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Genus
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<emphasis id="95C7A425FFC1FFC2F289FF7CFE825CFF" bold="true" box="[238,380,151,178]" italics="true" pageId="12" pageNumber="413">Cyathidium</emphasis>
Steenstrup
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Michaelis &amp; Scherk, 1847
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<emphasis id="95C7A425FFC1FFC2F2F0FF30FEDB5CBE" bold="true" box="[151,293,219,243]" pageId="12" pageNumber="413">Type-species.</emphasis>
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<emphasis id="95C7A425FFC1FFC2F34BFF37FE065CB9" box="[300,504,220,244]" italics="true" pageId="12" pageNumber="413">Cyathidium holopus</emphasis>
Steenstrup
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Michaelis &amp; Scherk, 1847, by monotypy.
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<emphasis id="95C7A425FFC1FFC2F37CFF11FE6F5D5F" box="[283,401,250,274]" italics="true" pageId="12" pageNumber="413">Cyathidium</emphasis>
(
<bibRefCitation id="C32205C6FFC1FFC2F3C3FF11FDAF5D5F" author="Steenstrup, J. J." box="[420,593,250,274]" pageId="12" pageNumber="413" pagination="148 - 150" refId="ref23909" refString="Steenstrup, J. J. (1847) Uber Korallen in der Faxoer Kreide, Moltkia und Cyathidium. In: Michaelis, G. A. &amp; Scherk, H. F. (Eds.), Amtlicher Bericht uber die 24. Versammlung Deutscher Naturforscher und Arzte in Kiel, im September 1846. Akademische Buchhandlung, Kiel, pp. 148 - 150." type="book chapter" year="1847">Steenstrup 1847</bibRefCitation>
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in Michaelis &amp; Scherk, 1847: 119, 150;
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<emphasis id="95C7A425FFC1FFC2F642FF10FB455D5F" box="[1061,1211,251,274]" italics="true" pageId="12" pageNumber="413">Micropocrinus</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F6AFFF11FA665D5F" author="Michelin, H." box="[1224,1432,250,274]" pageId="12" pageNumber="413" pagination="93 - 94" refId="ref22506" refString="Michelin, H. (1851) Description d'un nouveau genre de la famille des Crinoides. Revue et magazin de zoologie pure et appliquee, 2 (3), 93 - 94." type="journal article" year="1851">Michelin, 1851: 93</bibRefCitation>
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;
<bibRefCitation id="C32205C6FFC1FFC2F2A0FEF2FE795D7C" author="Sieverts, H." box="[199,391,281,305]" pageId="12" pageNumber="413" pagination="159 - 176" refId="ref23808" refString="Sieverts, H. (1931) Cyathidium depressum aus dem Oberen Cenoman Rheinland-Westfalens. Jahrbuch der Preussischen Geologischen Landesanstalt zu Berlin, 52, 159 - 176." type="journal article" year="1931">Sieverts, 1931:173</bibRefCitation>
;
<taxonomicName id="60B303B4FFC1FFC2F3F5FEF2FD3F5D7C" authority="Valette" authorityName="Valette" authorityYear="1934" box="[402,705,281,305]" genus="Pseudocupressocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="413" phylum="Echinodermata" rank="genus">
<emphasis id="95C7A425FFC1FFC2F3F5FEF2FD8B5D7C" box="[402,629,281,305]" italics="true" pageId="12" pageNumber="413">Pseudocupressocrinus</emphasis>
Valette
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<emphasis id="95C7A425FFC1FFC2F0A0FEF1FD255D7C" box="[711,731,282,305]" italics="true" pageId="12" pageNumber="413">in</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F086FEF2FBFA5D7C" author="Lambert, J. &amp; Valette, D." box="[737,1028,281,305]" pageId="12" pageNumber="413" pagination="43 - 60" refId="ref21885" refString="Lambert, J. &amp; Valette, D. (1934) Etude sur quelque Echinodermes cretaces de Bugarach (Aude). Bulletin de la Societe Geologique de France, 5 e Serie, 4 (1 - 3), 43 - 60." type="journal article" year="1934">Lambert &amp; Valette, 1934: 59</bibRefCitation>
;
<taxonomicName id="60B303B4FFC1FFC2F668FEF2FA665D7C" authority="Sieverts-Doreck, 1938: 29" authorityName="Sieverts-Doreck" authorityPageNumber="29" authorityYear="1938" box="[1039,1432,281,305]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="genus">
<emphasis id="95C7A425FFC1FFC2F668FEF2FB7B5D7C" box="[1039,1157,281,305]" italics="true" pageId="12" pageNumber="413">Cyathidium</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F6ECFEF2FA665D7C" author="Sieverts-Doreck, H." box="[1163,1432,281,305]" pageId="12" pageNumber="413" pagination="29 - 32" refId="ref23837" refString="Sieverts-Doreck, H. (1938) Pseudocupressocrinus Valette, 1934, ein Synonym von Cyathidium Steenstrup, 1847. Zentralblatt fur Mineralogie, Geologie und Palantologie, Abteilung B: Geologie und Palantologie, 1938, 29 - 32." type="journal article" year="1938">Sieverts-Doreck, 1938: 29</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="60B303B4FFC1FFC2F2A0FED3FD2B5D1D" authority="Biese &amp; Sieverts-Doreck, 1939: 117" authorityName="Biese &amp; Sieverts-Doreck" authorityPageNumber="117" authorityYear="1939" box="[199,725,312,336]" genus="Micropocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="413" phylum="Echinodermata" rank="genus">
<emphasis id="95C7A425FFC1FFC2F2A0FED3FEA35D02" box="[199,349,312,335]" italics="true" pageId="12" pageNumber="413">Micropocrinus</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F304FED3FD2B5D1D" author="Biese, W. &amp; Sieverts-Doreck, H." box="[355,725,312,336]" pageId="12" pageNumber="413" refId="ref20269" refString="Biese, W. &amp; Sieverts-Doreck, H. (1939) Crinoidea caenozoica. In: Quenstedt, W. (Ed.), Fossilium Catalogus. I. Animalia. Vol. 80. W. Junk, ' s-Gravenhage, pp. 1 - 151." type="book" year="1939">Biese &amp; Sieverts-Doreck, 1939: 117</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="60B303B4FFC1FFC2F087FED3FB885D1D" authority="Rasmussen, 1961: 238 - 239" authorityName="Rasmussen" authorityPageNumber="238 - 239" authorityYear="1961" box="[736,1142,311,336]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="genus">
<emphasis id="95C7A425FFC1FFC2F087FED3FCA85D1D" box="[736,854,312,336]" italics="true" pageId="12" pageNumber="413">Cyathidium</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F13BFED3FB885D1D" author="Rasmussen, H. W." box="[860,1142,311,336]" pageId="12" pageNumber="413" pagination="1 - 428" refId="ref22859" refString="Rasmussen, H. W. (1961) A Monograph on the Cretaceous Crinoidea. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 12 (1), 1 - 428." type="journal article" year="1961">Rasmussen, 1961: 238239</bibRefCitation>
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;
<bibRefCitation id="C32205C6FFC1FFC2F6E6FED3FB495D1D" author="Rasmussen, H. W." box="[1153,1207,312,336]" pageId="12" pageNumber="413" refId="ref22933" refString="Rasmussen, H. W. (1978) Articulata. In: Moore, R. C. &amp; Teichert C. (Eds.), Treatise on Invertebrate Paleontology. Pt. T. Echinodermata 2. Vol. 3. The Geological Society of America, Inc. Boulder, Colorado and The University of Kansas Press, Lawrence, Kansas, pp. 813 - 928" type="book" year="1978">1978</bibRefCitation>
: T839.
</paragraph>
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<emphasis id="95C7A425FFC1FFC2F2F0FE95FEF55DD5" bold="true" box="[151,267,382,408]" pageId="12" pageNumber="413">Remarks.</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F375FE94FE355DD5" author="Zitt, J. &amp; Vodrazka, R. &amp; Zagorsek, K." box="[274,459,382,410]" pageId="12" pageNumber="413" pagination="56 - 69" refId="ref24389" refString="Zitt, J., Vodrazka, R. &amp; Zagorsek, K. (2014) New observations on the holopodid crinoid genus Cyathidium Steenstrup from the lower Turonian of the Bohemian Cretaceous Basin, Czech Republic. Cretaceous Research, 51, 56 - 69. https: // doi. org / 10.1016 / j. cretres. 2014.05.019" type="journal article" year="2014">
Žítt
<emphasis id="95C7A425FFC1FFC2F326FE94FE845DD5" box="[321,378,382,408]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
(2014)
</bibRefCitation>
thoroughly described the wide range of shape variation in a lower Turonian population of 136 aboral cups attributed to
<taxonomicName id="60B303B4FFC1FFC2F06CFE49FCC35DF1" authorityName="Sieverts" authorityYear="1932" box="[523,829,418,444]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" isUncertain="true" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="95C7A425FFC1FFC2F06CFE49FD705DF1" box="[523,654,418,444]" italics="true" pageId="12" pageNumber="413">Cyathidium</emphasis>
aff.
<emphasis id="95C7A425FFC1FFC2F0A1FE49FCC35DF1" box="[710,829,418,444]" italics="true" pageId="12" pageNumber="413">depressum</emphasis>
</taxonomicName>
. The ratio H/D varies from 0.2 to
<emphasis id="95C7A425FFC1FFC2F6B7FE48FB1D5DF1" box="[1232,1251,419,444]" italics="true" pageId="12" pageNumber="413">c.</emphasis>
1.0 and usually increases during growth (
<figureCitation id="3F8864B2FFC1FFC2F3D9FE2DFDF85DAD" box="[446,518,454,480]" captionStart="FIGURE 7" captionStartId="11.[151,250,810,834]" captionTargetBox="[210,1376,188,783]" captionTargetId="figure-19@11.[204,1383,181,785]" captionTargetPageId="11" captionText="FIGURE 7. Main quantitative characters of aboral cup and IBrax in fossil Holopodidae. S: Cyathidium holopus from Seymour Island. Straight lines: values of H/D and H/Wd. Other values in mm." figureDoi="http://doi.org/10.5281/zenodo.14241174" httpUri="https://zenodo.org/record/14241174/files/figure.png" pageId="12" pageNumber="413">Fig. 7</figureCitation>
). The cup curvature is also greatly variable with H/h from 1 to 2.7. Details of characters discriminating the extant species such as interradial morphology and fulcral ridge arrangement (
<bibRefCitation id="C32205C6FFC1FFC2F74EFE01FEEC5E65" author="Ameziane, N. &amp; Bourseau, J. - P. &amp; Heinzeller, T. &amp; Roux, M." pageId="12" pageNumber="413" pagination="439 - 470" refId="ref19538" refString="Ameziane, N., Bourseau, J. - P., Heinzeller, T. &amp; Roux, M. (1999) Les genres Cyathidium et Holopus au sein des Cyrtocrinida (Crinoidea; Echinodermata). Journal of Natural History, 33 (3), 439 - 470. https: // doi. org / 10.1080 / 002229399300335" type="journal article" year="1999">
Améziane
<emphasis id="95C7A425FFC1FFC2F2F0FDE4FF2F5E65" box="[151,209,526,552]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
1999
</bibRefCitation>
) are only preserved in a few specimens, and only parts of the external surface show growth lines and peculiar ornamentation. Therefore, attribution to a new species or to previously described species merely based on aboral cups requires well-preserved specimens and discriminating qualitative characters. Intact fossil specimens with a retracted crown are known exclusively in
<taxonomicName id="60B303B4FFC1FFC2F0A3FD91FCC15ED9" authorityName="Steenstrup" authorityYear="1847" box="[708,831,634,660]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="holopus">
<emphasis id="95C7A425FFC1FFC2F0A3FD91FCC15ED9" box="[708,831,634,660]" italics="true" pageId="12" pageNumber="413">C. holopus</emphasis>
</taxonomicName>
(
<bibRefCitation id="C32205C6FFC1FFC2F129FD91FB655ED9" author="Baumiller, T. K. &amp; Gazdzicki, A." box="[846,1179,634,660]" pageId="12" pageNumber="413" pagination="101 - 116" refId="ref19917" refString="Baumiller, T. K. &amp; Gazdzicki, A. (1996) New Crinoids from the Eocene La Meseta Formation of Seymour Island, Antarctic Peninsula. Palaeontologica Polonica, 55, 101 - 116." type="journal article" year="1996">Baumiller &amp; Gaździcki 1996</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F6CFFD91FF2D5EF5" author="Donovan, S. K. &amp; Jakobsen, S." pageId="12" pageNumber="413" pagination="407 - 415" refId="ref20922" refString="Donovan, S. K. &amp; Jakobsen, S. (2004) An unusual crinoid-barnacle association in the type area of the Danian (Paleocene), Denmark. Lethaia, 37 (4), 407 - 415. https: // doi. org / 10.1080 / 00241160410002126" type="journal article" year="2004">Donovan &amp; Jakobsen 2004</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F2BAFD75FE285EF5" author="Hess, H. &amp; Messing, C. G. &amp; Part T." box="[221,470,670,696]" pageId="12" pageNumber="413" refId="ref21626" refString="Hess, H. &amp; Messing, C. G. (Eds.) (2011) Treatise on Invertebrate Paleontology. Part T. Echinodermata 2. Crinoidea. Vol. 3. Revised. The University of Kansas Paleontological Institute, Lawrence, Kansas, 261 pp." type="book" year="2011">Hess &amp; Messing 2011</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F386FD75FD865EF5" author="Nielsen, K. B." box="[481,632,670,696]" pageId="12" pageNumber="413" pagination="1 - 120" refId="ref22580" refString="Nielsen, K. B. (1913) Crinoiderne i Danmarks Kridtaflejringer. Danmarks Geologiske UndersOgelse, 2 (26), 1 - 120. https: // doi. org / 10.34194 / raekke 2. v 26.6811" type="journal article" year="1913">Nielsen 1913</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F0E4FD75FCBD5EF5" author="Rasmussen, H. W." box="[643,835,670,696]" pageId="12" pageNumber="413" pagination="1 - 428" refId="ref22859" refString="Rasmussen, H. W. (1961) A Monograph on the Cretaceous Crinoidea. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 12 (1), 1 - 428." type="journal article" year="1961">Rasmussen 1961</bibRefCitation>
). The vault covering the distal arm series is composed solely of IIBr1, their height being substantially greater than their width. IIBr have not yet been described from the large species
