148 lines
16 KiB
XML
148 lines
16 KiB
XML
<document id="750FDD0646F0D5F50850AA1C9AC779BD" ID-CLB-Dataset="55466" ID-DOI="10.5281/zenodo.5169525" ID-GBIF-Dataset="4910761a-706e-4f53-a5a8-6c72379bdbc6" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169525" ID-ZooBank="DA6CBFE5-927E-45B6-9D05-69AC97AF7B76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277966057" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2017" docId="03DA133F7517B1314BC2FE6E8FAB6ABF" docLanguage="en" docName="InsectaMundi.0592.1-16.pdf" docOrigin="Insecta Mundi 2017 (592)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Agnelius Paulsen 2017, new genus" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFE36B47751BB13D4B02FFA38A0B6E57" masterDocTitle="Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="11" updateTime="1699056540890" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="D814AD06943653B08F05528063EA7A62" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="034F26DDED144F9AEA7628752E34AD19">
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<mods:title id="D9228184C1E6071986E0140DA131B8B4">Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
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<mods:name id="5BC7F0436CE653A5303EC49FE54CA54E" type="personal">
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<mods:roleTerm id="C4AE082CDE3E1BB2DACA5BDAD0A7340F">Author</mods:roleTerm>
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<mods:namePart id="189967CD2A42FF9C40068E7CF90635DD">Paulsen, M. J.</mods:namePart>
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<mods:affiliation id="A02DAEA2DD93840504AFFE1220578747">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:nameIdentifier id="BCC3A537CA6E4231B47CE2649AB9AA62" type="email">mjpaulsen@unl.edu</mods:nameIdentifier>
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<mods:typeOfResource id="0F709E03641318F47C42D6AA72B99304">text</mods:typeOfResource>
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<mods:title id="385D9BDBE1B0F8EAEF736F9F4B96AB60">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="4FA0DF63AF4D00E6150015DB6E2645F8">
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<mods:date id="F3B328D5446DA57BBE4363138EE741CA">2017</mods:date>
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<mods:number id="A9A3D3A050F39B8CB57C1B82FB2562BF">2017-12-29</mods:number>
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<mods:detail id="1C2C8381D5A803445E29B91BF8A0BD0C" type="volume">
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<mods:number id="02725ED947E59097CD3C1F52D8887113">2017</mods:number>
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</mods:detail>
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<mods:detail id="E73E6A91DC1C4B5A778B49764EFFD777" type="issue">
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<mods:number id="F2D398AFCE241B010300A3F6B48412AA">592</mods:number>
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<mods:classification id="0EB671940DB6AD42F62E2B009B23A644">journal article</mods:classification>
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<mods:identifier id="E437D4E7D7D35D3ABCE53FE9D72D877D" type="CLB-Dataset">55466</mods:identifier>
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<mods:identifier id="7D2809FFC8D8A144081E35AFF3AF6C21" type="DOI">10.5281/zenodo.5169525</mods:identifier>
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<mods:identifier id="152909FE033EAAE85224CBA35AFE9651" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03DA133F7517B1314BC2FE6E8FAB6ABF" ID-DOI="http://doi.org/10.5281/zenodo.6567180" ID-GBIF-Taxon="195537730" ID-Zenodo-Dep="6567180" LSID="urn:lsid:plazi:treatment:03DA133F7517B1314BC2FE6E8FAB6ABF" httpUri="http://treatment.plazi.org/id/03DA133F7517B1314BC2FE6E8FAB6ABF" lastPageNumber="11" pageId="12" pageNumber="11">
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<subSubSection id="C369F1A27517B1314BC2FE6E885B6FB1" box="[192,592,461,486]" pageId="12" pageNumber="11" type="nomenclature">
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<paragraph id="8BCCA2297517B1314BC2FE6E885B6FB1" blockId="12.[192,592,461,486]" box="[192,592,461,486]" pageId="12" pageNumber="11">
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<heading id="D08415457517B1314BC2FE6E885B6FB1" bold="true" box="[192,592,461,486]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
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<emphasis id="B9077E3B7517B1314BC2FE6E885B6FB1" bold="true" box="[192,592,461,486]" pageId="12" pageNumber="11">
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<taxonomicName id="4C73D9AA7517B1314BC2FE6E8BBB6FB1" ID-CoL="QT6" authority="Paulsen" authorityName="Paulsen" authorityYear="2017" box="[192,432,461,486]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9077E3B7517B1314BC2FE6E8B336FB2" bold="true" box="[192,312,461,485]" italics="true" pageId="12" pageNumber="11">Agnelius</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="A234C3407517B1314ABCFE6D885B6FB1" box="[446,592,462,486]" pageId="12" pageNumber="11" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369F1A27517B1314BC2FDB289806C7F" box="[192,907,528,553]" pageId="12" pageNumber="11" type="type_taxon">
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<paragraph id="8BCCA2297517B1314BC2FDB289806C7F" blockId="12.[192,907,528,553]" box="[192,907,528,553]" pageId="12" pageNumber="11">
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<emphasis id="B9077E3B7517B1314BC2FDB28B736C7F" bold="true" box="[192,376,528,553]" pageId="12" pageNumber="11">
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<typeStatus id="54C81C8B7517B1314BC2FDB28B0E6C7E" box="[192,261,529,553]" pageId="12" pageNumber="11">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C73D9AA7517B1314A7DFDB388BF6C7F" ID-CoL="73XVQ" authority="Kriesche" authorityName="Kriesche" authorityYear="1926" box="[383,692,528,552]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli">
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<emphasis id="B9077E3B7517B1314A7DFDB388356C7F" box="[383,574,528,552]" italics="true" pageId="12" pageNumber="11">Nigidius nageli</emphasis>
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Kriesche
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</taxonomicName>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369F1A27517B1314BC2FDF08EA56D63" pageId="12" pageNumber="11" type="description">
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<paragraph id="8BCCA2297517B1314BC2FDF08EA56D63" blockId="12.[190,1440,595,821]" pageId="12" pageNumber="11">
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<emphasis id="B9077E3B7517B1314BC2FDF08B606C3C" bold="true" box="[192,363,595,619]" pageId="12" pageNumber="11">Description.</emphasis>
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<taxonomicName id="4C73D9AA7517B1314A73FDF08BD76C3C" authorityName="Paulsen" authorityYear="2017" box="[369,476,595,619]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9077E3B7517B1314A73FDF08BD76C3C" box="[369,476,595,619]" italics="true" pageId="12" pageNumber="11">Agnelius</emphasis>
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</taxonomicName>
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are moderately sized (12.0–
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<quantity id="4C8B0FCC7517B1314831FDF789956C3B" box="[819,926,596,620]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="12" pageNumber="11" unit="mm" value="17.2">17.2 mm</quantity>
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) figuline species that lack sexually dimorphic mandibles. The mandibles of both sexes in this species are identical and are relatively simple but have a bifurcate dorsomedial tooth that is not found in other genera (
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<figureCitation id="1348BEAC7517B13148FCFD358E5E6CF8" box="[1022,1109,662,687]" captionStart="Figures 28–32" captionStartId="16.[192,302,1675,1699]" captionTargetBox="[230,1426,1030,1622]" captionTargetId="figure-48@16.[192,1440,1020,1634]" captionTargetPageId="16" captionText="Figures 28–32. Comparison of ocular canthi, females. 28) Ganelius madagascariensis. 29) G. oberndorferi. 30) G. gnamptus. 31) G. zombi. 32) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169539" httpUri="https://zenodo.org/record/5169539/files/figure.png" pageId="12" pageNumber="11">Fig. 32</figureCitation>
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). In both sexes the smooth, minutely punctate pronotum has a weak anterior tubercle, but the anterior margin is simply concave and not produced on either side of the tubercle as in
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<emphasis id="B9077E3B7517B1314841FD7A89B26CA6" box="[835,953,729,753]" italics="true" pageId="12" pageNumber="11">
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<taxonomicName id="4C73D9AA7517B1314841FD7A89BE6CA6" authorityName="Benesh" authorityYear="1955" box="[835,949,729,753]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">Ganelius</taxonomicName>
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.
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</emphasis>
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The pronotum disc is almost flat, never elevated anteriorly, and the longitudinal midline is weakly indicated by a slight depression. The male genitalia have a gradually narrowing flagellum that is less than
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<quantity id="4C8B0FCC7517B13148D9FCBE8E3B6D63" box="[987,1072,797,821]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="11" unit="mm" value="20.0">20 mm</quantity>
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in length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369F1A27517B1314BC2FCC388756AD4" pageId="12" pageNumber="11" type="discussion">
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<paragraph id="8BCCA2297517B1314BC2FCC388756AD4" blockId="12.[190,1441,863,1156]" pageId="12" pageNumber="11">
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<emphasis id="B9077E3B7517B1314BC2FCC38B486D2F" bold="true" box="[192,323,864,888]" pageId="12" pageNumber="11">Remarks.</emphasis>
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The species described as
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<taxonomicName id="4C73D9AA7517B131496DFCFC89916D20" authority="Kriesche" authorityName="Kriesche" authorityYear="1926" box="[623,922,863,887]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli">
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<emphasis id="B9077E3B7517B131496DFCFC89216D20" box="[623,810,863,887]" italics="true" pageId="12" pageNumber="11">Nigidius nageli</emphasis>
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Kriesche
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</taxonomicName>
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differs morphologically from
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<taxonomicName id="4C73D9AA7517B1314FF8FCFC8AE56DCE" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B9077E3B7517B1314FF8FCFC8F6C6D20" box="[1274,1383,863,887]" italics="true" pageId="12" pageNumber="11">Ganelius</emphasis>
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species
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</taxonomicName>
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in significant ways, and as indicated by
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<bibRefCitation id="EFE2DFD87517B13149D6FC2289796DCE" author="Fujita, H." box="[724,882,897,921]" pageId="12" pageNumber="11" refId="ref7468" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo. 472 p." type="book" year="2010">Fujita (2010)</bibRefCitation>
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its placement within
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<taxonomicName id="4C73D9AA7517B1314F7AFC228EED6DCE" authorityName="Benesh" authorityYear="1955" box="[1144,1254,897,921]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9077E3B7517B1314F7AFC228EED6DCE" box="[1144,1254,897,921]" italics="true" pageId="12" pageNumber="11">Ganelius</emphasis>
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</taxonomicName>
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is problematic. Among the
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<taxonomicName id="4C73D9AA7517B1314A48FC018BA46DED" box="[330,431,930,954]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1" phylum="Arthropoda" rank="tribe" tribe="Figulini">Figulini</taxonomicName>
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, it shares the long flagellum of the male genitalia with
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<taxonomicName id="4C73D9AA7517B1314F52FC018F146DED" box="[1104,1311,930,954]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B9077E3B7517B1314F52FC018EB56DED" box="[1104,1214,930,954]" italics="true" pageId="12" pageNumber="11">Ganelius</emphasis>
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species
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</taxonomicName>
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, as well as a trilobed conjoined clypeus and labrum. However, unlike
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<emphasis id="B9077E3B7517B1314889FC678E0A6D8B" box="[907,1025,964,988]" italics="true" pageId="12" pageNumber="11">
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<taxonomicName id="4C73D9AA7517B1314889FC6789F76D8B" authorityName="Benesh" authorityYear="1955" box="[907,1020,964,988]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">Ganelius</taxonomicName>
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,
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</emphasis>
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it lacks sexual dimorphism in the shape of the mandibles (males lacking the vertical ramus), the pronotum has a simply concave anterior margin with a weak anteromedial tubercle (vs. bisinuate margin and usually stronger tubercle), and its body is distinctly flattened and more elongate (
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<figureCitation id="1348BEAC7517B1314863FB8B89ED6A17" box="[865,998,1064,1088]" captionStart="Figures 21-25" captionStartId="15.[192,288,1875,1897]" captionTargetBox="[272,1360,229,1821]" captionTargetId="figure-38@15.[272,1360,213,1846]" captionTargetPageId="15" captionText="Figures 21-25. Lateral view, males. 21) Ganelius madagascariensis. 22) G. oberndorferi. 23) G. gnamptus. 24) G. zombi. 25) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169535" httpUri="https://zenodo.org/record/5169535/files/figure.png" pageId="12" pageNumber="11">Fig. 21–25</figureCitation>
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). The protibial dentition is different than in
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<emphasis id="B9077E3B7517B1314A23FBE98B9C6A35" box="[289,407,1098,1122]" italics="true" pageId="12" pageNumber="11">
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<taxonomicName id="4C73D9AA7517B1314A23FBE98B996A35" authorityName="Benesh" authorityYear="1955" box="[289,402,1098,1122]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">Ganelius</taxonomicName>
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,
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</emphasis>
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with large, contiguous teeth rather than isolated smaller teeth. For these reasons a new genus is erected for the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369F1A27517B1314BC2FB0C8FAB6ABF" pageId="12" pageNumber="11" type="etymology">
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<paragraph id="8BCCA2297517B1314BC2FB0C8FAB6ABF" blockId="12.[192,1440,1198,1256]" pageId="12" pageNumber="11">
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<emphasis id="B9077E3B7517B1314BC2FB0C8B576A90" bold="true" box="[192,348,1199,1223]" pageId="12" pageNumber="11">Etymology.</emphasis>
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The name, gender masculine, is formed by corruption of
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<taxonomicName id="4C73D9AA7517B1314F03FB0D8E646A91" authorityName="Benesh" authorityYear="1955" box="[1025,1135,1198,1222]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9077E3B7517B1314F03FB0D8E646A91" box="[1025,1135,1198,1222]" italics="true" pageId="12" pageNumber="11">Ganelius</emphasis>
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</taxonomicName>
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through rearranging letters to echo the name of the nearby and somewhat similar Mascarene figuline genus
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<taxonomicName id="4C73D9AA7517B1314FBBFB738FAB6ABF" authority="Burmeister." authorityName="Burmeister." box="[1209,1440,1232,1256]" class="Insecta" family="Lucanidae" genus="Agnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9077E3B7517B1314FBBFB738F0D6AB0" box="[1209,1286,1232,1255]" italics="true" pageId="12" pageNumber="11">Agnus</emphasis>
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Burmeister.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |