276 lines
36 KiB
XML
276 lines
36 KiB
XML
<document id="1DE53DFFDAE642258363BB2B1D1D6429" ID-DOI="10.1051/parasite/2023058" ID-ISSN="1776-1042" ID-PMC="PMC10732140" ID-PubMed="38117273" ID-Zenodo-Dep="12628842" ID-ZooBank="DC25665A-E218-496B-974E-B813F69395E5" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1719954839333" checkinUser="felipe" docAuthor="Durette-Desset, Marie-Claude & Digiani, María Celina" docDate="2023" docId="03ED87FEFF90FFD52440FC0BFD21F9F1" docLanguage="en" docName="Parasite.30.63.pdf" docOrigin="Parasite (Paris, France) 30 (63)" docSource="http://dx.doi.org/10.1051/parasite/2023058" docStyle="DocumentStyle:A40D95A4C09C797DBB807E438D2BF437.2:Parasite.2013-.journal_article" docStyleId="A40D95A4C09C797DBB807E438D2BF437" docStyleName="Parasite.2013-.journal_article" docStyleVersion="2" docTitle="Rodentanema aenigma Smales 2016" docType="treatment" docVersion="1" lastPageNumber="20" masterDocId="FFD4FF86FF81FFC6276DFFE6FF8BFF8C" masterDocTitle="Revision of the genera of Heligmonellidae (Nematoda, Heligmosomoidea), parasitic in Muridae from New Guinea" masterLastPageNumber="34" masterPageNumber="1" pageNumber="18" updateTime="1728046454050" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="4EABCB44D9E8FA7CE504D78E2481B22F">
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<mods:title id="E47F416350DCF1716059318879FEC6DD">Revision of the genera of Heligmonellidae (Nematoda, Heligmosomoidea), parasitic in Muridae from New Guinea</mods:title>
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<mods:name id="B4F6040808327C7EAF0FD59626A8E13B" type="personal">
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<mods:roleTerm id="EC46EC8C49C7D3052255F8EEEDE61203">Author</mods:roleTerm>
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<mods:namePart id="0B05ECB2AC6B594F3AF870D7AF03E9AA">Durette-Desset, Marie-Claude</mods:namePart>
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<mods:affiliation id="20DB4FDD400073DB3BA74D19BE0336E4">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum National d’Histoire Naturelle, CNRS, Sorbonne Université,</mods:affiliation>
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<mods:roleTerm id="801683D5488BDBDAAC94C33F17049F2E">Author</mods:roleTerm>
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<mods:namePart id="02861D7C50B4F6052C71CDDF72AAC68D">Digiani, María Celina</mods:namePart>
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<mods:affiliation id="C40B2CF51FCF13EFA4356FC571A08768">CONICET-Consejo Nacional de Investigaciones Científicas y Técnicas, Argentina & División Zoología Invertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata,</mods:affiliation>
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<mods:title id="3C9ACDB23A3C642D1244AAB817D3FEBE">Parasite</mods:title>
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<mods:date id="ADF733C4E828F9AEE435B661BC0BB71E">2023</mods:date>
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<mods:title id="E2E82B2FF9B3A3FADA253BBDEC44CB39">Paris, France</mods:title>
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<mods:number id="48E786292275F7CCFE69A004F7D7C027">2023-12-20</mods:number>
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<mods:number id="D32FD5F76AB96FB4ACFC9FDC65C1DB11">30</mods:number>
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<mods:number id="16E10C87EED8B650B87BDEC4224AF674">63</mods:number>
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<treatment id="03ED87FEFF90FFD52440FC0BFD21F9F1" LSID="urn:lsid:plazi:treatment:03ED87FEFF90FFD52440FC0BFD21F9F1" httpUri="http://treatment.plazi.org/id/03ED87FEFF90FFD52440FC0BFD21F9F1" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="C35E6563FF90FFD72440FC0BFBEAFB91" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BFB36E8FF90FFD72440FC0BFBEAFB91" blockId="17.[813,1359,1005,1053]" pageId="17" pageNumber="18">
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<emphasis id="B930EAFAFF90FFD72440FC0BFBEAFB91" bold="true" pageId="17" pageNumber="18">
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3.6.1 Analysis of data and difficulties encountered
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<taxonomicName id="4C444D6BFF90FFD72404FBEFFBEAFB91" ID-CoL="9BWL8" authority="Smales, 2016" authorityName="Smales" authorityYear="2016" box="[873,1121,1033,1053]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="17" pageNumber="18" phylum="Nematoda" rank="species" species="aenigma">
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<emphasis id="B930EAFAFF90FFD72404FBEFFBEAFB91" bold="true" box="[873,1121,1033,1053]" italics="true" pageId="17" pageNumber="18">Rodentanema aenigma</emphasis>
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35E6563FF90FFD52440FBA4FD21F9F1" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BFB36E8FF90FFD72440FBA4FC38FBDA" blockId="17.[813,1467,1090,1202]" box="[813,947,1090,1110]" pageId="17" pageNumber="18">
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<heading id="D0B38184FF90FFD72440FBA4FC38FBDA" box="[813,947,1090,1110]" fontSize="9" level="2" pageId="17" pageNumber="18" reason="4">
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<emphasis id="B930EAFAFF90FFD72440FBA4FC38FBDA" bold="true" box="[813,947,1090,1110]" italics="true" pageId="17" pageNumber="18">3.6.1.1 Head</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD72438FBBBFAB4FB3D" blockId="17.[813,1467,1090,1202]" pageId="17" pageNumber="18">Illustrations analyzed herein are proximal extremity in lateral (Fig. 27/6A) and apical (28/6B) views. Buccal capsule (Fig. 27/6A) and five lips (Fig. 28/6B), illustrated.</paragraph>
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<paragraph id="8BFB36E8FF90FFD72440FB02FBBEFA9F" blockId="17.[813,1467,1251,1728]" pageId="17" pageNumber="18">
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<emphasis id="B930EAFAFF90FFD72440FB02FAE3FB74" bold="true" box="[813,1384,1251,1272]" italics="true" pageId="17" pageNumber="18">
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3.6.1.2 Synlophe (based on sections from
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<specimenCount id="9D42FD61FF90FFD72394FB02FAE3FB74" box="[1273,1384,1252,1272]" count="2" pageId="17" pageNumber="18" type="male">two males</specimenCount>
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</emphasis>
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<emphasis id="B930EAFAFF90FFD72411FB19FBBEFA9F" bold="true" box="[892,1077,1279,1299]" italics="true" pageId="17" pageNumber="18">
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and
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<specimenCount id="9D42FD61FF90FFD724C6FB19FBA6FA9F" box="[939,1069,1279,1299]" count="2" pageId="17" pageNumber="18" type="female">two females</specimenCount>
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)
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</emphasis>
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD72438FAFDFC10FAE3" blockId="17.[813,1467,1251,1728]" pageId="17" pageNumber="18">Sections analyzed herein are at midbody: male (Figs. 31/6C) and female (32/6D). Lateral cords not illustrated; ridges not numbered.</paragraph>
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<paragraph id="8BFB36E8FF90FFD72438FA91FBF9FA47" blockId="17.[813,1467,1251,1728]" pageId="17" pageNumber="18">
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<emphasis id="B930EAFAFF90FFD72438FA91FC5DFA02" box="[853,982,1399,1422]" italics="true" pageId="17" pageNumber="18">At midbody:</emphasis>
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in both sexes, careen absent and ridges subequal, medium-sized. Ridges irregularly spaced, separated by ridge-free spaces (arrowheads).
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD72438FA35FBE3F9C9" blockId="17.[813,1467,1251,1728]" pageId="17" pageNumber="18">
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<figureCitation id="137F2A6DFF90FFD72438FA35FC4AFA66" box="[853,961,1491,1514]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="17" pageNumber="18">Figure 6C</figureCitation>
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(male) six ridges separated by three ridge-free spaces (arrowheads): two ridges mid- dorsal oriented from right to left, one ridge right-ventral, three ridges left-ventral; all these latter with unclear orientation.
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD72438F9ABFBC5F94C" blockId="17.[813,1467,1251,1728]" pageId="17" pageNumber="18">
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<figureCitation id="137F2A6DFF90FFD72438F9ABFC36F9E8" box="[853,957,1613,1636]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="17" pageNumber="18">Figure 6D</figureCitation>
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(female) seven ridges separated by two ridge-free spaces (arrowheads): one ridge mid-dorsal oriented to left, six ventral ridges (three right-ventral, two left-ventral); tips of these two latter groups divergent.
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD72440F914FA9DF8AE" blockId="17.[813,1467,1778,2040]" pageId="17" pageNumber="18">
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<emphasis id="B930EAFAFF90FFD72440F914FAF6F88A" bold="true" box="[813,1405,1778,1798]" italics="true" pageId="17" pageNumber="18">3.6.1.3 Bursa (number of worms studied not specified,</emphasis>
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<emphasis id="B930EAFAFF90FFD72411F8E8FA9DF8AE" bold="true" box="[892,1302,1806,1826]" italics="true" pageId="17" pageNumber="18">
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illustrated in [
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<bibRefCitation id="EFD54B19FF90FFD7237FF8E8FBA0F8AE" author="Smales LR" box="[1042,1067,1806,1826]" pageId="17" pageNumber="18" pagination="183 - 197" refId="ref29907" refString="38. Smales LR. 2016. The gastrointestinal helminths of Rattus niobe (Rodentia: Muridae) with descriptions of two new genera and three new species (Nematoda) from Papua New Guinea and Papua Indonesia. Zootaxa, 4117, 183 - 197." type="journal article" year="2016">38</bibRefCitation>
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]: Figs. 30, 33 and 34)
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</emphasis>
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</paragraph>
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<paragraph id="8BFB36E8FF90FFD42438F8CFFE26FB6A" blockId="17.[813,1467,1778,2040]" lastBlockId="18.[113,767,1200,1254]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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Figure 30: distal extremity including closed bursa, left lateral view, only rays 3-8 illustrated, rays 4-6 joined up to extremities. Figure 33: bursa “partially unrolled in left lateral and dorsal aspects”, only dorsal ray and left rays 4-8 illustrated, extremities of left rays 4-6 diverging. Figure 34: dorsal lobe, orientation not specified. Right lobe not illustrated. From generic definition [
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<bibRefCitation id="EFD54B19FF90FFD72497F807FB9FF874" author="Smales LR" box="[1018,1044,2017,2040]" pageId="17" pageNumber="18" pagination="183 - 197" refId="ref29907" refString="38. Smales LR. 2016. The gastrointestinal helminths of Rattus niobe (Rodentia: Muridae) with descriptions of two new genera and three new species (Nematoda) from Papua New Guinea and Papua Indonesia. Zootaxa, 4117, 183 - 197." type="journal article" year="2016">38</bibRefCitation>
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], bursa dissymmetrical with right lobe larger; whereas from description, left lobe larger. From generic definition, pattern of
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<typeStatus id="54FF884AFF93FFD42622FB29FEF0FB6A" box="[335,379,1231,1254]" pageId="18" pageNumber="19">type</typeStatus>
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1-4.
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</paragraph>
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<caption id="DF3B6660FF93FFD4271CFBFDFA8DFBE4" ID-DOI="http://doi.org/10.5281/zenodo.12628858" ID-Zenodo-Dep="12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19" startId="18.[113,180,1051,1072]" targetBox="[144,1428,238,1029]" targetPageId="18" targetType="figure">
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<paragraph id="8BFB36E8FF93FFD4271CFBFDFA8DFBE4" blockId="18.[113,1467,1051,1128]" pageId="18" pageNumber="19">
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<emphasis id="B930EAFAFF93FFD4271CFBFDFF44FBBC" bold="true" box="[113,207,1051,1072]" pageId="18" pageNumber="19">Figure 6.</emphasis>
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Genus
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<taxonomicName id="4C444D6BFF93FFD4264DFBFDFDBBFBBC" authority="Smales, 2016" authorityName="Smales" authorityYear="2016" box="[288,560,1051,1072]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
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<emphasis id="B930EAFAFF93FFD4264DFBFDFE28FBBC" box="[288,419,1051,1072]" italics="true" pageId="18" pageNumber="19">Rodentanema</emphasis>
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||
Smales, 2016
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</taxonomicName>
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. A–D’.
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<taxonomicName id="4C444D6BFF93FFD42512FBFDFC61FBBC" authority="Smales, 2016" authorityName="Smales" authorityYear="2016" box="[639,1002,1051,1072]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="species" species="aenigma">
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<emphasis id="B930EAFAFF93FFD42512FBFDFCD7FBBC" box="[639,860,1051,1072]" italics="true" pageId="18" pageNumber="19">Rodentanema aenigma</emphasis>
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Smales, 2016
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</taxonomicName>
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. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [
|
||
<bibRefCitation id="EFD54B19FF93FFD427C2FBB5FF4CFBE4" author="Smales LR" box="[175,199,1107,1128]" pageId="18" pageNumber="19" pagination="183 - 197" refId="ref29907" refString="38. Smales LR. 2016. The gastrointestinal helminths of Rattus niobe (Rodentia: Muridae) with descriptions of two new genera and three new species (Nematoda) from Papua New Guinea and Papua Indonesia. Zootaxa, 4117, 183 - 197." type="journal article" year="2016">38</bibRefCitation>
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]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original.
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</paragraph>
|
||
</caption>
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<paragraph id="8BFB36E8FF93FFD4271CFAE3FEA9FA95" blockId="18.[113,290,1284,1305]" box="[113,290,1284,1305]" pageId="18" pageNumber="19">
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<emphasis id="B930EAFAFF93FFD4271CFAE3FEA9FA95" bold="true" box="[113,290,1284,1305]" pageId="18" pageNumber="19">3.6.2 Comments</emphasis>
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</paragraph>
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<paragraph id="8BFB36E8FF93FFD4271CFAC8FF7CFACE" blockId="18.[113,767,1326,1529]" box="[113,247,1326,1346]" pageId="18" pageNumber="19">
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<heading id="D0B38184FF93FFD4271CFAC8FF7CFACE" box="[113,247,1326,1346]" fontSize="9" level="2" pageId="18" pageNumber="19" reason="4">
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<emphasis id="B930EAFAFF93FFD4271CFAC8FF7CFACE" bold="true" box="[113,247,1326,1346]" italics="true" pageId="18" pageNumber="19">3.6.2.1 Head</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BFB36E8FF93FFD427F4FAACFF1DFA75" blockId="18.[113,767,1326,1529]" pageId="18" pageNumber="19">
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In the original written description, the cephalic vesicle is indicated as present but is not illustrated. On the other hand, lips and teeth are indicated as absent, but the illustration in apical view shows clearly five lips (Fig.
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[
|
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<bibRefCitation id="EFD54B19FF93FFD42563FA43FDA3FA30" author="Smales LR" box="[526,552,1445,1468]" pageId="18" pageNumber="19" pagination="183 - 197" refId="ref29907" refString="38. Smales LR. 2016. The gastrointestinal helminths of Rattus niobe (Rodentia: Muridae) with descriptions of two new genera and three new species (Nematoda) from Papua New Guinea and Papua Indonesia. Zootaxa, 4117, 183 - 197." type="journal article" year="2016">38</bibRefCitation>
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]), which is undoubtedly a drawing flaw since these structures appear in number of six.
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD4271CF9C7FEADF9B9" blockId="18.[113,768,1568,2047]" box="[113,294,1568,1589]" pageId="18" pageNumber="19">
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<emphasis id="B930EAFAFF93FFD4271CF9C7FEADF9B9" bold="true" box="[113,294,1568,1589]" italics="true" pageId="18" pageNumber="19">3.6.2.2 Synlophe</emphasis>
|
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</paragraph>
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<paragraph id="8BFB36E8FF93FFD427F4F9DBFE50F9FE" blockId="18.[113,768,1568,2047]" pageId="18" pageNumber="19">The position of the ridges and the inclination of the axis are very different between both sexes.</paragraph>
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||
<paragraph id="8BFB36E8FF93FFD427F4F99CFDABF8A6" blockId="18.[113,768,1568,2047]" pageId="18" pageNumber="19">
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In
|
||
<figureCitation id="137F2A6DFF93FFD427D5F99CFEAFF91D" box="[184,292,1658,1681]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19">Figure 6C</figureCitation>
|
||
(male), the tip of the only ridge situated on right-ventral quadrant is perpendicular to the body surface but clearly divergent from the tips of the dorsal ridges. The remaining ventral ridges are also oriented perpendicularly to body surface, there are no convergent tips and the inclination of the axis of orientation remains uncertain.
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD427F4F8D7FD66F828" blockId="18.[113,768,1568,2047]" pageId="18" pageNumber="19">
|
||
<figureCitation id="137F2A6DFF93FFD427F4F8D7FE88F8C4" box="[153,259,1841,1864]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19">Figure 6D</figureCitation>
|
||
(female); if the section is reversed on its sagittal axis, then turned about 75° counterclockwise we obtain a section whose synlophe is similar to that of other Heligmosomoidea in which the axis is subfrontal (
|
||
<figureCitation id="137F2A6DFF93FFD425EBF86BFD55F828" box="[646,734,1932,1956]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19">Fig. 6D’</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD427F4F84DFB31FA88" blockId="18.[113,768,1568,2047]" lastBlockId="18.[813,1467,1199,1895]" pageId="18" pageNumber="19">
|
||
Based on these observations,
|
||
<specimenCount id="9D42FD61FF93FFD426BFF84DFDB3F84E" box="[466,568,1963,1986]" count="2" pageId="18" pageNumber="19" type="generic" typeStatus="types">two types</specimenCount>
|
||
of synlophe could be described:
|
||
<typeStatus id="54FF884AFF93FFD42666F82CFECBF86D" box="[267,320,1994,2017]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
I (
|
||
<figureCitation id="137F2A6DFF93FFD42605F82CFE37F86D" box="[360,444,1994,2017]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19">Fig. 6C</figureCitation>
|
||
) characterized by 6 ridges at midbody grouped into three sets with inclination of axis of orientation uncertain.
|
||
<typeStatus id="54FF884AFF93FFD4234DFB56FBDEFB4B" box="[1056,1109,1200,1223]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
II (
|
||
<figureCitation id="137F2A6DFF93FFD423E2FB56FB7BFB4B" box="[1167,1264,1199,1223]" captionStart="Figure 6" captionStartId="18.[113,180,1051,1072]" captionTargetBox="[144,1428,238,1029]" captionTargetId="figure-461@18.[144,1429,238,1029]" captionTargetPageId="18" captionText="Figure 6. Genus Rodentanema Smales, 2016. A–D’. Rodentanema aenigma Smales, 2016. A–B’ head. A lateral or median view showing buccal capsule. B, B’ apical view. C, D sections at midbody. C male, D, D’ female. Abbreviations: bc buccal capsule. Source: A–D redrawn from [38]. B’, D’ modified figures: B’ six lips instead five. D’ rotated 90° counterclockwise with respect to the original." figureDoi="http://doi.org/10.5281/zenodo.12628858" httpUri="https://zenodo.org/record/12628858/files/figure.png" pageId="18" pageNumber="19">Fig. 6D’</figureCitation>
|
||
) characterized by 7 ridges at midbody grouped into two sets with inclination of axis of orientation perhaps subfrontal.
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD42438FAEAFB5CF9FF" blockId="18.[813,1467,1199,1895]" pageId="18" pageNumber="19">
|
||
The presence of a cephalic vesicle, a developed buccal capsule, and six lips indicate that the specimens studied do not belong to the
|
||
<taxonomicName id="4C444D6BFF93FFD424D3FAAFFBE3FAEC" authorityName="Skrjabin & Schikobalova" authorityYear="1952" box="[958,1128,1353,1376]" class="Chromadorea" family="Heligmonellidae" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="family">Heligmonellidae</taxonomicName>
|
||
but should rather be assimilated with the
|
||
<taxonomicName id="4C444D6BFF93FFD424E4FA81FBD6FAF2" box="[905,1117,1383,1406]" class="Chromadorea" family="Herpetostrongylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="family">Herpetostrongylidae</taxonomicName>
|
||
: Herpetostrongylinae (parasitic in reptiles, Australian marsupials and exceptionally in rodents). The Herpetostrongylinae possess the same cephalic characters described above, plus 1-3 oesophageal teeth (not observed by the author in
|
||
<taxonomicName id="4C444D6BFF93FFD424AFFA04FBDBFA75" authorityName="Smales" authorityYear="2016" box="[962,1104,1506,1529]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF93FFD424AFFA04FBDBFA75" box="[962,1104,1506,1529]" italics="true" pageId="18" pageNumber="19">Rodentanema</emphasis>
|
||
</taxonomicName>
|
||
). Besides the cephalic characters, the Herpetostrongylinae possess a synlophe with an axis of orientation oblique or subfrontal, bursae of different
|
||
<typeStatus id="54FF884AFF93FFD422E9F9F9FA31F9BA" box="[1412,1466,1567,1590]" pageId="18" pageNumber="19">types</typeStatus>
|
||
including the
|
||
<typeStatus id="54FF884AFF93FFD424A6F9DBFC73F9D8" box="[971,1016,1597,1620]" pageId="18" pageNumber="19">type</typeStatus>
|
||
1-4, females didelphic or monodelphic, and female tails with or without a spine.
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD42438F99CFB6AF8EB" blockId="18.[813,1467,1199,1895]" pageId="18" pageNumber="19">
|
||
Among the Herpetostrongylinae, only two genera are characterized by females which are monodelphic and without a caudal spine:
|
||
<taxonomicName id="4C444D6BFF93FFD424D0F95EFAF2F942" authority="Humphery-Smith, 1981" authorityName="Humphery-Smith" authorityYear="1981" box="[957,1401,1719,1743]" class="Chromadorea" family="Heligmosomidae" genus="Dessetostrongylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF93FFD424D0F95EFBF2F943" box="[957,1145,1720,1743]" italics="true" pageId="18" pageNumber="19">Dessetostrongylus</emphasis>
|
||
Humphery-Smith, 1981
|
||
</taxonomicName>
|
||
, parasitic in
|
||
<taxonomicName id="4C444D6BFF93FFD424E8F930FB19F961" authority="(Marsupialia)" baseAuthorityName="Marsupialia" box="[901,1170,1750,1773]" class="Mammalia" family="Dasyuridae" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="19" phylum="Chordata" rank="family">Dasyuridae (Marsupialia)</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3537678FF93FFD423ACF930FAAAF961" box="[1217,1313,1750,1773]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C444D6BFF93FFD42208F930FBA8F880" authority="Smales, 2010" authorityName="Smales" authorityYear="2010" class="Chromadorea" family="Herpetostrongylidae" genus="Papuastrongylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF93FFD42208F930FC05F880" italics="true" pageId="18" pageNumber="19">Papuastrongylus</emphasis>
|
||
Smales, 2010
|
||
</taxonomicName>
|
||
parasitic in
|
||
<taxonomicName id="4C444D6BFF93FFD423CAF913FB75F880" box="[1191,1278,1781,1804]" class="Mammalia" family="Muridae" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="family">Muridae</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3537678FF93FFD4222FF913FCF0F8A6" name="Papua New Guinea" pageId="18" pageNumber="19">Papua New Guinea</collectingCountry>
|
||
. The two genera are differentiated from the specimens described as
|
||
<taxonomicName id="4C444D6BFF93FFD424D1F8D4FBC1F8C5" authorityName="Smales" authorityYear="2016" box="[956,1098,1842,1865]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF93FFD424D1F8D4FBC1F8C5" box="[956,1098,1842,1865]" italics="true" pageId="18" pageNumber="19">Rodentanema</emphasis>
|
||
</taxonomicName>
|
||
by having well-developed buccal capsules and a different synlophe pattern.
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD42440F869FC37F82F" blockId="18.[813,1467,1935,2047]" box="[813,956,1935,1955]" pageId="18" pageNumber="19">
|
||
<emphasis id="B930EAFAFF93FFD42440F869FC37F82F" bold="true" box="[813,956,1935,1955]" italics="true" pageId="18" pageNumber="19">3.6.2.3 Bursa</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF93FFD52438F84DFDF3FA86" blockId="18.[813,1467,1935,2047]" lastBlockId="19.[113,768,1115,1290]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
According to [
|
||
<bibRefCitation id="EFD54B19FF93FFD42487F84DFB8FF84E" author="Smales LR" box="[1002,1028,1963,1986]" pageId="18" pageNumber="19" pagination="183 - 197" refId="ref29907" refString="38. Smales LR. 2016. The gastrointestinal helminths of Rattus niobe (Rodentia: Muridae) with descriptions of two new genera and three new species (Nematoda) from Papua New Guinea and Papua Indonesia. Zootaxa, 4117, 183 - 197." type="journal article" year="2016">38</bibRefCitation>
|
||
], Figures 30 and 33 illustrate the left lobe. But both figures are clearly different, particularly concerning left ray 8, which in Figure 33 is seen mostly parallel to ray 6 and in Figure 30 it diverges proximally from it. Figures 30 and 33 correspond clearly to different bursae, which may imply the presence of two different species among the
|
||
<typeStatus id="54FF884AFF92FFD5253CFB7EFDF6FB23" box="[593,637,1176,1199]" pageId="19" pageNumber="20">type</typeStatus>
|
||
material. In addition, the right lobe is not illustrated, the description of the bursa is very brief and ambiguous and in the absence of an illustration of ray 2, the pattern cannot be determined.
|
||
</paragraph>
|
||
<caption id="DF3B6660FF92FFD5271CFC30FCBFFBAF" ID-DOI="http://doi.org/10.5281/zenodo.12628860" ID-Zenodo-Dep="12628860" httpUri="https://zenodo.org/record/12628860/files/figure.png" pageId="19" pageNumber="20" startId="19.[113,180,982,1003]" targetBox="[161,1435,239,961]" targetPageId="19" targetType="figure">
|
||
<paragraph id="8BFB36E8FF92FFD5271CFC30FCBFFBAF" blockId="19.[113,1467,981,1059]" pageId="19" pageNumber="20">
|
||
<emphasis id="B930EAFAFF92FFD5271CFC30FF44FC67" bold="true" box="[113,207,982,1003]" pageId="19" pageNumber="20">Figure 7.</emphasis>
|
||
Genus
|
||
<taxonomicName id="4C444D6BFF92FFD5264DFC30FD9DFC67" authority="Smales, 2017" authorityName="Smales" authorityYear="2017" box="[288,534,982,1003]" class="Chromadorea" family="Heligmosomidae" genus="Parvinema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF92FFD5264DFC30FE03FC67" box="[288,392,982,1003]" italics="true" pageId="19" pageNumber="20">Parvinema</emphasis>
|
||
Smales, 2017
|
||
</taxonomicName>
|
||
. Midbody sections. A, B
|
||
<taxonomicName id="4C444D6BFF92FFD52479FC30FB1DFC67" authority="Smales, 2017" authorityName="Smales" authorityYear="2017" box="[788,1174,982,1003]" class="Chromadorea" family="Heligmosomidae" genus="Parvinema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="species" species="bafunminense">
|
||
<emphasis id="B930EAFAFF92FFD52479FC30FB83FC67" box="[788,1032,982,1003]" italics="true" pageId="19" pageNumber="20">Parvinema bafunminense</emphasis>
|
||
Smales, 2017
|
||
</taxonomicName>
|
||
. A male, B, B’ female. C, C’
|
||
<taxonomicName id="4C444D6BFF92FFD5271CFC14FE25FB8B" authority="Smales, 2017" authorityName="Smales" authorityYear="2017" box="[113,430,1010,1031]" class="Chromadorea" family="Heligmosomidae" genus="Parvinema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="species" species="helgeni">
|
||
<emphasis id="B930EAFAFF92FFD5271CFC14FEADFB8B" box="[113,294,1010,1031]" italics="true" pageId="19" pageNumber="20">Parvinema helgeni</emphasis>
|
||
Smales, 2017
|
||
</taxonomicName>
|
||
, male. Source: A–C redrawn from [
|
||
<bibRefCitation id="EFD54B19FF92FFD5246EFC14FC90FB8B" author="Smales LR" box="[771,795,1010,1031]" pageId="19" pageNumber="20" pagination="762 - 771" refId="ref29998" refString="40. Smales LR. 2017. The gastrointestinal nematodes of Paramelomys lorentzii and Mammelomys spp. (Rodentia: Muridae) with descriptions of a new genus and three new species (Heligmonellidae) from Papua New Guinea and Indonesia. Acta Parasitologica, 62, 762 - 771." type="journal article" year="2017">40</bibRefCitation>
|
||
]. B’, C’ modified figures with respect to the original: B’ numbering of ridges added. C’ reversed on its frontal axis, then rotated 30° clockwise.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFB36E8FF92FFD5271CFADFFEA2FAC2" blockId="19.[113,297,1337,1358]" box="[113,297,1337,1358]" pageId="19" pageNumber="20">
|
||
<emphasis id="B930EAFAFF92FFD5271CFADFFEA2FAC2" bold="true" box="[113,297,1337,1358]" pageId="19" pageNumber="20">3.6.3 Conclusion</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFB36E8FF92FFD527F4FA94FD21F9F1" blockId="19.[113,768,1394,1662]" pageId="19" pageNumber="20">
|
||
There seem to be two taxa concerned in the description of this species (each characterized by a different synlophe): “
|
||
<taxonomicName id="4C444D6BFF92FFD52710FA49FF7FFA4A" authorityName="Smales" authorityYear="2016" box="[125,244,1455,1478]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="species" species="aenigma">
|
||
<emphasis id="B930EAFAFF92FFD52710FA49FF7FFA4A" box="[125,244,1455,1478]" italics="true" pageId="19" pageNumber="20">R. aenigma</emphasis>
|
||
</taxonomicName>
|
||
” and a
|
||
<taxonomicName id="4C444D6BFF92FFD5262AFA49FD93FA4A" box="[327,536,1455,1478]" class="Chromadorea" family="Herpetostrongylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="family">Herpetostrongylidae</taxonomicName>
|
||
<emphasis id="B930EAFAFF92FFD52572FA49FDB6FA4A" box="[543,573,1455,1478]" italics="true" pageId="19" pageNumber="20">i.s.</emphasis>
|
||
The uncertain orientation of the ridges, the incomplete description of the bursa, plus the fact that we do not know what type of synlophe the
|
||
<typeStatus id="54FF884AFF92FFD5271CF9EDFF40F9AE" box="[113,203,1547,1570]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
corresponds to,
|
||
<taxonomicName id="4C444D6BFF92FFD52610F9EDFDE6F9AE" authorityName="Smales" authorityYear="2016" box="[381,621,1547,1570]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="species" species="aenigma">
|
||
<emphasis id="B930EAFAFF92FFD52610F9EDFDE6F9AE" box="[381,621,1547,1570]" italics="true" pageId="19" pageNumber="20">Rodentanema aenigma</emphasis>
|
||
</taxonomicName>
|
||
is considered a
|
||
<emphasis id="B930EAFAFF92FFD527E4F9CFFEC4F9CC" box="[137,335,1577,1600]" italics="true" pageId="19" pageNumber="20">species inquirenda</emphasis>
|
||
. Being the type and unique species of the genus, it is impossible to give a precise definition of it. We thus consider
|
||
<taxonomicName id="4C444D6BFF92FFD5265CF981FE34F9F2" authorityName="Smales" authorityYear="2016" box="[305,447,1639,1662]" class="Chromadorea" family="Heligmosomidae" genus="Rodentanema" kingdom="Animalia" order="Rhabditida" pageId="19" pageNumber="20" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B930EAFAFF92FFD5265CF981FE34F9F2" box="[305,447,1639,1662]" italics="true" pageId="19" pageNumber="20">Rodentanema</emphasis>
|
||
</taxonomicName>
|
||
a
|
||
<emphasis id="B930EAFAFF92FFD526B6F981FD28F9F2" box="[475,675,1639,1662]" italics="true" pageId="19" pageNumber="20">genus inquirendum</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |