treatments-xml/data/03/A7/D7/03A7D746F075FFA7FF3DFE7AFCFEFA00.xml

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<mods:title id="7FE7566E11938893936F0933406F0E93">Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda</mods:title>
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<mods:namePart id="1C01D6000D5D172F847D67E24237F8B0">Mandic, Oleg</mods:namePart>
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<treatment id="03A7D746F075FFA7FF3DFE7AFCFEFA00" ID-DOI="http://doi.org/10.5281/zenodo.5076343" ID-GBIF-Taxon="183399417" ID-Zenodo-Dep="5076343" LSID="urn:lsid:plazi:treatment:03A7D746F075FFA7FF3DFE7AFCFEFA00" httpUri="http://treatment.plazi.org/id/03A7D746F075FFA7FF3DFE7AFCFEFA00" lastPageNumber="460" pageId="7" pageNumber="460" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_order="Littorinimorpha" scope_subClass="Caenogastropoda" scope_subFamily="Micromelaniinae" scope_superFamily="Rissooidea">
<subSubSection id="C31435DBF075FFA7FF3DFE7AFDD8FE36" box="[151,611,402,428]" pageId="7" pageNumber="460" type="nomenclature">
<paragraph id="8BB16650F075FFA7FF3DFE7AFDD8FE36" blockId="7.[151,611,402,428]" box="[151,611,402,428]" pageId="7" pageNumber="460">
<heading id="D0F9D13CF075FFA7FF3DFE7AFDD8FE36" bold="true" box="[151,611,402,428]" fontSize="11" level="1" pageId="7" pageNumber="460" reason="1">
<emphasis id="B97ABA42F075FFA7FF3DFE7AFDD8FE36" bold="true" box="[151,611,402,428]" pageId="7" pageNumber="460">
Genus
<taxonomicName id="4C0E1DD3F075FFA7FF47FE7AFDD8FE36" authority="Sandberger, 1875" authorityName="Sandberger" authorityYear="1875" box="[237,611,402,428]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="genus">
<emphasis id="B97ABA42F075FFA7FF47FE7AFE3AFE36" bold="true" box="[237,385,402,428]" italics="true" pageId="7" pageNumber="460">Goniochilus</emphasis>
Sandberger, 1875
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31435DBF075FFA7FF3DFE31FCB8FD1B" pageId="7" pageNumber="460" type="type_taxon">
<paragraph id="8BB16650F075FFA7FF3DFE31FF52FD8C" blockId="7.[151,1436,473,534]" pageId="7" pageNumber="460">
<materialsCitation id="3B666C0DF075FFA7FF3DFE31FF5FFD8C" ID-GBIF-Occurrence="3314665315" country="Romania" county="Wenz" location="Late Miocene" municipality="Subsequent" pageId="7" pageNumber="460" specimenCount="1" typeStatus="holotype">
<emphasis id="B97ABA42F075FFA7FF3DFE31FE82FE68" bold="true" box="[151,313,473,498]" pageId="7" pageNumber="460">
<typeStatus id="54B5D8F2F075FFA7FF3DFE31FF69FE68" box="[151,210,473,498]" pageId="7" pageNumber="460">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C0E1DD3F075FFA7FEEEFE31FD5FFE68" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[324,740,473,498]" class="Gastropoda" family="Pleuroceridae" genus="Pleurocera" kingdom="Animalia" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="costulatum">
<emphasis id="B97ABA42F075FFA7FEEEFE31FDFCFE68" box="[324,583,473,498]" italics="true" pageId="7" pageNumber="460">Pleurocera costulatum</emphasis>
Fuchs, 1870
</taxonomicName>
.
<location id="8ED1308BF075FFA7FD59FE32FC2EFE68" LSID="urn:lsid:plazi:treatment:03A7D746F075FFA7FF3DFE7AFCFEFA00:8ED1308BF075FFA7FD59FE32FC2EFE68" box="[755,917,473,498]" country="Romania" county="Wenz" municipality="Subsequent" name="Late Miocene" pageId="7" pageNumber="460">Late Miocene</location>
,
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.
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designation by
<collectingCounty id="62D01EDCF075FFA7FAF7FE32FA27FE6B" box="[1373,1436,474,497]" pageId="7" pageNumber="460">Wenz</collectingCounty>
(1926)
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.
