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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1123.90141" ID-GBIF-Dataset="d6d98bd3-2433-44ff-b560-dffef5ebd40e" ID-Pensoft-Pub="1313-2970-1123-1" ID-Pensoft-UUID="5378DBF1DEED5922910377DB1774B3B4" ID-ZooBank="B80C686A001C4A729E8A7DED3FEE9515" ModsDocID="1313-2970-1123-1" checkinTime="1664474094714" checkinUser="pensoft" docAuthor="Schneider, Scott A. &amp; LaPolla, John S." docDate="2022" docId="62993AF42C175FD4A2680988B45DC7A7" docLanguage="en" docName="ZooKeys 1123: 1-30" docOrigin="ZooKeys 1123" docPubDate="2022-09-29" docSource="http://dx.doi.org/10.3897/zookeys.1123.90141" docTitle="Ripersiella colombiensis" docType="treatment" docVersion="2" id="5378DBF1DEED5922910377DB1774B3B4" lastPageNumber="1" masterDocId="5378DBF1DEED5922910377DB1774B3B4" masterDocTitle="A Neotropical complex of Ripersiella species (Hemiptera, Coccomorpha, Rhizoecidae) collected from the nests of Acropyga ants (Hymenoptera, Formicidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="1" updateTime="1664474713734" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A Neotropical complex of Ripersiella species (Hemiptera, Coccomorpha, Rhizoecidae) collected from the nests of Acropyga ants (Hymenoptera, Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schneider, Scott A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9679-8437</mods:nameIdentifier>
<mods:affiliation>United States Department of Agriculture, Agricultural Research Service, Henry A. Wallace Beltsville Agricultural Research Center, Systematic Entomology Laboratory, 10300 Baltimore Avenue, Beltsville, MD, 20705, USA</mods:affiliation>
<mods:nameIdentifier type="email">scott.schneider@ars.usda.gov</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>LaPolla, John S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7602-1527</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, Towson University, 8000 York Road, Towson, MD, 21252, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>30</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1123.90141</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1123.90141</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1123-1</mods:identifier>
<mods:identifier type="ZooBank">B80C686A001C4A729E8A7DED3FEE9515</mods:identifier>
<mods:identifier type="Pensoft-UUID">5378DBF1DEED5922910377DB1774B3B4</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:62993AF42C175FD4A2680988B45DC7A7" httpUri="http://treatment.plazi.org/id/62993AF42C175FD4A2680988B45DC7A7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="62993AF4-2C17-5FD4-A268-0988B45DC7A7" authority="(Hambleton)" baseAuthorityName="Hambleton" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">Ripersiella colombiensis (Hambleton)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ripersiella colombiensis (Hambleton). Adult female, full body view, re-illustrated by JSL from Williams and Granara de Willink (1992) and Kozar and Konczne Benedicty (2007), with edits by SAS." figureDoi="10.3897/zookeys.1123.90141.figure4" httpUri="https://binary.pensoft.net/fig/750175" pageId="0" pageNumber="1">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Hambleton" authorityYear="1946" class="Insecta" family="Rhizoecidae" genus="Neorhizoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neorhizoecus colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">Neorhizoecus colombiensis</taxonomicName>
Hambleton, 1946: 43.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Hambleton" baseAuthorityYear="1946" class="Insecta" family="Rhizoecidae" genus="Rhizoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhizoecus colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">Rhizoecus colombiensis</taxonomicName>
(Hambleton);
<bibRefCitation author="Hambleton, EJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="367 - 376" refId="B11" refString="Hambleton, EJ, 1977. Notes on the species of Neorhizoecus Hambleton, a synonym of Rhizoecus Kuenckel D'Herculais (Homoptera: Pseudococcidae). Proceedings of the Entomological Society of Washington 79: 367 - 376" title="Notes on the species of Neorhizoecus Hambleton, a synonym of Rhizoecus Kuenckel D'Herculais (Homoptera: Pseudococcidae)." volume="79" year="1977">Hambleton 1977</bibRefCitation>
: 372.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">Ripersiella colombiensis</taxonomicName>
(Hambleton);
