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<document id="85185D2C43E0A2B938EA3FBE0E939FF4" ID-CLB-Dataset="50579" ID-DOI="10.11646/zootaxa.4097.4.5" ID-GBIF-Dataset="536bad92-81ae-49dc-aeff-ecfb2731274c" ID-ISSN="1175-5326" ID-Zenodo-Dep="259066" ID-ZooBank="CEB0F19C-15FF-4FEF-A334-2E03EB6A077F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459767938066" checkinUser="plazi" docAuthor="Cuccodoro, Giulio &amp; Liu, Zhiping" docDate="2016" docId="621B87E9FFC9FFB374CDFD00BFDEFA71" docLanguage="en" docName="zootaxa.4097.4.5.pdf.imf" docOrigin="Zootaxa 4097 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megarthrus" docType="key" docVersion="12" lastPageNumber="531" masterDocId="9E22FF91FFC8FFB2745AFFB1BA42FFBC" masterDocTitle="Megarthrus of southern India and Sri Lanka, with notes on their phylogenetic and biogeographical relationships (Coleoptera: Staphylinidae: Proteininae)" masterLastPageNumber="544" masterPageNumber="530" pageNumber="531" updateTime="1698223148651" updateUser="ExternalLinkService">
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<mods:title id="E8BDA7E50829DE72E00D924FD1E9ADE5">Megarthrus of southern India and Sri Lanka, with notes on their phylogenetic and biogeographical relationships (Coleoptera: Staphylinidae: Proteininae)</mods:title>
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<mods:namePart id="9AC9100ADC09F13D314FEEBDE9BEC8BF">Cuccodoro, Giulio</mods:namePart>
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<mods:namePart id="196F6B5B43B39BD55E33F6B1FB18B08B">Liu, Zhiping</mods:namePart>
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<mods:date id="6FD1A806BA769F18F5907BF4BD67C7E3">2016</mods:date>
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<treatment id="621B87E9FFC9FFB374CDFD00BFDEFA71" ID-DOI="http://doi.org/10.5281/zenodo.6074537" ID-GBIF-Taxon="119346548" ID-Zenodo-Dep="6074537" LSID="urn:lsid:plazi:treatment:621B87E9FFC9FFB374CDFD00BFDEFA71" httpUri="http://treatment.plazi.org/id/621B87E9FFC9FFB374CDFD00BFDEFA71" lastPageNumber="531" pageId="1" pageNumber="531">
<subSubSection id="A2A86574FFC9FFB374CDFD00B9AFFD77" box="[151,1005,689,715]" pageId="1" pageNumber="531" type="nomenclature">
<paragraph id="EA0D36FFFFC9FFB374CDFD00B9AFFD77" blockId="1.[151,1005,689,715]" box="[151,1005,689,715]" pageId="1" pageNumber="531">
<heading id="B1458193FFC9FFB374CDFD00B9AFFD77" bold="true" box="[151,1005,689,715]" fontSize="11" level="1" pageId="1" pageNumber="531" reason="1">
<emphasis id="D8C6EAEDFFC9FFB374CDFD00B9AFFD77" bold="true" box="[151,1005,689,715]" pageId="1" pageNumber="531">
Key to the species of
<taxonomicName id="2DB24D7CFFC9FFB375C6FD00B869FD77" ID-CoL="8VRTJ" box="[412,555,689,715]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="D8C6EAEDFFC9FFB375C6FD00B869FD77" bold="true" box="[412,555,689,715]" italics="true" pageId="1" pageNumber="531">Megarthrus</emphasis>
</taxonomicName>
from southern
<collectingCountry id="92A5766FFFC9FFB376B7FD00B972FD77" box="[749,816,689,715]" name="India" pageId="1" pageNumber="531">India</collectingCountry>
and
<collectingCountry id="92A5766FFFC9FFB37734FD00B9AFFD77" box="[878,1005,689,715]" name="Sri Lanka" pageId="1" pageNumber="531">
Sri
<taxonomicName id="2DB24D7CFFC9FFB377C1FD00B9AFFD77" box="[923,1005,689,715]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="543" phylum="Arthropoda" rank="species" species="lanka">Lanka</taxonomicName>
</collectingCountry>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A2A86574FFC9FFB374CDFD46BFDEFA71" pageId="1" pageNumber="531" type="key">
<keyStep id="51462E5AFFC9FFB374CDFD46BFD9FC23" pageId="1" pageNumber="531">
<paragraph id="EA0D36FFFFC9FFB374CDFD46BFD9FCFB" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFD46BFD9FCFB" pageId="1" pageNumber="531">
1 Body predominantly rust brown or chestnut brown; elytra unicolored; antennomeres 9 and 10 strongly asymmetrical (fig. 63); frons with lateral outlines oblique to apex broadly rounded, or subtruncate. Male: protarsomeres 5 with conspicuous adventral apophysis; mesotibiae markedly arcuate (fig. 56) (
<taxonomicName id="2DB24D7CFFC9FFB37699FC80B915FCFB" ID-CoL="3YXC2" box="[707,855,817,839]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="rufomarginatus">
<emphasis id="D8C6EAEDFFC9FFB37699FC80B915FCFB" box="[707,855,817,839]" italics="true" pageId="1" pageNumber="531">rufomarginatus</emphasis>
</taxonomicName>
-complex)....................................... 2
</keyLead>
</paragraph>
<paragraph id="EA0D36FFFFC9FFB374CDFCFEBFD9FC23" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFCFEBFD9FC23" pageId="1" pageNumber="531">
- Body predominantly yellow brown; elytra bicoloured; antennomeres 9 and 10 symmetrical; frons with lateral outlines markedly concave to apex truncate. Male: protarsomeres 5 without adventral apophysis; mesotibiae fairly straight (
<figureCitation id="72892A7AFFC9FFB3710BFCDDBFD6FC3E" box="[1361,1428,876,898]" captionStart="FIGURES 15 22" captionStartId="5.[151,264,1377,1400]" captionTargetBox="[343,1235,201,1347]" captionTargetId="figure@5.[340,1247,193,1357]" captionText="FIGURES 15 22. Megarthrus narendrani sp. n., male; aedeagus in ventral (15, 18) and lateral (16, 17) views; abdominal sternite VIII in ventral view (19); apex of abdominal tergite 8 in dorsal view (22); mesotibia (21); mesofemur and mesotrochanter (22). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/291814/files/figure.png" pageId="1" pageNumber="531">fig. 20</figureCitation>
) (
<taxonomicName id="2DB24D7CFFC9FFB374B3FC38BB1CFC23" ID-CoL="3YX8S" box="[233,350,905,927]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="D8C6EAEDFFC9FFB374B3FC38BB1CFC23" box="[233,350,905,927]" italics="true" pageId="1" pageNumber="531">bimaculatus</emphasis>
</taxonomicName>
-complex)................................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51462E5AFFC9FFB374CDFC16BFDEFB79" pageId="1" pageNumber="531">
<paragraph id="EA0D36FFFFC9FFB374CDFC16BFD9FB8E" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFC16BFD9FB8E" pageId="1" pageNumber="531">
2 Body chestnut brown, except legs and antennae paler; antennae unicoloured. Male: aedeagus (figs 6668) with sclerotized dorsal valve longer than half the distance from its posterior edge to the aedeagal tip; internal sac with a very large subbasal sclerite; abdominal sternite V with medial area slightly swollen and bearing a short transverse ridge. Female: valvifers in dorsal view gradually tapering, with lateral outline gently convex (fig. 71); dorsal outline of valvifers almost straight (fig. 72); genital tergites with a medial macula cordiform (fig. 70)...........................
<taxonomicName id="2DB24D7CFFC9FFB37012FBADBF16FB8E" ID-CoL="3YXC2" box="[1096,1364,1052,1074]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="rufomarginatus">
<emphasis id="D8C6EAEDFFC9FFB37012FBADBF16FB8E" box="[1096,1364,1052,1074]" italics="true" pageId="1" pageNumber="531">Megarthrus rufomarginatus</emphasis>
</taxonomicName>
(fig. 2)
</keyLead>
</paragraph>
<paragraph id="EA0D36FFFFC9FFB374CDFB8BBFDEFB79" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFB8BBFDEFB79" pageId="1" pageNumber="531">
- Body rust brown, concolorous with legs and antennae, except antennomeres 11 markedly paler than the others. Male: aedeagus (figs 4851) with sclerotized dorsal valve not longer than half the distance from its posterior edge to the aedeagal tip; internal sac with a subbasal group of strong spines; abdominal sternite V with medial area slightly swollen and bearing a short transverse ridge. Female: valvifers in dorsal view evenly tapering, with lateral outline almost straight (fig. 58); dorsal outline of valvifers angulate subapically (fig. 59); genital tergites with a medial macula oval (fig. 60).....
<taxonomicName id="2DB24D7CFFC9FFB370FDFB1EBF11FB79" ID-CoL="72VVF" box="[1191,1363,1199,1221]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="lanka">
<emphasis id="D8C6EAEDFFC9FFB370FDFB1EBF11FB79" box="[1191,1363,1199,1221]" italics="true" pageId="1" pageNumber="531">Megarthrus lanka</emphasis>
</taxonomicName>
(fig. 1)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51462E5AFFC9FFB374CDFB7DBFD9FA86" pageId="1" pageNumber="531">
<paragraph id="EA0D36FFFFC9FFB374CDFB7DBFDEFB43" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFB7DBFDEFB43" pageId="1" pageNumber="531">
<specimenCount id="FCB4FD76FFC9FFB374CDFB7DBB54FB5E" box="[151,278,1228,1250]" pageId="1" pageNumber="531" type="male">3 Male</specimenCount>
: aedeagus with ventral wall subapically right angled, apical portion conspicuously projecting ventrally (figs 32, 33). Female: valvifers each with a dorsal apophysis locate at level of dorsobasal margin (fig. 41) …
<taxonomicName id="2DB24D7CFFC9FFB3702BFB58BF16FB43" ID-CoL="3YXBB" box="[1137,1364,1257,1279]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="nilgiriensis">
<emphasis id="D8C6EAEDFFC9FFB3702BFB58BF16FB43" box="[1137,1364,1257,1279]" italics="true" pageId="1" pageNumber="531">Megarthrus nilgiriensis</emphasis>
</taxonomicName>
(fig. 5)
</keyLead>
</paragraph>
<paragraph id="EA0D36FFFFC9FFB374CDFAB6BFD9FA86" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFAB6BFD9FA86" pageId="1" pageNumber="531">- Male: aedeagus with ventral wall subapically not right angled. Female: valvifers each with a dorsal apophysis located near midlength (figs 12, 24)................................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="51462E5AFFC9FFB374CDFAF0BFDEFA71" pageId="1" pageNumber="531">
<paragraph id="EA0D36FFFFC9FFB374CDFAF0BFD9FA2E" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFAF0BFD9FA2E" pageId="1" pageNumber="531">
<specimenCount id="FCB4FD76FFC9FFB374CDFAF0BB54FAEB" box="[151,278,1345,1367]" pageId="1" pageNumber="531" type="male">4 Male</specimenCount>
: aedeagus with ventral outline of ventral wall sinuate to tip truncate (
<figureCitation id="72892A7AFFC9FFB377E9FAF0BE5CFAEB" box="[947,1054,1345,1367]" captionStart="FIGURES 15 22" captionStartId="5.[151,264,1377,1400]" captionTargetBox="[343,1235,201,1347]" captionTargetId="figure@5.[340,1247,193,1357]" captionText="FIGURES 15 22. Megarthrus narendrani sp. n., male; aedeagus in ventral (15, 18) and lateral (16, 17) views; abdominal sternite VIII in ventral view (19); apex of abdominal tergite 8 in dorsal view (22); mesotibia (21); mesofemur and mesotrochanter (22). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/291814/files/figure.png" pageId="1" pageNumber="531">figs 16, 17</figureCitation>
). Female: lateral outline of middle portion of valvifers converging (fig. 23); female: genital tergites with two transverse subbasal sclerites (fig. 25).........................................................................................
<taxonomicName id="2DB24D7CFFC9FFB3702BFACDBF11FA2E" ID-CoL="3YXB6" box="[1137,1363,1404,1426]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="narendrani">
<emphasis id="D8C6EAEDFFC9FFB3702BFACDBF11FA2E" box="[1137,1363,1404,1426]" italics="true" pageId="1" pageNumber="531">Megarthrus narendrani</emphasis>
</taxonomicName>
(fig. 4)
</keyLead>
</paragraph>
<paragraph id="EA0D36FFFFC9FFB374CDFA2BBFDEFA71" blockId="1.[151,1436,759,1485]" pageId="1" pageNumber="531">
<keyLead id="514395CAFFC9FFB374CDFA2BBFDEFA71" pageId="1" pageNumber="531">
- Male: aedeagus with ventral outline of ventral wall arcuate to tip blunt (figs 7, 8). Female: lateral outline of middle portion of valvifers parallel (fig. 11); female: genital tergites without subbasal sclerites (fig. 3).......
<taxonomicName id="2DB24D7CFFC9FFB3703DFA06BF11FA71" ID-CoL="3YX8S" box="[1127,1363,1463,1485]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="531" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="D8C6EAEDFFC9FFB3703DFA06BF11FA71" box="[1127,1363,1463,1485]" italics="true" pageId="1" pageNumber="531">Megarthrus bimaculatus</emphasis>
</taxonomicName>
(fig. 3)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>