</paragraph>
<paragraph id="8BB16650F075FFA7FF3DFD81FCB8FD1B" blockId="7.[151,771,617,642]" box="[151,771,617,642]" pageId="7" pageNumber="460">
<emphasis id="B97ABA42F075FFA7FF3DFD81FDF7FD18" bold="true" box="[151,588,617,642]" pageId="7" pageNumber="460">
&quot;
<taxonomicName id="4C0E1DD3F075FFA7FF0CFD81FDFBFD18" authority="Pana, 1990" authorityName="Pana" authorityYear="1990" box="[166,576,617,642]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="species" species="lecrenthey">
<emphasis id="B97ABA42F075FFA7FF0CFD81FE08FD18" bold="true" box="[166,435,617,642]" italics="true" pageId="7" pageNumber="460">Goniochilus lecrenthey</emphasis>
<bibRefCitation id="EF9F1BA1F075FFA7FE10FD81FDFBFD18" author="Pana, I." box="[442,576,617,642]" pageId="7" pageNumber="460" pagination="61 - 69" refId="ref10948" refString="Pana, I. (1990) Nouvelles especes des nannogastropodes pontiens. Revue Roumaine de Geologie, Geophysique et Geographie, 34, 61 - 69." type="journal article" year="1990">Pană, 1990</bibRefCitation>
</taxonomicName>
&quot;
</emphasis>
[
<emphasis id="B97ABA42F075FFA7FDF6FD82FD41FD18" box="[604,762,617,642]" italics="true" pageId="7" pageNumber="460">nomen nudum</emphasis>
]
</paragraph>
</subSubSection>
<subSubSection id="C31435DBF075FFA7FF3DFD59FCFEFA00" pageId="7" pageNumber="460" type="discussion">
<paragraph id="8BB16650F075FFA7FF3DFD59FBDFFC38" blockId="7.[151,1437,689,1434]" pageId="7" pageNumber="460">
<emphasis id="B97ABA42F075FFA7FF3DFD59FEA7FD50" bold="true" box="[151,284,689,714]" inLineHeading="true" pageId="7" pageNumber="460" reason="1">Discussion.</emphasis>
There are several problems with this species. The first and most crucial point is that it is formally a
<emphasis id="B97ABA42F075FFA7FF3DFD3EFE8EFD74" box="[151,309,725,750]" italics="true" pageId="7" pageNumber="460">nomen nudum</emphasis>
. While
<bibRefCitation id="EF9F1BA1F075FFA7FE3AFD3EFDA3FD74" author="Pana, I." box="[400,536,725,750]" pageId="7" pageNumber="460" pagination="61 - 69" refId="ref10948" refString="Pana, I. (1990) Nouvelles especes des nannogastropodes pontiens. Revue Roumaine de Geologie, Geophysique et Geographie, 34, 61 - 69." type="journal article" year="1990">Pană (1990)</bibRefCitation>
introduced two new subspecies, she did not describe the species itself. The two subspecies are still validly introduced as species-group names. Secondly, there are many misspellings (lecrenthey, loerenthey, loeerenthey) in the publication. Concerning the
<emphasis id="B97ABA42F075FFA7FC90FCF5FBBBFCAF" box="[826,1024,797,822]" italics="true" pageId="7" pageNumber="460">derivatio nominis</emphasis>
section and considering the rules of the ICZN the name should be &quot;
<emphasis id="B97ABA42F075FFA7FDB8FCA9FD39FCC0" box="[530,642,833,858]" italics="true" pageId="7" pageNumber="460">lorentheyi</emphasis>
&quot;. According to the
<bibRefCitation id="EF9F1BA1F075FFA7FCD9FCA9FBBFFCC0" author="ICZN" box="[883,1028,833,858]" pageId="7" pageNumber="460" refId="ref9970" refString="ICZN (1999) International Code of Zoological Nomenclature. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999</bibRefCitation>
, Articles 32.5.1 and 33.2.2), these incorrect original spellings (being either
<emphasis id="B97ABA42F075FFA7FDF7FC8DFD4CFCE4" box="[605,759,869,894]" italics="true" pageId="7" pageNumber="460">lapsus calami</emphasis>
or copyist's or printer's errors) would have to be corrected— which is not necessary here, as the name is a
<emphasis id="B97ABA42F075FFA7FD3BFC62FC94FC38" box="[657,815,905,930]" italics="true" pageId="7" pageNumber="460">nomen nudum</emphasis>
and formally not available.
</paragraph>
<paragraph id="8BB16650F075FFA7FF6DFC45FB76FAEC" blockId="7.[151,1437,689,1434]" pageId="7" pageNumber="460">
A similar case is mentioned by
<bibRefCitation id="EF9F1BA1F075FFA7FD85FC45FC96FC5C" author="Welter-Schultes, F. W." box="[559,813,941,966]" pageId="7" pageNumber="460" refId="ref12081" refString="Welter-Schultes, F. W. (2012) European non-marine molluscs, a guide for species identification. Planet Poster Editions, Gottingen, 679 pp." type="book" year="2012">Welter-Schultes (2012</bibRefCitation>
, p. 382) for the terrestrial snail
<taxonomicName id="4C0E1DD3F075FFA7FB3CFC45FE97FC70" authority="(Beck, 1837)" baseAuthorityName="Beck" baseAuthorityYear="1837" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="draparnaudi">
<emphasis id="B97ABA42F075FFA7FB3CFC45FA27FC5C" box="[1174,1436,941,966]" italics="true" pageId="7" pageNumber="460">Oxychilus draparnaudi</emphasis>
(Beck, 1837)
</taxonomicName>
. This species was neither described nor indicated in the original publication (Beck 1837, p. 6), while the varieties
<emphasis id="B97ABA42F075FFA7FE8FFC1DFEC8FB94" box="[293,371,1013,1038]" italics="true" pageId="7" pageNumber="460">gallica</emphasis>
and
<emphasis id="B97ABA42F075FFA7FE03FC1DFE4AFB94" box="[425,497,1013,1038]" italics="true" pageId="7" pageNumber="460">italica</emphasis>
were indeed indicated with bibliographic references.
<bibRefCitation id="EF9F1BA1F075FFA7FBE0FC1DFAF6FB94" author="Welter-Schultes, F. W." box="[1098,1357,1013,1038]" pageId="7" pageNumber="460" refId="ref12081" refString="Welter-Schultes, F. W. (2012) European non-marine molluscs, a guide for species identification. Planet Poster Editions, Gottingen, 679 pp." type="book" year="2012">Welter-Schultes (2012)</bibRefCitation>
argues that the name
<taxonomicName id="4C0E1DD3F075FFA7FEEFFBF1FE68FBA8" baseAuthorityName="Beck" baseAuthorityYear="1837" box="[325,467,1049,1074]" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="draparnaudi">
<emphasis id="B97ABA42F075FFA7FEEFFBF1FE68FBA8" box="[325,467,1049,1074]" italics="true" pageId="7" pageNumber="460">draparnaudi</emphasis>
</taxonomicName>
was made available under ICZN Article 12.1, because it had an indication for a subordinate taxon that was included originally and this article does not demand a description for the nominal taxon. While it is true that the Code states in the glossary that &quot;a taxon encompasses all taxa of lower rank&quot;, we would like to point out that the situation is a bit more complicated. According to the Principle of Coordination (Article 46), all taxa in the species group are deemed to be simultaneously established at every possible rank within the species group. Therefore, the names
<taxonomicName id="4C0E1DD3F075FFA7FE4BFB26FDF0FB7F" authorityName="Pana" authorityYear="1990" box="[481,587,1230,1253]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="species" species="carinatus">
<emphasis id="B97ABA42F075FFA7FE4BFB26FDF0FB7F" box="[481,587,1230,1253]" italics="true" pageId="7" pageNumber="460">carinatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E1DD3F075FFA7FD2FFB25FD5AFB7C" authorityName="Pana" authorityYear="1990" box="[645,737,1229,1254]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="species" species="nodosus">
<emphasis id="B97ABA42F075FFA7FD2FFB25FD5AFB7C" box="[645,737,1229,1254]" italics="true" pageId="7" pageNumber="460">nodosus</emphasis>
</taxonomicName>
, originally proposed as subspecies of
<emphasis id="B97ABA42F075FFA7FB3CFB25FABDFB7C" box="[1174,1286,1229,1254]" italics="true" pageId="7" pageNumber="460">lorentheyi</emphasis>
, are speciesgroup taxa too and thus not subordinate taxa of
<emphasis id="B97ABA42F075FFA7FD7AFB19FCFAFA90" box="[720,833,1265,1290]" italics="true" pageId="7" pageNumber="460">lorentheyi</emphasis>
in a nomenclatural sense. Hence, their descriptions cannot serve as definition for the undescribed nominal form
<emphasis id="B97ABA42F075FFA7FCEBFAFDFC09FAB4" box="[833,946,1301,1326]" italics="true" pageId="7" pageNumber="460">lorentheyi</emphasis>
. To do so would imply that one or both are synonymized with
<emphasis id="B97ABA42F075FFA7FEDDFAD1FE5CFAC8" box="[375,487,1337,1362]" italics="true" pageId="7" pageNumber="460">lorentheyi</emphasis>
, which would be a taxonomic but not a nomenclatural act. The situation in
<taxonomicName id="4C0E1DD3F075FFA7FA2BFAD2FE9FFAEC" baseAuthorityName="Beck" baseAuthorityYear="1837" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="draparnaudi">
<emphasis id="B97ABA42F075FFA7FA2BFAD2FE9FFAEC" italics="true" pageId="7" pageNumber="460">O. draparnaudi</emphasis>
</taxonomicName>
, in contrast, was solved by
<bibRefCitation id="EF9F1BA1F075FFA7FDF1FAB5FD5CFAEC" author="ICZN" box="[603,743,1373,1398]" pageId="7" pageNumber="460" pagination="77 - 108" refId="ref9924" refString="ICZN (1955) Opinion 336: Addition to the &quot; Official list of specific names in Zoology &quot; of the specific names of one hundred and twenty-two non-marine species of the phylum Mollusca. Bulletin of Zoological Nomenclature, 10, 77 - 108." type="journal article" year="1955">ICZN (1955</bibRefCitation>
, Op. 336), which made the name available.
</paragraph>
<paragraph id="8BB16650F075FFA7FF6DFA69FCFEFA00" blockId="7.[151,1437,689,1434]" box="[199,837,1409,1434]" pageId="7" pageNumber="460">Consequently, the only two taxa available are as follows:</paragraph>
</subSubSection>
</treatment>
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