322 lines
35 KiB
XML
322 lines
35 KiB
XML
<document id="2845F0AB56F14DBD7463DAD4E9343911" ID-DOI="10.5852/ejt.2024.947.2625" ID-ISSN="2118-9773" ID-Zenodo-Dep="13310019" ID-ZooBank="CBC9DA40-EC6D-4280-8BBC-6826D72A291E" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1723540771803" checkinUser="plazi" docAuthor="Humara-Gil, Karla J., Granja-Fernández, Rebeca, Bautista-Guerrero, Eric, Solís-Marín, Francisco A. & Rodríguez-Troncoso, Alma P." docDate="2024" docId="03F3204BFFAD823957F3B005601FFCB6" docLanguage="en" docName="EJT.2024.947.130-174.pdf" docOrigin="European Journal of Taxonomy 947" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2625/12067" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophioderma Muller & Troschel 1840" docType="key" docVersion="1" lastPageNumber="167" masterDocId="FFCA5833FF89821C574EB3146569FFB7" masterDocTitle="Delimitation of Ophioderma teres (Lyman, 1860) and Ophioderma unicolor H. L. Clark, 1940 stat. nov. (Echinodermata: Ophiuroidea), including the description of two new species" masterLastPageNumber="174" masterPageNumber="130" pageNumber="166" updateTime="1725019940605" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D831C52DA96CBA3A50C2CD77A604B988">Delimitation of Ophioderma teres (Lyman, 1860) and Ophioderma unicolor H. L. Clark, 1940 stat. nov. (Echinodermata: Ophiuroidea), including the description of two new species</mods:title>
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<mods:namePart id="1D966EF306F4D20DACECBD070355F19E">Humara-Gil, Karla J.</mods:namePart>
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<mods:affiliation id="C3FF30D7591E66904A5C7F4ABDFA8AD8">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="968FC13EE5C0EB4BD1263CD0BD0B4ED6" type="email">k.humaragil@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5A64544E0045503964A3BC912A5188B4">Granja-Fernández, Rebeca</mods:namePart>
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<mods:affiliation id="677F5F39BCFA8C9076D0BC30A5C60E4B">Postdoctoral researcher (CONAHCYT) associated with Programa de Maestría en Biosistemática y Manejo de Recursos Naturales y Agrícolas (BIMARENA) / Laboratorio de Ecología Molecular, Microbiología y Taxonomía (LEMITAX). Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Zapopan, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="570DFD2FEC5AE9605C069DCD15237A40" type="email">beckygranja@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BDCC2B1E1EE544CBF5A50679008398D5">Bautista-Guerrero, Eric</mods:namePart>
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<mods:affiliation id="0BEA65576BC42B69DA8615C646A1BC26">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="59DFE297F03E129F1A9B7F8C2523367E" type="email">eric.bautista0177@academicos.udg.mx</mods:nameIdentifier>
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<mods:namePart id="F6FC776F811829CC0698F785D96EDB87">Solís-Marín, Francisco A.</mods:namePart>
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<mods:affiliation id="DB4A48AE7C83051BEEC652D32372A1B6">Colección Nacional de Equinodermos “ Dra. Ma. Elena Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de México (UNAM), Mexico City, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="AE14DF0CA0F89733BBE329FA13BEE9D2" type="email">fasolis@cmarl.unam.mx</mods:nameIdentifier>
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<mods:namePart id="6D703E218DCCA877C0C78C0CE93B6E84">Rodríguez-Troncoso, Alma P.</mods:namePart>
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<mods:affiliation id="C7C03D9D45BACB31862BE9DCD989926D">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="6E9E26C0199500ADD9200A918B9CC176" type="email">alma.rtroncoso@academicos.udg.mx</mods:nameIdentifier>
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<mods:title id="3AA96B6C63A534252EEFD3FEF5E041FA">European Journal of Taxonomy</mods:title>
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<mods:part id="269E49204CE23C511240DE424C1A4A15">
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<mods:date id="29D2FBF9DFD036289E0EB341F4D07A8B">2024</mods:date>
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<mods:number id="85BA050157E4D1D93E60B90F5D7F1916">2024-08-06</mods:number>
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<mods:number id="C90E1ADF05685242F3B6EC9329D83B9C">947</mods:number>
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<mods:url id="82B4FB162C27FE82411A22A6411A4168">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2625/12067</mods:url>
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<mods:classification id="CCB63B94B737DD1ECCED42BB6C1725E3">journal article</mods:classification>
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<subSubSection id="C340C2D6FFAD823857F3B0056656FC9A" box="[189,831,784,813]" pageId="36" pageNumber="166" type="nomenclature">
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<paragraph id="8BE5915DFFAD823857F3B0056656FC9A" blockId="36.[189,831,784,813]" box="[189,831,784,813]" pageId="36" pageNumber="166">
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<emphasis id="B92E4D4FFFAD823857F3B0056656FC9A" bold="true" box="[189,831,784,813]" italics="true" pageId="36" pageNumber="166">
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Key to the eastern Pacific species of
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<taxonomicName id="4C5AEADEFFAD823855D4B0046656FC9A" ID-CoL="7PND2" authority="Müller & Troschel, 1840" authorityName="Muller & Troschel" authorityYear="1840" box="[666,831,784,813]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="genus">Ophioderma</taxonomicName>
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<subSubSection id="C340C2D6FFAD823857F3B0506470FBE2" pageId="36" pageNumber="166" type="discussion">
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<paragraph id="8BE5915DFFAD823857F3B0506470FBE2" blockId="36.[188,1401,836,1109]" pageId="36" pageNumber="166">
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This key serves as a reference for the identification of valid nominal species of
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<taxonomicName id="4C5AEADEFFAD82385327B0506194FCE9" authorityName="Muller & Troschel" authorityYear="1840" box="[1129,1277,836,862]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="genus">
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<emphasis id="B92E4D4FFFAD82385327B0506194FCE9" box="[1129,1277,836,862]" italics="true" pageId="36" pageNumber="166">Ophioderma</emphasis>
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</taxonomicName>
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in the EP. Before use, the following should be considered: 1) possible new undescribed species have been reported in the region, particularly in California, Baja California (Pacific side), and
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<collectingCountry id="F34DD1CDFFAD8238536DB09F6186FC12" box="[1059,1263,907,933]" name="Clipperton Island" pageId="36" pageNumber="166">Clipperton Island</collectingCountry>
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(
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<bibRefCitation id="EFCBECACFFAD823853B0B09F640EFC7F" author="Lessios & Hendler" firstAuthor="Lessios" pageId="36" pageNumber="166" refId="ref25842" refString="Lessios H. & Hendler G. 2022. Mitochondrial phylogeny of the brittle star genus Ophioderma. Scientific Reports 12: 5304. https: // doi. org / 10.1038 / s 41598 - 022 - 08944 - 0" type="journal volume" year="2022">Lessios & Hendler 2022</bibRefCitation>
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). Caution should be used when employing this key to identify specimens from these locations; 2) some diagnostic characters vary according to size (e.g., granulation extent, fragmentation of DAPs, number of arm spines) (
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<bibRefCitation id="EFCBECACFFAD82385518B0E066D0FBB8" author="Granja-Fernandez R. & Hooker Y." box="[598,953,1012,1039]" journalOrPublisher="Zootaxa" pageId="36" pageNumber="166" pagination="539 - 556" part="4766" publicationUrl="https://doi.org/10.11646/zootaxa.4766.4.2" refId="ref24899" refString="Granja-Fernandez R. & Hooker Y. 2020. Revisiting the diversity and distribution of the ophiuroids (Echinodermata: Ophiuroidea) from Peru. Zootaxa 4766: 539 - 556. https: // doi. org / 10.11646 / zootaxa. 4766.4.2" title="Revisiting the diversity and distribution of the ophiuroids (Echinodermata: Ophiuroidea) from Peru" type="journal article" year="2020">
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||
Granja-Fernández
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||
<emphasis id="B92E4D4FFFAD82385479B0E1661BFBB9" box="[823,882,1012,1038]" italics="true" pageId="36" pageNumber="166">et al.</emphasis>
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||
2020
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||
</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFCBECACFFAD82385489B0E061FFFBB8" author="Stohr" box="[967,1174,1012,1039]" etAl="et al." firstAuthor="Stohr" pageId="36" pageNumber="166" pagination="1 - 37" refId="ref26857" refString="Stohr S., Weber A. A. - T., Boissin E. & Chenuil A. 2020. Resolving the Ophioderma longicauda (Echinodermata: Ophiuroidea) cryptic species complex: five sisters, three of them new. European Journal of Taxonomy 600: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.600" type="journal article" year="2020">
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||
Stöhr
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||
<emphasis id="B92E4D4FFFAD8238535EB0E16124FBB9" box="[1040,1101,1012,1038]" italics="true" pageId="36" pageNumber="166">et al.</emphasis>
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||
2020
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||
</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFCBECACFFAD823853EBB0E06596FB85" author="Humara-Gil" etAl="et al." firstAuthor="Humara-Gil" pageId="36" pageNumber="166" pagination="365 - 384" refId="ref25718" refString="Humara-Gil K. J., Granja-Fernandez R., Bautista-Guerrero E. & Rodriguez-Troncoso A. P. 2022. Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific. Journal of Natural History 56 (5 - 8): 365 - 384. https: // doi. org / 10.1080 / 00222933.2022.2071179" type="journal article" year="2022">
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Humara-Gil
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<emphasis id="B92E4D4FFFAD82385272B0E1601EFBB9" box="[1340,1399,1012,1038]" italics="true" pageId="36" pageNumber="166">et al.</emphasis>
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2022
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||
</bibRefCitation>
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||
). Therefore, small individuals (= younger) may not have developed all the characters included in the key.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C340C2D6FFAD823957F3B769601FFCB6" lastPageId="37" lastPageNumber="167" pageId="36" pageNumber="166" type="key">
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<key id="FFCBC66CFFAD823957F3B769601FFCB6" lastPageId="37" lastPageNumber="167" pageId="36" pageNumber="166">
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<keyStep id="30AE89F8FFAD823857F3B769601EFB0D" pageId="36" pageNumber="166">
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<paragraph id="8BE5915DFFAD823857F3B769601EFB21" blockId="36.[189,1399,1148,1210]" box="[189,1399,1148,1175]" pageId="36" pageNumber="166">
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||
<keyLead id="30AB3268FFAD823857F3B769601EFB21" box="[189,1399,1148,1175]" pageId="36" pageNumber="166">
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||
1. Disc covered with flattened, brick-like granules ......................................
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||
<taxonomicName id="4C5AEADEFFAD82385372B768601EFB21" authority="Hendler, 1996" authorityName="Hendler" authorityYear="1996" box="[1084,1399,1148,1174]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="species" species="vansyoci">
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||
<emphasis id="B92E4D4FFFAD82385372B76861A1FB21" bold="true" box="[1084,1224,1148,1174]" italics="true" pageId="36" pageNumber="166">O. vansyoci</emphasis>
|
||
<bibRefCitation id="EFCBECACFFAD8238539EB768601EFB21" author="Hendler" box="[1232,1399,1148,1174]" firstAuthor="Hendler" pageId="36" pageNumber="166" pagination="319 - 338" refId="ref25250" refString="Hendler G. 1996. Echinodermata collected at Rocas Alijos. In: Schmieder R. W. (ed.) Rocas Alijos. Scientific Results from the Cordell Expeditions: 319 - 338. Springer, Dordrecht. https: // doi. org / 10.1007 / 978 - 94 - 017 - 2917 - 8 _ 27" type="book chapter" year="1996">Hendler, 1996</bibRefCitation>
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</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
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||
<paragraph id="8BE5915DFFAD8238578FB7B4601EFB0D" blockId="36.[189,1399,1148,1210]" box="[193,1399,1184,1210]" pageId="36" pageNumber="166">
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||
<keyLead id="30AB3268FFAD8238578FB7B4601EFB0D" box="[193,1399,1184,1210]" pageId="36" pageNumber="166">– Disc covered with rounded granules ................................................................................................. 2</keyLead>
|
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</paragraph>
|
||
</keyStep>
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||
<keyStep id="30AE89F8FFAD823857F3B7F5601EFAA9" pageId="36" pageNumber="166">
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<paragraph id="8BE5915DFFAD823857F3B7F5601EFB4C" blockId="36.[189,1399,1249,1311]" box="[189,1399,1249,1276]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD823857F3B7F5601EFB4C" box="[189,1399,1249,1276]" pageId="36" pageNumber="166">2. Dorsal arm plates mostly or always entire ........................................................................................ 3</keyLead>
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</paragraph>
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||
<paragraph id="8BE5915DFFAD8238578FB611601EFAA9" blockId="36.[189,1399,1249,1311]" box="[193,1399,1284,1311]" pageId="36" pageNumber="166">
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||
<keyLead id="30AB3268FFAD8238578FB611601EFAA9" box="[193,1399,1284,1311]" pageId="36" pageNumber="166">– Dorsal arm plates fragmented into two or more pieces .................................................................... 6</keyLead>
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||
</paragraph>
|
||
</keyStep>
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||
<keyStep id="30AE89F8FFAD823857F3B652601EFA7E" pageId="36" pageNumber="166">
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<paragraph id="8BE5915DFFAD823857F3B652601EFA10" blockId="36.[189,1399,1350,1482]" pageId="36" pageNumber="166">
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<keyLead id="30AB3268FFAD823857F3B652601EFA10" pageId="36" pageNumber="166">
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||
3. Dorsal arm plates mostly entire; a few can be divided into up to two pieces. Radial shields naked. Adoral shields covered with granules. Arms with transverse bands (in vivo and preserved) ............ ................................................................................................................
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<taxonomicName id="4C5AEADEFFAD82385368B698601EFA10" authority="Lutken, 1859" authorityName="Lutken" authorityYear="1859" box="[1062,1399,1420,1447]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B92E4D4FFFAD82385368B69861BAFA11" bold="true" box="[1062,1235,1420,1446]" pageId="36" pageNumber="166">
|
||
<emphasis id="B92E4D4FFFAD82385368B6986152FA11" bold="true" box="[1062,1083,1420,1446]" italics="true" pageId="36" pageNumber="166">O</emphasis>
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||
.
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||
<emphasis id="B92E4D4FFFAD82385304B69861BAFA11" bold="true" box="[1098,1235,1420,1446]" italics="true" pageId="36" pageNumber="166">panamense</emphasis>
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||
</emphasis>
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||
Lütken, 1859
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</taxonomicName>
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||
</keyLead>
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||
</paragraph>
|
||
<paragraph id="8BE5915DFFAD8238578FB6A4601EFA7E" blockId="36.[189,1399,1350,1482]" box="[193,1399,1455,1482]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD8238578FB6A4601EFA7E" box="[193,1399,1455,1482]" pageId="36" pageNumber="166">– Dorsal arm plates entire. Radial shields covered with granules. Adoral shields naked .................... 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAD823857F3B6E5601EF92F" pageId="36" pageNumber="166">
|
||
<paragraph id="8BE5915DFFAD823857F3B6E5601EF9E5" blockId="36.[189,1399,1521,1688]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD823857F3B6E5601EF9E5" pageId="36" pageNumber="166">
|
||
4. Distal genital slit with granule-bearing scales only. Disc olive green with dark pink or red (in vivo), or light brown (preserved); arms with olive green and dark pink/red (in vivo) or dark brown transverse bands (preserved) ...................................................................................
|
||
<taxonomicName id="4C5AEADEFFAD82385368B523601EF9E5" authority="Lutken, 1856" authorityName="Lutken" authorityYear="1856" box="[1062,1399,1591,1618]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B92E4D4FFFAD82385368B52361BAF9E6" bold="true" box="[1062,1235,1591,1617]" pageId="36" pageNumber="166">
|
||
<emphasis id="B92E4D4FFFAD82385368B5236152F9E6" bold="true" box="[1062,1083,1591,1617]" italics="true" pageId="36" pageNumber="166">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAD82385304B52361BAF9E6" bold="true" box="[1098,1235,1591,1617]" italics="true" pageId="36" pageNumber="166">variegatum</emphasis>
|
||
</emphasis>
|
||
Lütken, 1856
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAD8238578FB54F601EF92F" blockId="36.[189,1399,1521,1688]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD8238578FB54F601EF92F" pageId="36" pageNumber="166">– Distal genital slit with naked and granule-bearing scales. Disc brown (preserved); arms with transverse bands (in vivo and preserved) .......................................................................................... 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAD823857F3B5AB601EF8F4" pageId="36" pageNumber="166">
|
||
<paragraph id="8BE5915DFFAD823857F3B5AB601EF94B" blockId="36.[189,1399,1727,1859]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD823857F3B5AB601EF94B" pageId="36" pageNumber="166">
|
||
5. Disc dark brown with small white spots. Up to 11 arm spines ........................................................... ..........................................................................................
|
||
<taxonomicName id="4C5AEADEFFAD823854C9B5F6601EF94B" authority="Granja-Fernandez et al., 2020" authorityName="Granja-Fernandez" authorityYear="2020" box="[903,1399,1762,1789]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B92E4D4FFFAD823854C9B5F66179F94A" bold="true" box="[903,1040,1762,1789]" pageId="36" pageNumber="166">
|
||
<emphasis id="B92E4D4FFFAD823854C9B5F666F5F94B" bold="true" box="[903,924,1762,1788]" italics="true" pageId="36" pageNumber="166">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAD823854E5B5F66179F94A" bold="true" box="[939,1040,1762,1789]" italics="true" pageId="36" pageNumber="166">hendleri</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EFCBECACFFAD82385359B5F6601EF94B" author="Granja-Fernandez R. & Hooker Y." box="[1047,1399,1762,1789]" journalOrPublisher="Zootaxa" pageId="36" pageNumber="166" pagination="539 - 556" part="4766" publicationUrl="https://doi.org/10.11646/zootaxa.4766.4.2" refId="ref24899" refString="Granja-Fernandez R. & Hooker Y. 2020. Revisiting the diversity and distribution of the ophiuroids (Echinodermata: Ophiuroidea) from Peru. Zootaxa 4766: 539 - 556. https: // doi. org / 10.11646 / zootaxa. 4766.4.2" title="Revisiting the diversity and distribution of the ophiuroids (Echinodermata: Ophiuroidea) from Peru" type="journal article" year="2020">
|
||
Granja-Fernández
|
||
<emphasis id="B92E4D4FFFAD823853BCB5F76044F94B" box="[1266,1325,1762,1788]" italics="true" pageId="36" pageNumber="166">et al.</emphasis>
|
||
, 2020
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAD8238578FB412601EF8F4" blockId="36.[189,1399,1727,1859]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD8238578FB412601EF8F4" pageId="36" pageNumber="166">
|
||
– Disc orange (in vivo) or brown (preserved), with dark blotches. Up to six arm spines ..................... .....................................................................................................
|
||
<taxonomicName id="4C5AEADEFFAD8238549BB43C601EF8F4" authority="H. L. Clark, 1917" authorityName="H. L. Clark" authorityYear="1917" box="[981,1399,1832,1859]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="36" pageNumber="166" phylum="Echinodermata" rank="species" species="pentacanthum">
|
||
<emphasis id="B92E4D4FFFAD8238549BB43C61CEF8F4" bold="true" box="[981,1191,1832,1859]" pageId="36" pageNumber="166">
|
||
<emphasis id="B92E4D4FFFAD8238549BB43C6683F8F5" bold="true" box="[981,1002,1832,1858]" italics="true" pageId="36" pageNumber="166">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAD823854B7B43C61CEF8F4" bold="true" box="[1017,1191,1832,1859]" italics="true" pageId="36" pageNumber="166">pentacanthum</emphasis>
|
||
</emphasis>
|
||
H.L. Clark, 1917
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAD823857F3B47E601EF810" pageId="36" pageNumber="166">
|
||
<paragraph id="8BE5915DFFAD823857F3B47E601EF832" blockId="36.[189,1399,1898,1960]" box="[189,1399,1898,1925]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD823857F3B47E601EF832" box="[189,1399,1898,1925]" pageId="36" pageNumber="166">6. Coloration uniform brown on disc and arms .................................................................................... 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAD8238578FB49A601EF810" blockId="36.[189,1399,1898,1960]" box="[193,1399,1933,1960]" pageId="36" pageNumber="166">
|
||
<keyLead id="30AB3268FFAD8238578FB49A601EF810" box="[193,1399,1933,1960]" pageId="36" pageNumber="166">– Coloration with characteristic patterns on disc or arms .................................................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAC823957F3B21D601EFEF0" pageId="37" pageNumber="167">
|
||
<paragraph id="8BE5915DFFAC823957F3B21D601EFE94" blockId="37.[189,1399,264,327]" box="[189,1399,264,291]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC823957F3B21D601EFE94" box="[189,1399,264,291]" pageId="37" pageNumber="167">
|
||
7. Distal genital slit with granule-bearing scales only ..................................................
|
||
<taxonomicName id="4C5AEADEFFAC823953FBB21D607CFE94" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1205,1301,264,291]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="bichi" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFFAC823953FBB21D607CFE94" bold="true" box="[1205,1301,264,291]" pageId="37" pageNumber="167">
|
||
<emphasis id="B92E4D4FFFAC823953FBB21D61A3FE94" bold="true" box="[1205,1226,265,291]" italics="true" pageId="37" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAC82395397B21C607CFE94" bold="true" box="[1241,1301,264,291]" italics="true" pageId="37" pageNumber="167">bichi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FFAC82395252B21D601EFE94" box="[1308,1399,265,291]" pageId="37" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAC8239578FB239601EFEF0" blockId="37.[189,1399,264,327]" box="[193,1399,300,327]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC8239578FB239601EFEF0" box="[193,1399,300,327]" pageId="37" pageNumber="167">
|
||
– Distal genital slit with naked and granule-bearing scales .....
|
||
<taxonomicName id="4C5AEADEFFAC823954E3B238606EFEF1" authority="H. L. Clark, 1940" authorityName="H. L. Clark" authorityYear="1940" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[941,1287,300,327]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="unicolor" status="stat. nov.">
|
||
<emphasis id="B92E4D4FFFAC823954E3B2386151FEF0" bold="true" box="[941,1080,300,327]" pageId="37" pageNumber="167">
|
||
<emphasis id="B92E4D4FFFAC823954E3B23866ABFEF1" bold="true" box="[941,962,300,326]" italics="true" pageId="37" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAC8239549FB2386151FEF0" bold="true" box="[977,1080,300,327]" italics="true" pageId="37" pageNumber="167">unicolor</emphasis>
|
||
</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="EFCBECACFFAC82395332B238606EFEF1" author="Clark" box="[1148,1287,300,326]" firstAuthor="Clark" pageId="37" pageNumber="167" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FFAC82395240B239601EFEF0" box="[1294,1399,301,327]" pageId="37" pageNumber="167">stat. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAC823957F3B27A601EFE46" pageId="37" pageNumber="167">
|
||
<paragraph id="8BE5915DFFAC823957F3B27A601EFE79" blockId="37.[189,1400,366,498]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC823957F3B27A601EFE79" pageId="37" pageNumber="167">
|
||
8. Disc dark olive green to brown (in vivo), or dark gray to brown (preserved); arms with transverse bands (more subtle in preserved specimens) and parallel white bands in the median and distal arm sections ....................................................................................
|
||
<taxonomicName id="4C5AEADEFFAC8239548CB2A0601EFE79" authority="Humara-Gil et al., 2022" authorityName="Humara-Gil" authorityYear="2022" box="[962,1399,435,463]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="occultum">
|
||
<emphasis id="B92E4D4FFFAC8239548CB2A0613FFE79" bold="true" box="[962,1110,435,462]" pageId="37" pageNumber="167">
|
||
<emphasis id="B92E4D4FFFAC8239548CB2A066BEFE79" bold="true" box="[962,983,436,462]" italics="true" pageId="37" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAC823954A8B2A7613FFE79" bold="true" box="[998,1110,435,462]" italics="true" pageId="37" pageNumber="167">occultum</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EFCBECACFFAC82395313B2A0601EFE79" author="Humara-Gil" box="[1117,1399,436,463]" etAl="et al." firstAuthor="Humara-Gil" pageId="37" pageNumber="167" pagination="365 - 384" refId="ref25718" refString="Humara-Gil K. J., Granja-Fernandez R., Bautista-Guerrero E. & Rodriguez-Troncoso A. P. 2022. Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific. Journal of Natural History 56 (5 - 8): 365 - 384. https: // doi. org / 10.1080 / 00222933.2022.2071179" type="journal article" year="2022">
|
||
Humara-Gil
|
||
<emphasis id="B92E4D4FFFAC823953BCB2A16044FE79" box="[1266,1325,436,462]" italics="true" pageId="37" pageNumber="167">et al.</emphasis>
|
||
, 2022
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAC8239578FB2CC601EFE46" blockId="37.[189,1400,366,498]" box="[193,1399,471,498]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC8239578FB2CC601EFE46" box="[193,1399,471,498]" pageId="37" pageNumber="167">– Disc with specks or rings; arms without bands ................................................................................ 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAC823957F3B10D601FFDCE" pageId="37" pageNumber="167">
|
||
<paragraph id="8BE5915DFFAC823957F3B10D601EFDE1" blockId="37.[189,1399,537,634]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC823957F3B10D601EFDE1" pageId="37" pageNumber="167">
|
||
9. Disc dark brown with darker, sinuous rings on both dorsal and ventral sides. Radial shields naked .....................................................................................................................................
|
||
<taxonomicName id="4C5AEADEFFAC8239538DB128607CFDE1" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1219,1301,572,598]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFFAC8239538DB128607CFDE1" bold="true" box="[1219,1301,572,598]" pageId="37" pageNumber="167">
|
||
<emphasis id="B92E4D4FFFAC8239538DB12861B1FDE1" bold="true" box="[1219,1240,572,598]" italics="true" pageId="37" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAC823953A9B128607CFDE1" bold="true" box="[1255,1301,572,598]" italics="true" pageId="37" pageNumber="167">aija</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FFAC82395252B128601EFDE1" box="[1308,1399,572,598]" pageId="37" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAC8239578FB174601FFDCE" blockId="37.[189,1399,537,634]" box="[193,1398,607,634]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC8239578FB174601FFDCE" box="[193,1398,607,634]" pageId="37" pageNumber="167">– Disc speckled. Radial shields covered or naked ............................................................................. 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AE89F8FFAC823957F3B1B5601FFCB6" pageId="37" pageNumber="167">
|
||
<paragraph id="8BE5915DFFAC823957F3B1B5601EFD69" blockId="37.[189,1399,672,769]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC823957F3B1B5601EFD69" pageId="37" pageNumber="167">
|
||
10. Granules present on the disc and the distal part of dorsal arm plates in the proximalmost and median arm sections ....................................................................
|
||
<taxonomicName id="4C5AEADEFFAC82395431B1D0601EFD69" authority="Pineda-Enriquez et al., 2013" authorityName="Pineda-Enriquez" authorityYear="2013" box="[895,1399,708,734]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="peruanum">
|
||
<emphasis id="B92E4D4FFFAC82395431B1D06148FD69" bold="true" box="[895,1057,708,734]" pageId="37" pageNumber="167">
|
||
<emphasis id="B92E4D4FFFAC82395431B1D066FDFD69" bold="true" box="[895,916,708,734]" italics="true" pageId="37" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B92E4D4FFFAC823954EDB1D06148FD69" bold="true" box="[931,1057,708,734]" italics="true" pageId="37" pageNumber="167">peruanum</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EFCBECACFFAC82395366B1D0601EFD69" author="Pineda-Enriquez" box="[1064,1399,708,734]" etAl="et al." firstAuthor="Pineda-Enriquez" pageId="37" pageNumber="167" pagination="53 - 65" refId="ref26453" refString="Pineda-Enriquez T., Solis-Marin F. A., Hooker Y. & Laguarda-Figueras A. 2013. Ophioderma peruana, a new species of brittlestar (Echinodermata, Ophiuroidea, Ophiodermatidae) from the Peruvian coast. ZooKeys 357: 53 - 65. https: // doi. org / 10.3897 / zookeys. 357.6176" type="journal article" year="2013">
|
||
Pineda-Enríquez
|
||
<emphasis id="B92E4D4FFFAC823953BCB1D06044FD69" box="[1266,1325,708,734]" italics="true" pageId="37" pageNumber="167">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFFAC8239578FB1F3601FFCB6" blockId="37.[189,1399,672,769]" box="[193,1398,743,769]" pageId="37" pageNumber="167">
|
||
<keyLead id="30AB3268FFAC8239578FB1F3601FFCB6" box="[193,1398,743,769]" pageId="37" pageNumber="167">
|
||
– Granules limited to the disc and arm base .....................................................
|
||
<taxonomicName id="4C5AEADEFFAC8239532AB1F3601FFCB6" authority="(Lyman, 1860)" authorityName="" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[1124,1398,743,769]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="37" pageNumber="167" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFFAC8239532AB1F361A8FCB6" bold="true" box="[1124,1217,743,769]" italics="true" pageId="37" pageNumber="167">O. teres</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCBECACFFAC8239539EB1F36007FCB6" author="Lyman" box="[1232,1390,743,769]" firstAuthor="Lyman" pageId="37" pageNumber="167" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |