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<document id="7653006BF01ED3DC87B1DF61D42F487F" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="D5298002FCF96ADC19A543AE9E9A5BFE" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Bamaroodes Gueorguiev, 2014, gen. n." docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="90" updateTime="1731807077730" updateUser="admin">
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<mods:title id="156DA0107F220AD6B9C168D2D810911D">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="4B64978036AB5FBB1AC51EA3E1F8E76E">Gueorguiev, Borislav</mods:namePart>
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<mods:classification id="0AF04E751EBA72A945E32E575CDD9945">journal article</mods:classification>
<mods:identifier id="A367DA26ECDBEA85C80FEA9B08B30BC6" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
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<treatment id="D5298002FCF96ADC19A543AE9E9A5BFE" ID-GBIF-Taxon="127864671" LSID="urn:lsid:plazi:treatment:D5298002FCF96ADC19A543AE9E9A5BFE" httpUri="http://treatment.plazi.org/id/D5298002FCF96ADC19A543AE9E9A5BFE" lastPageNumber="90" pageId="3" pageNumber="90">
<subSubSection id="84476FD3A5C4B2B2B40CCCC26FAF810C" pageId="3" pageNumber="90" type="multiple">
<paragraph id="2C37FAA204AA297EFFC422EED00282C1" pageId="3" pageNumber="90">
<pageBreakToken id="086F5494F71AFD79C9920067E319B949" pageId="3" pageNumber="90" start="start">Taxon</pageBreakToken>
classification Animalia
<taxonomicName id="4FB15613C2D0C213049B6F2907046168" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="D8054DF0C154668DA83BBFB71B1DBE8C" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="17755DB8CCE9AD5E5BDAD9429F80C6B4" pageId="3" pageNumber="90" type="nomenclature">
<paragraph id="D8E812CB509224939BAA580FE4306A0E" pageId="3" pageNumber="90">
<taxonomicName id="1104F96312E2CE3366B454560E9EB26F" ID-CoL="8KVJ2" LSID="http://zoobank.org/0209BE0A-64A2-46B9-BAFB-92BC758D1189" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="8F274CAA5B59B95A6E77BE3CFDCE8418" pageId="3" pageNumber="90" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="134B4FC3EE9D6A8ECC2ABA9923B045DC" pageId="3" pageNumber="90" type="type species">
<paragraph id="06E83861B982916FC744CF096738A906" pageId="3" pageNumber="90">Type species.</paragraph>
<paragraph id="8C27AF968702D28F07E8015DCA402591" pageId="3" pageNumber="90">
<taxonomicName id="E75D6C9D26982E84035F749874FF7228" authority="Facchini, 2011" authorityName="Facchini" authorityYear="2011" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus Facchini, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="487D5D42AC2B8382944DEBBADCB733D9" pageId="3" pageNumber="90" type="diagnosis">
<paragraph id="CE346A1E4FB9BCE03FB1DC24A8F7BBCD" pageId="3" pageNumber="90">Diagnosis.</paragraph>
<paragraph id="0B9ABA2A3F7CB8755E285BD5365CC3CC" pageId="3" pageNumber="90">
Medium-sized species (8.0-8.5 mm) for Oriental oodines; habitus semi-elongate; color
<taxonomicName id="862F955AC5BC2626C58713E1C84DC5D7" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cyaneus">cyaneus</taxonomicName>
on head and pronotum, piceocyaneus on elytra, with yellowish apical band, piceous to rufopiceous on ventral surface, scape, palpi, tibiae, and tarsomeres red-yellow to yellow; microsculpture isodiametric; integument of body glabrous and impunctate dorsally and ventrally, (only disc of head moderately punctate and striate laterally); head with conspicuously large eyes and minute tempora; frons with punctiform impressions and one pair of punctures; labrum with six setae along anterior margin; clypeus with two setae; labial palpomere glabrous; mentum tooth rounded anteriorly, bordered posteriorly; pronotum moderately transverse, without protruded angles, with anterior and posterior margins of similar width; sides of pronotum finely bordered throughout; anterolateral setae lacking, posterolateral setae present; laterobasal impressions of pronotum faint, sublinear; elytra with fine striae, 1-7 obliterated apically, 5-7 obliterated basally; stria 7 separate to apex (not joined to stria 8), scarcely noticeable apically; stria 8 deeper than other striae, moderately grooved along posterior half; parascutellar striola well-developed, long, located between suture and stria 1; intervals uniformly flat and smooth throughout; intervals 7 and 8 not fused posteriorly, interval 8 not forming ridge externally; interval 9 transformed into marginal gutter at anterior two fifths of elytra, distinct on posterior three fifths; parascutellar pore present; discal pores in interval 3 lacking; interval 7 with two subapical pores; prosternal process unbordered; marginal gutter ended before apex of elytron; mesosternum concave; metepisternum laterally coadunate with elytral epipleuron; sterna 4-5 with pair of minute ambulatory setae; apical sternum with two marginal pores in female; mesocoxa without lateral seta, with posteromedial one; metacoxa with anterior pore; mesofemur without posterior setae; metafemur glabrous or with single pore on dorsal surface; tarsomere 5 of all legs setose ventrally.
