288 lines
36 KiB
XML
288 lines
36 KiB
XML
<document id="5C9F2107F14BD227E51A162862A8AD2E" ID-DOI="10.11646/zootaxa.5543.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386385" ID-ZooBank="8A21E43A-5605-4238-97B2-81CCA432A503" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1733918705237" checkinUser="plazi" docAuthor="Omelko, Mikhail M. & Fomichev, Alexander A." docDate="2024" docId="B20419093769FF9569A0FDAD01F69D2B" docLanguage="en" docName="zootaxa.5543.3.5.pdf" docOrigin="Zootaxa 5543 (3)" docSource="https://doi.org/10.11646/zootaxa.5543.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bowie tangalunga Omelko & Fomichev, 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="395" masterDocId="4E3D61713763FF996937FF91026F9F4A" masterDocTitle="New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia" masterLastPageNumber="403" masterPageNumber="383" pageNumber="393" updateTime="1734117937186" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:mods id="B402C5906172C50E905C95D3F1DE4F51" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CBA0255446151F9C130418C223261E32">
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<mods:title id="033A41AB3AC8A384C26F5FB09218A7AC">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name id="68D890B5FB5CAAB54254CF9FC4D3130D" type="personal">
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<mods:roleTerm id="8CDD5F08D45B964B928B1B471784F6F0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="124DB3A58F6604C7D65FA81C3257E310">Omelko, Mikhail M.</mods:namePart>
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<mods:nameIdentifier id="AF3FD38077CA750FF0CD94C33723A353" type="ORCID">0000-0002-1556-6248</mods:nameIdentifier>
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<mods:affiliation id="431BD2D073BE23BEC15777B53EFC55C2">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="95F05FF040E51031F148A6B01322DEBA" type="email">omelkom@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="7CFA368C7F361BE3E91053601A2FDE79" type="personal">
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<mods:roleTerm id="DC370698944372C5C9619E914592BDED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4798C2F81CD0A272576BB9ECA1D8FCBC">Fomichev, Alexander A.</mods:namePart>
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<mods:nameIdentifier id="AFE7ED8BE31F4F13D268703763CD0BD0" type="ORCID">0000-0001-9268-622X</mods:nameIdentifier>
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<mods:affiliation id="C1CD02F9BE2030C96395494410046E1E">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. & Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
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<mods:nameIdentifier id="6CC21D1CDB034BD6892BABAACB36CD2C" type="email">a.fomichov@mail.ru</mods:nameIdentifier>
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<mods:title id="6E83E0D9290126787A9401B8490E9A5C">Zootaxa</mods:title>
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<mods:part id="E3106BDA5012E3C6B905312BFEE0BB2D">
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<mods:date id="DFADC3B41F70D605E614174489FFF64A">2024</mods:date>
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<mods:number id="8CCB83926DFBF230E4A0A7705E32EDA6">2024-12-04</mods:number>
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<mods:detail id="36E5EEB1CC74B4E1CBB04EE0303CA080" type="volume">
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<mods:number id="AE3F4C827302E5F95ED7E4B07AFE3CCC">5543</mods:number>
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<mods:url id="A00B7A342BB0C179914E05B0654A5C8A">https://doi.org/10.11646/zootaxa.5543.3.5</mods:url>
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<mods:classification id="28E781324CC8306C544A51F641A90204">journal article</mods:classification>
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</mods:mods>
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<treatment id="B20419093769FF9569A0FDAD01F69D2B" LSID="urn:lsid:plazi:treatment:B20419093769FF9569A0FDAD01F69D2B" httpUri="http://treatment.plazi.org/id/B20419093769FF9569A0FDAD01F69D2B" lastPageId="12" lastPageNumber="395" pageId="10" pageNumber="393">
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<subSubSection id="72B7FB943769FF9369A0FDAD03B69D1C" box="[151,473,571,598]" pageId="10" pageNumber="393" type="nomenclature">
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<paragraph id="3A12A81F3769FF9369A0FDAD03B69D1C" blockId="10.[151,473,571,634]" box="[151,473,571,598]" pageId="10" pageNumber="393">
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<emphasis id="08D9740D3769FF9369A0FDAD03B69D1C" bold="true" box="[151,473,571,598]" pageId="10" pageNumber="393">
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<taxonomicName id="FDADD39C3769FF9369A0FDAD031B9D1C" authorityName="Omelko & Fomichev" authorityYear="2024" box="[151,372,571,598]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
