Add updates up until 2024-11-17 01:31:41

This commit is contained in:
ggserver 2024-11-17 01:37:45 +00:00
parent 4a0056d11b
commit 077617f2e5
9 changed files with 1065 additions and 980 deletions

View file

@ -1,118 +1,125 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864667" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="1682DAE484E103570B6BE6340FCAF0FF" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Hololeius LaFerte-Senectere 1851" docType="treatment" docVersion="5" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="BF9755C3827F2D976610BFC83E22371A" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="1682DAE484E103570B6BE6340FCAF0FF" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Hololeius LaFerte-Senectere 1851" docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="88" updateTime="1731807077730" updateUser="admin">
<mods:mods id="C1D5306F4F0E54C0E681D5CF7A4F1495" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E79EEDBF37961847794E2CF64EA1B013">
<mods:title id="76D03CBDD6760731E71421969A03C797">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="7A1EA06F73FF4DF7CFDFDB742546AA66" type="personal">
<mods:role id="659385E991864B8D017A3D68582F609C">
<mods:roleTerm id="483BF925E7F68ECAB3A0A38982E9617B">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="FA2609951601FCE8A1BE6CCF41BEB239">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="B1196990C7868611696A2B7CA235EF74">text</mods:typeOfResource>
<mods:relatedItem id="89B255F214CFB928ABCABDC3B9D6338B" type="host">
<mods:titleInfo id="DDD65D9D35B7BC3A2F727E785052047F">
<mods:title id="B49732B469D9AB59A78D33C017C77417">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="330049B9245A23C1D45B6011AF566578">
<mods:date id="B872ACE91EF9CD8368401FF8554B7B64">2014</mods:date>
<mods:detail id="3C72147DF5630A2A7C01CD6DBF8D8AC6" type="volume">
<mods:number id="971E06B44F048B0363EC37B50A158D5C">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="98AE7CBC6AD775FF66F65D140C99F881" type="issue">
<mods:number id="37A54BEE15DFEC67302239D5B1BB0C7B">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="A8A85CF0B2543AC8D1AF6E601B5E28D9" unit="page">
<mods:start id="E6208CA5FAB1B5D2D8E1347078EF72AF">87</mods:start>
<mods:end id="0906ED1199254C18406B599675CC21C5">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="D798FE87812020E206BBBBA10F109A0E">
<mods:url id="19D063D035F0188BCCCA119D71740EF2">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="96CE862ACEAE1B14291EA270390ACE46">journal article</mods:classification>
<mods:identifier id="199D983394F5F5DB4A1AE85C0DF878F8" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="7EFF693D911B31B919E206E2228E5FD2" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864667" LSID="urn:lsid:plazi:treatment:1682DAE484E103570B6BE6340FCAF0FF" httpUri="http://treatment.plazi.org/id/1682DAE484E103570B6BE6340FCAF0FF" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="multiple">
<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
<treatment id="1682DAE484E103570B6BE6340FCAF0FF" ID-GBIF-Taxon="127864667" LSID="urn:lsid:plazi:treatment:1682DAE484E103570B6BE6340FCAF0FF" httpUri="http://treatment.plazi.org/id/1682DAE484E103570B6BE6340FCAF0FF" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection id="C3FE913B4EDD3345F38811AC82E47791" pageId="1" pageNumber="88" type="multiple">
<paragraph id="4C1D74AC8847C6DFF1D1153BAEE31232" pageId="1" pageNumber="88">
Taxon classification Animalia
<taxonomicName id="BAA2EC7136C487E0C159749BE3DD615E" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="B37921225111F7D571031D67F89284A4" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
<subSubSection id="C635E7D01C7A880315D65651CEA5BDF6" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="D9D8E9C33D8C204E2FFD8953D4C606E9" pageId="1" pageNumber="88">
<taxonomicName id="C85C0C65E03B1DD00F285615E367730F" ID-CoL="8KX8N" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
Hololeius
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
<normalizedToken id="937630C5ECBD9A114B4D2BDBE2C438F7" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
, 1851
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="type species">
<paragraph pageId="1" pageNumber="88">Type species.</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus</taxonomicName>
Dejean, 1826
<subSubSection id="5A487C7C67F0B988F74C25A582CC2AC6" pageId="1" pageNumber="88" type="type species">
<paragraph id="5D866AEED83FFA02FFFCE91BE82FFDE9" pageId="1" pageNumber="88">Type species.</paragraph>
<paragraph id="00A745662B2276A2AD987B371100FD1A" pageId="1" pageNumber="88">
<taxonomicName id="B709B1E0256672F77A420E635B702999" authority="Dejean, 1826" authorityName="Dejean" authorityYear="1826" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus Dejean, 1826</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="diagnosis">
<paragraph pageId="1" pageNumber="88">Diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="88">Small to medium-sized specimens (11-13 mm) for Oriental chlaeniines, with the following characters: habitus semi-elongate; color mostly olivaceous; integument punctate and pubescent, pubescence denser on sides of elytra and on sides of abdomen, less developed, scattered or lacking on rest of body; head with conspicuously large eyes and minute tempora; penultimate segment of labial palpomere without setae; antennomeres 1-3 lighter than other antennomeres, scape and pedicel glabrous, antennomere 3 with a few fine and scattered setae; pronotum subquadrate, without protruded angles, with basal margin laterally oblique towards posterior angle; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; striae 1-7 of elytra punctiform; striae 5-7 obliterated anteriorly; stria 8 linear, somewhat more impressed than other striae; mesocoxa with two long setae, one lateral seta and one posteromedial seta; sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae; last abdominal sternum with two pores in male, four pores in female; male protarsomeres 1-3 longer than wide and with protarsomere 1 longer than each of following two protarsomeres separately; tarsomere 5 of all legs setose ventrally.</paragraph>
<subSubSection id="1D51AC8E96C4F388135A60E83C23C543" pageId="1" pageNumber="88" type="diagnosis">
<paragraph id="48906EDEA69F7F54A3E0BEEA90C240A8" pageId="1" pageNumber="88">Diagnosis.</paragraph>
<paragraph id="8966415A3C2FF9919EC1A9CA3A676686" pageId="1" pageNumber="88">Small to medium-sized specimens (11-13 mm) for Oriental chlaeniines, with the following characters: habitus semi-elongate; color mostly olivaceous; integument punctate and pubescent, pubescence denser on sides of elytra and on sides of abdomen, less developed, scattered or lacking on rest of body; head with conspicuously large eyes and minute tempora; penultimate segment of labial palpomere without setae; antennomeres 1-3 lighter than other antennomeres, scape and pedicel glabrous, antennomere 3 with a few fine and scattered setae; pronotum subquadrate, without protruded angles, with basal margin laterally oblique towards posterior angle; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; striae 1-7 of elytra punctiform; striae 5-7 obliterated anteriorly; stria 8 linear, somewhat more impressed than other striae; mesocoxa with two long setae, one lateral seta and one posteromedial seta; sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae; last abdominal sternum with two pores in male, four pores in female; male protarsomeres 1-3 longer than wide and with protarsomere 1 longer than each of following two protarsomeres separately; tarsomere 5 of all legs setose ventrally.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="taxonomic position">
<paragraph pageId="1" pageNumber="88">Taxonomic position.</paragraph>
<paragraph pageId="1" pageNumber="88">
<bibRefCitation author="LaFerte-Senectere, F de" journalOrPublisher="Annales de la Societe Entomologique de France (2 e serie)" pageId="11" pageNumber="98" pagination="209 - 294" title="Revision de la tribu des patellimanes de Dejean, Coleopteres pentameres de lades carabiques." url="http://www.biodiversitylibrary.org/part/16123/summary" volume="9" year="1851">
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
<subSubSection id="49212A05412B699925D1F6814654CCE5" pageId="1" pageNumber="88" type="taxonomic position">
<paragraph id="4651C0BEEB68BE734959BE4B554D2466" pageId="1" pageNumber="88">Taxonomic position.</paragraph>
<paragraph id="8253452DE8DD3E58411F80EB093FE742" pageId="1" pageNumber="88">
<bibRefCitation id="F42D458FE13931644F48904474F603ED" author="LaFerte-Senectere, F de" journalOrPublisher="Annales de la Societe Entomologique de France (2 e serie)" pageId="11" pageNumber="98" pagination="209 - 294" title="Revision de la tribu des patellimanes de Dejean, Coleopteres pentameres de lades carabiques." url="http://www.biodiversitylibrary.org/part/16123/summary" volume="9" year="1851">
<normalizedToken id="F819DB733F08BFA5890AD9EC16A38DDC" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
(1851
</bibRefCitation>
: 274) proposed
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<taxonomicName id="1FEE717BF8408ADFDBCF3A93F9F018F9" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
for
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus</taxonomicName>
Dejean, 1826, due to: 1, pronotum at the base narrower than the base of elytra; 2, lack of pubescence; 3, antennomere 3 not longer than subsequent antennomeres. He placed the genus in tribe
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
, 1851, concluding:
<normalizedToken originalValue="“Cest-à-dire">&quot;C'est-a-dire</normalizedToken>
<normalizedToken originalValue="quà">qu'a</normalizedToken>
<normalizedToken originalValue="lexception">l'exception</normalizedToken>
<taxonomicName id="4D14E8CEE17B69482D63C9346196BCE5" authority="Dejean, 1826" authorityName="Dejean" authorityYear="1826" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus Dejean, 1826</taxonomicName>
, due to: 1, pronotum at the base narrower than the base of elytra; 2, lack of pubescence; 3, antennomere 3 not longer than subsequent antennomeres. He placed the genus in tribe
<taxonomicName id="07BC17F78D0977AE804C8549156B65FA" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">
Oodini
<normalizedToken id="51E6D7326A51BC1379E168A681ACA5FA" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
, 1851
</taxonomicName>
, concluding:
<normalizedToken id="5C001CB9FBE6D085D655F55FC4AC8B81" originalValue="“Cest-à-dire">&quot;C'est-a-dire</normalizedToken>
<normalizedToken id="BA22918733807F44C93EA74DF5280025" originalValue="quà">qu'a</normalizedToken>
<normalizedToken id="254342AA1ABDB7DD556916DFAE6172EC" originalValue="lexception">l'exception</normalizedToken>
de la largeur du corselet, cet insecte
<normalizedToken originalValue="réunit">reunit</normalizedToken>
<normalizedToken id="100212EA79CBA6AAC50F53B53F1792A9" originalValue="réunit">reunit</normalizedToken>
tout les
<normalizedToken originalValue="charactèrs">characters</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken originalValue="laide">l'aide</normalizedToken>
<normalizedToken id="7F5980D4808CB67D12049B001854B31E" originalValue="charactèrs">characters</normalizedToken>
<normalizedToken id="B08F45A94D459573FFAC9F41C642F5BA" originalValue="à">a</normalizedToken>
<normalizedToken id="6570CE2EB9D24DE3DFB8826FFBF76702" originalValue="laide">l'aide</normalizedToken>
desquels nous avons
<normalizedToken originalValue="séparé">separe</normalizedToken>
<normalizedToken id="EFCECE6A185AC47B3AA5E37E7B548196" originalValue="séparé">separe</normalizedToken>
les Oodites des
<normalizedToken originalValue="Chlaœnides.”">Chlaoenides.&quot;</normalizedToken>
<normalizedToken id="553C7137C8852C7AD4FBA04FE1D9C7E6" originalValue="Chlaœnides.”">Chlaoenides.&quot;</normalizedToken>
. In terms of the present knowledge, I can state that the first and third characters are true, but not the second one. When examined it was found that
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<taxonomicName id="5F8F8B5EFB398C8E144779D1F77D83EE" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
has the integument partially and diffusely punctate and pubescent, both dorsally and ventrally. This state is opposed to one of the basic features of the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="9C2E9439B5454EF1CA781537FAF040A9" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, namely the lack of pubescence and lack of extensive punctation of the integument (
<bibRefCitation author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
<bibRefCitation id="0167231B1C4122A98FCF8827C329F8E4" author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
: 448). Hence, the three characters
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
<normalizedToken id="2417E4CDCE414310CA582CFFA3896580" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
mentioned are typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="96991110C4FF72CC1E5C6498093DE269" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. The length and ratio of antennomere 3 to the following antennomeres is not of tribal significance.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<paragraph id="0547C711A3C700F1A45D7A1AD7DBE938" pageId="1" pageNumber="88">
<taxonomicName id="97773DCCC047C182CE54E929464BB637" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
is usually placed in the beginning of the genera of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="34D27C5BE87138CADD428574E185B9A0" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
(
<bibRefCitation author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="3C0797117AE4A0047CD584BAFF41016F" author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="1998">Lorenz 1998</bibRefCitation>
,
<bibRefCitation author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">2005</bibRefCitation>
<bibRefCitation id="8BA1C2D4019BDB88EB25E9E0ED9295BF" author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">2005</bibRefCitation>
), presumably because of some odd features that it possesses. Actually, the genus combines characters distinctive for either one or the other tribe and the importance of each is discussed below (see
<normalizedToken originalValue="Affinities">'Affinities'</normalizedToken>
<normalizedToken id="D2CB4519D2FC62E1B737A19FBCCE0906" originalValue="Affinities">'Affinities'</normalizedToken>
under
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n.,
<normalizedToken originalValue="Discussion">'Discussion'</normalizedToken>
<taxonomicName id="6DB58F4FE315024FB9B87FD4BD902F27" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="25DEB8619171D7EB8B290F79962E7643" rank="genus">gen. n.</taxonomicNameLabel>
,
<normalizedToken id="C262250E3A71C0AF2A80DF49E45E67C8" originalValue="Discussion">'Discussion'</normalizedToken>
).
</paragraph>
</subSubSection>

View file

@ -1,97 +1,99 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864668" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="22552D11258D6BD17CA2A1A2BCF7963C" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Hololeius ceylanicus Nietner 1856" docType="treatment" docVersion="5" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="3A16069E52381BECD59CC726CEBFB63D" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="22552D11258D6BD17CA2A1A2BCF7963C" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Hololeius ceylanicus Nietner 1856" docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="88" updateTime="1731807077730" updateUser="admin">
<mods:mods id="943ADB1B170CE89A5C0B9B9E3095F19E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48CA4415E721918993E6BC7FD2D601DC">
<mods:title id="5988B4D0CA4875C6D0F88A6EF629D4F4">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="19BF59498DF68314365767FD9A1B24D9" type="personal">
<mods:role id="046743E615E4CC73C082B570ABB0AA9D">
<mods:roleTerm id="E3DDB0829537F3D15BF666CE099CC249">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="62CB2C4E08CC49F6B08557D6488A23A7">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="A92C3C15E8B6FC3B48F9AF725304CCAB">text</mods:typeOfResource>
<mods:relatedItem id="5BA6ECD20C2563A965D55AD119CE7A2A" type="host">
<mods:titleInfo id="2E8B94B5282A8D811929E6548AE21417">
<mods:title id="4141ECFA2714615240E0C0D4EE554276">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="A1EA9731D3635D1693F9440699C2F8AE">
<mods:date id="AC8612DCF68CBC0C3E80B300A4232503">2014</mods:date>
<mods:detail id="73ED228C190101357755BE1BE519DA4B" type="volume">
<mods:number id="0648A1C8863D4AB4120001B9141B3A2D">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="2A0AD78114CE87764E2263AE4D071E82" type="issue">
<mods:number id="237162D97791A4DC0D552955990900A9">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="2E18FA6A8AA40E5D27711E845BE7E5D7" unit="page">
<mods:start id="992F587323BC6541CC922BF6AF0327C5">87</mods:start>
<mods:end id="E051D8856AE72AF8C36549D7C4C1C93F">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="AE147D96A5E434C0D981D187681AA66E">
<mods:url id="70CB51A0AD9823740AD7CFC78D7E38EB">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="0D1F5222FA58C6DABC841EDBB360D7F2">journal article</mods:classification>
<mods:identifier id="4561673250F647BFE69CBE6681FD5546" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="9FCB34FA9FE130B02F9714F1B346978D" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864668" LSID="urn:lsid:plazi:treatment:22552D11258D6BD17CA2A1A2BCF7963C" httpUri="http://treatment.plazi.org/id/22552D11258D6BD17CA2A1A2BCF7963C" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="multiple">
<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
<treatment id="22552D11258D6BD17CA2A1A2BCF7963C" ID-GBIF-Taxon="127864668" LSID="urn:lsid:plazi:treatment:22552D11258D6BD17CA2A1A2BCF7963C" httpUri="http://treatment.plazi.org/id/22552D11258D6BD17CA2A1A2BCF7963C" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection id="4286D4E4EE64FB60D8045AA7DEAFCC1F" pageId="1" pageNumber="88" type="multiple">
<paragraph id="B1A9923F111925421784E6AF70913EF4" pageId="1" pageNumber="88">
Taxon classification Animalia
<taxonomicName id="A75432069A6B2C85C498AFD2E1D3CCBA" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="1E9446679FB16FF5887A076C88F8F63C" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Nietner, 1856" authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus (Nietner, 1856)</taxonomicName>
<subSubSection id="A565A13785E441B620BCFBF066486679" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="34A7745922033E0064FCB7EDFF06B9AD" pageId="1" pageNumber="88">
<taxonomicName id="D522C3F657A128C517657DE8A6DE3ACC" ID-CoL="3M8YG" authority="(Nietner, 1856)" authorityName="Nietner" authorityYear="1856" baseAuthorityName="Nietner" baseAuthorityYear="1856" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus (Nietner, 1856)</taxonomicName>
Figs 1-11
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="reference_group">
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus</taxonomicName>
Dejean, 1826: 341 [nec Schrank, 1781]
<subSubSection id="163F99C43693EFDE6A64D5FA96D9B6F6" pageId="1" pageNumber="88" type="reference_group">
<paragraph id="A37C9A55B6694C21A59E346A84BD55A1" pageId="1" pageNumber="88">
<taxonomicName id="8E1E75E98C845F62A566D49A3884F120" authority="Dejean, 1826: 341" authorityName="Dejean" authorityPageNumber="341" authorityYear="1826" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus Dejean, 1826: 341</taxonomicName>
[nec Schrank, 1781]
</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Chlaenius ceylanicus</taxonomicName>
Nietner, 1856: 385
<paragraph id="A3C0BE166B2F381ECE0EE18032B05F5B" pageId="1" pageNumber="88">
<taxonomicName id="9D70D76E756BCAA0C3653CA0C3253A8C" authority="Nietner, 1856: 385" authorityName="Nietner" authorityPageNumber="385" authorityYear="1856" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Chlaenius ceylanicus Nietner, 1856: 385</taxonomicName>
</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Chaudoir, 1857" authorityName="Chaudoir" authorityYear="1857" class="Insecta" family="Carabidae" genus="Hololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hololius punctulatus subsp. punctulatus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="punctulatus" subSpecies="punctulatus" uncertainty-rank="sic!">Hololius (sic!) punctulatus</taxonomicName>
Chaudoir, 1857: 10
<paragraph id="8812252FD04E6633908D9ACA9F23C587" pageId="1" pageNumber="88">
<taxonomicName id="BF0F73D1666277CBA4C5711D67309579" authority="Chaudoir, 1857: 10" authorityName="Chaudoir" authorityPageNumber="10" authorityYear="1857" class="Insecta" family="Carabidae" genus="Hololius" higherTaxonomySource="GBIF,CoL" isUncertain="true" kingdom="Animalia" lsidName="Hololius punctulatus subsp. punctulatus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="punctulatus" uncertainty-rank="sic!">Hololius (sic!) punctulatus Chaudoir, 1857: 10</taxonomicName>
</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Poecilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Poecilus ornatus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ornatus">Poecilus ornatus</taxonomicName>
Tryon, 1890: 109.
<paragraph id="89A23693627E44967A46659D0FC3BE10" pageId="1" pageNumber="88">
<taxonomicName id="0D3B7D34CF969376B299CCF2F2DE6A75" authority="Tryon, 1890: 109" authorityName="Tryon" authorityPageNumber="109" authorityYear="1890" class="Insecta" family="Carabidae" genus="Poecilus" kingdom="Animalia" lsidName="Poecilus ornatus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ornatus">Poecilus ornatus Tryon, 1890: 109</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="material examined">
<paragraph pageId="1" pageNumber="88">Material examined.</paragraph>
<paragraph pageId="1" pageNumber="88">1♂, 1♀, Indonesia, Bali, south shore of Danau Buyan Lake, 1250 m, 11.VII.1991, Balke &amp; Hendrich leg. (NMNHS).</paragraph>
<subSubSection id="D5D3E50CC50126B122E3166FAA65D0F2" pageId="1" pageNumber="88" type="material examined">
<paragraph id="5EC79D2BFE9C11E04A05991684602BFB" pageId="1" pageNumber="88">Material examined.</paragraph>
<paragraph id="4BE0BAF37937FEB7C687146CD3C2548E" pageId="1" pageNumber="88">1♂, 1♀, Indonesia, Bali, south shore of Danau Buyan Lake, 1250 m, 11.VII.1991, Balke &amp; Hendrich leg. (NMNHS).</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="diagnosis">
<paragraph pageId="1" pageNumber="88">Diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="88">Same as the generic diagnosis.</paragraph>
<subSubSection id="178DFE6002C22D34329BD4A9E5C95328" pageId="1" pageNumber="88" type="diagnosis">
<paragraph id="EC5048BE24C6722CFBB016D0BFE35261" pageId="1" pageNumber="88">Diagnosis.</paragraph>
<paragraph id="70BAFB40AD5AD202DEE5E09C9F47B616" pageId="1" pageNumber="88">Same as the generic diagnosis.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="redescription">
<paragraph pageId="1" pageNumber="88">Redescription.</paragraph>
<paragraph pageId="1" pageNumber="88">Habitus. Semi-elongate (Fig. 1). Color. Uniformly olive-green on most of dorsal surface, elytra light green to coppery at apex; venter black; elytral epipleura reddish; palpi, legs, antennomeres 1-3 and base of 4 red-yellow. Microsculpture. Reduced on most of surface, isodiametric on interval 9 and on medial intervals posteriorly, transverse on proepisternum. Punctation and pubescence. Head finely and densely punctate dorsally, slightly rugose at sides and posteriorly; pronotum and elytra with punctures coarser and more scattered than punctures on head; pronotum more densely punctate laterally and basally, less densely punctate apically, with a few hairs on disc; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; prosternum, mesepisternum, metasternum, and metepisternum glabrous medially, sparsely to moderately punctate and pubescent laterally; abdominal sterna 1-2 sparsely punctate and pubescent, sterna 3-6 sparsely punctate and pubescent medially, more densely punctate and pubescent laterally. Head. Frons with punctiform impressions and one pair of setiferous punctures. Labrum with six setae along anterior margin. Clypeus with a pair of setae. Labial palpomere glabrous. Mentum tooth truncate apically, bordered posteriorly. Pronotum. Subrectangular, without protruded angles, with anterior margin slightly shorter than posterior margin, posterior margin laterally oblique towards posterior angle; margins finely bordered laterally and lateroapically, border obliterated on anterior margin medially, basal margin not bordered; anterolateral setae lacking, posterolateral setae present; two laterobasal impressions faint, sublinear. Elytra. Striae 1-7 fine, punctiform (Figs 1-3); striae 5-7 obliterated basally (Fig. 2); stria 7 perceptible to apex (Fig. 3); stria 8 linear, not or hardly deeper than other striae, well-impressed along most of its length, depressed only apically (Figs 2-3); parascutellar striola fine, punctiform like striae 1-7, located between suture and stria 1. Intervals 7 and 8 separate to apex (Fig. 3); interval 9 distinct nearly to apex (Figs 2-3). Parascutellar pore present; discal setiferous pores lacking. Ventral surface (thorax and abdomen). Intercoxal process of prosternum unbordered, glabrous and impunctate. Mesosternum profoundly concave. Metepisternum laterally coadunate with elytral epipleuron. Sternum 3 without ambulatory setae, sterna 4-5 with pair of long ambulatory setae; apical sternum with two setae in male, four in female. Legs. Relatively long and slender. Mesocoxa with two setae, one lateral and one posteromedail. Metacoxa with anterior pore; metafemur with a few short setae on dorsal surface. Tarsomere 5 setose ventrally. Male genitalia. Median lobe of aedeagus large, strongly curved laterally, with complex internal structure (Figs 4-5); basal part small, narrow, with small bulb and concave orifice in lateral aspect, curved rectangularly towards massive and broadened medial part, from there toward apical lamella gradually narrowed, and then slightly raised distally (Fig. 4); median lobe long, straight, slightly asymmetrical in dorsal aspect, with left side obtusely angled and more concave before apex and right side undulating, slightly concave in middle and before apex, apical lamella narrow proximally and widened distally (Fig. 5); inner sac with three well-chitinized, differently shaped structures: proximal paddle-like sclerite, long medial threadlike filament and small distal sclerite on the right side easily visible in dorsal aspect (Figs 4-5); parameres different in shape, right paramere elongate, thick, saddle-like, with short and oblique process internally (Figs 6-7), left paramere conchoid, with a thick, strongly chitinized and oblique process internally (Figs 8-9). Female genitalia. Ovipositor consists of valvifer and stylomere (Figs 10-11); distal margin of valvifer with 11-12 setae of varying lengths; basal stylomere conical, 1.5 times longer than apical stylomere, its ventral surface with 10 long and thin setae directed toward apical stylomere; apical stylomere subtriangular, with rounded blade, as wide as basal stylomere, provided with 36 long and well-chitinized ensiform setae (3 dorsomedial, 6 dorsolateral and 27 ventral) and with two thick nematiform setae, slightly shorter than ensiform setae.</paragraph>
<caption pageId="1" pageNumber="88">
<paragraph pageId="1" pageNumber="88">
<subSubSection id="C2A660759795707BB94BDBC3C4898831" pageId="1" pageNumber="88" type="redescription">
<paragraph id="FABAA1A33C8B66BE24E9D5A89436EC17" pageId="1" pageNumber="88">Redescription.</paragraph>
<paragraph id="D1280908F5C850709B7FC467557789BD" pageId="1" pageNumber="88">Habitus. Semi-elongate (Fig. 1). Color. Uniformly olive-green on most of dorsal surface, elytra light green to coppery at apex; venter black; elytral epipleura reddish; palpi, legs, antennomeres 1-3 and base of 4 red-yellow. Microsculpture. Reduced on most of surface, isodiametric on interval 9 and on medial intervals posteriorly, transverse on proepisternum. Punctation and pubescence. Head finely and densely punctate dorsally, slightly rugose at sides and posteriorly; pronotum and elytra with punctures coarser and more scattered than punctures on head; pronotum more densely punctate laterally and basally, less densely punctate apically, with a few hairs on disc; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; prosternum, mesepisternum, metasternum, and metepisternum glabrous medially, sparsely to moderately punctate and pubescent laterally; abdominal sterna 1-2 sparsely punctate and pubescent, sterna 3-6 sparsely punctate and pubescent medially, more densely punctate and pubescent laterally. Head. Frons with punctiform impressions and one pair of setiferous punctures. Labrum with six setae along anterior margin. Clypeus with a pair of setae. Labial palpomere glabrous. Mentum tooth truncate apically, bordered posteriorly. Pronotum. Subrectangular, without protruded angles, with anterior margin slightly shorter than posterior margin, posterior margin laterally oblique towards posterior angle; margins finely bordered laterally and lateroapically, border obliterated on anterior margin medially, basal margin not bordered; anterolateral setae lacking, posterolateral setae present; two laterobasal impressions faint, sublinear. Elytra. Striae 1-7 fine, punctiform (Figs 1-3); striae 5-7 obliterated basally (Fig. 2); stria 7 perceptible to apex (Fig. 3); stria 8 linear, not or hardly deeper than other striae, well-impressed along most of its length, depressed only apically (Figs 2-3); parascutellar striola fine, punctiform like striae 1-7, located between suture and stria 1. Intervals 7 and 8 separate to apex (Fig. 3); interval 9 distinct nearly to apex (Figs 2-3). Parascutellar pore present; discal setiferous pores lacking. Ventral surface (thorax and abdomen). Intercoxal process of prosternum unbordered, glabrous and impunctate. Mesosternum profoundly concave. Metepisternum laterally coadunate with elytral epipleuron. Sternum 3 without ambulatory setae, sterna 4-5 with pair of long ambulatory setae; apical sternum with two setae in male, four in female. Legs. Relatively long and slender. Mesocoxa with two setae, one lateral and one posteromedail. Metacoxa with anterior pore; metafemur with a few short setae on dorsal surface. Tarsomere 5 setose ventrally. Male genitalia. Median lobe of aedeagus large, strongly curved laterally, with complex internal structure (Figs 4-5); basal part small, narrow, with small bulb and concave orifice in lateral aspect, curved rectangularly towards massive and broadened medial part, from there toward apical lamella gradually narrowed, and then slightly raised distally (Fig. 4); median lobe long, straight, slightly asymmetrical in dorsal aspect, with left side obtusely angled and more concave before apex and right side undulating, slightly concave in middle and before apex, apical lamella narrow proximally and widened distally (Fig. 5); inner sac with three well-chitinized, differently shaped structures: proximal paddle-like sclerite, long medial threadlike filament and small distal sclerite on the right side easily visible in dorsal aspect (Figs 4-5); parameres different in shape, right paramere elongate, thick, saddle-like, with short and oblique process internally (Figs 6-7), left paramere conchoid, with a thick, strongly chitinized and oblique process internally (Figs 8-9). Female genitalia. Ovipositor consists of valvifer and stylomere (Figs 10-11); distal margin of valvifer with 11-12 setae of varying lengths; basal stylomere conical, 1.5 times longer than apical stylomere, its ventral surface with 10 long and thin setae directed toward apical stylomere; apical stylomere subtriangular, with rounded blade, as wide as basal stylomere, provided with 36 long and well-chitinized ensiform setae (3 dorsomedial, 6 dorsolateral and 27 ventral) and with two thick nematiform setae, slightly shorter than ensiform setae.</paragraph>
<caption id="3CBF91FB9A53A0FB1814725017AD215A" pageId="1" pageNumber="88">
<paragraph id="252D07F47F3211F6704130BB8823B8C1" pageId="1" pageNumber="88">
Figures 1-3.
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
(Niether, 1856) from Danau Buyan Lake, Bali, Indonesia, female. 1. Habitus; 2-3. Elytra (2. left anterolateral view; a - stria 8; b - interval 9; 3. left posterolateral view; a - stria 7; b - interval 9). Scale bars = 2 mm (Fig. 1); = 1 mm (Figs 2-3).
<taxonomicName id="AB312E2F1E52FABAC51FF5D3D42E70E6" authority="(Niether, 1856)" baseAuthorityName="Niether" baseAuthorityYear="1856" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus (Niether, 1856)</taxonomicName>
from Danau Buyan Lake, Bali, Indonesia, female. 1. Habitus; 2-3. Elytra (2. left anterolateral view; a - stria 8; b - interval 9; 3. left posterolateral view; a - stria 7; b - interval 9). Scale bars = 2 mm (Fig. 1); = 1 mm (Figs 2-3).
</paragraph>
</caption>
