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<document id="0F4CB15D3FA11E2AACBCDC5528A099D7" ID-DOI="10.11646/zootaxa.5543.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386385" ID-ZooBank="8A21E43A-5605-4238-97B2-81CCA432A503" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733918705237" checkinUser="plazi" docAuthor="Omelko, Mikhail M. &amp; Fomichev, Alexander A." docDate="2024" docId="B20419093762FF9C69A0F89901EC99E0" docLanguage="en" docName="zootaxa.5543.3.5.pdf" docOrigin="Zootaxa 5543 (3)" docSource="https://doi.org/10.11646/zootaxa.5543.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bowie neukoeln Jager 2022" docType="treatment" docVersion="2" lastPageNumber="388" masterDocId="4E3D61713763FF996937FF91026F9F4A" masterDocTitle="New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia" masterLastPageNumber="403" masterPageNumber="383" pageNumber="384" updateTime="1733919598295" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CDCEC5A5B74E257008D944F21DA7874E">
<mods:title id="EC6F4EACCDC61BC6AC669922F1949F2C">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
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<mods:namePart id="F18F0DC4CAC7D046C8125939ACC480F7">Omelko, Mikhail M.</mods:namePart>
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<mods:affiliation id="BB9A6FBB48508AF4E4C26DA6327C21F4">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
<mods:nameIdentifier id="BCECAEDA0D3FE8A84438619154175D10" type="email">omelkom@gmail.com</mods:nameIdentifier>
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<mods:namePart id="00CF19B7EAD70569A21D0B6A7DCAD1FC">Fomichev, Alexander A.</mods:namePart>
<mods:nameIdentifier id="649E496CF05D18F4161A7FB9325F16A6" type="ORCID">0000-0001-9268-622X</mods:nameIdentifier>
<mods:affiliation id="36BDAF5095793B0194AE7D938A3B5D62">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
<mods:nameIdentifier id="32FBE472E4EE3B4145408008919FC866" type="email">a.fomichov@mail.ru</mods:nameIdentifier>
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<mods:title id="D5BFE5E1BE56D8E3F53496C56A543154">Zootaxa</mods:title>
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<mods:date id="15B68A1F61F9AA946DCBB2C25A9CBFC6">2024</mods:date>
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<mods:number id="D1DB3A7BCBD4A2145A1918AD0C1C616C">2024-12-04</mods:number>
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<mods:start id="07F8617837DA6F1E779275B0801C9B3C">383</mods:start>
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<treatment id="B20419093762FF9C69A0F89901EC99E0" LSID="urn:lsid:plazi:treatment:B20419093762FF9C69A0F89901EC99E0" httpUri="http://treatment.plazi.org/id/B20419093762FF9C69A0F89901EC99E0" lastPageId="5" lastPageNumber="388" pageId="1" pageNumber="384">
<subSubSection id="72B7FB943762FF9869A0F89903819868" box="[151,494,1799,1826]" pageId="1" pageNumber="384" type="nomenclature">
<paragraph id="3A12A81F3762FF9869A0F89903819868" blockId="1.[151,494,1799,1863]" box="[151,494,1799,1826]" pageId="1" pageNumber="384">
<heading id="615A1F733762FF9869A0F89903819868" bold="true" box="[151,494,1799,1826]" fontSize="11" level="1" pageId="1" pageNumber="384" reason="1">
<taxonomicName id="FDADD39C3762FF9869A0F89903819868" ID-CoL="C6LJM" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[151,494,1799,1826]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3762FF9869A0F89903819868" bold="true" box="[151,494,1799,1826]" pageId="1" pageNumber="384">
<emphasis id="08D9740D3762FF9869A0F89903369868" bold="true" box="[151,345,1799,1826]" italics="true" pageId="1" pageNumber="384">Bowie neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3762FF986856F89903819868" author="Jager, P." box="[353,494,1800,1826]" pageId="1" pageNumber="384" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943762FF9869A0F8BD03F8980C" box="[151,407,1836,1863]" pageId="1" pageNumber="384" type="description">
<paragraph id="3A12A81F3762FF9869A0F8BD03F8980C" blockId="1.[151,494,1799,1863]" box="[151,407,1836,1863]" pageId="1" pageNumber="384">
(
<figureCitation id="A296B49A3762FF9869A8F8BD036B980C" box="[159,260,1836,1862]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="1" pageNumber="384">Figs 1–2</figureCitation>
,
<figureCitation id="A296B49A3762FF986827F8BD0353980D" box="[272,316,1836,1863]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="1" pageNumber="384">5–8</figureCitation>
,
<figureCitation id="A296B49A3762FF98687FF8BD03E0980C" box="[328,399,1836,1862]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="1" pageNumber="384">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943762FF9869A0F8E001EE98C2" box="[151,897,1904,1929]" pageId="1" pageNumber="384" type="reference_group">
<paragraph id="3A12A81F3762FF9869A0F8E001EE98C2" blockId="1.[151,897,1904,1929]" box="[151,897,1904,1929]" pageId="1" pageNumber="384">
<treatmentCitationGroup id="1ABD8F313762FF9869A0F8E001EE98C2" box="[151,897,1904,1929]" pageId="1" pageNumber="384">
<treatmentCitation id="BB0C8E0E3762FF9869A0F8E003DE98C3" author="Jager, P." box="[151,433,1904,1929]" page="98" pageId="1" pageNumber="384" year="2022">
<taxonomicName id="FDADD39C3762FF9869A0F8E003DE98C3" ID-CoL="C6LJM" authority="Jager, 2022: 98" authorityName="Jager" authorityPageNumber="98" authorityYear="2022" box="[151,433,1904,1929]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3762FF9869A0F8E0036198C2" box="[151,270,1904,1928]" italics="true" pageId="1" pageNumber="384">B. neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3762FF986822F8E003DE98C3" author="Jager, P." box="[277,433,1905,1929]" pageId="1" pageNumber="384" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 440–446, 448–460 (
<specimenCount id="2CAB63963762FF986B81F8E000A398C3" box="[694,716,1905,1929]" count="1" pageId="1" pageNumber="384" type="male">♂</specimenCount>
<specimenCount id="2CAB63963762FF986BFBF8E000B498C3" box="[716,731,1905,1929]" count="1" pageId="1" pageNumber="384" type="female">♀</specimenCount>
), not examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943762FF9869A0F82700AF98BF" pageId="1" pageNumber="384" type="materials_examined">
<paragraph id="3A12A81F3762FF9869A0F82700AF98BF" blockId="1.[151,1436,1974,2037]" pageId="1" pageNumber="384">
<emphasis id="08D9740D3762FF9869A0F82703EE989A" bold="true" box="[151,385,1974,2000]" pageId="1" pageNumber="384">Material examined.</emphasis>
<materialsCitation id="8AC5A2423762FF9868BEF82700AF98BF" collectingDate="1993-02-28" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1050" latitude="5.4" location="Crocker Range Ridge" longLatPrecision="1305" longitude="116.1" municipality="Borneo" pageId="1" pageNumber="384" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<emphasis id="08D9740D3762FF9868BEF82700AC989B" bold="true" box="[393,707,1974,2001]" pageId="1" pageNumber="384">
<collectingCountry id="42BAE88F3762FF9868BEF827004C989A" box="[393,547,1974,2000]" name="Malaysia" pageId="1" pageNumber="384">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D3762FF986B07F82700D5989B" bold="true" box="[560,698,1974,2001]" italics="true" pageId="1" pageNumber="384">
<collectingRegion id="F86966FD3762FF986B07F82700D5989B" box="[560,698,1974,2001]" country="Malaysia" name="Sabah" pageId="1" pageNumber="384">
<taxonomicName id="FDADD39C3762FF986B07F8270016989B" authorityName="Li &amp; Yao" authorityYear="2022" box="[560,633,1974,2001]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="2CAB63963762FF986BFCF826008F989B" box="[715,736,1975,2001]" count="1" pageId="1" pageNumber="384" type="female">♀</specimenCount>
(
<collectionCode id="5CBC30DA3762FF986BD8F8260123989B" box="[751,844,1975,2001]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="384" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653762FF986A57F82601DA989B" box="[864,949,1975,2001]" pageId="1" pageNumber="384">Borneo</collectingMunicipality>
,
<location id="3F72FEC43762FF986AF6F82606DA989B" LSID="urn:lsid:plazi:treatment:B20419093762FF9C69A0F89901EC99E0:3F72FEC43762FF986AF6F82606DA989B" box="[961,1205,1975,2001]" country="Malaysia" latitude="5.4" longLatPrecision="1305" longitude="116.1" municipality="Borneo" name="Crocker Range Ridge" pageId="1" pageNumber="384" stateProvince="Sabah">Crocker Range Ridge</location>
[
<geoCoordinate id="5F99CED83762FF986DF3F8270774989B" box="[1220,1307,1974,2001]" degrees="5" direction="north" minutes="24" orientation="latitude" pageId="1" pageNumber="384" precision="925" value="5.4">5°24’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83762FF986C1FF82607E0989B" box="[1320,1423,1975,2001]" degrees="116" direction="east" minutes="6" orientation="longitude" pageId="1" pageNumber="384" precision="925" value="116.1">116° 6’E</geoCoordinate>
],
<quantity id="FD5505FA3762FF9869A0F84B028198BF" box="[151,238,2010,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="384" unit="m" value="1050.0">
<elevation id="B1804F2C3762FF9869A0F84B028198BF" box="[151,238,2010,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="384" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="4E138EDF3762FF9869CDF84A03AB98BF" box="[250,452,2011,2037]" pageId="1" pageNumber="384" value="1993-02-28">
<collectingDate id="5E5777373762FF9869CDF84A03AB98BF" box="[250,452,2011,2037]" pageId="1" pageNumber="384" value="1993-02-28">28 February 1993</collectingDate>
</date>
,
<collectorName id="9758CDC93762FF9868FAF84A00FF98BF" box="[461,656,2011,2037]" pageId="1" pageNumber="384">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="6ED2F8973761FF9B69A0FB2F03219BB0" ID-DOI="http://doi.org/10.5281/zenodo.14386389" ID-Zenodo-Dep="14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="2" pageNumber="385" startId="2.[151,264,1214,1238]" targetBox="[183,1407,180,1182]" targetPageId="2" targetType="figure">
<paragraph id="3A12A81F3761FF9B69A0FB2F03219BB0" blockId="2.[151,1437,1214,1274]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69A0FB2F03539B9C" bold="true" box="[151,316,1214,1238]" pageId="2" pageNumber="385">FIGURES 1–4.</emphasis>
Female habitus of
<taxonomicName id="FDADD39C3761FF9B6B32FB2E008B9B9D" authority="Jager" authorityName="Jager" box="[517,740,1214,1239]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3761FF9B6B32FB2E00C89B9C" box="[517,679,1214,1238]" italics="true" pageId="2" pageNumber="385">Bowie neukoeln</emphasis>
Jäger
</taxonomicName>
(1–2) and
<taxonomicName id="FDADD39C3761FF9B6A6FFB2E01B69B9C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[856,985,1215,1238]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3761FF9B6A6FFB2E01B69B9C" box="[856,985,1215,1238]" italics="true" pageId="2" pageNumber="385">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3761FF9B6AD6FB2F065C9B9C" bold="true" box="[993,1075,1214,1238]" pageId="2" pageNumber="385">
<taxonomicNameLabel id="13EAC9763761FF9B6AD6FB2F065C9B9C" box="[993,1075,1214,1238]" pageId="2" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="72B7FB943761FF9B69F0FADD07779ABD" pageId="2" pageNumber="385" type="diagnosis">
<paragraph id="3A12A81F3761FF9B69F0FADD07779ABD" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69F0FADD032F9A2C" bold="true" box="[199,320,1356,1382]" pageId="2" pageNumber="385">Diagnosis.</emphasis>
Females of
<taxonomicName id="FDADD39C3761FF9B68F2FADC00299A2C" box="[453,582,1356,1382]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3761FF9B68F2FADC00299A2C" box="[453,582,1356,1382]" italics="true" pageId="2" pageNumber="385">B. neukoeln</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="FDADD39C3761FF9B6A08FADC06109A2D" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[831,1151,1357,1383]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3761FF9B6A08FADC01939A2C" box="[831,1020,1357,1382]" italics="true" pageId="2" pageNumber="385">B. scarymonsters</emphasis>
<bibRefCitation id="5E3CD5EE3761FF9B6D35FADC06109A2D" author="Jager, P." box="[1026,1151,1357,1383]" pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FDADD39C3761FF9B6D84FADC03099AC1" authority="Omelko &amp; Fomichev, 2023" authorityName="Omelko &amp; Fomichev" authorityYear="2023" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="binturong">
<emphasis id="08D9740D3761FF9B6D84FADC07529A2C" box="[1203,1341,1356,1382]" italics="true" pageId="2" pageNumber="385">B. binturong</emphasis>
<bibRefCitation id="5E3CD5EE3761FF9B6C74FADD03099AC1" author="Omelko, M. M. &amp; Fomichev, A. A." pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev, 2023</bibRefCitation>
</taxonomicName>
by shape of median plate (
<emphasis id="08D9740D3761FF9B6BA2FAE000D29AC0" box="[661,701,1393,1418]" italics="true" pageId="2" pageNumber="385">MP</emphasis>
) and spermathecae (
<emphasis id="08D9740D3761FF9B6A93FAE001AF9AC1" box="[932,960,1393,1419]" italics="true" pageId="2" pageNumber="385">Sp</emphasis>
) as well as position of lateral teeth (
<emphasis id="08D9740D3761FF9B6C61FAE0071B9AC0" box="[1366,1396,1393,1418]" italics="true" pageId="2" pageNumber="385">LT</emphasis>
). It can be distinguished from both species by 1) having posterior pocket (pit) of epigyne (vs. absence; cf.
<figureCitation id="A296B49A3761FF9B6DD8FA0507409AE5" box="[1263,1327,1428,1455]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="2" pageNumber="385">Fig. 8</figureCitation>
and
<bibRefCitation id="5E3CD5EE3761FF9B6C55FA0402BC9A99" author="Jager, P." pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 467,
<bibRefCitation id="5E3CD5EE3761FF9B6809FA2900059A99" author="Omelko, M. M. &amp; Fomichev, A. A." box="[318,618,1464,1491]" pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev 2023</bibRefCitation>
: fig. 54); 2) spermathecae separated by gap equal to ca. 0.5 their width (vs. 1 and 0.3 of width, respectively; cf.
<figureCitation id="A296B49A3761FF9B6B11FA4D00089ABD" box="[550,615,1500,1527]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="2" pageNumber="385">Fig. 7</figureCitation>
and
<bibRefCitation id="5E3CD5EE3761FF9B6BABFA4C01759ABD" author="Jager, P." box="[668,794,1501,1527]" pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 468,
<bibRefCitation id="5E3CD5EE3761FF9B6AB0FA4D06D99ABD" author="Omelko, M. M. &amp; Fomichev, A. A." box="[903,1206,1500,1527]" pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev 2023</bibRefCitation>
: fig. 53).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943761FF9D69F0F99107309E0B" lastPageId="4" lastPageNumber="387" pageId="2" pageNumber="385" type="description">
<paragraph id="3A12A81F3761FF9B69F0F99101FB99B8" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69F0F99103369950" bold="true" box="[199,345,1536,1562]" pageId="2" pageNumber="385">Description.</emphasis>
<emphasis id="08D9740D3761FF9B6855F99103D99950" box="[354,438,1536,1562]" italics="true" pageId="2" pageNumber="385">Female</emphasis>
(
<figureCitation id="A296B49A3761FF9B68F0F99100419951" box="[455,558,1536,1563]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="2" pageNumber="385">Figs 1–2</figureCitation>
). Total length 11.75. Prosoma 6.16 long, 4.68 wide. Opisthosoma 5.56 long, 2.93 wide. Prosoma brown, with light brown median band. Median band widened and with projections near ocular area. Lateral bands almost invisible. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum brown, without pattern. Labium dark brown. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma dark brown with distinct yellowish cardiac mark and a series of black spots. Lateral sides of opisthosoma yellowish brown. Ventral part of opisthosoma dark brown with small yellowish spots forming two V-marks. Spinnerets dark brown.
</paragraph>
<paragraph id="3A12A81F3761FF9B69F0F96C07109871" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">Eye diameters: AME 0.26, ALE 0.19, PME 0.40, PLE 0.34; interdistances: AME–AME 0.21, AME–ALE 0.41, PME–PME 0.25, PME–PLE 0.47, AME–PME 0.11, ALE–PLE 0.24. Clypeus height at AME 0.14, at ALE 0.54.</paragraph>
<paragraph id="3A12A81F3761FF9B69F0F8D4029E98A5" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
For palp and legs measurements see Table 1. Palp coloration: all joints light brown. Leg coloration: femora I brown with yellow longitudinal stripe retrolaterally; II–IV brown with poorly visible yellow spots. Patellae I–IV brown. Tibiae I–II light brown; III–IV light brown with poorly visible yellow semi-rings. Metatarsi I–II light brown; III–IV light brown with poorly visible yellow semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA43761FF9B69A0F845028298A5" box="[151,237,2004,2031]" captionStart="TABLE 2" captionStartId="3.[151,239,502,526]" captionText="TABLE 2. Palp and legs spination for the female of Bowie neukoeln Jäger, 2022" pageId="2" pageNumber="385">Table 2</tableCitation>
.
</paragraph>
<paragraph id="3A12A81F3761FF9B69BDF7A307F39701" blockId="2.[138,1436,2094,2123]" box="[138,1436,2094,2123]" pageId="2" pageNumber="385">
NEW AND POORLY KNOWN SPECIES OF
<taxonomicName id="FDADD39C3761FF9B6B16F7A300C8970F" authority="FROM" authorityName="FROM" box="[545,679,2098,2117]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="08D9740D3761FF9B6B16F7A3000B970F" box="[545,612,2098,2117]" italics="true" pageId="2" pageNumber="385">BOWIE</emphasis>
FROM
</taxonomicName>
SOUTHEAST ASIA
<emphasis id="08D9740D3761FF9B6AB9F7A2018A9700" box="[910,997,2099,2122]" italics="true" pageId="2" pageNumber="385">Zootaxa</emphasis>
5543 (3) © 2024 Magnolia Press
<emphasis id="08D9740D3761FF9B6C60F7A3070F9700" bold="true" box="[1367,1376,2098,2122]" pageId="2" pageNumber="385">·</emphasis>
385
</paragraph>
<caption id="6ED2F8973760FF9A69A0FE67019E9D44" box="[151,1009,502,527]" pageId="3" pageNumber="386" startId="3.[151,239,502,526]" targetType="table">
<paragraph id="3A12A81F3760FF9A69A0FE67019E9D44" blockId="3.[151,1009,502,527]" box="[151,1009,502,527]" pageId="3" pageNumber="386">
<emphasis id="08D9740D3760FF9A69A0FE6703659D44" bold="true" box="[151,266,502,526]" pageId="3" pageNumber="386">TABLE 2.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C3760FF9A6BF0FE66019E9D44" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[711,1009,502,527]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3760FF9A6BF0FE66011E9D44" box="[711,881,502,526]" italics="true" pageId="3" pageNumber="386">Bowie neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3760FF9A6A4FFE66019E9D44" author="Jager, P." box="[888,1009,502,527]" pageId="3" pageNumber="386" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F3760FF9A68B8FD8E07399C4A" pageId="3" pageNumber="386">
<table id="48AD5ABF376000666994FD8E07399C4A" box="[163,1366,543,768]" gridcols="5" gridrows="6" pageId="3" pageNumber="386">
<tr id="849DAA5D376000666994FD8E07399D7D" box="[163,1366,543,567]" gridrow="0" pageId="3" pageNumber="386" rowspan-0="1">
<th id="C74CC321376000666854FD8E03BA9D7D" box="[355,469,543,567]" gridcol="1" gridrow="0" pageId="3" pageNumber="386">Fe</th>
<th id="C74CC321376000666BA3FD8E00809D7D" box="[660,751,543,567]" gridcol="2" gridrow="0" pageId="3" pageNumber="386">Pa</th>
<th id="C74CC321376000666A4DFD8E063C9D7D" box="[890,1107,543,567]" gridcol="3" gridrow="0" pageId="3" pageNumber="386">Ti</th>
<th id="C74CC321376000666DF5FD8E07399D7D" box="[1218,1366,543,567]" gridcol="4" gridrow="0" pageId="3" pageNumber="386">Mt</th>
</tr>
<tr id="849DAA5D376000666994FDD607399D15" box="[163,1366,583,607]" gridrow="1" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FDD602BF9D15" box="[163,208,583,607]" gridcol="0" gridrow="1" pageId="3" pageNumber="386">Palp</th>
<td id="C74CC321376000666854FDD603BA9D15" box="[355,469,583,607]" gridcol="1" gridrow="1" pageId="3" pageNumber="386">3d 1p 1r</td>
<td id="C74CC321376000666BA3FDD600809D15" box="[660,751,583,607]" gridcol="2" gridrow="1" pageId="3" pageNumber="386">1p</td>
<td id="C74CC321376000666A4DFDD6063C9D15" box="[890,1107,583,607]" gridcol="3" gridrow="1" pageId="3" pageNumber="386">2p 2r</td>
<td id="C74CC321376000666DF5FDD607399D15" box="[1218,1366,583,607]" gridcol="4" gridrow="1" pageId="3" pageNumber="386">-</td>
</tr>
<tr id="849DAA5D376000666994FDFE07399DCD" box="[163,1366,623,647]" gridrow="2" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FDFE02BF9DCD" box="[163,208,623,647]" gridcol="0" gridrow="2" pageId="3" pageNumber="386">I</th>
<td id="C74CC321376000666854FDFE03BA9DCD" box="[355,469,623,647]" gridcol="1" gridrow="2" pageId="3" pageNumber="386">3d 3p 3(2)r</td>
<td id="C74CC321376000666BA3FDFE00809DCD" box="[660,751,623,647]" gridcol="2" gridrow="2" pageId="3" pageNumber="386">spineless</td>
<td id="C74CC321376000666A4DFDFE063C9DCD" box="[890,1107,623,647]" gridcol="3" gridrow="2" pageId="3" pageNumber="386">2-2-2-2-2v</td>
<td id="C74CC321376000666DF5FDFE07399DCD" box="[1218,1366,623,647]" gridcol="4" gridrow="2" pageId="3" pageNumber="386">2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD0607399DE5" box="[163,1366,663,687]" gridrow="3" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD0602BF9DE5" box="[163,208,663,687]" gridcol="0" gridrow="3" pageId="3" pageNumber="386">II</th>
<td id="C74CC321376000666854FD0603BA9DE5" box="[355,469,663,687]" gridcol="1" gridrow="3" pageId="3" pageNumber="386">3d 4(2) 2r</td>
<td id="C74CC321376000666BA3FD0600809DE5" box="[660,751,663,687]" gridcol="2" gridrow="3" pageId="3" pageNumber="386">spineless</td>
<td id="C74CC321376000666A4DFD06063C9DE5" box="[890,1107,663,687]" gridcol="3" gridrow="3" pageId="3" pageNumber="386">1r 2-2-2-2-2v</td>
<td id="C74CC321376000666DF5FD0607399DE5" box="[1218,1366,663,687]" gridcol="4" gridrow="3" pageId="3" pageNumber="386">2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD2E07399D92" box="[163,1366,703,728]" gridrow="4" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD2E02BF9D92" box="[163,208,703,728]" gridcol="0" gridrow="4" pageId="3" pageNumber="386">III</th>
<td id="C74CC321376000666854FD2E03BA9D92" box="[355,469,703,728]" gridcol="1" gridrow="4" pageId="3" pageNumber="386">3d 4p 3(4)r</td>
<td id="C74CC321376000666BA3FD2E00809D92" box="[660,751,703,728]" gridcol="2" gridrow="4" pageId="3" pageNumber="386">1p 1r</td>
<td id="C74CC321376000666A4DFD2E063C9D92" box="[890,1107,703,728]" gridcol="3" gridrow="4" pageId="3" pageNumber="386">2d 2p 2r 2-2-2v</td>
<td id="C74CC321376000666DF5FD2E07399D92" box="[1218,1366,703,728]" gridcol="4" gridrow="4" pageId="3" pageNumber="386">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD7907399C4A" box="[163,1366,744,768]" gridrow="5" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD7902BF9C4A" box="[163,208,744,768]" gridcol="0" gridrow="5" pageId="3" pageNumber="386">IV</th>
<td id="C74CC321376000666854FD7903BA9C4A" box="[355,469,744,768]" gridcol="1" gridrow="5" pageId="3" pageNumber="386">3d 3(4)p 2r</td>
<td id="C74CC321376000666BA3FD7900809C4A" box="[660,751,744,768]" gridcol="2" gridrow="5" pageId="3" pageNumber="386">1p 1r</td>
<td id="C74CC321376000666A4DFD79063C9C4A" box="[890,1107,744,768]" gridcol="3" gridrow="5" pageId="3" pageNumber="386">1(3)d 2p 2r 2(1)-2-2v</td>
<td id="C74CC321376000666DF5FD7907399C4A" box="[1218,1366,744,768]" gridcol="4" gridrow="5" pageId="3" pageNumber="386">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973760FF9A69A0F81E061B98A5" ID-DOI="http://doi.org/10.5281/zenodo.14386393" ID-Zenodo-Dep="14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="3" pageNumber="386" startId="3.[151,264,1935,1959]" targetBox="[151,1435,829,1910]" targetPageId="3" targetType="figure">
<paragraph id="3A12A81F3760FF9A69A0F81E061B98A5" blockId="3.[151,1436,1935,2031]" pageId="3" pageNumber="386">
<emphasis id="08D9740D3760FF9A69A0F81E032F98ED" bold="true" box="[151,320,1935,1959]" pageId="3" pageNumber="386">FIGURES 5–8.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3760FF9A68F9F80100D398ED" authority="Jager." authorityName="Jager." box="[462,700,1935,1959]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3760FF9A68F9F801001B98ED" box="[462,628,1935,1959]" italics="true" pageId="3" pageNumber="386">Bowie neukoeln</emphasis>
Jäger.
</taxonomicName>
5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3760FF9A6A5FF82501E59881" box="[872,906,1972,1995]" italics="true" pageId="3" pageNumber="386">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3760FF9A6D60F825061C9881" box="[1111,1139,1972,1995]" italics="true" pageId="3" pageNumber="386">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3760FF9A6C26F82207229880" box="[1297,1357,1971,1994]" italics="true" pageId="3" pageNumber="386">MC—</emphasis>
anterior constriction of median plate,
<emphasis id="08D9740D3760FF9A6888F849038B98A5" box="[447,484,2008,2031]" italics="true" pageId="3" pageNumber="386">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3760FF9A6BC9F846017898A4" box="[766,791,2007,2030]" italics="true" pageId="3" pageNumber="386">Sp</emphasis>
—spermatheca,
<emphasis id="08D9740D3760FF9A6A8BF84901B498A5" box="[956,987,2008,2031]" italics="true" pageId="3" pageNumber="386">VF</emphasis>
—vulval folds.
</paragraph>
</caption>
<paragraph id="3A12A81F3767FF9D69F0FF0607709E57" blockId="4.[151,1436,150,357]" pageId="4" pageNumber="387">
Female genitalia as shown in
<figureCitation id="A296B49A3767FF9D6B2CFF0600EE9FFB" box="[539,641,150,177]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Figs 5–8</figureCitation>
. Epigyne 1.4 times wider than long. Median plate (
<emphasis id="08D9740D3767FF9D6DFBFF06069B9FFA" box="[1228,1268,151,176]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
) with rounded lateral sides, without anterior keel. Lateral teeth (
<emphasis id="08D9740D3767FF9D6B88FF2A00B19F9E" box="[703,734,187,212]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
) well developed, large, located posterior of median plate, with blunt tips (in
<figureCitation id="A296B49A3767FF9D6801FF4E03F59FB3" box="[310,410,223,249]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Figs 6–7</figureCitation>
shape and position of
<emphasis id="08D9740D3767FF9D6B91FF4E00AA9FB2" box="[678,709,223,248]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
changed due to maceration). Spermathecae (
<emphasis id="08D9740D3767FF9D6DF8FF4E06849FB3" box="[1231,1259,223,249]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
) bottle-shaped, separated by ca. half spermatheca’s length (position of
<emphasis id="08D9740D3767FF9D6A37FE9201739E57" box="[768,796,259,285]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
could be slightly affected due to maceration).
</paragraph>
<paragraph id="3A12A81F3767FF9D69F0FEB607309E0B" blockId="4.[151,1436,150,357]" box="[199,1375,295,321]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FEB6037E9E0B" bold="true" box="[199,273,295,321]" pageId="4" pageNumber="387">Notes.</emphasis>
<taxonomicName id="FDADD39C3767FF9D682FFEB603A59E0B" box="[280,458,295,321]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3767FF9D682FFEB603A59E0B" box="[280,458,295,321]" italics="true" pageId="4" pageNumber="387">Bowie neukoeln</emphasis>
</taxonomicName>
was previously known from Malay Peninsula only, this record is first for Borneo.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69F0FEDB03CA9EA0" pageId="4" pageNumber="387" type="distribution">
<paragraph id="3A12A81F3767FF9D69F0FEDB067C9E2F" blockId="4.[151,1436,150,357]" box="[199,1043,330,357]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FEDB030E9E2E" bold="true" box="[199,353,330,356]" pageId="4" pageNumber="387">Distribution.</emphasis>
<collectingCountry id="42BAE88F3767FF9D6850FEDA03A19E2F" box="[359,462,331,357]" name="Malaysia" pageId="4" pageNumber="387">Malaysia</collectingCountry>
(Malay Peninsula and Borneo:
<collectingRegion id="F86966FD3767FF9D6A06FEDA01169E2F" box="[817,889,331,357]" country="Malaysia" name="Sabah" pageId="4" pageNumber="387">Sabah</collectingRegion>
;
<figureCitation id="A296B49A3767FF9D6AB2FEDA06689E2F" box="[901,1031,331,357]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="4" pageNumber="387">Figs 68–69</figureCitation>
).
</paragraph>
<paragraph id="3A12A81F3767FF9D69A0FE3D03AA9E8C" blockId="4.[151,453,428,490]" box="[151,453,428,454]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0FE3D03AA9E8C" bold="true" box="[151,453,428,454]" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D69A0FE3D030F9E8C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,352,428,454]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D69A0FE3D030F9E8C" bold="true" box="[151,352,428,454]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763767FF9D685FFE3D03AA9E8C" box="[360,453,428,454]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="3A12A81F3767FF9D69A0FE4103CA9EA0" blockId="4.[151,453,428,490]" box="[151,421,464,490]" pageId="4" pageNumber="387">
(
<figureCitation id="A296B49A3767FF9D69A8FE41036B9EA0" box="[159,260,464,490]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6827FE4103259EA0" box="[272,330,464,490]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">9–12</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6861FE4103F29EA0" box="[342,413,464,490]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="4" pageNumber="387">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69A0FD8906F49D1C" pageId="4" pageNumber="387" type="materials_examined">
<paragraph id="3A12A81F3767FF9D69A0FD8906F49D1C" blockId="4.[151,1436,535,671]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0FD8903299D78" bold="true" box="[151,326,536,562]" pageId="4" pageNumber="387">Type material.</emphasis>
<materialsCitation id="8AC5A2423767FF9D687AFD8900049D1C" collectingDate="1994-08-30" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1020" latitude="4.4666667" location="Lucia Mt." longLatPrecision="1306" longitude="117.933334" municipality="Borneo" pageId="4" pageNumber="387" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="08D9740D3767FF9D687AFD8900939D78" bold="true" box="[333,764,535,562]" pageId="4" pageNumber="387">
<collectingCountry id="42BAE88F3767FF9D687AFD8903889D78" box="[333,487,536,562]" name="Malaysia" pageId="4" pageNumber="387">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D3767FF9D68C3FD8600119D78" bold="true" box="[500,638,535,562]" italics="true" pageId="4" pageNumber="387">
<collectingRegion id="F86966FD3767FF9D68C3FD8600119D78" box="[500,638,535,562]" country="Malaysia" name="Sabah" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D68C3FD8600529D78" authorityName="Li &amp; Yao" authorityYear="2022" box="[500,573,535,562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
<typeStatus id="E51616BD3767FF9D6BB8FD8900939D78" box="[655,764,536,562]" pageId="4" pageNumber="387" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="2CAB63963767FF9D6A3CFD88014F9D79" box="[779,800,537,563]" count="1" pageId="4" pageNumber="387" type="female">♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A18FD8901E39D78" box="[815,908,536,562]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="387" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653767FF9D6A97FD88019A9D79" box="[928,1013,537,563]" pageId="4" pageNumber="387">Borneo</collectingMunicipality>
,
<location id="3F72FEC43767FF9D6D36FD89061F9D79" LSID="urn:lsid:plazi:treatment:B20419093762FF9C69A0F89901EC99E0:3F72FEC43767FF9D6D36FD89061F9D79" box="[1025,1136,536,563]" country="Malaysia" latitude="4.4666667" longLatPrecision="1306" longitude="117.933334" municipality="Borneo" name="Lucia Mt." pageId="4" pageNumber="387" stateProvince="Sabah">Lucia Mt.</location>
[
<geoCoordinate id="5F99CED83767FF9D6D48FD8906BA9D78" box="[1151,1237,536,562]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="387" precision="925" value="4.4666667">4°28’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83767FF9D6DD5FD89073E9D79" box="[1250,1361,536,563]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="4" pageNumber="387" precision="925" value="117.933334">117°56’E</geoCoordinate>
],
<quantity id="FD5505FA3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">
<elevation id="B1804F2C3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<date id="4E138EDF3767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">
<collectingDate id="5E5777373767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">30 August 1994</collectingDate>
</date>
,
<collectorName id="9758CDC93767FF9D684EFDAD00549D1C" box="[377,571,572,598]" pageId="4" pageNumber="387">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8AC5A2423767FF9D6B45FDAC06F89D1C" box="[626,1175,572,599]" collectionCode="IBSS" country="Malaysia" pageId="4" pageNumber="387" specimenCount="2" specimenCount-female="2" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="E51616BD3767FF9D6B45FDAC008C9D1D" box="[626,739,573,599]" pageId="4" pageNumber="387" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CAB63963767FF9D6BDAFDAD017F9D1C" box="[749,784,572,598]" count="2" pageId="4" pageNumber="387" type="female">2♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A28FDAD01329D1C" box="[799,861,572,598]" pageId="4" pageNumber="387">IBSS</collectionCode>
) together with the
<typeStatus id="E51616BD3767FF9D6D06FDAD06F89D1C" box="[1073,1175,572,598]" pageId="4" pageNumber="387" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69F0FDF100059DD4" pageId="4" pageNumber="387" type="etymology">
<paragraph id="3A12A81F3767FF9D69F0FDF100059DD4" blockId="4.[151,1436,535,671]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FDF103219D30" bold="true" box="[199,334,608,634]" pageId="4" pageNumber="387">Etymology.</emphasis>
The specific name is derived from the generic name of small wild cat from Southeast Asia,
<emphasis id="08D9740D3767FF9D69A0FD1503639DD4" box="[151,268,644,670]" italics="true" pageId="4" pageNumber="387">Catopuma</emphasis>
Severtzov; noun in apposition.