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<emphasis id="08D9740D3769FF9369A0FDAD031B9D1C" bold="true" box="[151,372,571,598]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="13EAC9763769FF93684CFDAD03B69D1C" box="[379,473,572,598]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="72B7FB943769FF9369A0FDF103149D30" box="[151,379,608,634]" pageId="10" pageNumber="393" type="description">
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<paragraph id="3A12A81F3769FF9369A0FDF103149D30" blockId="10.[151,473,571,634]" box="[151,379,608,634]" pageId="10" pageNumber="393">
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(
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||
<figureCitation id="A296B49A3769FF9369A8FDF1034E9D30" box="[159,289,608,634]" captionStart-0="FIGURES 31–32" captionStart-1="FIGURES 33–41" captionStartId-0="10.[151,264,1302,1326]" captionStartId-1="11.[151,264,1864,1888]" captionTargetBox-0="[153,1431,770,1275]" captionTargetBox-1="[163,1435,194,1834]" captionTargetId-0="figure-216@10.[151,1435,766,1278]" captionTargetId-1="figure-19@11.[151,1435,181,1840]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 31–32. Male habitus of Bowie tangalunga sp. nov. 31—dorsal; 32—ventral. Scale bar: 2 mm." captionText-1="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi-0="http://doi.org/10.5281/zenodo.14386413" figureDoi-1="http://doi.org/10.5281/zenodo.14386415" httpUri-0="https://zenodo.org/record/14386413/files/figure.png" httpUri-1="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Figs 31–41</figureCitation>
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,
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<figureCitation id="A296B49A3769FF93681BFDF1031C9D30" box="[300,371,608,634]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="10" pageNumber="393">68–69</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="72B7FB943769FF9369A0FD3907FA9DAC" pageId="10" pageNumber="393" type="materials_examined">
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||
<paragraph id="3A12A81F3769FF9369A0FD3907FA9DAC" blockId="10.[151,1437,679,743]" pageId="10" pageNumber="393">
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||
<emphasis id="08D9740D3769FF9369A0FD39032B9D88" bold="true" box="[151,324,680,706]" pageId="10" pageNumber="393">Type material.</emphasis>
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<materialsCitation id="8AC5A2423769FF93687DFD3901309DAC" collectingDate="1995-01-26" collectingDateMax="1995-01-30" collectingDateMin="1995-01-26" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Philippines" elevation="1350" latitude="7.016667" location="Apo Mt." longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" pageId="10" pageNumber="393" specimenCount="1" specimenCount-male="1" stateProvince="Cotabato Province" typeStatus="holotype">
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||
<emphasis id="08D9740D3769FF93687DFD3901309D88" bold="true" box="[330,863,679,706]" pageId="10" pageNumber="393">
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||
<collectingCountry id="42BAE88F3769FF93687DFD39006E9D88" box="[330,513,680,706]" name="Philippines" pageId="10" pageNumber="393">PHILIPPINES</collectingCountry>
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||
:
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||
<emphasis id="08D9740D3769FF936B3AFD36008C9D88" bold="true" box="[525,739,679,706]" italics="true" pageId="10" pageNumber="393">
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<collectingRegion id="F86966FD3769FF936B3AFD36008C9D88" box="[525,739,679,706]" country="Philippines" name="South Cotabato" pageId="10" pageNumber="393">Cotabato Province</collectingRegion>
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||
</emphasis>
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||
:
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||
<typeStatus id="E51616BD3769FF936BC5FD3901309D88" box="[754,863,680,706]" pageId="10" pageNumber="393" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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||
<specimenCount id="2CAB63963769FF936A5AFD3901FF9D88" box="[877,912,680,706]" count="1" pageId="10" pageNumber="393" type="male">1♂</specimenCount>
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||
(
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||
<collectionCode id="5CBC30DA3769FF936AA9FD3901949D88" box="[926,1019,680,706]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="393" type="Museum">ZMMU</collectionCode>
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||
),
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||
<collectingMunicipality id="DA7632653769FF936D3AFD3906A39D88" box="[1037,1228,680,706]" pageId="10" pageNumber="393">Mindanao Island</collectingMunicipality>
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||
,
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||
<location id="3F72FEC43769FF936DE2FD3907599D89" LSID="urn:lsid:plazi:treatment:B20419093769FF9569A0FDAD01F69D2B:3F72FEC43769FF936DE2FD3907599D89" box="[1237,1334,680,707]" country="Philippines" latitude="7.016667" longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" name="Apo Mt." pageId="10" pageNumber="393" stateProvince="Cotabato Province">Apo Mt.</location>
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, [
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||
<geoCoordinate id="5F99CED83769FF936C7EFD3907F89D88" box="[1353,1431,680,706]" degrees="7" direction="north" minutes="1" orientation="latitude" pageId="10" pageNumber="393" precision="925" value="7.016667">7° 1’N</geoCoordinate>
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,
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<geoCoordinate id="5F99CED83769FF9369A0FD5D03689DAD" box="[151,263,716,743]" degrees="125" direction="east" minutes="13" orientation="longitude" pageId="10" pageNumber="393" precision="925" value="125.21667">125°13’E</geoCoordinate>
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],
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||
<quantity id="FD5505FA3769FF93682DFD5D031E9DAD" box="[282,369,716,743]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="393" unit="m" value="1350.0">
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<elevation id="B1804F2C3769FF93682DFD5D031E9DAD" box="[282,369,716,743]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="393" unit="m" value="1350.0">1350 m</elevation>
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</quantity>
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||
,
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||
<date id="4E138EDF3769FF93684AFD5D000C9DAD" box="[381,611,716,743]" pageId="10" pageNumber="393" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">
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<collectingDate id="5E5777373769FF93684AFD5D000C9DAD" box="[381,611,716,743]" pageId="10" pageNumber="393" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">26–30 January 1995</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="9758CDC93769FF936B5BFD5D01409DAC" box="[620,815,716,742]" pageId="10" pageNumber="393">A.M. Emelyanov</collectorName>
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leg.
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</materialsCitation>
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||
<materialsCitation id="8AC5A2423769FF936A52FD5D07FE9DAC" box="[869,1425,716,743]" collectingDate="1995-01-26" collectingDateMax="1995-01-30" collectingDateMin="1995-01-26" collectionCode="IBSS" collectorName="A. M. Emelyanov" country="Philippines" elevation="1350" latitude="7.016667" location="Apo Mt." longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" pageId="10" pageNumber="393" specimenCount="1" specimenCount-male="1" stateProvince="Paratype" typeStatus="paratype">
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||
<emphasis id="08D9740D3769FF936A52FD5D01BD9DAC" bold="true" box="[869,978,716,742]" pageId="10" pageNumber="393">
|
||
<typeStatus id="E51616BD3769FF936A52FD5D01BD9DAC" box="[869,978,716,742]" pageId="10" pageNumber="393" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="2CAB63963769FF936AD6FD5D066B9DAC" box="[993,1028,716,742]" count="1" pageId="10" pageNumber="393" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="5CBC30DA3769FF936D24FD5D063C9DAC" box="[1043,1107,716,742]" collectionName="IBSS" pageId="10" pageNumber="393">IBSS</collectionCode>
|
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), together with the holotype
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6ED2F8973769FF9369A0FA8706A99A67" ID-DOI="http://doi.org/10.5281/zenodo.14386413" ID-Zenodo-Dep="14386413" box="[151,1222,1302,1326]" httpUri="https://zenodo.org/record/14386413/files/figure.png" pageId="10" pageNumber="393" startId="10.[151,264,1302,1326]" targetBox="[153,1431,770,1275]" targetPageId="10" targetType="figure">
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<paragraph id="3A12A81F3769FF9369A0FA8706A99A67" blockId="10.[151,1222,1302,1326]" box="[151,1222,1302,1326]" pageId="10" pageNumber="393">
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||
<emphasis id="08D9740D3769FF9369A0FA87033B9A64" bold="true" box="[151,340,1302,1326]" pageId="10" pageNumber="393">FIGURES 31–32.</emphasis>
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Male habitus of
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<taxonomicName id="FDADD39C3769FF9368C8FA8600D79A64" authorityName="Omelko & Fomichev" authorityYear="2024" box="[511,696,1302,1326]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
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||
<emphasis id="08D9740D3769FF9368C8FA8600D79A64" box="[511,696,1302,1326]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
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||
</taxonomicName>
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||
<emphasis id="08D9740D3769FF936B88FA8701609A64" bold="true" box="[703,783,1302,1326]" pageId="10" pageNumber="393">
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||
<taxonomicNameLabel id="13EAC9763769FF936B88FA8701609A64" box="[703,783,1302,1326]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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31—dorsal; 32—ventral. Scale bar: 2 mm.
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</paragraph>
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||
</caption>
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||
<subSubSection id="72B7FB943769FF9369F0FAE701F49AF9" pageId="10" pageNumber="393" type="etymology">
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||
<paragraph id="3A12A81F3769FF9369F0FAE701F49AF9" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
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||
<emphasis id="08D9740D3769FF9369F0FAE703279ADA" bold="true" box="[199,328,1398,1424]" pageId="10" pageNumber="393">Etymology</emphasis>
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. The specific name is derived from the local name of the Malayan civet [
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<taxonomicName id="FDADD39C3769FF936D43FAE707FE9ADA" authority="Gray" authorityName="Gray" authorityYear="1832" box="[1140,1425,1397,1424]" class="Mammalia" family="Viverridae" genus="Viverra" kingdom="Animalia" order="Carnivora" pageId="10" pageNumber="393" phylum="Chordata" rank="species" species="tangalunga">
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<emphasis id="08D9740D3769FF936D43FAE707219AC5" box="[1140,1358,1397,1423]" italics="true" pageId="10" pageNumber="393">Viverra tangalunga</emphasis>
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Gray
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</taxonomicName>
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], a small predatory beast native to Southeast Asia; name in apposition.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="72B7FB943769FF9369F0FA2C0683992D" pageId="10" pageNumber="393" type="diagnosis">
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||
<paragraph id="3A12A81F3769FF9369F0FA2C0683992D" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
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||
<emphasis id="08D9740D3769FF9369F0FA2C03559A9D" bold="true" box="[199,314,1469,1495]" pageId="10" pageNumber="393">Diagnosis</emphasis>
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. Males of
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<taxonomicName id="FDADD39C3769FF936886FA2F00119A9D" authorityName="Omelko & Fomichev" authorityYear="2024" box="[433,638,1469,1495]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
|
||
<emphasis id="08D9740D3769FF936886FA2F00119A9D" box="[433,638,1469,1495]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="08D9740D3769FF936BB3FA2F00B29A92" bold="true" box="[644,733,1470,1496]" pageId="10" pageNumber="393">
|
||
<taxonomicNameLabel id="13EAC9763769FF936BB3FA2F00B29A92" box="[644,733,1470,1496]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resemble those of
|
||
<taxonomicName id="FDADD39C3769FF936A87FA2F06019A9D" authorityName="Jager" authorityYear="2022" box="[944,1134,1469,1495]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="criminalworld">
|
||
<emphasis id="08D9740D3769FF936A87FA2F06019A9D" box="[944,1134,1469,1495]" italics="true" pageId="10" pageNumber="393">B. criminalworld</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the tegular apophysis (
|
||
<emphasis id="08D9740D3769FF93682DFA7303569AB6" box="[282,313,1506,1532]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
|
||
) and the shape of the retro-proximal cymbial outgrowth (
|
||
<emphasis id="08D9740D3769FF936AEFFA7306619AB6" box="[984,1038,1506,1532]" italics="true" pageId="10" pageNumber="393">RPO</emphasis>
|
||
) in ventral view but can be easily distinguished by 1) embolus hidden behind the tegular apophysis to a large extent (vs. not hidden; cf.
|
||
<figureCitation id="A296B49A3769FF936C2AF99407F3996A" box="[1309,1436,1541,1568]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Figs 34, 38</figureCitation>
|
||
and
|
||
<bibRefCitation id="5E3CD5EE3769FF9369F0F9BB0329990E" author="Jager, P." box="[199,326,1578,1604]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
|
||
: fig. 769), 2) base of
|
||
<emphasis id="08D9740D3769FF936B00F9BB0039990E" box="[567,598,1578,1604]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
|
||
located almost medially (vs. retrolaterally; cf.
|
||
<figureCitation id="A296B49A3769FF936D52F9B806DA990E" box="[1125,1205,1577,1604]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Fig. 38</figureCitation>
|
||
and
|
||
<bibRefCitation id="5E3CD5EE3769FF936DDBF9BB0704990E" author="Jager, P." box="[1260,1387,1578,1604]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
|
||
: fig. 769), 3) tip of embolus (
|
||
<emphasis id="08D9740D3769FF93689EF9DF03A1992D" box="[425,462,1614,1639]" italics="true" pageId="10" pageNumber="393">Em</emphasis>
|
||
) extending
|
||
<emphasis id="08D9740D3769FF936B63F9DF001D9922" box="[596,626,1614,1640]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
|
||
(vs. not extending; cf.