<taxonomicName id="60B303B4FFC1FFC2F33DFD0DFD555F4D" authority="Jagt, 1986" authorityName="Jagt" authorityYear="1986" box="[346,683,742,768]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="vlieksi">
<emphasis id="95C7A425FFC1FFC2F33DFD0DFDD35F4D" box="[346,557,742,768]" italics="true" pageId="12" pageNumber="413">Cyathidium vlieksi</emphasis>
<bibRefCitation id="C32205C6FFC1FFC2F053FD0DFD555F4D" author="Jagt, J. W. M." box="[564,683,742,768]" pageId="12" pageNumber="413" pagination="215 - 221" refId="ref21737" refString="Jagt, J. W. M. (1986) Cyathidium vlieksi, a new holopodid crinoid from the Upper Maastrichtian (Late Cretaceous) of southern Limburg, The Netherlands. Geologie en Mijnbouw, 65 (3), 215 - 221." type="journal article" year="1986">Jagt, 1986</bibRefCitation>
</taxonomicName>
, but are known (Jagt pers. comm.). This species has an aboral cup seemingly very similar to some specimens of
<taxonomicName id="60B303B4FFC1FFC2F0F3FCE1FCF35F69" authorityName="Steenstrup" authorityYear="1847" box="[660,781,778,804]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="holopus">
<emphasis id="95C7A425FFC1FFC2F0F3FCE1FCF35F69" box="[660,781,778,804]" italics="true" pageId="12" pageNumber="413">C. holopus</emphasis>
</taxonomicName>
. Intraspecific variation of the aboral cup in
<taxonomicName id="60B303B4FFC1FFC2F692FCE1FA905F69" authorityName="Steenstrup" authorityYear="1847" box="[1269,1390,778,804]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="holopus">
<emphasis id="95C7A425FFC1FFC2F692FCE1FA905F69" box="[1269,1390,778,804]" italics="true" pageId="12" pageNumber="413">C. holopus</emphasis>
</taxonomicName>
and
<taxonomicName id="60B303B4FFC1FFC2F2F0FCC5FEFF5F05" authorityName="Jagt" authorityYear="1986" box="[151,257,814,840]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="vlieksi">
<emphasis id="95C7A425FFC1FFC2F2F0FCC5FEFF5F05" box="[151,257,814,840]" italics="true" pageId="12" pageNumber="413">C. vlieksi</emphasis>
</taxonomicName>
show two extreme morphotypes: one very depressed, like in
<emphasis id="95C7A425FFC1FFC2F1D0FCC5FBA85F05" box="[951,1110,814,840]" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="60B303B4FFC1FFC2F1D0FCC5FBAC5F05" authorityName="Sieverts" authorityYear="1932" box="[951,1106,814,840]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">C. depressum</taxonomicName>
,
</emphasis>
and another as high or taller than wide with conspicuous growth lines. The lower Santonian species,
<emphasis id="95C7A425FFC1FFC2F1ABFCB9FBAF5F21" box="[972,1105,850,876]" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="60B303B4FFC1FFC2F1ABFCB9FBB25F21" baseAuthorityName="Valette" baseAuthorityYear="1934" box="[972,1100,850,876]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="senessei">C. senessei</taxonomicName>
,
</emphasis>
differs from all other known congeners in having a stout and tumid curved IBrax, with relatively small distal articular facets facing inwards and subperpendicular to the upper plan of the IBr circlet (
<bibRefCitation id="C32205C6FFC1FFC2F17BFC71FBEE5FF9" author="Ameziane, N. &amp; Bourseau, J. - P. &amp; Heinzeller, T. &amp; Roux, M." box="[796,1040,922,948]" pageId="12" pageNumber="413" pagination="439 - 470" refId="ref19538" refString="Ameziane, N., Bourseau, J. - P., Heinzeller, T. &amp; Roux, M. (1999) Les genres Cyathidium et Holopus au sein des Cyrtocrinida (Crinoidea; Echinodermata). Journal of Natural History, 33 (3), 439 - 470. https: // doi. org / 10.1080 / 002229399300335" type="journal article" year="1999">
Améziane
<emphasis id="95C7A425FFC1FFC2F1F1FC70FC315FF9" box="[918,975,922,948]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F67BFC71FAEC5FF9" author="Heinzeller, T. &amp; Fricke, H. &amp; Bourseau, J. - P. &amp; Ameziane-Cominardi, N. &amp; Welsch, U." box="[1052,1298,922,948]" pageId="12" pageNumber="413" pagination="77 - 84" refId="ref21460" refString="Heinzeller, T., Fricke, H., Bourseau, J. - P., Ameziane-Cominardi, N. &amp; Welsch, U. (1996) Cyathidium plantei sp. n., an extant cyrtocrinid (Echinodermata, Crinoidea) morphologically identical to the fossil Cyathidium depressum (Cretaceous, Cenomanian). Zoologica Scripta, 25 (1), 77 - 84. https: // doi. org / 10.1111 / j. 1463 - 6409.1996. tb 00153. x" type="journal article" year="1996">
Heinzeller
<emphasis id="95C7A425FFC1FFC2F6FFFC70FB2F5FF9" box="[1176,1233,922,948]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
1996
</bibRefCitation>
;
<bibRefCitation id="C32205C6FFC1FFC2F779FC71FF2D5F95" author="Rasmussen, H. W." pageId="12" pageNumber="413" pagination="1 - 428" refId="ref22859" refString="Rasmussen, H. W. (1961) A Monograph on the Cretaceous Crinoidea. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 12 (1), 1 - 428." type="journal article" year="1961">Rasmussen 1961</bibRefCitation>
). The central space provided by the IBrax circlet (
<bibRefCitation id="C32205C6FFC1FFC2F179FC55FC1C5F95" author="Rasmussen, H. W." box="[798,994,958,984]" pageId="12" pageNumber="413" pagination="1 - 428" refId="ref22859" refString="Rasmussen, H. W. (1961) A Monograph on the Cretaceous Crinoidea. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 12 (1), 1 - 428." type="journal article" year="1961">Rasmussen 1961</bibRefCitation>
, pl. 34 fig. 7c) probably houses IIBr1 keystone plates, but these have not yet been described.
<bibRefCitation id="C32205C6FFC1FFC2F165FC08FC455FB1" author="Zitt, J. &amp; Vodrazka, R. &amp; Zagorsek, K." box="[770,955,994,1022]" pageId="12" pageNumber="413" pagination="56 - 69" refId="ref24389" refString="Zitt, J., Vodrazka, R. &amp; Zagorsek, K. (2014) New observations on the holopodid crinoid genus Cyathidium Steenstrup from the lower Turonian of the Bohemian Cretaceous Basin, Czech Republic. Cretaceous Research, 51, 56 - 69. https: // doi. org / 10.1016 / j. cretres. 2014.05.019" type="journal article" year="2014">
Žítt
<emphasis id="95C7A425FFC1FFC2F156FC08FC975FB1" box="[817,873,994,1020]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
(2014)
</bibRefCitation>
figured the main discriminating characters of dissociated brachials of
<taxonomicName id="60B303B4FFC1FFC2F3A5FBEDFD78586D" authorityName="Sieverts" authorityYear="1932" box="[450,646,1030,1056]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" isUncertain="true" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="95C7A425FFC1FFC2F3A5FBEDFE22586D" box="[450,476,1030,1056]" italics="true" pageId="12" pageNumber="413">C.</emphasis>
aff.
<emphasis id="95C7A425FFC1FFC2F068FBEDFD78586D" box="[527,646,1030,1056]" italics="true" pageId="12" pageNumber="413">depressum</emphasis>
</taxonomicName>
from the lower Turonian. Unfortunately, IIBr were not available in the original description of
<taxonomicName id="60B303B4FFC1FFC2F3FFFBC1FDD15809" authorityName="Sieverts" authorityYear="1932" box="[408,559,1066,1092]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="95C7A425FFC1FFC2F3FFFBC1FDD15809" box="[408,559,1066,1092]" italics="true" pageId="12" pageNumber="413">C. depressum</emphasis>
</taxonomicName>
from the upper Cenomanian but IBr1 were illustrated (
<bibRefCitation id="C32205C6FFC1FFC2F6FCFBC1FAA05809" author="Rasmussen, H. W." box="[1179,1374,1066,1092]" pageId="12" pageNumber="413" pagination="1 - 428" refId="ref22859" refString="Rasmussen, H. W. (1961) A Monograph on the Cretaceous Crinoidea. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 12 (1), 1 - 428." type="journal article" year="1961">Rasmussen 1961</bibRefCitation>
). The presence of granulation on the aboral cup appears highly variable among fossil assemblages and cannot be used as a specific discriminating character except when there are sufficient ossicles to demonstrate the stability of this trait. Another early Campanian form from southern
<collectingCountry id="DFA438A7FFC1FFC2F0C7FB7DFD0358FD" box="[672,765,1174,1200]" name="United Kingdom" pageId="12" pageNumber="413">England</collectingCountry>
and north-west
<collectingCountry id="DFA438A7FFC1FFC2F1D6FB7DFC0358FD" box="[945,1021,1174,1200]" name="France" pageId="12" pageNumber="413">France</collectingCountry>
will be published soon (Gale &amp; Jagt, in press).
</paragraph>
<paragraph id="A70C7837FFC1FFC2F2A0FB35FB5F59E1" blockId="12.[151,1437,382,1488]" pageId="12" pageNumber="413">
<emphasis id="95C7A425FFC1FFC2F2A0FB35FD7058B5" bold="true" box="[199,654,1245,1272]" pageId="12" pageNumber="413">Included fossil species and occurrence.</emphasis>
<taxonomicName id="60B303B4FFC1FFC2F0F3FB35FC4A58B5" authorityName="Martinez-Soares &amp; Roux &amp; Giusberti &amp; Gatto &amp; Eléaume &amp; Améziane" authorityYear="2024" box="[660,948,1246,1272]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="chiampoensis" status="sp. nov.">
<emphasis id="95C7A425FFC1FFC2F0F3FB35FC4A58B5" box="[660,948,1246,1272]" italics="true" pageId="12" pageNumber="413">Cyathidium chiampoensis</emphasis>
</taxonomicName>
<emphasis id="95C7A425FFC1FFC2F1DDFB35FBEC58B5" bold="true" box="[954,1042,1246,1272]" pageId="12" pageNumber="413">
<taxonomicNameLabel id="8EF4195EFFC1FFC2F1DDFB35FBEC58B5" box="[954,1042,1246,1272]" pageId="12" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(lower Lutetian of Chiampo valley, north-eastern
<collectingCountry id="DFA438A7FFC1FFC2F357FAE9FE995951" box="[304,359,1282,1308]" name="Italy" pageId="12" pageNumber="413">Italy</collectingCountry>
),
<taxonomicName id="60B303B4FFC1FFC2F31FFAE9FDF05951" authorityName="Sieverts" authorityYear="1932" box="[376,526,1282,1308]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="95C7A425FFC1FFC2F31FFAE9FDF05951" box="[376,526,1282,1308]" italics="true" pageId="12" pageNumber="413">C. depressum</emphasis>
</taxonomicName>
including
<taxonomicName id="60B303B4FFC1FFC2F0E6FAE9FCBF5951" authorityName="Sieverts" authorityYear="1932" box="[641,833,1282,1308]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" isUncertain="true" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="95C7A425FFC1FFC2F0E6FAE9FD655951" box="[641,667,1282,1308]" italics="true" pageId="12" pageNumber="413">C.</emphasis>
aff.
<emphasis id="95C7A425FFC1FFC2F0ADFAE9FCBF5951" box="[714,833,1282,1308]" italics="true" pageId="12" pageNumber="413">depressum</emphasis>
</taxonomicName>
first described by
<bibRefCitation id="C32205C6FFC1FFC2F66EFAE8FB905951" author="Zitt, J." box="[1033,1134,1282,1310]" pageId="12" pageNumber="413" pagination="233 - 239" refId="ref24348" refString="Zitt, J. (1996) Cyathidium Steenstrup (Crinoidea) in the Upper Cretaceous of Bohemia (Czech Republic). Journal of the Czech Geological Society, 41 (3 - 4), 233 - 239." type="journal article" year="1996">Žítt 1996</bibRefCitation>
(upper Cenomanianlower Turonian,
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and
<collectingCountry id="DFA438A7FFC1FFC2F3C0FACDFDEF590D" box="[423,529,1318,1344]" name="Czech Republic" pageId="12" pageNumber="413">Bohemia</collectingCountry>
),
<taxonomicName id="60B303B4FFC1FFC2F045FACDFD5A590D" baseAuthorityName="Michelin" baseAuthorityYear="1851" box="[546,676,1318,1344]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="gastaldii">
<emphasis id="95C7A425FFC1FFC2F045FACDFD5A590D" box="[546,676,1318,1344]" italics="true" pageId="12" pageNumber="413">C. gastaldii</emphasis>
</taxonomicName>
(Middle Miocene near Turin, northwestern
<collectingCountry id="DFA438A7FFC1FFC2F6ECFACDFB3C590D" box="[1163,1218,1318,1344]" name="Italy" pageId="12" pageNumber="413">Italy</collectingCountry>
),
<taxonomicName id="60B303B4FFC1FFC2F6B2FACDFAB3590D" authorityName="Steenstrup" authorityYear="1847" box="[1237,1357,1318,1344]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="holopus">
<emphasis id="95C7A425FFC1FFC2F6B2FACDFAB3590D" box="[1237,1357,1318,1344]" italics="true" pageId="12" pageNumber="413">C. holopus</emphasis>
</taxonomicName>
(upper Maastrichtian of
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, Danian of
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and
<collectingCountry id="DFA438A7FFC1FFC2F16CFAA1FC985929" box="[779,870,1354,1380]" name="Sweden" pageId="12" pageNumber="413">Sweden</collectingCountry>
, lower Eocene of Seymour Island,
<collectingCountry id="DFA438A7FFC1FFC2F692FAA1FA905929" box="[1269,1390,1354,1380]" name="Antarctica" pageId="12" pageNumber="413">Antarctica</collectingCountry>
),
<taxonomicName id="60B303B4FFC1FFC2F7E4FAA1FF0D59C5" baseAuthorityName="Valette" baseAuthorityYear="1934" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="senessei">
<emphasis id="95C7A425FFC1FFC2F7E4FAA1FF0D59C5" italics="true" pageId="12" pageNumber="413">C. senessei</emphasis>
</taxonomicName>
(lower Santonian of Bugarach, southern
<collectingCountry id="DFA438A7FFC1FFC2F0DDFA85FCF559C5" box="[698,779,1390,1416]" name="France" pageId="12" pageNumber="413">France</collectingCountry>
),
<taxonomicName id="60B303B4FFC1FFC2F17BFA85FC7A59C5" authorityName="Jagt" authorityYear="1986" box="[796,900,1390,1416]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="12" pageNumber="413" phylum="Echinodermata" rank="species" species="vlieksi">
<emphasis id="95C7A425FFC1FFC2F17BFA85FC7A59C5" box="[796,900,1390,1416]" italics="true" pageId="12" pageNumber="413">C. vlieksi</emphasis>
</taxonomicName>
(Maastrichtian of
<collectingCountry id="DFA438A7FFC1FFC2F637FA85FB4D59C5" box="[1104,1203,1390,1416]" name="Belgium" pageId="12" pageNumber="413">Belgium</collectingCountry>
,
<collectingCountry id="DFA438A7FFC1FFC2F6D9FA85FAB459C5" box="[1214,1354,1390,1416]" name="Netherlands" pageId="12" pageNumber="413">Netherlands</collectingCountry>
),
<emphasis id="95C7A425FFC1FFC2F73CFA85FA8B59C5" box="[1371,1397,1390,1416]" italics="true" pageId="12" pageNumber="413">C.</emphasis>
<emphasis id="95C7A425FFC1FFC2F71CFA85FF3659E1" bold="true" pageId="12" pageNumber="413">sp. nov.</emphasis>
(Gale &amp; Jagt, in press; lower Campanian, north-western
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and southern
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).
</paragraph>
</subSubSection>
<subSubSection id="EFA92BBCFFC1FFC2F2A0FA5EFCB6599D" box="[199,840,1461,1488]" pageId="12" pageNumber="413" type="distribution">
<paragraph id="A70C7837FFC1FFC2F2A0FA5EFCB6599D" blockId="12.[151,1437,382,1488]" box="[199,840,1461,1488]" pageId="12" pageNumber="413">
<emphasis id="95C7A425FFC1FFC2F2A0FA5EFDFD5982" bold="true" box="[199,515,1461,1487]" pageId="12" pageNumber="413">Stratigraphic distribution.</emphasis>
Upper CenomanianRecent.
</paragraph>
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</treatment>
</document>

View file

@ -1,88 +1,88 @@
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<figureCitation id="3F8864B2FFC1FFC2F2F0F9ADFEC65A2D" box="[151,312,1606,1632]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.AC: Cyathidium chiampoensis sp. nov., AB: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), DE: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; FG: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; HJ: aboral cups of Holopus fabianii sp. nov., HI: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="12" pageNumber="413">Figures 8AC</figureCitation>
;
<tableCitation id="EA314D8CFFC1FFC2F324F9ADFE685A2D" box="[323,406,1606,1632]" captionStart="TABLE 4" captionStartId="13.[152,244,764,788]" captionText="TABLE 4. Main quantitative characters of aboral cups in Cyathidium chiampoensis sp. nov. Abbreviations: see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33229; Paratype: MGP-PD 33230." pageId="12" pageNumber="413">Table 4</tableCitation>
<tableCitation id="EA314D8CFFC1FFC2F324F9ADFE685A2D" box="[323,406,1606,1632]" captionStart="TABLE 4" captionStartId="13.[152,244,764,788]" captionTargetPageId="13" captionText="TABLE 4. Main quantitative characters of aboral cups in Cyathidium chiampoensis sp. nov. Abbreviations: see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33229; Paratype: MGP-PD 33230." httpUri="http://table.plazi.org/id/F3CC28BFFFC0FFC3F2FFFD17FAE25F75" pageId="12" pageNumber="413" tableUuid="F3CC28BFFFC0FFC3F2FFFD17FAE25F75">Table 4</tableCitation>
</paragraph>
<paragraph id="A70C7837FFC1FFC2F2F0F961FD4D5AEF" blockId="12.[151,1436,1674,1760]" box="[151,691,1674,1698]" pageId="12" pageNumber="413">
<emphasis id="95C7A425FFC1FFC2F2F0F961FEF05AEF" bold="true" box="[151,270,1674,1698]" inLineHeading="true" pageId="12" pageNumber="413" reason="1">Synonymy.</emphasis>
@ -120,14 +120,24 @@ Frisone
<subSubSection id="EFA92BBCFFC1FFC2F2F0F942FE905AAD" pageId="12" pageNumber="413" type="materials_examined">
<paragraph id="A70C7837FFC1FFC2F2F0F942FE905AAD" blockId="12.[151,1436,1674,1760]" pageId="12" pageNumber="413">
<emphasis id="95C7A425FFC1FFC2F2F0F942FECA5A8C" bold="true" box="[151,308,1705,1729]" inLineHeading="true" pageId="12" pageNumber="413" reason="1">Type material.</emphasis>
<materialsCitation id="17DB726AFFC1FFC2F35CF942FE945AAD" pageId="12" pageNumber="413" specimenCode="MGP-PD 33229, MGP-PD 33230" specimenCount="2" typeStatus="holotype">
Two aboral cups including the
<materialsCitation id="17DB726AFFC1FFC2F35CF942FBF65A8C" box="[315,1032,1705,1729]" country="Italy" location="Cengio dell'Orbo quarry" municipality="Chiampo Valley" pageId="12" pageNumber="413" specimenCode="MGP-PD 33229" specimenCount="2" typeStatus="holotype">
<specimenCount id="B1B5B3BEFFC1FFC2F35CF942FE965A8D" box="[315,360,1705,1728]" count="2" pageId="12" pageNumber="413" type="generic">Two</specimenCount>
aboral cups including the
<typeStatus id="7808C695FFC1FFC2F013F942FD335A8C" box="[628,717,1705,1729]" pageId="12" pageNumber="413" type="holotype">holotype</typeStatus>
(
<specimenCode id="F715D04CFFC1FFC2F0BDF942FC7A5A8C" box="[730,900,1705,1729]" collectionCode="MGP-PD" pageId="12" pageNumber="413">MGP-PD 33229</specimenCode>
,
<figureCitation id="3F8864B2FFC1FFC2F1E9F942FC025A8C" box="[910,1020,1705,1729]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.AC: Cyathidium chiampoensis sp. nov., AB: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), DE: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; FG: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; HJ: aboral cups of Holopus fabianii sp. nov., HI: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="12" pageNumber="413">Fig. 8AB</figureCitation>
), one well-preserved
<figureCitation id="3F8864B2FFC1FFC2F1E9F942FC025A8C" box="[910,1020,1705,1729]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.AC: Cyathidium chiampoensis sp. nov., AB: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), DE: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; FG: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; HJ: aboral cups of Holopus fabianii sp. nov., HI: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="12" pageNumber="413">
Fig. 8
<collectionCode id="C1A2E0F2FFC1FFC2F1AFF942FC225A8C" box="[968,988,1705,1729]" pageId="12" pageNumber="413">A</collectionCode>
<collectionCode id="C1A2E0F2FFC1FFC2F18CF942FC025A8C" box="[1003,1020,1705,1729]" pageId="12" pageNumber="413">B</collectionCode>
</figureCitation>
),
</materialsCitation>
<materialsCitation id="17DB726AFFC1FFC2F669F942FE945AAD" pageId="12" pageNumber="413" specimenCode="MGP-PD 33230" specimenCount="1" typeStatus="paratype">
<specimenCount id="B1B5B3BEFFC1FFC2F669F942FBCD5A8D" box="[1038,1075,1705,1728]" count="1" pageId="12" pageNumber="413" type="generic">one</specimenCount>
well-preserved
<typeStatus id="7808C695FFC1FFC2F6B1F942FAD35A8C" box="[1238,1325,1705,1729]" pageId="12" pageNumber="413" type="paratype">paratype</typeStatus>
(
<specimenCode id="F715D04CFFC1FFC2F75CF942FEF75AAD" collectionCode="MGP-PD" pageId="12" pageNumber="413">MGP-PD 33230</specimenCode>
@ -156,28 +166,19 @@ Small species (D&lt;
</subSubSection>
<subSubSection id="EFA92BBCFFC1FFC2F2A0F875FAD35B91" pageId="12" pageNumber="413" type="materials_examined">
<paragraph id="A70C7837FFC1FFC2F2A0F875FD2C5BF5" blockId="12.[151,1436,1806,2012]" box="[199,722,1950,1976]" pageId="12" pageNumber="413">
<materialsCitation id="17DB726AFFC1FFC2F2A0F875FD305BF5" box="[199,718,1950,1976]" location="Lower Lutetian" pageId="12" pageNumber="413" specimenCount="1" typeStatus="holotype">