<caption pageId="1" pageNumber="88">
<paragraph pageId="1" pageNumber="88">
<caption id="FB5CE04F3105D0A3C4BEE5082E369B05" pageId="1" pageNumber="88">
<paragraph id="14F1FA138E6BDDD8ED25CD17B31B44C7" pageId="1" pageNumber="88">
Figures 4-11.
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
(Niether, 1856) from Danau Buyan Lake, Bali, Indonesia. 4-9. Male genitalia (4. median lobe of aedeagus, left lateral view; 5. median lobe of aedeagus, dorsal view; 6-7. right paramere, internal and external face; 8-9, left paramere, internal and external face); 10-11. Left ovipositor, ventral view (10. photograph; 11. line drawing; as - apical stylomere; bs - basal stylomere; dles - dorsolateral ensiform setae; dmes - dorsomedial ensiform setae; ns - nematiform setae; ves - ventral ensiform setae; vf - valvifer). Scale bars = 0.3 mm (Figs 4-9); = 0.1 mm (Figs 10-11).
<taxonomicName id="0DDC99F3952DE30CC7130858AF135564" authority="(Niether, 1856)" baseAuthorityName="Niether" baseAuthorityYear="1856" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus (Niether, 1856)</taxonomicName>
from Danau Buyan Lake, Bali, Indonesia. 4-9. Male genitalia (4. median lobe of aedeagus, left lateral view; 5. median lobe of aedeagus, dorsal view; 6-7. right paramere, internal and external face; 8-9, left paramere, internal and external face); 10-11. Left ovipositor, ventral view (10. photograph; 11. line drawing; as - apical stylomere; bs - basal stylomere; dles - dorsolateral ensiform setae; dmes - dorsomedial ensiform setae; ns - nematiform setae; ves - ventral ensiform setae; vf - valvifer). Scale bars = 0.3 mm (Figs 4-9); = 0.1 mm (Figs 10-11).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="distribution">
<paragraph pageId="1" pageNumber="88">Distribution.</paragraph>
<paragraph pageId="1" pageNumber="88">Palaearctic Region (Japan, East China), Oriental Region (Ceylon, India, South China, Taiwan, Philippines, Malaysia, Indonesia), Australian Region (New Guinea, northeast and southeast Australia).</paragraph>
<subSubSection id="A981F7B8410C9571619DE9B3B06399ED" pageId="1" pageNumber="88" type="distribution">
<paragraph id="A8012BB53DF80D51F75A90F44939AFA7" pageId="1" pageNumber="88">Distribution.</paragraph>
<paragraph id="FC3191C0E5DD8785E6A52792BB17EFE1" pageId="1" pageNumber="88">Palaearctic Region (Japan, East China), Oriental Region (Ceylon, India, South China, Taiwan, Philippines, Malaysia, Indonesia), Australian Region (New Guinea, northeast and southeast Australia).</paragraph>
</subSubSection>
</treatment>
</document>

File diff suppressed because one or more lines are too long

File diff suppressed because one or more lines are too long

View file

@ -1,105 +1,113 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864673" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="86370FFE15EE55B256D6226E9D7E4D8F" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Thaioodes Gueorguiev, 2014, gen. n." docType="treatment" docVersion="5" lastPageId="7" lastPageNumber="94" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="5" pageNumber="92" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="FC3F251FF500CC0870E771BA34DEB379" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="86370FFE15EE55B256D6226E9D7E4D8F" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Thaioodes Gueorguiev, 2014, gen. n." docType="treatment" docVersion="6" lastPageNumber="94" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="92" updateTime="1731807077730" updateUser="admin">
<mods:mods id="8F0BF20561F9BD7EF4CA94C51CC720BC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5B734354F2939879669D19663EDBF5FE">
<mods:title id="62A48E118D08B47D8CAEEC7266A6C21B">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="8CD4604E3B64CAFE7DF3FE73CCD38359" type="personal">
<mods:role id="895A839AE5308FE8060ADD64D82F2133">
<mods:roleTerm id="AF2EEA3A6B7E41A4D2859416EC406140">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="F1848281DBFB3FC5CDE8FED04BBFE76A">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="076156018D1BABD78BC772DEC695CA19">text</mods:typeOfResource>
<mods:relatedItem id="185CAC5E7A7F53FAD0E7CD1A84320321" type="host">
<mods:titleInfo id="9BF743F92621AB431150B9AABC5D8E7F">
<mods:title id="9C3214C797FB601135BB4D7F99CE5840">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="37417BFED32434C018C9D845D337331A">
<mods:date id="0F9F217A3C921B83F3DF2BE2C08CF18E">2014</mods:date>
<mods:detail id="54D561F2FD113A3862A374A2D3A663AE" type="volume">
<mods:number id="32A68B6F5D116785BB7923BB64A96BC2">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="BF90C3997659E69B3A6E5BC0B3AC9478" type="issue">
<mods:number id="F6C501C75CCCA35BE377C52CDA96935F">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="E50320E40CBEF4D10FF6AF2C2B40EAC7" unit="page">
<mods:start id="F302AFA50294F82C52DFE7381F326E55">87</mods:start>
<mods:end id="2354CF955CACEAB1BE05C0328F9CC5DF">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="2CE925ECCDBA85599A4C581F65612A61">
<mods:url id="969B9B50732B40FE53E149A55252811C">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="5B7969D9D82A9661B2A89CE16DEFA935">journal article</mods:classification>
<mods:identifier id="2AA7C3080B61C164767FF953C144485D" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="090E2987B2954B3D1616D0DCE6CDA361" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864673" LSID="urn:lsid:plazi:treatment:86370FFE15EE55B256D6226E9D7E4D8F" httpUri="http://treatment.plazi.org/id/86370FFE15EE55B256D6226E9D7E4D8F" lastPageId="7" lastPageNumber="94" pageId="5" pageNumber="92">
<subSubSection pageId="5" pageNumber="92" type="multiple">
<paragraph pageId="5" pageNumber="92">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="92" type="nomenclature">
<paragraph pageId="5" pageNumber="92">
<taxonomicName LSID="http://zoobank.org/605266A8-7B56-45DE-AD3F-B4B112FCD209" class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus">Thaioodes</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="92">gen. n.</taxonomicNameLabel>
<treatment id="86370FFE15EE55B256D6226E9D7E4D8F" ID-GBIF-Taxon="127864673" LSID="urn:lsid:plazi:treatment:86370FFE15EE55B256D6226E9D7E4D8F" httpUri="http://treatment.plazi.org/id/86370FFE15EE55B256D6226E9D7E4D8F" lastPageId="7" lastPageNumber="94" pageId="5" pageNumber="92">
<subSubSection id="8C353CD4505466F563DE583EC30A3F6A" pageId="5" pageNumber="92" type="multiple">
<paragraph id="791A31E9F6C7D6E6BF2A093E5226B15A" pageId="5" pageNumber="92">
Taxon classification Animalia
<taxonomicName id="ADF083D7D3047DB042E6D2A1D02A69AA" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="2B76D0318D2CFE76B308AE5DD7795AEA" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="92" type="type species">
<paragraph pageId="5" pageNumber="92">Type species.</paragraph>
<paragraph pageId="5" pageNumber="92">
<taxonomicName class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes piceus" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="piceus">Thaioodes piceus</taxonomicName>
B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
sp. n.
<subSubSection id="8DBAA9C0063AEB6A904A9B00AB905C9B" pageId="5" pageNumber="92" type="nomenclature">
<paragraph id="F1AFB30CAF5F27836CFEB1EC3D6BE6B2" pageId="5" pageNumber="92">
<taxonomicName id="8684EAB856AF15AB114F730D242CD715" ID-CoL="8KZ7L" LSID="http://zoobank.org/605266A8-7B56-45DE-AD3F-B4B112FCD209" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">Thaioodes</taxonomicName>
<taxonomicNameLabel id="E410FF6816F77BD1517074BCC86C7579" pageId="5" pageNumber="92" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92" type="diagnosis">
<paragraph pageId="5" pageNumber="92">Diagnosis.</paragraph>
<paragraph lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
<subSubSection id="4E4AF781F6537CE82907D963A259B1BD" pageId="5" pageNumber="92" type="type species">
<paragraph id="70C17619F3D875B1EBBC9D718BA6DB8E" pageId="5" pageNumber="92">Type species.</paragraph>
<paragraph id="DA4483F06600D1B4E24E4B8FD9E151F9" pageId="5" pageNumber="92">
<taxonomicName id="5A43EBD4769005A7216EA5F9319BBA38" authority="B. Gueorguiev" authorityName="B. Gueorguiev" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes piceus" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="piceus" status="sp. nov.">
Thaioodes piceus B.
<normalizedToken id="B035F0906A3335162E33D63E12B75044" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="C8E1AFC1E40030F989BC5BEE72DEF6F1" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="7DAC7C2EFBE6C3B21AED2DEE4CDC4ED6" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92" type="diagnosis">
<paragraph id="9FFD4E868304464DA28CBE7A1ED68CBA" pageId="5" pageNumber="92">Diagnosis.</paragraph>
<paragraph id="2462C6B7F6CC156C8BD35DC48064B853" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
Small to medium-sized species (6.7-6.9 mm) for Oriental
<taxonomicName lsidName="" pageId="5" pageNumber="92" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="5C40B36615CB2B4D421AC539A4B5EBA9" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, habitus semi-elongate; color piceous on dorsal and ventral surface, with slight bluish color dorsally, antennomeres 1-3 of antennae, palpi, tibiae, tarsomeres red-yellow to yellow; microsculpture isodiametric; tegument completely glabrous and impunctate, only disc of head lightly punctate; head with conspicuously large eyes and minute tempora; frons with punctiform impressions and one pair of supraorbital punctures; labrum with six setae along anterior margin; clypeus with two setae; labial palpomere glabrous; mentum tooth rounded anteriorly, bordered posteriorly; pronotum subquadrate, rounded, without protruded angles, with anterior and posterior margins of nearly equal width; sides of pronotum bordered apically and laterally, not bordered basally; anterolateral and posterolateral setae lacking; laterobasal impressions faint, sublinear; elytra with well-im
<pageBreakToken pageId="6" pageNumber="93" start="start">pressed</pageBreakToken>
<pageBreakToken id="C15696DF462604C3722135A94AD416F3" pageId="6" pageNumber="93" start="start">pressed</pageBreakToken>
striae, distinct to apex; striae 5-7 obliterated basally; stria 8 grooved along most of its length, deeper than striae 1-7; parascutellar striola well-developed, long, situated between suture and stria 1; intervals impunctate, smooth, subconvex in anterior half, convex posteriorly; interval 3 at apex nearly twice as broad as intervals 1 and 2; intervals 7 and 8 fused posteriorly or not to apex; interval 8 (or its substitute interval 7) forming outward ridge, moderately elevated, more distinct along posterior half of elytra; interval 9 transformed into marginal gutter throughout; marginal gutter distinct to apex of elytron; prosternal process bordered; mesosternum concave; metepisternum laterally coadunate with elytral epipleuron; sterna 4-5 with pair of minute ambulatory setae; apical sternum without setae in male; mesocoxa without lateral seta, with posteromedial seta; metacoxa with anterior pore; mesofemur without posterior setae; metafemur glabrous; protarsomeres 1 and 3 in males longer than wide, tarsomere 2 subquadrate, each with variable number of small round adhesive setae with oval discs underneath; tarsomere 5 of all legs setose ventrally.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="93" type="etymology">
<paragraph pageId="6" pageNumber="93">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="93">
<subSubSection id="79EDD8D3FD130FD683E7E45AF484B8E2" pageId="6" pageNumber="93" type="etymology">
<paragraph id="734A71672555F0ED564DB1F5161E300E" pageId="6" pageNumber="93">Etymology.</paragraph>
<paragraph id="32216F204B806E33EA6409DFBEEB6C49" pageId="6" pageNumber="93">
A compound word, based on the ethnic name of the predominating people in the country where the type species was found, Thai, and
<taxonomicName class="Insecta" family="Carabidae" genus="Oodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
<taxonomicName id="DF4F399C4985440C3A020076CE301281" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
(for the etymology of this name see
<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="97" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexiko." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
<bibRefCitation id="E8A759B395B4FBF275D3EFE3CB859B2C" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="97" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexiko." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
: 955). It is treated as a Latin masculine.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="94" pageId="6" pageNumber="93" type="affinities">
<paragraph pageId="6" pageNumber="93">Affinities.</paragraph>
<paragraph pageId="6" pageNumber="93">
<subSubSection id="53ED752E68F22B2C24942A346AD891A8" lastPageId="7" lastPageNumber="94" pageId="6" pageNumber="93" type="affinities">
<paragraph id="9F922CF7F94C9E97DB22FBF628B3B830" pageId="6" pageNumber="93">Affinities.</paragraph>
<paragraph id="E9BC840BDA68433D429D0BCEF56C6291" pageId="6" pageNumber="93">
Specific characters of the new genus correspond with the basic features of
<taxonomicName lsidName="" pageId="6" pageNumber="93" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="45AAE530A5A9923A4BEF34E00E5C6517" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
as far as these have been outlined by
<bibRefCitation author="Jeannel, R" journalOrPublisher="Office de la Recherche scientifique colonial, Paris" pageId="10" pageNumber="97" pagination="767 - 1146" title="Coleopteres carabiques de la region Malgache (troisieme partie). Faune de l'Empire Francais" year="1949 a">Jeannel (1949a)</bibRefCitation>
<bibRefCitation id="7C572A5765DCBD6DEAFEFD2F2F6ED74F" author="Jeannel, R" journalOrPublisher="Office de la Recherche scientifique colonial, Paris" pageId="10" pageNumber="97" pagination="767 - 1146" title="Coleopteres carabiques de la region Malgache (troisieme partie). Faune de l'Empire Francais" year="1949 a">Jeannel (1949a)</bibRefCitation>
and
<bibRefCitation author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet (1996)</bibRefCitation>
<bibRefCitation id="8F90AFA0B451E2F9E8C217A64D01E95A" author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet (1996)</bibRefCitation>
: 1, integument of body glabrous and largely impunctate; 2, antennomere 3 glabrous, excluding apical setae; 3, stria 8 deeper than other striae; 4, interval 8 forming outward ridge posteriorly; 5, interval 9 transformed into marginal gutter.
</paragraph>
<paragraph pageId="7" pageNumber="94">
<taxonomicName class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="7" pageNumber="94" start="start">Thaioodes</pageBreakToken>
<paragraph id="ED1C0496D00C815022D138CBDA774CE5" pageId="7" pageNumber="94">
<taxonomicName id="630F3CC11C681CD6CBC0D6B0FFEDEE0E" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus" status="gen. nov.">
<pageBreakToken id="6DAC982365DC7E308607E7E6AC94DC0C" pageId="7" pageNumber="94" start="start">Thaioodes</pageBreakToken>
</taxonomicName>
gen. n. is most closely allied to
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n. (see
<normalizedToken originalValue="Discussion">'Discussion'</normalizedToken>
<taxonomicNameLabel id="B961AE2C6D0F1BAB9ADB2C6B151D37CA" rank="genus">gen. n.</taxonomicNameLabel>
is most closely allied to
<taxonomicName id="1EA159D5BFC456771A2E04CA47954290" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="C36944F002CC312F58390C7EFB40652F" rank="genus">gen. n.</taxonomicNameLabel>
(see
<normalizedToken id="174430217252665933D1D3CBA5709188" originalValue="Discussion">'Discussion'</normalizedToken>
). The former genus differs from the latter genus by: 1, tegument unicolored dorsally (vs. tegument bicolored dorsally); 2, pronotum slightly wider than long (PW/PL: 1.18), with sides partly bordered (vs. pronotum distinctly wider than long, PW/PL: 1.24-1.37, with sides finely bordered throughout); 3, anterolateral and basolateral setae of pronotum lacking (vs. anterolateral seta lacking, basolateral setae present); 4, all striae of elytra distinct to apex (vs. elytral striae 1-7 obliterated before apex); 5, stria 7 ending before apex or joining stria 8, thus intervals 7 and 8 fused posteriorly (vs. stria 7 separate before apex, thus intervals 7 and 8 not fused posteriorly); 6, intervals convex posteriorly (vs. intervals uniformly flat throughout); 7, interval 8 forms ridge posteriorly (vs. interval 8 not forming ridge posteriorly); 8, interval 9 transformed into marginal gutter throughout (vs. interval 9 transformed into marginal gutter only at anterior two fifths of elytron); 9, marginal gutter distinct to apex of elytron (vs. marginal gutter ended before apex of elytron); 10, umbilicate series of elytra with 19-20 pores (vs. umbilicate series with 15-16 pores); 11, prosternal process bordered (vs. prosternal process unbordered).
</paragraph>
<paragraph pageId="7" pageNumber="94">
<paragraph id="16E8A70DEC0C28BF70DE56071B714E77" pageId="7" pageNumber="94">
In my estimation, the presented series of differences and their opposed states are enough to differentiate
<taxonomicName class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes piceus" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="piceus">Thaioodes piceus</taxonomicName>
sp. n. from
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes cyaneus" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="cyaneus">Bamaroodes cyaneus</taxonomicName>
<taxonomicName id="A7B84F506C29C5922D93354A58A84A13" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes piceus" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="piceus" status="sp. nov.">Thaioodes piceus</taxonomicName>
<taxonomicNameLabel id="067D204B224B21300736C8A0523FEAFA" rank="species">sp. n.</taxonomicNameLabel>
from
<taxonomicName id="6ECF7E046839C5283486C25D74EC8043" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes cyaneus" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="cyaneus">Bamaroodes cyaneus</taxonomicName>
at a generic level.
</paragraph>
</subSubSection>

View file

@ -1,287 +1,282 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864670" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="878EC324604D05FA2DEED99D93A59957" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="OODINI LaFerte-Senectere 1851" docType="treatment" docVersion="5" lastPageId="2" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="34F4CB4E811D5E148DDAC8983EC019CE" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="878EC324604D05FA2DEED99D93A59957" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Oodini LaFerte-Senectere 1851" docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="88" updateTime="1731807077730" updateUser="admin">
<mods:mods id="92778D3030CC04F06115C15AF7C3AE9F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6B44DB0E78CB20A45AD51AFBA218DEF7">
<mods:title id="04E9ECA969DBB0E50D2315B9F3F3DA55">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="B6FFB9222888F6D03442500F63A533E4" type="personal">
<mods:role id="60A634E9B11038FC9B8F74744181A85E">
<mods:roleTerm id="53E22EC743FB1BEAD749ACC1F8BBA086">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="38DD58AEE84E915049E64FFF79E6E7D1">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="FE263B3155A37CE872BA55B0CAE0AEFA">text</mods:typeOfResource>
<mods:relatedItem id="A6560CC0E59991BEB2EFB7B999525AB0" type="host">
<mods:titleInfo id="A6D6A80EE14F97248D27128C7610CE92">
<mods:title id="969A27469CF838C8E342813F1100DD2B">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="1F0FFBE758A9C19ADD29E99B665A459E">
<mods:date id="7C96B578CAF0A572F289AA654F5A3032">2014</mods:date>
<mods:detail id="5F3F0B658EDE89AE08E4E45F51CC6AFB" type="volume">
<mods:number id="A7AD28B86C8D9130B67764CCDAF2C041">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="4F4BAA80984E0C7981B68E654BBBA066" type="issue">
<mods:number id="C63E3C12920352CE647A7C215D4F0176">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="34B83E8943587593F11DFF316E55C692" unit="page">
<mods:start id="58A3731858DFB35F371508043A8C39EB">87</mods:start>
<mods:end id="2BBC322397390C65EBD38056578C2402">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="199C532D1AD51E346BF79A8492D0D351">
<mods:url id="50468C1505FC78FFE01B20ACF65AF7A9">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="A135B82A1E16F4D12FC45C55CA968CD2">journal article</mods:classification>
<mods:identifier id="7C012A0A3E8029747444C5AE654C304C" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="C71DE56B54430FE94C91C794F6C62633" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864670" LSID="urn:lsid:plazi:treatment:878EC324604D05FA2DEED99D93A59957" httpUri="http://treatment.plazi.org/id/878EC324604D05FA2DEED99D93A59957" lastPageId="2" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="multiple">
<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
<treatment id="878EC324604D05FA2DEED99D93A59957" ID-GBIF-Taxon="127864670" LSID="urn:lsid:plazi:treatment:878EC324604D05FA2DEED99D93A59957" httpUri="http://treatment.plazi.org/id/878EC324604D05FA2DEED99D93A59957" lastPageId="2" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection id="CFA2F4F0490405973A4227F9AD83DCD5" pageId="1" pageNumber="88" type="multiple">
<paragraph id="AD14EEDE35930C8D40D8A03E0D53171A" pageId="1" pageNumber="88">
Taxon classification Animalia
<taxonomicName id="D2E7805166CB63B304A9832FDCB61B75" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="FBF0F29060FD7F484A7F1EE11443E79C" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="OODINI">
<subSubSection id="7CDABA3B34CB6311BF8876FDC167E448" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="D92C2692BA7F5879B793E6504ACACFDA" pageId="1" pageNumber="88">
<taxonomicName id="856E61A67CFD8CE353027F37D149400A" ID-CoL="8KTWG" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">
OODINI
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
<normalizedToken id="9A6D47030DDB7BF75B640B891740C002" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
, 1851
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="tribal diagnosis">
<paragraph pageId="1" pageNumber="88">Tribal diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="88">
<subSubSection id="B0CF713B55ABD244F8857817FF6072B2" pageId="1" pageNumber="88" type="tribal diagnosis">
<paragraph id="F0727FC75B1AD8778F2E016650C8B879" pageId="1" pageNumber="88">Tribal diagnosis.</paragraph>
<paragraph id="2820B1A024C1644A8972EF6F3F4EB0BB" pageId="1" pageNumber="88">
Mandibular scrobe without seta near anterior margin of excavation, mesocoxal cavities conjunct (mesepimeron does not reach middle coxal cavity), metepisternum separated from metepimeron by distinct suture, and parameres without setae, with left paramere
<normalizedToken originalValue="“conchoid”">&quot;conchoid&quot;</normalizedToken>
<normalizedToken id="0A9CC561CA636867F93E22D7DA12A944" originalValue="“conchoid”">&quot;conchoid&quot;</normalizedToken>
, therefore a member taxon of Conchifera Jeannel, 1941 (
<bibRefCitation author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
<bibRefCitation id="8A5AD6C7F52E1C74F41A46E64E8C9E4F" author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
: 1047); head with one pair of supraorbital setae, its capsule
<normalizedToken originalValue="“normal”">&quot;normal&quot;</normalizedToken>
<normalizedToken id="109949A917DC0AE173889E4D14A2B42A" originalValue="“normal”">&quot;normal&quot;</normalizedToken>
, i.e. without annular constriction behind eyes and with ultimate maxillary palpomere elongate and well-centered with respect to penultimate palpomere, pronotum usually without anterolateral setae, anterior coxal cavities biperforate, elytron with internal plica, metepisternum laterally coadunate with elytral epipleuron, middle femur posterior margin without longer setae, metacoxa lacking posterior seta, and median lobe of aedeagus in lateral view long and considerably bent, with basal bulb decreased, indicate a member taxon of
<taxonomicName family="Callistidae" lsidName="" pageId="1" pageNumber="88" rank="family">Callistidae</taxonomicName>
<taxonomicName id="9F0797F34008F1E80EB98349FAA2EE73" family="Callistidae" lsidName="" pageId="1" pageNumber="88" rank="family">Callistidae</taxonomicName>
Jeannel, 1941 (
<bibRefCitation author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
<bibRefCitation id="9A8BA6CF4566874C7944DF1AC063DD7B" author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
: 1051); tegument glabrous, largely impunctate, antennomere 3 glabrous (excluding apical setae), posterior margin of pronotum not fringed, elytron with stria 7 ended before apex, stria 8 deeper than other striae, and interval 9 transformed into marginal gutter partly or throughout, add traits peculiar to tribe (all genera, except for
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n., have also elytron with intervals 7 and 8 fused and forming a ridge posteriorly).
<taxonomicName id="CED0ACD6CC29CB163C7DF4EC62BFE306" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="CFDC33D775DF3C7162A8BAF4E3424B9D" rank="genus">gen. n.</taxonomicNameLabel>
, have also elytron with intervals 7 and 8 fused and forming a ridge posteriorly).
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88" type="a key to the genera for adults of oriental oodini, including oriental chlaeniini with oodine facies">
<paragraph pageId="1" pageNumber="88">
<subSubSection id="5AFA11E7F4507D14FDBAA6BFD9C86CBE" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88" type="a key to the genera for adults of oriental oodini, including oriental chlaeniini with oodine facies">
<paragraph id="7D536720D5C8B1D5F9239E3AC8D6E50B" pageId="1" pageNumber="88">
A key to the genera for adults of Oriental
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="F9CA3B2C4B69D91B671EC7791E87516B" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, including Oriental
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="CD671D365562794EE58A17484E13E927" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
with oodine facies
</paragraph>
<paragraph lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
<table lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">1</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">
<paragraph id="045F4EC882ED631B6C02975DC86E687E" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
<table id="662B7E4B2813CD24CF357BE34975721F" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
<tr id="7867A4AF8093D5E8B1FCF500E130C275" pageId="1" pageNumber="88">
<td id="47F9ACB38A54DF10D209B53439407737" colspan="1" pageId="1" pageNumber="88" rowspan="1">1</td>
<td id="501B180EE50851491BE83E64D12145DF" colspan="1" pageId="1" pageNumber="88" rowspan="1">
Interval 9 of elytra distinct, not transformed into marginal gutter. Stria 8 neither deeper than other striae (modestly deeper only in
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<taxonomicName id="2F59D912EFB15DB3D78F3449AB48F51C" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
) nor grooved. Integument more or less extensively punctate and pubescent. Antennomere 3 with a few short, scattered setae (in addition to apical setae)
</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">
<td id="3FDD9BF9586D1D7F9F567B09EB9DBEBA" colspan="1" pageId="1" pageNumber="88" rowspan="1">
[
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="59A140A0FFE128BD54700395361F9F74" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
]...2
</td>
</tr>
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">Interval 9 of elytra transformed into marginal gutter throughout or at least along its anterior two-fifths. Stria 8 deeper than other striae, distinctly grooved posteriorly. Integument glabrous, impunctate or restrictedly punctate. Antennomere 3 glabrous (but with apical setae)</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">
<tr id="2C1A48459052601F798E754797627EC9" pageId="1" pageNumber="88">
<td id="B5710DA64E22E877DD0831E7562B2C4A" colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td id="BF705A8A11B0405B7FCC338C0FB7CCB6" colspan="1" pageId="1" pageNumber="88" rowspan="1">Interval 9 of elytra transformed into marginal gutter throughout or at least along its anterior two-fifths. Stria 8 deeper than other striae, distinctly grooved posteriorly. Integument glabrous, impunctate or restrictedly punctate. Antennomere 3 glabrous (but with apical setae)</td>
<td id="1BF5705027F4E9383F010C8870BE2B1B" colspan="1" pageId="1" pageNumber="88" rowspan="1">
[
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="8393A8CC186859AA20B8CC3796FCEA14" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
]...3
</td>
</tr>
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">2</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">Penultimate segment of labial palpomere with 2-4 spines at front margin. All striae of elytra linear, stria 8 shallower than or as deep as other striae. Basal margin of pronotum nearly straight, only briefly rounded at posterior angle. Sterna 3-5 with pair of long ambulatory setae</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
Semenov, 1889
<tr id="5DE7A3240D63D2864FAAA020F500EC3F" pageId="1" pageNumber="88">
<td id="AE151C386DC5390B8A7B3F07508CC1D6" colspan="1" pageId="1" pageNumber="88" rowspan="1">2</td>
<td id="3A2E007B6B7796FA00F061B69A397F38" colspan="1" pageId="1" pageNumber="88" rowspan="1">Penultimate segment of labial palpomere with 2-4 spines at front margin. All striae of elytra linear, stria 8 shallower than or as deep as other striae. Basal margin of pronotum nearly straight, only briefly rounded at posterior angle. Sterna 3-5 with pair of long ambulatory setae</td>
<td id="B848EA5EB3E60F366E004A125BA6213A" colspan="1" pageId="1" pageNumber="88" rowspan="1">
<taxonomicName id="04CA53C8796CC19221AAA14396122E85" authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma Semenov, 1889</taxonomicName>
</td>
</tr>
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">Penultimate segment of labial palpomere without seta. Striae 1-7 punctiform, stria 8 linear, somewhat deeper than other striae (Figs 2-3). Basal margin of pronotum laterally oblique towards posterior angle. Sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
<tr id="257CB7686841CA273204CC9969EAAD87" pageId="1" pageNumber="88">
<td id="A017B243E0619AD0DA2C956023D9A67F" colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td id="5EEF7DDE3D236249DE278BEF762EE931" colspan="1" pageId="1" pageNumber="88" rowspan="1">Penultimate segment of labial palpomere without seta. Striae 1-7 punctiform, stria 8 linear, somewhat deeper than other striae (Figs 2-3). Basal margin of pronotum laterally oblique towards posterior angle. Sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae</td>
<td id="6EC489EE49E04DC00B7A9D7623AFFD16" colspan="1" pageId="1" pageNumber="88" rowspan="1">
<taxonomicName id="873B9006E8EA3A83D5FED1D326E3D8F3" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
Hololeius
<normalizedToken id="276A5B481FE91842EA2BFA3470A9C386" originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
, 1851
</taxonomicName>
</td>
</tr>
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">3</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">Tarsomere 5 setose ventrally. Mesocoxa lateral margin without seta. Body semi-elongate. Anterior and posterior margins of pronotum with similar widths. Claws nearly parallel (Figs 15-16, 21-22)</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">4</td>
<tr id="38B29A9DABEB7035C645BA047CDD1954" pageId="1" pageNumber="88">
<td id="EFB274B388B339C1B30DDC9D4F3EAD63" colspan="1" pageId="1" pageNumber="88" rowspan="1">3</td>
<td id="1277D9089C6DCDED9D883F700D22053A" colspan="1" pageId="1" pageNumber="88" rowspan="1">Tarsomere 5 setose ventrally. Mesocoxa lateral margin without seta. Body semi-elongate. Anterior and posterior margins of pronotum with similar widths. Claws nearly parallel (Figs 15-16, 21-22)</td>
<td id="54421520E135BF5FC3E22F0D6E166A2F" colspan="1" pageId="1" pageNumber="88" rowspan="1">4</td>
</tr>
<tr pageId="1" pageNumber="88">
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">Tarsomere 5 glabrous ventrally. Mesocoxa lateral margin with one or a few setae. Body elliptic or broadly oval. Anterior and posterior margins of pronotum with dissimilar widths. Claws widely separated</td>
<td colspan="1" pageId="1" pageNumber="88" rowspan="1">5</td>
<tr id="361DB7E2C1BF66378BA07E5D89E58B71" pageId="1" pageNumber="88">
<td id="E2E4505A1FA12A377CA61137ABFC3F28" colspan="1" pageId="1" pageNumber="88" rowspan="1">-</td>
<td id="6796AC68B617E7145D107F650FD7FE29" colspan="1" pageId="1" pageNumber="88" rowspan="1">Tarsomere 5 glabrous ventrally. Mesocoxa lateral margin with one or a few setae. Body elliptic or broadly oval. Anterior and posterior margins of pronotum with dissimilar widths. Claws widely separated</td>
<td id="E0ED585002533BDE26C6B0FBCF6F0B64" colspan="1" pageId="1" pageNumber="88" rowspan="1">5</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<pageBreakToken pageId="2" pageNumber="89" start="start">4</pageBreakToken>
<tr id="773160271A52D69CCFD595238CB277DF" pageId="2" pageNumber="89">
<td id="415994311B2A0CFF1E9154469EF67467" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<pageBreakToken id="686AD11917EA533E02759E93CD75A1DE" pageId="2" pageNumber="89" start="start">4</pageBreakToken>
</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra two-colored, mostly black with bluish hue, with yellow apical band (Fig. 13). Striae 2-7 obliterated apically; intervals 7 and 8 not fused posteriorly (Figs 15-16). Intervals uniformly flat throughout; interval 8 not forming a ridge posteriorly; interval 9 transformed into marginal gutter only anteriorly, distinct on posterior three fifths of elytra; marginal gutter ended before apex of elytron (Figs 14-16). Pronotum distinctly wider than long (PW/PL: 1.24-1.37), with sides finely bordered throughout; posterolateral setae present. Prosternal process unbordered</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n.
<td id="241B0D0FB1F5621B2ACCF7F9DFCE5128" colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra two-colored, mostly black with bluish hue, with yellow apical band (Fig. 13). Striae 2-7 obliterated apically; intervals 7 and 8 not fused posteriorly (Figs 15-16). Intervals uniformly flat throughout; interval 8 not forming a ridge posteriorly; interval 9 transformed into marginal gutter only anteriorly, distinct on posterior three fifths of elytra; marginal gutter ended before apex of elytron (Figs 14-16). Pronotum distinctly wider than long (PW/PL: 1.24-1.37), with sides finely bordered throughout; posterolateral setae present. Prosternal process unbordered</td>
<td id="69C02ACDC3E61D02D41722758D8BE3C5" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="E73D51B54AF4EE3B429762AFA0DD5213" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="E043FF7698736D704F56FA2EC2AD9AA7" rank="genus">gen. n.</taxonomicNameLabel>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra unicolored, blackish (Fig. 19). Striae distinct toward apex; intervals 7 and 8 fused posteriorly (Figs 21-22). Intervals convex posteriorly; interval 8 forming a ridge posteriorly; interval 9 transformed into marginal gutter throughout; marginal gutter distinct to apex of elytron (Figs 20-22). Pronotum slightly wider than long (PW/PL: 1.18), with sides not bordered basally; posterolateral setae lacking. Prosternal process bordered.</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Thaioodes</taxonomicName>
gen. n.
<tr id="C435DE3BBC2841179456AF26033A59F0" pageId="2" pageNumber="89">
<td id="B3AB50648D94F6916B758EA083D30165" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="0A2EE1C610FA28055596430EEC282C25" colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra unicolored, blackish (Fig. 19). Striae distinct toward apex; intervals 7 and 8 fused posteriorly (Figs 21-22). Intervals convex posteriorly; interval 8 forming a ridge posteriorly; interval 9 transformed into marginal gutter throughout; marginal gutter distinct to apex of elytron (Figs 20-22). Pronotum slightly wider than long (PW/PL: 1.18), with sides not bordered basally; posterolateral setae lacking. Prosternal process bordered.</td>
<td id="54A517723798FFBA41319FAA6FDCCD83" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="173FD9E04029E6B1257839F276DA251F" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">Thaioodes</taxonomicName>
<taxonomicNameLabel id="E595B79494EA82AA7DD57168B9CE4E98" rank="genus">gen. n.</taxonomicNameLabel>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">5</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Labrum with three pores on anterior margin, central pore bearing two or four connate setae</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">6</td>
<tr id="7B6E37021DF6BBD12F137ED613969E2A" pageId="2" pageNumber="89">
<td id="EB029A1BE55D740C6561AA24E3B54CB1" colspan="1" pageId="2" pageNumber="89" rowspan="1">5</td>
<td id="47CED9504A4A7243A362B7A73A8CAAFF" colspan="1" pageId="2" pageNumber="89" rowspan="1">Labrum with three pores on anterior margin, central pore bearing two or four connate setae</td>
<td id="3AF5EDE81746911CE66DE33C9F4DFDC4" colspan="1" pageId="2" pageNumber="89" rowspan="1">6</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Labrum with six pores on anterior margin, each pore bearing a separate seta</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">10</td>
<tr id="1A3493EA317209892E2681ACF82A3336" pageId="2" pageNumber="89">
<td id="577D5892579ED3E269A29D7DDD15259E" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="59DE7DB61AE9C82C324FA120507DDD8E" colspan="1" pageId="2" pageNumber="89" rowspan="1">Labrum with six pores on anterior margin, each pore bearing a separate seta</td>
<td id="98A9B30479E08E744D9A2F86671204B2" colspan="1" pageId="2" pageNumber="89" rowspan="1">10</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">6</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus without setae. Mesepisternum without apodemal pit. Length of body 5-16 mm</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">7</td>
<tr id="11E1E62EC723B05F2E648C1FA78DFA4C" pageId="2" pageNumber="89">
<td id="74E9AAEFC4925CD3EF2E332D2B7B3A52" colspan="1" pageId="2" pageNumber="89" rowspan="1">6</td>
<td id="9A7EFC021BF9AEC37E15838FCA0E0834" colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus without setae. Mesepisternum without apodemal pit. Length of body 5-16 mm</td>
<td id="22E555EB9E3F882CF3889ADED5EF8E3A" colspan="1" pageId="2" pageNumber="89" rowspan="1">7</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<tr id="53756CB9F22152254CCBC4FC3715E295" pageId="2" pageNumber="89">
<td id="8C4AFD979B65331E03371FFDE9A2A309" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="EF557D54839C8C42393E557C2AC41843" colspan="1" pageId="2" pageNumber="89" rowspan="1">
Clypeus with two setae. Mesepisternum with apodemal pit (in
<taxonomicName class="Insecta" family="Carabidae" genus="Miltodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miltodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Miltodes</taxonomicName>
<taxonomicName id="3EF4B9B6E50FB4CFAB850A2879C949A4" class="Insecta" family="Carabidae" genus="Miltodes" kingdom="Animalia" lsidName="Miltodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Miltodes</taxonomicName>
pit very small). Length of body 3-5 mm
</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">9</td>
<td id="9F13C577327559F671265048B3C06E75" colspan="1" pageId="2" pageNumber="89" rowspan="1">9</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">7</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Smaller species, length of body less than 10 mm</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Nanodiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanodiodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Nanodiodes</taxonomicName>
Bousquet, 1996
<tr id="F30F6A67A4B93DDDD31D4E12CF2D7B97" pageId="2" pageNumber="89">
<td id="9257C848A69DE692F316108EA04EB56B" colspan="1" pageId="2" pageNumber="89" rowspan="1">7</td>
<td id="F1FA41F33EDCCC586C9725F997C595A2" colspan="1" pageId="2" pageNumber="89" rowspan="1">Smaller species, length of body less than 10 mm</td>
<td id="F168B447B599EBB854DE643C1DC8A200" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="91B8464DD4AAEE3ABC2ECFD0A1224B7A" authority="Bousquet, 1996" authorityName="Bousquet" authorityYear="1996" class="Insecta" family="Carabidae" genus="Nanodiodes" kingdom="Animalia" lsidName="Nanodiodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Nanodiodes Bousquet, 1996</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Larger species, length of body more than 13 mm</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">8</td>
<tr id="59506ADB4C00DFBD83406943523691B9" pageId="2" pageNumber="89">
<td id="D45D2CCA54579232ECFA6C1969AA7A08" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="D852423B2451D6B91DAE5C8AF8171792" colspan="1" pageId="2" pageNumber="89" rowspan="1">Larger species, length of body more than 13 mm</td>
<td id="092A2F7F19BFE8043AEA7A7BC0C45F60" colspan="1" pageId="2" pageNumber="89" rowspan="1">8</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">8</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra with striae 1-7 distinctly grooved, crenulated and complete; interval 3 with two discal punctures</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Systolocranius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systolocranius" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Systolocranius</taxonomicName>
Chaudoir, 1857
<tr id="D3D484FC41BD7482A71CA9C84BDCB224" pageId="2" pageNumber="89">
<td id="B037F70BB02C3EDFAD0DE4717D91EA24" colspan="1" pageId="2" pageNumber="89" rowspan="1">8</td>
<td id="246F62B4E8AF159D512DD22C6B3600CE" colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra with striae 1-7 distinctly grooved, crenulated and complete; interval 3 with two discal punctures</td>
<td id="625CC055C23132E0419BA65BF7667CF0" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="52297F6965411EE6A2C2BB7C7A1E423E" authority="Chaudoir, 1857" authorityName="Chaudoir" authorityYear="1857" class="Insecta" family="Carabidae" genus="Systolocranius" kingdom="Animalia" lsidName="Systolocranius" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Systolocranius Chaudoir, 1857</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra with striae extremely fine and smooth, stria 6 shortened, stria 7 obliterated; interval 3 without discal punctures</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Megaloodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaloodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Megaloodes</taxonomicName>
Lesne, 1896
<tr id="691822DD6ECEA865ED7E86754BC2FEF6" pageId="2" pageNumber="89">
<td id="A840A8B30E4A011B5E5C71F70B6C6985" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="C118CADC2B2080ECBC73C9AE8FBFB041" colspan="1" pageId="2" pageNumber="89" rowspan="1">Elytra with striae extremely fine and smooth, stria 6 shortened, stria 7 obliterated; interval 3 without discal punctures</td>
<td id="651C94EEB453E8A5A9BCDFF650CAB679" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="00E3D3B3046429B078783ED1BDF1E7CA" authority="Lesne, 1896" authorityName="Lesne" authorityYear="1896" class="Insecta" family="Carabidae" genus="Megaloodes" kingdom="Animalia" lsidName="Megaloodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Megaloodes Lesne, 1896</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">9</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Head around 2.5 times narrower than width of base of pronotum. Apodemal pits very small, indistinct, with minute apertures. Interval 3 of elytra without discal setiferous punctures</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Miltodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miltodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Miltodes</taxonomicName>
Andrewes, 1922
<tr id="053C84A7FF4E127BA875604A7AA6CA76" pageId="2" pageNumber="89">
<td id="5EE0C95C39A4295CFA98D82439E0902E" colspan="1" pageId="2" pageNumber="89" rowspan="1">9</td>
<td id="A438B54FB47F61D22EC8BD74F1E56CF6" colspan="1" pageId="2" pageNumber="89" rowspan="1">Head around 2.5 times narrower than width of base of pronotum. Apodemal pits very small, indistinct, with minute apertures. Interval 3 of elytra without discal setiferous punctures</td>
<td id="79820BAB9026E75576467B772E27AAA8" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="7E87C4438DDAC59C3693150644372521" authority="Andrewes, 1922" authorityName="Andrewes" authorityYear="1922" class="Insecta" family="Carabidae" genus="Miltodes" kingdom="Animalia" lsidName="Miltodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Miltodes Andrewes, 1922</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Head around two times narrower than width of base of pronotum. Apodemal pits deep, with wide apertures. Interval 3 of elytra with two small setiferous discal punctures</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Oodinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oodinus" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Oodinus</taxonomicName>
Motschulsky, 1864
<tr id="4746E423FA17620C16D20EFAE87B9B93" pageId="2" pageNumber="89">
<td id="2E7A52C0A11342CD01E85922B5A51524" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="EDD9F2DF51069358E3A420758B350433" colspan="1" pageId="2" pageNumber="89" rowspan="1">Head around two times narrower than width of base of pronotum. Apodemal pits deep, with wide apertures. Interval 3 of elytra with two small setiferous discal punctures</td>
<td id="B4E9F7FC9003C8378959235BAD500992" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="511CDEB1CE0EE713D137E6AE90BD1924" authority="Motschulsky, 1864" authorityName="Motschulsky" authorityYear="1864" class="Insecta" family="Carabidae" genus="Oodinus" kingdom="Animalia" lsidName="Oodinus" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Oodinus Motschulsky, 1864</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">10</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus without setae</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Anatrichis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anatrichis" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Anatrichis</taxonomicName>
LeConte, 1853
<tr id="203B95764233F107688A378D9E50BDE4" pageId="2" pageNumber="89">
<td id="11C9F578CA89C6CE46105012739738C2" colspan="1" pageId="2" pageNumber="89" rowspan="1">10</td>
<td id="F4F8C9FBAB312891D8D3F483C0A81F88" colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus without setae</td>
<td id="D27793EDAFF1BCA99864C114DF0E50C6" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="02622B4C57A8A76C6896CA3214335EEE" authority="LeConte, 1853" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Carabidae" genus="Anatrichis" kingdom="Animalia" lsidName="Anatrichis" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Anatrichis LeConte, 1853</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus with two setae</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">11</td>
<tr id="24F4F3E77011F726354CCBE42848738A" pageId="2" pageNumber="89">
<td id="1737C33CE5C6659C32889E10A42C3C20" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="3C56BE74F7F12AB9480E84C71F9D61AF" colspan="1" pageId="2" pageNumber="89" rowspan="1">Clypeus with two setae</td>
<td id="096FC56DD35621CA373AE512AD9AEDB2" colspan="1" pageId="2" pageNumber="89" rowspan="1">11</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">11</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Parascutellar striola of elytra vestigial or absent</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Pseudosphaerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosphaerodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Pseudosphaerodes</taxonomicName>
Jeannel, 1949
<tr id="66AE6ED8B4704037E82342107E74DD4D" pageId="2" pageNumber="89">
<td id="2552C75EAB6E09C9C220F76EEC76B405" colspan="1" pageId="2" pageNumber="89" rowspan="1">11</td>
<td id="48F95FA0A3BDDE127C16BB3E388C6817" colspan="1" pageId="2" pageNumber="89" rowspan="1">Parascutellar striola of elytra vestigial or absent</td>
<td id="7D322A2D980EAFF7DD8F8835FFED5978" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="18C30CB13445210A9087198240DD6694" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Pseudosphaerodes" kingdom="Animalia" lsidName="Pseudosphaerodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Pseudosphaerodes Jeannel, 1949</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Parascutellar striola of elytra complete</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">12</td>
<tr id="485042D1344C84AB23EB7C992DF21F9F" pageId="2" pageNumber="89">
<td id="86E31B1B63353098FFEE4635BFEC34D0" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="824ECC7DD012368DABCC53C1E343E476" colspan="1" pageId="2" pageNumber="89" rowspan="1">Parascutellar striola of elytra complete</td>
<td id="F781C11980E1A1C6624486A4ECEABD8E" colspan="1" pageId="2" pageNumber="89" rowspan="1">12</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">12</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Protibia with lateral groove on external side. Interval 8 of elytra distinctly narrower than interval 7 (half as much behind shoulder to much narrower toward apex)</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">13</td>
<tr id="0DC26759C995C59A3FA82B88763E37AE" pageId="2" pageNumber="89">
<td id="B25AB2F6A845AA238D14CBAEDED7E324" colspan="1" pageId="2" pageNumber="89" rowspan="1">12</td>
<td id="818E36E9BED73C0F7D604FB618521CB9" colspan="1" pageId="2" pageNumber="89" rowspan="1">Protibia with lateral groove on external side. Interval 8 of elytra distinctly narrower than interval 7 (half as much behind shoulder to much narrower toward apex)</td>
<td id="15D4D93D5332509754DC94C245297616" colspan="1" pageId="2" pageNumber="89" rowspan="1">13</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Protibia without lateral groove on external side. Interval 8 of elytra as wide as or only slightly narrower than interval 7</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">14</td>
<tr id="D65261C8FEF5CE9534CED4FDF38943D1" pageId="2" pageNumber="89">
<td id="9D9E64C4F7AE8DBE03B8848BFD2D0214" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="D1BF4A4D189ABD062B5A83D8554080F8" colspan="1" pageId="2" pageNumber="89" rowspan="1">Protibia without lateral groove on external side. Interval 8 of elytra as wide as or only slightly narrower than interval 7</td>
<td id="C67D73C916E4E122257EC84E4D3D0974" colspan="1" pageId="2" pageNumber="89" rowspan="1">14</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">13</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Interval 8 of elytra carinate behind shoulder</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Holcocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holcocoleus" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Holcocoleus</taxonomicName>
Chaudoir, 1883
<tr id="82E7C028896DBC2E1088DC6F19D8EBFA" pageId="2" pageNumber="89">
<td id="4326F017AD15AF7FC4C7D84EA0EB29B8" colspan="1" pageId="2" pageNumber="89" rowspan="1">13</td>
<td id="4F90AB080E87688D238EB375D9632972" colspan="1" pageId="2" pageNumber="89" rowspan="1">Interval 8 of elytra carinate behind shoulder</td>
<td id="0205262A828CEA09990B112A4638A910" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="BBB399370D8BB41545941C27EDDFD620" authority="Chaudoir, 1883" authorityName="Chaudoir" authorityYear="1883" class="Insecta" family="Carabidae" genus="Holcocoleus" kingdom="Animalia" lsidName="Holcocoleus" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Holcocoleus Chaudoir, 1883</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Interval 8 of elytra as flat as internal intervals</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Brachyodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Brachyodes</taxonomicName>
Jeannel, 1949
<tr id="CAEF4552D0CCB0FA56DDB2BEA75A432B" pageId="2" pageNumber="89">
<td id="04C478B91555113EC9FF6D28A006592B" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="5C97D9C0675C935E44260D23B0B8F7AF" colspan="1" pageId="2" pageNumber="89" rowspan="1">Interval 8 of elytra as flat as internal intervals</td>
<td id="F29489C785ED7DEAC27B01D911701943" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="147C119705FA5505EAFE142F81ECAA04" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Brachyodes" kingdom="Animalia" lsidName="Brachyodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Brachyodes Jeannel, 1949</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">14</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">Mentum tooth subtriangular or triangular, rounded or pointed anteriorly. Non-metallic coloration</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Oodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
Bonelli, 1810
<tr id="0B1E96AEB54A8AB2D0E83C1BBB31435F" pageId="2" pageNumber="89">
<td id="DBDAAC3AF8ECFBA390552EBDCDDC37E7" colspan="1" pageId="2" pageNumber="89" rowspan="1">14</td>
<td id="17CE98392BB323A44EECCEFCE14207CD" colspan="1" pageId="2" pageNumber="89" rowspan="1">Mentum tooth subtriangular or triangular, rounded or pointed anteriorly. Non-metallic coloration</td>
<td id="61D697DDAD577064677D0511659F9B8E" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="D4C72BB82B4EEB854947438EF4834537" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Oodes Bonelli, 1810</taxonomicName>
</td>
</tr>
<tr pageId="2" pageNumber="89">
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<tr id="35ABF6CA90088D08B8D2D6FF5B3006DF" pageId="2" pageNumber="89">
<td id="042BA9F79C09DAFF7936809C63D2F6C6" colspan="1" pageId="2" pageNumber="89" rowspan="1">-</td>
<td id="F84BED475FAACDCD9E0C0B7DD78A22E8" colspan="1" pageId="2" pageNumber="89" rowspan="1">
Mentum tooth sub-trapezoidal, truncate or emarginate anteriorly. Mostly metallic coloration (excl.
<taxonomicName class="Insecta" family="Carabidae" genus="Simous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simous nubilis" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="nubilis">Simous nubilis</taxonomicName>
Andrewes, 1933)
<taxonomicName id="384627C1A5BE653FA934A3DD0BE825A8" authority="Andrewes, 1933" authorityName="Andrewes" authorityYear="1933" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" lsidName="Simous nubilis" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="nubilis">Simous nubilis Andrewes, 1933</taxonomicName>
)
</td>
<td colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Simous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Simous</taxonomicName>
Chaudoir, 1882
<td id="F4603446412EFF473DFB2369379B700C" colspan="1" pageId="2" pageNumber="89" rowspan="1">
<taxonomicName id="FC5B73A1364A6566F9377EF9E2F9D2C0" authority="Chaudoir, 1882" authorityName="Chaudoir" authorityYear="1882" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Simous Chaudoir, 1882</taxonomicName>
</td>
</tr>
</table>

View file

@ -1,251 +1,261 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864671" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="D5298002FCF96ADC19A543AE9E9A5BFE" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Bamaroodes Gueorguiev, 2014, gen. n." docType="treatment" docVersion="5" lastPageNumber="90" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="3" pageNumber="90" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="7653006BF01ED3DC87B1DF61D42F487F" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="D5298002FCF96ADC19A543AE9E9A5BFE" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Bamaroodes Gueorguiev, 2014, gen. n." docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="90" updateTime="1731807077730" updateUser="admin">
<mods:mods id="DAAE25872F791D9F4D287B64CE1F57C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2E36D72C607579A3404A070CC4F50943">
<mods:title id="156DA0107F220AD6B9C168D2D810911D">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="3FFAFA2A5F7922A2175B006249CBB7FE" type="personal">
<mods:role id="6E89378A1E295EF490F0C35E07C67809">
<mods:roleTerm id="A167D939F60234F642F0371326FA9E11">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="4B64978036AB5FBB1AC51EA3E1F8E76E">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="7005A93F36B09E41D0AF635958E80D02">text</mods:typeOfResource>
<mods:relatedItem id="C7112A2FA007689041CAF33FEC22EC40" type="host">
<mods:titleInfo id="5AFDDA3163A5C7CC4C5F10288B7F8744">
<mods:title id="9ACED212F5C14985C7BCCFB3BDB7E552">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="8369346331D01AAB501E5B0D7468F549">
<mods:date id="577153466A74B2DC6681D373FAA373E6">2014</mods:date>
<mods:detail id="58203E4EB6CB1CF1D3509EAE6D147FD7" type="volume">
<mods:number id="14C92DE183E4D61A524D45173421F606">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="31EBCBACCE5170D64D3AF8E3E99EE89E" type="issue">
<mods:number id="FBF6EA1E2D12A596D1FC54ACECE6839E">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="B53CD6FC04473EF0005DDD88A9817A18" unit="page">
<mods:start id="51AD1778CB775B91E2A05B2D3DA2A75B">87</mods:start>
<mods:end id="3E49066F5055D46C037B26DCBF22E7A9">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="E57A7FA6444331CD1B442820D6D04C4A">
<mods:url id="DCAC7D3430157E45DD525498CB29B701">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="0AF04E751EBA72A945E32E575CDD9945">journal article</mods:classification>
<mods:identifier id="A367DA26ECDBEA85C80FEA9B08B30BC6" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="81CD2F9030A04CC86291A0D15BA5B317" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864671" LSID="urn:lsid:plazi:treatment:D5298002FCF96ADC19A543AE9E9A5BFE" httpUri="http://treatment.plazi.org/id/D5298002FCF96ADC19A543AE9E9A5BFE" lastPageNumber="90" pageId="3" pageNumber="90">
<subSubSection pageId="3" pageNumber="90" type="multiple">
<paragraph pageId="3" pageNumber="90">
<pageBreakToken pageId="3" pageNumber="90" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Carabidae
<treatment id="D5298002FCF96ADC19A543AE9E9A5BFE" ID-GBIF-Taxon="127864671" LSID="urn:lsid:plazi:treatment:D5298002FCF96ADC19A543AE9E9A5BFE" httpUri="http://treatment.plazi.org/id/D5298002FCF96ADC19A543AE9E9A5BFE" lastPageNumber="90" pageId="3" pageNumber="90">
<subSubSection id="84476FD3A5C4B2B2B40CCCC26FAF810C" pageId="3" pageNumber="90" type="multiple">
<paragraph id="2C37FAA204AA297EFFC422EED00282C1" pageId="3" pageNumber="90">
<pageBreakToken id="086F5494F71AFD79C9920067E319B949" pageId="3" pageNumber="90" start="start">Taxon</pageBreakToken>
classification Animalia
<taxonomicName id="4FB15613C2D0C213049B6F2907046168" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="D8054DF0C154668DA83BBFB71B1DBE8C" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="90" type="nomenclature">
<paragraph pageId="3" pageNumber="90">
<taxonomicName LSID="http://zoobank.org/0209BE0A-64A2-46B9-BAFB-92BC758D1189" class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="90">gen. n.</taxonomicNameLabel>
<subSubSection id="17755DB8CCE9AD5E5BDAD9429F80C6B4" pageId="3" pageNumber="90" type="nomenclature">
<paragraph id="D8E812CB509224939BAA580FE4306A0E" pageId="3" pageNumber="90">
<taxonomicName id="1104F96312E2CE3366B454560E9EB26F" ID-CoL="8KVJ2" LSID="http://zoobank.org/0209BE0A-64A2-46B9-BAFB-92BC758D1189" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="8F274CAA5B59B95A6E77BE3CFDCE8418" pageId="3" pageNumber="90" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="90" type="type species">
<paragraph pageId="3" pageNumber="90">Type species.</paragraph>
<paragraph pageId="3" pageNumber="90">
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
Facchini, 2011
<subSubSection id="134B4FC3EE9D6A8ECC2ABA9923B045DC" pageId="3" pageNumber="90" type="type species">
<paragraph id="06E83861B982916FC744CF096738A906" pageId="3" pageNumber="90">Type species.</paragraph>
<paragraph id="8C27AF968702D28F07E8015DCA402591" pageId="3" pageNumber="90">
<taxonomicName id="E75D6C9D26982E84035F749874FF7228" authority="Facchini, 2011" authorityName="Facchini" authorityYear="2011" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus Facchini, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="90" type="diagnosis">
<paragraph pageId="3" pageNumber="90">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="90">Medium-sized species (8.0-8.5 mm) for Oriental oodines; habitus semi-elongate; color cyaneus on head and pronotum, piceocyaneus on elytra, with yellowish apical band, piceous to rufopiceous on ventral surface, scape, palpi, tibiae, and tarsomeres red-yellow to yellow; microsculpture isodiametric; integument of body glabrous and impunctate dorsally and ventrally, (only disc of head moderately punctate and striate laterally); head with conspicuously large eyes and minute tempora; frons with punctiform impressions and one pair of punctures; labrum with six setae along anterior margin; clypeus with two setae; labial palpomere glabrous; mentum tooth rounded anteriorly, bordered posteriorly; pronotum moderately transverse, without protruded angles, with anterior and posterior margins of similar width; sides of pronotum finely bordered throughout; anterolateral setae lacking, posterolateral setae present; laterobasal impressions of pronotum faint, sublinear; elytra with fine striae, 1-7 obliterated apically, 5-7 obliterated basally; stria 7 separate to apex (not joined to stria 8), scarcely noticeable apically; stria 8 deeper than other striae, moderately grooved along posterior half; parascutellar striola well-developed, long, located between suture and stria 1; intervals uniformly flat and smooth throughout; intervals 7 and 8 not fused posteriorly, interval 8 not forming ridge externally; interval 9 transformed into marginal gutter at anterior two fifths of elytra, distinct on posterior three fifths; parascutellar pore present; discal pores in interval 3 lacking; interval 7 with two subapical pores; prosternal process unbordered; marginal gutter ended before apex of elytron; mesosternum concave; metepisternum laterally coadunate with elytral epipleuron; sterna 4-5 with pair of minute ambulatory setae; apical sternum with two marginal pores in female; mesocoxa without lateral seta, with posteromedial one; metacoxa with anterior pore; mesofemur without posterior setae; metafemur glabrous or with single pore on dorsal surface; tarsomere 5 of all legs setose ventrally.</paragraph>
<subSubSection id="487D5D42AC2B8382944DEBBADCB733D9" pageId="3" pageNumber="90" type="diagnosis">
<paragraph id="CE346A1E4FB9BCE03FB1DC24A8F7BBCD" pageId="3" pageNumber="90">Diagnosis.</paragraph>
<paragraph id="0B9ABA2A3F7CB8755E285BD5365CC3CC" pageId="3" pageNumber="90">
Medium-sized species (8.0-8.5 mm) for Oriental oodines; habitus semi-elongate; color
<taxonomicName id="862F955AC5BC2626C58713E1C84DC5D7" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cyaneus">cyaneus</taxonomicName>
on head and pronotum, piceocyaneus on elytra, with yellowish apical band, piceous to rufopiceous on ventral surface, scape, palpi, tibiae, and tarsomeres red-yellow to yellow; microsculpture isodiametric; integument of body glabrous and impunctate dorsally and ventrally, (only disc of head moderately punctate and striate laterally); head with conspicuously large eyes and minute tempora; frons with punctiform impressions and one pair of punctures; labrum with six setae along anterior margin; clypeus with two setae; labial palpomere glabrous; mentum tooth rounded anteriorly, bordered posteriorly; pronotum moderately transverse, without protruded angles, with anterior and posterior margins of similar width; sides of pronotum finely bordered throughout; anterolateral setae lacking, posterolateral setae present; laterobasal impressions of pronotum faint, sublinear; elytra with fine striae, 1-7 obliterated apically, 5-7 obliterated basally; stria 7 separate to apex (not joined to stria 8), scarcely noticeable apically; stria 8 deeper than other striae, moderately grooved along posterior half; parascutellar striola well-developed, long, located between suture and stria 1; intervals uniformly flat and smooth throughout; intervals 7 and 8 not fused posteriorly, interval 8 not forming ridge externally; interval 9 transformed into marginal gutter at anterior two fifths of elytra, distinct on posterior three fifths; parascutellar pore present; discal pores in interval 3 lacking; interval 7 with two subapical pores; prosternal process unbordered; marginal gutter ended before apex of elytron; mesosternum concave; metepisternum laterally coadunate with elytral epipleuron; sterna 4-5 with pair of minute ambulatory setae; apical sternum with two marginal pores in female; mesocoxa without lateral seta, with posteromedial one; metacoxa with anterior pore; mesofemur without posterior setae; metafemur glabrous or with single pore on dorsal surface; tarsomere 5 of all legs setose ventrally.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="90" type="etymology">
<paragraph pageId="3" pageNumber="90">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="90">
<subSubSection id="A7057911BCC20394ECF972A3703B8C7A" pageId="3" pageNumber="90" type="etymology">
<paragraph id="B3795596886D122E1A44D74167A5A326" pageId="3" pageNumber="90">Etymology.</paragraph>
<paragraph id="D7B4D9B467FEDC1B6E9162B24106BD83" pageId="3" pageNumber="90">
A compound word, based on the ethnic name of the people in the region where the type species was first found, Bamar, and
<taxonomicName class="Insecta" family="Carabidae" genus="Oodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
<taxonomicName id="E36F6E45FEDB7F99E57079C13D883BCE" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
(for its etymology see
<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="97" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexiko." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
<bibRefCitation id="554DB3AB4DA7751A8084FA5BEE723581" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="97" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexiko." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
: 955). It is treated as a Latin masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="90" type="affinities">
<paragraph pageId="3" pageNumber="90">Affinities.</paragraph>
<paragraph pageId="3" pageNumber="90">
<subSubSection id="2F63C531FFED04B6F2DA4C673962A137" pageId="3" pageNumber="90" type="affinities">
<paragraph id="1D755FC003F0F95CE8500B6A5549BFB5" pageId="3" pageNumber="90">Affinities.</paragraph>
<paragraph id="FD21F36F9ECE6EB0B90BFAC7500A40D3" pageId="3" pageNumber="90">
In 2011, Facchini described eight new species of
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="71A83701B5A545776910C709995F1EF8" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
from the Afrotropical and Oriental region, among them
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="0B6374A70C4576D524F86C745CD96222" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
from Myanmar (
<bibRefCitation author="Facchini, S" journalOrPublisher="Giornale Italiano di Entomologia" pageId="10" pageNumber="97" pagination="381 - 394" title="Otto nuove specie e un nuovo genere di Chlaeniini Brulle, 1834 delle Regioni Afrotropicale e Orientale (Coleoptera Carabidae, Chlaeniini)." volume="12 / 56" year="2011">Facchini 2011</bibRefCitation>
<bibRefCitation id="98CC655AE70FC10BD2FE092C821E2BD7" author="Facchini, S" journalOrPublisher="Giornale Italiano di Entomologia" pageId="10" pageNumber="97" pagination="381 - 394" title="Otto nuove specie e un nuovo genere di Chlaeniini Brulle, 1834 delle Regioni Afrotropicale e Orientale (Coleoptera Carabidae, Chlaeniini)." volume="12 / 56" year="2011">Facchini 2011</bibRefCitation>
: 350-351). The author noted also that the holotype of the species has sympatrically been collected with specimens of
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<taxonomicName id="BAB5CB70515104E69DE01130602BC08E" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
, the type species of the genus. Facchini differentiated the latter from the former by: 1, size of the body; 2, coloration of the integument, including the color of femora and epipleura; 3, shape of the pronotum; 4, shape and striation of the elytra; 5, punctuation of the elytral intervals; 6, chaetotaxy of the last abdominal sternum in the females; 7, distance between the joints of the claws of tarsomere 5. Examination of the paratype and another specimen of
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="A2E3FC741520FB3559A9E6F8EA2587E6" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
from Thailand confirms that these distinguishing features are valid (except for the color of epipleura, which is not markedly different) and well-chosen for ready differentiation of these taxa. However, they are inadequate to give an idea of a more precise systematic position of the species, though some of the differences noted by Facchini are of generic value.
</paragraph>
<paragraph pageId="3" pageNumber="90">
<paragraph id="1EE0A89BBB63B563CD41DBE39C07DB8C" pageId="3" pageNumber="90">
Careful study ascertained significant structural differences between
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="764EF2D820020DCC868F322E3B10F6BF" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<taxonomicName id="FA0EE4465DF94CBFFEB87EF442E26CFD" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
(Table 2).
</paragraph>
<caption ID-Table-UUID="99BBBD73C36DFF01E82A73D81BFEA8E2" httpUri="http://table.plazi.org/id/99BBBD73C36DFF01E82A73D81BFEA8E2" pageId="3" pageNumber="90">
<paragraph pageId="3" pageNumber="90">
<caption id="99BBBD73C36DFF01E82A73D81BFEA8E2" ID-Table-UUID="99BBBD73C36DFF01E82A73D81BFEA8E2" httpUri="http://table.plazi.org/id/99BBBD73C36DFF01E82A73D81BFEA8E2" pageId="3" pageNumber="90">
<paragraph id="6F22983FAFB968890AF40C0C49DA863C" pageId="3" pageNumber="90">
Table 2. Diagnostic character state combinations shown by exemplars of
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<taxonomicName id="1842117E3DC47AABC51E286496BAFDA4" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="95AB7157E37E660E373D242E71DA27D3" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="90">
<table pageId="3" pageNumber="90">
<tr pageId="3" pageNumber="90">
<th colspan="1" pageId="3" pageNumber="90" rowspan="1">No.</th>
<th colspan="1" pageId="3" pageNumber="90" rowspan="1">Characters</th>
<th colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<paragraph id="F6864E398644592DF8BE3FAEE14B4088" pageId="3" pageNumber="90">
<table id="EF6530291B39A362E9E1C233DE1523B7" pageId="3" pageNumber="90">
<tr id="0839D3BCBBD188604436357233F45F9E" pageId="3" pageNumber="90">
<th id="D6B0D6D6492E2926CB2F617A3AD06BF0" colspan="1" pageId="3" pageNumber="90" rowspan="1">No.</th>
<th id="95478EA4E8621A83C31B55401E6C9279" colspan="1" pageId="3" pageNumber="90" rowspan="1">Characters</th>
<th id="223B10D8916EEE5A0B91FFBF76FA880D" colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName id="849C8BEC94387C4F5CC6C8EDDBAB5F2D" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
</th>
<th colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<th id="BC592B72BAB434DFA16FB22504AC2DB9" colspan="1" pageId="3" pageNumber="90" rowspan="1">
<taxonomicName id="FFA516DD4A2C2A56F2B704B7FA7BFD56" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
</th>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">01</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Pubescence of integument</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">partly pubescent</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
<tr id="00BDCF332CAD80193D8E1B4A3FEC0A94" pageId="3" pageNumber="90">
<td id="CF844DDF3099B911982BF3C6614DC80E" colspan="1" pageId="3" pageNumber="90" rowspan="1">01</td>
<td id="FF7BD0AE3783C6B44B02E39AD175C6F1" colspan="1" pageId="3" pageNumber="90" rowspan="1">Pubescence of integument</td>
<td id="7DF06F1BD60C61D8355485ED75383914" colspan="1" pageId="3" pageNumber="90" rowspan="1">partly pubescent</td>
<td id="4C294ED74B79346532F3ECFBBF7CD86F" colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">02</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Punctuation of integument</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 1-3)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 13-16)</td>
<tr id="CF0DBBE59D0DBA5965EE590B24B94208" pageId="3" pageNumber="90">
<td id="9107C69B83EFA2E876F8CDBBAD7A3190" colspan="1" pageId="3" pageNumber="90" rowspan="1">02</td>
<td id="8F3AA88FD68E02C60596E6C270E40E97" colspan="1" pageId="3" pageNumber="90" rowspan="1">Punctuation of integument</td>
<td id="6AE8715B5687E894C1158D0900487C6D" colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 1-3)</td>
<td id="1832BC510C19246EE33F6D2D0CAADDA3" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 13-16)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">03</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Antennomere 3</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">with a few short and scattered setae</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
<tr id="E304700E9810539F05D45060CF59FC31" pageId="3" pageNumber="90">
<td id="E35DC8EC2541817C8BCAB8BAFE884742" colspan="1" pageId="3" pageNumber="90" rowspan="1">03</td>
<td id="39C92F64E27FEE5C175061E056E5579F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Antennomere 3</td>
<td id="AD279B482E19CDBBC5938717033EABC5" colspan="1" pageId="3" pageNumber="90" rowspan="1">with a few short and scattered setae</td>
<td id="82B084E8DBDE0C7E29490FB76B1FCC95" colspan="1" pageId="3" pageNumber="90" rowspan="1">glabrous</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">04</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Mentum tooth form anteriorly</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">truncate</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">rounded</td>
<tr id="A948A51A3E9F705A3D030309317E4BB6" pageId="3" pageNumber="90">
<td id="910A0522DE3CA15EC639E47F6C654C8B" colspan="1" pageId="3" pageNumber="90" rowspan="1">04</td>
<td id="48201F850331EEA9E226D9565EA088DB" colspan="1" pageId="3" pageNumber="90" rowspan="1">Mentum tooth form anteriorly</td>
<td id="B5D912B880A5DF38B560B8A03C730216" colspan="1" pageId="3" pageNumber="90" rowspan="1">truncate</td>
<td id="0D0626BC6602DFE1773966A4C450D03B" colspan="1" pageId="3" pageNumber="90" rowspan="1">rounded</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">05</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">PW/PL</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">1.14-1.15 (Fig. 1)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">1.24-1.37 (Fig. 13)</td>
<tr id="CE9B2692810D9F6E51DD1CB5B495BF6B" pageId="3" pageNumber="90">
<td id="F5944EA1D986E9E0D03E7E5B6747AD1A" colspan="1" pageId="3" pageNumber="90" rowspan="1">05</td>
<td id="251FD0F78F4713CDD2BB9CEC1CD9D5E2" colspan="1" pageId="3" pageNumber="90" rowspan="1">PW/PL</td>
<td id="F204306DC3EA46EF438E4A85BEF58501" colspan="1" pageId="3" pageNumber="90" rowspan="1">1.14-1.15 (Fig. 1)</td>
<td id="A059F3B778F540CB04035D662441D674" colspan="1" pageId="3" pageNumber="90" rowspan="1">1.24-1.37 (Fig. 13)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">06</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Basal margin of pronotum at posterior angles</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">oblique (Fig. 1)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">gradually rounded (Fig. 13)</td>
<tr id="EE77714E8B825484CB6AC4B51E377CBB" pageId="3" pageNumber="90">
<td id="C631B5463679C0CCF0D38F9FD9F3BB40" colspan="1" pageId="3" pageNumber="90" rowspan="1">06</td>
<td id="ED50C207C43DAB01588C47E55D0C065C" colspan="1" pageId="3" pageNumber="90" rowspan="1">Basal margin of pronotum at posterior angles</td>
<td id="0E56ACE258EB5063FE6840C8276FFA0C" colspan="1" pageId="3" pageNumber="90" rowspan="1">oblique (Fig. 1)</td>
<td id="B27F5797FE153E7AD5BEBB594340983E" colspan="1" pageId="3" pageNumber="90" rowspan="1">gradually rounded (Fig. 13)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">07</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Pronotum</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered laterally and lateroapically, not bordered medioapically and basally (Fig. 1)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered throughout (Fig. 13)</td>
<tr id="060FC7B25905A393F83901304BBCFFC9" pageId="3" pageNumber="90">
<td id="7FD85CD8B48308725C66D68234240D08" colspan="1" pageId="3" pageNumber="90" rowspan="1">07</td>
<td id="C50F66EC471B04138D03E4A2D7A19513" colspan="1" pageId="3" pageNumber="90" rowspan="1">Pronotum</td>
<td id="6261DD22DEF0BAA31A1AFDDE9B3D2CAA" colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered laterally and lateroapically, not bordered medioapically and basally (Fig. 1)</td>
<td id="334736F25C09388EBCDA5BC7AACEABB2" colspan="1" pageId="3" pageNumber="90" rowspan="1">bordered throughout (Fig. 13)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">08</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral striae 1-8</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">1-7 punctate, 8 impunctate (Fig. 2)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Fig. 14)</td>
<tr id="90A8A7EDA228FABB673438DB99F6DE65" pageId="3" pageNumber="90">
<td id="9FB63F79EB2634E6B841668D7217151E" colspan="1" pageId="3" pageNumber="90" rowspan="1">08</td>
<td id="5A13AD71A2CBA7208E63C0BA765683CA" colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral striae 1-8</td>
<td id="283CACA735371376B0379F80269EA201" colspan="1" pageId="3" pageNumber="90" rowspan="1">1-7 punctate, 8 impunctate (Fig. 2)</td>
<td id="2731544B78BFA4B3C53BA8714AAAFCBC" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Fig. 14)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">09</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Stria 8</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">not or hardly deeper than other striae (Figs 2-3)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">evidently deeper than other striae (Figs 14-16)</td>
<tr id="726BD1379C5FD754174AA0350E0C2AB8" pageId="3" pageNumber="90">
<td id="0B0C481EC3A2B590D5C91D8A7640D472" colspan="1" pageId="3" pageNumber="90" rowspan="1">09</td>
<td id="355BE19CB9157793D3AA3EA7BBBD00D5" colspan="1" pageId="3" pageNumber="90" rowspan="1">Stria 8</td>
<td id="657F4132603DD8D0F925C2084F12177B" colspan="1" pageId="3" pageNumber="90" rowspan="1">not or hardly deeper than other striae (Figs 2-3)</td>
<td id="147DE81FD010B384E23C239992F849A2" colspan="1" pageId="3" pageNumber="90" rowspan="1">evidently deeper than other striae (Figs 14-16)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">10</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral intervals</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 2-3)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 14-16)</td>
<tr id="1682451D3EF4B34F63039F48947B52BC" pageId="3" pageNumber="90">
<td id="4B677207293E2FFBEAA839FB4A1AC1A2" colspan="1" pageId="3" pageNumber="90" rowspan="1">10</td>
<td id="C281E05D5B3B8166FE4531E522EB3B75" colspan="1" pageId="3" pageNumber="90" rowspan="1">Elytral intervals</td>
<td id="AAAD75B4C7B55D8D446F7749EECE6914" colspan="1" pageId="3" pageNumber="90" rowspan="1">punctate (Figs 2-3)</td>
<td id="A7ABD8FF540411B8DA8F0E9668958045" colspan="1" pageId="3" pageNumber="90" rowspan="1">impunctate (Figs 14-16)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">11</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Interval 9</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">distinct throughout</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">transformed into marginal gutter along anterior 2/5 of elytra, distinct in posterior 3/5</td>
<tr id="65B0F1C702A1A7D4ECFB7CEFEB3D9974" pageId="3" pageNumber="90">
<td id="8446DA3F253748E8CAD6A0121B224326" colspan="1" pageId="3" pageNumber="90" rowspan="1">11</td>
<td id="316DC5417F3E6DB1EEB292A88C3E03FD" colspan="1" pageId="3" pageNumber="90" rowspan="1">Interval 9</td>
<td id="0DFF992C821CAE64B4F57DF3AE66BC74" colspan="1" pageId="3" pageNumber="90" rowspan="1">distinct throughout</td>
<td id="345640A70BC2428FB90779E5888D192C" colspan="1" pageId="3" pageNumber="90" rowspan="1">transformed into marginal gutter along anterior 2/5 of elytra, distinct in posterior 3/5</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">12</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of umbilicate pores</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">23-25</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">15-16</td>
<tr id="7A3DF24850D48224123CDC980C0E86ED" pageId="3" pageNumber="90">
<td id="471EF020DB52F8A3E98C9A2B08F65940" colspan="1" pageId="3" pageNumber="90" rowspan="1">12</td>
<td id="97758029C6C7DB6B210596040C5414B4" colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of umbilicate pores</td>
<td id="068C1E9BA2F7F42F06070675B18D0DC0" colspan="1" pageId="3" pageNumber="90" rowspan="1">23-25</td>
<td id="F7762B1FA618508B47434BC42FB2CD52" colspan="1" pageId="3" pageNumber="90" rowspan="1">15-16</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">13</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Mesocoxa lateral margin</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">with one long seta</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">without seta</td>
<tr id="365091F0178A47C3A77FDB992BDDEBF8" pageId="3" pageNumber="90">
<td id="AAC431D31C15A7A3261721CE199054B8" colspan="1" pageId="3" pageNumber="90" rowspan="1">13</td>
<td id="F4D200014CAC1982965077A7B21B207F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Mesocoxa lateral margin</td>
<td id="0CE02AE9C0AD1068DE6092654BFD7B41" colspan="1" pageId="3" pageNumber="90" rowspan="1">with one long seta</td>
<td id="4006BCDE74B3D16057410A4B1B5843B6" colspan="1" pageId="3" pageNumber="90" rowspan="1">without seta</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">14</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Joins of claws</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">opposite, distantly situated from each other</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">nearly parallel, closely situated to each other (Figs 14-15)</td>
<tr id="21A47FA336DD66E0064CDAFAA45956B7" pageId="3" pageNumber="90">
<td id="4B3B21E381A19066A98EFE34E8B2B74C" colspan="1" pageId="3" pageNumber="90" rowspan="1">14</td>
<td id="038BDF83EE24FF28CFED820C190D044F" colspan="1" pageId="3" pageNumber="90" rowspan="1">Joins of claws</td>
<td id="25F12DDF271070F57603BB6ED388ECC0" colspan="1" pageId="3" pageNumber="90" rowspan="1">opposite, distantly situated from each other</td>
<td id="E30D2CEB84F67A34E0CB7610383893B1" colspan="1" pageId="3" pageNumber="90" rowspan="1">nearly parallel, closely situated to each other (Figs 14-15)</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">15</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of pores on last sternum in female</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">four pores</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">two pores</td>
<tr id="33A9DE33BD2A1007AF0642CA5CAA14E2" pageId="3" pageNumber="90">
<td id="16E862C44C2E23935A71454DE7689AF5" colspan="1" pageId="3" pageNumber="90" rowspan="1">15</td>
<td id="F4B4C0F48497DF6FA1AE223FDC8BB854" colspan="1" pageId="3" pageNumber="90" rowspan="1">Number of pores on last sternum in female</td>
<td id="BD77F3186B7E582F9405EEBFE444F3AA" colspan="1" pageId="3" pageNumber="90" rowspan="1">four pores</td>
<td id="A00D400FCB71C11E37D442499475B910" colspan="1" pageId="3" pageNumber="90" rowspan="1">two pores</td>
</tr>
<tr pageId="3" pageNumber="90">
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">16</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">Apical stylomere of ovipositor</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">subtriangular, as wide as basal stylomere, with 36 long ensiform setae at dorsomedial, dorsolateral and ventral position, and with nematiform setae, shorter than ensiform setae (Figs 10-11)</td>
<td colspan="1" pageId="3" pageNumber="90" rowspan="1">subelongate, nearly twice as narrow as basal stylomere, with 11 short ensiform setae at dorsomedial and dorsolateral position, and with nematiform setae, nearly twice as long as ensiform setae (Figs 17-18)</td>
<tr id="6F051EBD26E4761104D7CBFD864420D3" pageId="3" pageNumber="90">
<td id="0E21256873336B66AB240C4E40BA3CE5" colspan="1" pageId="3" pageNumber="90" rowspan="1">16</td>
<td id="811BC5F1C518D521DFF414154160DDED" colspan="1" pageId="3" pageNumber="90" rowspan="1">Apical stylomere of ovipositor</td>
<td id="74AAD7FE57B01635292E6A01ABE400FE" colspan="1" pageId="3" pageNumber="90" rowspan="1">subtriangular, as wide as basal stylomere, with 36 long ensiform setae at dorsomedial, dorsolateral and ventral position, and with nematiform setae, shorter than ensiform setae (Figs 10-11)</td>
<td id="5AD531FD970148E9E0B6777369C256EF" colspan="1" pageId="3" pageNumber="90" rowspan="1">subelongate, nearly twice as narrow as basal stylomere, with 11 short ensiform setae at dorsomedial and dorsolateral position, and with nematiform setae, nearly twice as long as ensiform setae (Figs 17-18)</td>
</tr>
</table>
</paragraph>
<paragraph pageId="3" pageNumber="90">
<paragraph id="737BD20E1702F161E87C1DBEC2EB965E" pageId="3" pageNumber="90">
Five of the listed character states (1, 2, 3, 9, and 11) are of tribal magnitude, though one of them (i.e., attribute 9) shows transitional conditions in the two species. All the states are characteristic of
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="53455AE12FA2F900AEACB3D7A7A0DB95" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
and atypical for
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="E7653104505ACA30DDC9EE8CB08299D9" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
(
<bibRefCitation author="Jeannel, R" journalOrPublisher="Office de la Recherche scientifique colonial, Paris" pageId="10" pageNumber="97" pagination="767 - 1146" title="Coleopteres carabiques de la region Malgache (troisieme partie). Faune de l'Empire Francais" year="1949 a">Jeannel 1949a</bibRefCitation>
<bibRefCitation id="6B33DE063D18435936FCCB5B08CD8C5E" author="Jeannel, R" journalOrPublisher="Office de la Recherche scientifique colonial, Paris" pageId="10" pageNumber="97" pagination="767 - 1146" title="Coleopteres carabiques de la region Malgache (troisieme partie). Faune de l'Empire Francais" year="1949 a">Jeannel 1949a</bibRefCitation>
,
<bibRefCitation author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
<bibRefCitation id="60AD40C807F3B9096C51831D1939C4E2" author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
). Point 11 exhibits a unique condition in
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="EE6C56BEF37564004B95E0B1110C0F5D" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
that is hitherto unknown in the two tribes. However, a complete character state transformation of this feature occurs in the
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="1ECBA5DA689F71EF01A40E831B9CE79B" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Except for
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n., all other examined oodines possess a ninth interval of elytra completely transformed into a marginal gutter throughout. In my view, this special feature may be morphological evidence for transition between the two tribes, but this needs further examination. The losses of the mesocoxa lateral seta (point 13) and parallel position of the joints of the tarsal claws (attribute 14) are conditions hitherto not found together in the aforementioned tribes. Among the
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="91D93B085A5460911FCC793C16D9D6F6" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="CC2B322DC0B6A3823A4DD4412D5048B3" rank="genus">gen. n.</taxonomicNameLabel>
, all other examined oodines possess a ninth interval of elytra completely transformed into a marginal gutter throughout. In my view, this special feature may be morphological evidence for transition between the two tribes, but this needs further examination. The losses of the mesocoxa lateral seta (point 13) and parallel position of the joints of the tarsal claws (attribute 14) are conditions hitherto not found together in the aforementioned tribes. Among the
<taxonomicName id="76900E41CF6D72FCFBC6D0F0F595B6A3" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, the three character states occur together only in
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="97A7E5397758D8E6E2BE3DBCA3213327" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Thaioodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Thaioodes</taxonomicName>
gen. n. piceus sp. n. I believe that they have arisen as a consequence of a specific adaptation and survival strategy for an aquatic manner of living. The remaining character states (i.e., 4, 5, 6, 7, 8, 10, 12, 15, and 16) demonstrate marked differences of a grade higher than the grade usual for species from one and the same genus. I treat each of these nine differences as of generic significance.
<taxonomicName id="0FC39F3DA36BA5F04A1C733DCD9CB25A" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" lsidName="Thaioodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Thaioodes</taxonomicName>
<taxonomicNameLabel id="0AF9831B2D531F3D08B3DADC3FE253AF" rank="genus">gen. n.</taxonomicNameLabel>
<taxonomicName id="132E4EC9B034652ADD9E6958188085A4" class="Insecta" family="Carabidae" genus="Thaioodes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="piceus" status="sp. nov.">piceus</taxonomicName>
<taxonomicNameLabel id="20B812D4196D84FB080DF4F3BE702D9B" rank="species">sp. n.</taxonomicNameLabel>
I believe that they have arisen as a consequence of a specific adaptation and survival strategy for an aquatic manner of living. The remaining character states (i.e., 4, 5, 6, 7, 8, 10, 12, 15, and 16) demonstrate marked differences of a grade higher than the grade usual for species from one and the same genus. I treat each of these nine differences as of generic significance.
</paragraph>
<paragraph pageId="3" pageNumber="90">
<paragraph id="F8A67C14C08100DD64E0025DE1656F95" pageId="3" pageNumber="90">
In conclusion, there are quite a number of considerable morphological differences between
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
<taxonomicName id="8B2C3E6F40A0F22A8CFB6D73C3A16242" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
<taxonomicName id="FF7D639CB591B24A84C1AA149CFD18A6" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Hololeius cyaneus</taxonomicName>
, which taken together are cause for the separation of the latter in a separate genus. That is why
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n. is proposed to accommodate this species. The new taxon does not share main tribal characters of chlaeniines but shows important similarities with oodines.
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Bamaroodes cyaneus</taxonomicName>
<taxonomicName id="557C5D2678CFDCE9E2BCDADBACB27162" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="4406F75A4E34A0CB0711FDF6BF335110" rank="genus">gen. n.</taxonomicNameLabel>
is proposed to accommodate this species. The new taxon does not share main tribal characters of chlaeniines but shows important similarities with oodines.
<taxonomicName id="165BF415B867F52F2CEA7717DAE8BF51" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes cyaneus" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="cyaneus">Bamaroodes cyaneus</taxonomicName>
is therefore removed to a new tribal placement within
<taxonomicName lsidName="" pageId="3" pageNumber="90" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="B02EAA7A34134B58E2F523B7A0BA24A1" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. The relationships of the new genus are discussed below (see
<normalizedToken originalValue="Discussion">'Discussion'</normalizedToken>
<normalizedToken id="C406DC350D80233E43698550294B2BE5" originalValue="Discussion">'Discussion'</normalizedToken>
).
</paragraph>
</subSubSection>

View file

@ -1,373 +1,385 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864664" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="DD79EC34BB254F9CB1A988B135B86F75" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Holosoma Semenov 1889" docType="treatment" docVersion="5" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="754D6666782B7BFF37530E74B47D60B4" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="DD79EC34BB254F9CB1A988B135B86F75" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Holosoma Semenov 1889" docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="88" updateTime="1731807077730" updateUser="admin">
<mods:mods id="9C5FFB2CDF3DCD84DC14838AC6399AC3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D596453B4F49DC30DF4FF3497ED45F14">
<mods:title id="C4E1CEC4996EC92173EEF2EF2FB50B40">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="F6EC87E088016B511895E3B365DCC02D" type="personal">
<mods:role id="BE53FDF290F5204F8303188E358FBDA7">
<mods:roleTerm id="E160884C9C96C46CC50E59C632B91FAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="4E4B9AB9218418E915C61F102C77F8F7">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="1F1120CD3C337AC5B0846F323BBDE6BF">text</mods:typeOfResource>
<mods:relatedItem id="715C7F1A3556B12EEAF3728416272180" type="host">
<mods:titleInfo id="FA2099AE200CC6B6FA5D6BBDDF51BF52">
<mods:title id="C7DB0982D5AE54C8DFC1F8D1E4515452">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="59967CB09A886511998433EFC62C5DE2">
<mods:date id="0C2417A023F310C608C18ACB9963EDE5">2014</mods:date>
<mods:detail id="E23A8F7B8D855204EF86E8D269E8DB3A" type="volume">
<mods:number id="D74CA09F085B0FAE0C7D1EEC6424550D">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="406375E1B4AB48889FE449E59CDE16DD" type="issue">
<mods:number id="39478FC32328667F94BE56B4131DA32C">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="BFB6BD1170DA2F46209DAA261063DB46" unit="page">
<mods:start id="E985C069CD817BC7EB8F5FAB7CDA02A9">87</mods:start>
<mods:end id="95DE50AD81A3DC10B8E4894C164EBCD9">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="15003C6D36D491B518F45906BA03DCB4">
<mods:url id="85309AB859DED6D4E31122181AD9ECA7">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="0EBF1DF0EBD63C2F164DAEF24E3E2F3E">journal article</mods:classification>
<mods:identifier id="BC4FCF7CFC4B051346C17ED993D577F1" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="67CEB58B5410CBF0EC761C809CF1A441" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864664" LSID="urn:lsid:plazi:treatment:DD79EC34BB254F9CB1A988B135B86F75" httpUri="http://treatment.plazi.org/id/DD79EC34BB254F9CB1A988B135B86F75" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="multiple">
<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma Semenov, 1889</taxonomicName>
<treatment id="DD79EC34BB254F9CB1A988B135B86F75" ID-GBIF-Taxon="127864664" LSID="urn:lsid:plazi:treatment:DD79EC34BB254F9CB1A988B135B86F75" httpUri="http://treatment.plazi.org/id/DD79EC34BB254F9CB1A988B135B86F75" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection id="CA33B2F1E3EB7966DFCD330EF0A84ABB" pageId="1" pageNumber="88" type="multiple">
<paragraph id="88B2ABE1ACC0E853C4BE2C52922492DA" pageId="1" pageNumber="88">
Taxon classification Animalia
<taxonomicName id="1D32566AAD51674BFA9F40C82C07E962" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="2A72E195B93CDC506A9C0700222382DA" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="reference_group">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
Heller, 1923: 66 (type species
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Parahololius weigoldi</taxonomicName>
Heller, 1923)
<subSubSection id="B289E15875E62EF856369ACA4D0173BB" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="E31A375EB41797B8498C767D7E8F03AA" pageId="1" pageNumber="88">
<taxonomicName id="E459DFC055FA63C55219204DB050CBB6" ID-CoL="8KWFN" authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma Semenov, 1889</taxonomicName>
</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Semenov, 1927" authorityName="Semenov" authorityYear="1927" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius</taxonomicName>
Semenov, 1927: 232 [unjustified emendation of
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
</subSubSection>
<subSubSection id="044AEB23350A1D26425D58FD052D2E22" pageId="1" pageNumber="88" type="reference_group">
<paragraph id="C81DDB4052FB3CF59B148DDF6063F771" pageId="1" pageNumber="88">
<taxonomicName id="313F64194F50DE2A2C064E191B6439CD" authority="Heller, 1923: 66" authorityName="Heller" authorityPageNumber="66" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius Heller, 1923: 66</taxonomicName>
(type species
<taxonomicName id="92F818405C1D88B8BCAD674E48211A1C" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Parahololius weigoldi Heller, 1923</taxonomicName>
)
</paragraph>
<paragraph id="40CD222BC277208A289FD5189EB47595" pageId="1" pageNumber="88">
<taxonomicName id="000127F53E8CD4E886314972D4CDF2E3" authority="Semenov, 1927: 232" authorityName="Semenov" authorityPageNumber="232" authorityYear="1927" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius Semenov, 1927: 232</taxonomicName>
[unjustified emendation of
<taxonomicName id="8992F7A09B241DAF7C5F01163E981AC7" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
, not in prevailing usage]
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="type species">
<paragraph pageId="1" pageNumber="88">Type species.</paragraph>
<paragraph pageId="1" pageNumber="88">
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum</taxonomicName>
Semenov, 1889
<subSubSection id="9178E80C73E4B5F1F4E283A001E9A4C0" pageId="1" pageNumber="88" type="type species">
<paragraph id="3E8FC99C156DDBBE49664CF80191A42F" pageId="1" pageNumber="88">Type species.</paragraph>
<paragraph id="C3FDCCCD0CCF81BE57F0665E8005AC5B" pageId="1" pageNumber="88">
<taxonomicName id="BFAB2EC92588A224B93C76A18A70A9F9" authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum Semenov, 1889</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="historical remarks">
<paragraph pageId="1" pageNumber="88">Historical remarks.</paragraph>
<paragraph pageId="1" pageNumber="88">
<bibRefCitation author="Semenov, AP" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou (nouvelle serie)" pageId="11" pageNumber="98" pagination="686 - 693" title="Apercu des genres palearctiques de la tribu des anchomenides (famille des carabiques)." volume="2" year="1889">Semenov (1889</bibRefCitation>
<subSubSection id="89EB04BCEF347B21BE8B761136553870" pageId="1" pageNumber="88" type="historical remarks">
<paragraph id="841598231886AC1E84397B6C7D7DA831" pageId="1" pageNumber="88">Historical remarks.</paragraph>
<paragraph id="3DE17D4A87FF11DAEE9A0DAEB7D10AA3" pageId="1" pageNumber="88">
<bibRefCitation id="4B224BE11B2D22738CCD9183608DB9BC" author="Semenov, AP" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou (nouvelle serie)" pageId="11" pageNumber="98" pagination="686 - 693" title="Apercu des genres palearctiques de la tribu des anchomenides (famille des carabiques)." volume="2" year="1889">Semenov (1889</bibRefCitation>
: 388) proposed the generic name
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="3CF2A3D0AE8FE1619E3B42DEC3BC4775" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
for
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum</taxonomicName>
<taxonomicName id="E2DC0F9EE8A6D1668E2B67F40C1EC33F" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum</taxonomicName>
from South Gansu, China. The author placed the genus in &quot;subtribum Oodidarum, prope genera
<taxonomicName class="Insecta" family="Carabidae" genus="Oodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Oodes</taxonomicName>
Bon. et
<taxonomicName class="Insecta" family="Carabidae" genus="Simous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Simous</taxonomicName>
Chaud, collocandum&quot; (ibid.: 389). This tribal placement has been subsequently accepted (
<bibRefCitation author="Jakobson, GG" editor="Devrien, AF" journalOrPublisher="A. F. Devrien, Sankt-Petersburg" pageId="10" pageNumber="97" pagination="241 - 320" title="Zhuki Rossii i Zapadnoi Evropy" year="1906">Jakobson 1906</bibRefCitation>
<taxonomicName id="751599A709E06B5916AC5B22C44797B1" authority="Bon." authorityName="Bon." class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" lsidName="Oodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Oodes Bon.</taxonomicName>
et
<taxonomicName id="FCB493419697451C6A8F604ECF30E400" authority="Chaud" authorityName="Chaud" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Simous Chaud</taxonomicName>
, collocandum&quot; (ibid.: 389). This tribal placement has been subsequently accepted (
<bibRefCitation id="33D124A4571C73230BF933E4A8D82E21" author="Jakobson, GG" editor="Devrien, AF" journalOrPublisher="A. F. Devrien, Sankt-Petersburg" pageId="10" pageNumber="97" pagination="241 - 320" title="Zhuki Rossii i Zapadnoi Evropy" year="1906">Jakobson 1906</bibRefCitation>
: 310-311). In 1923,
<bibRefCitation author="Heller, KM" journalOrPublisher="Entomologischen Blaetter" pageId="10" pageNumber="97" pagination="61 - 79" title="Die Coleopterenausbeute der Stotzerschen Sze-Tschwan-Expedition." volume="19" year="1923">Heller (1923</bibRefCitation>
<bibRefCitation id="FF0B60B7D33D19F16D4BB64C658A0F68" author="Heller, KM" journalOrPublisher="Entomologischen Blaetter" pageId="10" pageNumber="97" pagination="61 - 79" title="Die Coleopterenausbeute der Stotzerschen Sze-Tschwan-Expedition." volume="19" year="1923">Heller (1923</bibRefCitation>
: 66) described the genus
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
Heller, 1923, for
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Parahololius weigoldi</taxonomicName>
Heller, 1923, from Sichuan, China. He placed this genus in
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="955855CE4662E8D027D93F5872870DDE" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius Heller, 1923</taxonomicName>
, for
<taxonomicName id="75DF3A22C3EFA1751D5157C7CF16325C" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Parahololius weigoldi Heller, 1923</taxonomicName>
, from Sichuan, China. He placed this genus in
<taxonomicName id="8320B541128BD9EAB1011522D6AAEC68" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
, near to
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<taxonomicName id="A0B035AD6FBCF5DDDB01814112E60BC7" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
. Shortly afterwards,
<bibRefCitation author="Semenov, AP" journalOrPublisher="Russkoe Entomologicheskoe Obozrenie" pageId="11" pageNumber="98" pagination="230 - 239" title="Analecta coleopterologica. Koleopterologicheskie zametki. XX." volume="21" year="1927">Semenov (1927</bibRefCitation>
<bibRefCitation id="5AAEFF8A83F07EC4624CE589A1CCB99B" author="Semenov, AP" journalOrPublisher="Russkoe Entomologicheskoe Obozrenie" pageId="11" pageNumber="98" pagination="230 - 239" title="Analecta coleopterologica. Koleopterologicheskie zametki. XX." volume="21" year="1927">Semenov (1927</bibRefCitation>
: 232) proposed [not
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Institut des Parcs Nationaux du Congo Belge" pageId="10" pageNumber="97" pagination="3 - 252" title="Exploration du Parc National de l'Upemba. Mission G. F. De Witte. Carabidae (Coleoptera Adephaga)" year="1953">Basilewsky 1953</bibRefCitation>
<bibRefCitation id="B5D61AAF25A95C4C617ED3C1B5707D42" author="Basilewsky, P" journalOrPublisher="Institut des Parcs Nationaux du Congo Belge" pageId="10" pageNumber="97" pagination="3 - 252" title="Exploration du Parc National de l'Upemba. Mission G. F. De Witte. Carabidae (Coleoptera Adephaga)" year="1953">Basilewsky 1953</bibRefCitation>
, as
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
<bibRefCitation id="6A3EF0768C0FEAC834F0A70C7048DCC1" author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
: 77 stated] the synonymy of
<taxonomicName authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Parahololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
<taxonomicName id="AD866B395CC8CE3822F4FB1CF221FF5E" class="Insecta" family="Carabidae" genus="Parahololius" kingdom="Animalia" lsidName="Parahololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parahololius</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="A51B4D1B12F261C799C178E07DB43E2C" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
and emended the former name to
<taxonomicName authority="Semenov, 1927" authorityName="Semenov" authorityYear="1927" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius</taxonomicName>
<taxonomicName id="62090C10C74DB81096431781C67E2C0B" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius</taxonomicName>
[according to the Article 32.2.3 of
<bibRefCitation author="Heller, KM" journalOrPublisher="International Trust for Zoological Nomenclature, London" pageId="10" pageNumber="97" title="International Code on Zoological Nomenclature, fourth edition, adopted by the International Union of Biological Sciences" url="http://www.iczn.org/iczn" year="1999">ICZN 1999</bibRefCitation>
<bibRefCitation id="EE71167CFF25A423F665F788917BF2CB" author="Heller, KM" journalOrPublisher="International Trust for Zoological Nomenclature, London" pageId="10" pageNumber="97" title="International Code on Zoological Nomenclature, fourth edition, adopted by the International Union of Biological Sciences" url="http://www.iczn.org/iczn" year="1999">ICZN 1999</bibRefCitation>
, the change of the original name to
<taxonomicName authority="Semenov, 1927" authorityName="Semenov" authorityYear="1927" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius</taxonomicName>
<taxonomicName id="F87EF6754186F6DF0C9F984854F14C83" class="Insecta" family="Carabidae" genus="Parololius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Parololius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Parololius</taxonomicName>
is an &quot;unjustified emendation].
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="B97A5BCC3EB226FD433C11986BFA45D7" author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken id="79A9172CF812110847AF332C0700F351" originalValue="Jedlička">Jedlicka</normalizedToken>
(1931
</bibRefCitation>
: 21-22) described
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">Holosoma rambouseki</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1931 from Sichuan, China.
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Arkiv foer Zoologi" pageId="10" pageNumber="97" pagination="8 - 12" title="Schwedisch-chinesische wissenschaftliche Expeditionennachden Nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-Chang. Insecten gesammelt von schwedischen Artz der Expedition Dr. David Hummel 1927 - 1930. 18. Coleoptera. 4. Carabidae und Cicindelidae. C. Harpalinae: Apotomini, Broscini, Pogonini, and Chlaeniini." volume="27 A" year="1935">Andrewes (1935)</bibRefCitation>
<taxonomicName id="C971C4D79318C8353040A79BAFC52469" authority="Jedlicka, 1931" authorityName="Jedlicka" authorityYear="1931" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">
Holosoma rambouseki
<normalizedToken id="ADA58DEB998919D20E1CE2FAA860B469" originalValue="Jedlička">Jedlicka</normalizedToken>
, 1931
</taxonomicName>
from Sichuan, China.
<bibRefCitation id="F740CC88527DDD5FF1FA4EF9CDD15B6F" author="Andrewes, HE" journalOrPublisher="Arkiv foer Zoologi" pageId="10" pageNumber="97" pagination="8 - 12" title="Schwedisch-chinesische wissenschaftliche Expeditionennachden Nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-Chang. Insecten gesammelt von schwedischen Artz der Expedition Dr. David Hummel 1927 - 1930. 18. Coleoptera. 4. Carabidae und Cicindelidae. C. Harpalinae: Apotomini, Broscini, Pogonini, and Chlaeniini." volume="27 A" year="1935">Andrewes (1935)</bibRefCitation>
described
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Chlaenius hedini</taxonomicName>
Andrewes, 1935, from North Gansu and Southeast Sichuan, China. He noted, &quot;It does not appear to be nearly allied to any other Asiatic species.&quot; [of
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
Bonelli, 1810].
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologisches Nachrichtenbaltt" pageId="10" pageNumber="97" pagination="51 - 55" title="Neue Carabiden aus Ostasien. (XI. Teil)." volume="10" year="1936">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<taxonomicName id="1BC785A29E16801663E371CE027980C8" authority="Andrewes, 1935" authorityName="Andrewes" authorityYear="1935" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Chlaenius hedini Andrewes, 1935</taxonomicName>
, from North Gansu and Southeast Sichuan, China. He noted, &quot;It does not appear to be nearly allied to any other Asiatic species.&quot; [of
<taxonomicName id="7D6EEF1437BC6774C317390AF926C971" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaenius Bonelli, 1810</taxonomicName>
].
<bibRefCitation id="78C369888152B6B6ABCD85C75D32F427" author="Jedlicka, A" journalOrPublisher="Entomologisches Nachrichtenbaltt" pageId="10" pageNumber="97" pagination="51 - 55" title="Neue Carabiden aus Ostasien. (XI. Teil)." volume="10" year="1936">
<normalizedToken id="7D93A3197E09DF38F6073DAE2EE73C84" originalValue="Jedlička">Jedlicka</normalizedToken>
(1936
</bibRefCitation>
: 51) described
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1936, from the Philippines, which is the only known extra-Palaearctic record for the genus. Subsequent authors dealing with the genus, except for
<bibRefCitation author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz (2005)</bibRefCitation>
<taxonomicName id="30F914736DAC1BB85505F4C9E7223D67" authority="Jedlicka, 1936" authorityName="Jedlicka" authorityYear="1936" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">
Holosoma boettcheri
<normalizedToken id="5D6900776EC018F8F8AABEF08EE52EFA" originalValue="Jedlička">Jedlicka</normalizedToken>
, 1936
</taxonomicName>
, from the Philippines, which is the only known extra-Palaearctic record for the genus. Subsequent authors dealing with the genus, except for
<bibRefCitation id="75804F8586ED4551BF8C630CB4B692F0" author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz (2005)</bibRefCitation>
, have omitted this species. I have seen the holotype of
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
<taxonomicName id="179DCB15AE7FB9D9B9C1D68D3A9C3758" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
in BMNH and found that it belongs to a different group of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="E3801EC98C0023EAA8EB187C3942B1DD" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Later,
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Institut des Parcs Nationaux du Congo Belge" pageId="10" pageNumber="97" pagination="3 - 252" title="Exploration du Parc National de l'Upemba. Mission G. F. De Witte. Carabidae (Coleoptera Adephaga)" year="1953">Basilewsky (1953</bibRefCitation>
<bibRefCitation id="43C92FBCEA1D36EFEBCF2ED4FAFD7FE5" author="Basilewsky, P" journalOrPublisher="Institut des Parcs Nationaux du Congo Belge" pageId="10" pageNumber="97" pagination="3 - 252" title="Exploration du Parc National de l'Upemba. Mission G. F. De Witte. Carabidae (Coleoptera Adephaga)" year="1953">Basilewsky (1953</bibRefCitation>
: 153) included
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="D42B9F31DD3971E25C1185BD0E043FF6" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
to the tribe
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Simoini">Simoini</taxonomicName>
Basilewsky, 1953, of subfamily
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="subFamily" subFamily="Oodinae">Oodinae</taxonomicName>
<taxonomicName id="062303361F87EB7D9263DBE9DFC717D8" authority="Basilewsky, 1953" authorityName="Basilewsky" authorityYear="1953" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Simoini">Simoini Basilewsky, 1953</taxonomicName>
, of subfamily
<taxonomicName id="B437F4B0F9129030417325703CDF290F" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="subFamily" subFamily="Oodinae">Oodinae</taxonomicName>
(sensu Jeannel, 1949a).
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer (1995)</bibRefCitation>
<bibRefCitation id="1DAA0DD4B20A1E666D091FB9F0B999CA" author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer (1995)</bibRefCitation>
reviewed the known species (excl.
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Chlaenius hedini</taxonomicName>
<taxonomicName id="874EB0A956260A1CBDA407E998F45B37" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Chlaenius hedini</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
<taxonomicName id="28E25F176C45B78DA2EC009B18B70DF6" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
), adding three more species and retaining the tribal affiliation of the genus. Later, he synonymized one of his added taxa with
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Holosoma hedini</taxonomicName>
(
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="10" pageNumber="97" pagination="317 - 332" title="Neue Chlaeniinae der palaearktischen und orientalischen Region (Coleoptera, Carabidae)." url="http://www.landesmuseum.at/pdf_frei_remote/ENT_0019_0317-0329.pdf" volume="19 / 20" year="1998">Kirschenhofer 1998</bibRefCitation>
). Recently,
<bibRefCitation author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito (2003</bibRefCitation>
<taxonomicName id="C5012F9C5EBC2599B40DC063EA038F66" authority="(Kirschenhofer 1998)" baseAuthorityName="Kirschenhofer" baseAuthorityYear="1998" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">
Holosoma hedini (
<bibRefCitation id="9B018113F4B4DE5E4D5D228C312E7807" author="Kirschenhofer, E" journalOrPublisher="Entomofauna" pageId="10" pageNumber="97" pagination="317 - 332" title="Neue Chlaeniinae der palaearktischen und orientalischen Region (Coleoptera, Carabidae)." url="http://www.landesmuseum.at/pdf_frei_remote/ENT_0019_0317-0329.pdf" volume="19 / 20" year="1998">Kirschenhofer 1998</bibRefCitation>
)
</taxonomicName>
. Recently,
<bibRefCitation id="C439CBFC46BF816AAD33CD8485E283C2" author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito (2003</bibRefCitation>
,
<bibRefCitation author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">2012</bibRefCitation>
<bibRefCitation id="E2723146EE2DAA5DE33FD43994E51EA2" author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">2012</bibRefCitation>
) added five more species and one subspecies to the genus and keyed all species known at that time. He retained the position of the genus within the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="D367CA4C49EE1551A5420CC3B007EDC6" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="taxonomic position">
<paragraph pageId="1" pageNumber="88">Taxonomic position.</paragraph>
<paragraph pageId="1" pageNumber="88">
<subSubSection id="8BDC191443FDEE7D495F9A7D22EAFFF7" pageId="1" pageNumber="88" type="taxonomic position">
<paragraph id="20910CA8641A92609A2B40F82D1590C5" pageId="1" pageNumber="88">Taxonomic position.</paragraph>
<paragraph id="741197EA8E15DE4DA76AD6A391019CF8" pageId="1" pageNumber="88">
The discussion here is based on all the generic and species descriptions (
<bibRefCitation author="Semenov, AP" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou (nouvelle serie)" pageId="11" pageNumber="98" pagination="686 - 693" title="Apercu des genres palearctiques de la tribu des anchomenides (famille des carabiques)." volume="2" year="1889">Semenov 1889</bibRefCitation>
<bibRefCitation id="08378EC051CAB3F7DB5E5EA03832DBD4" author="Semenov, AP" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou (nouvelle serie)" pageId="11" pageNumber="98" pagination="686 - 693" title="Apercu des genres palearctiques de la tribu des anchomenides (famille des carabiques)." volume="2" year="1889">Semenov 1889</bibRefCitation>
,
<bibRefCitation author="Heller, KM" journalOrPublisher="Entomologischen Blaetter" pageId="10" pageNumber="97" pagination="61 - 79" title="Die Coleopterenausbeute der Stotzerschen Sze-Tschwan-Expedition." volume="19" year="1923">Heller 1923</bibRefCitation>
<bibRefCitation id="C8153273B7233CFA5A2198CC8A0CFB5E" author="Heller, KM" journalOrPublisher="Entomologischen Blaetter" pageId="10" pageNumber="97" pagination="61 - 79" title="Die Coleopterenausbeute der Stotzerschen Sze-Tschwan-Expedition." volume="19" year="1923">Heller 1923</bibRefCitation>
,
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="E2652F7744FCE03EE39827D9305DB144" author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken id="9C6BAAD59A01D44B467BBEEE538EF4E4" originalValue="Jedlička">Jedlicka</normalizedToken>
1931
</bibRefCitation>
,
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Arkiv foer Zoologi" pageId="10" pageNumber="97" pagination="8 - 12" title="Schwedisch-chinesische wissenschaftliche Expeditionennachden Nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-Chang. Insecten gesammelt von schwedischen Artz der Expedition Dr. David Hummel 1927 - 1930. 18. Coleoptera. 4. Carabidae und Cicindelidae. C. Harpalinae: Apotomini, Broscini, Pogonini, and Chlaeniini." volume="27 A" year="1935">Andrewes 1935</bibRefCitation>
<bibRefCitation id="613FE419D596AA8D5728B8D155F0FF54" author="Andrewes, HE" journalOrPublisher="Arkiv foer Zoologi" pageId="10" pageNumber="97" pagination="8 - 12" title="Schwedisch-chinesische wissenschaftliche Expeditionennachden Nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-Chang. Insecten gesammelt von schwedischen Artz der Expedition Dr. David Hummel 1927 - 1930. 18. Coleoptera. 4. Carabidae und Cicindelidae. C. Harpalinae: Apotomini, Broscini, Pogonini, and Chlaeniini." volume="27 A" year="1935">Andrewes 1935</bibRefCitation>
,
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
<bibRefCitation id="580FF115A1853B42D22F4CFCE81BDCAA" author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
,
<bibRefCitation author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito 2003</bibRefCitation>
<bibRefCitation id="C0D4550B2A278628A2480725A45EB477" author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito 2003</bibRefCitation>
,
<bibRefCitation author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">2012</bibRefCitation>
<bibRefCitation id="61A093DBEC9DFB6F84491D4CFAF96B72" author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">2012</bibRefCitation>
) and on the detailed examination of two specimens. It considers only the characters that are significant for the tribal position of
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="C08DC4B2F3BEC6E986BDE233D1C44158" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
. Excluding
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
<taxonomicName id="92CC1B5FAD7ED3C1961DF7CED2ECA7ED" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma boettcheri" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boettcheri">Holosoma boettcheri</taxonomicName>
, I am aware that the group is homogeneous and the main structural features are uniform among the species.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="71237A022727041DFF1BDB904B688697" pageId="1" pageNumber="88">
Pronotum posterior margin as wide as basal margin of elytra, thus habitus seems semi-oval rather than elongate. Atypical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="9EEB424695A25237C752E595F4FDD9E6" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
(but occurs in a few species, such as the Nearctic
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius tomentosus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="tomentosus">Chlaenius tomentosus</taxonomicName>
(Say, 1823); remark by R. Davidson). Typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="94C93CC60D765A10664829CF473CD8B9" authority="(Say, 1823)" baseAuthorityName="Say" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius tomentosus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="tomentosus">Chlaenius tomentosus (Say, 1823)</taxonomicName>
; remark by R. Davidson). Typical of
<taxonomicName id="C2DD8CC21E4B15F7EEE597BD7C5C2AD0" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, except for the new genera. This condition is probably a derived trend within the genus because it occurs in most, but not all, species.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="A7B8C43E02C7A589147C6631DB8BCE5F" pageId="1" pageNumber="88">
Body dorsally with metallic lustre (greenish, turquoise, bluish, violet to black-blue) on dorsal surface. Habitual to
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="9FB0F7B607C0D347F2033FF268773F53" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
, since many species from this tribe are metallic colored. In the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="C73A65120BD0A97B28260BC49427F2E8" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, a metallic hue is present only in
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n., most species of
<taxonomicName class="Insecta" family="Carabidae" genus="Simous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Simous</taxonomicName>
<taxonomicName id="84D259195DE2A9FFCF8CEE56A97D8EE9" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="078FCCF11D5263EE4E75389A7D0F12BC" rank="genus">gen. n.</taxonomicNameLabel>
, most species of
<taxonomicName id="40E81CEBCF1FE1E1E2D9F07C69CD9286" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" lsidName="Simous" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Simous</taxonomicName>
and a few taxa of
<taxonomicName class="Insecta" family="Carabidae" genus="Stenocrepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocrepis" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Stenocrepis</taxonomicName>
Chaudoir, 1857.
<taxonomicName id="80731021D5DC13231E79BB45B7AD76B4" authority="Chaudoir, 1857" authorityName="Chaudoir" authorityYear="1857" class="Insecta" family="Carabidae" genus="Stenocrepis" kingdom="Animalia" lsidName="Stenocrepis" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Stenocrepis Chaudoir, 1857</taxonomicName>
.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="E08DFE610634A59FF0F4913CCFA3F682" pageId="1" pageNumber="88">
Integument sparsely pubescent dorsally and ventrally. The character is distinctive of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="9FA317544D9832D61668CA4B6B7B9CA8" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
, but it is unknown in the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="4E1372DD10E8F54286467157C0AF9DAF" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. In the species of
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="1D61CAD1B1A03FB2D0E03C887DFBF7EC" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
, the dorsal surface of the head, antennomere 3 (excl. apical setae), pronotum, intervals 8-9 of elytra, prosternum, mesepisternum, mesocoxa, mesofemur, metasternum, metepisternum, and abdomen all have rather fine and scattered punctures (see also
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
<bibRefCitation id="4FC1F7B68A5BEE16F3A27E180A2DE696" author="Kirschenhofer, E" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="10" pageNumber="97" pagination="77 - 84" title="Neue Arten Gattung Holosoma Semenov, 1889 aus China (Col. Carabidae, Oodiinae, Trib. Simoini)." url="http://www.landesmuseum.at/pdf_frei_remote/ZAOE_47_0077-0084.pdf" volume="47" year="1995">Kirschenhofer 1995</bibRefCitation>
,
<bibRefCitation author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito 2003</bibRefCitation>
<bibRefCitation id="D90FDBC3266E14B242BC9A744780A76C" author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito 2003</bibRefCitation>
). Most of the punctures are provided with short, yellowish hairs, usually well visible under higher magnification. Sparse pubescence is also present on the medial elytral intervals posteriorly, though it is much more sporadic than on the intervals 8-9.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="A2BA937A8EAB8FCE0C9BED276956D013" pageId="1" pageNumber="88">
Labrum with six setae along anterior margin. This feature is typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="DBEFC36521C0EBCB14FCF336C9A7A30C" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Although it is present in most
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="51BCD8415A4F34A955A37850F87E6883" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, several groups have different setation of the labrum.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="97C07949B43D710775F0A9AAC4004AE6" pageId="1" pageNumber="88">
Clypeus with a pair of setae. The condition is usual for
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="12F4355B6016ECD4F4836F2E6FD65AD3" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Although it occurs in most
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="1E65AA146311A74AE652ECA869FFF14F" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, several groups lack clypeal setae.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="5966C05DF4F42C78FC108EB1D9BC61BD" pageId="1" pageNumber="88">
Penultimate labial palpomere with 2-4 spines at front margin. Indicative of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="F6E85A6B6407BB9A92458552082BF7DB" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
(occurs in the most of the species). This feature is unknown in the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="E080AA194966DC5712382069A5384BBB" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
.
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="6A34148C92F0B5314F143F1B7ED83C40" author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovnske Spolecnosti Entomologicke" pageId="10" pageNumber="97" pagination="21 - 30" title="Neue Carabiden aus China-Szetschuan." volume="1 / 2" year="1931">
<normalizedToken id="447F9A61C2E72D37DE2594082F99C9AE" originalValue="Jedlička">Jedlicka</normalizedToken>
(1931
</bibRefCitation>
: 22) has noted that the penultimate labial palpomere in
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">Holosoma rambouseki</taxonomicName>
<taxonomicName id="08126F5069DECA5DF03D41AF19934FFB" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">Holosoma rambouseki</taxonomicName>
lack setae, but this fact needs verification.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="A0D01005AE647D6F2B28E1917985439D" pageId="1" pageNumber="88">
Terminal labial palpomere with a few fine and short setae on lateral margin (see also
<bibRefCitation author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">Ito 2012</bibRefCitation>
<bibRefCitation id="E525B4EF1D6CE96B20A532DB0D6BCCB6" author="Ito, N" journalOrPublisher="Elytra, Tokyo (New Series)" pageId="10" pageNumber="97" pagination="303 - 310" title="Two New Species and One Subspecies of the Genus Holosoma from China (Coleoptera, Carabidae, Oodini)." volume="2" year="2012">Ito 2012</bibRefCitation>
: 303). Occurs in some
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="CE3E0498D444037F184C43CFF5C7BEBB" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Unknown in the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="D56930215977C805C5B4FFDAF5B4ED9A" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="87D8946F5A414BB07496DCFD51DC2C42" pageId="1" pageNumber="88">
Elytral stria 8 shallower than, or as deep as striae 1-7. Typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="3AD2B534FCD39CFAC2ABF10AE11DD7C6" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
, except for
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
<taxonomicName id="0AE037031C73C027F28493E1BD43DE90" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
. All taxa of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="3753E7280F03501233BBC4E03D88EFC5" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
I have studied have stria 8 more or less grooved along its extent and deeper than other striae.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="B2050A6155748BD0155A35939ABE0A7B" pageId="1" pageNumber="88">
Discal setiferous punctures situated in elytral intervals 3 and 5, or in intervals 3, 5 and 7. There is no data for this condition in
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="60D645D3188D5A80BCC44E7745EA2CEB" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
, but it is also atypical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="3D77435E0EEF3C599E887E8DE2816034" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Like point 1, it can be an apotypic trend within the genus since it occurs in several, but not in all species of
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="EB1FDFB76F85350DA1189D8EA27D80E4" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
. For example,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Holosoma hedini</taxonomicName>
<taxonomicName id="77E0E260AF48236AA7A920C20FD586B0" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma hedini" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="hedini">Holosoma hedini</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma heros" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="heros">Holosoma heros</taxonomicName>
Kirschenhofer, 1995, and the specimens from Wenxian possess setiferous punctures in intervals 3, 5 and 7.
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma namikoae" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="namikoae">Holosoma namikoae</taxonomicName>
Ito, 2012, has such punctures only in intervals 3 and 5, and
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">Holosoma rambouseki</taxonomicName>
<taxonomicName id="84D828B60040258671F9DE84816E21C0" authority="Kirschenhofer, 1995" authorityName="Kirschenhofer" authorityYear="1995" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma heros" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="heros">Holosoma heros Kirschenhofer, 1995</taxonomicName>
, and the specimens from Wenxian possess setiferous punctures in intervals 3, 5 and 7.
<taxonomicName id="897039C8E61AB0D1998E7905E7C68C74" authority="Ito, 2012" authorityName="Ito" authorityYear="2012" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma namikoae" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="namikoae">Holosoma namikoae Ito, 2012</taxonomicName>
, has such punctures only in intervals 3 and 5, and
<taxonomicName id="160FA75E76F5A0D8B7338870273C7AB9" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma rambouseki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="rambouseki">Holosoma rambouseki</taxonomicName>
solely in interval 3. Discal punctures are lacking in
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum</taxonomicName>
<taxonomicName id="37EDF2B4EDE810677F53081456A5514F" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma opacum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="opacum">Holosoma opacum</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma nigritum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nigritum">Holosoma nigritum</taxonomicName>
Ito, 2003,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma imurai" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="imurai">Holosoma imurai</taxonomicName>
<taxonomicName id="67FA91BB3329AE6B0FF01DBDF7C6D34A" authority="Ito, 2003" authorityName="Ito" authorityYear="2003" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma nigritum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nigritum">Holosoma nigritum Ito, 2003</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma speciosum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="speciosum">Holosoma speciosum</taxonomicName>
Ito, 2003, and
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma misaoae" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="misaoae">Holosoma misaoae</taxonomicName>
Ito, 2012. This character has not been described for
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Holosoma weigoldi</taxonomicName>
<taxonomicName id="F1680EEA0F6BE80A67EFCAF56BA9F6DA" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma imurai" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="imurai">Holosoma imurai</taxonomicName>
,
<taxonomicName id="FF920665766273EF33D49733D33E1CA6" authority="Ito, 2003" authorityName="Ito" authorityYear="2003" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma speciosum" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="speciosum">Holosoma speciosum Ito, 2003</taxonomicName>
, and
<taxonomicName id="729F7183348308C06DFE857FE1DFE387" authority="Ito, 2012" authorityName="Ito" authorityYear="2012" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma misaoae" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="misaoae">Holosoma misaoae Ito, 2012</taxonomicName>
. This character has not been described for
<taxonomicName id="6BBE828E8C77A0FF425C9A9A67F9C00F" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma weigoldi" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="weigoldi">Holosoma weigoldi</taxonomicName>
.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="2DF323B342C0F9CA5C2BC00456B434FB" pageId="1" pageNumber="88">
Elytral intervals 7 and 8 separate (e.g., not fused) posteriorly, thus stria 7 perceptible to apex. Typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="E685F4B22AF628071F928223B3D9AF59" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Unusual for
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="C98CD62ED9ACB87ADDC7E1E69E70A27D" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, except for
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n.
<taxonomicName id="E64C7B1D5A0F82348DCCA4062A750941" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="37224ED6DB1D5B51D7980087B9A2D031" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="5E09964BCAABCA19F4C7AD58E337E7A1" pageId="1" pageNumber="88">
Elytral interval 8 not forming ridge laterally. Typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="859B99904B824A1759FBBBF9B2BEA208" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Unusual for
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="9B7BE52F3795D39A0B4918E243A31B98" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, except for
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
gen. n.
<taxonomicName id="7C46C5424A7AF9CEE216A5A56DCFC9F8" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
<taxonomicNameLabel id="1855FDEF53CA9FCB8D99A95E34EE7AF8" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="9A738039A82DB3EF8D184DA2D2D82962" pageId="1" pageNumber="88">
Tarsomere 5 of all legs setose ventrally. Typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="278B08028338C9718C22EBB07B82C607" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Unusual for
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="2462E90EA01CAB35FF368492C67CA601" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
, except for the two new genera and a few species of
<taxonomicName class="Insecta" family="Carabidae" genus="Systolocranius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systolocranius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Systolocranius</taxonomicName>
<taxonomicName id="062BAE73577311304D71A2A7ED9DC9C0" class="Insecta" family="Carabidae" genus="Systolocranius" kingdom="Animalia" lsidName="Systolocranius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Systolocranius</taxonomicName>
. Among the species of
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="DD58BA1FFFAEDB2A84E84980A20611B3" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
, the number of the setae varies from two to six on each side of tarsomere 5.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="FD559AC11FE3B41223FF1D9D3BEAABBA" pageId="1" pageNumber="88">
Quinone-like smell defensive secretion. This is one of three groups of compounds used for defence in the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="C585E4329DBC58509EBF1C7AF6A985A8" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. It is not found in
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="51CBD96D1CEE2E0284C9A7891D2A8B2E" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
.
<bibRefCitation author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito (2003</bibRefCitation>
<bibRefCitation id="B8A021ED59861FC0A482AE6ADDFDDE41" author="Ito, N" journalOrPublisher="The Entomological Review of Japan" pageId="10" pageNumber="97" pagination="95 - 104" title="Note on Species of the Genus Holosoma (Coleoptera: Carabidae: Oodinae: Simoini)." volume="58" year="2003">Ito (2003</bibRefCitation>
: 95) noted that the defensive chemical in
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="0752673A9996397835A18E5F80CD1460" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
is &quot;also the same as that of the genus
<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
<taxonomicName id="97AB9EE37A3FB21C721ADE9129A597E7" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
&quot;. I noticed this pungent smell many times when was taking the specimens from Wenxian out of the test-tube and handling them. The odor is identical or similar to that existing in the European species of
<taxonomicName authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" class="Insecta" family="Carabidae" genus="Chlaeniellus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chlaeniellus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaeniellus</taxonomicName>
Reitter, 1908 (
<bibRefCitation author="Bousquet, Y" journalOrPublisher="The Coleopterists Bulletin" pageId="10" pageNumber="97" pagination="165 - 166" title="Notes about the relationships of the Callistini (= Chlaeniini) (Coleoptera: Carabidae)." volume="41" year="1987">Bousquet 1987</bibRefCitation>
<taxonomicName id="C6D1792ABB7D5072CB3AD40D6DCAA4BF" authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" class="Insecta" family="Carabidae" genus="Chlaeniellus" kingdom="Animalia" lsidName="Chlaeniellus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Chlaeniellus Reitter, 1908</taxonomicName>
(
<bibRefCitation id="1AF15779B10BE2DDB0F9E7C211EE32C0" author="Bousquet, Y" journalOrPublisher="The Coleopterists Bulletin" pageId="10" pageNumber="97" pagination="165 - 166" title="Notes about the relationships of the Callistini (= Chlaeniini) (Coleoptera: Carabidae)." volume="41" year="1987">Bousquet 1987</bibRefCitation>
).
<bibRefCitation author="Moore, BP" editor="Erwin, TL" journalOrPublisher="Junk, The Hague" pageId="11" pageNumber="98" pagination="193 - 203" title="Chemical defence in carabids and its bearing on phylogeny." url="http://link.springer.com/chapter/10.1007/978-94-009-9628-1_9" volumeTitle="Carabid beetles: their evolution, natural history, and classification" year="1979">Moore (1979</bibRefCitation>
<bibRefCitation id="F1AA8A81C77A4F75ADC59EE472CAEE41" author="Moore, BP" editor="Erwin, TL" journalOrPublisher="Junk, The Hague" pageId="11" pageNumber="98" pagination="193 - 203" title="Chemical defence in carabids and its bearing on phylogeny." url="http://link.springer.com/chapter/10.1007/978-94-009-9628-1_9" volumeTitle="Carabid beetles: their evolution, natural history, and classification" year="1979">Moore (1979</bibRefCitation>
: 198-199) regarded the quinones as one of the most elaborate defensive strategies in the ground beetles.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="33337E82A14333E7EA1007504BC48989" pageId="1" pageNumber="88">
Thirteen character states are considered. Number 9 is not counted due to deficient data about its presence among other taxa. Six character states, i.e., 2, 3, 6, 7, 8, 13, are typical for
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="85316F198EBC3E341F7B4B5AB1827B7E" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
and are unknown to
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="EA3B6AB369E9A3ED708CAD6CC68CFF4D" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Characters 10, 11 and 12 are also typical of the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="CB390F2A4CBF17452B4FCD39BAF9F22A" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
and have a few exceptions in the
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="53F3A2D153096828011B44C409345C68" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
. Similarly, characters 4 and 5 are always indicative of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="3C979A6E425E7450EFE799A8990C3A04" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. Most genera and species of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="DF0069FC912582483369BFF63B8F99D0" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
also share these two conditions, but there are some important exceptions. Character 1 is the only one characteristic of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
<taxonomicName id="E79CA71EB6DBA84F8D0C0F5BC7E7F191" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
and not typical of
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="EB8CFC5BA6A10B7FF38BB278A632BCDD" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
.
</paragraph>
<paragraph pageId="1" pageNumber="88">
<paragraph id="EF4471BA0EAE2B1C87993496D9693B91" pageId="1" pageNumber="88">
In conclusion,
<taxonomicName class="Insecta" family="Carabidae" genus="Holosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
<taxonomicName id="4C07801BF97E37CB5B357EB3483A10D1" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" lsidName="Holosoma" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Holosoma</taxonomicName>
lacks oodine characters but does share important traits with the chlaeniines. It is therefore removed to a new tribal placement incertae sedis within
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
<taxonomicName id="0DC967A2CF1A50895F0D0750739F9DD2" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
. The precise affinity of the genus within the tribe remains unresolved.
</paragraph>
</subSubSection>

View file

@ -1,66 +1,70 @@
<document ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-GBIF-Taxon="127864669" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="FA624B2097B6EE5E5DB8DA9C4F741108" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="CHLAENIINI Brulle 1834" docType="treatment" docVersion="5" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1643525315145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
<document id="BF8D981F6C737F7EC2BCBDB26FFC4206" ID-DOI="http://dx.doi.org/10.3897/dez.61.7754" ID-GBIF-Dataset="fc82b1a1-2343-4735-a7e2-11e37ab26281" ID-Pensoft-Pub="1860-1324-2" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-1324-2" ModsDocOrigin="Deutsche Entomologische Zeitschrift 61 (2)" ModsDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" checkinTime="1418151589113" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav" docDate="2014" docId="FA624B2097B6EE5E5DB8DA9C4F741108" docLanguage="en" docName="DeutEntomolZeit 61(2): 87-104" docOrigin="Deutsche Entomologische Zeitschrift 61 (2)" docSource="http://dx.doi.org/10.3897/dez.61.7754" docTitle="Chlaeniini Brulle 1834" docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="40197A1CFF8EFFD1FFEDB042E16CFF8D" masterDocTitle="Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="88" updateTime="1731807077730" updateUser="admin">
<mods:mods id="457D8AEDC07BEEC9B7C6B39899CCA4CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="47E8A2A8D33B9E4B95C3098BBAD57A88">
<mods:title id="CB37700838DE970BEE42CA3F03AAB006">Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="F8E95D3104FD9EE1FAABAFD8EE5D73F9" type="personal">
<mods:role id="CA21B9CCF5F7CA979AC51E8D03EE8C4C">
<mods:roleTerm id="D5BFE0933C911F1284AF46F1E90A706D">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:namePart id="4D8C8E4AB2357B60BE3DA826F227CF95">Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
<mods:typeOfResource id="7908A4A765A791F7AF143077A8683A70">text</mods:typeOfResource>
<mods:relatedItem id="67D7BDB6E31C8B7EFA15C922B9D67FA0" type="host">
<mods:titleInfo id="28ED26A381EA09E8D3C5304FC8CAEB05">
<mods:title id="8EE813114F59A6C4BAD9A2776FABFFC2">Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>61</mods:number>
<mods:part id="6D391C55E33EB3FA4F29FAC2974F756F">
<mods:date id="45196A3D8866FA658281A00FEF9B38EA">2014</mods:date>
<mods:detail id="8C59354AA80A6E008636CAC1F342F1B9" type="volume">
<mods:number id="1393AC7154BE5392A7766D7960BD14F1">61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
<mods:detail id="9454AAD6269C2010C3A4514E7BC6DF08" type="issue">
<mods:number id="1B28141257AA6E261D4BC447BC0AA625">2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>104</mods:end>
<mods:extent id="D4C01FF9349FC29121688F07C6D7D6AB" unit="page">
<mods:start id="C09258EE83A37C4F1C29C50AB2E69427">87</mods:start>
<mods:end id="EE9B923577A239C1F35568D69EA368F0">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.61.7754</mods:url>
<mods:location id="3935A5E124453040BBF2CBF79280A6F9">
<mods:url id="DF255C3D2BBA6FCE67F908C582361CC3">http://dx.doi.org/10.3897/dez.61.7754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2</mods:identifier>
<mods:classification id="FD4C78D78CD4D0E1D810F229010A8963">journal article</mods:classification>
<mods:identifier id="AAA88B8D0D30566654AA5BF8E676CB0A" type="DOI">http://dx.doi.org/10.3897/dez.61.7754</mods:identifier>
<mods:identifier id="648BCE38B85E2289D102AC8EB1C11B8D" type="Pensoft-Pub">1860-1324-2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864669" LSID="urn:lsid:plazi:treatment:FA624B2097B6EE5E5DB8DA9C4F741108" httpUri="http://treatment.plazi.org/id/FA624B2097B6EE5E5DB8DA9C4F741108" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="multiple">
<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
<treatment id="FA624B2097B6EE5E5DB8DA9C4F741108" ID-GBIF-Taxon="127864669" LSID="urn:lsid:plazi:treatment:FA624B2097B6EE5E5DB8DA9C4F741108" httpUri="http://treatment.plazi.org/id/FA624B2097B6EE5E5DB8DA9C4F741108" lastPageNumber="88" pageId="1" pageNumber="88">
<subSubSection id="B7E8739FEBE769DB4917599EF7B4A5D9" pageId="1" pageNumber="88" type="multiple">
<paragraph id="7BEE80C6D195D5BD40DA18DDB9EA877A" pageId="1" pageNumber="88">
Taxon classification Animalia
<taxonomicName id="84B651D42FB6BA7F4CF7FCA7A9D76C9B" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="90E25C34A01B948EDE024281D745057E" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph pageId="1" pageNumber="88">
<taxonomicName authority="Brulle, 1834" authorityName="Brulle" authorityYear="1834" lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="CHLAENIINI">
<subSubSection id="B72395A416D011373E35344C1C1A25A0" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="41A28FEF80F83F6E12BD7125B123707F" pageId="1" pageNumber="88">
<taxonomicName id="E71CAC8713E8B01DC859528BFB98519E" ID-CoL="8KTTH" authority="Brulle, 1834" authorityName="Brulle" authorityYear="1834" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">
CHLAENIINI
<normalizedToken originalValue="Brullé">Brulle</normalizedToken>
<normalizedToken id="3D9125BEFF8D689D18770F57B450E572" originalValue="Brullé">Brulle</normalizedToken>
, 1834
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="tribal diagnosis">
<paragraph pageId="1" pageNumber="88">Tribal diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="88">
<subSubSection id="5216088B39F4B3AAFFD2F5E60E282993" pageId="1" pageNumber="88" type="tribal diagnosis">
<paragraph id="D792F6CB868B0848F91CF69356E54528" pageId="1" pageNumber="88">Tribal diagnosis.</paragraph>
<paragraph id="DBAE81E8AAF02DE1C960C9319520490A" pageId="1" pageNumber="88">
Mandibular scrobe without seta near anterior margin of excavation, mesocoxal cavities conjunct (mesepimeron does not reach middle coxal cavity), metepisternum separated from metepimeron by distinct suture, and parameres without setae, with left paramere
<normalizedToken originalValue="“conchoid”">&quot;conchoid&quot;</normalizedToken>
<normalizedToken id="66386AC589402BE8203BFED6045931B0" originalValue="“conchoid”">&quot;conchoid&quot;</normalizedToken>
, therefore a member taxon of Conchifera Jeannel, 1941 (
<bibRefCitation author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
<bibRefCitation id="2B7928984904D25C061A194BD2B80E57" author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
: 1047); head with one pair of supraorbital setae, its capsule
<normalizedToken originalValue="“normal”">&quot;normal&quot;</normalizedToken>
<normalizedToken id="D46D815FE909E9C2D461EAD4AC97F4A1" originalValue="“normal”">&quot;normal&quot;</normalizedToken>
, i.e. without annular constriction behind eyes and with ultimate maxillary palpomere elongate and well-centered with respect to penultimate palpomere, pronotum usually without anterolateral setae, anterior coxal cavities biperforate, elytron with internal plica, metepisternum laterally coadunate with elytral epipleuron, middle femur posterior margin without longer setae, metacoxa lacking posterior seta, and median lobe of aedeagus in lateral view long and considerably bent, with basal bulb decreased, indicate a member taxon of
<taxonomicName family="Callistidae" lsidName="" pageId="1" pageNumber="88" rank="family">Callistidae</taxonomicName>
<taxonomicName id="59DAE00025D594676585AFB3178ED849" family="Callistidae" lsidName="" pageId="1" pageNumber="88" rank="family">Callistidae</taxonomicName>
Jeannel, 1941 (
<bibRefCitation author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
<bibRefCitation id="A5FC7B0524645C8FEC1E2EF34B0B2801" author="Jeannel, R" editor="Grasse, P-P" journalOrPublisher="Masson" pageId="10" pageNumber="97" pagination="1026 - 1077" title="Ordre des Coleopteres (Coleoptera Linne, 1758). Partie Systematique (part)." volumeTitle="Traite de Zoologie. Anatomie, Systematique, Biologie" year="1949 b">Jeannel 1949b</bibRefCitation>
: 1051); tegument largely punctate and pubescent, antennomere 3 densely pubescent or at least with a few short, scattered setae (in addition to apical setae), elytron with intervals 7 and 8 not fused and not forming ridge posteriorly, stria 7 distinct to apex, stria 8 not deeper than other striae, and interval 9 normal (not transformed), wider or narrower, are other features peculiar to tribe.
</paragraph>
</subSubSection>