|
||
<figureCitation id="A296B49A3769FF936A42F9DC01AB9922" box="[885,964,1613,1640]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Fig. 38</figureCitation>
|
||
and
|
||
<bibRefCitation id="5E3CD5EE3769FF936ACCF9DF06169922" author="Jager, P." box="[1019,1145,1614,1640]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
|
||
: fig. 769).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72B7FB943769FF9569F0F9E000D09D77" lastPageId="12" lastPageNumber="395" pageId="10" pageNumber="393" type="description">
|
||
<paragraph id="3A12A81F3769FF9369F0F9E0037098CD" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
|
||
<emphasis id="08D9740D3769FF9369F0F9E0033699C1" bold="true" box="[199,345,1649,1675]" pageId="10" pageNumber="393">Description.</emphasis>
|
||
<emphasis id="08D9740D3769FF936855F9E003F399C1" box="[354,412,1649,1675]" italics="true" pageId="10" pageNumber="393">Male</emphasis>
|
||
(
|
||
<figureCitation id="A296B49A3769FF93689BF9E0005E99C6" box="[428,561,1649,1676]" captionStart="FIGURES 31–32" captionStartId="10.[151,264,1302,1326]" captionTargetBox="[153,1431,770,1275]" captionTargetId="figure-216@10.[151,1435,766,1278]" captionTargetPageId="10" captionText="FIGURES 31–32. Male habitus of Bowie tangalunga sp. nov. 31—dorsal; 32—ventral. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386413" httpUri="https://zenodo.org/record/14386413/files/figure.png" pageId="10" pageNumber="393">Figs 31–32</figureCitation>
|
||
). Total length 13.11. Prosoma 7.10 long, 5.23 wide. Opisthosoma 6.05 long, 4.05 wide. Prosoma black, with light brown median band, slightly widened near ocular area. Lateral bands disrupted into several whitish spots. Fovea thin, black. Clypeus light brown. Chelicerae dark brown with brown longitudinal stripes anteriorly, 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum brown without pattern. Labium dark brown. Endites brown with yellow distal edge. Dorsal part of opisthosoma black with short yellowish cardiac mark and series of transverse stripes posterior of it. Lateral sides of opisthosoma black with lots of tiny yellow spots. Ventral part of opisthosoma black with yellow spots forming two V-marks (inner mark poorly visible). Spinnerets dark brown.
|
||
</paragraph>
|
||
<paragraph id="3A12A81F3769FF9369F0F80307EF989A" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">Eye diameters: AME 0.28, ALE 0.21, PME 0.38, PLE 0.32; interdistances: AME–AME 0.16, AME–ALE 0.38, PME–PME 0.23, PME–PLE 0.35, AME–PME 0.10, ALE–PLE 0.18. Clypeus height at AME 0.20, at ALE 0.59.</paragraph>
|
||
<caption id="6ED2F8973768FF9269A0F8D9008D9886" ID-DOI="http://doi.org/10.5281/zenodo.14386415" ID-Zenodo-Dep="14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="11" pageNumber="394" startId="11.[151,264,1864,1888]" targetBox="[163,1435,194,1834]" targetPageId="11" targetType="figure">
|
||
<paragraph id="3A12A81F3768FF9269A0F8D9008D9886" blockId="11.[151,1436,1864,1996]" pageId="11" pageNumber="394">
|
||
<emphasis id="08D9740D3768FF9269A0F8D9033C982A" bold="true" box="[151,339,1864,1888]" pageId="11" pageNumber="394">FIGURES 33–41.</emphasis>
|
||
Palp (33–36), bulb (37–40) and palpal tibia (41) of male of
|
||
<taxonomicName id="FDADD39C3768FF926A91F8D80632982A" authorityName="Omelko & Fomichev" authorityYear="2024" box="[934,1117,1864,1888]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="394" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
|
||