<emphasis id="95C7A425FFC1FFC2F2A0F875FE915BF5" bold="true" box="[199,367,1950,1976]" pageId="12" pageNumber="413">
<typeStatus id="7808C695FFC1FFC2F2A0F875FEFC5BF5" box="[199,258,1950,1976]" pageId="12" pageNumber="413">Type</typeStatus>
stratum.
</emphasis>
<location id="A26C2EECFFC1FFC2F311F875FDDD5BF5" LSID="urn:lsid:plazi:treatment:2F1AC921FFC1FFC3F2F0F9CAFC495E81:A26C2EECFFC1FFC2F311F875FDDD5BF5" box="[374,547,1950,1976]" name="Lower Lutetian" pageId="12" pageNumber="413">Lower Lutetian</location>
tuffite horizon
</materialsCitation>
.
<emphasis id="95C7A425FFC1FFC2F2A0F875FE915BF5" bold="true" box="[199,367,1950,1976]" pageId="12" pageNumber="413">Type stratum.</emphasis>
<geologicalTimeScale id="D593F89CFFC1FFC2F311F875FDDD5BF5" box="[374,547,1950,1976]" pageId="12" pageNumber="413">Lower Lutetian</geologicalTimeScale>
tuffite horizon.
</paragraph>
<paragraph id="A70C7837FFC1FFC2F2A0F829FAD35B91" blockId="12.[151,1436,1806,2012]" box="[199,1325,1985,2012]" pageId="12" pageNumber="413">
<materialsCitation id="17DB726AFFC1FFC2F2A0F829FAD75B91" box="[199,1321,1985,2012]" country="Italy" location="Cengio" municipality="Chiampo Valley" pageId="12" pageNumber="413" specimenCount="1" typeStatus="holotype">
<emphasis id="95C7A425FFC1FFC2F2A0F829FE985B96" bold="true" box="[199,358,1985,2012]" pageId="12" pageNumber="413">
<typeStatus id="7808C695FFC1FFC2F2A0F829FEFC5B91" box="[199,258,1986,2012]" pageId="12" pageNumber="413">Type</typeStatus>
locality.
</emphasis>
<location id="A26C2EECFFC1FFC2F30BF829FE435B91" LSID="urn:lsid:plazi:treatment:2F1AC921FFC1FFC3F2F0F9CAFC495E81:A26C2EECFFC1FFC2F30BF829FE435B91" box="[364,445,1986,2012]" country="Italy" municipality="Chiampo Valley" name="Cengio" pageId="12" pageNumber="413">Cengio</location>
dellOrbo quarry (Cava Boschetto) in the
<emphasis id="95C7A425FFC1FFC2F2A0F829FE985B96" bold="true" box="[199,358,1985,2012]" pageId="12" pageNumber="413">Type locality.</emphasis>
<location id="A26C2EECFFC1FFC2F30BF829FD7F5B91" LSID="urn:lsid:plazi:treatment:2F1AC921FFC1FFC3F2F0F9CAFC495E81:A26C2EECFFC1FFC2F30BF829FD7F5B91" box="[364,641,1986,2012]" country="Italy" municipality="Chiampo Valley" name="Cengio dell'Orbo quarry" pageId="12" pageNumber="413">Cengio dellOrbo quarry</location>
(
<locationDeviation id="B6BE6115FFC1FFC2F0F7F829FCBA5B91" box="[656,836,1986,2012]" pageId="12" pageNumber="413">Cava Boschetto</locationDeviation>
) in the
<collectingMunicipality id="4768E24DFFC1FFC2F1FFF829FBB05B91" box="[920,1102,1986,2012]" pageId="12" pageNumber="413">Chiampo Valley</collectingMunicipality>
, north-eastern
<collectingCountry id="DFA438A7FFC1FFC2F693F829FAD75B91" box="[1268,1321,1986,2012]" name="Italy" pageId="12" pageNumber="413">Italy</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -187,7 +188,7 @@ dellOrbo quarry (Cava Boschetto) in the
Aboral cups belonging to small specimens with cup diameter up to
<quantity id="604BD5D2FFC0FFC3F6BEFF7DFAC35CFC" box="[1241,1341,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.72" pageId="13" pageNumber="414" unit="mm" value="2.72">2.72 mm</quantity>
, H equal or substantially lower than D (H/D 0.42), usually D greater than Db. Quantitative characters of the type series listed in
<tableCitation id="EA314D8CFFC0FFC3F2D2FF35FEF25CB5" box="[181,268,222,248]" captionStart="TABLE 4" captionStartId="13.[152,244,764,788]" captionText="TABLE 4. Main quantitative characters of aboral cups in Cyathidium chiampoensis sp. nov. Abbreviations: see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33229; Paratype: MGP-PD 33230." pageId="13" pageNumber="414">Table 4</tableCitation>
<tableCitation id="EA314D8CFFC0FFC3F2D2FF35FEF25CB5" box="[181,268,222,248]" captionStart="TABLE 4" captionStartId="13.[152,244,764,788]" captionTargetPageId="13" captionText="TABLE 4. Main quantitative characters of aboral cups in Cyathidium chiampoensis sp. nov. Abbreviations: see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MGP-PD 33229; Paratype: MGP-PD 33230." httpUri="http://table.plazi.org/id/F3CC28BFFFC0FFC3F2FFFD17FAE25F75" pageId="13" pageNumber="414" tableUuid="F3CC28BFFFC0FFC3F2FFFD17FAE25F75">Table 4</tableCitation>
. The
<typeStatus id="7808C695FFC0FFC3F32CFF35FE4E5CB5" box="[331,432,222,248]" pageId="13" pageNumber="414" type="holotype">holotype</typeStatus>
, which is the largest specimen, has a curved aboral cup and an external surface showing 89 conspicuous parallel growth lines interrupting numerous perpendicular ridges; base smaller than D straight on one side (

View file

@ -1,88 +1,88 @@
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<mods:namePart id="BEB1E8BC86D7D3C65D643DAAE8DE6009">Gatto, Roberto</mods:namePart>
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<mods:namePart id="5827C2BB3BA34640815AA6C5B7618CFA">Eléaume, Marc</mods:namePart>
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<mods:affiliation id="A5BAD2553A7DE564AC76EC1F821386AD">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, 57 rue Cuvier, 75005 Paris, France. &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
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<mods:namePart id="4F0ACBC71208777D9B92757CB836A5FA">Eléaume, Marc</mods:namePart>
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<mods:affiliation id="14E2429CD8ADAB20073C5E33F5F24CC6">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, 57 rue Cuvier, 75005 Paris, France. &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
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<mods:namePart id="EB400CE8CF51CF819A8F5074420C2AD6">Améziane, Nadia</mods:namePart>
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<mods:affiliation id="1A822A4EBB0F8C2BDA0766EA7F5D7064">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
<mods:nameIdentifier id="5F37AC3980989CBDE65CB71C4AFD1B02" type="email">nadia.ameziane@mnhn.fr</mods:nameIdentifier>
<mods:namePart id="EA8C80EE9565A0A5F32E3CE4DFA870EA">Améziane, Nadia</mods:namePart>
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<mods:affiliation id="6F69354BD48B777568EF0ED404AB62DA">Institut de Systématique, Evolution, Biodiversité (ISYEB) UMR 7205 CNRS-MNHN-SU-EPHE-UA, Muséum national dHistoire naturelle, BP 225, 29182 Concarneau Cedex, France.</mods:affiliation>
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<treatment id="2F1AC921FFC3FFC0F2F0FCAAFB345BA2" ID-DOI="http://doi.org/10.5281/zenodo.14248391" ID-Zenodo-Dep="14248391" LSID="urn:lsid:plazi:treatment:2F1AC921FFC3FFC0F2F0FCAAFB345BA2" httpUri="http://treatment.plazi.org/id/2F1AC921FFC3FFC0F2F0FCAAFB345BA2" lastPageNumber="415" pageId="14" pageNumber="415">
<subSubSection id="EFA92BBCFFC3FFC0F2F0FCAAFD985F16" box="[151,614,833,860]" pageId="14" pageNumber="415" type="nomenclature">
@ -103,7 +103,7 @@
<paragraph id="A70C7837FFC3FFC0F2F0FC8EFE6F5F32" blockId="14.[151,614,833,896]" box="[151,401,869,896]" pageId="14" pageNumber="415">
<figureCitation id="3F8864B2FFC3FFC0F2F0FC8EFECD5FCD" box="[151,307,869,896]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1775]" captionTargetId="figure-19@15.[151,1436,181,1775]" captionTargetPageId="15" captionText="FIGURE 8. Holopodid aboral cups from Chiampo.AC: Cyathidium chiampoensis sp. nov., AB: holotype (MGP-PD 33229), A: oblique lateral view, B: opposite side, C: paratype 1 oblique lateral view (MGP-PD 33230), DE: Cyathidium sp., D: lateral view MGP-PD 31469, E: lateral view MGP-PD 31471; FG: holotype of Cyathidium gastaldii (MNHN-F-R68284), F: lateral adoral view, G: close-up of one arm articulation; HJ: aboral cups of Holopus fabianii sp. nov., HI: holotype (MGP-PD 33232), H: oblique lateral view, I: opposite side; J: lateral aboral view of juvenile paratype 1 (MGP-PD 33233); p: interradial process, r: radial ridge. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14241176" httpUri="https://zenodo.org/record/14241176/files/figure.png" pageId="14" pageNumber="415">Figures 8F, G</figureCitation>
;
<tableCitation id="EA314D8CFFC3FFC0F359FC8EFE6F5F32" box="[318,401,869,895]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." pageId="14" pageNumber="415">Table 6</tableCitation>
<tableCitation id="EA314D8CFFC3FFC0F359FC8EFE6F5F32" box="[318,401,869,895]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." httpUri="http://table.plazi.org/id/F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA" pageId="14" pageNumber="415" tableUuid="F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA">Table 6</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="EFA92BBCFFC3FFC0F2F0FC41FCD65FB2" pageId="14" pageNumber="415" type="reference_group">
@ -144,22 +144,18 @@ Non
<subSubSection id="EFA92BBCFFC3FFC0F2F0FBC5FACE589A" pageId="14" pageNumber="415" type="materials_examined">
<paragraph id="A70C7837FFC3FFC0F2F0FBC5FACE589A" blockId="14.[151,1437,1069,2032]" pageId="14" pageNumber="415">
<emphasis id="95C7A425FFC3FFC0F2F0FBC5FEB9580A" bold="true" box="[151,327,1069,1096]" pageId="14" pageNumber="415">Type material.</emphasis>
A single aboral cup (
A
<materialsCitation id="17DB726AFFC3FFC0F30DFBC6FAD2589A" collectionCode="MPURLS" country="Italy" location="Superga Hill" pageId="14" pageNumber="415" specimenCode="MPURLS NS 6/800" specimenCount="1" stateProvince="Turin" typeStatus="holotype">
single aboral cup (
<typeStatus id="7808C695FFC3FFC0F059FBC6FD5D580A" box="[574,675,1069,1095]" pageId="14" pageNumber="415" type="holotype">holotype</typeStatus>
) from the Miocene of the Superga Hill near Turin and two plaster casts are housed in the palaeontological collection of the
<collectionCode id="C1A2E0F2FFC3FFC0F172FBB9FC955821" box="[789,875,1106,1132]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="415">MNHN</collectionCode>
in Paris (MNHN-F-R68284). Several casts of this
<typeStatus id="7808C695FFC3FFC0F2F0FB9EFF0758C2" box="[151,249,1141,1167]" pageId="14" pageNumber="415" type="holotype">holotype</typeStatus>
were sent to collections outside of
<collectingCountry id="DFA438A7FFC3FFC0F0F5FB9DFD1C58DD" box="[658,738,1142,1168]" name="France" pageId="14" pageNumber="415">France</collectingCountry>
. Two of them are housed in the Palaeontological Museum of Turin University (
) from the Miocene of the Superga Hill near Turin and two plaster casts are housed in the palaeontological collection of the MNHN in Paris (MNHN-F-R68284). Several casts of this holotype were sent to collections outside of France. Two of them are housed in the Palaeontological Museum of Turin University (
<bibRefCitation id="C32205C6FFC3FFC0F306FB72FD9958F9" author="Manni, R. &amp; Pacioni, E." box="[353,615,1177,1204]" pageId="14" pageNumber="415" pagination="65 - 77" refId="ref22121" refString="Manni, R. &amp; Pacioni, E. (2021) New records of the cyrtocrinid crinoids Cyathidium gastaldii (Michelin, 1851) and Holopus spileccense (Schluter, 1878) from Eocene strata in Veneto, north-east Italy. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen, 301 (1), 65 - 77. https: // doi. org / 10.1127 / njgpa / 2021 / 0997" type="journal article" year="2021">Manni &amp; Pacioni 2021</bibRefCitation>
). The cast of an aboral cup (
<collectionCode id="C1A2E0F2FFC3FFC0F1C3FB71FBBF58F9" box="[932,1089,1178,1204]" pageId="14" pageNumber="415">MPURLS NS</collectionCode>
6/800) having nearly the same size and belonging to the Palaeontological Museum of Rome (
<specimenCode id="F715D04CFFC3FFC0F1C3FB71FB7758FE" box="[932,1161,1177,1204]" pageId="14" pageNumber="415">MPURLS NS 6/800</specimenCode>
) having nearly the same size and belonging to the Palaeontological Museum of Rome (
<bibRefCitation id="C32205C6FFC3FFC0F136FB56FC225895" author="Manni, R." box="[849,988,1213,1240]" pageId="14" pageNumber="415" pagination="211 - 218" refId="ref22041" refString="Manni, R. (2005) The non-isocrinid crinoids of the Michelotti Collection. Bollettino della Societa Paleontologica Italiana, 44 (3), 211 - 218." type="journal article" year="2005">Manni 2005</bibRefCitation>
) is also a cast of the
<typeStatus id="7808C695FFC3FFC0F6A1FB56FAD2589A" box="[1222,1324,1213,1239]" pageId="14" pageNumber="415" type="holotype">holotype</typeStatus>
) is also a cast of the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -179,7 +175,7 @@ Description of the
.
</emphasis>
Aboral cup rounded subpentagonal, with granulated external surface as indicated in Michelins diagnosis. Quantitative characters listed in
<tableCitation id="EA314D8CFFC3FFC0F142FA9AFC8359C6" box="[805,893,1393,1419]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." pageId="14" pageNumber="415">Table 6</tableCitation>
<tableCitation id="EA314D8CFFC3FFC0F142FA9AFC8359C6" box="[805,893,1393,1419]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." httpUri="http://table.plazi.org/id/F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA" pageId="14" pageNumber="415" tableUuid="F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA">Table 6</tableCitation>
. Wall relatively thick (
<quantity id="604BD5D2FFC3FFC0F6E3FA9AFAED59C1" box="[1156,1299,1393,1420]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="14" pageNumber="415" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
); interradial processes conspicuous but irregularly preserved; synarthrial articulation of radials worn, aboral ligament pit inconspicuous (
@ -192,7 +188,7 @@ Aboral cup rounded subpentagonal, with granulated external surface as indicated
<emphasis id="95C7A425FFC3FFC0F2A0F9A2FE735A2E" bold="true" box="[199,397,1609,1635]" pageId="14" pageNumber="415">Other specimen.</emphasis>
<bibRefCitation id="C32205C6FFC3FFC0F3F0F9A2FDCA5A29" author="Manni, R." box="[407,564,1609,1636]" pageId="14" pageNumber="415" pagination="211 - 218" refId="ref22041" refString="Manni, R. (2005) The non-isocrinid crinoids of the Michelotti Collection. Bollettino della Societa Paleontologica Italiana, 44 (3), 211 - 218." type="journal article" year="2005">Manni (2005)</bibRefCitation>
figured a small aboral cup (quantitative characters listed in
<tableCitation id="EA314D8CFFC3FFC0F68BF9A2FABD5A2E" box="[1260,1347,1609,1635]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." pageId="14" pageNumber="415">Table 6</tableCitation>
<tableCitation id="EA314D8CFFC3FFC0F68BF9A2FABD5A2E" box="[1260,1347,1609,1635]" captionStart="TABLE 6" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 6. Quantitative characters of the aboral cups of Cyathidium gastaldii. Measurements of MPURLS 2534 are based on text and figures in Manni (2005: fig. 4 and pl. 1 fig. 8). Abbreviations see Fig. 1A. D used as growth index. Values in mm except ratios. Holotype: MNHN-F-R68284." httpUri="http://table.plazi.org/id/F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA" pageId="14" pageNumber="415" tableUuid="F3CC28BFFFDDFFDEF2FFFF7CFCFB5CBA">Table 6</tableCitation>
) with a granulated external surface belonging to the Michelotti Collection housed in the Museum of Palaeontology of “La Sapienza” University of Rome (MPURLS 2534). He attributed it to a juvenile of
<taxonomicName id="60B303B4FFC3FFC0F64CF979FB505AE6" authorityName="Manni &amp; Pacioni" authorityYear="2021" box="[1067,1198,1681,1708]" class="Crinoidea" family="Holopodidae" genus="Cyathidium" kingdom="Animalia" order="Cyrtocrinida" pageId="14" pageNumber="415" phylum="Echinodermata" rank="species" species="gastaldii">
<emphasis id="95C7A425FFC3FFC0F64CF979FB505AE6" box="[1067,1198,1681,1708]" italics="true" pageId="14" pageNumber="415">C. gastaldii</emphasis>
@ -246,7 +242,12 @@ Boschetto (= Cengio dellOrbo quarry, Chiampo Valley, Lutetian), specimens her
<subSubSection id="EFA92BBCFFC3FFC0F2A0F83EFB345BA2" box="[199,1226,2005,2032]" pageId="14" pageNumber="415" type="distribution">
<paragraph id="A70C7837FFC3FFC0F2A0F83EFB345BA2" blockId="14.[151,1437,1069,2032]" box="[199,1226,2005,2032]" pageId="14" pageNumber="415">
<emphasis id="95C7A425FFC3FFC0F2A0F83EFEA45BA2" bold="true" box="[199,346,2005,2031]" pageId="14" pageNumber="415">Occurrence.</emphasis>
Serravallian (Middle Miocene) of Superga Hill near Turin, northwestern
<geologicalTimeScale id="D593F89CFFC3FFC0F306F83EFD455BA2" box="[353,699,2005,2031]" pageId="14" pageNumber="415">Serravallian (Middle Miocene)</geologicalTimeScale>
of
<location id="A26C2EECFFC3FFC0F086F83DFC915BA2" LSID="urn:lsid:plazi:treatment:2F1AC921FFC3FFC0F2F0FCAAFB345BA2:A26C2EECFFC3FFC0F086F83DFC915BA2" box="[737,879,2005,2032]" country="Italy" name="Superga Hill" pageId="14" pageNumber="415" stateProvince="Turin">Superga Hill</location>
near
<collectingRegion id="6577B6D5FFC3FFC0F1CBF83EFC125BA2" box="[940,1004,2005,2031]" pageId="14" pageNumber="415">Turin</collectingRegion>
, northwestern
<collectingCountry id="DFA438A7FFC3FFC0F6F6F83EFB385BA2" box="[1169,1222,2005,2031]" name="Italy" pageId="14" pageNumber="415">Italy</collectingCountry>
.
</paragraph>

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