</paragraph>
</subSubSection>
<caption id="6ED2F8973767FF9D69A0F9310172984A" ID-DOI="http://doi.org/10.5281/zenodo.14386395" ID-Zenodo-Dep="14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387" startId="4.[151,264,1696,1720]" targetBox="[151,1435,694,1672]" targetPageId="4" targetType="figure">
<paragraph id="3A12A81F3767FF9D69A0F9310172984A" blockId="4.[151,1436,1696,1792]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0F931032999F2" bold="true" box="[151,326,1696,1720]" pageId="4" pageNumber="387">FIGURES 9–12.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3767FF9D688AF930000A99F2" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[445,613,1697,1720]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D688AF930000A99F2" box="[445,613,1697,1720]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6B5DF93100D699F2" bold="true" box="[618,697,1696,1720]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6B5DF93100D699F2" box="[618,697,1696,1720]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3767FF9D6BFBF95400819996" box="[716,750,1733,1756]" italics="true" pageId="4" pageNumber="387">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3767FF9D6A8BF95401B79996" box="[956,984,1733,1756]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3767FF9D6D40F95506DC9991" box="[1143,1203,1732,1755]" italics="true" pageId="4" pageNumber="387">MC—</emphasis>
anterior constriction of median plate,
<emphasis id="08D9740D3767FF9D6811F9780324984A" box="[294,331,1769,1792]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3767FF9D6B52F979001199B5" box="[613,638,1768,1791]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
—spermatheca.
</paragraph>
</caption>
<subSubSection id="72B7FB943767FF9D69F0F8D6073198E3" pageId="4" pageNumber="387" type="diagnosis">
<paragraph id="3A12A81F3767FF9D69F0F8D6073198E3" blockId="4.[151,1437,1863,2034]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F8D6032D982B" bold="true" box="[199,322,1863,1889]" pageId="4" pageNumber="387">Diagnosis.</emphasis>
By shape of the median plate (
<emphasis id="08D9740D3767FF9D6BA1F8D900D1982B" box="[662,702,1864,1889]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
) widest anteriorly females of
<taxonomicName id="FDADD39C3767FF9D6D3EF8D906AD982B" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1033,1218,1864,1889]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D6D3EF8D906AD982B" box="[1033,1218,1864,1889]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6DFEF8D9074E9828" bold="true" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6DFEF8D9074E9828" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="FDADD39C3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" italics="true" pageId="4" pageNumber="387">B. neukoeln</emphasis>
</taxonomicName>
, but may be easily distinguished by 1)
<emphasis id="08D9740D3767FF9D6A74F8FD010498CF" box="[835,875,1900,1925]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
with long anterior constriction (
<emphasis id="08D9740D3767FF9D6DE6F8FD069498CC" box="[1233,1275,1900,1926]" italics="true" pageId="4" pageNumber="387">MC</emphasis>
) (vs. short; cf.
<figureCitation id="A296B49A3767FF9D69A0F80102B698E0" box="[151,217,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 9</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6838F801033D98E0" box="[271,338,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 5</figureCitation>
); 2) vulval folds (
<emphasis id="08D9740D3767FF9D6B2BF801005198E3" box="[540,574,1936,1961]" italics="true" pageId="4" pageNumber="387">VF</emphasis>
) almost touching each other (vs. widely spaced; cf.
<figureCitation id="A296B49A3767FF9D6DB1F80106B998E0" box="[1158,1238,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 10</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6C3BF801072098E3" box="[1292,1359,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9C69F0F822073999CC" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387" type="description">
<paragraph id="3A12A81F3767FF9D69F0F82207F398B8" blockId="4.[151,1437,1863,2034]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F82203369887" bold="true" box="[199,345,1971,1997]" pageId="4" pageNumber="387">Description.</emphasis>
<emphasis id="08D9740D3767FF9D6856F82503D29884" box="[353,445,1972,1998]" italics="true" pageId="4" pageNumber="387">Female.</emphasis>
(
<figureCitation id="A296B49A3767FF9D68FAF825005A9884" box="[461,565,1972,1998]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
). Total length 11.76. Prosoma 5.79 long, 4.28 wide. Opisthosoma 5.37 long, 3.44 wide. Prosoma reddish brown, with brown median band. Median band widened and with projections near
</paragraph>
<paragraph id="3A12A81F3767FF9D69BDF7A307F39701" blockId="4.[138,1436,2094,2123]" box="[138,1436,2094,2123]" pageId="4" pageNumber="387">
NEW AND POORLY KNOWN SPECIES OF
<taxonomicName id="FDADD39C3767FF9D6B16F7A300C8970F" authority="FROM" authorityName="FROM" box="[545,679,2098,2117]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="08D9740D3767FF9D6B16F7A3000B970F" box="[545,612,2098,2117]" italics="true" pageId="4" pageNumber="387">BOWIE</emphasis>
FROM
</taxonomicName>
SOUTHEAST ASIA
<emphasis id="08D9740D3767FF9D6AB9F7A2018A9700" box="[910,997,2099,2122]" italics="true" pageId="4" pageNumber="387">Zootaxa</emphasis>
5543 (3) © 2024 Magnolia Press
<emphasis id="08D9740D3767FF9D6C60F7A3070F9700" bold="true" box="[1367,1376,2098,2122]" pageId="4" pageNumber="387">·</emphasis>
387
</paragraph>
<paragraph id="3A12A81F3766FF9C69A0FF0606FB9E0B" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">ocular area. Lateral bands absent. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum light brown without pattern. Labium dark brown. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma light brown with yellowish cardiac mark and series of poorly visible gray spots. Lateral sides of opisthosoma light brown with blurry spots. Ventral part of opisthosoma light brown with yellow spots forming two poorly visible V-marks. Spinnerets light brown.</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FEDA07109EC3" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">Eye diameters: AME 0.22, ALE 0.18, PME 0.31, PLE 0.32; interdistances: AME–AME 0.19, AME–ALE 0.38, PME–PME 0.26, PME–PLE 0.42, AME–PME 0.11, ALE–PLE 0.17. Clypeus height at AME 0.14, at ALE 0.51.</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FE0207629D53" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">
For palp and legs measurements see
<tableCitation id="772F9DA43766FF9C6B5AFE0200AC9EE7" box="[621,707,403,429]" captionStart="TABLE 3" captionStartId="5.[151,239,620,644]" captionText="TABLE 3. Palp and legs measurements of female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 3</tableCitation>
. Palp coloration: femora, patellae, tibiae light brown; tarsi light brown with darker tip. Leg coloration: femora I–IV light brown with poorly visible spots. Patellae I–IV light brown. Tibiae I–II light brown; III–IV light brown with blurred brown semi-rings. Metatarsi I–II light brown; III–IV light brown with blurred brown semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA43766FF9C6D84FE6E07669D53" box="[1203,1289,511,537]" captionStart="TABLE 4" captionStartId="5.[151,239,971,995]" captionText="TABLE 4. Palp and legs spination for the female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 4</tableCitation>
.
</paragraph>
<caption id="6ED2F8973766FF9C69A0FDFD01B29DCF" box="[151,989,620,645]" pageId="5" pageNumber="388" startId="5.[151,239,620,644]" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FDFD01B29DCF" blockId="5.[151,989,620,645]" box="[151,989,620,645]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FDFD03659DCE" bold="true" box="[151,266,620,644]" pageId="5" pageNumber="388">TABLE 3.</emphasis>
Palp and legs measurements of female of
<taxonomicName id="FDADD39C3766FF9C6BF9FDFC01EF9DCE" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[718,896,621,644]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF9FDFC01EF9DCE" box="[718,896,621,644]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB0FDFC01B29DCF" box="[903,989,621,645]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C684EFD0407359C3C" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FD0407359C3C" box="[163,1370,661,886]" gridcols="7" gridrows="6" pageId="5" pageNumber="388" rowsContinueIn="5.[163,1366,1011,1237]">
<tr id="849DAA5D376600666994FD0407359DE7" box="[163,1370,661,685]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666858FD0403F49DE7" box="[367,411,661,685]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666B19FD0400359DE7" box="[558,602,661,685]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666BDBFD0401779DE7" box="[748,792,661,685]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666A9CFD0401B89DE7" box="[939,983,661,685]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
<th id="C74CC321376600666D5EFD0406FA9DE7" box="[1129,1173,661,685]" gridcol="5" gridrow="0" pageId="5" pageNumber="388">Ta</th>
<th id="C74CC321376600666C16FD0407359DE7" box="[1313,1370,661,685]" gridcol="6" gridrow="0" pageId="5" pageNumber="388">Total</th>
</tr>
<tr id="849DAA5D376600666994FD2C07359D9F" box="[163,1370,701,725]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD2C02BF9D9F" box="[163,208,701,725]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666858FD2C03F49D9F" box="[367,411,701,725]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">2.00</td>
<td id="C74CC321376600666B19FD2C00359D9F" box="[558,602,701,725]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1.06</td>
<td id="C74CC321376600666BDBFD2C01779D9F" box="[748,792,701,725]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1.19</td>
<td id="C74CC321376600666A9CFD2C01B89D9F" box="[939,983,701,725]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
<td id="C74CC321376600666D5EFD2C06FA9D9F" box="[1129,1173,701,725]" gridcol="5" gridrow="1" pageId="5" pageNumber="388">1.35</td>
<td id="C74CC321376600666C16FD2C07359D9F" box="[1313,1370,701,725]" gridcol="6" gridrow="1" pageId="5" pageNumber="388">5.6</td>
</tr>
<tr id="849DAA5D376600666994FD7407359DB7" box="[163,1370,741,765]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD7402BF9DB7" box="[163,208,741,765]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666858FD7403F49DB7" box="[367,411,741,765]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">4.08</td>
<td id="C74CC321376600666B19FD7400359DB7" box="[558,602,741,765]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">2.17</td>
<td id="C74CC321376600666BDBFD7401779DB7" box="[748,792,741,765]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">3.77</td>
<td id="C74CC321376600666A9CFD7401B89DB7" box="[939,983,741,765]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">3.10</td>
<td id="C74CC321376600666D5EFD7406FA9DB7" box="[1129,1173,741,765]" gridcol="5" gridrow="2" pageId="5" pageNumber="388">1.00</td>
<td id="C74CC321376600666C16FD7407359DB7" box="[1313,1370,741,765]" gridcol="6" gridrow="2" pageId="5" pageNumber="388">14.12</td>
</tr>
<tr id="849DAA5D376600666994FC9C07359C6C" box="[163,1370,781,806]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FC9C02BF9C6C" box="[163,208,781,806]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666858FC9C03F49C6C" box="[367,411,781,806]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3.94</td>
<td id="C74CC321376600666B19FC9C00359C6C" box="[558,602,781,806]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">2.14</td>
<td id="C74CC321376600666BDBFC9C01779C6C" box="[748,792,781,806]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">3.32</td>
<td id="C74CC321376600666A9CFC9C01B89C6C" box="[939,983,781,806]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2.92</td>
<td id="C74CC321376600666D5EFC9C06FA9C6C" box="[1129,1173,781,806]" gridcol="5" gridrow="3" pageId="5" pageNumber="388">1.05</td>
<td id="C74CC321376600666C16FC9C07359C6C" box="[1313,1370,781,806]" gridcol="6" gridrow="3" pageId="5" pageNumber="388">13.37</td>
</tr>
<tr id="849DAA5D376600666994FCA407359C04" box="[163,1370,821,846]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCA402BF9C04" box="[163,208,821,846]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666858FCA403F49C04" box="[367,411,821,846]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3.31</td>
<td id="C74CC321376600666B19FCA400359C04" box="[558,602,821,846]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1.71</td>
<td id="C74CC321376600666BDBFCA401779C04" box="[748,792,821,846]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">2.45</td>
<td id="C74CC321376600666A9CFCA401B89C04" box="[939,983,821,846]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">2.91</td>
<td id="C74CC321376600666D5EFCA406FA9C04" box="[1129,1173,821,846]" gridcol="5" gridrow="4" pageId="5" pageNumber="388">1.09</td>
<td id="C74CC321376600666C16FCA407359C04" box="[1313,1370,821,846]" gridcol="6" gridrow="4" pageId="5" pageNumber="388">11.47</td>
</tr>
<tr id="849DAA5D376600666994FCCF07359C3C" box="[163,1370,862,886]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCCF02BF9C3C" box="[163,208,862,886]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666858FCCF03F49C3C" box="[367,411,862,886]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">4.61</td>
<td id="C74CC321376600666B19FCCF00359C3C" box="[558,602,862,886]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1.77</td>
<td id="C74CC321376600666BDBFCCF01779C3C" box="[748,792,862,886]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">4.00</td>
<td id="C74CC321376600666A9CFCCF01B89C3C" box="[939,983,862,886]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">5.34</td>
<td id="C74CC321376600666D5EFCCF06FA9C3C" box="[1129,1173,862,886]" gridcol="5" gridrow="5" pageId="5" pageNumber="388">1.52</td>
<td id="C74CC321376600666C16FCCF07359C3C" box="[1313,1370,862,886]" gridcol="6" gridrow="5" pageId="5" pageNumber="388">17.24</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973766FF9C69A0FC5A01B99CA9" box="[151,982,971,995]" pageId="5" pageNumber="388" startId="5.[151,239,971,995]" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FC5A01B99CA9" blockId="5.[151,982,971,995]" box="[151,982,971,995]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FC5A03659CA9" bold="true" box="[151,266,971,995]" pageId="5" pageNumber="388">TABLE 4.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C3766FF9C6BF0FC5D01159CA9" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[711,890,972,995]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF0FC5D01159CA9" box="[711,890,972,995]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB7FC5A01B99CA9" box="[896,982,971,995]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C68B8FC6507399B9F" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FC6207399B9F" box="[163,1366,1011,1237]" gridcols="5" gridrows="6" pageId="5" pageNumber="388" rowsContinueFrom="5.[163,1370,661,886]">
<tr id="849DAA5D376600666994FC6207399B41" box="[163,1366,1011,1035]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666854FC6203BA9B41" box="[355,469,1011,1035]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666BA3FC6200809B41" box="[660,751,1011,1035]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666ABEFC62062A9B41" box="[905,1093,1011,1035]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666DF5FC6207399B41" box="[1218,1366,1011,1035]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
</tr>
<tr id="849DAA5D376600666994FB8D07399B7E" box="[163,1366,1052,1076]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB8D02BF9B7E" box="[163,208,1052,1076]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666854FB8D03BA9B7E" box="[355,469,1052,1076]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">3d 1p 1r</td>
<td id="C74CC321376600666BA3FB8D00809B7E" box="[660,751,1052,1076]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1(0)p</td>
<td id="C74CC321376600666ABEFB8D062A9B7E" box="[905,1093,1052,1076]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1d 2p 1r</td>
<td id="C74CC321376600666DF5FB8D07399B7E" box="[1218,1366,1052,1076]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
</tr>
<tr id="849DAA5D376600666994FBD507399B16" box="[163,1366,1092,1116]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBD502BF9B16" box="[163,208,1092,1116]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666854FBD503BA9B16" box="[355,469,1092,1116]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">3d 3p 3(2)r</td>
<td id="C74CC321376600666BA3FBD500809B16" box="[660,751,1092,1116]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBD5062A9B16" box="[905,1093,1092,1116]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBD507399B16" box="[1218,1366,1092,1116]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FBFD07399BCE" box="[163,1366,1132,1156]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBFD02BF9BCE" box="[163,208,1132,1156]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666854FBFD03BA9BCE" box="[355,469,1132,1156]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FBFD00809BCE" box="[660,751,1132,1156]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBFD062A9BCE" box="[905,1093,1132,1156]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBFD07399BCE" box="[1218,1366,1132,1156]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB0507399BE7" box="[163,1366,1172,1197]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB0502BF9BE7" box="[163,208,1172,1197]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666854FB0503BA9BE7" box="[355,469,1172,1197]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3d 4p 4r</td>
<td id="C74CC321376600666BA3FB0500809BE7" box="[660,751,1172,1197]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB05062A9BE7" box="[905,1093,1172,1197]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">3d 2p 2r 2-2v</td>
<td id="C74CC321376600666DF5FB0507399BE7" box="[1218,1366,1172,1197]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">4p 5r 2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB2C07399B9F" box="[163,1366,1213,1237]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB2C02BF9B9F" box="[163,208,1213,1237]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666854FB2C03BA9B9F" box="[355,469,1213,1237]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FB2C00809B9F" box="[660,751,1213,1237]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB2C062A9B9F" box="[905,1093,1213,1237]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">3(2)d 2p 2r 2-2-2v</td>
<td id="C74CC321376600666DF5FB2C07399B9F" box="[1218,1366,1213,1237]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FAB903E59AE4" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
Female genitalia as shown in
<figureCitation id="A296B49A3766FF9C6B2BFAB900FE9A08" box="[540,657,1320,1346]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="5" pageNumber="388">Figs 9–12</figureCitation>
. Epigyne 1.5 times wider than long. Median plate (
<emphasis id="08D9740D3766FF9C6DE9FAB807699A08" box="[1246,1286,1321,1346]" italics="true" pageId="5" pageNumber="388">MP</emphasis>
) trapezoidal, strongly sclerotized anteriorly. Lateral teeth (
<emphasis id="08D9740D3766FF9C6B87FADC00A09A2C" box="[688,719,1357,1382]" italics="true" pageId="5" pageNumber="388">LT</emphasis>
) well developed, located posterior of median plate, slightly extending beyond epigastric furrow, with rounded tips. Spermathecae (
<emphasis id="08D9740D3766FF9C6A8BFAE001B79AC1" box="[956,984,1393,1419]" italics="true" pageId="5" pageNumber="388">Sp</emphasis>
) large, piriform, separated by a third of spermatheca’s length.
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FA29073999CC" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0FA29037E9A98" bold="true" box="[199,273,1464,1490]" pageId="5" pageNumber="388">Notes.</emphasis>
Since males
<taxonomicName id="FDADD39C3766FF9C689DFA2800579A98" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[426,568,1465,1490]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C689DFA2800579A98" box="[426,568,1465,1490]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6B76FA2900F39A98" bold="true" box="[577,668,1464,1490]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6B76FA2900F39A98" box="[577,668,1464,1490]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are unknown, it is difficult to place it in a species group. There are species with a similar shape of median plate and lateral teeth situated at the posterior margin of the epigyne in
<emphasis id="08D9740D3766FF9C69A0F99102919950" box="[151,254,1536,1562]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
and
<taxonomicName id="FDADD39C3766FF9C680EF99003B49950" authorityName="Jager" authorityYear="2022" box="[313,475,1537,1562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C680EF99003B49950" box="[313,475,1537,1562]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
species groups. Because the new species have spermathecae consisting of only one compartment, while those in
<emphasis id="08D9740D3766FF9C68E8F9B500299974" box="[479,582,1572,1598]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
-group have particularly structured spermathecae, we place
<taxonomicName id="FDADD39C3766FF9C6DD2F9B4071C9974" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1253,1395,1573,1598]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6DD2F9B4071C9974" box="[1253,1395,1573,1598]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6C4CF9B502A79928" bold="true" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6C4CF9B502A79928" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the
<taxonomicName id="FDADD39C3766FF9C6820F9D803D69928" authorityName="Jager" authorityYear="2022" box="[279,441,1609,1634]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C6820F9D803D69928" box="[279,441,1609,1634]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
-species group. Thus, with the description of the new species, this group now consists of five species:
<taxonomicName id="FDADD39C3766FF9C681EF9FC038C99CC" authorityName="Omelko &amp; Fomichev" authorityYear="2023" box="[297,483,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="binturong">
<emphasis id="08D9740D3766FF9C681EF9FC038C99CC" box="[297,483,1644,1670]" italics="true" pageId="5" pageNumber="388">Bowie binturong</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C68C5F9FC001099CC" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[498,639,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C68C5F9FC001099CC" box="[498,639,1645,1670]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6BB1F9FD008F99CC" bold="true" box="[646,736,1644,1670]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6BB1F9FD008F99CC" box="[646,736,1644,1670]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="FDADD39C3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" italics="true" pageId="5" pageNumber="388">B. neukoeln</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C6AB7F9FC063B99CD" authority="Li &amp; Yao" authorityName="Li &amp; Yao" authorityYear="2022" box="[896,1108,1644,1671]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="sabah">
<emphasis id="08D9740D3766FF9C6AB7F9FC018D99CC" box="[896,994,1644,1670]" italics="true" pageId="5" pageNumber="388">B. sabah</emphasis>
Li &amp; Yao
</taxonomicName>
, and
<emphasis id="08D9740D3766FF9C6DB9F9FC073999CC" box="[1166,1366,1645,1670]" italics="true" pageId="5" pageNumber="388">
<taxonomicName id="FDADD39C3766FF9C6DB9F9FC073D99CC" authorityName="Jager" authorityYear="2022" box="[1166,1362,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">B. scarymonsters</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943766FF9C69F0F90101EC99E0" box="[199,899,1680,1707]" pageId="5" pageNumber="388" type="distribution">
<paragraph id="3A12A81F3766FF9C69F0F90101EC99E0" blockId="5.[151,1436,1320,1707]" box="[199,899,1680,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0F901030E99E0" bold="true" box="[199,353,1680,1706]" pageId="5" pageNumber="388">Distribution.</emphasis>
<typeStatus id="E51616BD3766FF9C6850F90003F099E1" box="[359,415,1681,1707]" pageId="5" pageNumber="388">Type</typeStatus>
locality only, Borneo:
<collectingRegion id="F86966FD3766FF9C6B97F901008B99E0" box="[672,740,1680,1706]" country="Malaysia" name="Sabah" pageId="5" pageNumber="388">Sabah</collectingRegion>
(
<figureCitation id="A296B49A3766FF9C6BC4F901011899E0" box="[755,887,1680,1706]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="5" pageNumber="388">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<mods:affiliation id="F8921491F2686B7E766F4156207A5563">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
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<paragraph id="3A12A81F3762FF9869A0F89903819868" blockId="1.[151,494,1799,1863]" box="[151,494,1799,1826]" pageId="1" pageNumber="384">
<heading id="615A1F733762FF9869A0F89903819868" bold="true" box="[151,494,1799,1826]" fontSize="11" level="1" pageId="1" pageNumber="384" reason="1">
<taxonomicName id="FDADD39C3762FF9869A0F89903819868" ID-CoL="C6LJM" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[151,494,1799,1826]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3762FF9869A0F89903819868" bold="true" box="[151,494,1799,1826]" pageId="1" pageNumber="384">
<emphasis id="08D9740D3762FF9869A0F89903369868" bold="true" box="[151,345,1799,1826]" italics="true" pageId="1" pageNumber="384">Bowie neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3762FF986856F89903819868" author="Jager, P." box="[353,494,1800,1826]" pageId="1" pageNumber="384" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
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<paragraph id="3A12A81F3762FF9869A0F8BD03F8980C" blockId="1.[151,494,1799,1863]" box="[151,407,1836,1863]" pageId="1" pageNumber="384">
(
<figureCitation id="A296B49A3762FF9869A8F8BD036B980C" box="[159,260,1836,1862]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="1" pageNumber="384">Figs 1–2</figureCitation>
,
<figureCitation id="A296B49A3762FF986827F8BD0353980D" box="[272,316,1836,1863]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="1" pageNumber="384">5–8</figureCitation>
,
<figureCitation id="A296B49A3762FF98687FF8BD03E0980C" box="[328,399,1836,1862]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="1" pageNumber="384">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943762FF9869A0F8E001EE98C2" box="[151,897,1904,1929]" pageId="1" pageNumber="384" type="reference_group">
<paragraph id="3A12A81F3762FF9869A0F8E001EE98C2" blockId="1.[151,897,1904,1929]" box="[151,897,1904,1929]" pageId="1" pageNumber="384">
<treatmentCitationGroup id="1ABD8F313762FF9869A0F8E001EE98C2" box="[151,897,1904,1929]" pageId="1" pageNumber="384">
<treatmentCitation id="BB0C8E0E3762FF9869A0F8E003DE98C3" author="Jager, P." box="[151,433,1904,1929]" page="98" pageId="1" pageNumber="384" year="2022">
<taxonomicName id="FDADD39C3762FF9869A0F8E003DE98C3" ID-CoL="C6LJM" authority="Jager, 2022: 98" authorityName="Jager" authorityPageNumber="98" authorityYear="2022" box="[151,433,1904,1929]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3762FF9869A0F8E0036198C2" box="[151,270,1904,1928]" italics="true" pageId="1" pageNumber="384">B. neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3762FF986822F8E003DE98C3" author="Jager, P." box="[277,433,1905,1929]" pageId="1" pageNumber="384" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 440–446, 448–460 (
<specimenCount id="2CAB63963762FF986B81F8E000A398C3" box="[694,716,1905,1929]" count="1" pageId="1" pageNumber="384" type="male">♂</specimenCount>
<specimenCount id="2CAB63963762FF986BFBF8E000B498C3" box="[716,731,1905,1929]" count="1" pageId="1" pageNumber="384" type="female">♀</specimenCount>
), not examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943762FF9869A0F82700AF98BF" pageId="1" pageNumber="384" type="materials_examined">
<paragraph id="3A12A81F3762FF9869A0F82700AF98BF" blockId="1.[151,1436,1974,2037]" pageId="1" pageNumber="384">
<emphasis id="08D9740D3762FF9869A0F82703EE989A" bold="true" box="[151,385,1974,2000]" pageId="1" pageNumber="384">Material examined.</emphasis>
<materialsCitation id="8AC5A2423762FF9868BEF82700AF98BF" collectingDate="1993-02-28" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1050" latitude="5.4" location="Crocker Range Ridge" longLatPrecision="1305" longitude="116.1" municipality="Borneo" pageId="1" pageNumber="384" specimenCount="1" specimenCount-female="1" stateProvince="Sabah State">
<emphasis id="08D9740D3762FF9868BEF82700AC989B" bold="true" box="[393,707,1974,2001]" pageId="1" pageNumber="384">
<collectingCountry id="42BAE88F3762FF9868BEF827004C989A" box="[393,547,1974,2000]" name="Malaysia" pageId="1" pageNumber="384">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D3762FF986B07F82700D5989B" bold="true" box="[560,698,1974,2001]" italics="true" pageId="1" pageNumber="384">
<collectingRegion id="F86966FD3762FF986B07F82700D5989B" box="[560,698,1974,2001]" country="Malaysia" name="Sabah" pageId="1" pageNumber="384">
<taxonomicName id="FDADD39C3762FF986B07F8270016989B" authorityName="Li &amp; Yao" authorityYear="2022" box="[560,633,1974,2001]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="2CAB63963762FF986BFCF826008F989B" box="[715,736,1975,2001]" count="1" pageId="1" pageNumber="384" type="female">♀</specimenCount>
(
<collectionCode id="5CBC30DA3762FF986BD8F8260123989B" box="[751,844,1975,2001]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="384" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653762FF986A57F82601DA989B" box="[864,949,1975,2001]" pageId="1" pageNumber="384">Borneo</collectingMunicipality>
,
<location id="3F72FEC43762FF986AF6F82606DA989B" LSID="urn:lsid:plazi:treatment:B20419093762FF9D69A0F899067C9E2F:3F72FEC43762FF986AF6F82606DA989B" box="[961,1205,1975,2001]" country="Malaysia" latitude="5.4" longLatPrecision="1305" longitude="116.1" municipality="Borneo" name="Crocker Range Ridge" pageId="1" pageNumber="384" stateProvince="Sabah State">Crocker Range Ridge</location>
[
<geoCoordinate id="5F99CED83762FF986DF3F8270774989B" box="[1220,1307,1974,2001]" degrees="5" direction="north" minutes="24" orientation="latitude" pageId="1" pageNumber="384" precision="925" value="5.4">5°24’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83762FF986C1FF82607E0989B" box="[1320,1423,1975,2001]" degrees="116" direction="east" minutes="6" orientation="longitude" pageId="1" pageNumber="384" precision="925" value="116.1">116° 6’E</geoCoordinate>
],
<quantity id="FD5505FA3762FF9869A0F84B028198BF" box="[151,238,2010,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="384" unit="m" value="1050.0">
<elevation id="B1804F2C3762FF9869A0F84B028198BF" box="[151,238,2010,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="384" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="4E138EDF3762FF9869CDF84A03AB98BF" box="[250,452,2011,2037]" pageId="1" pageNumber="384" value="1993-02-28">
<collectingDate id="5E5777373762FF9869CDF84A03AB98BF" box="[250,452,2011,2037]" pageId="1" pageNumber="384" value="1993-02-28">28 February 1993</collectingDate>
</date>
,
<collectorName id="9758CDC93762FF9868FAF84A00FF98BF" box="[461,656,2011,2037]" pageId="1" pageNumber="384">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="6ED2F8973761FF9B69A0FB2F03219BB0" ID-DOI="http://doi.org/10.5281/zenodo.14386389" ID-Zenodo-Dep="14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="2" pageNumber="385" startId="2.[151,264,1214,1238]" targetBox="[183,1407,180,1182]" targetPageId="2" targetType="figure">
<paragraph id="3A12A81F3761FF9B69A0FB2F03219BB0" blockId="2.[151,1437,1214,1274]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69A0FB2F03539B9C" bold="true" box="[151,316,1214,1238]" pageId="2" pageNumber="385">FIGURES 1–4.</emphasis>
Female habitus of
<taxonomicName id="FDADD39C3761FF9B6B32FB2E008B9B9D" authority="Jager" authorityName="Jager" box="[517,740,1214,1239]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3761FF9B6B32FB2E00C89B9C" box="[517,679,1214,1238]" italics="true" pageId="2" pageNumber="385">Bowie neukoeln</emphasis>
Jäger
</taxonomicName>
(1–2) and
<taxonomicName id="FDADD39C3761FF9B6A6FFB2E01B69B9C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[856,985,1215,1238]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3761FF9B6A6FFB2E01B69B9C" box="[856,985,1215,1238]" italics="true" pageId="2" pageNumber="385">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3761FF9B6AD6FB2F065C9B9C" bold="true" box="[993,1075,1214,1238]" pageId="2" pageNumber="385">
<taxonomicNameLabel id="13EAC9763761FF9B6AD6FB2F065C9B9C" box="[993,1075,1214,1238]" pageId="2" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="72B7FB943761FF9B69F0FADD07779ABD" pageId="2" pageNumber="385" type="diagnosis">
<paragraph id="3A12A81F3761FF9B69F0FADD07779ABD" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69F0FADD032F9A2C" bold="true" box="[199,320,1356,1382]" pageId="2" pageNumber="385">Diagnosis.</emphasis>
Females of
<taxonomicName id="FDADD39C3761FF9B68F2FADC00299A2C" box="[453,582,1356,1382]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3761FF9B68F2FADC00299A2C" box="[453,582,1356,1382]" italics="true" pageId="2" pageNumber="385">B. neukoeln</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="FDADD39C3761FF9B6A08FADC06109A2D" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[831,1151,1357,1383]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3761FF9B6A08FADC01939A2C" box="[831,1020,1357,1382]" italics="true" pageId="2" pageNumber="385">B. scarymonsters</emphasis>
<bibRefCitation id="5E3CD5EE3761FF9B6D35FADC06109A2D" author="Jager, P." box="[1026,1151,1357,1383]" pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FDADD39C3761FF9B6D84FADC03099AC1" authority="Omelko &amp; Fomichev, 2023" authorityName="Omelko &amp; Fomichev" authorityYear="2023" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="binturong">
<emphasis id="08D9740D3761FF9B6D84FADC07529A2C" box="[1203,1341,1356,1382]" italics="true" pageId="2" pageNumber="385">B. binturong</emphasis>
<bibRefCitation id="5E3CD5EE3761FF9B6C74FADD03099AC1" author="Omelko, M. M. &amp; Fomichev, A. A." pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev, 2023</bibRefCitation>
</taxonomicName>
by shape of median plate (
<emphasis id="08D9740D3761FF9B6BA2FAE000D29AC0" box="[661,701,1393,1418]" italics="true" pageId="2" pageNumber="385">MP</emphasis>
) and spermathecae (
<emphasis id="08D9740D3761FF9B6A93FAE001AF9AC1" box="[932,960,1393,1419]" italics="true" pageId="2" pageNumber="385">Sp</emphasis>
) as well as position of lateral teeth (
<emphasis id="08D9740D3761FF9B6C61FAE0071B9AC0" box="[1366,1396,1393,1418]" italics="true" pageId="2" pageNumber="385">LT</emphasis>
). It can be distinguished from both species by 1) having posterior pocket (pit) of epigyne (vs. absence; cf.
<figureCitation id="A296B49A3761FF9B6DD8FA0507409AE5" box="[1263,1327,1428,1455]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="2" pageNumber="385">Fig. 8</figureCitation>
and
<bibRefCitation id="5E3CD5EE3761FF9B6C55FA0402BC9A99" author="Jager, P." pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 467,
<bibRefCitation id="5E3CD5EE3761FF9B6809FA2900059A99" author="Omelko, M. M. &amp; Fomichev, A. A." box="[318,618,1464,1491]" pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev 2023</bibRefCitation>
: fig. 54); 2) spermathecae separated by gap equal to ca. 0.5 their width (vs. 1 and 0.3 of width, respectively; cf.
<figureCitation id="A296B49A3761FF9B6B11FA4D00089ABD" box="[550,615,1500,1527]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="2" pageNumber="385">Fig. 7</figureCitation>
and
<bibRefCitation id="5E3CD5EE3761FF9B6BABFA4C01759ABD" author="Jager, P." box="[668,794,1501,1527]" pageId="2" pageNumber="385" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 468,
<bibRefCitation id="5E3CD5EE3761FF9B6AB0FA4D06D99ABD" author="Omelko, M. M. &amp; Fomichev, A. A." box="[903,1206,1500,1527]" pageId="2" pageNumber="385" pagination="101 - 116" refId="ref10243" refString="Omelko, M. M. &amp; Fomichev, A. A. (2023) A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jager, 2022. Zootaxa, 5353 (2), 101 - 116. https: // doi. org / 10.11646 / zootaxa. 5353.2.1" type="journal article" year="2023">Omelko &amp; Fomichev 2023</bibRefCitation>
: fig. 53).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943761FF9D69F0F99107309E0B" lastPageId="4" lastPageNumber="387" pageId="2" pageNumber="385" type="description">
<paragraph id="3A12A81F3761FF9B69F0F99101FB99B8" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
<emphasis id="08D9740D3761FF9B69F0F99103369950" bold="true" box="[199,345,1536,1562]" pageId="2" pageNumber="385">Description.</emphasis>
<emphasis id="08D9740D3761FF9B6855F99103D99950" box="[354,438,1536,1562]" italics="true" pageId="2" pageNumber="385">Female</emphasis>
(
<figureCitation id="A296B49A3761FF9B68F0F99100419951" box="[455,558,1536,1563]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="2" pageNumber="385">Figs 1–2</figureCitation>
). Total length 11.75. Prosoma 6.16 long, 4.68 wide. Opisthosoma 5.56 long, 2.93 wide. Prosoma brown, with light brown median band. Median band widened and with projections near ocular area. Lateral bands almost invisible. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum brown, without pattern. Labium dark brown. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma dark brown with distinct yellowish cardiac mark and a series of black spots. Lateral sides of opisthosoma yellowish brown. Ventral part of opisthosoma dark brown with small yellowish spots forming two V-marks. Spinnerets dark brown.
</paragraph>
<paragraph id="3A12A81F3761FF9B69F0F96C07109871" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">Eye diameters: AME 0.26, ALE 0.19, PME 0.40, PLE 0.34; interdistances: AME–AME 0.21, AME–ALE 0.41, PME–PME 0.25, PME–PLE 0.47, AME–PME 0.11, ALE–PLE 0.24. Clypeus height at AME 0.14, at ALE 0.54.</paragraph>
<paragraph id="3A12A81F3761FF9B69F0F8D4029E98A5" blockId="2.[151,1437,1356,2031]" pageId="2" pageNumber="385">
For palp and legs measurements see
<tableCitation id="772F9DA43761FF9B6B5EF8D500D19815" box="[617,702,1860,1887]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetBox="[163,1370,192,417]" captionTargetId="graphics-30@3.[151,1437,180,423]" captionTargetPageId="3" captionText="TABLE 1. Palp and legs measurements of female of Bowie neukoeln Jäger, 2022" pageId="2" pageNumber="385">Table 1</tableCitation>
. Palp coloration: all joints light brown. Leg coloration: femora I brown with yellow longitudinal stripe retrolaterally; II–IV brown with poorly visible yellow spots. Patellae I–IV brown. Tibiae I–II light brown; III–IV light brown with poorly visible yellow semi-rings. Metatarsi I–II light brown; III–IV light brown with poorly visible yellow semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA43761FF9B69A0F845028298A5" box="[151,237,2004,2031]" captionStart="TABLE 2" captionStartId="3.[151,239,502,526]" captionTargetBox="[163,1370,192,417]" captionTargetId="graphics-30@3.[151,1437,180,423]" captionText="TABLE 2. Palp and legs spination for the female of Bowie neukoeln Jäger, 2022" pageId="2" pageNumber="385">Table 2</tableCitation>
.
</paragraph>
<caption id="6ED2F8973760FF9A69A0FF0601979FFA" box="[151,1016,151,176]" pageId="3" pageNumber="386" startId="3.[151,239,151,175]" targetBox="[163,1370,192,417]" targetIsTable="true" targetPageId="3">
<paragraph id="3A12A81F3760FF9A69A0FF0601979FFA" box="[151,1016,151,176]" pageId="3" pageNumber="386">
<emphasis id="08D9740D3760FF9A69A0FF0603659FE5" bold="true" box="[151,266,151,175]" pageId="3" pageNumber="386">TABLE 1.</emphasis>
Palp and legs measurements of female of
<emphasis id="08D9740D3760FF9A6BF9FF0901189FE5" box="[718,887,151,175]" italics="true" pageId="3" pageNumber="386">Bowie neukoeln</emphasis>
Jäger, 2022
</paragraph>
</caption>
<paragraph id="3A12A81F3760FF9A684EFF5107359EEB" pageId="3" pageNumber="386">
<table id="48AD5ABF376000666994FF5107359EEB" box="[163,1370,192,417]" gridcols="7" gridrows="6" pageId="3" pageNumber="386">
<tr id="849DAA5D376000666994FF5107359F92" box="[163,1370,192,216]" gridrow="0" pageId="3" pageNumber="386" rowspan-0="1">
<th id="C74CC321376000666858FF5103F49F92" box="[367,411,192,216]" gridcol="1" gridrow="0" pageId="3" pageNumber="386">Fe</th>
<th id="C74CC321376000666B1AFF5100369F92" box="[557,601,192,216]" gridcol="2" gridrow="0" pageId="3" pageNumber="386">Pa</th>
<th id="C74CC321376000666BDBFF5101779F92" box="[748,792,192,216]" gridcol="3" gridrow="0" pageId="3" pageNumber="386">Ti</th>
<th id="C74CC321376000666A9DFF5101B99F92" box="[938,982,192,216]" gridcol="4" gridrow="0" pageId="3" pageNumber="386">Mt</th>
<th id="C74CC321376000666D5EFF5106FA9F92" box="[1129,1173,192,216]" gridcol="5" gridrow="0" pageId="3" pageNumber="386">Ta</th>
<th id="C74CC321376000666C16FF5107359F92" box="[1313,1370,192,216]" gridcol="6" gridrow="0" pageId="3" pageNumber="386">Total</th>
</tr>
<tr id="849DAA5D376000666994FF7907359E4A" box="[163,1370,232,256]" gridrow="1" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FF7902BF9E4A" box="[163,208,232,256]" gridcol="0" gridrow="1" pageId="3" pageNumber="386">Palp</th>
<td id="C74CC321376000666858FF7903F49E4A" box="[367,411,232,256]" gridcol="1" gridrow="1" pageId="3" pageNumber="386">2.17</td>
<td id="C74CC321376000666B1AFF7900369E4A" box="[557,601,232,256]" gridcol="2" gridrow="1" pageId="3" pageNumber="386">1.03</td>
<td id="C74CC321376000666BDBFF7901779E4A" box="[748,792,232,256]" gridcol="3" gridrow="1" pageId="3" pageNumber="386">1.30</td>
<td id="C74CC321376000666A9DFF7901B99E4A" box="[938,982,232,256]" gridcol="4" gridrow="1" pageId="3" pageNumber="386">-</td>
<td id="C74CC321376000666D5EFF7906FA9E4A" box="[1129,1173,232,256]" gridcol="5" gridrow="1" pageId="3" pageNumber="386">1.42</td>
<td id="C74CC321376000666C16FF7907359E4A" box="[1313,1370,232,256]" gridcol="6" gridrow="1" pageId="3" pageNumber="386">5.92</td>
</tr>
<tr id="849DAA5D376000666994FE8107359E62" box="[163,1370,272,296]" gridrow="2" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FE8102BF9E62" box="[163,208,272,296]" gridcol="0" gridrow="2" pageId="3" pageNumber="386">I</th>
<td id="C74CC321376000666858FE8103F49E62" box="[367,411,272,296]" gridcol="1" gridrow="2" pageId="3" pageNumber="386">4.17</td>
<td id="C74CC321376000666B1AFE8100369E62" box="[557,601,272,296]" gridcol="2" gridrow="2" pageId="3" pageNumber="386">2.20</td>
<td id="C74CC321376000666BDBFE8101779E62" box="[748,792,272,296]" gridcol="3" gridrow="2" pageId="3" pageNumber="386">3.77</td>
<td id="C74CC321376000666A9DFE8101B99E62" box="[938,982,272,296]" gridcol="4" gridrow="2" pageId="3" pageNumber="386">3.15</td>
<td id="C74CC321376000666D5EFE8106FA9E62" box="[1129,1173,272,296]" gridcol="5" gridrow="2" pageId="3" pageNumber="386">1.28</td>
<td id="C74CC321376000666C16FE8107359E62" box="[1313,1370,272,296]" gridcol="6" gridrow="2" pageId="3" pageNumber="386">14.57</td>
</tr>
<tr id="849DAA5D376000666994FEA907359E1B" box="[163,1370,312,337]" gridrow="3" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FEA902BF9E1B" box="[163,208,312,337]" gridcol="0" gridrow="3" pageId="3" pageNumber="386">II</th>
<td id="C74CC321376000666858FEA903F49E1B" box="[367,411,312,337]" gridcol="1" gridrow="3" pageId="3" pageNumber="386">3.97</td>
<td id="C74CC321376000666B1AFEA900369E1B" box="[557,601,312,337]" gridcol="2" gridrow="3" pageId="3" pageNumber="386">2.24</td>
<td id="C74CC321376000666BDBFEA901779E1B" box="[748,792,312,337]" gridcol="3" gridrow="3" pageId="3" pageNumber="386">3.36</td>
<td id="C74CC321376000666A9DFEA901B99E1B" box="[938,982,312,337]" gridcol="4" gridrow="3" pageId="3" pageNumber="386">2.93</td>
<td id="C74CC321376000666D5EFEA906FA9E1B" box="[1129,1173,312,337]" gridcol="5" gridrow="3" pageId="3" pageNumber="386">1.13</td>
<td id="C74CC321376000666C16FEA907359E1B" box="[1313,1370,312,337]" gridcol="6" gridrow="3" pageId="3" pageNumber="386">13.63</td>
</tr>
<tr id="849DAA5D376000666994FEF007359E33" box="[163,1370,353,377]" gridrow="4" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FEF002BF9E33" box="[163,208,353,377]" gridcol="0" gridrow="4" pageId="3" pageNumber="386">III</th>
<td id="C74CC321376000666858FEF003F49E33" box="[367,411,353,377]" gridcol="1" gridrow="4" pageId="3" pageNumber="386">2.29</td>
<td id="C74CC321376000666B1AFEF000369E33" box="[557,601,353,377]" gridcol="2" gridrow="4" pageId="3" pageNumber="386">1.67</td>
<td id="C74CC321376000666BDBFEF001779E33" box="[748,792,353,377]" gridcol="3" gridrow="4" pageId="3" pageNumber="386">2.61</td>
<td id="C74CC321376000666A9DFEF001B99E33" box="[938,982,353,377]" gridcol="4" gridrow="4" pageId="3" pageNumber="386">2.79</td>
<td id="C74CC321376000666D5EFEF006FA9E33" box="[1129,1173,353,377]" gridcol="5" gridrow="4" pageId="3" pageNumber="386">1.22</td>
<td id="C74CC321376000666C16FEF007359E33" box="[1313,1370,353,377]" gridcol="6" gridrow="4" pageId="3" pageNumber="386">10.58</td>
</tr>
<tr id="849DAA5D376000666994FE1807359EEB" box="[163,1370,393,417]" gridrow="5" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FE1802BF9EEB" box="[163,208,393,417]" gridcol="0" gridrow="5" pageId="3" pageNumber="386">IV</th>
<td id="C74CC321376000666858FE1803F49EEB" box="[367,411,393,417]" gridcol="1" gridrow="5" pageId="3" pageNumber="386">4.54</td>
<td id="C74CC321376000666B1AFE1800369EEB" box="[557,601,393,417]" gridcol="2" gridrow="5" pageId="3" pageNumber="386">1.83</td>
<td id="C74CC321376000666BDBFE1801779EEB" box="[748,792,393,417]" gridcol="3" gridrow="5" pageId="3" pageNumber="386">4.00</td>
<td id="C74CC321376000666A9DFE1801B99EEB" box="[938,982,393,417]" gridcol="4" gridrow="5" pageId="3" pageNumber="386">4.83</td>
<td id="C74CC321376000666D5EFE1806FA9EEB" box="[1129,1173,393,417]" gridcol="5" gridrow="5" pageId="3" pageNumber="386">1.37</td>
<td id="C74CC321376000666C16FE1807359EEB" box="[1313,1370,393,417]" gridcol="6" gridrow="5" pageId="3" pageNumber="386">16.57</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973760FF9A69A0FE67019E9D44" box="[151,1009,502,527]" pageId="3" pageNumber="386" startId="3.[151,239,502,526]" targetBox="[163,1370,192,417]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="3A12A81F3760FF9A69A0FE67019E9D44" blockId="3.[151,1009,502,527]" box="[151,1009,502,527]" pageId="3" pageNumber="386">
<emphasis id="08D9740D3760FF9A69A0FE6703659D44" bold="true" box="[151,266,502,526]" pageId="3" pageNumber="386">TABLE 2.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C3760FF9A6BF0FE66019E9D44" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[711,1009,502,527]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3760FF9A6BF0FE66011E9D44" box="[711,881,502,526]" italics="true" pageId="3" pageNumber="386">Bowie neukoeln</emphasis>
<bibRefCitation id="5E3CD5EE3760FF9A6A4FFE66019E9D44" author="Jager, P." box="[888,1009,502,527]" pageId="3" pageNumber="386" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F3760FF9A68B8FD8E07399C4A" pageId="3" pageNumber="386">
<table id="48AD5ABF376000666994FD8E07399C4A" box="[163,1366,543,768]" gridcols="5" gridrows="6" pageId="3" pageNumber="386">
<tr id="849DAA5D376000666994FD8E07399D7D" box="[163,1366,543,567]" gridrow="0" pageId="3" pageNumber="386" rowspan-0="1">
<th id="C74CC321376000666854FD8E03BA9D7D" box="[355,469,543,567]" gridcol="1" gridrow="0" pageId="3" pageNumber="386">Fe</th>
<th id="C74CC321376000666BA3FD8E00809D7D" box="[660,751,543,567]" gridcol="2" gridrow="0" pageId="3" pageNumber="386">Pa</th>
<th id="C74CC321376000666A4DFD8E063C9D7D" box="[890,1107,543,567]" gridcol="3" gridrow="0" pageId="3" pageNumber="386">Ti</th>
<th id="C74CC321376000666DF5FD8E07399D7D" box="[1218,1366,543,567]" gridcol="4" gridrow="0" pageId="3" pageNumber="386">Mt</th>
</tr>
<tr id="849DAA5D376000666994FDD607399D15" box="[163,1366,583,607]" gridrow="1" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FDD602BF9D15" box="[163,208,583,607]" gridcol="0" gridrow="1" pageId="3" pageNumber="386">Palp</th>
<td id="C74CC321376000666854FDD603BA9D15" box="[355,469,583,607]" gridcol="1" gridrow="1" pageId="3" pageNumber="386">3d 1p 1r</td>
<td id="C74CC321376000666BA3FDD600809D15" box="[660,751,583,607]" gridcol="2" gridrow="1" pageId="3" pageNumber="386">1p</td>
<td id="C74CC321376000666A4DFDD6063C9D15" box="[890,1107,583,607]" gridcol="3" gridrow="1" pageId="3" pageNumber="386">2p 2r</td>
<td id="C74CC321376000666DF5FDD607399D15" box="[1218,1366,583,607]" gridcol="4" gridrow="1" pageId="3" pageNumber="386">-</td>
</tr>
<tr id="849DAA5D376000666994FDFE07399DCD" box="[163,1366,623,647]" gridrow="2" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FDFE02BF9DCD" box="[163,208,623,647]" gridcol="0" gridrow="2" pageId="3" pageNumber="386">I</th>
<td id="C74CC321376000666854FDFE03BA9DCD" box="[355,469,623,647]" gridcol="1" gridrow="2" pageId="3" pageNumber="386">3d 3p 3(2)r</td>
<td id="C74CC321376000666BA3FDFE00809DCD" box="[660,751,623,647]" gridcol="2" gridrow="2" pageId="3" pageNumber="386">spineless</td>
<td id="C74CC321376000666A4DFDFE063C9DCD" box="[890,1107,623,647]" gridcol="3" gridrow="2" pageId="3" pageNumber="386">2-2-2-2-2v</td>
<td id="C74CC321376000666DF5FDFE07399DCD" box="[1218,1366,623,647]" gridcol="4" gridrow="2" pageId="3" pageNumber="386">2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD0607399DE5" box="[163,1366,663,687]" gridrow="3" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD0602BF9DE5" box="[163,208,663,687]" gridcol="0" gridrow="3" pageId="3" pageNumber="386">II</th>
<td id="C74CC321376000666854FD0603BA9DE5" box="[355,469,663,687]" gridcol="1" gridrow="3" pageId="3" pageNumber="386">3d 4(2) 2r</td>
<td id="C74CC321376000666BA3FD0600809DE5" box="[660,751,663,687]" gridcol="2" gridrow="3" pageId="3" pageNumber="386">spineless</td>
<td id="C74CC321376000666A4DFD06063C9DE5" box="[890,1107,663,687]" gridcol="3" gridrow="3" pageId="3" pageNumber="386">1r 2-2-2-2-2v</td>
<td id="C74CC321376000666DF5FD0607399DE5" box="[1218,1366,663,687]" gridcol="4" gridrow="3" pageId="3" pageNumber="386">2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD2E07399D92" box="[163,1366,703,728]" gridrow="4" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD2E02BF9D92" box="[163,208,703,728]" gridcol="0" gridrow="4" pageId="3" pageNumber="386">III</th>
<td id="C74CC321376000666854FD2E03BA9D92" box="[355,469,703,728]" gridcol="1" gridrow="4" pageId="3" pageNumber="386">3d 4p 3(4)r</td>
<td id="C74CC321376000666BA3FD2E00809D92" box="[660,751,703,728]" gridcol="2" gridrow="4" pageId="3" pageNumber="386">1p 1r</td>
<td id="C74CC321376000666A4DFD2E063C9D92" box="[890,1107,703,728]" gridcol="3" gridrow="4" pageId="3" pageNumber="386">2d 2p 2r 2-2-2v</td>
<td id="C74CC321376000666DF5FD2E07399D92" box="[1218,1366,703,728]" gridcol="4" gridrow="4" pageId="3" pageNumber="386">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376000666994FD7907399C4A" box="[163,1366,744,768]" gridrow="5" pageId="3" pageNumber="386">
<th id="C74CC321376000666994FD7902BF9C4A" box="[163,208,744,768]" gridcol="0" gridrow="5" pageId="3" pageNumber="386">IV</th>
<td id="C74CC321376000666854FD7903BA9C4A" box="[355,469,744,768]" gridcol="1" gridrow="5" pageId="3" pageNumber="386">3d 3(4)p 2r</td>
<td id="C74CC321376000666BA3FD7900809C4A" box="[660,751,744,768]" gridcol="2" gridrow="5" pageId="3" pageNumber="386">1p 1r</td>
<td id="C74CC321376000666A4DFD79063C9C4A" box="[890,1107,744,768]" gridcol="3" gridrow="5" pageId="3" pageNumber="386">1(3)d 2p 2r 2(1)-2-2v</td>
<td id="C74CC321376000666DF5FD7907399C4A" box="[1218,1366,744,768]" gridcol="4" gridrow="5" pageId="3" pageNumber="386">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973760FF9A69A0F81E061B98A5" ID-DOI="http://doi.org/10.5281/zenodo.14386393" ID-Zenodo-Dep="14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="3" pageNumber="386" startId="3.[151,264,1935,1959]" targetBox="[151,1435,829,1910]" targetPageId="3" targetType="figure">
<paragraph id="3A12A81F3760FF9A69A0F81E061B98A5" blockId="3.[151,1436,1935,2031]" pageId="3" pageNumber="386">
<emphasis id="08D9740D3760FF9A69A0F81E032F98ED" bold="true" box="[151,320,1935,1959]" pageId="3" pageNumber="386">FIGURES 5–8.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3760FF9A68F9F80100D398ED" authority="Jager." authorityName="Jager." box="[462,700,1935,1959]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3760FF9A68F9F801001B98ED" box="[462,628,1935,1959]" italics="true" pageId="3" pageNumber="386">Bowie neukoeln</emphasis>
Jäger.
</taxonomicName>
5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3760FF9A6A5FF82501E59881" box="[872,906,1972,1995]" italics="true" pageId="3" pageNumber="386">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3760FF9A6D60F825061C9881" box="[1111,1139,1972,1995]" italics="true" pageId="3" pageNumber="386">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3760FF9A6C26F82207229880" box="[1297,1357,1971,1994]" italics="true" pageId="3" pageNumber="386">MC—</emphasis>
anterior constriction of median plate,
<emphasis id="08D9740D3760FF9A6888F849038B98A5" box="[447,484,2008,2031]" italics="true" pageId="3" pageNumber="386">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3760FF9A6BC9F846017898A4" box="[766,791,2007,2030]" italics="true" pageId="3" pageNumber="386">Sp</emphasis>
—spermatheca,
<emphasis id="08D9740D3760FF9A6A8BF84901B498A5" box="[956,987,2008,2031]" italics="true" pageId="3" pageNumber="386">VF</emphasis>
—vulval folds.
</paragraph>
</caption>
<paragraph id="3A12A81F3767FF9D69F0FF0607709E57" blockId="4.[151,1436,150,357]" pageId="4" pageNumber="387">
Female genitalia as shown in
<figureCitation id="A296B49A3767FF9D6B2CFF0600EE9FFB" box="[539,641,150,177]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Figs 5–8</figureCitation>
. Epigyne 1.4 times wider than long. Median plate (
<emphasis id="08D9740D3767FF9D6DFBFF06069B9FFA" box="[1228,1268,151,176]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
) with rounded lateral sides, without anterior keel. Lateral teeth (
<emphasis id="08D9740D3767FF9D6B88FF2A00B19F9E" box="[703,734,187,212]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
) well developed, large, located posterior of median plate, with blunt tips (in
<figureCitation id="A296B49A3767FF9D6801FF4E03F59FB3" box="[310,410,223,249]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Figs 6–7</figureCitation>
shape and position of
<emphasis id="08D9740D3767FF9D6B91FF4E00AA9FB2" box="[678,709,223,248]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
changed due to maceration). Spermathecae (
<emphasis id="08D9740D3767FF9D6DF8FF4E06849FB3" box="[1231,1259,223,249]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
) bottle-shaped, separated by ca. half spermatheca’s length (position of
<emphasis id="08D9740D3767FF9D6A37FE9201739E57" box="[768,796,259,285]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
could be slightly affected due to maceration).
</paragraph>
<paragraph id="3A12A81F3767FF9D69F0FEB607309E0B" blockId="4.[151,1436,150,357]" box="[199,1375,295,321]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FEB6037E9E0B" bold="true" box="[199,273,295,321]" pageId="4" pageNumber="387">Notes.</emphasis>
<taxonomicName id="FDADD39C3767FF9D682FFEB603A59E0B" box="[280,458,295,321]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3767FF9D682FFEB603A59E0B" box="[280,458,295,321]" italics="true" pageId="4" pageNumber="387">Bowie neukoeln</emphasis>
</taxonomicName>
was previously known from Malay Peninsula only, this record is first for Borneo.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69F0FEDB067C9E2F" box="[199,1043,330,357]" pageId="4" pageNumber="387" type="distribution">
<paragraph id="3A12A81F3767FF9D69F0FEDB067C9E2F" blockId="4.[151,1436,150,357]" box="[199,1043,330,357]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FEDB030E9E2E" bold="true" box="[199,353,330,356]" pageId="4" pageNumber="387">Distribution.</emphasis>
<collectingCountry id="42BAE88F3767FF9D6850FEDA03A19E2F" box="[359,462,331,357]" name="Malaysia" pageId="4" pageNumber="387">Malaysia</collectingCountry>
(Malay Peninsula and Borneo:
<collectingRegion id="F86966FD3767FF9D6A06FEDA01169E2F" box="[817,889,331,357]" country="Malaysia" name="Sabah" pageId="4" pageNumber="387">Sabah</collectingRegion>
;
<figureCitation id="A296B49A3767FF9D6AB2FEDA06689E2F" box="[901,1031,331,357]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="4" pageNumber="387">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="46239131328DFBDD8C70C67413F53029">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
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<mods:affiliation id="BABB8A1FB42D85D5A7E11AFC5DCD9B5C">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
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<paragraph id="3A12A81F3766FF9C69A0F8D503C09814" blockId="5.[151,431,1860,1922]" box="[151,431,1860,1887]" pageId="5" pageNumber="388">
<heading id="615A1F733766FF9C69A0F8D503C09814" bold="true" box="[151,431,1860,1887]" fontSize="11" level="1" pageId="5" pageNumber="388" reason="1">
<emphasis id="08D9740D3766FF9C69A0F8D503C09814" bold="true" box="[151,431,1860,1887]" pageId="5" pageNumber="388">
<taxonomicName id="FDADD39C3766FF9C69A0F8D503269815" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,329,1860,1887]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3766FF9C69A0F8D503269815" bold="true" box="[151,329,1860,1887]" italics="true" pageId="5" pageNumber="388">Bowie musang</emphasis>
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<taxonomicNameLabel id="13EAC9763766FF9C6866F8D503C09814" box="[337,431,1860,1886]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="72B7FB943766FF9C69A0F8F9031498C8" box="[151,379,1896,1922]" pageId="5" pageNumber="388" type="description">
<paragraph id="3A12A81F3766FF9C69A0F8F9031498C8" blockId="5.[151,431,1860,1922]" box="[151,379,1896,1922]" pageId="5" pageNumber="388">
(
<figureCitation id="A296B49A3766FF9C69A8F8F9034F98C8" box="[159,288,1896,1922]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–26" captionStart-2="FIGURES 27–30" captionStartId-0="6.[151,264,1068,1092]" captionStartId-1="7.[151,264,1764,1788]" captionStartId-2="8.[151,264,1382,1406]" captionTargetBox-0="[179,1408,241,1039]" captionTargetBox-1="[160,1430,198,1735]" captionTargetBox-2="[151,1435,181,1357]" captionTargetId-0="figure-51@6.[177,1411,237,1044]" captionTargetId-1="figure-19@7.[151,1435,181,1741]" captionTargetId-2="figure-30@8.[151,1435,181,1357]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 13–16. Habitus of male (13–14) and female (15–16) of Bowie musang sp. nov. 13, 15—dorsal; 14, 16—ventral. Scale bars: 2 mm." captionText-1="FIGURES 17–26. Palp (17–20), bulb (21–24), embolus (25) and palpal tibia (26) of male of Bowie musang sp. nov. 17, 21—prolateral; 18, 22, 25—ventral; 19, 23, 26—retrolateral; 20—dorsal, 24—apical. Scale bars: 17–20 = 0.5 mm; 21–24, 26 = 0.2 mm; 25 = 0.1 mm. Abbreviations: AA—apical arm of embolus, BA—basal arm of embolus, C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." captionText-2="FIGURES 27–30. Epigyne of Bowie musang sp. nov. 27—intact, ventral; 28—macerated, ventral; 29—internal genitalia, dorsal; 30—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.14386399" figureDoi-1="http://doi.org/10.5281/zenodo.14386403" figureDoi-2="http://doi.org/10.5281/zenodo.14386405" httpUri-0="https://zenodo.org/record/14386399/files/figure.png" httpUri-1="https://zenodo.org/record/14386403/files/figure.png" httpUri-2="https://zenodo.org/record/14386405/files/figure.png" pageId="5" pageNumber="388">Figs 13–30</figureCitation>
,
<figureCitation id="A296B49A3766FF9C681BF8F9031C98C8" box="[300,371,1896,1922]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="5" pageNumber="388">68–69</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="3A12A81F3766FF9F69A0F821032F9F9F" blockId="5.[151,1437,1968,2031]" lastBlockId="6.[151,1436,150,213]" lastPageId="6" lastPageNumber="389" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0F821032A9880" bold="true" box="[151,325,1968,1994]" pageId="5" pageNumber="388">Type material.</emphasis>
<materialsCitation id="8AC5A2423766FF9C687CF82101A198A4" collectingDate="1995-02-04" collectingDateMax="1995-02-09" collectingDateMin="1995-02-04" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Philippines" elevation="1430" latitude="8.183333" location="Kitanglad Range." longLatPrecision="1302" longitude="124.85" municipality="Mindanao Island" pageId="5" pageNumber="388" specimenCount="1" specimenCount-male="1" stateProvince="Bukidnon Province" typeStatus="holotype">
<emphasis id="08D9740D3766FF9C687CF82101039880" bold="true" box="[331,876,1968,1995]" pageId="5" pageNumber="388">
<collectingCountry id="42BAE88F3766FF9C687CF821006D9880" box="[331,514,1968,1994]" name="Philippines" pageId="5" pageNumber="388">PHILIPPINES</collectingCountry>
:
<emphasis id="08D9740D3766FF9C6B39F82100809880" bold="true" box="[526,751,1968,1995]" italics="true" pageId="5" pageNumber="388">
<collectingRegion id="F86966FD3766FF9C6B39F82100809880" box="[526,751,1968,1995]" country="Philippines" name="Bukidnon" pageId="5" pageNumber="388">Bukidnon Province</collectingRegion>
</emphasis>
:
<typeStatus id="E51616BD3766FF9C6BC8F82101039880" box="[767,876,1968,1994]" pageId="5" pageNumber="388" type="holotype">Holotype</typeStatus>
</emphasis>
:
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(
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),
<collectingMunicipality id="DA7632653766FF9C6D39F82106A29880" box="[1038,1229,1968,1994]" pageId="5" pageNumber="388">Mindanao Island</collectingMunicipality>
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<collectorName id="9758CDC93766FF9C6BE0F84501F398A4" box="[727,924,2004,2030]" pageId="5" pageNumber="388">A.M. Emelyanov</collectorName>
leg.
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<collectingRegion id="F86966FD3766FF9C6AE0F845062098A4" box="[983,1103,2004,2030]" country="Philippines" name="Paratypes" pageId="5" pageNumber="388">Paratypes</collectingRegion>
</emphasis>
:
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(
<collectionCode id="5CBC30DA3766FF9C6DF9F845074498A4" box="[1230,1323,2004,2030]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="388" type="Museum">ZMMU</collectionCode>
), together with the holotype
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.
<materialsCitation id="8AC5A2423765FF9F6852FF0703539F9F" collectingDate="1995-01-26" collectingDateMax="1995-01-30" collectingDateMin="1995-01-26" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Philippines" elevation="1350" latitude="7.016667" location="Apo Mt." longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" pageId="6" pageNumber="389" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Cotabato Province" typeStatus="paratype">
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<emphasis id="08D9740D3765FF9F6852FF0700559FFB" bold="true" box="[357,570,150,177]" italics="true" pageId="6" pageNumber="389">
<collectingRegion id="F86966FD3765FF9F6852FF0700559FFB" box="[357,570,150,177]" country="Philippines" name="South Cotabato" pageId="6" pageNumber="389">Cotabato Province</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="2CAB63963765FF9F6B7EFF0700039FFB" box="[585,620,150,177]" count="5" pageId="6" pageNumber="389" type="male">5♂</specimenCount>
<specimenCount id="2CAB63963765FF9F6B45FF0600FB9FFB" box="[626,660,151,177]" count="6" pageId="6" pageNumber="389" type="female">6♀</specimenCount>
(
<collectionCode id="5CBC30DA3765FF9F6B95FF0600929FFB" box="[674,765,151,177]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="6" pageNumber="389" type="Museum">ZMMU</collectionCode>
)
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, [
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],
<quantity id="FD5505FA3765FF9F6D62FF0706C59FFB" box="[1109,1194,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="389" unit="m" value="1350.0">
<elevation id="B1804F2C3765FF9F6D62FF0706C59FFB" box="[1109,1194,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="389" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<date id="4E138EDF3765FF9F6D82FF0607F79FFB" box="[1205,1432,150,177]" pageId="6" pageNumber="389" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">
<collectingDate id="5E5777373765FF9F6D82FF0607F79FFB" box="[1205,1432,150,177]" pageId="6" pageNumber="389" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">26–30 January 1995</collectingDate>
</date>
, same collector
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6ED2F8973765FF9F69A0FBBD03219B22" ID-DOI="http://doi.org/10.5281/zenodo.14386399" ID-Zenodo-Dep="14386399" httpUri="https://zenodo.org/record/14386399/files/figure.png" pageId="6" pageNumber="389" startId="6.[151,264,1068,1092]" targetBox="[179,1408,241,1039]" targetPageId="6" targetType="figure">
<paragraph id="3A12A81F3765FF9F69A0FBBD03219B22" blockId="6.[151,1437,1068,1128]" pageId="6" pageNumber="389">
<emphasis id="08D9740D3765FF9F69A0FBBD03399B0E" bold="true" box="[151,342,1068,1092]" pageId="6" pageNumber="389">FIGURES 13–16.</emphasis>
Habitus of male (13–14) and female (15–16) of
<taxonomicName id="FDADD39C3765FF9F6A63FBBC01859B0E" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[852,1002,1069,1092]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3765FF9F6A63FBBC01859B0E" box="[852,1002,1069,1092]" italics="true" pageId="6" pageNumber="389">Bowie musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D3765FF9F6AC5FBBD062A9B0E" bold="true" box="[1010,1093,1068,1092]" pageId="6" pageNumber="389">
<taxonomicNameLabel id="13EAC9763765FF9F6AC5FBBD062A9B0E" box="[1010,1093,1068,1092]" pageId="6" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
13, 15—dorsal; 14, 16—ventral. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="72B7FB943765FF9F69F0FB2A019E9BB2" pageId="6" pageNumber="389" type="etymology">
<paragraph id="3A12A81F3765FF9F69F0FB2A019E9BB2" blockId="6.[151,1437,1210,1993]" pageId="6" pageNumber="389">
<emphasis id="08D9740D3765FF9F69F0FB2A03239B9F" bold="true" box="[199,332,1211,1237]" pageId="6" pageNumber="389">Etymology.</emphasis>
The specific name is derived from the local name of the Asian palm civet [
<taxonomicName id="FDADD39C3765FF9F6D6CFB2B02809BB2" authority="(Pallas)" baseAuthorityName="Pallas" baseAuthorityYear="1777" class="Mammalia" family="Viverridae" genus="Paradoxurus" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="389" phylum="Chordata" rank="species" species="hermaphroditus">
<emphasis id="08D9740D3765FF9F6D6CFB2B07F39B9E" box="[1115,1436,1210,1236]" italics="true" pageId="6" pageNumber="389">Paradoxurus hermaphroditus</emphasis>
(Pallas)
</taxonomicName>
], a small predatory beast native to Southeast Asia; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943765FF9F69F0FA9307029977" pageId="6" pageNumber="389" type="diagnosis">
<paragraph id="3A12A81F3765FF9F69F0FA9307029977" blockId="6.[151,1437,1210,1993]" pageId="6" pageNumber="389">
<emphasis id="08D9740D3765FF9F69F0FA9303509A56" bold="true" box="[199,319,1282,1308]" pageId="6" pageNumber="389">Diagnosis.</emphasis>
Males of
<taxonomicName id="FDADD39C3765FF9F688FFA9200319A56" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[440,606,1283,1308]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3765FF9F688FFA9200319A56" box="[440,606,1283,1308]" italics="true" pageId="6" pageNumber="389">Bowie musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D3765FF9F6B5FFA9200AC9A57" bold="true" box="[616,707,1283,1309]" pageId="6" pageNumber="389">
<taxonomicNameLabel id="13EAC9763765FF9F6B5FFA9200AC9A57" box="[616,707,1283,1309]" pageId="6" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="FDADD39C3765FF9F6A96FA9206C09A57" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[929,1199,1282,1309]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="bluejean">
<emphasis id="08D9740D3765FF9F6A96FA92064E9A56" box="[929,1057,1282,1308]" italics="true" pageId="6" pageNumber="389">B. bluejean</emphasis>
<bibRefCitation id="5E3CD5EE3765FF9F6D1BFA9206C09A57" author="Jager, P." box="[1068,1199,1283,1309]" pageId="6" pageNumber="389" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
, by the shape of the retro-proximal cymbial outgrowth (
<emphasis id="08D9740D3765FF9F6B13FAB600359A0B" box="[548,602,1319,1345]" italics="true" pageId="6" pageNumber="389">RPO</emphasis>
) in ventral view. They are also similar to
<taxonomicName id="FDADD39C3765FF9F6D0FFAB603319A2F" authority="(F. O. Pickard-Cambridge, 1897)" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1897" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="08D9740D3765FF9F6D0FFAB606829A0A" box="[1080,1261,1318,1344]" italics="true" pageId="6" pageNumber="389">B. sarawakensis</emphasis>
(F. O.
<bibRefCitation id="5E3CD5EE3765FF9F6C08FAB703399A2F" author="Pickard-Cambridge, F. O." pageId="6" pageNumber="389" pagination="329 - 356" refId="ref10347" refString="Pickard-Cambridge, F. O. (1897) On the cteniform spiders of Ceylon, Burmah and the Indian archipelago, west and north of Wallace's line; with bibliography and list of those from Australia, south and east of Wallace's line. Annals and Magazine of Natural History (6), 20 (118), 329 - 356, pl. 4. https: // doi. org / 10.1080 / 00222939708680638" type="journal article" year="1897">Pickard-Cambridge, 1897</bibRefCitation>
)
</taxonomicName>
in having a large, diagonally oriented tegular apophysis (
<emphasis id="08D9740D3765FF9F6AFEFADA01889A2F" box="[969,999,1355,1381]" italics="true" pageId="6" pageNumber="389">TA</emphasis>
). Males of the new species can be easily distinguished from both species by the tip of the embolus (
<emphasis id="08D9740D3765FF9F6A07FAFE013A9AC2" box="[816,853,1391,1416]" italics="true" pageId="6" pageNumber="389">Em</emphasis>
) divided into long apical (
<emphasis id="08D9740D3765FF9F6DB4FAFE06CA9AC3" box="[1155,1189,1391,1417]" italics="true" pageId="6" pageNumber="389">AA</emphasis>
) and short basal (
<emphasis id="08D9740D3765FF9F6C46FAFE07FC9AC3" box="[1393,1427,1391,1417]" italics="true" pageId="6" pageNumber="389">BA</emphasis>
) arms (vs. single tip; cf.
<figureCitation id="A296B49A3765FF9F68AFFA03007D9AE7" box="[408,530,1426,1453]" captionStart="FIGURES 17–26" captionStartId="7.[151,264,1764,1788]" captionTargetBox="[160,1430,198,1735]" captionTargetId="figure-19@7.[151,1435,181,1741]" captionTargetPageId="7" captionText="FIGURES 17–26. Palp (17–20), bulb (21–24), embolus (25) and palpal tibia (26) of male of Bowie musang sp. nov. 17, 21—prolateral; 18, 22, 25—ventral; 19, 23, 26—retrolateral; 20—dorsal, 24—apical. Scale bars: 17–20 = 0.5 mm; 21–24, 26 = 0.2 mm; 25 = 0.1 mm. Abbreviations: AA—apical arm of embolus, BA—basal arm of embolus, C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386403" httpUri="https://zenodo.org/record/14386403/files/figure.png" pageId="6" pageNumber="389">Figs 22, 25</figureCitation>
and
<bibRefCitation id="5E3CD5EE3765FF9F6B7FFA0200AB9AE7" author="Jager, P." box="[584,708,1427,1453]" pageId="6" pageNumber="389" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: figs 754, 766). By the shape of the median plate (
<emphasis id="08D9740D3765FF9F6DDCFA02077C9AE6" box="[1259,1299,1427,1452]" italics="true" pageId="6" pageNumber="389">MP</emphasis>
), females of
<taxonomicName id="FDADD39C3765FF9F69A0FA2603579A9A" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,312,1463,1488]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3765FF9F69A0FA2603579A9A" box="[151,312,1463,1488]" italics="true" pageId="6" pageNumber="389">Bowie musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D3765FF9F6809FA2603FA9A9B" bold="true" box="[318,405,1463,1489]" pageId="6" pageNumber="389">
<taxonomicNameLabel id="13EAC9763765FF9F6809FA2603FA9A9B" box="[318,405,1463,1489]" pageId="6" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="FDADD39C3765FF9F6B57FA2601F29A9B" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[608,925,1462,1489]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="criminalworld">
<emphasis id="08D9740D3765FF9F6B57FA2601749A9A" box="[608,795,1462,1488]" italics="true" pageId="6" pageNumber="389">B. criminalworld</emphasis>
<bibRefCitation id="5E3CD5EE3765FF9F6A16FA2601F29A9B" author="Jager, P." box="[801,925,1463,1489]" pageId="6" pageNumber="389" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
by the shape of epigyne median plate, but may be distinguished by 1) significantly shorter carapace and body length (6.11 and
<quantity id="FD5505FA3765FF9F6D34FA4A061D9ABF" box="[1027,1138,1499,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.197" pageId="6" pageNumber="389" unit="mm" value="11.97">11.97 mm</quantity>
vs. 8 and
<quantity id="FD5505FA3765FF9F6DD4FA4B07269ABF" box="[1251,1353,1498,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="6" pageNumber="389" unit="mm" value="16.9">16.9 mm</quantity>
), 2) the median plate’s width/length ratio 1.6 (vs. 1.2; cf.
<figureCitation id="A296B49A3765FF9F6B86FA6F00919953" box="[689,766,1534,1561]" captionStart="FIGURES 27–30" captionStartId="8.[151,264,1382,1406]" captionTargetBox="[151,1435,181,1357]" captionTargetId="figure-30@8.[151,1435,181,1357]" captionTargetPageId="8" captionText="FIGURES 27–30. Epigyne of Bowie musang sp. nov. 27—intact, ventral; 28—macerated, ventral; 29—internal genitalia, dorsal; 30—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386405" httpUri="https://zenodo.org/record/14386405/files/figure.png" pageId="6" pageNumber="389">Fig. 27</figureCitation>
and
<bibRefCitation id="5E3CD5EE3765FF9F6A03FA6E01C09953" author="Jager, P." box="[820,943,1535,1561]" pageId="6" pageNumber="389" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 772), 3) spermathecae (
<emphasis id="08D9740D3765FF9F6DD0FA6E076D9953" box="[1255,1282,1535,1561]" italics="true" pageId="6" pageNumber="389">Sp</emphasis>
) separated by a distance 2.7 times less than a the length of a spermatheca (vs. 4 times less; cf.
<figureCitation id="A296B49A3765FF9F6D37F9B306239977" box="[1024,1100,1570,1597]" captionStart="FIGURES 27–30" captionStartId="8.[151,264,1382,1406]" captionTargetBox="[151,1435,181,1357]" captionTargetId="figure-30@8.[151,1435,181,1357]" captionTargetPageId="8" captionText="FIGURES 27–30. Epigyne of Bowie musang sp. nov. 27—intact, ventral; 28—macerated, ventral; 29—internal genitalia, dorsal; 30—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386405" httpUri="https://zenodo.org/record/14386405/files/figure.png" pageId="6" pageNumber="389">Fig. 29</figureCitation>
and
<bibRefCitation id="5E3CD5EE3765FF9F6DB5F9B206929977" author="Jager, P." box="[1154,1277,1571,1597]" pageId="6" pageNumber="389" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 774).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943765FF9369F0F9D703DB9E9B" lastPageId="10" lastPageNumber="393" pageId="6" pageNumber="389" type="description">
<paragraph id="3A12A81F3765FF9F69F0F9D703EF98CA" blockId="6.[151,1437,1210,1993]" pageId="6" pageNumber="389">
<emphasis id="08D9740D3765FF9F69F0F9D70336992A" bold="true" box="[199,345,1606,1632]" pageId="6" pageNumber="389">Description.</emphasis>
<emphasis id="08D9740D3765FF9F6855F9D703F3992A" box="[354,412,1606,1632]" italics="true" pageId="6" pageNumber="389">Male</emphasis>
(
<figureCitation id="A296B49A3765FF9F689BF9D7005F992B" box="[428,560,1606,1633]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1068,1092]" captionTargetBox="[179,1408,241,1039]" captionTargetId="figure-51@6.[177,1411,237,1044]" captionTargetPageId="6" captionText="FIGURES 13–16. Habitus of male (13–14) and female (15–16) of Bowie musang sp. nov. 13, 15—dorsal; 14, 16—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386399" httpUri="https://zenodo.org/record/14386399/files/figure.png" pageId="6" pageNumber="389">Figs 13–14</figureCitation>
). Total length 10.70. Prosoma 6.06 long, 4.59 wide. Opisthosoma 4.82 long, 2.99 wide. Prosoma dark brown, with light brown median band. Median band near ocular area widened, with projections and covered with short white setae, bearing couple of gray spots. Lateral bands narrow, light brown. Fovea thin, black. Clypeus light brown. Chelicerae brown proximally, dark brown distally, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum brown, somewhat lighter in the middle. Labium brown. Endites brown with yellowish distal edge. Dorsal part of opisthosoma black with distinct yellowish cardiac mark and series of transverse stripes posterior of it. Lateral sides of opisthosoma gray with yellowish spots. Ventral part of opisthosoma black with yellow spots forming two V-marks (inner mark poorly visible). Spinnerets yellow ventrally, dark brown laterally.
</paragraph>
<paragraph id="3A12A81F3765FF9F69F0F81A07EF9883" blockId="6.[151,1437,1210,1993]" pageId="6" pageNumber="389">Eye diameters: AME 0.25, ALE 0.15, PME 0.39, PLE 0.32; interdistances: AME–AME 0.11, AME–ALE 0.35, PME–PME 0.22, PME–PLE 0.35, AME–PME 0.06, ALE–PLE 0.19. Clypeus height at AME 0.19, at ALE 0.44.</paragraph>
<caption id="6ED2F8973764FF9E69A0F975038098C7" ID-DOI="http://doi.org/10.5281/zenodo.14386403" ID-Zenodo-Dep="14386403" httpUri="https://zenodo.org/record/14386403/files/figure.png" pageId="7" pageNumber="390" startId="7.[151,264,1764,1788]" targetBox="[160,1430,198,1735]" targetPageId="7" targetType="figure">
<paragraph id="3A12A81F3764FF9E69A0F975038098C7" blockId="7.[151,1437,1764,1933]" pageId="7" pageNumber="390">
<emphasis id="08D9740D3764FF9E69A0F975033799B7" bold="true" box="[151,344,1764,1789]" pageId="7" pageNumber="390">FIGURES 17–26.</emphasis>
Palp (17–20), bulb (21–24), embolus (25) and palpal tibia (26) of male of
<taxonomicName id="FDADD39C3764FF9E6D4AF974077A99B6" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1149,1301,1765,1788]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="390" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3764FF9E6D4AF974077A99B6" box="[1149,1301,1765,1788]" italics="true" pageId="7" pageNumber="390">Bowie musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D3764FF9E6C29F974071D99B7" bold="true" box="[1310,1394,1765,1789]" pageId="7" pageNumber="390">
<taxonomicNameLabel id="13EAC9763764FF9E6C29F974071D99B7" box="[1310,1394,1765,1789]" pageId="7" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
17, 21—prolateral; 18, 22, 25—ventral; 19, 23, 26—retrolateral; 20—dorsal, 24—apical. Scale bars: 17–20 = 0.5 mm; 21–24, 26 = 0.2 mm; 25 = 0.1 mm. Abbreviations:
<emphasis id="08D9740D3764FF9E6B07F8BC0020980E" box="[560,591,1837,1860]" italics="true" pageId="7" pageNumber="390">AA</emphasis>
—apical arm of embolus,
<emphasis id="08D9740D3764FF9E6A61F8BC011A980E" box="[854,885,1837,1860]" italics="true" pageId="7" pageNumber="390">BA</emphasis>
—basal arm of embolus,
<emphasis id="08D9740D3764FF9E6D43F8BC06EA980E" box="[1140,1157,1837,1860]" italics="true" pageId="7" pageNumber="390">C</emphasis>
—conductor,
<emphasis id="08D9740D3764FF9E6C27F8BC075D980E" box="[1296,1330,1837,1860]" italics="true" pageId="7" pageNumber="390">EH</emphasis>
—embolic hook,
<emphasis id="08D9740D3764FF9E69EDF8C002939822" box="[218,252,1873,1896]" italics="true" pageId="7" pageNumber="390">Em</emphasis>
—embolus,
<emphasis id="08D9740D3764FF9E684BF8C003C29822" box="[380,429,1873,1896]" italics="true" pageId="7" pageNumber="390">RPO</emphasis>
—retro-proximal cymbial outgrowth,
<emphasis id="08D9740D3764FF9E6A0EF8C0010B9822" box="[825,868,1873,1896]" italics="true" pageId="7" pageNumber="390">RTA</emphasis>
—retrolateral tibial apophysis,
<emphasis id="08D9740D3764FF9E6D9CF8C006A89822" box="[1195,1223,1873,1896]" italics="true" pageId="7" pageNumber="390">TA</emphasis>
—tegular apophysis,
<emphasis id="08D9740D3764FF9E69A0F8E402C198C6" box="[151,174,1909,1932]" italics="true" pageId="7" pageNumber="390">Te</emphasis>
—tegulum,
<emphasis id="08D9740D3764FF9E6810F8E4032898C6" box="[295,327,1909,1932]" italics="true" pageId="7" pageNumber="390">TH</emphasis>
—tegular hump.
</paragraph>
</caption>
<caption id="6ED2F897376BFF9169A0FAF703C29A8C" ID-DOI="http://doi.org/10.5281/zenodo.14386405" ID-Zenodo-Dep="14386405" httpUri="https://zenodo.org/record/14386405/files/figure.png" pageId="8" pageNumber="391" startId="8.[151,264,1382,1406]" targetBox="[151,1435,181,1357]" targetPageId="8" targetType="figure">
<paragraph id="3A12A81F376BFF9169A0FAF703C29A8C" blockId="8.[151,1436,1382,1478]" pageId="8" pageNumber="391">
<emphasis id="08D9740D376BFF9169A0FAF703379A34" bold="true" box="[151,344,1382,1406]" pageId="8" pageNumber="391">FIGURES 27–30.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C376BFF9168E8FAF600189A34" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[479,631,1383,1406]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376BFF9168E8FAF600189A34" box="[479,631,1383,1406]" italics="true" pageId="8" pageNumber="391">Bowie musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D376BFF916BB6FAF700BA9A34" bold="true" box="[641,725,1382,1406]" pageId="8" pageNumber="391">
<taxonomicNameLabel id="13EAC976376BFF916BB6FAF700BA9A34" box="[641,725,1382,1406]" pageId="8" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
27—intact, ventral; 28—macerated, ventral; 29—internal genitalia, dorsal; 30—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D376BFF916A21FA1A01579AE8" box="[790,824,1419,1442]" italics="true" pageId="8" pageNumber="391">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D376BFF916D31FA1A064D9AE8" box="[1030,1058,1419,1442]" italics="true" pageId="8" pageNumber="391">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D376BFF916DF6FA1A06899AE8" box="[1217,1254,1419,1442]" italics="true" pageId="8" pageNumber="391">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D376BFF9169C3FA3E03629A8C" box="[244,269,1455,1478]" italics="true" pageId="8" pageNumber="391">Sp</emphasis>
—spermatheca.
</paragraph>
</caption>
<paragraph id="3A12A81F376BFF9169F0F98806C499D4" blockId="8.[151,1437,1560,2019]" pageId="8" pageNumber="391">
For palp and legs measurements see
<tableCitation id="772F9DA4376BFF916B6CF98900C09979" box="[603,687,1560,1587]" captionStart="TABLE 5" captionStartId="9.[151,239,584,608]" captionTargetPageId="9" captionText="TABLE 5. Palp and legs measurements of male of Bowie musang sp. nov." pageId="8" pageNumber="391">Table 5</tableCitation>
. Palp coloration: femora and patellae light brown with blurry spots; tibiae and cymbium brown. Legs coloration: femora I–IV light brown with blurry semi-rings; patellae I–IV light brown; tibiae I–II light brown, III–IV light brown with blurry semi-rings; metatarsi I–II light brown, III–IV light brown with blurry semi-rings; tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA4376BFF916D65F91506C799D4" box="[1106,1192,1668,1694]" captionStart="TABLE 6" captionStartId="9.[151,239,935,959]" captionTargetPageId="9" captionText="TABLE 6. Palp and legs spination for the male of Bowie musang sp. nov." pageId="8" pageNumber="391">Table 6</tableCitation>
.
</paragraph>
<paragraph id="3A12A81F376BFF9169F0F93900379819" blockId="8.[151,1437,1560,2019]" pageId="8" pageNumber="391">
Palp as shown in
<figureCitation id="A296B49A376BFF9168B0F93900689988" box="[391,519,1704,1730]" captionStart="FIGURES 17–26" captionStartId="7.[151,264,1764,1788]" captionTargetBox="[160,1430,198,1735]" captionTargetId="figure-19@7.[151,1435,181,1741]" captionTargetPageId="7" captionText="FIGURES 17–26. Palp (17–20), bulb (21–24), embolus (25) and palpal tibia (26) of male of Bowie musang sp. nov. 17, 21—prolateral; 18, 22, 25—ventral; 19, 23, 26—retrolateral; 20—dorsal, 24—apical. Scale bars: 17–20 = 0.5 mm; 21–24, 26 = 0.2 mm; 25 = 0.1 mm. Abbreviations: AA—apical arm of embolus, BA—basal arm of embolus, C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386403" httpUri="https://zenodo.org/record/14386403/files/figure.png" pageId="8" pageNumber="391">Figs 17–26</figureCitation>
.
<emphasis id="08D9740D376BFF916B26F938002E9989" box="[529,577,1705,1731]" italics="true" pageId="8" pageNumber="391">RTA</emphasis>
arising from middle of tibia, curved, with pointed tip. Cymbium with pointed retro-proximal cymbial outgrowth (
<emphasis id="08D9740D376BFF916B02F95C000499AD" box="[565,619,1741,1767]" italics="true" pageId="8" pageNumber="391">RPO</emphasis>
). Tegulum (
<emphasis id="08D9740D376BFF916BC8F95C017799AC" box="[767,792,1741,1766]" italics="true" pageId="8" pageNumber="391">Te</emphasis>
) with rounded hump (
<emphasis id="08D9740D376BFF916D1FF95C062399AC" box="[1064,1100,1741,1766]" italics="true" pageId="8" pageNumber="391">TH</emphasis>
) in retrolateral half. Tegular apophysis (
<emphasis id="08D9740D376BFF91682EF96003579841" box="[281,312,1777,1803]" italics="true" pageId="8" pageNumber="391">TA</emphasis>
) large, elongated, originating in a 5 o’clock position. Conductor (
<emphasis id="08D9740D376BFF916D1BF96006509841" box="[1068,1087,1777,1803]" italics="true" pageId="8" pageNumber="391">C</emphasis>
) large, membranous. Embolus (
<emphasis id="08D9740D376BFF916996F88402A99864" box="[161,198,1813,1838]" italics="true" pageId="8" pageNumber="391">Em</emphasis>
) originating from tegulum in 7-o’clock-position, its tip divided in long apical (
<emphasis id="08D9740D376BFF916D77F884060D9865" box="[1088,1122,1813,1839]" italics="true" pageId="8" pageNumber="391">AA</emphasis>
) and short basal (
<emphasis id="08D9740D376BFF916C1BF88407219865" box="[1324,1358,1813,1839]" italics="true" pageId="8" pageNumber="391">BA</emphasis>
) arms. Embolus with large embolic hook (
<emphasis id="08D9740D376BFF916B15F8A800289818" box="[546,583,1849,1874]" italics="true" pageId="8" pageNumber="391">EH</emphasis>
).
</paragraph>
<paragraph id="3A12A81F376BFF9069F0F8CD06929E57" blockId="8.[151,1437,1560,2019]" lastBlockId="9.[151,1437,151,501]" lastPageId="9" lastPageNumber="392" pageId="8" pageNumber="391">
<emphasis id="08D9740D376BFF9169F0F8CD034C983C" box="[199,291,1884,1910]" italics="true" pageId="8" pageNumber="391">Female.</emphasis>
(
<figureCitation id="A296B49A376BFF916804F8CD03D8983D" box="[307,439,1884,1911]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1068,1092]" captionTargetBox="[179,1408,241,1039]" captionTargetId="figure-51@6.[177,1411,237,1044]" captionTargetPageId="6" captionText="FIGURES 13–16. Habitus of male (13–14) and female (15–16) of Bowie musang sp. nov. 13, 15—dorsal; 14, 16—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386399" httpUri="https://zenodo.org/record/14386399/files/figure.png" pageId="8" pageNumber="391">Figs 15–16</figureCitation>
). Total length 11.97. Prosoma 6.11 long, 4.72 wide. Opisthosoma 5.87 long, 4.01 wide. Prosoma dark brown (darker than in males), with brown median band. Median band near ocular area widened, bearing couple of gray spots. Lateral bands narrow, light brown, poorly visible. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum dark brown with short longitudinal strap anteriorly. Labium dark brown, almost black. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma black with poorly visible yellowish cardiac mark and almost invisible series of transverse stripes posterior of it. Lateral sides of opisthosoma gray with yellowish spots. Ventral part of opisthosoma dark gray with yellow spots forming two V-marks. Spinnerets yellow ventrally, dark brown laterally.
</paragraph>
<paragraph id="3A12A81F376AFF9069F0FEB607EF9E2F" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="392">Eye diameters: AME 0.23, ALE 0.18, PME 0.36, PLE 0.33; interdistances: AME–AME 0.20, AME–ALE 0.47, PME–PME 0.23, PME–PLE 0.49, AME–PME 0.10, ALE–PLE 0.25. Clypeus height at AME 0.17, at ALE 0.54.</paragraph>
<paragraph id="3A12A81F376AFF9069F0FEFE07539EBF" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="392">
For palp and legs measurements see
<tableCitation id="772F9DA4376AFF906B59FEFE00A99EC3" box="[622,710,367,393]" captionStart="TABLE 7" captionStartId="9.[151,239,1286,1310]" captionTargetPageId="9" captionText="TABLE 7. Palp and legs measurements of female of Bowie musang sp. nov." pageId="9" pageNumber="392">Table 7</tableCitation>
. Palp coloration. Femora light brown; other joints brown. Legs coloration. Femora I brown with light brown longitudinal stripes laterally; II–IV brown with dark brown blurry spots. Patellae I–IV brown. Tibiae I–II brown; III–IV brown with dark brown semi-rings. Metatarsi I–II brown; III–IV brown with dark brown semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA4376AFF906DD5FE4A07589EBF" box="[1250,1335,475,501]" captionStart="TABLE 8" captionStartId="9.[151,239,1637,1661]" captionTargetPageId="9" captionText="TABLE 8. Palp and legs spination for the female of Bowie musang sp. nov." pageId="9" pageNumber="392">Table 8</tableCitation>
.
</paragraph>
<caption id="6ED2F897376AFF9069A0FDD901DD9D2B" box="[151,946,584,609]" pageId="9" pageNumber="392" startId="9.[151,239,584,608]" targetBox="[163,1370,625,850]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="3A12A81F376AFF9069A0FDD901DD9D2B" blockId="9.[151,946,584,609]" box="[151,946,584,609]" pageId="9" pageNumber="392">
<emphasis id="08D9740D376AFF9069A0FDD903659D2B" bold="true" box="[151,266,584,609]" pageId="9" pageNumber="392">TABLE 5.</emphasis>
Palp and legs measurements of male of
<taxonomicName id="FDADD39C376AFF906B8EFDD801399D2A" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[697,854,585,608]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376AFF906B8EFDD801399D2A" box="[697,854,585,608]" italics="true" pageId="9" pageNumber="392">Bowie musang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC976376AFF906A6BFDD801DD9D2B" box="[860,946,585,609]" pageId="9" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F376AFF90684EFDE007359C18" pageId="9" pageNumber="392">
<table id="48AD5ABF376A00666994FDE007359C18" box="[163,1370,625,850]" gridcols="7" gridrows="6" pageId="9" pageNumber="392">
<tr id="849DAA5D376A00666994FDE007359DC3" box="[163,1370,625,649]" gridrow="0" pageId="9" pageNumber="392" rowspan-0="1">
<th id="C74CC321376A00666858FDE003F49DC3" box="[367,411,625,649]" gridcol="1" gridrow="0" pageId="9" pageNumber="392">Fe</th>
<th id="C74CC321376A00666B1AFDE000369DC3" box="[557,601,625,649]" gridcol="2" gridrow="0" pageId="9" pageNumber="392">Pa</th>
<th id="C74CC321376A00666BDBFDE001779DC3" box="[748,792,625,649]" gridcol="3" gridrow="0" pageId="9" pageNumber="392">Ti</th>
<th id="C74CC321376A00666A9DFDE001B99DC3" box="[938,982,625,649]" gridcol="4" gridrow="0" pageId="9" pageNumber="392">Mt</th>
<th id="C74CC321376A00666D5EFDE006FA9DC3" box="[1129,1173,625,649]" gridcol="5" gridrow="0" pageId="9" pageNumber="392">Ta</th>
<th id="C74CC321376A00666C16FDE007359DC3" box="[1313,1370,625,649]" gridcol="6" gridrow="0" pageId="9" pageNumber="392">Total</th>
</tr>
<tr id="849DAA5D376A00666994FD0807359DFB" box="[163,1370,665,689]" gridrow="1" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FD0802BF9DFB" box="[163,208,665,689]" gridcol="0" gridrow="1" pageId="9" pageNumber="392">Palp</th>
<td id="C74CC321376A00666858FD0803F49DFB" box="[367,411,665,689]" gridcol="1" gridrow="1" pageId="9" pageNumber="392">2.22</td>
<td id="C74CC321376A00666B1AFD0800369DFB" box="[557,601,665,689]" gridcol="2" gridrow="1" pageId="9" pageNumber="392">0.99</td>
<td id="C74CC321376A00666BDBFD0801779DFB" box="[748,792,665,689]" gridcol="3" gridrow="1" pageId="9" pageNumber="392">0.95</td>
<td id="C74CC321376A00666A9DFD0801B99DFB" box="[938,982,665,689]" gridcol="4" gridrow="1" pageId="9" pageNumber="392">-</td>
<td id="C74CC321376A00666D5EFD0806FA9DFB" box="[1129,1173,665,689]" gridcol="5" gridrow="1" pageId="9" pageNumber="392">2.02</td>
<td id="C74CC321376A00666C16FD0807359DFB" box="[1313,1370,665,689]" gridcol="6" gridrow="1" pageId="9" pageNumber="392">6.18</td>
</tr>
<tr id="849DAA5D376A00666994FD5007359D93" box="[163,1370,705,729]" gridrow="2" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FD5002BF9D93" box="[163,208,705,729]" gridcol="0" gridrow="2" pageId="9" pageNumber="392">I</th>
<td id="C74CC321376A00666858FD5003F49D93" box="[367,411,705,729]" gridcol="1" gridrow="2" pageId="9" pageNumber="392">4.43</td>
<td id="C74CC321376A00666B1AFD5000369D93" box="[557,601,705,729]" gridcol="2" gridrow="2" pageId="9" pageNumber="392">2.08</td>
<td id="C74CC321376A00666BDBFD5001779D93" box="[748,792,705,729]" gridcol="3" gridrow="2" pageId="9" pageNumber="392">4.28</td>
<td id="C74CC321376A00666A9DFD5001B99D93" box="[938,982,705,729]" gridcol="4" gridrow="2" pageId="9" pageNumber="392">3.95</td>
<td id="C74CC321376A00666D5EFD5006FA9D93" box="[1129,1173,705,729]" gridcol="5" gridrow="2" pageId="9" pageNumber="392">1.34</td>
<td id="C74CC321376A00666C16FD5007359D93" box="[1313,1370,705,729]" gridcol="6" gridrow="2" pageId="9" pageNumber="392">16.08</td>
</tr>
<tr id="849DAA5D376A00666994FD7807359C48" box="[163,1370,745,770]" gridrow="3" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FD7802BF9C48" box="[163,208,745,770]" gridcol="0" gridrow="3" pageId="9" pageNumber="392">II</th>
<td id="C74CC321376A00666858FD7803F49C48" box="[367,411,745,770]" gridcol="1" gridrow="3" pageId="9" pageNumber="392">4.10</td>
<td id="C74CC321376A00666B1AFD7800369C48" box="[557,601,745,770]" gridcol="2" gridrow="3" pageId="9" pageNumber="392">2.12</td>
<td id="C74CC321376A00666BDBFD7801779C48" box="[748,792,745,770]" gridcol="3" gridrow="3" pageId="9" pageNumber="392">3.66</td>
<td id="C74CC321376A00666A9DFD7801B99C48" box="[938,982,745,770]" gridcol="4" gridrow="3" pageId="9" pageNumber="392">3.43</td>
<td id="C74CC321376A00666D5EFD7806FA9C48" box="[1129,1173,745,770]" gridcol="5" gridrow="3" pageId="9" pageNumber="392">1.24</td>
<td id="C74CC321376A00666C16FD7807359C48" box="[1313,1370,745,770]" gridcol="6" gridrow="3" pageId="9" pageNumber="392">14.55</td>
</tr>
<tr id="849DAA5D376A00666994FC8007359C60" box="[163,1370,785,810]" gridrow="4" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FC8002BF9C60" box="[163,208,785,810]" gridcol="0" gridrow="4" pageId="9" pageNumber="392">III</th>
<td id="C74CC321376A00666858FC8003F49C60" box="[367,411,785,810]" gridcol="1" gridrow="4" pageId="9" pageNumber="392">3.59</td>
<td id="C74CC321376A00666B1AFC8000369C60" box="[557,601,785,810]" gridcol="2" gridrow="4" pageId="9" pageNumber="392">1.87</td>
<td id="C74CC321376A00666BDBFC8001779C60" box="[748,792,785,810]" gridcol="3" gridrow="4" pageId="9" pageNumber="392">2.72</td>
<td id="C74CC321376A00666A9DFC8001B99C60" box="[938,982,785,810]" gridcol="4" gridrow="4" pageId="9" pageNumber="392">3.30</td>
<td id="C74CC321376A00666D5EFC8006FA9C60" box="[1129,1173,785,810]" gridcol="5" gridrow="4" pageId="9" pageNumber="392">1.18</td>
<td id="C74CC321376A00666C16FC8007359C60" box="[1313,1370,785,810]" gridcol="6" gridrow="4" pageId="9" pageNumber="392">12.66</td>
</tr>
<tr id="849DAA5D376A00666994FCAB07359C18" box="[163,1370,826,850]" gridrow="5" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FCAB02BF9C18" box="[163,208,826,850]" gridcol="0" gridrow="5" pageId="9" pageNumber="392">IV</th>
<td id="C74CC321376A00666858FCAB03F49C18" box="[367,411,826,850]" gridcol="1" gridrow="5" pageId="9" pageNumber="392">4.84</td>
<td id="C74CC321376A00666B1AFCAB00369C18" box="[557,601,826,850]" gridcol="2" gridrow="5" pageId="9" pageNumber="392">1.92</td>
<td id="C74CC321376A00666BDBFCAB01779C18" box="[748,792,826,850]" gridcol="3" gridrow="5" pageId="9" pageNumber="392">4.19</td>
<td id="C74CC321376A00666A9DFCAB01B99C18" box="[938,982,826,850]" gridcol="4" gridrow="5" pageId="9" pageNumber="392">5.45</td>
<td id="C74CC321376A00666D5EFCAB06FA9C18" box="[1129,1173,826,850]" gridcol="5" gridrow="5" pageId="9" pageNumber="392">1.61</td>
<td id="C74CC321376A00666C16FCAB07359C18" box="[1313,1370,826,850]" gridcol="6" gridrow="5" pageId="9" pageNumber="392">18.01</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376AFF9069A0FC3601C49CF5" box="[151,939,935,959]" pageId="9" pageNumber="392" startId="9.[151,239,935,959]" targetBox="[163,1385,975,1201]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="3A12A81F376AFF9069A0FC3601C49CF5" blockId="9.[151,939,935,959]" box="[151,939,935,959]" pageId="9" pageNumber="392">
<emphasis id="08D9740D376AFF9069A0FC3603659CF5" bold="true" box="[151,266,935,959]" pageId="9" pageNumber="392">TABLE 6.</emphasis>
Palp and legs spination for the male of
<taxonomicName id="FDADD39C376AFF906B84FC3901209CF5" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[691,847,936,959]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376AFF906B84FC3901209CF5" box="[691,847,936,959]" italics="true" pageId="9" pageNumber="392">Bowie musang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC976376AFF906A61FC3601C49CF5" box="[854,939,935,959]" pageId="9" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F376AFF9068B8FC4107069BFB" pageId="9" pageNumber="392">
<table id="48AD5ABF376A00666994FC5E07069BFB" box="[163,1385,975,1201]" gridcols="5" gridrows="6" pageId="9" pageNumber="392">
<tr id="849DAA5D376A00666994FC5E07069CAD" box="[163,1385,975,999]" gridrow="0" pageId="9" pageNumber="392" rowspan-0="1">
<th id="C74CC321376A00666855FC5E03BA9CAD" box="[354,469,975,999]" gridcol="1" gridrow="0" pageId="9" pageNumber="392">Fe</th>
<th id="C74CC321376A00666B91FC5E00B59CAD" box="[678,730,975,999]" gridcol="2" gridrow="0" pageId="9" pageNumber="392">Pa</th>
<th id="C74CC321376A00666AB6FC5E06259CAD" box="[897,1098,975,999]" gridcol="3" gridrow="0" pageId="9" pageNumber="392">Ti</th>
<th id="C74CC321376A00666D9CFC5E07069CAD" box="[1195,1385,975,999]" gridcol="4" gridrow="0" pageId="9" pageNumber="392">Mt</th>
</tr>
<tr id="849DAA5D376A00666994FC6907069B5A" box="[163,1385,1016,1040]" gridrow="1" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FC6902BF9B5A" box="[163,208,1016,1040]" gridcol="0" gridrow="1" pageId="9" pageNumber="392">Palp</th>
<td id="C74CC321376A00666855FC6903BA9B5A" box="[354,469,1016,1040]" gridcol="1" gridrow="1" pageId="9" pageNumber="392">5d 1p 1r</td>
<td id="C74CC321376A00666B91FC6900B59B5A" box="[678,730,1016,1040]" gridcol="2" gridrow="1" pageId="9" pageNumber="392">1p</td>
<td id="C74CC321376A00666AB6FC6906259B5A" box="[897,1098,1016,1040]" gridcol="3" gridrow="1" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666D9CFC6907069B5A" box="[1195,1385,1016,1040]" gridcol="4" gridrow="1" pageId="9" pageNumber="392">-</td>
</tr>
<tr id="849DAA5D376A00666994FBB107069B72" box="[163,1385,1056,1080]" gridrow="2" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FBB102BF9B72" box="[163,208,1056,1080]" gridcol="0" gridrow="2" pageId="9" pageNumber="392">I</th>
<td id="C74CC321376A00666855FBB103BA9B72" box="[354,469,1056,1080]" gridcol="1" gridrow="2" pageId="9" pageNumber="392">3d 3p 4r</td>
<td id="C74CC321376A00666B91FBB100B59B72" box="[678,730,1056,1080]" gridcol="2" gridrow="2" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AB6FBB106259B72" box="[897,1098,1056,1080]" gridcol="3" gridrow="2" pageId="9" pageNumber="392">3d 2p 2r 2-2-2-2-2v</td>
<td id="C74CC321376A00666D9CFBB107069B72" box="[1195,1385,1056,1080]" gridcol="4" gridrow="2" pageId="9" pageNumber="392">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994FBD907069B2A" box="[163,1385,1096,1120]" gridrow="3" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FBD902BF9B2A" box="[163,208,1096,1120]" gridcol="0" gridrow="3" pageId="9" pageNumber="392">II</th>
<td id="C74CC321376A00666855FBD903BA9B2A" box="[354,469,1096,1120]" gridcol="1" gridrow="3" pageId="9" pageNumber="392">3d 3(4)p 4r</td>
<td id="C74CC321376A00666B91FBD900B59B2A" box="[678,730,1096,1120]" gridcol="2" gridrow="3" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AB6FBD906259B2A" box="[897,1098,1096,1120]" gridcol="3" gridrow="3" pageId="9" pageNumber="392">3d 2p 2r 2-2-2-2-2v</td>
<td id="C74CC321376A00666D9CFBD907069B2A" box="[1195,1385,1096,1120]" gridcol="4" gridrow="3" pageId="9" pageNumber="392">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994FBE107069BC3" box="[163,1385,1136,1161]" gridrow="4" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FBE102BF9BC3" box="[163,208,1136,1161]" gridcol="0" gridrow="4" pageId="9" pageNumber="392">III</th>
<td id="C74CC321376A00666855FBE103BA9BC3" box="[354,469,1136,1161]" gridcol="1" gridrow="4" pageId="9" pageNumber="392">3d 4p 4r</td>
<td id="C74CC321376A00666B91FBE100B59BC3" box="[678,730,1136,1161]" gridcol="2" gridrow="4" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AB6FBE106259BC3" box="[897,1098,1136,1161]" gridcol="3" gridrow="4" pageId="9" pageNumber="392">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376A00666D9CFBE107069BC3" box="[1195,1385,1136,1161]" gridcol="4" gridrow="4" pageId="9" pageNumber="392">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994FB0907069BFB" box="[163,1385,1176,1201]" gridrow="5" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FB0902BF9BFB" box="[163,208,1176,1201]" gridcol="0" gridrow="5" pageId="9" pageNumber="392">IV</th>
<td id="C74CC321376A00666855FB0903BA9BFB" box="[354,469,1176,1201]" gridcol="1" gridrow="5" pageId="9" pageNumber="392">3d 4p 3r</td>
<td id="C74CC321376A00666B91FB0900B59BFB" box="[678,730,1176,1201]" gridcol="2" gridrow="5" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AB6FB0906259BFB" box="[897,1098,1176,1201]" gridcol="3" gridrow="5" pageId="9" pageNumber="392">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376A00666D9CFB0907069BFB" box="[1195,1385,1176,1201]" gridcol="4" gridrow="5" pageId="9" pageNumber="392">4p 5r 1-1-1-1-1-2v</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376AFF9069A0FA9701A99A54" box="[151,966,1286,1310]" pageId="9" pageNumber="392" startId="9.[151,239,1286,1310]" targetBox="[163,1372,1326,1551]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="3A12A81F376AFF9069A0FA9701A99A54" blockId="9.[151,966,1286,1310]" box="[151,966,1286,1310]" pageId="9" pageNumber="392">
<emphasis id="08D9740D376AFF9069A0FA9703659A54" bold="true" box="[151,266,1286,1310]" pageId="9" pageNumber="392">TABLE 7.</emphasis>
Palp and legs measurements of female of
<taxonomicName id="FDADD39C376AFF906BF9FA9601059A54" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[718,874,1287,1310]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376AFF906BF9FA9601059A54" box="[718,874,1287,1310]" italics="true" pageId="9" pageNumber="392">Bowie musang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC976376AFF906A46FA9701A99A54" box="[881,966,1286,1310]" pageId="9" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F376AFF90684EFABE07339945" pageId="9" pageNumber="392">
<table id="48AD5ABF376A00666994FABF07339945" box="[163,1372,1326,1551]" gridcols="7" gridrows="6" pageId="9" pageNumber="392">
<tr id="849DAA5D376A00666994FABF07339A0C" box="[163,1372,1326,1350]" gridrow="0" pageId="9" pageNumber="392" rowspan-0="1">
<th id="C74CC321376A00666858FABF03F49A0C" box="[367,411,1326,1350]" gridcol="1" gridrow="0" pageId="9" pageNumber="392">Fe</th>
<th id="C74CC321376A00666B19FABF00359A0C" box="[558,602,1326,1350]" gridcol="2" gridrow="0" pageId="9" pageNumber="392">Pa</th>
<th id="C74CC321376A00666BDAFABF01769A0C" box="[749,793,1326,1350]" gridcol="3" gridrow="0" pageId="9" pageNumber="392">Ti</th>
<th id="C74CC321376A00666A9BFABF01B79A0C" box="[940,984,1326,1350]" gridcol="4" gridrow="0" pageId="9" pageNumber="392">Mt</th>
<th id="C74CC321376A00666D5DFABF06F99A0C" box="[1130,1174,1326,1350]" gridcol="5" gridrow="0" pageId="9" pageNumber="392">Ta</th>
<th id="C74CC321376A00666C14FABF07339A0C" box="[1315,1372,1326,1350]" gridcol="6" gridrow="0" pageId="9" pageNumber="392">Total</th>
</tr>
<tr id="849DAA5D376A00666994FAC707339A25" box="[163,1372,1366,1391]" gridrow="1" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FAC702BF9A25" box="[163,208,1366,1391]" gridcol="0" gridrow="1" pageId="9" pageNumber="392">Palp</th>
<td id="C74CC321376A00666858FAC703F49A25" box="[367,411,1366,1391]" gridcol="1" gridrow="1" pageId="9" pageNumber="392">2.00</td>
<td id="C74CC321376A00666B19FAC700359A25" box="[558,602,1366,1391]" gridcol="2" gridrow="1" pageId="9" pageNumber="392">0.89</td>
<td id="C74CC321376A00666BDAFAC701769A25" box="[749,793,1366,1391]" gridcol="3" gridrow="1" pageId="9" pageNumber="392">1.32</td>
<td id="C74CC321376A00666A9BFAC701B79A25" box="[940,984,1366,1391]" gridcol="4" gridrow="1" pageId="9" pageNumber="392">-</td>
<td id="C74CC321376A00666D5DFAC706F99A25" box="[1130,1174,1366,1391]" gridcol="5" gridrow="1" pageId="9" pageNumber="392">1.48</td>
<td id="C74CC321376A00666C14FAC707339A25" box="[1315,1372,1366,1391]" gridcol="6" gridrow="1" pageId="9" pageNumber="392">5.69</td>
</tr>
<tr id="849DAA5D376A00666994FAEE07339ADD" box="[163,1372,1407,1431]" gridrow="2" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FAEE02BF9ADD" box="[163,208,1407,1431]" gridcol="0" gridrow="2" pageId="9" pageNumber="392">I</th>
<td id="C74CC321376A00666858FAEE03F49ADD" box="[367,411,1407,1431]" gridcol="1" gridrow="2" pageId="9" pageNumber="392">3.94</td>
<td id="C74CC321376A00666B19FAEE00359ADD" box="[558,602,1407,1431]" gridcol="2" gridrow="2" pageId="9" pageNumber="392">2.20</td>
<td id="C74CC321376A00666BDAFAEE01769ADD" box="[749,793,1407,1431]" gridcol="3" gridrow="2" pageId="9" pageNumber="392">3.36</td>
<td id="C74CC321376A00666A9BFAEE01B79ADD" box="[940,984,1407,1431]" gridcol="4" gridrow="2" pageId="9" pageNumber="392">2.82</td>
<td id="C74CC321376A00666D5DFAEE06F99ADD" box="[1130,1174,1407,1431]" gridcol="5" gridrow="2" pageId="9" pageNumber="392">1.11</td>
<td id="C74CC321376A00666C14FAEE07339ADD" box="[1315,1372,1407,1431]" gridcol="6" gridrow="2" pageId="9" pageNumber="392">13.43</td>
</tr>
<tr id="849DAA5D376A00666994FA3607339AF5" box="[163,1372,1447,1471]" gridrow="3" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FA3602BF9AF5" box="[163,208,1447,1471]" gridcol="0" gridrow="3" pageId="9" pageNumber="392">II</th>
<td id="C74CC321376A00666858FA3603F49AF5" box="[367,411,1447,1471]" gridcol="1" gridrow="3" pageId="9" pageNumber="392">3.78</td>
<td id="C74CC321376A00666B19FA3600359AF5" box="[558,602,1447,1471]" gridcol="2" gridrow="3" pageId="9" pageNumber="392">2.07</td>
<td id="C74CC321376A00666BDAFA3601769AF5" box="[749,793,1447,1471]" gridcol="3" gridrow="3" pageId="9" pageNumber="392">2.98</td>
<td id="C74CC321376A00666A9BFA3601B79AF5" box="[940,984,1447,1471]" gridcol="4" gridrow="3" pageId="9" pageNumber="392">2.68</td>
<td id="C74CC321376A00666D5DFA3606F99AF5" box="[1130,1174,1447,1471]" gridcol="5" gridrow="3" pageId="9" pageNumber="392">1.02</td>
<td id="C74CC321376A00666C14FA3607339AF5" box="[1315,1372,1447,1471]" gridcol="6" gridrow="3" pageId="9" pageNumber="392">12.53</td>
</tr>
<tr id="849DAA5D376A00666994FA5E07339AAD" box="[163,1372,1487,1511]" gridrow="4" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FA5E02BF9AAD" box="[163,208,1487,1511]" gridcol="0" gridrow="4" pageId="9" pageNumber="392">III</th>
<td id="C74CC321376A00666858FA5E03F49AAD" box="[367,411,1487,1511]" gridcol="1" gridrow="4" pageId="9" pageNumber="392">3.13</td>
<td id="C74CC321376A00666B19FA5E00359AAD" box="[558,602,1487,1511]" gridcol="2" gridrow="4" pageId="9" pageNumber="392">1.73</td>
<td id="C74CC321376A00666BDAFA5E01769AAD" box="[749,793,1487,1511]" gridcol="3" gridrow="4" pageId="9" pageNumber="392">2.34</td>
<td id="C74CC321376A00666A9BFA5E01B79AAD" box="[940,984,1487,1511]" gridcol="4" gridrow="4" pageId="9" pageNumber="392">2.69</td>
<td id="C74CC321376A00666D5DFA5E06F99AAD" box="[1130,1174,1487,1511]" gridcol="5" gridrow="4" pageId="9" pageNumber="392">1.16</td>
<td id="C74CC321376A00666C14FA5E07339AAD" box="[1315,1372,1487,1511]" gridcol="6" gridrow="4" pageId="9" pageNumber="392">11.05</td>
</tr>
<tr id="849DAA5D376A00666994FA6607339945" box="[163,1372,1527,1551]" gridrow="5" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994FA6602BF9945" box="[163,208,1527,1551]" gridcol="0" gridrow="5" pageId="9" pageNumber="392">IV</th>
<td id="C74CC321376A00666858FA6603F49945" box="[367,411,1527,1551]" gridcol="1" gridrow="5" pageId="9" pageNumber="392">4.25</td>
<td id="C74CC321376A00666B19FA6600359945" box="[558,602,1527,1551]" gridcol="2" gridrow="5" pageId="9" pageNumber="392">1.86</td>
<td id="C74CC321376A00666BDAFA6601769945" box="[749,793,1527,1551]" gridcol="3" gridrow="5" pageId="9" pageNumber="392">3.50</td>
<td id="C74CC321376A00666A9BFA6601B79945" box="[940,984,1527,1551]" gridcol="4" gridrow="5" pageId="9" pageNumber="392">4.52</td>
<td id="C74CC321376A00666D5DFA6606F99945" box="[1130,1174,1527,1551]" gridcol="5" gridrow="5" pageId="9" pageNumber="392">1.51</td>
<td id="C74CC321376A00666C14FA6607339945" box="[1315,1372,1527,1551]" gridcol="6" gridrow="5" pageId="9" pageNumber="392">15.64</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376AFF9069A0F9F401AF9937" box="[151,960,1637,1661]" pageId="9" pageNumber="392" startId="9.[151,239,1637,1661]" targetBox="[163,1385,1677,1902]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="3A12A81F376AFF9069A0F9F401AF9937" blockId="9.[151,960,1637,1661]" box="[151,960,1637,1661]" pageId="9" pageNumber="392">
<emphasis id="08D9740D376AFF9069A0F9F403649937" bold="true" box="[151,267,1637,1661]" pageId="9" pageNumber="392">TABLE 8.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C376AFF906BF0F9F4010B9937" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[711,868,1637,1661]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376AFF906BF0F9F4010B9937" box="[711,868,1637,1661]" italics="true" pageId="9" pageNumber="392">Bowie musang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC976376AFF906A5DF9F401AF9937" box="[874,960,1637,1661]" pageId="9" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F376AFF9068B8F91F07069824" pageId="9" pageNumber="392">
<table id="48AD5ABF376A00666994F91C07069824" box="[163,1385,1677,1902]" gridcols="5" gridrows="6" pageId="9" pageNumber="392">
<tr id="849DAA5D376A00666994F91C070699EF" box="[163,1385,1677,1701]" gridrow="0" pageId="9" pageNumber="392" rowspan-0="1">
<th id="C74CC321376A00666846F91C03A999EF" box="[369,454,1677,1701]" gridcol="1" gridrow="0" pageId="9" pageNumber="392">Fe</th>
<th id="C74CC321376A00666BA5F91C008299EF" box="[658,749,1677,1701]" gridcol="2" gridrow="0" pageId="9" pageNumber="392">Pa</th>
<th id="C74CC321376A00666AA1F91C065A99EF" box="[918,1077,1677,1701]" gridcol="3" gridrow="0" pageId="9" pageNumber="392">Ti</th>
<th id="C74CC321376A00666D9CF91C070699EF" box="[1195,1385,1677,1701]" gridcol="4" gridrow="0" pageId="9" pageNumber="392">Mt</th>
</tr>
<tr id="849DAA5D376A00666994F92407069987" box="[163,1385,1717,1741]" gridrow="1" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994F92402BF9987" box="[163,208,1717,1741]" gridcol="0" gridrow="1" pageId="9" pageNumber="392">Palp</th>
<td id="C74CC321376A00666846F92403A99987" box="[369,454,1717,1741]" gridcol="1" gridrow="1" pageId="9" pageNumber="392">3d 1p 1r</td>
<td id="C74CC321376A00666BA5F92400829987" box="[658,749,1717,1741]" gridcol="2" gridrow="1" pageId="9" pageNumber="392">1p</td>
<td id="C74CC321376A00666AA1F924065A9987" box="[918,1077,1717,1741]" gridcol="3" gridrow="1" pageId="9" pageNumber="392">2p 1r</td>
<td id="C74CC321376A00666D9CF92407069987" box="[1195,1385,1717,1741]" gridcol="4" gridrow="1" pageId="9" pageNumber="392">-</td>
</tr>
<tr id="849DAA5D376A00666994F94C070699BC" box="[163,1385,1757,1782]" gridrow="2" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994F94C02BF99BC" box="[163,208,1757,1782]" gridcol="0" gridrow="2" pageId="9" pageNumber="392">I</th>
<td id="C74CC321376A00666846F94C03A999BC" box="[369,454,1757,1782]" gridcol="1" gridrow="2" pageId="9" pageNumber="392">3d 3p 3r</td>
<td id="C74CC321376A00666BA5F94C008299BC" box="[658,749,1757,1782]" gridcol="2" gridrow="2" pageId="9" pageNumber="392">spineless</td>
<td id="C74CC321376A00666AA1F94C065A99BC" box="[918,1077,1757,1782]" gridcol="3" gridrow="2" pageId="9" pageNumber="392">2-2-2-2-2v</td>
<td id="C74CC321376A00666D9CF94C070699BC" box="[1195,1385,1757,1782]" gridcol="4" gridrow="2" pageId="9" pageNumber="392">2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994F89707069854" box="[163,1385,1798,1822]" gridrow="3" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994F89702BF9854" box="[163,208,1798,1822]" gridcol="0" gridrow="3" pageId="9" pageNumber="392">II</th>
<td id="C74CC321376A00666846F89703A99854" box="[369,454,1798,1822]" gridcol="1" gridrow="3" pageId="9" pageNumber="392">3d 4p 3r</td>
<td id="C74CC321376A00666BA5F89700829854" box="[658,749,1798,1822]" gridcol="2" gridrow="3" pageId="9" pageNumber="392">spineless</td>
<td id="C74CC321376A00666AA1F897065A9854" box="[918,1077,1798,1822]" gridcol="3" gridrow="3" pageId="9" pageNumber="392">2-2-2-2-2v</td>
<td id="C74CC321376A00666D9CF89707069854" box="[1195,1385,1798,1822]" gridcol="4" gridrow="3" pageId="9" pageNumber="392">2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994F8BF0706980C" box="[163,1385,1838,1862]" gridrow="4" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994F8BF02BF980C" box="[163,208,1838,1862]" gridcol="0" gridrow="4" pageId="9" pageNumber="392">III</th>
<td id="C74CC321376A00666846F8BF03A9980C" box="[369,454,1838,1862]" gridcol="1" gridrow="4" pageId="9" pageNumber="392">3d 4p 4r</td>
<td id="C74CC321376A00666BA5F8BF0082980C" box="[658,749,1838,1862]" gridcol="2" gridrow="4" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AA1F8BF065A980C" box="[918,1077,1838,1862]" gridcol="3" gridrow="4" pageId="9" pageNumber="392">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376A00666D9CF8BF0706980C" box="[1195,1385,1838,1862]" gridcol="4" gridrow="4" pageId="9" pageNumber="392">4p 5r 2-2-2v</td>
</tr>
<tr id="849DAA5D376A00666994F8C707069824" box="[163,1385,1878,1902]" gridrow="5" pageId="9" pageNumber="392">
<th id="C74CC321376A00666994F8C702BF9824" box="[163,208,1878,1902]" gridcol="0" gridrow="5" pageId="9" pageNumber="392">IV</th>
<td id="C74CC321376A00666846F8C703A99824" box="[369,454,1878,1902]" gridcol="1" gridrow="5" pageId="9" pageNumber="392">3d 4p 2r</td>
<td id="C74CC321376A00666BA5F8C700829824" box="[658,749,1878,1902]" gridcol="2" gridrow="5" pageId="9" pageNumber="392">1p 1r</td>
<td id="C74CC321376A00666AA1F8C7065A9824" box="[918,1077,1878,1902]" gridcol="3" gridrow="5" pageId="9" pageNumber="392">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376A00666D9CF8C707069824" box="[1195,1385,1878,1902]" gridcol="4" gridrow="5" pageId="9" pageNumber="392">4p 5r 1-1-1-1-1-2v</td>
</tr>
</table>
</paragraph>
<paragraph id="3A12A81F3769FF9369F0FF06032D9FB3" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="393">
Female genitalia as shown in
<figureCitation id="A296B49A3769FF936B31FF0600EB9FFB" box="[518,644,151,177]" captionStart="FIGURES 27–30" captionStartId="8.[151,264,1382,1406]" captionTargetBox="[151,1435,181,1357]" captionTargetId="figure-30@8.[151,1435,181,1357]" captionTargetPageId="8" captionText="FIGURES 27–30. Epigyne of Bowie musang sp. nov. 27—intact, ventral; 28—macerated, ventral; 29—internal genitalia, dorsal; 30—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386405" httpUri="https://zenodo.org/record/14386405/files/figure.png" pageId="10" pageNumber="393">Figs 27–30</figureCitation>
. Epigyne slightly wider than long. Median plate (
<emphasis id="08D9740D3769FF936DADFF0606AD9FFA" box="[1178,1218,151,176]" italics="true" pageId="10" pageNumber="393">MP</emphasis>
) widest posteriorly. Lateral teeth (
<emphasis id="08D9740D3769FF936804FF2A033D9F9E" box="[307,338,187,212]" italics="true" pageId="10" pageNumber="393">LT</emphasis>
) large, located posterior of median plate, with rounded tips. Spermathecae (
<emphasis id="08D9740D3769FF936DA8FF2A06D49F9F" box="[1183,1211,187,213]" italics="true" pageId="10" pageNumber="393">Sp</emphasis>
) small, more or less kidney-shaped.
</paragraph>
<paragraph id="3A12A81F3769FF9369F0FE9203DB9E9B" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369F0FE92037E9E57" bold="true" box="[199,273,259,285]" pageId="10" pageNumber="393">Notes.</emphasis>
According to the structure of the male copulatory organs, such as the hook-like
<emphasis id="08D9740D3769FF936DA5FE9206AD9E57" box="[1170,1218,259,285]" italics="true" pageId="10" pageNumber="393">RTA</emphasis>
arising medially as well as diagonally oriented tegular apophysis covering part of the embolus,
<taxonomicName id="FDADD39C3769FF936AE0FEB606529E0A" box="[983,1085,295,320]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="musan">
<emphasis id="08D9740D3769FF936AE0FEB606529E0A" box="[983,1085,295,320]" italics="true" pageId="10" pageNumber="393">B. musan</emphasis>
</taxonomicName>
g
<emphasis id="08D9740D3769FF936D66FEB606E79E0B" bold="true" box="[1105,1160,295,321]" pageId="10" pageNumber="393">sp. n</emphasis>
ov. belongs to the
<taxonomicName id="FDADD39C3769FF936C7AFEB607F39E0B" box="[1357,1436,295,321]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D3769FF936C7AFEB607F39E0B" box="[1357,1436,295,321]" italics="true" pageId="10" pageNumber="393">shakeit</emphasis>
</taxonomicName>
species-group sensu
<bibRefCitation id="5E3CD5EE3769FF9368B6FEDA007D9E2F" author="Jager, P." box="[385,530,331,357]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger (2022)</bibRefCitation>
. Thus, with the description of the new species, this group now consists of seven species:
<taxonomicName id="FDADD39C3769FF9369C0FEFE000D9EC3" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[247,610,367,393]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="ashestoashes">
<emphasis id="08D9740D3769FF9369C0FEFE03B89EC3" box="[247,471,367,393]" italics="true" pageId="10" pageNumber="393">Bowie ashestoashes</emphasis>
<bibRefCitation id="5E3CD5EE3769FF9368E9FEFE000D9EC3" author="Jager, P." box="[478,610,367,393]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDADD39C3769FF936B5AFEFE00829EC3" authorityName="Jager" authorityYear="2022" box="[621,749,367,393]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="bluejean">
<emphasis id="08D9740D3769FF936B5AFEFE00829EC3" box="[621,749,367,393]" italics="true" pageId="10" pageNumber="393">B. bluejean</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3769FF936BCBFEFE01D49EC3" authorityName="Jager" authorityYear="2022" box="[764,955,367,393]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="criminalworld">
<emphasis id="08D9740D3769FF936BCBFEFE01D49EC3" box="[764,955,367,393]" italics="true" pageId="10" pageNumber="393">B. criminalworld</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3769FF936AFDFEFE062E9EC2" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[970,1089,367,392]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D3769FF936AFDFEFE062E9EC2" box="[970,1089,367,392]" italics="true" pageId="10" pageNumber="393">B. musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D3769FF936D7FFEFE06CD9EC3" bold="true" box="[1096,1186,367,393]" pageId="10" pageNumber="393">
<taxonomicNameLabel id="13EAC9763769FF936D7FFEFE06CD9EC3" box="[1096,1186,367,393]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="FDADD39C3769FF936D86FEFE07FA9EC3" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1897" box="[1201,1429,367,393]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="08D9740D3769FF936D86FEFE07FA9EC3" box="[1201,1429,367,393]" italics="true" pageId="10" pageNumber="393">Bowie sarawakensis</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3769FF9369A0FE0203AB9EE7" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[151,452,403,429]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D3769FF9369A0FE02035A9EE7" box="[151,309,403,429]" italics="true" pageId="10" pageNumber="393">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE3769FF936808FE0203AB9EE7" author="Jager, P." box="[319,452,403,429]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="FDADD39C3769FF936B34FE0200BD9EE7" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[515,722,403,429]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D3769FF936B34FE0200BD9EE7" box="[515,722,403,429]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
</taxonomicName>
<emphasis id="08D9740D3769FF936BEDFE0201599EE7" bold="true" box="[730,822,403,429]" pageId="10" pageNumber="393">
<taxonomicNameLabel id="13EAC9763769FF936BEDFE0201599EE7" box="[730,822,403,429]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
All previous records of species from this group were restricted to Borneo only.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943769FF9369F0FE4B01429EBF" box="[199,813,474,501]" pageId="10" pageNumber="393" type="distribution">
<paragraph id="3A12A81F3769FF9369F0FE4B01429EBF" blockId="10.[151,1437,151,501]" box="[199,813,474,501]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369F0FE4B030E9EBE" bold="true" box="[199,353,474,500]" pageId="10" pageNumber="393">Distribution.</emphasis>
Mindanao Isl.,
<collectingCountry id="42BAE88F3769FF936B26FE4A00E09EBF" box="[529,655,475,501]" name="Philippines" pageId="10" pageNumber="393">Philippines</collectingCountry>
(
<figureCitation id="A296B49A3769FF936BA9FE4A014D9EBF" box="[670,802,475,501]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="10" pageNumber="393">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<mods:affiliation id="8CF6FDA500057C4898E46C93B5DCBF87">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
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<paragraph id="3A12A81F3767FF9D69A0FE3D03AA9E8C" blockId="4.[151,453,428,490]" box="[151,453,428,454]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0FE3D03AA9E8C" bold="true" box="[151,453,428,454]" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D69A0FE3D030F9E8C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,352,428,454]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D69A0FE3D030F9E8C" bold="true" box="[151,352,428,454]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
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<paragraph id="3A12A81F3767FF9D69A0FE4103CA9EA0" blockId="4.[151,453,428,490]" box="[151,421,464,490]" pageId="4" pageNumber="387">
(
<figureCitation id="A296B49A3767FF9D69A8FE41036B9EA0" box="[159,260,464,490]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6827FE4103259EA0" box="[272,330,464,490]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">9–12</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6861FE4103F29EA0" box="[342,413,464,490]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="4" pageNumber="387">68–69</figureCitation>
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<emphasis id="08D9740D3767FF9D69A0FD8903299D78" bold="true" box="[151,326,536,562]" pageId="4" pageNumber="387">Type material.</emphasis>
<materialsCitation id="8AC5A2423767FF9D687AFD8900049D1C" collectingDate="1994-08-30" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1020" latitude="4.4666667" location="Lucia Mt." longLatPrecision="1306" longitude="117.933334" municipality="Borneo" pageId="4" pageNumber="387" specimenCount="1" specimenCount-female="1" stateProvince="Sabah State" typeStatus="holotype">
<emphasis id="08D9740D3767FF9D687AFD8900939D78" bold="true" box="[333,764,535,562]" pageId="4" pageNumber="387">
<collectingCountry id="42BAE88F3767FF9D687AFD8903889D78" box="[333,487,536,562]" name="Malaysia" pageId="4" pageNumber="387">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D3767FF9D68C3FD8600119D78" bold="true" box="[500,638,535,562]" italics="true" pageId="4" pageNumber="387">
<collectingRegion id="F86966FD3767FF9D68C3FD8600119D78" box="[500,638,535,562]" country="Malaysia" name="Sabah" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D68C3FD8600529D78" authorityName="Li &amp; Yao" authorityYear="2022" box="[500,573,535,562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
<typeStatus id="E51616BD3767FF9D6BB8FD8900939D78" box="[655,764,536,562]" pageId="4" pageNumber="387" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="2CAB63963767FF9D6A3CFD88014F9D79" box="[779,800,537,563]" count="1" pageId="4" pageNumber="387" type="female">♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A18FD8901E39D78" box="[815,908,536,562]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="387" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653767FF9D6A97FD88019A9D79" box="[928,1013,537,563]" pageId="4" pageNumber="387">Borneo</collectingMunicipality>
,
<location id="3F72FEC43767FF9D6D36FD89061F9D79" LSID="urn:lsid:plazi:treatment:B20419093767FF9C69A0FE3D01EC99E0:3F72FEC43767FF9D6D36FD89061F9D79" box="[1025,1136,536,563]" country="Malaysia" latitude="4.4666667" longLatPrecision="1306" longitude="117.933334" municipality="Borneo" name="Lucia Mt." pageId="4" pageNumber="387" stateProvince="Sabah State">Lucia Mt.</location>
[
<geoCoordinate id="5F99CED83767FF9D6D48FD8906BA9D78" box="[1151,1237,536,562]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="387" precision="925" value="4.4666667">4°28’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83767FF9D6DD5FD89073E9D79" box="[1250,1361,536,563]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="4" pageNumber="387" precision="925" value="117.933334">117°56’E</geoCoordinate>
],
<quantity id="FD5505FA3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">
<elevation id="B1804F2C3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<date id="4E138EDF3767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">
<collectingDate id="5E5777373767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">30 August 1994</collectingDate>
</date>
,
<collectorName id="9758CDC93767FF9D684EFDAD00549D1C" box="[377,571,572,598]" pageId="4" pageNumber="387">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8AC5A2423767FF9D6B45FDAC06F89D1C" box="[626,1175,572,599]" collectingDate="1994-08-30" collectionCode="IBSS" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1020" latitude="4.4666667" location="Lucia Mt." longLatPrecision="1306" longitude="117.933334" municipality="Borneo" pageId="4" pageNumber="387" specimenCount="2" specimenCount-female="2" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="E51616BD3767FF9D6B45FDAC008C9D1D" box="[626,739,573,599]" pageId="4" pageNumber="387" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CAB63963767FF9D6BDAFDAD017F9D1C" box="[749,784,572,598]" count="2" pageId="4" pageNumber="387" type="female">2♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A28FDAD01329D1C" box="[799,861,572,598]" collectionName="IBSS" pageId="4" pageNumber="387">IBSS</collectionCode>
) together with the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69F0FDF100059DD4" pageId="4" pageNumber="387" type="etymology">
<paragraph id="3A12A81F3767FF9D69F0FDF100059DD4" blockId="4.[151,1436,535,671]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FDF103219D30" bold="true" box="[199,334,608,634]" pageId="4" pageNumber="387">Etymology.</emphasis>
The specific name is derived from the generic name of small wild cat from Southeast Asia,
<emphasis id="08D9740D3767FF9D69A0FD1503639DD4" box="[151,268,644,670]" italics="true" pageId="4" pageNumber="387">Catopuma</emphasis>
Severtzov; noun in apposition.
</paragraph>
</subSubSection>
<caption id="6ED2F8973767FF9D69A0F9310172984A" ID-DOI="http://doi.org/10.5281/zenodo.14386395" ID-Zenodo-Dep="14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387" startId="4.[151,264,1696,1720]" targetBox="[151,1435,694,1672]" targetPageId="4" targetType="figure">
<paragraph id="3A12A81F3767FF9D69A0F9310172984A" blockId="4.[151,1436,1696,1792]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0F931032999F2" bold="true" box="[151,326,1696,1720]" pageId="4" pageNumber="387">FIGURES 9–12.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3767FF9D688AF930000A99F2" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[445,613,1697,1720]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D688AF930000A99F2" box="[445,613,1697,1720]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6B5DF93100D699F2" bold="true" box="[618,697,1696,1720]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6B5DF93100D699F2" box="[618,697,1696,1720]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3767FF9D6BFBF95400819996" box="[716,750,1733,1756]" italics="true" pageId="4" pageNumber="387">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3767FF9D6A8BF95401B79996" box="[956,984,1733,1756]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3767FF9D6D40F95506DC9991" box="[1143,1203,1732,1755]" italics="true" pageId="4" pageNumber="387">MC—</emphasis>
anterior constriction of median plate,
<emphasis id="08D9740D3767FF9D6811F9780324984A" box="[294,331,1769,1792]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3767FF9D6B52F979001199B5" box="[613,638,1768,1791]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
—spermatheca.
</paragraph>
</caption>
<subSubSection id="72B7FB943767FF9D69F0F8D6073198E3" pageId="4" pageNumber="387" type="diagnosis">
<paragraph id="3A12A81F3767FF9D69F0F8D6073198E3" blockId="4.[151,1437,1863,2034]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F8D6032D982B" bold="true" box="[199,322,1863,1889]" pageId="4" pageNumber="387">Diagnosis.</emphasis>
By shape of the median plate (
<emphasis id="08D9740D3767FF9D6BA1F8D900D1982B" box="[662,702,1864,1889]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
) widest anteriorly females of
<taxonomicName id="FDADD39C3767FF9D6D3EF8D906AD982B" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1033,1218,1864,1889]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D6D3EF8D906AD982B" box="[1033,1218,1864,1889]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6DFEF8D9074E9828" bold="true" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6DFEF8D9074E9828" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="FDADD39C3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" italics="true" pageId="4" pageNumber="387">B. neukoeln</emphasis>
</taxonomicName>
, but may be easily distinguished by 1)
<emphasis id="08D9740D3767FF9D6A74F8FD010498CF" box="[835,875,1900,1925]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
with long anterior constriction (
<emphasis id="08D9740D3767FF9D6DE6F8FD069498CC" box="[1233,1275,1900,1926]" italics="true" pageId="4" pageNumber="387">MC</emphasis>
) (vs. short; cf.
<figureCitation id="A296B49A3767FF9D69A0F80102B698E0" box="[151,217,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 9</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6838F801033D98E0" box="[271,338,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 5</figureCitation>
); 2) vulval folds (
<emphasis id="08D9740D3767FF9D6B2BF801005198E3" box="[540,574,1936,1961]" italics="true" pageId="4" pageNumber="387">VF</emphasis>
) almost touching each other (vs. widely spaced; cf.
<figureCitation id="A296B49A3767FF9D6DB1F80106B998E0" box="[1158,1238,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 10</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6C3BF801072098E3" box="[1292,1359,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9C69F0F822073999CC" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387" type="description">
<paragraph id="3A12A81F3767FF9C69F0F82206FB9E0B" blockId="4.[151,1437,1863,2034]" lastBlockId="5.[151,1437,151,537]" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F82203369887" bold="true" box="[199,345,1971,1997]" pageId="4" pageNumber="387">Description.</emphasis>
<emphasis id="08D9740D3767FF9D6856F82503D29884" box="[353,445,1972,1998]" italics="true" pageId="4" pageNumber="387">Female.</emphasis>
(
<figureCitation id="A296B49A3767FF9D68FAF825005A9884" box="[461,565,1972,1998]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
). Total length 11.76. Prosoma 5.79 long, 4.28 wide. Opisthosoma 5.37 long, 3.44 wide. Prosoma reddish brown, with brown median band. Median band widened and with projections near ocular area. Lateral bands absent. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum light brown without pattern. Labium dark brown. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma light brown with yellowish cardiac mark and series of poorly visible gray spots. Lateral sides of opisthosoma light brown with blurry spots. Ventral part of opisthosoma light brown with yellow spots forming two poorly visible V-marks. Spinnerets light brown.
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FEDA07109EC3" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">Eye diameters: AME 0.22, ALE 0.18, PME 0.31, PLE 0.32; interdistances: AME–AME 0.19, AME–ALE 0.38, PME–PME 0.26, PME–PLE 0.42, AME–PME 0.11, ALE–PLE 0.17. Clypeus height at AME 0.14, at ALE 0.51.</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FE0207629D53" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">
For palp and legs measurements see
<tableCitation id="772F9DA43766FF9C6B5AFE0200AC9EE7" box="[621,707,403,429]" captionStart="TABLE 3" captionStartId="5.[151,239,620,644]" captionTargetPageId="5" captionText="TABLE 3. Palp and legs measurements of female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 3</tableCitation>
. Palp coloration: femora, patellae, tibiae light brown; tarsi light brown with darker tip. Leg coloration: femora I–IV light brown with poorly visible spots. Patellae I–IV light brown. Tibiae I–II light brown; III–IV light brown with blurred brown semi-rings. Metatarsi I–II light brown; III–IV light brown with blurred brown semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA43766FF9C6D84FE6E07669D53" box="[1203,1289,511,537]" captionStart="TABLE 4" captionStartId="5.[151,239,971,995]" captionTargetPageId="5" captionText="TABLE 4. Palp and legs spination for the female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 4</tableCitation>
.
</paragraph>
<caption id="6ED2F8973766FF9C69A0FDFD01B29DCF" box="[151,989,620,645]" pageId="5" pageNumber="388" startId="5.[151,239,620,644]" targetBox="[163,1370,661,886]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FDFD01B29DCF" blockId="5.[151,989,620,645]" box="[151,989,620,645]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FDFD03659DCE" bold="true" box="[151,266,620,644]" pageId="5" pageNumber="388">TABLE 3.</emphasis>
Palp and legs measurements of female of
<taxonomicName id="FDADD39C3766FF9C6BF9FDFC01EF9DCE" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[718,896,621,644]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF9FDFC01EF9DCE" box="[718,896,621,644]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB0FDFC01B29DCF" box="[903,989,621,645]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C684EFD0407359C3C" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FD0407359C3C" box="[163,1370,661,886]" gridcols="7" gridrows="6" pageId="5" pageNumber="388">
<tr id="849DAA5D376600666994FD0407359DE7" box="[163,1370,661,685]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666858FD0403F49DE7" box="[367,411,661,685]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666B19FD0400359DE7" box="[558,602,661,685]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666BDBFD0401779DE7" box="[748,792,661,685]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666A9CFD0401B89DE7" box="[939,983,661,685]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
<th id="C74CC321376600666D5EFD0406FA9DE7" box="[1129,1173,661,685]" gridcol="5" gridrow="0" pageId="5" pageNumber="388">Ta</th>
<th id="C74CC321376600666C16FD0407359DE7" box="[1313,1370,661,685]" gridcol="6" gridrow="0" pageId="5" pageNumber="388">Total</th>
</tr>
<tr id="849DAA5D376600666994FD2C07359D9F" box="[163,1370,701,725]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD2C02BF9D9F" box="[163,208,701,725]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666858FD2C03F49D9F" box="[367,411,701,725]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">2.00</td>
<td id="C74CC321376600666B19FD2C00359D9F" box="[558,602,701,725]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1.06</td>
<td id="C74CC321376600666BDBFD2C01779D9F" box="[748,792,701,725]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1.19</td>
<td id="C74CC321376600666A9CFD2C01B89D9F" box="[939,983,701,725]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
<td id="C74CC321376600666D5EFD2C06FA9D9F" box="[1129,1173,701,725]" gridcol="5" gridrow="1" pageId="5" pageNumber="388">1.35</td>
<td id="C74CC321376600666C16FD2C07359D9F" box="[1313,1370,701,725]" gridcol="6" gridrow="1" pageId="5" pageNumber="388">5.6</td>
</tr>
<tr id="849DAA5D376600666994FD7407359DB7" box="[163,1370,741,765]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD7402BF9DB7" box="[163,208,741,765]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666858FD7403F49DB7" box="[367,411,741,765]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">4.08</td>
<td id="C74CC321376600666B19FD7400359DB7" box="[558,602,741,765]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">2.17</td>
<td id="C74CC321376600666BDBFD7401779DB7" box="[748,792,741,765]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">3.77</td>
<td id="C74CC321376600666A9CFD7401B89DB7" box="[939,983,741,765]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">3.10</td>
<td id="C74CC321376600666D5EFD7406FA9DB7" box="[1129,1173,741,765]" gridcol="5" gridrow="2" pageId="5" pageNumber="388">1.00</td>
<td id="C74CC321376600666C16FD7407359DB7" box="[1313,1370,741,765]" gridcol="6" gridrow="2" pageId="5" pageNumber="388">14.12</td>
</tr>
<tr id="849DAA5D376600666994FC9C07359C6C" box="[163,1370,781,806]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FC9C02BF9C6C" box="[163,208,781,806]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666858FC9C03F49C6C" box="[367,411,781,806]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3.94</td>
<td id="C74CC321376600666B19FC9C00359C6C" box="[558,602,781,806]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">2.14</td>
<td id="C74CC321376600666BDBFC9C01779C6C" box="[748,792,781,806]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">3.32</td>
<td id="C74CC321376600666A9CFC9C01B89C6C" box="[939,983,781,806]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2.92</td>
<td id="C74CC321376600666D5EFC9C06FA9C6C" box="[1129,1173,781,806]" gridcol="5" gridrow="3" pageId="5" pageNumber="388">1.05</td>
<td id="C74CC321376600666C16FC9C07359C6C" box="[1313,1370,781,806]" gridcol="6" gridrow="3" pageId="5" pageNumber="388">13.37</td>
</tr>
<tr id="849DAA5D376600666994FCA407359C04" box="[163,1370,821,846]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCA402BF9C04" box="[163,208,821,846]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666858FCA403F49C04" box="[367,411,821,846]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3.31</td>
<td id="C74CC321376600666B19FCA400359C04" box="[558,602,821,846]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1.71</td>
<td id="C74CC321376600666BDBFCA401779C04" box="[748,792,821,846]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">2.45</td>
<td id="C74CC321376600666A9CFCA401B89C04" box="[939,983,821,846]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">2.91</td>
<td id="C74CC321376600666D5EFCA406FA9C04" box="[1129,1173,821,846]" gridcol="5" gridrow="4" pageId="5" pageNumber="388">1.09</td>
<td id="C74CC321376600666C16FCA407359C04" box="[1313,1370,821,846]" gridcol="6" gridrow="4" pageId="5" pageNumber="388">11.47</td>
</tr>
<tr id="849DAA5D376600666994FCCF07359C3C" box="[163,1370,862,886]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCCF02BF9C3C" box="[163,208,862,886]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666858FCCF03F49C3C" box="[367,411,862,886]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">4.61</td>
<td id="C74CC321376600666B19FCCF00359C3C" box="[558,602,862,886]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1.77</td>
<td id="C74CC321376600666BDBFCCF01779C3C" box="[748,792,862,886]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">4.00</td>
<td id="C74CC321376600666A9CFCCF01B89C3C" box="[939,983,862,886]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">5.34</td>
<td id="C74CC321376600666D5EFCCF06FA9C3C" box="[1129,1173,862,886]" gridcol="5" gridrow="5" pageId="5" pageNumber="388">1.52</td>
<td id="C74CC321376600666C16FCCF07359C3C" box="[1313,1370,862,886]" gridcol="6" gridrow="5" pageId="5" pageNumber="388">17.24</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973766FF9C69A0FC5A01B99CA9" box="[151,982,971,995]" pageId="5" pageNumber="388" startId="5.[151,239,971,995]" targetBox="[163,1366,1011,1237]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FC5A01B99CA9" blockId="5.[151,982,971,995]" box="[151,982,971,995]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FC5A03659CA9" bold="true" box="[151,266,971,995]" pageId="5" pageNumber="388">TABLE 4.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C3766FF9C6BF0FC5D01159CA9" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[711,890,972,995]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF0FC5D01159CA9" box="[711,890,972,995]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB7FC5A01B99CA9" box="[896,982,971,995]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C68B8FC6507399B9F" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FC6207399B9F" box="[163,1366,1011,1237]" gridcols="5" gridrows="6" pageId="5" pageNumber="388">
<tr id="849DAA5D376600666994FC6207399B41" box="[163,1366,1011,1035]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666854FC6203BA9B41" box="[355,469,1011,1035]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666BA3FC6200809B41" box="[660,751,1011,1035]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666ABEFC62062A9B41" box="[905,1093,1011,1035]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666DF5FC6207399B41" box="[1218,1366,1011,1035]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
</tr>
<tr id="849DAA5D376600666994FB8D07399B7E" box="[163,1366,1052,1076]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB8D02BF9B7E" box="[163,208,1052,1076]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666854FB8D03BA9B7E" box="[355,469,1052,1076]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">3d 1p 1r</td>
<td id="C74CC321376600666BA3FB8D00809B7E" box="[660,751,1052,1076]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1(0)p</td>
<td id="C74CC321376600666ABEFB8D062A9B7E" box="[905,1093,1052,1076]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1d 2p 1r</td>
<td id="C74CC321376600666DF5FB8D07399B7E" box="[1218,1366,1052,1076]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
</tr>
<tr id="849DAA5D376600666994FBD507399B16" box="[163,1366,1092,1116]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBD502BF9B16" box="[163,208,1092,1116]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666854FBD503BA9B16" box="[355,469,1092,1116]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">3d 3p 3(2)r</td>
<td id="C74CC321376600666BA3FBD500809B16" box="[660,751,1092,1116]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBD5062A9B16" box="[905,1093,1092,1116]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBD507399B16" box="[1218,1366,1092,1116]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FBFD07399BCE" box="[163,1366,1132,1156]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBFD02BF9BCE" box="[163,208,1132,1156]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666854FBFD03BA9BCE" box="[355,469,1132,1156]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FBFD00809BCE" box="[660,751,1132,1156]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBFD062A9BCE" box="[905,1093,1132,1156]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBFD07399BCE" box="[1218,1366,1132,1156]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB0507399BE7" box="[163,1366,1172,1197]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB0502BF9BE7" box="[163,208,1172,1197]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666854FB0503BA9BE7" box="[355,469,1172,1197]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3d 4p 4r</td>
<td id="C74CC321376600666BA3FB0500809BE7" box="[660,751,1172,1197]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB05062A9BE7" box="[905,1093,1172,1197]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">3d 2p 2r 2-2v</td>
<td id="C74CC321376600666DF5FB0507399BE7" box="[1218,1366,1172,1197]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">4p 5r 2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB2C07399B9F" box="[163,1366,1213,1237]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB2C02BF9B9F" box="[163,208,1213,1237]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666854FB2C03BA9B9F" box="[355,469,1213,1237]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FB2C00809B9F" box="[660,751,1213,1237]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB2C062A9B9F" box="[905,1093,1213,1237]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">3(2)d 2p 2r 2-2-2v</td>
<td id="C74CC321376600666DF5FB2C07399B9F" box="[1218,1366,1213,1237]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FAB903E59AE4" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
Female genitalia as shown in
<figureCitation id="A296B49A3766FF9C6B2BFAB900FE9A08" box="[540,657,1320,1346]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="5" pageNumber="388">Figs 9–12</figureCitation>
. Epigyne 1.5 times wider than long. Median plate (
<emphasis id="08D9740D3766FF9C6DE9FAB807699A08" box="[1246,1286,1321,1346]" italics="true" pageId="5" pageNumber="388">MP</emphasis>
) trapezoidal, strongly sclerotized anteriorly. Lateral teeth (
<emphasis id="08D9740D3766FF9C6B87FADC00A09A2C" box="[688,719,1357,1382]" italics="true" pageId="5" pageNumber="388">LT</emphasis>
) well developed, located posterior of median plate, slightly extending beyond epigastric furrow, with rounded tips. Spermathecae (
<emphasis id="08D9740D3766FF9C6A8BFAE001B79AC1" box="[956,984,1393,1419]" italics="true" pageId="5" pageNumber="388">Sp</emphasis>
) large, piriform, separated by a third of spermatheca’s length.
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FA29073999CC" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0FA29037E9A98" bold="true" box="[199,273,1464,1490]" pageId="5" pageNumber="388">Notes.</emphasis>
Since males
<taxonomicName id="FDADD39C3766FF9C689DFA2800579A98" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[426,568,1465,1490]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C689DFA2800579A98" box="[426,568,1465,1490]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6B76FA2900F39A98" bold="true" box="[577,668,1464,1490]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6B76FA2900F39A98" box="[577,668,1464,1490]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are unknown, it is difficult to place it in a species group. There are species with a similar shape of median plate and lateral teeth situated at the posterior margin of the epigyne in
<emphasis id="08D9740D3766FF9C69A0F99102919950" box="[151,254,1536,1562]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
and
<taxonomicName id="FDADD39C3766FF9C680EF99003B49950" authorityName="Jager" authorityYear="2022" box="[313,475,1537,1562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C680EF99003B49950" box="[313,475,1537,1562]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
species groups. Because the new species have spermathecae consisting of only one compartment, while those in
<emphasis id="08D9740D3766FF9C68E8F9B500299974" box="[479,582,1572,1598]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
-group have particularly structured spermathecae, we place
<taxonomicName id="FDADD39C3766FF9C6DD2F9B4071C9974" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1253,1395,1573,1598]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6DD2F9B4071C9974" box="[1253,1395,1573,1598]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6C4CF9B502A79928" bold="true" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6C4CF9B502A79928" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the
<taxonomicName id="FDADD39C3766FF9C6820F9D803D69928" authorityName="Jager" authorityYear="2022" box="[279,441,1609,1634]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C6820F9D803D69928" box="[279,441,1609,1634]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
-species group. Thus, with the description of the new species, this group now consists of five species:
<taxonomicName id="FDADD39C3766FF9C681EF9FC038C99CC" authorityName="Omelko &amp; Fomichev" authorityYear="2023" box="[297,483,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="binturong">
<emphasis id="08D9740D3766FF9C681EF9FC038C99CC" box="[297,483,1644,1670]" italics="true" pageId="5" pageNumber="388">Bowie binturong</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C68C5F9FC001099CC" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[498,639,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C68C5F9FC001099CC" box="[498,639,1645,1670]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6BB1F9FD008F99CC" bold="true" box="[646,736,1644,1670]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6BB1F9FD008F99CC" box="[646,736,1644,1670]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="FDADD39C3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" italics="true" pageId="5" pageNumber="388">B. neukoeln</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C6AB7F9FC063B99CD" authority="Li &amp; Yao" authorityName="Li &amp; Yao" authorityYear="2022" box="[896,1108,1644,1671]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="sabah">
<emphasis id="08D9740D3766FF9C6AB7F9FC018D99CC" box="[896,994,1644,1670]" italics="true" pageId="5" pageNumber="388">B. sabah</emphasis>
Li &amp; Yao
</taxonomicName>
, and
<emphasis id="08D9740D3766FF9C6DB9F9FC073999CC" box="[1166,1366,1645,1670]" italics="true" pageId="5" pageNumber="388">
<taxonomicName id="FDADD39C3766FF9C6DB9F9FC073D99CC" authorityName="Jager" authorityYear="2022" box="[1166,1362,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">B. scarymonsters</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943766FF9C69F0F90101EC99E0" box="[199,899,1680,1707]" pageId="5" pageNumber="388" type="distribution">
<paragraph id="3A12A81F3766FF9C69F0F90101EC99E0" blockId="5.[151,1436,1320,1707]" box="[199,899,1680,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0F901030E99E0" bold="true" box="[199,353,1680,1706]" pageId="5" pageNumber="388">Distribution.</emphasis>
<typeStatus id="E51616BD3766FF9C6850F90003F099E1" box="[359,415,1681,1707]" pageId="5" pageNumber="388">Type</typeStatus>
locality only, Borneo:
<collectingRegion id="F86966FD3766FF9C6B97F901008B99E0" box="[672,740,1680,1706]" country="Malaysia" name="Sabah" pageId="5" pageNumber="388">Sabah</collectingRegion>
(
<figureCitation id="A296B49A3766FF9C6BC4F901011899E0" box="[755,887,1680,1706]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="5" pageNumber="388">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

288
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@ -0,0 +1,288 @@
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<mods:title id="033A41AB3AC8A384C26F5FB09218A7AC">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
</mods:titleInfo>
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<mods:namePart id="124DB3A58F6604C7D65FA81C3257E310">Omelko, Mikhail M.</mods:namePart>
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<mods:affiliation id="431BD2D073BE23BEC15777B53EFC55C2">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
<mods:nameIdentifier id="95F05FF040E51031F148A6B01322DEBA" type="email">omelkom@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4798C2F81CD0A272576BB9ECA1D8FCBC">Fomichev, Alexander A.</mods:namePart>
<mods:nameIdentifier id="AFE7ED8BE31F4F13D268703763CD0BD0" type="ORCID">0000-0001-9268-622X</mods:nameIdentifier>
<mods:affiliation id="C1CD02F9BE2030C96395494410046E1E">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
<mods:nameIdentifier id="6CC21D1CDB034BD6892BABAACB36CD2C" type="email">a.fomichov@mail.ru</mods:nameIdentifier>
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<mods:date id="DFADC3B41F70D605E614174489FFF64A">2024</mods:date>
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<treatment id="B20419093769FF9569A0FDAD01F69D2B" LSID="urn:lsid:plazi:treatment:B20419093769FF9569A0FDAD01F69D2B" httpUri="http://treatment.plazi.org/id/B20419093769FF9569A0FDAD01F69D2B" lastPageId="12" lastPageNumber="395" pageId="10" pageNumber="393">
<subSubSection id="72B7FB943769FF9369A0FDAD03B69D1C" box="[151,473,571,598]" pageId="10" pageNumber="393" type="nomenclature">
<paragraph id="3A12A81F3769FF9369A0FDAD03B69D1C" blockId="10.[151,473,571,634]" box="[151,473,571,598]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369A0FDAD03B69D1C" bold="true" box="[151,473,571,598]" pageId="10" pageNumber="393">
<taxonomicName id="FDADD39C3769FF9369A0FDAD031B9D1C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,372,571,598]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D3769FF9369A0FDAD031B9D1C" bold="true" box="[151,372,571,598]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763769FF93684CFDAD03B69D1C" box="[379,473,572,598]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943769FF9369A0FDF103149D30" box="[151,379,608,634]" pageId="10" pageNumber="393" type="description">
<paragraph id="3A12A81F3769FF9369A0FDF103149D30" blockId="10.[151,473,571,634]" box="[151,379,608,634]" pageId="10" pageNumber="393">
(
<figureCitation id="A296B49A3769FF9369A8FDF1034E9D30" box="[159,289,608,634]" captionStart-0="FIGURES 31–32" captionStart-1="FIGURES 33–41" captionStartId-0="10.[151,264,1302,1326]" captionStartId-1="11.[151,264,1864,1888]" captionTargetBox-0="[153,1431,770,1275]" captionTargetBox-1="[163,1435,194,1834]" captionTargetId-0="figure-216@10.[151,1435,766,1278]" captionTargetId-1="figure-19@11.[151,1435,181,1840]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 31–32. Male habitus of Bowie tangalunga sp. nov. 31—dorsal; 32—ventral. Scale bar: 2 mm." captionText-1="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi-0="http://doi.org/10.5281/zenodo.14386413" figureDoi-1="http://doi.org/10.5281/zenodo.14386415" httpUri-0="https://zenodo.org/record/14386413/files/figure.png" httpUri-1="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Figs 31–41</figureCitation>
,
<figureCitation id="A296B49A3769FF93681BFDF1031C9D30" box="[300,371,608,634]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="10" pageNumber="393">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943769FF9369A0FD3907FA9DAC" pageId="10" pageNumber="393" type="materials_examined">
<paragraph id="3A12A81F3769FF9369A0FD3907FA9DAC" blockId="10.[151,1437,679,743]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369A0FD39032B9D88" bold="true" box="[151,324,680,706]" pageId="10" pageNumber="393">Type material.</emphasis>
<materialsCitation id="8AC5A2423769FF93687DFD3901309DAC" collectingDate="1995-01-26" collectingDateMax="1995-01-30" collectingDateMin="1995-01-26" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Philippines" elevation="1350" latitude="7.016667" location="Apo Mt." longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" pageId="10" pageNumber="393" specimenCount="1" specimenCount-male="1" stateProvince="Cotabato Province" typeStatus="holotype">
<emphasis id="08D9740D3769FF93687DFD3901309D88" bold="true" box="[330,863,679,706]" pageId="10" pageNumber="393">
<collectingCountry id="42BAE88F3769FF93687DFD39006E9D88" box="[330,513,680,706]" name="Philippines" pageId="10" pageNumber="393">PHILIPPINES</collectingCountry>
:
<emphasis id="08D9740D3769FF936B3AFD36008C9D88" bold="true" box="[525,739,679,706]" italics="true" pageId="10" pageNumber="393">
<collectingRegion id="F86966FD3769FF936B3AFD36008C9D88" box="[525,739,679,706]" country="Philippines" name="South Cotabato" pageId="10" pageNumber="393">Cotabato Province</collectingRegion>
</emphasis>
:
<typeStatus id="E51616BD3769FF936BC5FD3901309D88" box="[754,863,680,706]" pageId="10" pageNumber="393" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="2CAB63963769FF936A5AFD3901FF9D88" box="[877,912,680,706]" count="1" pageId="10" pageNumber="393" type="male">1♂</specimenCount>
(
<collectionCode id="5CBC30DA3769FF936AA9FD3901949D88" box="[926,1019,680,706]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="393" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653769FF936D3AFD3906A39D88" box="[1037,1228,680,706]" pageId="10" pageNumber="393">Mindanao Island</collectingMunicipality>
,
<location id="3F72FEC43769FF936DE2FD3907599D89" LSID="urn:lsid:plazi:treatment:B20419093769FF9569A0FDAD01F69D2B:3F72FEC43769FF936DE2FD3907599D89" box="[1237,1334,680,707]" country="Philippines" latitude="7.016667" longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" name="Apo Mt." pageId="10" pageNumber="393" stateProvince="Cotabato Province">Apo Mt.</location>
, [
<geoCoordinate id="5F99CED83769FF936C7EFD3907F89D88" box="[1353,1431,680,706]" degrees="7" direction="north" minutes="1" orientation="latitude" pageId="10" pageNumber="393" precision="925" value="7.016667">7° 1’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83769FF9369A0FD5D03689DAD" box="[151,263,716,743]" degrees="125" direction="east" minutes="13" orientation="longitude" pageId="10" pageNumber="393" precision="925" value="125.21667">125°13’E</geoCoordinate>
],
<quantity id="FD5505FA3769FF93682DFD5D031E9DAD" box="[282,369,716,743]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="393" unit="m" value="1350.0">
<elevation id="B1804F2C3769FF93682DFD5D031E9DAD" box="[282,369,716,743]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="393" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<date id="4E138EDF3769FF93684AFD5D000C9DAD" box="[381,611,716,743]" pageId="10" pageNumber="393" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">
<collectingDate id="5E5777373769FF93684AFD5D000C9DAD" box="[381,611,716,743]" pageId="10" pageNumber="393" value="1995-01-26" valueMax="1995-01-30" valueMin="1995-01-26">26–30 January 1995</collectingDate>
</date>
,
<collectorName id="9758CDC93769FF936B5BFD5D01409DAC" box="[620,815,716,742]" pageId="10" pageNumber="393">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8AC5A2423769FF936A52FD5D07FE9DAC" box="[869,1425,716,743]" collectingDate="1995-01-26" collectingDateMax="1995-01-30" collectingDateMin="1995-01-26" collectionCode="IBSS" collectorName="A. M. Emelyanov" country="Philippines" elevation="1350" latitude="7.016667" location="Apo Mt." longLatPrecision="1303" longitude="125.21667" municipality="Mindanao Island" pageId="10" pageNumber="393" specimenCount="1" specimenCount-male="1" stateProvince="Paratype" typeStatus="paratype">
<emphasis id="08D9740D3769FF936A52FD5D01BD9DAC" bold="true" box="[869,978,716,742]" pageId="10" pageNumber="393">
<typeStatus id="E51616BD3769FF936A52FD5D01BD9DAC" box="[869,978,716,742]" pageId="10" pageNumber="393" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="2CAB63963769FF936AD6FD5D066B9DAC" box="[993,1028,716,742]" count="1" pageId="10" pageNumber="393" type="male">1♂</specimenCount>
(
<collectionCode id="5CBC30DA3769FF936D24FD5D063C9DAC" box="[1043,1107,716,742]" collectionName="IBSS" pageId="10" pageNumber="393">IBSS</collectionCode>
), together with the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6ED2F8973769FF9369A0FA8706A99A67" ID-DOI="http://doi.org/10.5281/zenodo.14386413" ID-Zenodo-Dep="14386413" box="[151,1222,1302,1326]" httpUri="https://zenodo.org/record/14386413/files/figure.png" pageId="10" pageNumber="393" startId="10.[151,264,1302,1326]" targetBox="[153,1431,770,1275]" targetPageId="10" targetType="figure">
<paragraph id="3A12A81F3769FF9369A0FA8706A99A67" blockId="10.[151,1222,1302,1326]" box="[151,1222,1302,1326]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369A0FA87033B9A64" bold="true" box="[151,340,1302,1326]" pageId="10" pageNumber="393">FIGURES 31–32.</emphasis>
Male habitus of
<taxonomicName id="FDADD39C3769FF9368C8FA8600D79A64" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[511,696,1302,1326]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D3769FF9368C8FA8600D79A64" box="[511,696,1302,1326]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
</taxonomicName>
<emphasis id="08D9740D3769FF936B88FA8701609A64" bold="true" box="[703,783,1302,1326]" pageId="10" pageNumber="393">
<taxonomicNameLabel id="13EAC9763769FF936B88FA8701609A64" box="[703,783,1302,1326]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
31—dorsal; 32—ventral. Scale bar: 2 mm.
</paragraph>
</caption>
<subSubSection id="72B7FB943769FF9369F0FAE701F49AF9" pageId="10" pageNumber="393" type="etymology">
<paragraph id="3A12A81F3769FF9369F0FAE701F49AF9" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369F0FAE703279ADA" bold="true" box="[199,328,1398,1424]" pageId="10" pageNumber="393">Etymology</emphasis>
. The specific name is derived from the local name of the Malayan civet [
<taxonomicName id="FDADD39C3769FF936D43FAE707FE9ADA" authority="Gray" authorityName="Gray" authorityYear="1832" box="[1140,1425,1397,1424]" class="Mammalia" family="Viverridae" genus="Viverra" kingdom="Animalia" order="Carnivora" pageId="10" pageNumber="393" phylum="Chordata" rank="species" species="tangalunga">
<emphasis id="08D9740D3769FF936D43FAE707219AC5" box="[1140,1358,1397,1423]" italics="true" pageId="10" pageNumber="393">Viverra tangalunga</emphasis>
Gray
</taxonomicName>
], a small predatory beast native to Southeast Asia; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943769FF9369F0FA2C0683992D" pageId="10" pageNumber="393" type="diagnosis">
<paragraph id="3A12A81F3769FF9369F0FA2C0683992D" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369F0FA2C03559A9D" bold="true" box="[199,314,1469,1495]" pageId="10" pageNumber="393">Diagnosis</emphasis>
. Males of
<taxonomicName id="FDADD39C3769FF936886FA2F00119A9D" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[433,638,1469,1495]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D3769FF936886FA2F00119A9D" box="[433,638,1469,1495]" italics="true" pageId="10" pageNumber="393">Bowie tangalunga</emphasis>
</taxonomicName>
<emphasis id="08D9740D3769FF936BB3FA2F00B29A92" bold="true" box="[644,733,1470,1496]" pageId="10" pageNumber="393">
<taxonomicNameLabel id="13EAC9763769FF936BB3FA2F00B29A92" box="[644,733,1470,1496]" pageId="10" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="FDADD39C3769FF936A87FA2F06019A9D" authorityName="Jager" authorityYear="2022" box="[944,1134,1469,1495]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="criminalworld">
<emphasis id="08D9740D3769FF936A87FA2F06019A9D" box="[944,1134,1469,1495]" italics="true" pageId="10" pageNumber="393">B. criminalworld</emphasis>
</taxonomicName>
by the shape of the tegular apophysis (
<emphasis id="08D9740D3769FF93682DFA7303569AB6" box="[282,313,1506,1532]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
) and the shape of the retro-proximal cymbial outgrowth (
<emphasis id="08D9740D3769FF936AEFFA7306619AB6" box="[984,1038,1506,1532]" italics="true" pageId="10" pageNumber="393">RPO</emphasis>
) in ventral view but can be easily distinguished by 1) embolus hidden behind the tegular apophysis to a large extent (vs. not hidden; cf.
<figureCitation id="A296B49A3769FF936C2AF99407F3996A" box="[1309,1436,1541,1568]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Figs 34, 38</figureCitation>
and
<bibRefCitation id="5E3CD5EE3769FF9369F0F9BB0329990E" author="Jager, P." box="[199,326,1578,1604]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 769), 2) base of
<emphasis id="08D9740D3769FF936B00F9BB0039990E" box="[567,598,1578,1604]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
located almost medially (vs. retrolaterally; cf.
<figureCitation id="A296B49A3769FF936D52F9B806DA990E" box="[1125,1205,1577,1604]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Fig. 38</figureCitation>
and
<bibRefCitation id="5E3CD5EE3769FF936DDBF9BB0704990E" author="Jager, P." box="[1260,1387,1578,1604]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 769), 3) tip of embolus (
<emphasis id="08D9740D3769FF93689EF9DF03A1992D" box="[425,462,1614,1639]" italics="true" pageId="10" pageNumber="393">Em</emphasis>
) extending
<emphasis id="08D9740D3769FF936B63F9DF001D9922" box="[596,626,1614,1640]" italics="true" pageId="10" pageNumber="393">TA</emphasis>
(vs. not extending; cf.
<figureCitation id="A296B49A3769FF936A42F9DC01AB9922" box="[885,964,1613,1640]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="10" pageNumber="393">Fig. 38</figureCitation>
and
<bibRefCitation id="5E3CD5EE3769FF936ACCF9DF06169922" author="Jager, P." box="[1019,1145,1614,1640]" pageId="10" pageNumber="393" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 769).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943769FF9569F0F9E000D09D77" lastPageId="12" lastPageNumber="395" pageId="10" pageNumber="393" type="description">
<paragraph id="3A12A81F3769FF9369F0F9E0037098CD" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">
<emphasis id="08D9740D3769FF9369F0F9E0033699C1" bold="true" box="[199,345,1649,1675]" pageId="10" pageNumber="393">Description.</emphasis>
<emphasis id="08D9740D3769FF936855F9E003F399C1" box="[354,412,1649,1675]" italics="true" pageId="10" pageNumber="393">Male</emphasis>
(
<figureCitation id="A296B49A3769FF93689BF9E0005E99C6" box="[428,561,1649,1676]" captionStart="FIGURES 31–32" captionStartId="10.[151,264,1302,1326]" captionTargetBox="[153,1431,770,1275]" captionTargetId="figure-216@10.[151,1435,766,1278]" captionTargetPageId="10" captionText="FIGURES 31–32. Male habitus of Bowie tangalunga sp. nov. 31—dorsal; 32—ventral. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386413" httpUri="https://zenodo.org/record/14386413/files/figure.png" pageId="10" pageNumber="393">Figs 31–32</figureCitation>
). Total length 13.11. Prosoma 7.10 long, 5.23 wide. Opisthosoma 6.05 long, 4.05 wide. Prosoma black, with light brown median band, slightly widened near ocular area. Lateral bands disrupted into several whitish spots. Fovea thin, black. Clypeus light brown. Chelicerae dark brown with brown longitudinal stripes anteriorly, 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum brown without pattern. Labium dark brown. Endites brown with yellow distal edge. Dorsal part of opisthosoma black with short yellowish cardiac mark and series of transverse stripes posterior of it. Lateral sides of opisthosoma black with lots of tiny yellow spots. Ventral part of opisthosoma black with yellow spots forming two V-marks (inner mark poorly visible). Spinnerets dark brown.
</paragraph>
<paragraph id="3A12A81F3769FF9369F0F80307EF989A" blockId="10.[151,1437,1397,2000]" pageId="10" pageNumber="393">Eye diameters: AME 0.28, ALE 0.21, PME 0.38, PLE 0.32; interdistances: AME–AME 0.16, AME–ALE 0.38, PME–PME 0.23, PME–PLE 0.35, AME–PME 0.10, ALE–PLE 0.18. Clypeus height at AME 0.20, at ALE 0.59.</paragraph>
<caption id="6ED2F8973768FF9269A0F8D9008D9886" ID-DOI="http://doi.org/10.5281/zenodo.14386415" ID-Zenodo-Dep="14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="11" pageNumber="394" startId="11.[151,264,1864,1888]" targetBox="[163,1435,194,1834]" targetPageId="11" targetType="figure">
<paragraph id="3A12A81F3768FF9269A0F8D9008D9886" blockId="11.[151,1436,1864,1996]" pageId="11" pageNumber="394">
<emphasis id="08D9740D3768FF9269A0F8D9033C982A" bold="true" box="[151,339,1864,1888]" pageId="11" pageNumber="394">FIGURES 33–41.</emphasis>
Palp (33–36), bulb (37–40) and palpal tibia (41) of male of
<taxonomicName id="FDADD39C3768FF926A91F8D80632982A" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[934,1117,1864,1888]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="394" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D3768FF926A91F8D80632982A" box="[934,1117,1864,1888]" italics="true" pageId="11" pageNumber="394">Bowie tangalunga</emphasis>
</taxonomicName>
<emphasis id="08D9740D3768FF926D54F8D906DD982A" bold="true" box="[1123,1202,1864,1888]" pageId="11" pageNumber="394">
<taxonomicNameLabel id="13EAC9763768FF926D54F8D906DD982A" box="[1123,1202,1864,1888]" pageId="11" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm. Abbreviations:
<emphasis id="08D9740D3768FF926C45F8FD07EC98C9" box="[1394,1411,1900,1923]" italics="true" pageId="11" pageNumber="394">C</emphasis>
— conductor,
<emphasis id="08D9740D3768FF92683DF800034398E2" box="[266,300,1937,1960]" italics="true" pageId="11" pageNumber="394">EH</emphasis>
—embolic hook,
<emphasis id="08D9740D3768FF9268D6F800006C98E2" box="[481,515,1937,1960]" italics="true" pageId="11" pageNumber="394">Em</emphasis>
—embolus,
<emphasis id="08D9740D3768FF926BB7F80100DE98ED" box="[640,689,1936,1959]" italics="true" pageId="11" pageNumber="394">RPO</emphasis>
—retro-proximal cymbial outgrowth,
<emphasis id="08D9740D3768FF926D0FF801060C98ED" box="[1080,1123,1936,1959]" italics="true" pageId="11" pageNumber="394">RTA</emphasis>
—retrolateral tibial apophysis,
<emphasis id="08D9740D3768FF9269A0F82502DC9881" box="[151,179,1972,1995]" italics="true" pageId="11" pageNumber="394">TA</emphasis>
—tegular apophysis,
<emphasis id="08D9740D3768FF9268BCF82403CD9886" box="[395,418,1973,1996]" italics="true" pageId="11" pageNumber="394">Te</emphasis>
—tegulum,
<emphasis id="08D9740D3768FF926B2CF82400549886" box="[539,571,1973,1996]" italics="true" pageId="11" pageNumber="394">TH</emphasis>
—tegular hump.
</paragraph>
</caption>
<paragraph id="3A12A81F376FFF9569F0FF0600DA9E2F" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
For palp and legs measurements see
<tableCitation id="772F9DA4376FFF956B59FF0600A99FFB" box="[622,710,151,177]" captionStart="TABLE 9" captionStartId="12.[151,239,656,680]" captionTargetPageId="12" captionText="TABLE 9. Palp and legs measurements of male of Bowie tangalunga sp. nov." pageId="12" pageNumber="395">Table 9</tableCitation>
. Palp coloration: femora, patellae, tibiae brown; cymbium dark brown. Legs coloration: femora I dark brown dorsally and laterally, brown ventrally, with light brown longitudinal stripe retrolaterally; II dark brown dorsally and prolaterally, brown retrolaterally and ventrally; III–IV dark brown dorsally and laterally, brown ventrally, with light brown semi-rings. Patellae I–IV brown. Tibiae I–II, IV light brown with white setae; III light brown. Metatarsi I–II brown; III–IV brown with light brow semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA4376FFF956B7AFEDA00DE9E2F" box="[589,689,331,357]" captionStart="TABLE 10" captionStartId="12.[151,239,1007,1031]" captionTargetPageId="12" captionText="TABLE 10. Palp and legs spination for the male of Bowie tangalunga sp. nov." pageId="12" pageNumber="395">Table 10</tableCitation>
.
</paragraph>
<paragraph id="3A12A81F376FFF9569F0FEFE07F49EBF" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
Palp as shown in
<figureCitation id="A296B49A376FFF9568BFFEFE00679EC3" box="[392,520,367,393]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1864,1888]" captionTargetBox="[163,1435,194,1834]" captionTargetId="figure-19@11.[151,1435,181,1840]" captionTargetPageId="11" captionText="FIGURES 33–41. Palp (33–36), bulb (37–40) and palpal tibia (41) of male of Bowie tangalunga sp. nov. 33, 37—prolateral; 34, 38—ventral; 35, 39, 41—retrolateral; 36—dorsal; 40—apical. Scale bars: 33–36 = 0.5 mm; 37–41 = 0.2 mm.Abbreviations: C— conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386415" httpUri="https://zenodo.org/record/14386415/files/figure.png" pageId="12" pageNumber="395">Figs 33–41</figureCitation>
.
<emphasis id="08D9740D376FFF956B26FEFE002E9EC3" box="[529,577,367,393]" italics="true" pageId="12" pageNumber="395">RTA</emphasis>
arising from middle of tibia, short, curved, with sharply pointed tip in ventral view. Cymbium with short, blunt retro-proximal cymbial outgrowth (
<emphasis id="08D9740D376FFF956AAFFE0201A19EE7" box="[920,974,403,429]" italics="true" pageId="12" pageNumber="395">RPO</emphasis>
). Tegulum (
<emphasis id="08D9740D376FFF956D61FE0206009EE6" box="[1110,1135,403,428]" italics="true" pageId="12" pageNumber="395">Te</emphasis>
) with rounded hump (
<emphasis id="08D9740D376FFF956C5FFE0207E39EE6" box="[1384,1420,403,428]" italics="true" pageId="12" pageNumber="395">TH</emphasis>
). Tegular apophysis (
<emphasis id="08D9740D376FFF956843FE2603FC9E9B" box="[372,403,439,465]" italics="true" pageId="12" pageNumber="395">TA</emphasis>
) oval, originating in a 5 o’clock position. Conductor (
<emphasis id="08D9740D376FFF956AC5FE26066A9E9B" box="[1010,1029,439,465]" italics="true" pageId="12" pageNumber="395">C</emphasis>
) large, membranous. Embolus (
<emphasis id="08D9740D376FFF956C59FE2607FC9E9A" box="[1390,1427,439,464]" italics="true" pageId="12" pageNumber="395">Em</emphasis>
) originating from tegulum in 6.30-o’clock-position, with pointed tip. Embolus with large basal embolus hook (
<emphasis id="08D9740D376FFF956C51FE4A07E49EBE" box="[1382,1419,475,500]" italics="true" pageId="12" pageNumber="395">EH</emphasis>
).
</paragraph>
<paragraph id="3A12A81F376FFF9569F0FE6E00D09D77" blockId="12.[151,1437,151,609]" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569F0FE6E037E9D53" bold="true" box="[199,273,511,537]" pageId="12" pageNumber="395">Notes.</emphasis>
According to the structure of the copulatory organs,
<taxonomicName id="FDADD39C376FFF956A5AFE6E06549D53" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[877,1083,511,537]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="tangalunga" status="sp. nov.">
<emphasis id="08D9740D376FFF956A5AFE6E06549D53" box="[877,1083,511,537]" italics="true" pageId="12" pageNumber="395">Bowie tangalunga</emphasis>
</taxonomicName>
<emphasis id="08D9740D376FFF956D74FE6E06F29D53" bold="true" box="[1091,1181,511,537]" pageId="12" pageNumber="395">
<taxonomicNameLabel id="13EAC976376FFF956D74FE6E06F29D53" box="[1091,1181,511,537]" pageId="12" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="FDADD39C376FFF956C7AFE6E07F39D53" box="[1357,1436,511,537]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376FFF956C7AFE6E07F39D53" box="[1357,1436,511,537]" italics="true" pageId="12" pageNumber="395">shakeit</emphasis>
</taxonomicName>
species-group (see notes for
<taxonomicName id="FDADD39C376FFF9568EFFDB200219D76" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[472,590,547,572]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="musang" status="sp. nov.">
<emphasis id="08D9740D376FFF9568EFFDB200219D76" box="[472,590,547,572]" italics="true" pageId="12" pageNumber="395">B. musang</emphasis>
</taxonomicName>
<emphasis id="08D9740D376FFF956B62FDB200C19D77" bold="true" box="[597,686,547,573]" pageId="12" pageNumber="395">
<taxonomicNameLabel id="13EAC976376FFF956B62FDB200C19D77" box="[597,686,547,573]" pageId="12" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF9569F0FDD701F69D2B" box="[199,921,582,609]" pageId="12" pageNumber="395" type="distribution">
<paragraph id="3A12A81F376FFF9569F0FDD701F69D2B" blockId="12.[151,1437,151,609]" box="[199,921,582,609]" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569F0FDD7030E9D2A" bold="true" box="[199,353,582,608]" pageId="12" pageNumber="395">Distribution.</emphasis>
Mindanao Isl., Apo Mt.,
<collectingCountry id="42BAE88F376FFF956B4BFDD600959D2B" box="[636,762,583,609]" name="Philippines" pageId="12" pageNumber="395">Philippines</collectingCountry>
(
<figureCitation id="A296B49A376FFF956A3EFDD601E29D2B" box="[777,909,583,609]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="12" pageNumber="395">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="CFDABE726E2E7E61C5A1D604EBC47C1F">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
</mods:titleInfo>
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<mods:namePart id="076985095640CCEFFD1FD7A642EABC9D">Omelko, Mikhail M.</mods:namePart>
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<mods:affiliation id="90C36ED17F64A43413D5B0A22817DF28">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
<mods:nameIdentifier id="667AFF58BF864F94869430368662E5E5" type="email">omelkom@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E01B3809EA05D0782FFFE901AE445249">Fomichev, Alexander A.</mods:namePart>
<mods:nameIdentifier id="CB2BB844BB608BBA816075FB87EDC2AB" type="ORCID">0000-0001-9268-622X</mods:nameIdentifier>
<mods:affiliation id="1FA629EEDFDEE2BB5FE17162494CF566">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
<mods:nameIdentifier id="4FA613AE8A94529CC20B0342792BB7CB" type="email">a.fomichov@mail.ru</mods:nameIdentifier>
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<mods:typeOfResource id="77EEA43FF5CC27D60A854D5E9E248036">text</mods:typeOfResource>
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<mods:title id="553E760D066C0F586BD2AE374BC79E2D">Zootaxa</mods:title>
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<mods:part id="8062743777324165ADDB84B2983A3A41">
<mods:date id="7166F2D969A5CDE22C1247DE291EFF4B">2024</mods:date>
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<mods:number id="F20E7D72778654ECEF01163480BA8F94">2024-12-04</mods:number>
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<mods:number id="8094820227C775EC9117F4638F2344C8">5543</mods:number>
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<mods:number id="115191908C8B7FD7772D9E8A705A2A18">3</mods:number>
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<mods:start id="22A25D4B21C64C8FC8B6737D0D036595">383</mods:start>
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<mods:identifier id="96439D6A717437ACA7CAAC785E22E853" type="DOI">10.11646/zootaxa.5543.3.5</mods:identifier>
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<subSubSection id="72B7FB94376FFF9569A0FADD03BB9A2C" box="[151,468,1356,1383]" pageId="12" pageNumber="395" type="nomenclature">
<paragraph id="3A12A81F376FFF9569A0FADD03BB9A2C" blockId="12.[151,468,1356,1419]" box="[151,468,1356,1383]" pageId="12" pageNumber="395">
<taxonomicName id="FDADD39C376FFF9569A0FADD03BB9A2C" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[151,468,1356,1383]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376FFF9569A0FADD03BB9A2C" bold="true" box="[151,468,1356,1383]" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569A0FADD03509A2D" bold="true" box="[151,319,1356,1383]" italics="true" pageId="12" pageNumber="395">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376FFF956870FADD03BB9A2C" author="Jager, P." box="[327,468,1356,1382]" pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF9569A0FAE103149AC0" box="[151,379,1392,1419]" pageId="12" pageNumber="395" type="description">
<paragraph id="3A12A81F376FFF9569A0FAE103149AC0" blockId="12.[151,468,1356,1419]" box="[151,379,1392,1419]" pageId="12" pageNumber="395">
(
<figureCitation id="A296B49A376FFF9569A8FAE1034F9AC1" box="[159,288,1392,1419]" captionStart-0="FIGURES 42–45" captionStart-1="FIGURES 46–53" captionStart-2="FIGURES 54–57" captionStartId-0="13.[151,264,1340,1364]" captionStartId-1="15.[151,264,1813,1837]" captionStartId-2="16.[151,264,1581,1605]" captionTargetBox-0="[155,1429,241,1309]" captionTargetBox-1="[166,1435,189,1784]" captionTargetBox-2="[151,1435,181,1557]" captionTargetId-0="figure-45@13.[151,1435,237,1317]" captionTargetId-1="figure-19@15.[151,1435,181,1788]" captionTargetId-2="figure-30@16.[151,1435,181,1557]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURES 42–45. Habitus of male (42–43) and female (44–45) of Bowie shakeit Jäger. 42, 44—dorsal; 43, 45—ventral. Scale bars: 2 mm." captionText-1="FIGURES 46–53. Palp (46–49) and bulb (50–53) of male of Bowie shakeit Jäger. 46, 50—prolateral; 47, 51—ventral; 48, 52— retrolateral; 49—dorsal; 53—apical. Scale bars: 46–49 = 0.5 mm; 50–53 = 0.2 mm.Abbreviations: C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." captionText-2="FIGURES 54–57. Epigyne of Bowie shakeit Jäger. 54—intact, ventral; 55—macerated, ventral; 56—internal genitalia, dorsal; 57—ditto, posterior. Scale bar: 0.2 mm.Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi-0="http://doi.org/10.5281/zenodo.14386417" figureDoi-1="http://doi.org/10.5281/zenodo.14386421" figureDoi-2="http://doi.org/10.5281/zenodo.14386423" httpUri-0="https://zenodo.org/record/14386417/files/figure.png" httpUri-1="https://zenodo.org/record/14386421/files/figure.png" httpUri-2="https://zenodo.org/record/14386423/files/figure.png" pageId="12" pageNumber="395">Figs 42–57</figureCitation>
,
<figureCitation id="A296B49A376FFF95681BFAE1031C9AC0" box="[300,371,1392,1418]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="12" pageNumber="395">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF9569A0FA2401159A87" box="[151,890,1461,1485]" pageId="12" pageNumber="395" type="reference_group">
<paragraph id="3A12A81F376FFF9569A0FA2401159A87" blockId="12.[151,890,1461,1485]" box="[151,890,1461,1485]" pageId="12" pageNumber="395">
<treatmentCitationGroup id="1ABD8F31376FFF9569A0FA2401159A87" box="[151,890,1461,1485]" pageId="12" pageNumber="395">
<treatmentCitation id="BB0C8E0E376FFF9569A0FA2403C49A87" author="Jager, P." box="[151,427,1461,1485]" page="165" pageId="12" pageNumber="395" year="2022">
<taxonomicName id="FDADD39C376FFF9569A0FA2403C49A87" ID-CoL="C6LK8" authority="Jager, 2022: 165" authorityName="Jager" authorityPageNumber="165" authorityYear="2022" box="[151,427,1461,1485]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376FFF9569A0FA2402949A87" box="[151,251,1461,1485]" italics="true" pageId="12" pageNumber="395">B. shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376FFF956836FA2403C49A87" author="Jager, P." box="[257,427,1461,1485]" pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 745–748, 775–780 (
<specimenCount id="2CAB6396376FFF956B98FA2400AA9A87" box="[687,709,1461,1485]" count="1" pageId="12" pageNumber="395" type="male">♂</specimenCount>
<specimenCount id="2CAB6396376FFF956BF2FA2400BB9A87" box="[709,724,1461,1485]" count="1" pageId="12" pageNumber="395" type="female">♀</specimenCount>
), not examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF9569A0FA6A01589973" pageId="12" pageNumber="395" type="materials_examined">
<paragraph id="3A12A81F376FFF9569A0FA6A01589973" blockId="12.[151,1436,1530,2025]" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569A0FA6A03EE995F" bold="true" box="[151,385,1531,1557]" pageId="12" pageNumber="395">Material examined.</emphasis>
<materialsCitation id="8AC5A242376FFF9568BFFA6A01589973" collectingDate="1994-08-30" collectingDateMax="1994-09-07" collectingDateMin="1994-08-30" collectionCode="IBSS" collectorName="A. M. Emelyanov" country="Malaysia" elevation="300" latitude="4.4" location="Tawau" longLatPrecision="1306" longitude="117.88333" municipality="Borneo" pageId="12" pageNumber="395" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah">
<emphasis id="08D9740D376FFF9568BFFA6A00AE995F" bold="true" box="[392,705,1530,1557]" pageId="12" pageNumber="395">
<collectingCountry id="42BAE88F376FFF9568BFFA6A004D995F" box="[392,546,1531,1557]" name="Malaysia" pageId="12" pageNumber="395">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D376FFF956B19FA6B00D7995F" bold="true" box="[558,696,1530,1557]" italics="true" pageId="12" pageNumber="395">
<collectingRegion id="F86966FD376FFF956B19FA6B00D7995F" box="[558,696,1530,1557]" country="Malaysia" name="Sabah" pageId="12" pageNumber="395">
<taxonomicName id="FDADD39C376FFF956B19FA6B0018995F" authorityName="Li &amp; Yao" authorityYear="2022" box="[558,631,1530,1557]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="2CAB6396376FFF956BFFFA6A0081995F" box="[712,750,1531,1557]" count="2" pageId="12" pageNumber="395" type="male">2♂</specimenCount>
,
<specimenCount id="2CAB6396376FFF956BCDFA6A0173995F" box="[762,796,1531,1557]" count="1" pageId="12" pageNumber="395" type="female">1♀</specimenCount>
(
<collectionCode id="5CBC30DA376FFF956A1BFA6A0103995F" box="[812,876,1531,1557]" pageId="12" pageNumber="395">IBSS</collectionCode>
),
<collectingMunicipality id="DA763265376FFF956AB7FA6A01BA995F" box="[896,981,1531,1557]" pageId="12" pageNumber="395">Borneo</collectingMunicipality>
, environs of
<location id="3F72FEC4376FFF956D51FA6A06DF995F" LSID="urn:lsid:plazi:treatment:B2041909376FFF8969A0FADD010F98BF:3F72FEC4376FFF956D51FA6A06DF995F" box="[1126,1200,1531,1557]" country="Malaysia" latitude="4.4" longLatPrecision="1306" longitude="117.88333" municipality="Borneo" name="Tawau" pageId="12" pageNumber="395" stateProvince="Sabah">Tawau</location>
[
<geoCoordinate id="5F99CED8376FFF956D89FA6A077A995F" box="[1214,1301,1531,1557]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="12" pageNumber="395" precision="925" value="4.4">4°24’N</geoCoordinate>
,
<geoCoordinate id="5F99CED8376FFF956C16FA6A07FF995C" box="[1313,1424,1531,1558]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="12" pageNumber="395" precision="925" value="117.88333">117°53’E</geoCoordinate>
],
<quantity id="FD5505FA376FFF9569A0F98E028F9973" box="[151,224,1567,1593]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="395" unit="m" value="300.0">
<elevation id="B1804F2C376FFF9569A0F98E028F9973" box="[151,224,1567,1593]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="395" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="4E138EDF376FFF9569DBF98E00549973" box="[236,571,1567,1593]" pageId="12" pageNumber="395" value="1994-08-30" valueMax="1994-09-07" valueMin="1994-08-30">
<collectingDate id="5E577737376FFF9569DBF98E00549973" box="[236,571,1567,1593]" pageId="12" pageNumber="395" value="1994-08-30" valueMax="1994-09-07" valueMin="1994-08-30">30 August–7 September 1994</collectingDate>
</date>
,
<collectorName id="9758CDC9376FFF956B72F98E01689973" box="[581,775,1567,1593]" pageId="12" pageNumber="395">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF9569F0F9D20286987F" pageId="12" pageNumber="395" type="diagnosis">
<paragraph id="3A12A81F376FFF9569F0F9D20286987F" blockId="12.[151,1436,1530,2025]" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569F0F9D203559917" bold="true" box="[199,314,1603,1629]" pageId="12" pageNumber="395">Diagnosis</emphasis>
. Males of
<taxonomicName id="FDADD39C376FFF956886F9D500229917" box="[433,589,1603,1629]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376FFF956886F9D500229917" box="[433,589,1603,1629]" italics="true" pageId="12" pageNumber="395">Bowie shakeit</emphasis>
</taxonomicName>
closely resemble those of
<taxonomicName id="FDADD39C376FFF956A41F9D5065A9917" authorityName="Jager" authorityYear="2022" box="[886,1077,1603,1629]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="criminalworld">
<emphasis id="08D9740D376FFF956A41F9D5065A9917" box="[886,1077,1603,1629]" italics="true" pageId="12" pageNumber="395">B. criminalworld</emphasis>
</taxonomicName>
by the shape and size of
<emphasis id="08D9740D376FFF956C79F9D207119917" box="[1358,1406,1603,1629]" italics="true" pageId="12" pageNumber="395">RTA</emphasis>
as well as tegular apophysis (
<emphasis id="08D9740D376FFF9568F1F9F6038A99CB" box="[454,485,1639,1665]" italics="true" pageId="12" pageNumber="395">TA</emphasis>
) but can be distinguished by 1) retro-proximal cymbial outgrowth (
<emphasis id="08D9740D376FFF956DD1F9F6077399CB" box="[1254,1308,1639,1665]" italics="true" pageId="12" pageNumber="395">RPO</emphasis>
) extending
<emphasis id="08D9740D376FFF9569A0F91A02A899EF" box="[151,199,1675,1701]" italics="true" pageId="12" pageNumber="395">RTA</emphasis>
in ventral view (vs. not extending; cf.
<figureCitation id="A296B49A376FFF956B41F91A00AA99EF" box="[630,709,1675,1701]" captionStart="FIGURES 46–53" captionStartId="15.[151,264,1813,1837]" captionTargetBox="[166,1435,189,1784]" captionTargetId="figure-19@15.[151,1435,181,1788]" captionTargetPageId="15" captionText="FIGURES 46–53. Palp (46–49) and bulb (50–53) of male of Bowie shakeit Jäger. 46, 50—prolateral; 47, 51—ventral; 48, 52— retrolateral; 49—dorsal; 53—apical. Scale bars: 46–49 = 0.5 mm; 50–53 = 0.2 mm.Abbreviations: C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386421" httpUri="https://zenodo.org/record/14386421/files/figure.png" pageId="12" pageNumber="395">Fig. 47</figureCitation>
and
<bibRefCitation id="5E3CD5EE376FFF956BCDF91A011799EF" author="Jager, P." box="[762,888,1675,1701]" pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 769), 2) tegular hump (
<emphasis id="08D9740D376FFF956D85F91D06B999EF" box="[1202,1238,1676,1701]" italics="true" pageId="12" pageNumber="395">TH</emphasis>
) touching tegular apophysis (
<emphasis id="08D9740D376FFF95682EF93E03579983" box="[281,312,1711,1737]" italics="true" pageId="12" pageNumber="395">TA</emphasis>
) (vs. not touching; cf.
<figureCitation id="A296B49A376FFF956B08F93E00FF9980" box="[575,656,1711,1738]" captionStart="FIGURES 46–53" captionStartId="15.[151,264,1813,1837]" captionTargetBox="[166,1435,189,1784]" captionTargetId="figure-19@15.[151,1435,181,1788]" captionTargetPageId="15" captionText="FIGURES 46–53. Palp (46–49) and bulb (50–53) of male of Bowie shakeit Jäger. 46, 50—prolateral; 47, 51—ventral; 48, 52— retrolateral; 49—dorsal; 53—apical. Scale bars: 46–49 = 0.5 mm; 50–53 = 0.2 mm.Abbreviations: C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386421" httpUri="https://zenodo.org/record/14386421/files/figure.png" pageId="12" pageNumber="395">Fig. 51</figureCitation>
and
<bibRefCitation id="5E3CD5EE376FFF956BFDF93E01269983" author="Jager, P." box="[714,841,1711,1737]" pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 769). Females of
<taxonomicName id="FDADD39C376FFF956D7AF92106849983" box="[1101,1259,1711,1737]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376FFF956D7AF92106849983" box="[1101,1259,1711,1737]" italics="true" pageId="12" pageNumber="395">Bowie shakeit</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="FDADD39C376FFF95698FF945031699A7" authorityName="Jager" authorityYear="2022" box="[184,377,1747,1773]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="395" phylum="Arthropoda" rank="species" species="criminalworld">
<emphasis id="08D9740D376FFF95698FF945031699A7" box="[184,377,1747,1773]" italics="true" pageId="12" pageNumber="395">B. criminalworld</emphasis>
</taxonomicName>
by the shape of median plate (
<emphasis id="08D9740D376FFF956BD6F945016699A7" box="[737,777,1748,1773]" italics="true" pageId="12" pageNumber="395">MP</emphasis>
) but can be easily distinguished by widely spaced vulval folds (
<emphasis id="08D9740D376FFF9569D7F969036D985B" box="[224,258,1784,1809]" italics="true" pageId="12" pageNumber="395">VF</emphasis>
) (vs. almost touching each other; cf.
<figureCitation id="A296B49A376FFF956B9FF96600979858" box="[680,760,1783,1810]" captionStart="FIGURES 54–57" captionStartId="16.[151,264,1581,1605]" captionTargetBox="[151,1435,181,1557]" captionTargetId="figure-30@16.[151,1435,181,1557]" captionTargetPageId="16" captionText="FIGURES 54–57. Epigyne of Bowie shakeit Jäger. 54—intact, ventral; 55—macerated, ventral; 56—internal genitalia, dorsal; 57—ditto, posterior. Scale bar: 0.2 mm.Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386423" httpUri="https://zenodo.org/record/14386423/files/figure.png" pageId="12" pageNumber="395">Fig. 56</figureCitation>
and
<bibRefCitation id="5E3CD5EE376FFF956A07F96601C1985B" author="Jager, P." box="[816,942,1783,1809]" pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger 2022</bibRefCitation>
: fig. 774). For extended diagnosis see
<bibRefCitation id="5E3CD5EE376FFF956C56F966028A987F" author="Jager, P." pageId="12" pageNumber="395" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB94376FFF8969F0F8AE02A0989B" lastPageId="16" lastPageNumber="399" pageId="12" pageNumber="395" type="description">
<paragraph id="3A12A81F376FFF9469F0F8AE01A29F9F" blockId="12.[151,1436,1530,2025]" lastBlockId="13.[151,1437,151,213]" lastPageId="13" lastPageNumber="396" pageId="12" pageNumber="395">
<emphasis id="08D9740D376FFF9569F0F8AE03369813" bold="true" box="[199,345,1855,1881]" pageId="12" pageNumber="395">Description.</emphasis>
<emphasis id="08D9740D376FFF956855F8AE03F39813" box="[354,412,1855,1881]" italics="true" pageId="12" pageNumber="395">Male</emphasis>
(
<figureCitation id="A296B49A376FFF95689BF8AE005D9813" box="[428,562,1855,1881]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1340,1364]" captionTargetBox="[155,1429,241,1309]" captionTargetId="figure-45@13.[151,1435,237,1317]" captionTargetPageId="13" captionText="FIGURES 42–45. Habitus of male (42–43) and female (44–45) of Bowie shakeit Jäger. 42, 44—dorsal; 43, 45—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386417" httpUri="https://zenodo.org/record/14386417/files/figure.png" pageId="12" pageNumber="395">Figs 42–43</figureCitation>
). Total length 16.20. Prosoma 9.03 long, 6.68 wide. Opisthosoma 6.90 long, 4.30 wide. Prosoma dark brown, with light brown median band, widened and bearing two dark spots nearly ocular area. Lateral bands absent. Fovea thin, black. Clypeus light brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum light brown, without pattern. Labium dark brown. Endites dark brown with yellow distal edge. Dorsal part of opisthosoma yellowish with poorly visible cardiac mark and series of black spots. Lateral sides of opisthosoma greenish yellow. Ventral part of opisthosoma dark gray with yellow spots forming single V-mark. Spinnerets yellow ventrally, dark brown laterally.
</paragraph>
<caption id="6ED2F897376EFF9469A0FAAD037F9A3D" ID-DOI="http://doi.org/10.5281/zenodo.14386417" ID-Zenodo-Dep="14386417" httpUri="https://zenodo.org/record/14386417/files/figure.png" pageId="13" pageNumber="396" startId="13.[151,264,1340,1364]" targetBox="[155,1429,241,1309]" targetPageId="13" targetType="figure">
<paragraph id="3A12A81F376EFF9469A0FAAD037F9A3D" blockId="13.[151,1436,1340,1400]" pageId="13" pageNumber="396">
<emphasis id="08D9740D376EFF9469A0FAAD033B9A1E" bold="true" box="[151,340,1340,1364]" pageId="13" pageNumber="396">FIGURES 42–45.</emphasis>
Habitus of male (42–43) and female (44–45) of
<taxonomicName id="FDADD39C376EFF946A76FAAC067F9A1E" authority="Jager." authorityName="Jager." box="[833,1040,1340,1364]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376EFF946A76FAAC01A09A1E" box="[833,975,1340,1364]" italics="true" pageId="13" pageNumber="396">Bowie shakeit</emphasis>
Jäger.
</taxonomicName>
42, 44—dorsal; 43, 45—ventral. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="3A12A81F376EFF9469F0FA5A07EF9943" blockId="13.[151,1437,1482,2013]" pageId="13" pageNumber="396">Eye diameters: AME 0.38, ALE 0.24, PME 0.47, PLE 0.37; interdistances: AME–AME 0.17, AME–ALE 0.41, PME–PME 0.23, PME–PLE 0.46, AME–PME 0.09, ALE–PLE 0.19. Clypeus height at AME 0.29, at ALE 0.75.</paragraph>
<paragraph id="3A12A81F376EFF9469F0F98206D199D2" blockId="13.[151,1437,1482,2013]" pageId="13" pageNumber="396">
For palp and legs measurements see
<tableCitation id="772F9DA4376EFF946B5EF98300A29966" box="[617,717,1554,1580]" captionStart="TABLE 11" captionStartId="14.[151,239,548,572]" captionTargetPageId="14" captionText="TABLE 11. Palp and legs measurements of male of Bowie shakeit Jäger, 2022" pageId="13" pageNumber="396">Table 11</tableCitation>
. Palp coloration: femora, patellae, tibiae brown; cymbium dark brown. Legs coloration: femora I–IV brown with dark brown blurry spots dorsally, brown laterally, light brown ventrally. Patellae I–IV brown. Tibiae I–II brown proximally, light brown with short white setae distally; III–IV brown. Metatarsi I–IV dark brown. Tarsi I–IV brown. For palp and legs spination see
<tableCitation id="772F9DA4376EFF946D60F9EF06D599D2" box="[1111,1210,1662,1688]" captionStart="TABLE 12" captionStartId="14.[151,239,899,923]" captionTargetPageId="14" captionText="TABLE 12. Palp and legs spination for the male of Bowie shakeit Jäger, 2022" pageId="13" pageNumber="396">Table 12</tableCitation>
.
</paragraph>
<paragraph id="3A12A81F376EFF9469F0F933037A9806" blockId="13.[151,1437,1482,2013]" pageId="13" pageNumber="396">
Palp as shown in
<figureCitation id="A296B49A376EFF9468A5F933007B99F7" box="[402,532,1698,1725]" captionStart="FIGURES 46–53" captionStartId="15.[151,264,1813,1837]" captionTargetBox="[166,1435,189,1784]" captionTargetId="figure-19@15.[151,1435,181,1788]" captionTargetPageId="15" captionText="FIGURES 46–53. Palp (46–49) and bulb (50–53) of male of Bowie shakeit Jäger. 46, 50—prolateral; 47, 51—ventral; 48, 52— retrolateral; 49—dorsal; 53—apical. Scale bars: 46–49 = 0.5 mm; 50–53 = 0.2 mm.Abbreviations: C—conductor, EH—embolic hook, Em—embolus, RPO—retro-proximal cymbial outgrowth, RTA—retrolateral tibial apophysis, TA—tegular apophysis, Te—tegulum, TH—tegular hump." figureDoi="http://doi.org/10.5281/zenodo.14386421" httpUri="https://zenodo.org/record/14386421/files/figure.png" pageId="13" pageNumber="396">Figs 46–53</figureCitation>
.
<emphasis id="08D9740D376EFF946B17F932003F99F7" box="[544,592,1699,1725]" italics="true" pageId="13" pageNumber="396">RTA</emphasis>
arising from the middle of tibia, short, curved, with pointed tip in ventral view. Cymbium with large, blunt retro-proximal cymbial outgrowth (
<emphasis id="08D9740D376EFF946AA0F95601A299AB" box="[919,973,1735,1761]" italics="true" pageId="13" pageNumber="396">RPO</emphasis>
). Tegulum (
<emphasis id="08D9740D376EFF946D61F956060099AA" box="[1110,1135,1735,1760]" italics="true" pageId="13" pageNumber="396">Te</emphasis>
) with rounded hump (
<emphasis id="08D9740D376EFF946C5FF95607E399AA" box="[1384,1420,1735,1760]" italics="true" pageId="13" pageNumber="396">TH</emphasis>
). Tegular apophysis (
<emphasis id="08D9740D376EFF94684FF97A03F8984F" box="[376,407,1771,1797]" italics="true" pageId="13" pageNumber="396">TA</emphasis>
) oval, originating in a 5.30 o’clock position. Conductor (
<emphasis id="08D9740D376EFF946D1CF97A0651984F" box="[1067,1086,1771,1797]" italics="true" pageId="13" pageNumber="396">C</emphasis>
) large, membranous. Embolus (
<emphasis id="08D9740D376EFF946996F89E02A99862" box="[161,198,1807,1832]" italics="true" pageId="13" pageNumber="396">Em</emphasis>
) originating from tegulum in 7-o’clock-position, short, with pointed tip. Embolus with large basal embolic hook (
<emphasis id="08D9740D376EFF9469D7F8A2036A9806" box="[224,261,1843,1868]" italics="true" pageId="13" pageNumber="396">EH</emphasis>
).
</paragraph>
<paragraph id="3A12A81F376EFF9769F0F8C706489FB3" blockId="13.[151,1437,1482,2013]" lastBlockId="14.[151,1436,151,465]" lastPageId="14" lastPageNumber="397" pageId="13" pageNumber="396">
<emphasis id="08D9740D376EFF9469F0F8C7034C983A" box="[199,291,1878,1904]" italics="true" pageId="13" pageNumber="396">Female.</emphasis>
(
<figureCitation id="A296B49A376EFF946805F8C703D8983B" box="[306,439,1878,1905]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1340,1364]" captionTargetBox="[155,1429,241,1309]" captionTargetId="figure-45@13.[151,1435,237,1317]" captionTargetPageId="13" captionText="FIGURES 42–45. Habitus of male (42–43) and female (44–45) of Bowie shakeit Jäger. 42, 44—dorsal; 43, 45—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386417" httpUri="https://zenodo.org/record/14386417/files/figure.png" pageId="13" pageNumber="396">Figs 44–45</figureCitation>
). Total length 13.99. Prosoma 7.62 long, 5.93 wide. Opisthosoma 6.46 long, 3.92 wide. Prosoma brown, with light brown median band, widened and bearing two dark spots nearly ocular area. Lateral bands poorly visible. Fovea thin, black. Clypeus light brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum light brown, without pattern. Labium dark brown. Endites brown with yellow distal edge. Dorsal part of opisthosoma dark gray with distinct yellowish cardiac mark and series of dark brown transversal stripes. Lateral sides of opisthosoma dark gray. Ventral part of opisthosoma dark gray with yellow spots forming single V-marks. Spinnerets yellow with brown lateral sides.
</paragraph>
<paragraph id="3A12A81F376DFF9769F0FE9207109E0B" blockId="14.[151,1436,151,465]" pageId="14" pageNumber="397">Eye diameters: AME 0.37, ALE 0.23, PME 0.44, PLE 0.41; interdistances: AME–AME 0.22, AME–ALE 0.46, PME–PME 0.24, PME–PLE 0.52, AME–PME 0.11, ALE–PLE 0.15. Clypeus height at AME 0.30, at ALE 0.71.</paragraph>
<paragraph id="3A12A81F376DFF9769F0FEDA03959E9B" blockId="14.[151,1436,151,465]" pageId="14" pageNumber="397">
For palp and legs measurements see
<tableCitation id="772F9DA4376DFF976B51FEDA00A49E2F" box="[614,715,331,357]" captionStart="TABLE 13" captionStartId="14.[151,239,1250,1274]" captionTargetPageId="14" captionText="TABLE 13. Palp and legs measurements of female of Bowie shakeit Jäger, 2022" pageId="14" pageNumber="397">Table 13</tableCitation>
. Palp coloration: all joints light brown. Legs coloration: femora I–IV light brown. Patellae I–IV light brown. Tibiae I–II light brown; III– IV light brown with poorly visible semi-rings. Metatarsi I–II light brown; III–IV light brown with poorly visible semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA4376DFF9768A4FE2603999E9B" box="[403,502,439,465]" captionStart="TABLE 14" captionStartId="14.[151,239,1601,1625]" captionTargetPageId="14" captionText="TABLE 14. Palp and legs spination for the female of Bowie shakeit Jäger, 2022" pageId="14" pageNumber="397">Table 14</tableCitation>
.
</paragraph>
<caption id="6ED2F897376DFF9769A0FDB501B59D77" box="[151,986,548,573]" pageId="14" pageNumber="397" startId="14.[151,239,548,572]" targetBox="[163,1370,589,814]" targetIsTable="true" targetPageId="14" targetType="table">
<paragraph id="3A12A81F376DFF9769A0FDB501B59D77" blockId="14.[151,986,548,573]" box="[151,986,548,573]" pageId="14" pageNumber="397">
<emphasis id="08D9740D376DFF9769A0FDB503799D76" bold="true" box="[151,278,548,572]" pageId="14" pageNumber="397">TABLE 11.</emphasis>
Palp and legs measurements of male of
<taxonomicName id="FDADD39C376DFF976BF2FDB401B59D77" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[709,986,548,573]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376DFF976BF2FDB401359D76" box="[709,858,548,572]" italics="true" pageId="14" pageNumber="397">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376DFF976A56FDB401B59D77" author="Jager, P." box="[865,986,549,573]" pageId="14" pageNumber="397" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F376DFF97684EFDDC07359C64" pageId="14" pageNumber="397">
<table id="48AD5ABF376D00666994FDDC07359C64" box="[163,1370,589,814]" gridcols="7" gridrows="6" pageId="14" pageNumber="397">
<tr id="849DAA5D376D00666994FDDC07359D2F" box="[163,1370,589,613]" gridrow="0" pageId="14" pageNumber="397" rowspan-0="1">
<th id="C74CC321376D00666858FDDC03F49D2F" box="[367,411,589,613]" gridcol="1" gridrow="0" pageId="14" pageNumber="397">Fe</th>
<th id="C74CC321376D00666B1AFDDC00369D2F" box="[557,601,589,613]" gridcol="2" gridrow="0" pageId="14" pageNumber="397">Pa</th>
<th id="C74CC321376D00666BDBFDDC01779D2F" box="[748,792,589,613]" gridcol="3" gridrow="0" pageId="14" pageNumber="397">Ti</th>
<th id="C74CC321376D00666A9DFDDC01B99D2F" box="[938,982,589,613]" gridcol="4" gridrow="0" pageId="14" pageNumber="397">Mt</th>
<th id="C74CC321376D00666D5EFDDC06FA9D2F" box="[1129,1173,589,613]" gridcol="5" gridrow="0" pageId="14" pageNumber="397">Ta</th>
<th id="C74CC321376D00666C16FDDC07359D2F" box="[1313,1370,589,613]" gridcol="6" gridrow="0" pageId="14" pageNumber="397">Total</th>
</tr>
<tr id="849DAA5D376D00666994FDE407359DC7" box="[163,1370,629,653]" gridrow="1" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FDE402BF9DC7" box="[163,208,629,653]" gridcol="0" gridrow="1" pageId="14" pageNumber="397">Palp</th>
<td id="C74CC321376D00666858FDE403F49DC7" box="[367,411,629,653]" gridcol="1" gridrow="1" pageId="14" pageNumber="397">3.51</td>
<td id="C74CC321376D00666B1AFDE400369DC7" box="[557,601,629,653]" gridcol="2" gridrow="1" pageId="14" pageNumber="397">1.42</td>
<td id="C74CC321376D00666BDBFDE401779DC7" box="[748,792,629,653]" gridcol="3" gridrow="1" pageId="14" pageNumber="397">1.61</td>
<td id="C74CC321376D00666A9DFDE401B99DC7" box="[938,982,629,653]" gridcol="4" gridrow="1" pageId="14" pageNumber="397">0</td>
<td id="C74CC321376D00666D5EFDE406FA9DC7" box="[1129,1173,629,653]" gridcol="5" gridrow="1" pageId="14" pageNumber="397">2.82</td>
<td id="C74CC321376D00666C16FDE407359DC7" box="[1313,1370,629,653]" gridcol="6" gridrow="1" pageId="14" pageNumber="397">9.36</td>
</tr>
<tr id="849DAA5D376D00666994FD0C07359DFF" box="[163,1370,669,693]" gridrow="2" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FD0C02BF9DFF" box="[163,208,669,693]" gridcol="0" gridrow="2" pageId="14" pageNumber="397">I</th>
<td id="C74CC321376D00666858FD0C03F49DFF" box="[367,411,669,693]" gridcol="1" gridrow="2" pageId="14" pageNumber="397">7.25</td>
<td id="C74CC321376D00666B1AFD0C00369DFF" box="[557,601,669,693]" gridcol="2" gridrow="2" pageId="14" pageNumber="397">3.45</td>
<td id="C74CC321376D00666BDBFD0C01779DFF" box="[748,792,669,693]" gridcol="3" gridrow="2" pageId="14" pageNumber="397">6.93</td>
<td id="C74CC321376D00666A9DFD0C01B99DFF" box="[938,982,669,693]" gridcol="4" gridrow="2" pageId="14" pageNumber="397">6.24</td>
<td id="C74CC321376D00666D5EFD0C06FA9DFF" box="[1129,1173,669,693]" gridcol="5" gridrow="2" pageId="14" pageNumber="397">1.93</td>
<td id="C74CC321376D00666C16FD0C07359DFF" box="[1313,1370,669,693]" gridcol="6" gridrow="2" pageId="14" pageNumber="397">25.8</td>
</tr>
<tr id="849DAA5D376D00666994FD5407359D94" box="[163,1370,709,734]" gridrow="3" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FD5402BF9D94" box="[163,208,709,734]" gridcol="0" gridrow="3" pageId="14" pageNumber="397">II</th>
<td id="C74CC321376D00666858FD5403F49D94" box="[367,411,709,734]" gridcol="1" gridrow="3" pageId="14" pageNumber="397">6.51</td>
<td id="C74CC321376D00666B1AFD5400369D94" box="[557,601,709,734]" gridcol="2" gridrow="3" pageId="14" pageNumber="397">3.05</td>
<td id="C74CC321376D00666BDBFD5401779D94" box="[748,792,709,734]" gridcol="3" gridrow="3" pageId="14" pageNumber="397">5.72</td>
<td id="C74CC321376D00666A9DFD5401B99D94" box="[938,982,709,734]" gridcol="4" gridrow="3" pageId="14" pageNumber="397">5.26</td>
<td id="C74CC321376D00666D5EFD5406FA9D94" box="[1129,1173,709,734]" gridcol="5" gridrow="3" pageId="14" pageNumber="397">1.73</td>
<td id="C74CC321376D00666C16FD5407359D94" box="[1313,1370,709,734]" gridcol="6" gridrow="3" pageId="14" pageNumber="397">22.27</td>
</tr>
<tr id="849DAA5D376D00666994FD7C07359C4C" box="[163,1370,749,774]" gridrow="4" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FD7C02BF9C4C" box="[163,208,749,774]" gridcol="0" gridrow="4" pageId="14" pageNumber="397">III</th>
<td id="C74CC321376D00666858FD7C03F49C4C" box="[367,411,749,774]" gridcol="1" gridrow="4" pageId="14" pageNumber="397">5.84</td>
<td id="C74CC321376D00666B1AFD7C00369C4C" box="[557,601,749,774]" gridcol="2" gridrow="4" pageId="14" pageNumber="397">2.35</td>
<td id="C74CC321376D00666BDBFD7C01779C4C" box="[748,792,749,774]" gridcol="3" gridrow="4" pageId="14" pageNumber="397">4.54</td>
<td id="C74CC321376D00666A9DFD7C01B99C4C" box="[938,982,749,774]" gridcol="4" gridrow="4" pageId="14" pageNumber="397">4.75</td>
<td id="C74CC321376D00666D5EFD7C06FA9C4C" box="[1129,1173,749,774]" gridcol="5" gridrow="4" pageId="14" pageNumber="397">1.50</td>
<td id="C74CC321376D00666C16FD7C07359C4C" box="[1313,1370,749,774]" gridcol="6" gridrow="4" pageId="14" pageNumber="397">18.98</td>
</tr>
<tr id="849DAA5D376D00666994FC8707359C64" box="[163,1370,790,814]" gridrow="5" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FC8702BF9C64" box="[163,208,790,814]" gridcol="0" gridrow="5" pageId="14" pageNumber="397">IV</th>
<td id="C74CC321376D00666858FC8703F49C64" box="[367,411,790,814]" gridcol="1" gridrow="5" pageId="14" pageNumber="397">7.68</td>
<td id="C74CC321376D00666B1AFC8700369C64" box="[557,601,790,814]" gridcol="2" gridrow="5" pageId="14" pageNumber="397">2.94</td>
<td id="C74CC321376D00666BDBFC8701779C64" box="[748,792,790,814]" gridcol="3" gridrow="5" pageId="14" pageNumber="397">6.77</td>
<td id="C74CC321376D00666A9DFC8701B99C64" box="[938,982,790,814]" gridcol="4" gridrow="5" pageId="14" pageNumber="397">9.07</td>
<td id="C74CC321376D00666D5EFC8706FA9C64" box="[1129,1173,790,814]" gridcol="5" gridrow="5" pageId="14" pageNumber="397">2.06</td>
<td id="C74CC321376D00666C16FC8707359C64" box="[1313,1370,790,814]" gridcol="6" gridrow="5" pageId="14" pageNumber="397">28.52</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376DFF9769A0FC1201BA9CD1" box="[151,981,899,924]" pageId="14" pageNumber="397" startId="14.[151,239,899,923]" targetBox="[163,1385,939,1165]" targetIsTable="true" targetPageId="14" targetType="table">
<paragraph id="3A12A81F376DFF9769A0FC1201BA9CD1" blockId="14.[151,981,899,924]" box="[151,981,899,924]" pageId="14" pageNumber="397">
<emphasis id="08D9740D376DFF9769A0FC1203789CD1" bold="true" box="[151,279,899,923]" pageId="14" pageNumber="397">TABLE 12.</emphasis>
Palp and legs spination for the male of
<taxonomicName id="FDADD39C376DFF976BF7FC1501BA9CD1" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[704,981,899,924]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376DFF976BF7FC15013A9CD1" box="[704,853,899,923]" italics="true" pageId="14" pageNumber="397">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376DFF976A6CFC1501BA9CD1" author="Jager, P." box="[859,981,899,924]" pageId="14" pageNumber="397" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F376DFF9768B8FC3D07069BC7" pageId="14" pageNumber="397">
<table id="48AD5ABF376D00666994FC3A07069BC7" box="[163,1385,939,1165]" gridcols="5" gridrows="6" pageId="14" pageNumber="397">
<tr id="849DAA5D376D00666994FC3A07069C89" box="[163,1385,939,963]" gridrow="0" pageId="14" pageNumber="397" rowspan-0="1">
<th id="C74CC321376D00666846FC3A03A99C89" box="[369,454,939,963]" gridcol="1" gridrow="0" pageId="14" pageNumber="397">Fe</th>
<th id="C74CC321376D00666B91FC3A00B59C89" box="[678,730,939,963]" gridcol="2" gridrow="0" pageId="14" pageNumber="397">Pa</th>
<th id="C74CC321376D00666A4BFC3A06219C89" box="[892,1102,939,963]" gridcol="3" gridrow="0" pageId="14" pageNumber="397">Ti</th>
<th id="C74CC321376D00666D9CFC3A07069C89" box="[1195,1385,939,963]" gridcol="4" gridrow="0" pageId="14" pageNumber="397">Mt</th>
</tr>
<tr id="849DAA5D376D00666994FC4507069CA6" box="[163,1385,980,1004]" gridrow="1" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FC4502BF9CA6" box="[163,208,980,1004]" gridcol="0" gridrow="1" pageId="14" pageNumber="397">Palp</th>
<td id="C74CC321376D00666846FC4503A99CA6" box="[369,454,980,1004]" gridcol="1" gridrow="1" pageId="14" pageNumber="397">4d 1p 1r</td>
<td id="C74CC321376D00666B91FC4500B59CA6" box="[678,730,980,1004]" gridcol="2" gridrow="1" pageId="14" pageNumber="397">1p</td>
<td id="C74CC321376D00666A4BFC4506219CA6" box="[892,1102,980,1004]" gridcol="3" gridrow="1" pageId="14" pageNumber="397">2p 1r</td>
<td id="C74CC321376D00666D9CFC4507069CA6" box="[1195,1385,980,1004]" gridcol="4" gridrow="1" pageId="14" pageNumber="397">-</td>
</tr>
<tr id="849DAA5D376D00666994FC6D07069B5E" box="[163,1385,1020,1044]" gridrow="2" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FC6D02BF9B5E" box="[163,208,1020,1044]" gridcol="0" gridrow="2" pageId="14" pageNumber="397">I</th>
<td id="C74CC321376D00666846FC6D03A99B5E" box="[369,454,1020,1044]" gridcol="1" gridrow="2" pageId="14" pageNumber="397">3d 3p 4r</td>
<td id="C74CC321376D00666B91FC6D00B59B5E" box="[678,730,1020,1044]" gridcol="2" gridrow="2" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666A4BFC6D06219B5E" box="[892,1102,1020,1044]" gridcol="3" gridrow="2" pageId="14" pageNumber="397">3d 2p 2(3)r 2-2-2-2v</td>
<td id="C74CC321376D00666D9CFC6D07069B5E" box="[1195,1385,1020,1044]" gridcol="4" gridrow="2" pageId="14" pageNumber="397">1d 3p 3r 2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994FBB507069B76" box="[163,1385,1060,1084]" gridrow="3" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FBB502BF9B76" box="[163,208,1060,1084]" gridcol="0" gridrow="3" pageId="14" pageNumber="397">II</th>
<td id="C74CC321376D00666846FBB503A99B76" box="[369,454,1060,1084]" gridcol="1" gridrow="3" pageId="14" pageNumber="397">3d 4p 4r</td>
<td id="C74CC321376D00666B91FBB500B59B76" box="[678,730,1060,1084]" gridcol="2" gridrow="3" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666A4BFBB506219B76" box="[892,1102,1060,1084]" gridcol="3" gridrow="3" pageId="14" pageNumber="397">3d 3p 3r 2-2-2-2v</td>
<td id="C74CC321376D00666D9CFBB507069B76" box="[1195,1385,1060,1084]" gridcol="4" gridrow="3" pageId="14" pageNumber="397">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994FBDD07069B2F" box="[163,1385,1100,1125]" gridrow="4" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FBDD02BF9B2F" box="[163,208,1100,1125]" gridcol="0" gridrow="4" pageId="14" pageNumber="397">III</th>
<td id="C74CC321376D00666846FBDD03A99B2F" box="[369,454,1100,1125]" gridcol="1" gridrow="4" pageId="14" pageNumber="397">3d 4p 4r</td>
<td id="C74CC321376D00666B91FBDD00B59B2F" box="[678,730,1100,1125]" gridcol="2" gridrow="4" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666A4BFBDD06219B2F" box="[892,1102,1100,1125]" gridcol="3" gridrow="4" pageId="14" pageNumber="397">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376D00666D9CFBDD07069B2F" box="[1195,1385,1100,1125]" gridcol="4" gridrow="4" pageId="14" pageNumber="397">4p 4r 2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994FBE407069BC7" box="[163,1385,1141,1165]" gridrow="5" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FBE402BF9BC7" box="[163,208,1141,1165]" gridcol="0" gridrow="5" pageId="14" pageNumber="397">IV</th>
<td id="C74CC321376D00666846FBE403A99BC7" box="[369,454,1141,1165]" gridcol="1" gridrow="5" pageId="14" pageNumber="397">3d 4p 3r</td>
<td id="C74CC321376D00666B91FBE400B59BC7" box="[678,730,1141,1165]" gridcol="2" gridrow="5" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666A4BFBE406219BC7" box="[892,1102,1141,1165]" gridcol="3" gridrow="5" pageId="14" pageNumber="397">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376D00666D9CFBE407069BC7" box="[1195,1385,1141,1165]" gridcol="4" gridrow="5" pageId="14" pageNumber="397">4p 4r 1-1-1-1-1-2v</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376DFF9769A0FB73019F9BB0" box="[151,1008,1250,1275]" pageId="14" pageNumber="397" startId="14.[151,239,1250,1274]" targetBox="[163,1370,1290,1516]" targetIsTable="true" targetPageId="14" targetType="table">
<paragraph id="3A12A81F376DFF9769A0FB73019F9BB0" blockId="14.[151,1008,1250,1275]" box="[151,1008,1250,1275]" pageId="14" pageNumber="397">
<emphasis id="08D9740D376DFF9769A0FB7303789BB0" bold="true" box="[151,279,1250,1274]" pageId="14" pageNumber="397">TABLE 13.</emphasis>
Palp and legs measurements of female of
<taxonomicName id="FDADD39C376DFF976BECFB72019F9BB0" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[731,1008,1250,1275]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376DFF976BECFB72011E9BB0" box="[731,881,1250,1274]" italics="true" pageId="14" pageNumber="397">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376DFF976A40FB72019F9BB0" author="Jager, P." box="[887,1008,1250,1275]" pageId="14" pageNumber="397" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F376DFF97684EFA9A07359AA6" pageId="14" pageNumber="397">
<table id="48AD5ABF376D00666994FA9B07359AA6" box="[163,1370,1290,1516]" gridcols="7" gridrows="6" pageId="14" pageNumber="397">
<tr id="849DAA5D376D00666994FA9B07359A68" box="[163,1370,1290,1314]" gridrow="0" pageId="14" pageNumber="397" rowspan-0="1">
<th id="C74CC321376D00666858FA9B03F49A68" box="[367,411,1290,1314]" gridcol="1" gridrow="0" pageId="14" pageNumber="397">Fe</th>
<th id="C74CC321376D00666B1AFA9B00369A68" box="[557,601,1290,1314]" gridcol="2" gridrow="0" pageId="14" pageNumber="397">Pa</th>
<th id="C74CC321376D00666BDBFA9B01779A68" box="[748,792,1290,1314]" gridcol="3" gridrow="0" pageId="14" pageNumber="397">Ti</th>
<th id="C74CC321376D00666A9DFA9B01B99A68" box="[938,982,1290,1314]" gridcol="4" gridrow="0" pageId="14" pageNumber="397">Mt</th>
<th id="C74CC321376D00666D5EFA9B06FA9A68" box="[1129,1173,1290,1314]" gridcol="5" gridrow="0" pageId="14" pageNumber="397">Ta</th>
<th id="C74CC321376D00666C16FA9B07359A68" box="[1313,1370,1290,1314]" gridcol="6" gridrow="0" pageId="14" pageNumber="397">Total</th>
</tr>
<tr id="849DAA5D376D00666994FAA307359A01" box="[163,1370,1330,1355]" gridrow="1" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FAA302BF9A01" box="[163,208,1330,1355]" gridcol="0" gridrow="1" pageId="14" pageNumber="397">Palp</th>
<td id="C74CC321376D00666858FAA303F49A01" box="[367,411,1330,1355]" gridcol="1" gridrow="1" pageId="14" pageNumber="397">2.68</td>
<td id="C74CC321376D00666B1AFAA300369A01" box="[557,601,1330,1355]" gridcol="2" gridrow="1" pageId="14" pageNumber="397">1.36</td>
<td id="C74CC321376D00666BDBFAA301779A01" box="[748,792,1330,1355]" gridcol="3" gridrow="1" pageId="14" pageNumber="397">1.74</td>
<td id="C74CC321376D00666A9DFAA301B99A01" box="[938,982,1330,1355]" gridcol="4" gridrow="1" pageId="14" pageNumber="397">-</td>
<td id="C74CC321376D00666D5EFAA306FA9A01" box="[1129,1173,1330,1355]" gridcol="5" gridrow="1" pageId="14" pageNumber="397">2.03</td>
<td id="C74CC321376D00666C16FAA307359A01" box="[1313,1370,1330,1355]" gridcol="6" gridrow="1" pageId="14" pageNumber="397">7.81</td>
</tr>
<tr id="849DAA5D376D00666994FACA07359A39" box="[163,1370,1371,1395]" gridrow="2" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FACA02BF9A39" box="[163,208,1371,1395]" gridcol="0" gridrow="2" pageId="14" pageNumber="397">I</th>
<td id="C74CC321376D00666858FACA03F49A39" box="[367,411,1371,1395]" gridcol="1" gridrow="2" pageId="14" pageNumber="397">5.09</td>
<td id="C74CC321376D00666B1AFACA00369A39" box="[557,601,1371,1395]" gridcol="2" gridrow="2" pageId="14" pageNumber="397">2.93</td>
<td id="C74CC321376D00666BDBFACA01779A39" box="[748,792,1371,1395]" gridcol="3" gridrow="2" pageId="14" pageNumber="397">4.89</td>
<td id="C74CC321376D00666A9DFACA01B99A39" box="[938,982,1371,1395]" gridcol="4" gridrow="2" pageId="14" pageNumber="397">3.96</td>
<td id="C74CC321376D00666D5EFACA06FA9A39" box="[1129,1173,1371,1395]" gridcol="5" gridrow="2" pageId="14" pageNumber="397">1.40</td>
<td id="C74CC321376D00666C16FACA07359A39" box="[1313,1370,1371,1395]" gridcol="6" gridrow="2" pageId="14" pageNumber="397">18.27</td>
</tr>
<tr id="849DAA5D376D00666994FA1207359AD1" box="[163,1370,1411,1435]" gridrow="3" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FA1202BF9AD1" box="[163,208,1411,1435]" gridcol="0" gridrow="3" pageId="14" pageNumber="397">II</th>
<td id="C74CC321376D00666858FA1203F49AD1" box="[367,411,1411,1435]" gridcol="1" gridrow="3" pageId="14" pageNumber="397">5.05</td>
<td id="C74CC321376D00666B1AFA1200369AD1" box="[557,601,1411,1435]" gridcol="2" gridrow="3" pageId="14" pageNumber="397">2.76</td>
<td id="C74CC321376D00666BDBFA1201779AD1" box="[748,792,1411,1435]" gridcol="3" gridrow="3" pageId="14" pageNumber="397">4.29</td>
<td id="C74CC321376D00666A9DFA1201B99AD1" box="[938,982,1411,1435]" gridcol="4" gridrow="3" pageId="14" pageNumber="397">3.71</td>
<td id="C74CC321376D00666D5EFA1206FA9AD1" box="[1129,1173,1411,1435]" gridcol="5" gridrow="3" pageId="14" pageNumber="397">1.39</td>
<td id="C74CC321376D00666C16FA1207359AD1" box="[1313,1370,1411,1435]" gridcol="6" gridrow="3" pageId="14" pageNumber="397">17.2</td>
</tr>
<tr id="849DAA5D376D00666994FA3A07359A89" box="[163,1370,1451,1475]" gridrow="4" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FA3A02BF9A89" box="[163,208,1451,1475]" gridcol="0" gridrow="4" pageId="14" pageNumber="397">III</th>
<td id="C74CC321376D00666858FA3A03F49A89" box="[367,411,1451,1475]" gridcol="1" gridrow="4" pageId="14" pageNumber="397">4.22</td>
<td id="C74CC321376D00666B1AFA3A00369A89" box="[557,601,1451,1475]" gridcol="2" gridrow="4" pageId="14" pageNumber="397">2.34</td>
<td id="C74CC321376D00666BDBFA3A01779A89" box="[748,792,1451,1475]" gridcol="3" gridrow="4" pageId="14" pageNumber="397">3.14</td>
<td id="C74CC321376D00666A9DFA3A01B99A89" box="[938,982,1451,1475]" gridcol="4" gridrow="4" pageId="14" pageNumber="397">3.76</td>
<td id="C74CC321376D00666D5EFA3A06FA9A89" box="[1129,1173,1451,1475]" gridcol="5" gridrow="4" pageId="14" pageNumber="397">1.44</td>
<td id="C74CC321376D00666C16FA3A07359A89" box="[1313,1370,1451,1475]" gridcol="6" gridrow="4" pageId="14" pageNumber="397">14.9</td>
</tr>
<tr id="849DAA5D376D00666994FA4207359AA6" box="[163,1370,1491,1516]" gridrow="5" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994FA4202BF9AA6" box="[163,208,1491,1516]" gridcol="0" gridrow="5" pageId="14" pageNumber="397">IV</th>
<td id="C74CC321376D00666858FA4203F49AA6" box="[367,411,1491,1516]" gridcol="1" gridrow="5" pageId="14" pageNumber="397">5.79</td>
<td id="C74CC321376D00666B1AFA4200369AA6" box="[557,601,1491,1516]" gridcol="2" gridrow="5" pageId="14" pageNumber="397">2.25</td>
<td id="C74CC321376D00666BDBFA4201779AA6" box="[748,792,1491,1516]" gridcol="3" gridrow="5" pageId="14" pageNumber="397">4.98</td>
<td id="C74CC321376D00666A9DFA4201B99AA6" box="[938,982,1491,1516]" gridcol="4" gridrow="5" pageId="14" pageNumber="397">6.66</td>
<td id="C74CC321376D00666D5EFA4206FA9AA6" box="[1129,1173,1491,1516]" gridcol="5" gridrow="5" pageId="14" pageNumber="397">1.76</td>
<td id="C74CC321376D00666C16FA4207359AA6" box="[1313,1370,1491,1516]" gridcol="6" gridrow="5" pageId="14" pageNumber="397">21.44</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376DFF9769A0F9D001859913" box="[151,1002,1601,1625]" pageId="14" pageNumber="397" startId="14.[151,239,1601,1625]" targetBox="[163,1363,1641,1866]" targetIsTable="true" targetPageId="14" targetType="table">
<paragraph id="3A12A81F376DFF9769A0F9D001859913" blockId="14.[151,1002,1601,1625]" box="[151,1002,1601,1625]" pageId="14" pageNumber="397">
<emphasis id="08D9740D376DFF9769A0F9D003789913" bold="true" box="[151,279,1601,1625]" pageId="14" pageNumber="397">TABLE 14.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C376DFF976BE2F9D001859913" authority="Jager, 2022" authorityName="Jager" authorityYear="2022" box="[725,1002,1601,1625]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376DFF976BE2F9D001059913" box="[725,874,1601,1625]" italics="true" pageId="14" pageNumber="397">Bowie shakeit</emphasis>
<bibRefCitation id="5E3CD5EE376DFF976A47F9D001859913" author="Jager, P." box="[880,1002,1601,1625]" pageId="14" pageNumber="397" pagination="1 - 200" refId="ref9888" refString="Jager, P. (2022) Bowie gen. nov., a diverse lineage of ground-dwelling spiders occurring from the Himalayas to Papua New Guinea and northern Australia (Araneae: Ctenidae: Cteninae). Zootaxa, 5170 (1), 1 - 200. https: // doi. org / 10.11646 / zootaxa. 5170.1.1" type="journal article" year="2022">Jäger, 2022</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="3A12A81F376DFF9768B8F9FB073C9800" pageId="14" pageNumber="397">
<table id="48AD5ABF376D00666994F9F8073C9800" box="[163,1363,1641,1866]" gridcols="5" gridrows="6" pageId="14" pageNumber="397">
<tr id="849DAA5D376D00666994F9F8073C99CB" box="[163,1363,1641,1665]" gridrow="0" pageId="14" pageNumber="397" rowspan-0="1">
<th id="C74CC321376D00666857F9F803B899CB" box="[352,471,1641,1665]" gridcol="1" gridrow="0" pageId="14" pageNumber="397">Fe</th>
<th id="C74CC321376D00666BA5F9F8008299CB" box="[658,749,1641,1665]" gridcol="2" gridrow="0" pageId="14" pageNumber="397">Pa</th>
<th id="C74CC321376D00666AA1F9F8065A99CB" box="[918,1077,1641,1665]" gridcol="3" gridrow="0" pageId="14" pageNumber="397">Ti</th>
<th id="C74CC321376D00666DF7F9F8073C99CB" box="[1216,1363,1641,1665]" gridcol="4" gridrow="0" pageId="14" pageNumber="397">Mt</th>
</tr>
<tr id="849DAA5D376D00666994F900073C99E3" box="[163,1363,1681,1705]" gridrow="1" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994F90002BF99E3" box="[163,208,1681,1705]" gridcol="0" gridrow="1" pageId="14" pageNumber="397">Palp</th>
<td id="C74CC321376D00666857F90003B899E3" box="[352,471,1681,1705]" gridcol="1" gridrow="1" pageId="14" pageNumber="397">3d 1p 1r</td>
<td id="C74CC321376D00666BA5F900008299E3" box="[658,749,1681,1705]" gridcol="2" gridrow="1" pageId="14" pageNumber="397">1p</td>
<td id="C74CC321376D00666AA1F900065A99E3" box="[918,1077,1681,1705]" gridcol="3" gridrow="1" pageId="14" pageNumber="397">2p 2r</td>
<td id="C74CC321376D00666DF7F900073C99E3" box="[1216,1363,1681,1705]" gridcol="4" gridrow="1" pageId="14" pageNumber="397">-</td>
</tr>
<tr id="849DAA5D376D00666994F928073C9998" box="[163,1363,1721,1746]" gridrow="2" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994F92802BF9998" box="[163,208,1721,1746]" gridcol="0" gridrow="2" pageId="14" pageNumber="397">I</th>
<td id="C74CC321376D00666857F92803B89998" box="[352,471,1721,1746]" gridcol="1" gridrow="2" pageId="14" pageNumber="397">3d 3p 3(4)p</td>
<td id="C74CC321376D00666BA5F92800829998" box="[658,749,1721,1746]" gridcol="2" gridrow="2" pageId="14" pageNumber="397">spineless</td>
<td id="C74CC321376D00666AA1F928065A9998" box="[918,1077,1721,1746]" gridcol="3" gridrow="2" pageId="14" pageNumber="397">2-2-2-2-2v</td>
<td id="C74CC321376D00666DF7F928073C9998" box="[1216,1363,1721,1746]" gridcol="4" gridrow="2" pageId="14" pageNumber="397">2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994F973073C99B0" box="[163,1363,1762,1786]" gridrow="3" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994F97302BF99B0" box="[163,208,1762,1786]" gridcol="0" gridrow="3" pageId="14" pageNumber="397">II</th>
<td id="C74CC321376D00666857F97303B899B0" box="[352,471,1762,1786]" gridcol="1" gridrow="3" pageId="14" pageNumber="397">3d 4p 3r</td>
<td id="C74CC321376D00666BA5F973008299B0" box="[658,749,1762,1786]" gridcol="2" gridrow="3" pageId="14" pageNumber="397">spineless</td>
<td id="C74CC321376D00666AA1F973065A99B0" box="[918,1077,1762,1786]" gridcol="3" gridrow="3" pageId="14" pageNumber="397">2-2-2-2-2v</td>
<td id="C74CC321376D00666DF7F973073C99B0" box="[1216,1363,1762,1786]" gridcol="4" gridrow="3" pageId="14" pageNumber="397">2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994F89B073C9868" box="[163,1363,1802,1826]" gridrow="4" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994F89B02BF9868" box="[163,208,1802,1826]" gridcol="0" gridrow="4" pageId="14" pageNumber="397">III</th>
<td id="C74CC321376D00666857F89B03B89868" box="[352,471,1802,1826]" gridcol="1" gridrow="4" pageId="14" pageNumber="397">3d 2(4)p 4r</td>
<td id="C74CC321376D00666BA5F89B00829868" box="[658,749,1802,1826]" gridcol="2" gridrow="4" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666AA1F89B065A9868" box="[918,1077,1802,1826]" gridcol="3" gridrow="4" pageId="14" pageNumber="397">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376D00666DF7F89B073C9868" box="[1216,1363,1802,1826]" gridcol="4" gridrow="4" pageId="14" pageNumber="397">5p 5r 2-2-2v</td>
</tr>
<tr id="849DAA5D376D00666994F8A3073C9800" box="[163,1363,1842,1866]" gridrow="5" pageId="14" pageNumber="397">
<th id="C74CC321376D00666994F8A302BF9800" box="[163,208,1842,1866]" gridcol="0" gridrow="5" pageId="14" pageNumber="397">IV</th>
<td id="C74CC321376D00666857F8A303B89800" box="[352,471,1842,1866]" gridcol="1" gridrow="5" pageId="14" pageNumber="397">3d 4(5)p 2r</td>
<td id="C74CC321376D00666BA5F8A300829800" box="[658,749,1842,1866]" gridcol="2" gridrow="5" pageId="14" pageNumber="397">1p 1r</td>
<td id="C74CC321376D00666AA1F8A3065A9800" box="[918,1077,1842,1866]" gridcol="3" gridrow="5" pageId="14" pageNumber="397">3d 2p 2r 2-2-2v</td>
<td id="C74CC321376D00666DF7F8A3073C9800" box="[1216,1363,1842,1866]" gridcol="4" gridrow="5" pageId="14" pageNumber="397">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F897376CFF9669A0F884038098D3" ID-DOI="http://doi.org/10.5281/zenodo.14386421" ID-Zenodo-Dep="14386421" httpUri="https://zenodo.org/record/14386421/files/figure.png" pageId="15" pageNumber="398" startId="15.[151,264,1813,1837]" targetBox="[166,1435,189,1784]" targetPageId="15" targetType="figure">
<paragraph id="3A12A81F376CFF9669A0F884038098D3" blockId="15.[151,1436,1813,1945]" pageId="15" pageNumber="398">
<emphasis id="08D9740D376CFF9669A0F884033B9867" bold="true" box="[151,340,1813,1837]" pageId="15" pageNumber="398">FIGURES 46–53.</emphasis>
Palp (46–49) and bulb (50–53) of male of
<taxonomicName id="FDADD39C376CFF966A36F88701A09864" authority="Jager." authorityName="Jager." box="[769,975,1813,1838]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D376CFF966A36F88701E19867" box="[769,910,1813,1837]" italics="true" pageId="15" pageNumber="398">Bowie shakeit</emphasis>
Jäger.
</taxonomicName>
46, 50—prolateral; 47, 51—ventral; 48, 52— retrolateral; 49—dorsal; 53—apical. Scale bars: 46–49 = 0.5 mm; 50–53 = 0.2 mm. Abbreviations:
<emphasis id="08D9740D376CFF966D41F8AB06E8981B" box="[1142,1159,1850,1873]" italics="true" pageId="15" pageNumber="398">C</emphasis>
—conductor,
<emphasis id="08D9740D376CFF966C27F8AB075D981B" box="[1296,1330,1850,1873]" italics="true" pageId="15" pageNumber="398">EH</emphasis>
—embolic hook,
<emphasis id="08D9740D376CFF9669EDF8CF0293983F" box="[218,252,1886,1909]" italics="true" pageId="15" pageNumber="398">Em</emphasis>
—embolus,
<emphasis id="08D9740D376CFF96684BF8CF03C2983F" box="[380,429,1886,1909]" italics="true" pageId="15" pageNumber="398">RPO</emphasis>
—retro-proximal cymbial outgrowth,
<emphasis id="08D9740D376CFF966A0EF8CF010B983F" box="[825,868,1886,1909]" italics="true" pageId="15" pageNumber="398">RTA</emphasis>
—retrolateral tibial apophysis,
<emphasis id="08D9740D376CFF966D9CF8CF06A8983F" box="[1195,1223,1886,1909]" italics="true" pageId="15" pageNumber="398">TA</emphasis>
—tegular apophysis,
<emphasis id="08D9740D376CFF9669A0F81302C198D3" box="[151,174,1922,1945]" italics="true" pageId="15" pageNumber="398">Te</emphasis>
—tegulum,
<emphasis id="08D9740D376CFF966810F813032898D3" box="[295,327,1922,1945]" italics="true" pageId="15" pageNumber="398">TH</emphasis>
—tegular hump.
</paragraph>
</caption>
<caption id="6ED2F8973773FF8969A0F9BC006299C7" ID-DOI="http://doi.org/10.5281/zenodo.14386423" ID-Zenodo-Dep="14386423" httpUri="https://zenodo.org/record/14386423/files/figure.png" pageId="16" pageNumber="399" startId="16.[151,264,1581,1605]" targetBox="[151,1435,181,1557]" targetPageId="16" targetType="figure">
<paragraph id="3A12A81F3773FF8969A0F9BC006299C7" blockId="16.[151,1436,1581,1677]" pageId="16" pageNumber="399">
<emphasis id="08D9740D3773FF8969A0F9BC033B990F" bold="true" box="[151,340,1581,1605]" pageId="16" pageNumber="399">FIGURES 54–57.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3773FF8968E6F9BF00CF990F" authority="Jager." authorityName="Jager." box="[465,672,1581,1605]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="399" phylum="Arthropoda" rank="species" species="shakeit">
<emphasis id="08D9740D3773FF8968E6F9BF0030990F" box="[465,607,1581,1605]" italics="true" pageId="16" pageNumber="399">Bowie shakeit</emphasis>
Jäger.
</taxonomicName>
54—intact, ventral; 55—macerated, ventral; 56—internal genitalia, dorsal; 57—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3773FF896BF5F9C3008B9923" box="[706,740,1618,1641]" italics="true" pageId="16" pageNumber="399">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3773FF896A98F9C301A49923" box="[943,971,1618,1641]" italics="true" pageId="16" pageNumber="399">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3773FF896D5FF9C306E29923" box="[1128,1165,1618,1641]" italics="true" pageId="16" pageNumber="399">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3773FF8969A0F9E402DF99C6" box="[151,176,1653,1676]" italics="true" pageId="16" pageNumber="399">Sp</emphasis>
—spermatheca,
<emphasis id="08D9740D3773FF896862F9E7031B99C7" box="[341,372,1654,1677]" italics="true" pageId="16" pageNumber="399">VF</emphasis>
—vulval folds.
</paragraph>
</caption>
<paragraph id="3A12A81F3773FF8969F0F94E0092980B" blockId="16.[151,1443,1759,2038]" pageId="16" pageNumber="399">
Female genitalia as shown in
<figureCitation id="A296B49A3773FF896B19F94E00DB99B0" box="[558,692,1759,1786]" captionStart="FIGURES 54–57" captionStartId="16.[151,264,1581,1605]" captionTargetBox="[151,1435,181,1557]" captionTargetId="figure-30@16.[151,1435,181,1557]" captionTargetPageId="16" captionText="FIGURES 54–57. Epigyne of Bowie shakeit Jäger. 54—intact, ventral; 55—macerated, ventral; 56—internal genitalia, dorsal; 57—ditto, posterior. Scale bar: 0.2 mm.Abbreviations: FD—fertilization duct, LT—lateral tooth, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386423" httpUri="https://zenodo.org/record/14386423/files/figure.png" pageId="16" pageNumber="399">Figs 54–57</figureCitation>
. Epigyne slightly longer than wide. Median plate (
<emphasis id="08D9740D3773FF896C22F971075299B3" box="[1301,1341,1760,1785]" italics="true" pageId="16" pageNumber="399">MP</emphasis>
) widest posteriorly. Lateral teeth (
<emphasis id="08D9740D3773FF896884F89503BD9857" box="[435,466,1796,1821]" italics="true" pageId="16" pageNumber="399">LT</emphasis>
) comparatively short, located posterior of median plate, with pointed tips. Spermathecae (
<emphasis id="08D9740D3773FF896996F8B602D2980B" box="[161,189,1831,1857]" italics="true" pageId="16" pageNumber="399">Sp</emphasis>
) small, kidney-shaped, almost touching each other.
</paragraph>
<paragraph id="3A12A81F3773FF8969F0F8DA02A0989B" blockId="16.[151,1443,1759,2038]" pageId="16" pageNumber="399">
<emphasis id="08D9740D3773FF8969F0F8DA037E982F" bold="true" box="[199,273,1867,1893]" pageId="16" pageNumber="399">Notes.</emphasis>
The species has been previously known from a single record from Maliau Basin (
<taxonomicName id="FDADD39C3773FF896DDAF8DA0715982C" authority="Prov." authorityName="Prov." box="[1261,1402,1867,1894]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="399" phylum="Arthropoda" rank="species" species="sabah">Sabah Prov.</taxonomicName>
of
<collectingCountry id="42BAE88F3773FF8969A0F8FE036C98C3" box="[151,259,1903,1929]" name="Malaysia" pageId="16" pageNumber="399">Malaysia</collectingCountry>
). The new record is located ca.
<quantity id="FD5505FA3773FF896B54F8FE00D798C3" box="[611,696,1903,1929]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="399" unit="km" value="100.0">100 km</quantity>
to the east of the
<typeStatus id="E51616BD3773FF896A49F8E101C198C0" box="[894,942,1904,1930]" pageId="16" pageNumber="399">type</typeStatus>
locality. Because only its copulatory organs have been illustrated by black-and-white drawings only, we provide digital photos along with the redescription here.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943773FF8969F0F84A010F98BF" box="[199,864,2011,2038]" pageId="16" pageNumber="399" type="distribution">
<paragraph id="3A12A81F3773FF8969F0F84A010F98BF" blockId="16.[151,1443,1759,2038]" box="[199,864,2011,2038]" pageId="16" pageNumber="399">
<emphasis id="08D9740D3773FF8969F0F84A030E98BF" bold="true" box="[199,353,2011,2037]" pageId="16" pageNumber="399">Distribution.</emphasis>
Borneo,
<collectingCountry id="42BAE88F3773FF8968FFF84A005E98BF" box="[456,561,2011,2037]" name="Malaysia" pageId="16" pageNumber="399">Malaysia</collectingCountry>
,
<taxonomicName id="FDADD39C3773FF896B0AF84A00AC98BC" authority="Prov." authorityName="Prov." box="[573,707,2011,2038]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="399" phylum="Arthropoda" rank="species" species="sabah">Sabah Prov.</taxonomicName>
(
<figureCitation id="A296B49A3773FF896BE6F84A013A98BF" box="[721,853,2011,2037]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="16" pageNumber="399">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>