<emphasis id="08D9740D3768FF926A91F8D80632982A" box="[934,1117,1864,1888]" italics="true" pageId="11" pageNumber="394">Bowie tangalunga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="08D9740D3768FF926D54F8D906DD982A" bold="true" box="[1123,1202,1864,1888]" pageId="11" pageNumber="394">
|
||
<taxonomicNameLabel id="13EAC9763768FF926D54F8D906DD982A" box="[1123,1202,1864,1888]" pageId="11" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm. Abbreviations:
|
||
<emphasis id="08D9740D3768FF926C45F8FD07EC98C9" box="[1394,1411,1900,1923]" italics="true" pageId="11" pageNumber="394">C</emphasis>
|
||
— conductor,
|
||
<emphasis id="08D9740D3768FF92683DF800034398E2" box="[266,300,1937,1960]" italics="true" pageId="11" pageNumber="394">EH</emphasis>
|
||
—embolic hook,
|
||
<emphasis id="08D9740D3768FF9268D6F800006C98E2" box="[481,515,1937,1960]" italics="true" pageId="11" pageNumber="394">Em</emphasis>
|
||
—embolus,
|
||
<emphasis id="08D9740D3768FF926BB7F80100DE98ED" box="[640,689,1936,1959]" italics="true" pageId="11" pageNumber="394">RPO</emphasis>
|
||
—retro-proximal cymbial outgrowth,
|
||
<emphasis id="08D9740D3768FF926D0FF801060C98ED" box="[1080,1123,1936,1959]" italics="true" pageId="11" pageNumber="394">RTA</emphasis>
|
||
—retrolateral tibial apophysis,
|
||
<emphasis id="08D9740D3768FF9269A0F82502DC9881" box="[151,179,1972,1995]" italics="true" pageId="11" pageNumber="394">TA</emphasis>
|
||
—tegular apophysis,
|
||
<emphasis id="08D9740D3768FF9268BCF82403CD9886" box="[395,418,1973,1996]" italics="true" pageId="11" pageNumber="394">Te</emphasis>
|
||
—tegulum,
|
||
<emphasis id="08D9740D3768FF926B2CF82400549886" box="[539,571,1973,1996]" italics="true" pageId="11" pageNumber="394">TH</emphasis>
|
||
—tegular hump.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3A12A81F376FFF9569F0FF0600DA9E2F" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
|
||
For palp and legs measurements see
|
||
<tableCitation id="772F9DA4376FFF956B59FF0600A99FFB" box="[622,710,151,177]" captionStart="TABLE 9" captionStartId="12.[151,239,656,680]" captionTargetPageId="12" captionText="TABLE 9. Palp and legs measurements of male of Bowie tangalunga sp. nov." pageId="12" pageNumber="395">Table 9</tableCitation>
|
||
. Palp coloration: femora, patellae, tibiae brown; cymbium dark brown. Legs coloration: femora I dark brown dorsally and laterally, brown ventrally, with light brown longitudinal stripe retrolaterally; II dark brown dorsally and prolaterally, brown retrolaterally and ventrally; III–IV dark brown dorsally and laterally, brown ventrally, with light brown semi-rings. Patellae I–IV brown. Tibiae I–II, IV light brown with white setae; III light brown. Metatarsi I–II brown; III–IV brown with light brow semi-rings. Tarsi I–IV light brown. For palp and legs spination see
|
||
<tableCitation id="772F9DA4376FFF956B7AFEDA00DE9E2F" box="[589,689,331,357]" captionStart="TABLE 10" captionStartId="12.[151,239,1007,1031]" captionTargetPageId="12" captionText="TABLE 10. Palp and legs spination for the male of Bowie tangalunga sp. nov." pageId="12" pageNumber="395">Table 10</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3A12A81F376FFF9569F0FEFE07F49EBF" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
|
||
Palp as shown in
|
||
<figureCitation id="A296B49A376FFF9568BFFEFE00679EC3" box="[392,520,367,393]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="12" pageNumber="395">Figs 33–41</figureCitation>
|
||
.