</paragraph>
</subSubSection>
<subSubSection id="A7057911BCC20394ECF972A3703B8C7A" pageId="3" pageNumber="90" type="etymology">
<paragraph id="B3795596886D122E1A44D74167A5A326" pageId="3" pageNumber="90">Etymology.</paragraph>
<paragraph id="D7B4D9B467FEDC1B6E9162B24106BD83" pageId="3" pageNumber="90">
A compound word, based on the ethnic name of the people in the region where the type species was first found, Bamar, and
<taxonomicName id="E36F6E45FEDB7F99E57079C13D883BCE" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
(for its etymology see
<bibRefCitation id="554DB3AB4DA7751A8084FA5BEE723581" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="97" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexiko." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
: 955). It is treated as a Latin masculine.
</paragraph>
</subSubSection>
<subSubSection id="2F63C531FFED04B6F2DA4C673962A137" pageId="3" pageNumber="90" type="affinities">
<paragraph id="1D755FC003F0F95CE8500B6A5549BFB5" pageId="3" pageNumber="90">Affinities.</paragraph>
<paragraph id="FD21F36F9ECE6EB0B90BFAC7500A40D3" pageId="3" pageNumber="90">
In 2011, Facchini described eight new species of
<taxonomicName id="71A83701B5A545776910C709995F1EF8" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
from the Afrotropical and Oriental region, among them
<taxonomicName id="0B6374A70C4576D524F86C745CD96222" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
from Myanmar (
<bibRefCitation id="98CC655AE70FC10BD2FE092C821E2BD7" author="Facchini, S" journalOrPublisher="Giornale Italiano di Entomologia" pageId="10" pageNumber="97" pagination="381 - 394" title="Otto nuove specie e un nuovo genere di Chlaeniini Brulle, 1834 delle Regioni Afrotropicale e Orientale (Coleoptera Carabidae, Chlaeniini)." volume="12 / 56" year="2011">Facchini 2011</bibRefCitation>
: 350-351). The author noted also that the holotype of the species has sympatrically been collected with specimens of
<taxonomicName id="BAB5CB70515104E69DE01130602BC08E" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
, the type species of the genus. Facchini differentiated the latter from the former by: 1, size of the body; 2, coloration of the integument, including the color of femora and epipleura; 3, shape of the pronotum; 4, shape and striation of the elytra; 5, punctuation of the elytral intervals; 6, chaetotaxy of the last abdominal sternum in the females; 7, distance between the joints of the claws of tarsomere 5. Examination of the paratype and another specimen of
<taxonomicName id="A2E3FC741520FB3559A9E6F8EA2587E6" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
from Thailand confirms that these distinguishing features are valid (except for the color of epipleura, which is not markedly different) and well-chosen for ready differentiation of these taxa. However, they are inadequate to give an idea of a more precise systematic position of the species, though some of the differences noted by Facchini are of generic value.
</paragraph>
<paragraph id="1EE0A89BBB63B563CD41DBE39C07DB8C" pageId="3" pageNumber="90">
Careful study ascertained significant structural differences between
<taxonomicName id="764EF2D820020DCC868F322E3B10F6BF" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
and
<taxonomicName id="FA0EE4465DF94CBFFEB87EF442E26CFD" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
(Table 2).
</paragraph>
<caption id="99BBBD73C36DFF01E82A73D81BFEA8E2" ID-Table-UUID="99BBBD73C36DFF01E82A73D81BFEA8E2" httpUri="http://table.plazi.org/id/99BBBD73C36DFF01E82A73D81BFEA8E2" pageId="3" pageNumber="90">
<paragraph id="6F22983FAFB968890AF40C0C49DA863C" pageId="3" pageNumber="90">
Table 2. Diagnostic character state combinations shown by exemplars of
<taxonomicName id="1842117E3DC47AABC51E286496BAFDA4" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
and
<taxonomicName id="95AB7157E37E660E373D242E71DA27D3" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F6864E398644592DF8BE3FAEE14B4088" pageId="3" pageNumber="90">
<table id="EF6530291B39A362E9E1C233DE1523B7" pageId="3" pageNumber="90">
<tr id="0839D3BCBBD188604436357233F45F9E" pageId="3" pageNumber="90">
<th id="D6B0D6D6492E2926CB2F617A3AD06BF0" colspan="1" pageId="3" pageNumber="90" rowspan="1">No.</th>
<th id="95478EA4E8621A83C31B55401E6C9279" colspan="1" pageId="3" pageNumber="90" rowspan="1">Characters</th>
<th id="223B10D8916EEE5A0B91FFBF76FA880D" colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName id="849C8BEC94387C4F5CC6C8EDDBAB5F2D" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
</th>
<th id="BC592B72BAB434DFA16FB22504AC2DB9" colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName id="FFA516DD4A2C2A56F2B704B7FA7BFD56" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
</th>
</tr>
<tr id="00BDCF332CAD80193D8E1B4A3FEC0A94" pageId="3" pageNumber="90">
<td id="CF844DDF3099B911982BF3C6614DC80E" colspan="1" pageId="3" pageNumber="90" rowspan="1">01</td>
<td id="FF7BD0AE3783C6B44B02E39AD175C6F1" colspan="1" pageId="3" pageNumber="90" rowspan="1">Pubescence of integument</td>
<td id="7DF06F1BD60C61D8355485ED75383914" colspan="1" pageId="3" pageNumber="90" rowspan="1">partly pubescent</td>
<td id="4C294ED74B79346532F3ECFBBF7CD86F" colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
</tr>
<tr id="CF0DBBE59D0DBA5965EE590B24B94208" pageId="3" pageNumber="90">
<td id="9107C69B83EFA2E876F8CDBBAD7A3190" colspan="1" pageId="3" pageNumber="90" rowspan="1">02</td>
<td id="8F3AA88FD68E02C60596E6C270E40E97" colspan="1" pageId="3" pageNumber="90" rowspan="1">Punctuation of integument</td>
<td id="6AE8715B5687E894C1158D0900487C6D" colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 1-3)</td>
<td id="1832BC510C19246EE33F6D2D0CAADDA3" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 13-16)</td>
</tr>
<tr id="E304700E9810539F05D45060CF59FC31" pageId="3" pageNumber="90">
<td id="E35DC8EC2541817C8BCAB8BAFE884742" colspan="1" pageId="3" pageNumber="90" rowspan="1">03</td>
<td id="39C92F64E27FEE5C175061E056E5579F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Antennomere 3</td>
<td id="AD279B482E19CDBBC5938717033EABC5" colspan="1" pageId="3" pageNumber="90" rowspan="1">with a few short and scattered setae</td>
<td id="82B084E8DBDE0C7E29490FB76B1FCC95" colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
</tr>
<tr id="A948A51A3E9F705A3D030309317E4BB6" pageId="3" pageNumber="90">
<td id="910A0522DE3CA15EC639E47F6C654C8B" colspan="1" pageId="3" pageNumber="90" rowspan="1">04</td>
<td id="48201F850331EEA9E226D9565EA088DB" colspan="1" pageId="3" pageNumber="90" rowspan="1">Mentum tooth form anteriorly</td>
<td id="B5D912B880A5DF38B560B8A03C730216" colspan="1" pageId="3" pageNumber="90" rowspan="1">truncate</td>
<td id="0D0626BC6602DFE1773966A4C450D03B" colspan="1" pageId="3" pageNumber="90" rowspan="1">rounded</td>
</tr>
<tr id="CE9B2692810D9F6E51DD1CB5B495BF6B" pageId="3" pageNumber="90">
<td id="F5944EA1D986E9E0D03E7E5B6747AD1A" colspan="1" pageId="3" pageNumber="90" rowspan="1">05</td>
<td id="251FD0F78F4713CDD2BB9CEC1CD9D5E2" colspan="1" pageId="3" pageNumber="90" rowspan="1">PW/PL</td>
<td id="F204306DC3EA46EF438E4A85BEF58501" colspan="1" pageId="3" pageNumber="90" rowspan="1">1.14-1.15 (Fig. 1)</td>
<td id="A059F3B778F540CB04035D662441D674" colspan="1" pageId="3" pageNumber="90" rowspan="1">1.24-1.37 (Fig. 13)</td>
</tr>
<tr id="EE77714E8B825484CB6AC4B51E377CBB" pageId="3" pageNumber="90">
<td id="C631B5463679C0CCF0D38F9FD9F3BB40" colspan="1" pageId="3" pageNumber="90" rowspan="1">06</td>
<td id="ED50C207C43DAB01588C47E55D0C065C" colspan="1" pageId="3" pageNumber="90" rowspan="1">Basal margin of pronotum at posterior angles</td>
<td id="0E56ACE258EB5063FE6840C8276FFA0C" colspan="1" pageId="3" pageNumber="90" rowspan="1">oblique (Fig. 1)</td>
<td id="B27F5797FE153E7AD5BEBB594340983E" colspan="1" pageId="3" pageNumber="90" rowspan="1">gradually rounded (Fig. 13)</td>
</tr>
<tr id="060FC7B25905A393F83901304BBCFFC9" pageId="3" pageNumber="90">
<td id="7FD85CD8B48308725C66D68234240D08" colspan="1" pageId="3" pageNumber="90" rowspan="1">07</td>
<td id="C50F66EC471B04138D03E4A2D7A19513" colspan="1" pageId="3" pageNumber="90" rowspan="1">Pronotum</td>
<td id="6261DD22DEF0BAA31A1AFDDE9B3D2CAA" colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered laterally and lateroapically, not bordered medioapically and basally (Fig. 1)</td>
<td id="334736F25C09388EBCDA5BC7AACEABB2" colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered throughout (Fig. 13)</td>
</tr>
<tr id="90A8A7EDA228FABB673438DB99F6DE65" pageId="3" pageNumber="90">
<td id="9FB63F79EB2634E6B841668D7217151E" colspan="1" pageId="3" pageNumber="90" rowspan="1">08</td>
<td id="5A13AD71A2CBA7208E63C0BA765683CA" colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral striae 1-8</td>
<td id="283CACA735371376B0379F80269EA201" colspan="1" pageId="3" pageNumber="90" rowspan="1">1-7 punctate, 8 impunctate (Fig. 2)</td>
<td id="2731544B78BFA4B3C53BA8714AAAFCBC" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Fig. 14)</td>
</tr>
<tr id="726BD1379C5FD754174AA0350E0C2AB8" pageId="3" pageNumber="90">
<td id="0B0C481EC3A2B590D5C91D8A7640D472" colspan="1" pageId="3" pageNumber="90" rowspan="1">09</td>
<td id="355BE19CB9157793D3AA3EA7BBBD00D5" colspan="1" pageId="3" pageNumber="90" rowspan="1">Stria 8</td>
<td id="657F4132603DD8D0F925C2084F12177B" colspan="1" pageId="3" pageNumber="90" rowspan="1">not or hardly deeper than other striae (Figs 2-3)</td>
<td id="147DE81FD010B384E23C239992F849A2" colspan="1" pageId="3" pageNumber="90" rowspan="1">evidently deeper than other striae (Figs 14-16)</td>
</tr>
<tr id="1682451D3EF4B34F63039F48947B52BC" pageId="3" pageNumber="90">
<td id="4B677207293E2FFBEAA839FB4A1AC1A2" colspan="1" pageId="3" pageNumber="90" rowspan="1">10</td>
<td id="C281E05D5B3B8166FE4531E522EB3B75" colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral intervals</td>
<td id="AAAD75B4C7B55D8D446F7749EECE6914" colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 2-3)</td>
<td id="A7ABD8FF540411B8DA8F0E9668958045" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 14-16)</td>
</tr>
<tr id="65B0F1C702A1A7D4ECFB7CEFEB3D9974" pageId="3" pageNumber="90">
<td id="8446DA3F253748E8CAD6A0121B224326" colspan="1" pageId="3" pageNumber="90" rowspan="1">11</td>
<td id="316DC5417F3E6DB1EEB292A88C3E03FD" colspan="1" pageId="3" pageNumber="90" rowspan="1">Interval 9</td>
<td id="0DFF992C821CAE64B4F57DF3AE66BC74" colspan="1" pageId="3" pageNumber="90" rowspan="1">distinct throughout</td>
<td id="345640A70BC2428FB90779E5888D192C" colspan="1" pageId="3" pageNumber="90" rowspan="1">transformed into marginal gutter along anterior 2/5 of elytra, distinct in posterior 3/5</td>
</tr>
<tr id="7A3DF24850D48224123CDC980C0E86ED" pageId="3" pageNumber="90">
<td id="471EF020DB52F8A3E98C9A2B08F65940" colspan="1" pageId="3" pageNumber="90" rowspan="1">12</td>
<td id="97758029C6C7DB6B210596040C5414B4" colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of umbilicate pores</td>
<td id="068C1E9BA2F7F42F06070675B18D0DC0" colspan="1" pageId="3" pageNumber="90" rowspan="1">23-25</td>
<td id="F7762B1FA618508B47434BC42FB2CD52" colspan="1" pageId="3" pageNumber="90" rowspan="1">15-16</td>
</tr>
<tr id="365091F0178A47C3A77FDB992BDDEBF8" pageId="3" pageNumber="90">
<td id="AAC431D31C15A7A3261721CE199054B8" colspan="1" pageId="3" pageNumber="90" rowspan="1">13</td>
<td id="F4D200014CAC1982965077A7B21B207F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Mesocoxa lateral margin</td>
<td id="0CE02AE9C0AD1068DE6092654BFD7B41" colspan="1" pageId="3" pageNumber="90" rowspan="1">with one long seta</td>
<td id="4006BCDE74B3D16057410A4B1B5843B6" colspan="1" pageId="3" pageNumber="90" rowspan="1">without seta</td>
</tr>
<tr id="21A47FA336DD66E0064CDAFAA45956B7" pageId="3" pageNumber="90">
<td id="4B3B21E381A19066A98EFE34E8B2B74C" colspan="1" pageId="3" pageNumber="90" rowspan="1">14</td>
<td id="038BDF83EE24FF28CFED820C190D044F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Joins of claws</td>
<td id="25F12DDF271070F57603BB6ED388ECC0" colspan="1" pageId="3" pageNumber="90" rowspan="1">opposite, distantly situated from each other</td>
<td id="E30D2CEB84F67A34E0CB7610383893B1" colspan="1" pageId="3" pageNumber="90" rowspan="1">nearly parallel, closely situated to each other (Figs 14-15)</td>
</tr>
<tr id="33A9DE33BD2A1007AF0642CA5CAA14E2" pageId="3" pageNumber="90">
<td id="16E862C44C2E23935A71454DE7689AF5" colspan="1" pageId="3" pageNumber="90" rowspan="1">15</td>
<td id="F4B4C0F48497DF6FA1AE223FDC8BB854" colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of pores on last sternum in female</td>
<td id="BD77F3186B7E582F9405EEBFE444F3AA" colspan="1" pageId="3" pageNumber="90" rowspan="1">four pores</td>
<td id="A00D400FCB71C11E37D442499475B910" colspan="1" pageId="3" pageNumber="90" rowspan="1">two pores</td>
</tr>
<tr id="6F051EBD26E4761104D7CBFD864420D3" pageId="3" pageNumber="90">
<td id="0E21256873336B66AB240C4E40BA3CE5" colspan="1" pageId="3" pageNumber="90" rowspan="1">16</td>
<td id="811BC5F1C518D521DFF414154160DDED" colspan="1" pageId="3" pageNumber="90" rowspan="1">Apical stylomere of ovipositor</td>
<td id="74AAD7FE57B01635292E6A01ABE400FE" colspan="1" pageId="3" pageNumber="90" rowspan="1">subtriangular, as wide as basal stylomere, with 36 long ensiform setae at dorsomedial, dorsolateral and ventral position, and with nematiform setae, shorter than ensiform setae (Figs 10-11)</td>
<td id="5AD531FD970148E9E0B6777369C256EF" colspan="1" pageId="3" pageNumber="90" rowspan="1">subelongate, nearly twice as narrow as basal stylomere, with 11 short ensiform setae at dorsomedial and dorsolateral position, and with nematiform setae, nearly twice as long as ensiform setae (Figs 17-18)</td>
</tr>
</table>
</paragraph>
<paragraph id="737BD20E1702F161E87C1DBEC2EB965E" pageId="3" pageNumber="90">
Five of the listed character states (1, 2, 3, 9, and 11) are of tribal magnitude, though one of them (i.e., attribute 9) shows transitional conditions in the two species. All the states are characteristic of
<taxonomicName id="53455AE12FA2F900AEACB3D7A7A0DB95" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
and atypical for
<taxonomicName id="E7653104505ACA30DDC9EE8CB08299D9" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
(
<bibRefCitation id="6B33DE063D18435936FCCB5B08CD8C5E" author="Jeannel, R" journalOrPublisher="Office de la Recherche scientifique colonial, Paris" pageId="10" pageNumber="97" pagination="767 - 1146" title="Coleopteres carabiques de la region Malgache (troisieme partie). Faune de l'Empire Francais" year="1949 a">Jeannel 1949a</bibRefCitation>
,
<bibRefCitation id="60AD40C807F3B9096C51831D1939C4E2" author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
). Point 11 exhibits a unique condition in
<taxonomicName id="EE6C56BEF37564004B95E0B1110C0F5D" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
that is hitherto unknown in the two tribes. However, a complete character state transformation of this feature occurs in the
<taxonomicName id="1ECBA5DA689F71EF01A40E831B9CE79B" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Except for
<taxonomicName id="91D93B085A5460911FCC793C16D9D6F6" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="CC2B322DC0B6A3823A4DD4412D5048B3" rank="genus">gen. n.</taxonomicNameLabel>
, all other examined oodines possess a ninth interval of elytra completely transformed into a marginal gutter throughout. In my view, this special feature may be morphological evidence for transition between the two tribes, but this needs further examination. The losses of the mesocoxa lateral seta (point 13) and parallel position of the joints of the tarsal claws (attribute 14) are conditions hitherto not found together in the aforementioned tribes. Among the
<taxonomicName id="76900E41CF6D72FCFBC6D0F0F595B6A3" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, the three character states occur together only in
<taxonomicName id="97A7E5397758D8E6E2BE3DBCA3213327" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
and
<taxonomicName id="0FC39F3DA36BA5F04A1C733DCD9CB25A" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Thaioodes</taxonomicName>
<taxonomicNameLabel id="0AF9831B2D531F3D08B3DADC3FE253AF" rank="genus">gen. n.</taxonomicNameLabel>
<taxonomicName id="132E4EC9B034652ADD9E6958188085A4" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="piceus" status="sp. nov.">piceus</taxonomicName>
<taxonomicNameLabel id="20B812D4196D84FB080DF4F3BE702D9B" rank="species">sp. n.</taxonomicNameLabel>
I believe that they have arisen as a consequence of a specific adaptation and survival strategy for an aquatic manner of living. The remaining character states (i.e., 4, 5, 6, 7, 8, 10, 12, 15, and 16) demonstrate marked differences of a grade higher than the grade usual for species from one and the same genus. I treat each of these nine differences as of generic significance.
</paragraph>
<paragraph id="F8A67C14C08100DD64E0025DE1656F95" pageId="3" pageNumber="90">
In conclusion, there are quite a number of considerable morphological differences between
<taxonomicName id="8B2C3E6F40A0F22A8CFB6D73C3A16242" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
and
<taxonomicName id="FF7D639CB591B24A84C1AA149CFD18A6" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
, which taken together are cause for the separation of the latter in a separate genus. That is why
<taxonomicName id="557C5D2678CFDCE9E2BCDADBACB27162" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="4406F75A4E34A0CB0711FDF6BF335110" rank="genus">gen. n.</taxonomicNameLabel>
is proposed to accommodate this species. The new taxon does not share main tribal characters of chlaeniines but shows important similarities with oodines.
<taxonomicName id="165BF415B867F52F2CEA7717DAE8BF51" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Bamaroodes cyaneus</taxonomicName>
is therefore removed to a new tribal placement within
<taxonomicName id="B02EAA7A34134B58E2F523B7A0BA24A1" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. The relationships of the new genus are discussed below (see
<normalizedToken id="C406DC350D80233E43698550294B2BE5" originalValue="‘Discussion’">'Discussion'</normalizedToken>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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