<bibRefCitation author="Kozar, F" journalOrPublisher="Bollettino di Zoologia Agraria e di Bachicoltura (Milano)" pageId="0" pageNumber="1" pagination="225 - 239" refId="B14" refString="Kozar, F, Konczne Benedicty, Z, 2003. Description of four new species from Australian, Austro-oriental, New Zealand and South Pacific regions (Homoptera, Coccoidea, Pseudococcidae, Rhizoecinae), with a review, and a key to the species Ripersiella. Bollettino di Zoologia Agraria e di Bachicoltura (Milano) 35: 225 - 239" title="Description of four new species from Australian, Austro-oriental, New Zealand and South Pacific regions (Homoptera, Coccoidea, Pseudococcidae, Rhizoecinae), with a review, and a key to the species Ripersiella." volume="35" year="2003">
<normalizedToken originalValue="Kozár">Kozar</normalizedToken>
and
<normalizedToken originalValue="Konczné">Konczne</normalizedToken>
Benedicty 2003
</bibRefCitation>
: 236.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1936-02" collectorName="La Esperanza, R Roba" country="Colombia" location="Colombia" specimenCount="1" specimenCount-adult="1" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Colombia">Colombia</collectingCountry>
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount></specimenCount>
;
<collectorName>La Esperanza</collectorName>
;
<collectingDate value="1936-02">ii.1936</collectingDate>
;
<collectorName>R Roba</collectorName>
coll.; USNM
</materialsCitation>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Other material</emphasis>
.
</emphasis>
<materialsCitation collectingDate="2001-08-05" collectingDateMax="2001-08-15" collectingDateMin="2001-08-05" collectorName="Chiricahua Mtns, SW, W. Portal, JS LaPolla" country="United States" county="Cochise Co." elevation="1646" latitude="31.8833" location="Cochise Co." longLatPrecision="6" longitude="-109.2063" specimenCount="1" specimenCount-adult="1" stateProvince="Arizona">
<collectingCountry name="United States of America">United States</collectingCountry>
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount></specimenCount>
;
<collectingRegion country="United States of America" name="Arizona">Arizona</collectingRegion>
,
<collectingCounty>Cochise Co.</collectingCounty>
,
<collectorName>Chiricahua Mtns, SW</collectorName>
Res. Sta.,
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.04672" unit="mi" value="5.0">5 miles</quantity>
<collectorName>W. Portal</collectorName>
;
<geoCoordinate degrees="31.8833" direction="north" orientation="latitude" precision="5" value="31.8833">31.8833</geoCoordinate>
,
<geoCoordinate degrees="109.2063" direction="west" orientation="longitude" precision="5" value="-109.2063">-109.2063</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.646" unit="m" value="1646.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.646" unit="m" value="1646.0">1646 m</elevation>
</quantity>
;
<collectingDate value="2001-08-05" valueMax="2001-08-15" valueMin="2001-08-05">5-15.viii.2001</collectingDate>
;
<collectorName>JS LaPolla</collectorName>
; with
<taxonomicName authorityName="Snelling" authorityYear="1973" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga epedana" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="epedana">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga epedana</emphasis>
</taxonomicName>
; USNM
</materialsCitation>
<materialsCitation collectingDate="2005-07-31" collectorName="CR Smith" country="United States" county="Cochise Co." elevation="1645" latitude="31.8838" location="Portal" longLatPrecision="6" longitude="-109.2229" specimenCount="1" specimenCount-adult="1" stateProvince="Arizona">
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount></specimenCount>
;
<collectingRegion country="United States of America" name="Arizona">Arizona</collectingRegion>
,
<collectingCounty>Cochise Co.</collectingCounty>
, near
<location LSID="urn:lsid:plazi:treatment:62993AF42C175FD4A2680988B45DC7A7:6813E2584FAF1D0DCB3C8C6346CABA87" country="United States" county="Cochise Co." latitude="31.8838" longLatPrecision="6" longitude="-109.2229" name="Portal" stateProvince="Arizona">Portal</location>
;
<geoCoordinate degrees="31.8838" direction="north" orientation="latitude" precision="5" value="31.8838">31.8838</geoCoordinate>
,
<geoCoordinate degrees="109.2229" direction="west" orientation="longitude" precision="5" value="-109.2229">-109.2229</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.645" unit="m" value="1645.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.645" unit="m" value="1645.0">1645 m</elevation>
</quantity>
;
<collectingDate value="2005-07-31">31.vii.2005</collectingDate>
;
<collectorName>CR Smith</collectorName>
; collected from colony of
<taxonomicName authorityName="Snelling" authorityYear="1973" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga epedana" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="epedana">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga epedana</emphasis>
</taxonomicName>
; USNM
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1123.90141.figure4" httpUri="https://binary.pensoft.net/fig/750175" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella colombiensis</emphasis>
</taxonomicName>
(Hambleton). Adult female, full body view, re-illustrated by JSL from
<bibRefCitation author="Williams, DJ" journalOrPublisher="CAB International, Wallingford" pageId="0" pageNumber="1" refId="B33" refString="Williams, DJ, Granara de Willink, MC, 1992. Mealybugs of Central and South America. CAB International, Wallingford" title="Mealybugs of Central and South America." year="1992">Williams and Granara de Willink (1992)</bibRefCitation>
and
<bibRefCitation author="Kozar, F" journalOrPublisher="Plant Protection Institute, Hungarian Academy of Sciences, Budapest" pageId="0" pageNumber="1" refId="B15" refString="Kozar, F, Konczne Benedicty, Z, 2007. Rhizoecinae of the World. Plant Protection Institute, Hungarian Academy of Sciences, Budapest" title="Rhizoecinae of the World." year="2007">
<normalizedToken originalValue="Kozár">Kozar</normalizedToken>
and
<normalizedToken originalValue="Konczné">Konczne</normalizedToken>
Benedicty (2007)
</bibRefCitation>
, with edits by SAS.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Adult female, based on holotype. Appearance in life not reported; extent of wax production unknown.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">General.</emphasis>
Mounted on microscope slide, body broadly oval and membranous, 1.09 mm long, widest at metathorax and abdominal segments I-II, 0.91 mm wide. Abdomen rounded and gently tapering toward posterior apex; abdominal segment VIII approximately 270
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at base. Anal lobes poorly developed with several flagellate setae on dorsum and venter, 18-28
<normalizedToken originalValue="μm">μm</normalizedToken>
long; lacking differentiated group of 3 long anal lobe setae. Body setae flagellate, 17-25
<normalizedToken originalValue="μm">μm</normalizedToken>
on head, 15-20
<normalizedToken originalValue="μm">μm</normalizedToken>
on thorax, 16-21
<normalizedToken originalValue="μm">μm</normalizedToken>
on abdominal segments. Trilocular pores scarcely distributed among body setae; multilocular disc pores absent; bitubular cerores absent; oral collar tubular ducts absent. Microtrichia present on abdominal segments and thorax. Eyes absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Venter</emphasis>
.
</emphasis>
Cephalic plate absent. Labium with 3 segments; 88-110
<normalizedToken originalValue="μm">μm</normalizedToken>
long and 69
<normalizedToken originalValue="μm">μm</normalizedToken>
at widest point. Antennae geniculate, 5-segmented, closely situated near midline on ventral submargin of head; overall length 182
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of segment I: 36
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment II: 26
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment III: 28
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment IV: 22
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment V: 70
<normalizedToken originalValue="μm">μm</normalizedToken>
; apical segment with 1 spine-like and 4 falcate sensory setate; flagellate setae on each antennal segment 18-40
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Legs well developed; overall length of hind leg 394
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind coxa 48
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind trochanter + femur 158
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind tibia + tarsus 152
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind claw 36
<normalizedToken originalValue="μm">μm</normalizedToken>
; each claw with setose digitule 4
<normalizedToken originalValue="μm">μm</normalizedToken>
long; flagellate or stout setae present on each segment 18-28
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Single conical circulus present between abdominal segments III and IV, 28
<normalizedToken originalValue="μm">μm</normalizedToken>
wide; inner margin of orifice slightly crenulated.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Dorsum</emphasis>
.
</emphasis>
Anal ring situated on dorsal surface separated from posterior body margin by approximately 1
<normalizedToken originalValue="×">x</normalizedToken>
diameter of ring, 80
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter; with oval cells lacking spicules; bearing three pairs of setae about 63
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Posterior pair of ostioles present, diameter of orifice measured along longitudinal axis approximately 68
<normalizedToken originalValue="μm">μm</normalizedToken>
; anterior pair of ostioles apparently absent.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella colombiensis</emphasis>
</taxonomicName>
is most similar in appearance to the newly described species
<taxonomicName lsidName="R. illicians" pageId="0" pageNumber="1" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">R. illicians</emphasis>
</taxonomicName>
. Consult the diagnosis of
<taxonomicName lsidName="R. illicians" pageId="0" pageNumber="1" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">R. illicians</emphasis>
</taxonomicName>
for a discussion on how to distinguish them.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella colombiensis" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella colombiensis</emphasis>
</taxonomicName>
is confirmed to associate with
<taxonomicName authorityName="Snelling" authorityYear="1973" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga epedana" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="epedana">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga epedana</emphasis>
</taxonomicName>
and is one of few species of
<taxonomicName authorityName="Williams" authorityYear="1969" class="Insecta" family="Rhizoecidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Rhizoecidae</taxonomicName>
with a published record of
<taxonomicName authorityName="Roger" authorityYear="1862" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga</emphasis>
</taxonomicName>
queens carrying gravid female trophobionts on their nuptial flight (
<bibRefCitation DOI="https://doi.org/10.1673/031.007.1101" author="Smith, CR" journalOrPublisher="Journal of Insect Science" pageId="0" pageNumber="1" pagination="1 - 5" refId="B26" refString="Smith, CR, Oettler, J, Kay, A, Deans, C, 2007. First recorded mating flight of the hypogeic ant, Acropyga epedana, with its obligate mutualist mealybug, Rhizoecus colombiensis. Journal of Insect Science 7 (11): 1 - 5, DOI: https://doi.org/10.1673/031.007.1101" title="First recorded mating flight of the hypogeic ant, Acropyga epedana, with its obligate mutualist mealybug, Rhizoecus colombiensis." url="https://doi.org/10.1673/031.007.1101" volume="7" year="2007">Smith et al. 2007</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4853.2.9" author="Schneider, SA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="283 - 291" refId="B24" refString="Schneider, SA, LaPolla, JS, 2020. Trophobiosis between a new species of Williamsrhizoecus (Hemiptera: Coccomorpha: Rhizoecidae) and Acropyga silvestrii (Hymenoptera: Formicidae) in Tanzania. Zootaxa 4853 (2): 283 - 291, DOI: https://doi.org/10.11646/zootaxa.4853.2.9" title="Trophobiosis between a new species of Williamsrhizoecus (Hemiptera: Coccomorpha: Rhizoecidae) and Acropyga silvestrii (Hymenoptera: Formicidae) in Tanzania." url="https://doi.org/10.11646/zootaxa.4853.2.9" volume="4853" year="2020">Schneider and LaPolla 2020</bibRefCitation>
). Specimens from Colombia and Arizona appear likely to be conspecific, although those from Arizona have noticeably longer and thinner legs relative to their body size.
</paragraph>
</subSubSection>
</treatment>
</document>