|
||
<emphasis id="08D9740D376FFF956B26FEFE002E9EC3" box="[529,577,367,393]" italics="true" pageId="12" pageNumber="395">RTA</emphasis>
|
||
arising from middle of tibia, short, curved, with sharply pointed tip in ventral view. Cymbium with short, blunt retro-proximal cymbial outgrowth (
|
||
<emphasis id="08D9740D376FFF956AAFFE0201A19EE7" box="[920,974,403,429]" italics="true" pageId="12" pageNumber="395">RPO</emphasis>
|
||
). Tegulum (
|
||
<emphasis id="08D9740D376FFF956D61FE0206009EE6" box="[1110,1135,403,428]" italics="true" pageId="12" pageNumber="395">Te</emphasis>
|
||
) with rounded hump (
|
||
<emphasis id="08D9740D376FFF956C5FFE0207E39EE6" box="[1384,1420,403,428]" italics="true" pageId="12" pageNumber="395">TH</emphasis>
|
||
). Tegular apophysis (
|
||
<emphasis id="08D9740D376FFF956843FE2603FC9E9B" box="[372,403,439,465]" italics="true" pageId="12" pageNumber="395">TA</emphasis>
|
||
) oval, originating in a 5 o’clock position. Conductor (
|
||
<emphasis id="08D9740D376FFF956AC5FE26066A9E9B" box="[1010,1029,439,465]" italics="true" pageId="12" pageNumber="395">C</emphasis>
|
||
) large, membranous. Embolus (
|
||
<emphasis id="08D9740D376FFF956C59FE2607FC9E9A" box="[1390,1427,439,464]" italics="true" pageId="12" pageNumber="395">Em</emphasis>
|
||
) originating from tegulum in 6.30-o’clock-position, with pointed tip. Embolus with large basal embolus hook (
|
||
<emphasis id="08D9740D376FFF956C51FE4A07E49EBE" box="[1382,1419,475,500]" italics="true" pageId="12" pageNumber="395">EH</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3A12A81F376FFF9569F0FE6E00D09D77" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
|
||
<emphasis id="08D9740D376FFF9569F0FE6E037E9D53" bold="true" box="[199,273,511,537]" pageId="12" pageNumber="395">Notes.</emphasis>
|
||
According to the structure of the copulatory organs,
|
||
<taxonomicName id="FDADD39C376FFF956A5AFE6E06549D53" authorityName="Omelko & Fomichev" authorityYear="2024" box="[877,1083,511,537]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
|
||
<emphasis id="08D9740D376FFF956A5AFE6E06549D53" box="[877,1083,511,537]" italics="true" pageId="12" pageNumber="395">Bowie tangalunga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="08D9740D376FFF956D74FE6E06F29D53" bold="true" box="[1091,1181,511,537]" pageId="12" pageNumber="395">
|
||
<taxonomicNameLabel id="13EAC976376FFF956D74FE6E06F29D53" box="[1091,1181,511,537]" pageId="12" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs to the
|
||
<taxonomicName id="FDADD39C376FFF956C7AFE6E07F39D53" box="[1357,1436,511,537]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
|
||
<emphasis id="08D9740D376FFF956C7AFE6E07F39D53" box="[1357,1436,511,537]" italics="true" pageId="12" pageNumber="395">shakeit</emphasis>
|
||
</taxonomicName>
|
||
species-group (see notes for
|
||
<taxonomicName id="FDADD39C376FFF9568EFFDB200219D76" authorityName="Omelko & Fomichev" authorityYear="2024" box="[472,590,547,572]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
|
||
<emphasis id="08D9740D376FFF9568EFFDB200219D76" box="[472,590,547,572]" italics="true" pageId="12" pageNumber="395">B. musang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="08D9740D376FFF956B62FDB200C19D77" bold="true" box="[597,686,547,573]" pageId="12" pageNumber="395">
|
||
<taxonomicNameLabel id="13EAC976376FFF956B62FDB200C19D77" box="[597,686,547,573]" pageId="12" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72B7FB94376FFF9569F0FDD701F69D2B" box="[199,921,582,609]" pageId="12" pageNumber="395" type="distribution">
|
||
<paragraph id="3A12A81F376FFF9569F0FDD701F69D2B" blockId="12.[151,1437,151,609]" box="[199,921,582,609]" pageId="12" pageNumber="395">
|
||
<emphasis id="08D9740D376FFF9569F0FDD7030E9D2A" bold="true" box="[199,353,582,608]" pageId="12" pageNumber="395">Distribution.</emphasis>
|
||
Mindanao Isl., Apo Mt.,
|
||
<collectingCountry id="42BAE88F376FFF956B4BFDD600959D2B" box="[636,762,583,609]" name="Philippines" pageId="12" pageNumber="395">Philippines</collectingCountry>
|
||
(
|
||
<figureCitation id="A296B49A376FFF956A3EFDD601E29D2B" box="[777,909,583,609]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="12" pageNumber="395">Figs 68–69</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |