id="5A521309FFD80031F0AFFD64FCEB5EEA" box="[712,789,655,679]" gridcol="5" gridrow="0" pageId="21" pageNumber="422">Db</th>
|
||
<th id="5A521309FFD80031F12FFD64FC915EEA" box="[840,879,655,679]" gridcol="6" gridrow="0" pageId="21" pageNumber="422">Dcs</th>
|
||
<th id="5A521309FFD80031F1C3FD64FC3A5EEA" box="[932,964,655,679]" gridcol="7" gridrow="0" pageId="21" pageNumber="422">Wr</th>
|
||
<th id="5A521309FFD80031F668FD64FBB45EEA" box="[1039,1098,655,679]" gridcol="8" gridrow="0" pageId="21" pageNumber="422">H/D</th>
|
||
<th id="5A521309FFD80031F610FD64FB5D5EEA" box="[1143,1187,655,679]" gridcol="9" gridrow="0" pageId="21" pageNumber="422">H/h</th>
|
||
<th id="5A521309FFD80031F6B1FD64FAFD5EEA" box="[1238,1283,655,679]" gridcol="10" gridrow="0" pageId="21" pageNumber="422">H/ht</th>
|
||
<th id="5A521309FFD80031F751FD64FA885EEA" box="[1334,1398,655,679]" gridcol="11" gridrow="0" pageId="21" pageNumber="422">D/Dcs</th>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FD2AFA885E97" box="[163,1398,705,730]" gridrow="1" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FD2AFEF05E97" box="[163,270,705,730]" gridcol="0" gridrow="1" pageId="21" pageNumber="422">Paratype 7</th>
|
||
<td id="5A521309FFD80031F331FD2AFE835E97" box="[342,381,705,730]" gridcol="1" gridrow="1" pageId="21" pageNumber="422">1.8</td>
|
||
<td id="5A521309FFD80031F3D5FD2AFE275E97" box="[434,473,705,730]" gridcol="2" gridrow="1" pageId="21" pageNumber="422">1.7</td>
|
||
<td id="5A521309FFD80031F068FD2AFDC85E97" box="[527,566,705,730]" gridcol="3" gridrow="1" pageId="21" pageNumber="422">1.7</td>
|
||
<td id="5A521309FFD80031F00DFD2AFD745E97" box="[618,650,705,730]" gridcol="4" gridrow="1" pageId="21" pageNumber="422">0.8</td>
|
||
<td id="5A521309FFD80031F0AFFD2AFCEB5E97" box="[712,789,705,730]" gridcol="5" gridrow="1" pageId="21" pageNumber="422">1.2–1.7</td>
|
||
<td id="5A521309FFD80031F12FFD2AFC915E97" box="[840,879,705,730]" gridcol="6" gridrow="1" pageId="21" pageNumber="422">1.6</td>
|
||
<td id="5A521309FFD80031F1C3FD2AFC3A5E97" box="[932,964,705,730]" gridcol="7" gridrow="1" pageId="21" pageNumber="422">1.2</td>
|
||
<td id="5A521309FFD80031F668FD2AFBB45E97" box="[1039,1098,705,730]" gridcol="8" gridrow="1" pageId="21" pageNumber="422">0.98</td>
|
||
<td id="5A521309FFD80031F610FD2AFB5D5E97" box="[1143,1187,705,730]" gridcol="9" gridrow="1" pageId="21" pageNumber="422">1.00</td>
|
||
<td id="5A521309FFD80031F6B1FD2AFAFD5E97" box="[1238,1283,705,730]" gridcol="10" gridrow="1" pageId="21" pageNumber="422">2.07</td>
|
||
<td id="5A521309FFD80031F751FD2AFA885E97" box="[1334,1398,705,730]" gridcol="11" gridrow="1" pageId="21" pageNumber="422">1.13</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FD18FA885F41" box="[163,1398,755,780]" gridrow="2" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FD18FEF05F41" box="[163,270,755,780]" gridcol="0" gridrow="2" pageId="21" pageNumber="422">Paratype 1</th>
|
||
<td id="5A521309FFD80031F331FD18FE835F41" box="[342,381,755,780]" gridcol="1" gridrow="2" pageId="21" pageNumber="422">2.5</td>
|
||
<td id="5A521309FFD80031F3D5FD18FE275F41" box="[434,473,755,780]" gridcol="2" gridrow="2" pageId="21" pageNumber="422">3.0</td>
|
||
<td id="5A521309FFD80031F068FD18FDC85F41" box="[527,566,755,780]" gridcol="3" gridrow="2" pageId="21" pageNumber="422">2.8</td>
|
||
<td id="5A521309FFD80031F00DFD18FD745F41" box="[618,650,755,780]" gridcol="4" gridrow="2" pageId="21" pageNumber="422">1.6</td>
|
||
<td id="5A521309FFD80031F0AFFD18FCEB5F41" box="[712,789,755,780]" gridcol="5" gridrow="2" pageId="21" pageNumber="422">1.7–2.3</td>
|
||
<td id="5A521309FFD80031F12FFD18FC915F41" box="[840,879,755,780]" gridcol="6" gridrow="2" pageId="21" pageNumber="422">1.7</td>
|
||
<td id="5A521309FFD80031F1C3FD18FC3A5F41" box="[932,964,755,780]" gridcol="7" gridrow="2" pageId="21" pageNumber="422">1.4</td>
|
||
<td id="5A521309FFD80031F668FD18FBB45F41" box="[1039,1098,755,780]" gridcol="8" gridrow="2" pageId="21" pageNumber="422">1.21</td>
|
||
<td id="5A521309FFD80031F610FD18FB5D5F41" box="[1143,1187,755,780]" gridcol="9" gridrow="2" pageId="21" pageNumber="422">1.06</td>
|
||
<td id="5A521309FFD80031F6B1FD18FAFD5F41" box="[1238,1283,755,780]" gridcol="10" gridrow="2" pageId="21" pageNumber="422">1.83</td>
|
||
<td id="5A521309FFD80031F751FD18FA885F41" box="[1334,1398,755,780]" gridcol="11" gridrow="2" pageId="21" pageNumber="422">1.49</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FCCEFA885F70" box="[163,1398,805,829]" gridrow="3" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FCCEFEF05F70" box="[163,270,805,829]" gridcol="0" gridrow="3" pageId="21" pageNumber="422">Holotype</th>
|
||
<td id="5A521309FFD80031F331FCCEFE835F70" box="[342,381,805,829]" gridcol="1" gridrow="3" pageId="21" pageNumber="422">2.7</td>
|
||
<td id="5A521309FFD80031F3D5FCCEFE275F70" box="[434,473,805,829]" gridcol="2" gridrow="3" pageId="21" pageNumber="422">3.9</td>
|
||
<td id="5A521309FFD80031F068FCCEFDC85F70" box="[527,566,805,829]" gridcol="3" gridrow="3" pageId="21" pageNumber="422">3.7</td>
|
||
<td id="5A521309FFD80031F00DFCCEFD745F70" box="[618,650,805,829]" gridcol="4" gridrow="3" pageId="21" pageNumber="422">2.5</td>
|
||
<td id="5A521309FFD80031F0AFFCCEFCEB5F70" box="[712,789,805,829]" gridcol="5" gridrow="3" pageId="21" pageNumber="422">3.1</td>
|
||
<td id="5A521309FFD80031F12FFCCEFC915F70" box="[840,879,805,829]" gridcol="6" gridrow="3" pageId="21" pageNumber="422">1.8</td>
|
||
<td id="5A521309FFD80031F1C3FCCEFC3A5F70" box="[932,964,805,829]" gridcol="7" gridrow="3" pageId="21" pageNumber="422">1.7</td>
|
||
<td id="5A521309FFD80031F668FCCEFBB45F70" box="[1039,1098,805,829]" gridcol="8" gridrow="3" pageId="21" pageNumber="422">1.39</td>
|
||
<td id="5A521309FFD80031F610FCCEFB5D5F70" box="[1143,1187,805,829]" gridcol="9" gridrow="3" pageId="21" pageNumber="422">1.02</td>
|
||
<td id="5A521309FFD80031F6B1FCCEFAFD5F70" box="[1238,1283,805,829]" gridcol="10" gridrow="3" pageId="21" pageNumber="422">1.53</td>
|
||
<td id="5A521309FFD80031F751FCCEFA885F70" box="[1334,1398,805,829]" gridcol="11" gridrow="3" pageId="21" pageNumber="422">1.49</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FCBCFA885F22" box="[163,1398,855,879]" gridrow="4" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FCBCFEF05F22" box="[163,270,855,879]" gridcol="0" gridrow="4" pageId="21" pageNumber="422">Paratype 6</th>
|
||
<td id="5A521309FFD80031F331FCBCFE835F22" box="[342,381,855,879]" gridcol="1" gridrow="4" pageId="21" pageNumber="422">>3*</td>
|
||
<td id="5A521309FFD80031F3D5FCBCFE275F22" box="[434,473,855,879]" gridcol="2" gridrow="4" pageId="21" pageNumber="422">4.4</td>
|
||
<td id="5A521309FFD80031F068FCBCFDC85F22" box="[527,566,855,879]" gridcol="3" gridrow="4" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F00DFCBCFD745F22" box="[618,650,855,879]" gridcol="4" gridrow="4" pageId="21" pageNumber="422">2.1</td>
|
||
<td id="5A521309FFD80031F0AFFCBCFCEB5F22" box="[712,789,855,879]" gridcol="5" gridrow="4" pageId="21" pageNumber="422">3.2–4.1</td>
|
||
<td id="5A521309FFD80031F12FFCBCFC915F22" box="[840,879,855,879]" gridcol="6" gridrow="4" pageId="21" pageNumber="422">3.1</td>
|
||
<td id="5A521309FFD80031F1C3FCBCFC3A5F22" box="[932,964,855,879]" gridcol="7" gridrow="4" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F668FCBCFBB45F22" box="[1039,1098,855,879]" gridcol="8" gridrow="4" pageId="21" pageNumber="422"><1.4*</td>
|
||
<td id="5A521309FFD80031F610FCBCFB5D5F22" box="[1143,1187,855,879]" gridcol="9" gridrow="4" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F6B1FCBCFAFD5F22" box="[1238,1283,855,879]" gridcol="10" gridrow="4" pageId="21" pageNumber="422">1.35</td>
|
||
<td id="5A521309FFD80031F751FCBCFA885F22" box="[1334,1398,855,879]" gridcol="11" gridrow="4" pageId="21" pageNumber="422">1.06</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FC62FA885FEC" box="[163,1398,905,929]" gridrow="5" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FC62FEF05FEC" box="[163,270,905,929]" gridcol="0" gridrow="5" pageId="21" pageNumber="422">Paratype 3</th>
|
||
<td id="5A521309FFD80031F331FC62FE835FEC" box="[342,381,905,929]" gridcol="1" gridrow="5" pageId="21" pageNumber="422">3.3</td>
|
||
<td id="5A521309FFD80031F3D5FC62FE275FEC" box="[434,473,905,929]" gridcol="2" gridrow="5" pageId="21" pageNumber="422">4.9</td>
|
||
<td id="5A521309FFD80031F068FC62FDC85FEC" box="[527,566,905,929]" gridcol="3" gridrow="5" pageId="21" pageNumber="422">3.8</td>
|
||
<td id="5A521309FFD80031F00DFC62FD745FEC" box="[618,650,905,929]" gridcol="4" gridrow="5" pageId="21" pageNumber="422">2.8</td>
|
||
<td id="5A521309FFD80031F0AFFC62FCEB5FEC" box="[712,789,905,929]" gridcol="5" gridrow="5" pageId="21" pageNumber="422">5.0</td>
|
||
<td id="5A521309FFD80031F12FFC62FC915FEC" box="[840,879,905,929]" gridcol="6" gridrow="5" pageId="21" pageNumber="422">3.2</td>
|
||
<td id="5A521309FFD80031F1C3FC62FC3A5FEC" box="[932,964,905,929]" gridcol="7" gridrow="5" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F668FC62FBB45FEC" box="[1039,1098,905,929]" gridcol="8" gridrow="5" pageId="21" pageNumber="422">1.48</td>
|
||
<td id="5A521309FFD80031F610FC62FB5D5FEC" box="[1143,1187,905,929]" gridcol="9" gridrow="5" pageId="21" pageNumber="422">1.29</td>
|
||
<td id="5A521309FFD80031F6B1FC62FAFD5FEC" box="[1238,1283,905,929]" gridcol="10" gridrow="5" pageId="21" pageNumber="422">1.74</td>
|
||
<td id="5A521309FFD80031F751FC62FA885FEC" box="[1334,1398,905,929]" gridcol="11" gridrow="5" pageId="21" pageNumber="422">1.03</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FC50FA885F9E" box="[163,1398,955,979]" gridrow="6" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FC50FEF05F9E" box="[163,270,955,979]" gridcol="0" gridrow="6" pageId="21" pageNumber="422">Paratype 5</th>
|
||
<td id="5A521309FFD80031F331FC50FE835F9E" box="[342,381,955,979]" gridcol="1" gridrow="6" pageId="21" pageNumber="422">>4*</td>
|
||
<td id="5A521309FFD80031F3D5FC50FE275F9E" box="[434,473,955,979]" gridcol="2" gridrow="6" pageId="21" pageNumber="422">5.1</td>
|
||
<td id="5A521309FFD80031F068FC50FDC85F9E" box="[527,566,955,979]" gridcol="3" gridrow="6" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F00DFC50FD745F9E" box="[618,650,955,979]" gridcol="4" gridrow="6" pageId="21" pageNumber="422">3.7</td>
|
||
<td id="5A521309FFD80031F0AFFC50FCEB5F9E" box="[712,789,955,979]" gridcol="5" gridrow="6" pageId="21" pageNumber="422">>2.7*</td>
|
||
<td id="5A521309FFD80031F12FFC50FC915F9E" box="[840,879,955,979]" gridcol="6" gridrow="6" pageId="21" pageNumber="422">2.4</td>
|
||
<td id="5A521309FFD80031F1C3FC50FC3A5F9E" box="[932,964,955,979]" gridcol="7" gridrow="6" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F668FC50FBB45F9E" box="[1039,1098,955,979]" gridcol="8" gridrow="6" pageId="21" pageNumber="422"><1.4*</td>
|
||
<td id="5A521309FFD80031F610FC50FB5D5F9E" box="[1143,1187,955,979]" gridcol="9" gridrow="6" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F6B1FC50FAFD5F9E" box="[1238,1283,955,979]" gridcol="10" gridrow="6" pageId="21" pageNumber="422">1.35</td>
|
||
<td id="5A521309FFD80031F751FC50FA885F9E" box="[1334,1398,955,979]" gridcol="11" gridrow="6" pageId="21" pageNumber="422">>1.5*</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FC07FA885848" box="[163,1398,1004,1029]" gridrow="7" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FC07FEF05848" box="[163,270,1004,1029]" gridcol="0" gridrow="7" pageId="21" pageNumber="422">Paratype 4</th>
|
||
<td id="5A521309FFD80031F331FC07FE835848" box="[342,381,1004,1029]" gridcol="1" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F3D5FC07FE275848" box="[434,473,1004,1029]" gridcol="2" gridrow="7" pageId="21" pageNumber="422">>5*</td>
|
||
<td id="5A521309FFD80031F068FC07FDC85848" box="[527,566,1004,1029]" gridcol="3" gridrow="7" pageId="21" pageNumber="422">>3*</td>
|
||
<td id="5A521309FFD80031F00DFC07FD745848" box="[618,650,1004,1029]" gridcol="4" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F0AFFC07FCEB5848" box="[712,789,1004,1029]" gridcol="5" gridrow="7" pageId="21" pageNumber="422">4.5–5.3</td>
|
||
<td id="5A521309FFD80031F12FFC07FC915848" box="[840,879,1004,1029]" gridcol="6" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F1C3FC07FC3A5848" box="[932,964,1004,1029]" gridcol="7" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F668FC07FBB45848" box="[1039,1098,1004,1029]" gridcol="8" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F610FC07FB5D5848" box="[1143,1187,1004,1029]" gridcol="9" gridrow="7" pageId="21" pageNumber="422">1.6*</td>
|
||
<td id="5A521309FFD80031F6B1FC07FAFD5848" box="[1238,1283,1004,1029]" gridcol="10" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F751FC07FA885848" box="[1334,1398,1004,1029]" gridcol="11" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FBF5FA88587A" box="[163,1398,1054,1079]" gridrow="8" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FBF5FEF0587A" box="[163,270,1054,1079]" gridcol="0" gridrow="8" pageId="21" pageNumber="422">Paratype 2</th>
|
||
<td id="5A521309FFD80031F331FBF5FE83587A" box="[342,381,1054,1079]" gridcol="1" gridrow="8" pageId="21" pageNumber="422">4.6</td>
|
||
<td id="5A521309FFD80031F3D5FBF5FE27587A" box="[434,473,1054,1079]" gridcol="2" gridrow="8" pageId="21" pageNumber="422">6.5</td>
|
||
<td id="5A521309FFD80031F068FBF5FDC8587A" box="[527,566,1054,1079]" gridcol="3" gridrow="8" pageId="21" pageNumber="422">6.5</td>
|
||
<td id="5A521309FFD80031F00DFBF5FD74587A" box="[618,650,1054,1079]" gridcol="4" gridrow="8" pageId="21" pageNumber="422">4.5</td>
|
||
<td id="5A521309FFD80031F0AFFBF5FCEB587A" box="[712,789,1054,1079]" gridcol="5" gridrow="8" pageId="21" pageNumber="422">7.3–8.1</td>
|
||
<td id="5A521309FFD80031F12FFBF5FC91587A" box="[840,879,1054,1079]" gridcol="6" gridrow="8" pageId="21" pageNumber="422">3.9</td>
|
||
<td id="5A521309FFD80031F1C3FBF5FC3A587A" box="[932,964,1054,1079]" gridcol="7" gridrow="8" pageId="21" pageNumber="422">2*</td>
|
||
<td id="5A521309FFD80031F668FBF5FBB4587A" box="[1039,1098,1054,1079]" gridcol="8" gridrow="8" pageId="21" pageNumber="422">1.42</td>
|
||
<td id="5A521309FFD80031F610FBF5FB5D587A" box="[1143,1187,1054,1079]" gridcol="9" gridrow="8" pageId="21" pageNumber="422">1.00</td>
|
||
<td id="5A521309FFD80031F6B1FBF5FAFD587A" box="[1238,1283,1054,1079]" gridcol="10" gridrow="8" pageId="21" pageNumber="422">1.44</td>
|
||
<td id="5A521309FFD80031F751FBF5FA88587A" box="[1334,1398,1054,1079]" gridcol="11" gridrow="8" pageId="21" pageNumber="422">1.17</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="F3CC28BFFFD8FFDBF2FFFB60FB30588A" ID-Table-UUID="F3CC28BFFFD8FFDBF2FFFB60FB30588A" httpUri="http://table.plazi.org/id/F3CC28BFFFD8FFDBF2FFFB60FB30588A" pageId="21" pageNumber="422" startId="21.[152,244,1163,1187]" targetBox="[163,1392,1239,1862]" targetIsTable="true" targetPageId="21" targetType="table">
|
||
<paragraph id="A70C7837FFD8FFDBF2FFFB60FB30588A" blockId="21.[151,1437,1163,1223]" pageId="21" pageNumber="422">
|
||
<emphasis id="95C7A425FFD8FFDBF2FFFB60FEEE58EE" bold="true" box="[152,272,1163,1187]" pageId="21" pageNumber="422">TABLE 8.</emphasis>
|
||
Quantitative characters of IBrax of
|
||
<taxonomicName id="60B303B4FFD8FFDBF0F5FB60FCB958EE" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[658,839,1163,1187]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF0F5FB60FCB958EE" box="[658,839,1163,1187]" italics="true" pageId="21" pageNumber="422">Holopus fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF12AFB60FC5D58EE" bold="true" box="[845,931,1163,1187]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF12AFB60FC5D58EE" box="[845,931,1163,1187]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belonging to figured paratypes. Abbreviations: see Fig. 1B. Wd used as growth index. Values in mm except ratios. Paratypes 8–19: MGP-PD 33235.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A70C7837FFD8FFDBF2C4FB3CFAB55B0B" pageId="21" pageNumber="422">
|
||
<table id="D5B38A97FFD80031F2C4FB3CFA8E5B0B" box="[163,1392,1239,1862]" gridcols="10" gridrows="13" pageId="21" pageNumber="422">
|
||
<tr id="19837A75FFD80031F2C4FB3CFA8E58A2" box="[163,1392,1239,1263]" gridrow="0" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FB3CFEE458A2" box="[163,282,1239,1263]" gridcol="0" gridrow="0" pageId="21" pageNumber="422">Specimen</th>
|
||
<th id="5A521309FFD80031F315FB3CFE4A58A2" box="[370,436,1239,1263]" gridcol="1" gridrow="0" pageId="21" pageNumber="422">Figure</th>
|
||
<th id="5A521309FFD80031F381FB3CFDF558A2" box="[486,523,1239,1263]" gridcol="2" gridrow="0" pageId="21" pageNumber="422">Wd</th>
|
||
<th id="5A521309FFD80031F035FB3CFD8C58A2" box="[594,626,1239,1263]" gridcol="3" gridrow="0" pageId="21" pageNumber="422">H</th>
|
||
<th id="5A521309FFD80031F0D9FB3CFD1D58A2" box="[702,739,1239,1263]" gridcol="4" gridrow="0" pageId="21" pageNumber="422">Wp</th>
|
||
<th id="5A521309FFD80031F155FB3CFCA058A2" box="[818,862,1239,1263]" gridcol="5" gridrow="0" pageId="21" pageNumber="422">Wm</th>
|
||
<th id="5A521309FFD80031F1C1FB3CFC1B58A2" box="[934,997,1239,1263]" gridcol="6" gridrow="0" pageId="21" pageNumber="422">H/Wd</th>
|
||
<th id="5A521309FFD80031F67CFB3CFBA758A2" box="[1051,1113,1239,1263]" gridcol="7" gridrow="0" pageId="21" pageNumber="422">H/Wp</th>
|
||
<th id="5A521309FFD80031F6E8FB3CFB1958A2" box="[1167,1255,1239,1263]" gridcol="8" gridrow="0" pageId="21" pageNumber="422">Wp/Wm</th>
|
||
<th id="5A521309FFD80031F778FB3CFA8E58A2" box="[1311,1392,1239,1263]" gridcol="9" gridrow="0" pageId="21" pageNumber="422">Wp/Wd</th>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FAE2FA8E596C" box="[163,1392,1289,1313]" gridrow="1" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FAE2FEE4596C" box="[163,282,1289,1313]" gridcol="0" gridrow="1" pageId="21" pageNumber="422">Paratype 8</th>
|
||
<td id="5A521309FFD80031F315FAE2FE4A596C" box="[370,436,1289,1313]" gridcol="1" gridrow="1" pageId="21" pageNumber="422">10A</td>
|
||
<td id="5A521309FFD80031F381FAE2FDF5596C" box="[486,523,1289,1313]" gridcol="2" gridrow="1" pageId="21" pageNumber="422">1.8</td>
|
||
<td id="5A521309FFD80031F035FAE2FD8C596C" box="[594,626,1289,1313]" gridcol="3" gridrow="1" pageId="21" pageNumber="422">1.8</td>
|
||
<td id="5A521309FFD80031F0D9FAE2FD1D596C" box="[702,739,1289,1313]" gridcol="4" gridrow="1" pageId="21" pageNumber="422">1.0</td>
|
||
<td id="5A521309FFD80031F155FAE2FCA0596C" box="[818,862,1289,1313]" gridcol="5" gridrow="1" pageId="21" pageNumber="422">1.1</td>
|
||
<td id="5A521309FFD80031F1C1FAE2FC1B596C" box="[934,997,1289,1313]" gridcol="6" gridrow="1" pageId="21" pageNumber="422">1.02</td>
|
||
<td id="5A521309FFD80031F67CFAE2FBA7596C" box="[1051,1113,1289,1313]" gridcol="7" gridrow="1" pageId="21" pageNumber="422">1.78</td>
|
||
<td id="5A521309FFD80031F6E8FAE2FB19596C" box="[1167,1255,1289,1313]" gridcol="8" gridrow="1" pageId="21" pageNumber="422">0.93</td>
|
||
<td id="5A521309FFD80031F778FAE2FA8E596C" box="[1311,1392,1289,1313]" gridcol="9" gridrow="1" pageId="21" pageNumber="422">0.57</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FAD0FA8E591E" box="[163,1392,1339,1363]" gridrow="2" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FAD0FEE4591E" box="[163,282,1339,1363]" gridcol="0" gridrow="2" pageId="21" pageNumber="422">Paratype 11</th>
|
||
<td id="5A521309FFD80031F315FAD0FE4A591E" box="[370,436,1339,1363]" gridcol="1" gridrow="2" pageId="21" pageNumber="422">10D</td>
|
||
<td id="5A521309FFD80031F381FAD0FDF5591E" box="[486,523,1339,1363]" gridcol="2" gridrow="2" pageId="21" pageNumber="422">1.9</td>
|
||
<td id="5A521309FFD80031F035FAD0FD8C591E" box="[594,626,1339,1363]" gridcol="3" gridrow="2" pageId="21" pageNumber="422">1.9</td>
|
||
<td id="5A521309FFD80031F0D9FAD0FD1D591E" box="[702,739,1339,1363]" gridcol="4" gridrow="2" pageId="21" pageNumber="422">1.0</td>
|
||
<td id="5A521309FFD80031F155FAD0FCA0591E" box="[818,862,1339,1363]" gridcol="5" gridrow="2" pageId="21" pageNumber="422">1.0</td>
|
||
<td id="5A521309FFD80031F1C1FAD0FC1B591E" box="[934,997,1339,1363]" gridcol="6" gridrow="2" pageId="21" pageNumber="422">1.03</td>
|
||
<td id="5A521309FFD80031F67CFAD0FBA7591E" box="[1051,1113,1339,1363]" gridcol="7" gridrow="2" pageId="21" pageNumber="422">1.90</td>
|
||
<td id="5A521309FFD80031F6E8FAD0FB19591E" box="[1167,1255,1339,1363]" gridcol="8" gridrow="2" pageId="21" pageNumber="422">0.97</td>
|
||
<td id="5A521309FFD80031F778FAD0FA8E591E" box="[1311,1392,1339,1363]" gridcol="9" gridrow="2" pageId="21" pageNumber="422">0.54</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FA86FA8E59C8" box="[163,1392,1389,1413]" gridrow="3" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FA86FEE459C8" box="[163,282,1389,1413]" gridcol="0" gridrow="3" pageId="21" pageNumber="422">Paratype 12</th>
|
||
<td id="5A521309FFD80031F315FA86FE4A59C8" box="[370,436,1389,1413]" gridcol="1" gridrow="3" pageId="21" pageNumber="422">10E</td>
|
||
<td id="5A521309FFD80031F381FA86FDF559C8" box="[486,523,1389,1413]" gridcol="2" gridrow="3" pageId="21" pageNumber="422">2.2</td>
|
||
<td id="5A521309FFD80031F035FA86FD8C59C8" box="[594,626,1389,1413]" gridcol="3" gridrow="3" pageId="21" pageNumber="422">2.2</td>
|
||
<td id="5A521309FFD80031F0D9FA86FD1D59C8" box="[702,739,1389,1413]" gridcol="4" gridrow="3" pageId="21" pageNumber="422">1.5</td>
|
||
<td id="5A521309FFD80031F155FA86FCA059C8" box="[818,862,1389,1413]" gridcol="5" gridrow="3" pageId="21" pageNumber="422">1.3</td>
|
||
<td id="5A521309FFD80031F1C1FA86FC1B59C8" box="[934,997,1389,1413]" gridcol="6" gridrow="3" pageId="21" pageNumber="422">1.01</td>
|
||
<td id="5A521309FFD80031F67CFA86FBA759C8" box="[1051,1113,1389,1413]" gridcol="7" gridrow="3" pageId="21" pageNumber="422">1.46</td>
|
||
<td id="5A521309FFD80031F6E8FA86FB1959C8" box="[1167,1255,1389,1413]" gridcol="8" gridrow="3" pageId="21" pageNumber="422">1.16</td>
|
||
<td id="5A521309FFD80031F778FA86FA8E59C8" box="[1311,1392,1389,1413]" gridcol="9" gridrow="3" pageId="21" pageNumber="422">0.69</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FA75FA8E59FA" box="[163,1392,1438,1463]" gridrow="4" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FA75FEE459FA" box="[163,282,1438,1463]" gridcol="0" gridrow="4" pageId="21" pageNumber="422">Paratype 9</th>
|
||
<td id="5A521309FFD80031F315FA75FE4A59FA" box="[370,436,1438,1463]" gridcol="1" gridrow="4" pageId="21" pageNumber="422">10B</td>
|
||
<td id="5A521309FFD80031F381FA75FDF559FA" box="[486,523,1438,1463]" gridcol="2" gridrow="4" pageId="21" pageNumber="422">2.2</td>
|
||
<td id="5A521309FFD80031F035FA75FD8C59FA" box="[594,626,1438,1463]" gridcol="3" gridrow="4" pageId="21" pageNumber="422">2.1</td>
|
||
<td id="5A521309FFD80031F0D9FA75FD1D59FA" box="[702,739,1438,1463]" gridcol="4" gridrow="4" pageId="21" pageNumber="422">1.1</td>
|
||
<td id="5A521309FFD80031F155FA75FCA059FA" box="[818,862,1438,1463]" gridcol="5" gridrow="4" pageId="21" pageNumber="422">1.3</td>
|
||
<td id="5A521309FFD80031F1C1FA75FC1B59FA" box="[934,997,1438,1463]" gridcol="6" gridrow="4" pageId="21" pageNumber="422">0.96</td>
|
||
<td id="5A521309FFD80031F67CFA75FBA759FA" box="[1051,1113,1438,1463]" gridcol="7" gridrow="4" pageId="21" pageNumber="422">1.96</td>
|
||
<td id="5A521309FFD80031F6E8FA75FB1959FA" box="[1167,1255,1438,1463]" gridcol="8" gridrow="4" pageId="21" pageNumber="422">0.85</td>
|
||
<td id="5A521309FFD80031F778FA75FA8E59FA" box="[1311,1392,1438,1463]" gridcol="9" gridrow="4" pageId="21" pageNumber="422">0.49</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4FA3BFA8E59A4" box="[163,1392,1488,1513]" gridrow="5" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4FA3BFEE459A4" box="[163,282,1488,1513]" gridcol="0" gridrow="5" pageId="21" pageNumber="422">Paratype 15</th>
|
||
<td id="5A521309FFD80031F315FA3BFE4A59A4" box="[370,436,1488,1513]" gridcol="1" gridrow="5" pageId="21" pageNumber="422">10J</td>
|
||
<td id="5A521309FFD80031F381FA3BFDF559A4" box="[486,523,1488,1513]" gridcol="2" gridrow="5" pageId="21" pageNumber="422">2.3</td>
|
||
<td id="5A521309FFD80031F035FA3BFD8C59A4" box="[594,626,1488,1513]" gridcol="3" gridrow="5" pageId="21" pageNumber="422">2.2</td>
|
||
<td id="5A521309FFD80031F0D9FA3BFD1D59A4" box="[702,739,1488,1513]" gridcol="4" gridrow="5" pageId="21" pageNumber="422">1.2</td>
|
||
<td id="5A521309FFD80031F155FA3BFCA059A4" box="[818,862,1488,1513]" gridcol="5" gridrow="5" pageId="21" pageNumber="422">1.2</td>
|
||
<td id="5A521309FFD80031F1C1FA3BFC1B59A4" box="[934,997,1488,1513]" gridcol="6" gridrow="5" pageId="21" pageNumber="422">0.95</td>
|
||
<td id="5A521309FFD80031F67CFA3BFBA759A4" box="[1051,1113,1488,1513]" gridcol="7" gridrow="5" pageId="21" pageNumber="422">1.82</td>
|
||
<td id="5A521309FFD80031F6E8FA3BFB1959A4" box="[1167,1255,1488,1513]" gridcol="8" gridrow="5" pageId="21" pageNumber="422">0.98</td>
|
||
<td id="5A521309FFD80031F778FA3BFA8E59A4" box="[1311,1392,1488,1513]" gridcol="9" gridrow="5" pageId="21" pageNumber="422">0.52</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F9E9FA8E5A56" box="[163,1392,1538,1563]" gridrow="6" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F9E9FEE45A56" box="[163,282,1538,1563]" gridcol="0" gridrow="6" pageId="21" pageNumber="422">Paratype 10</th>
|
||
<td id="5A521309FFD80031F315F9E9FE4A5A56" box="[370,436,1538,1563]" gridcol="1" gridrow="6" pageId="21" pageNumber="422">10C</td>
|
||
<td id="5A521309FFD80031F381F9E9FDF55A56" box="[486,523,1538,1563]" gridcol="2" gridrow="6" pageId="21" pageNumber="422">2.6</td>
|
||
<td id="5A521309FFD80031F035F9E9FD8C5A56" box="[594,626,1538,1563]" gridcol="3" gridrow="6" pageId="21" pageNumber="422">2.1</td>
|
||
<td id="5A521309FFD80031F0D9F9E9FD1D5A56" box="[702,739,1538,1563]" gridcol="4" gridrow="6" pageId="21" pageNumber="422">1.2</td>
|
||
<td id="5A521309FFD80031F155F9E9FCA05A56" box="[818,862,1538,1563]" gridcol="5" gridrow="6" pageId="21" pageNumber="422">1.3</td>
|
||
<td id="5A521309FFD80031F1C1F9E9FC1B5A56" box="[934,997,1538,1563]" gridcol="6" gridrow="6" pageId="21" pageNumber="422">0.82</td>
|
||
<td id="5A521309FFD80031F67CF9E9FBA75A56" box="[1051,1113,1538,1563]" gridcol="7" gridrow="6" pageId="21" pageNumber="422">1.76</td>
|
||
<td id="5A521309FFD80031F6E8F9E9FB195A56" box="[1167,1255,1538,1563]" gridcol="8" gridrow="6" pageId="21" pageNumber="422">0.96</td>
|
||
<td id="5A521309FFD80031F778F9E9FA8E5A56" box="[1311,1392,1538,1563]" gridcol="9" gridrow="6" pageId="21" pageNumber="422">0.46</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F9DFFA8E5A00" box="[163,1392,1588,1613]" gridrow="7" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F9DFFEE45A00" box="[163,282,1588,1613]" gridcol="0" gridrow="7" pageId="21" pageNumber="422">Paratype 14</th>
|
||
<td id="5A521309FFD80031F315F9DFFE4A5A00" box="[370,436,1588,1613]" gridcol="1" gridrow="7" pageId="21" pageNumber="422">10G</td>
|
||
<td id="5A521309FFD80031F381F9DFFDF55A00" box="[486,523,1588,1613]" gridcol="2" gridrow="7" pageId="21" pageNumber="422">2.7</td>
|
||
<td id="5A521309FFD80031F035F9DFFD8C5A00" box="[594,626,1588,1613]" gridcol="3" gridrow="7" pageId="21" pageNumber="422">2.4</td>
|
||
<td id="5A521309FFD80031F0D9F9DFFD1D5A00" box="[702,739,1588,1613]" gridcol="4" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F155F9DFFCA05A00" box="[818,862,1588,1613]" gridcol="5" gridrow="7" pageId="21" pageNumber="422">1.8</td>
|
||
<td id="5A521309FFD80031F1C1F9DFFC1B5A00" box="[934,997,1588,1613]" gridcol="6" gridrow="7" pageId="21" pageNumber="422">0.89</td>
|
||
<td id="5A521309FFD80031F67CF9DFFBA75A00" box="[1051,1113,1588,1613]" gridcol="7" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F6E8F9DFFB195A00" box="[1167,1255,1588,1613]" gridcol="8" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
<td id="5A521309FFD80031F778F9DFFA8E5A00" box="[1311,1392,1588,1613]" gridcol="9" gridrow="7" pageId="21" pageNumber="422">-</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F98DFA8E5A32" box="[163,1392,1638,1663]" gridrow="8" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F98DFEE45A32" box="[163,282,1638,1663]" gridcol="0" gridrow="8" pageId="21" pageNumber="422">Paratype 13</th>
|
||
<td id="5A521309FFD80031F315F98DFE4A5A32" box="[370,436,1638,1663]" gridcol="1" gridrow="8" pageId="21" pageNumber="422">10F</td>
|
||
<td id="5A521309FFD80031F381F98DFDF55A32" box="[486,523,1638,1663]" gridcol="2" gridrow="8" pageId="21" pageNumber="422">2.8</td>
|
||
<td id="5A521309FFD80031F035F98DFD8C5A32" box="[594,626,1638,1663]" gridcol="3" gridrow="8" pageId="21" pageNumber="422">2.6</td>
|
||
<td id="5A521309FFD80031F0D9F98DFD1D5A32" box="[702,739,1638,1663]" gridcol="4" gridrow="8" pageId="21" pageNumber="422">2.6</td>
|
||
<td id="5A521309FFD80031F155F98DFCA05A32" box="[818,862,1638,1663]" gridcol="5" gridrow="8" pageId="21" pageNumber="422">1.7</td>
|
||
<td id="5A521309FFD80031F1C1F98DFC1B5A32" box="[934,997,1638,1663]" gridcol="6" gridrow="8" pageId="21" pageNumber="422">0.92</td>
|
||
<td id="5A521309FFD80031F67CF98DFBA75A32" box="[1051,1113,1638,1663]" gridcol="7" gridrow="8" pageId="21" pageNumber="422">1.01</td>
|
||
<td id="5A521309FFD80031F6E8F98DFB195A32" box="[1167,1255,1638,1663]" gridcol="8" gridrow="8" pageId="21" pageNumber="422">1.51</td>
|
||
<td id="5A521309FFD80031F778F98DFA8E5A32" box="[1311,1392,1638,1663]" gridcol="9" gridrow="8" pageId="21" pageNumber="422">0.91</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F973FA8E5AFC" box="[163,1392,1688,1713]" gridrow="9" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F973FEE45AFC" box="[163,282,1688,1713]" gridcol="0" gridrow="9" pageId="21" pageNumber="422">Paratype 16</th>
|
||
<td id="5A521309FFD80031F315F973FE4A5AFC" box="[370,436,1688,1713]" gridcol="1" gridrow="9" pageId="21" pageNumber="422">10H</td>
|
||
<td id="5A521309FFD80031F381F973FDF55AFC" box="[486,523,1688,1713]" gridcol="2" gridrow="9" pageId="21" pageNumber="422">3.8</td>
|
||
<td id="5A521309FFD80031F035F973FD8C5AFC" box="[594,626,1688,1713]" gridcol="3" gridrow="9" pageId="21" pageNumber="422">3.4</td>
|
||
<td id="5A521309FFD80031F0D9F973FD1D5AFC" box="[702,739,1688,1713]" gridcol="4" gridrow="9" pageId="21" pageNumber="422">3.4</td>
|
||
<td id="5A521309FFD80031F155F973FCA05AFC" box="[818,862,1688,1713]" gridcol="5" gridrow="9" pageId="21" pageNumber="422">2.0</td>
|
||
<td id="5A521309FFD80031F1C1F973FC1B5AFC" box="[934,997,1688,1713]" gridcol="6" gridrow="9" pageId="21" pageNumber="422">0.90</td>
|
||
<td id="5A521309FFD80031F67CF973FBA75AFC" box="[1051,1113,1688,1713]" gridcol="7" gridrow="9" pageId="21" pageNumber="422">1.02</td>
|
||
<td id="5A521309FFD80031F6E8F973FB195AFC" box="[1167,1255,1688,1713]" gridcol="8" gridrow="9" pageId="21" pageNumber="422">1.64</td>
|
||
<td id="5A521309FFD80031F778F973FA8E5AFC" box="[1311,1392,1688,1713]" gridcol="9" gridrow="9" pageId="21" pageNumber="422">0.88</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F921FA8E5AAF" box="[163,1392,1738,1762]" gridrow="10" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F921FEE45AAF" box="[163,282,1738,1762]" gridcol="0" gridrow="10" pageId="21" pageNumber="422">Paratype 17</th>
|
||
<td id="5A521309FFD80031F315F921FE4A5AAF" box="[370,436,1738,1762]" gridcol="1" gridrow="10" pageId="21" pageNumber="422">10I</td>
|
||
<td id="5A521309FFD80031F381F921FDF55AAF" box="[486,523,1738,1762]" gridcol="2" gridrow="10" pageId="21" pageNumber="422">4.1</td>
|
||
<td id="5A521309FFD80031F035F921FD8C5AAF" box="[594,626,1738,1762]" gridcol="3" gridrow="10" pageId="21" pageNumber="422">4.0</td>
|
||
<td id="5A521309FFD80031F0D9F921FD1D5AAF" box="[702,739,1738,1762]" gridcol="4" gridrow="10" pageId="21" pageNumber="422">2.9</td>
|
||
<td id="5A521309FFD80031F155F921FCA05AAF" box="[818,862,1738,1762]" gridcol="5" gridrow="10" pageId="21" pageNumber="422">2.4</td>
|
||
<td id="5A521309FFD80031F1C1F921FC1B5AAF" box="[934,997,1738,1762]" gridcol="6" gridrow="10" pageId="21" pageNumber="422">0.97</td>
|
||
<td id="5A521309FFD80031F67CF921FBA75AAF" box="[1051,1113,1738,1762]" gridcol="7" gridrow="10" pageId="21" pageNumber="422">1.35</td>
|
||
<td id="5A521309FFD80031F6E8F921FB195AAF" box="[1167,1255,1738,1762]" gridcol="8" gridrow="10" pageId="21" pageNumber="422">1.24</td>
|
||
<td id="5A521309FFD80031F778F921FA8E5AAF" box="[1311,1392,1738,1762]" gridcol="9" gridrow="10" pageId="21" pageNumber="422">0.72</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F917FA8E5B59" box="[163,1392,1788,1812]" gridrow="11" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F917FEE45B59" box="[163,282,1788,1812]" gridcol="0" gridrow="11" pageId="21" pageNumber="422">Paratype 18</th>
|
||
<td id="5A521309FFD80031F315F917FE4A5B59" box="[370,436,1788,1812]" gridcol="1" gridrow="11" pageId="21" pageNumber="422">10K</td>
|
||
<td id="5A521309FFD80031F381F917FDF55B59" box="[486,523,1788,1812]" gridcol="2" gridrow="11" pageId="21" pageNumber="422">4.4</td>
|
||
<td id="5A521309FFD80031F035F917FD8C5B59" box="[594,626,1788,1812]" gridcol="3" gridrow="11" pageId="21" pageNumber="422">3.7</td>
|
||
<td id="5A521309FFD80031F0D9F917FD1D5B59" box="[702,739,1788,1812]" gridcol="4" gridrow="11" pageId="21" pageNumber="422">3.1</td>
|
||
<td id="5A521309FFD80031F155F917FCA05B59" box="[818,862,1788,1812]" gridcol="5" gridrow="11" pageId="21" pageNumber="422">2.4</td>
|
||
<td id="5A521309FFD80031F1C1F917FC1B5B59" box="[934,997,1788,1812]" gridcol="6" gridrow="11" pageId="21" pageNumber="422">0.84</td>
|
||
<td id="5A521309FFD80031F67CF917FBA75B59" box="[1051,1113,1788,1812]" gridcol="7" gridrow="11" pageId="21" pageNumber="422">1.18</td>
|
||
<td id="5A521309FFD80031F6E8F917FB195B59" box="[1167,1255,1788,1812]" gridcol="8" gridrow="11" pageId="21" pageNumber="422">1.30</td>
|
||
<td id="5A521309FFD80031F778F917FA8E5B59" box="[1311,1392,1788,1812]" gridcol="9" gridrow="11" pageId="21" pageNumber="422">0.71</td>
|
||
</tr>
|
||
<tr id="19837A75FFD80031F2C4F8C5FA8E5B0B" box="[163,1392,1838,1862]" gridrow="12" pageId="21" pageNumber="422">
|
||
<th id="5A521309FFD80031F2C4F8C5FEE45B0B" box="[163,282,1838,1862]" gridcol="0" gridrow="12" pageId="21" pageNumber="422">Paratype 19</th>
|
||
<td id="5A521309FFD80031F315F8C5FE4A5B0B" box="[370,436,1838,1862]" gridcol="1" gridrow="12" pageId="21" pageNumber="422">10L</td>
|
||
<td id="5A521309FFD80031F381F8C5FDF55B0B" box="[486,523,1838,1862]" gridcol="2" gridrow="12" pageId="21" pageNumber="422">4.9</td>
|
||
<td id="5A521309FFD80031F035F8C5FD8C5B0B" box="[594,626,1838,1862]" gridcol="3" gridrow="12" pageId="21" pageNumber="422">4.5</td>
|
||
<td id="5A521309FFD80031F0D9F8C5FD1D5B0B" box="[702,739,1838,1862]" gridcol="4" gridrow="12" pageId="21" pageNumber="422">3.9</td>
|
||
<td id="5A521309FFD80031F155F8C5FCA05B0B" box="[818,862,1838,1862]" gridcol="5" gridrow="12" pageId="21" pageNumber="422">2.4</td>
|
||
<td id="5A521309FFD80031F1C1F8C5FC1B5B0B" box="[934,997,1838,1862]" gridcol="6" gridrow="12" pageId="21" pageNumber="422">0.92</td>
|
||
<td id="5A521309FFD80031F67CF8C5FBA75B0B" box="[1051,1113,1838,1862]" gridcol="7" gridrow="12" pageId="21" pageNumber="422">1.17</td>
|
||
<td id="5A521309FFD80031F6E8F8C5FB195B0B" box="[1167,1255,1838,1862]" gridcol="8" gridrow="12" pageId="21" pageNumber="422">1.58</td>
|
||
<td id="5A521309FFD80031F778F8C5FA8E5B0B" box="[1311,1392,1838,1862]" gridcol="9" gridrow="12" pageId="21" pageNumber="422">0.78</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFD8FFD8F2A0F873FDA95D0C" blockId="21.[151,1437,1944,2043]" lastBlockId="22.[151,1437,150,357]" lastPageId="22" lastPageNumber="423" pageId="21" pageNumber="422">
|
||
<bibRefCitation id="C32205C6FFD8FFDBF2A0F873FE115BFF" author="Manni, R. & Pacioni, E." box="[199,495,1944,1970]" pageId="21" pageNumber="422" pagination="65 - 77" refId="ref22121" refString="Manni, R. & Pacioni, E. (2021) New records of the cyrtocrinid crinoids Cyathidium gastaldii (Michelin, 1851) and Holopus spileccense (Schluter, 1878) from Eocene strata in Veneto, north-east Italy. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen, 301 (1), 65 - 77. https: // doi. org / 10.1127 / njgpa / 2021 / 0997" type="journal article" year="2021">Manni and Pacioni (2021)</bibRefCitation>
|
||
attributed 30 brachials from
|
||
<collectingRegion id="6577B6D5FFD8FFDBF15DF873FC8A5BFF" box="[826,884,1944,1970]" country="Indonesia" name="Jawa Timur" pageId="21" pageNumber="422">Cava</collectingRegion>
|
||
Boschetto (= Cengio dell’Orbo quarry, the type-locality of
|
||
<taxonomicName id="60B303B4FFD8FFDBF375F856FE735B9B" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[274,397,1980,2006]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFD8FFDBF375F856FE735B9B" box="[274,397,1980,2006]" italics="true" pageId="21" pageNumber="422">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFD8FFDBF3F2F857FE115B9B" bold="true" box="[405,495,1980,2006]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="8EF4195EFFD8FFDBF3F2F857FE115B9B" box="[405,495,1980,2006]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) to
|
||
<taxonomicName id="60B303B4FFD8FFDBF079F856FCC25B9B" authority="(Schluter)" baseAuthorityName="Schluter" box="[542,828,1980,2006]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="21" pageNumber="422" phylum="Echinodermata" rank="species" species="spileccense">
|
||
<emphasis id="95C7A425FFD8FFDBF079F856FD3F5B9B" box="[542,705,1980,2006]" italics="true" pageId="21" pageNumber="422">H. spileccense</emphasis>
|
||
(Schlüter)
|
||
</taxonomicName>
|
||
, a taxon known only from cups found in the Ypresian strata of Spilecco Hill (see below). These 30 brachials are housed in the MGP-PD collections.
|
||
<bibRefCitation id="C32205C6FFD8FFD8F6A0F80BFF1C5CFC" author="Manni, R. & Pacioni, E." lastPageId="22" lastPageNumber="423" pageId="21" pageNumber="422" pagination="65 - 77" refId="ref22121" refString="Manni, R. & Pacioni, E. (2021) New records of the cyrtocrinid crinoids Cyathidium gastaldii (Michelin, 1851) and Holopus spileccense (Schluter, 1878) from Eocene strata in Veneto, north-east Italy. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen, 301 (1), 65 - 77. https: // doi. org / 10.1127 / njgpa / 2021 / 0997" type="journal article" year="2021">Manni and Pacioni (2021)</bibRefCitation>
|
||
figured the ornamentation of 5 IBrax and 6 IIBr from the proximal series but did not identify keystone plates. The variation of X-shaped ridges on IIBr was well-illustrated and corresponds to the range observed in the material here ascribed to
|
||
<taxonomicName id="60B303B4FFDBFFD8F32BFF34FE3A5CB4" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[332,452,223,249]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="22" pageNumber="423" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDBFFD8F32BFF34FE3A5CB4" box="[332,452,223,249]" italics="true" pageId="22" pageNumber="423">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDBFFD8F3ADFF34FDDC5CB4" bold="true" box="[458,546,223,249]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="8EF4195EFFDBFFD8F3ADFF34FDDC5CB4" box="[458,546,223,249]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The quantitative characters of these brachials also fall within the variation range of those of
|
||
<taxonomicName id="60B303B4FFDBFFD8F37BFEE8FE695D50" authorityName="Martinez-Soares & Roux & Giusberti & Gatto & Eléaume & Améziane" authorityYear="2024" box="[284,407,259,285]" class="Crinoidea" family="Holopodidae" genus="Holopus" kingdom="Animalia" order="Cyrtocrinida" pageId="22" pageNumber="423" phylum="Echinodermata" rank="species" species="fabianii" status="sp. nov.">
|
||
<emphasis id="95C7A425FFDBFFD8F37BFEE8FE695D50" box="[284,407,259,285]" italics="true" pageId="22" pageNumber="423">H. fabianii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95C7A425FFDBFFD8F3C7FEE8FE055D50" bold="true" box="[416,507,259,285]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="8EF4195EFFDBFFD8F3C7FEE8FE055D50" box="[416,507,259,285]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(except IBrax MGP-PD 31442 which presents the largest Wd,
|
||
<quantity id="604BD5D2FFDBFFD8F6B6FEE9FAD45D50" box="[1233,1322,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="22" pageNumber="423" unit="mm" value="5.3">5.3 mm</quantity>
|
||
) and they should thus be attributed to this species.
|
||
</paragraph>
|
||
<paragraph id="A70C7837FFDBFFD8F2A0FEA1FBC25D28" blockId="22.[151,1437,150,357]" box="[199,1084,330,357]" pageId="22" pageNumber="423">
|
||
<emphasis id="95C7A425FFDBFFD8F2A0FEA1FEA45D29" bold="true" box="[199,346,330,356]" pageId="22" pageNumber="423">Occurrence.</emphasis>
|
||
Middle Eocene (Lutetian) of Chiampo Valley, north-eastern
|
||
<collectingCountry id="DFA438A7FFDBFFD8F664FEA0FBC65D28" box="[1027,1080,331,357]" name="Italy" pageId="22" pageNumber="423">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |