Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)
Author
Paulsen, M. J.
Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546
mjpaulsen@unl.edu
text
Insecta Mundi
2017
2017-12-29
2017
592
1
16
journal article
55466
10.5281/zenodo.5169525
4910761a-706e-4f53-a5a8-6c72379bdbc6
1942-1354
5169525
DA6CBFE5-927E-45B6-9D05-69AC97AF7B76
Agnelius
Paulsen
,
new genus
Type
species.
Nigidius nageli
Kriesche
, here designated.
Description.
Agnelius
are moderately sized (12.0–
17.2 mm
) figuline species that lack sexually dimorphic mandibles. The mandibles of both sexes in this species are identical and are relatively simple but have a bifurcate dorsomedial tooth that is not found in other genera (
Fig. 32
). In both sexes the smooth, minutely punctate pronotum has a weak anterior tubercle, but the anterior margin is simply concave and not produced on either side of the tubercle as in
Ganelius
.
The pronotum disc is almost flat, never elevated anteriorly, and the longitudinal midline is weakly indicated by a slight depression. The male genitalia have a gradually narrowing flagellum that is less than
20 mm
in length.
Remarks.
The species described as
Nigidius nageli
Kriesche
differs morphologically from
Ganelius
species
in significant ways, and as indicated by
Fujita (2010)
its placement within
Ganelius
is problematic. Among the
Figulini
, it shares the long flagellum of the male genitalia with
Ganelius
species
, as well as a trilobed conjoined clypeus and labrum. However, unlike
Ganelius
,
it lacks sexual dimorphism in the shape of the mandibles (males lacking the vertical ramus), the pronotum has a simply concave anterior margin with a weak anteromedial tubercle (vs. bisinuate margin and usually stronger tubercle), and its body is distinctly flattened and more elongate (
Fig. 21–25
). The protibial dentition is different than in
Ganelius
,
with large, contiguous teeth rather than isolated smaller teeth. For these reasons a new genus is erected for the species.
Etymology.
The name, gender masculine, is formed by corruption of
Ganelius
through rearranging letters to echo the name of the nearby and somewhat similar Mascarene figuline genus
Agnus
Burmeister.