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data/00/49/C9/0049C912FFE0AE45FF32F8DD38B4F29B.xml
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[sic]
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<emphasis id="B97ABA42F075FFA7FF6DF90AFEF7F961" bold="true" box="[199,332,1762,1787]" inLineHeading="true" pageId="7" pageNumber="460" reason="1">Discussion.</emphasis>
<taxonomicName id="4C0E1DD3F075FFA7FEF7F90AFD41F961" authority="Pana, 1990" authorityName="Pana" authorityYear="1990" box="[349,762,1762,1787]" class="Gastropoda" family="Hydrobiidae" genus="Goniochilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="carinatus">
<taxonomicName id="4C0E1DD3F075FFA7FEF7F90AFD41F961" authority="Pana, 1990" authorityName="Pana" authorityYear="1990" box="[349,762,1762,1787]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="species" species="carinatus">
<emphasis id="B97ABA42F075FFA7FEF7F90AFDD9F960" box="[349,610,1762,1787]" italics="true" pageId="7" pageNumber="460">Goniochilus carinatus</emphasis>
<bibRefCitation id="EF9F1BA1F075FFA7FDD9F90BFD41F961" author="Pana, I." box="[627,762,1762,1787]" pageId="7" pageNumber="460" pagination="61 - 69" refId="ref10948" refString="Pana, I. (1990) Nouvelles especes des nannogastropodes pontiens. Revue Roumaine de Geologie, Geophysique et Geographie, 34, 61 - 69." type="journal article" year="1990">Pană, 1990</bibRefCitation>
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may become a secondary homonym. As
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<emphasis id="B97ABA42F075FFA7FAB9F90AFA27F961" box="[1299,1436,1762,1787]" italics="true" pageId="7" pageNumber="460">Goniochilus</emphasis>
Sandberger, 1875
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@ -156,7 +156,7 @@ is repeatedly treated as subgenus of
<bibRefCitation id="EF9F1BA1F075FFA7FBA0F8A6FB7AF8FD" author="Andrusov, N." box="[1034,1217,1870,1895]" pageId="7" pageNumber="460" pagination="193 - 344" refId="ref8180" refString="Andrusov, N. (1890) Kerchenskiy izvestniak i ego fauna. Zapiski Imperatorskago S. - Petersburgskago Mineralogicheskago Obshchestva, seriya 2, 26, 193 - 344." type="journal article" year="1890">Andrusov, 1890</bibRefCitation>
</taxonomicName>
(see discussion of respective species above). Currently,
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<taxonomicName id="4C0E1DD3F075FFA7FD93F89AFD79F811" authorityName="Sandberger" authorityYear="1875" box="[569,706,1906,1931]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" rank="genus">
<emphasis id="B97ABA42F075FFA7FD93F89AFD79F811" box="[569,706,1906,1931]" italics="true" pageId="7" pageNumber="460">Goniochilus</emphasis>
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is treated as separate genus (see

104
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Genus
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<emphasis id="B97ABA42F075FFA7FF47FE7AFE3AFE36" bold="true" box="[237,385,402,428]" italics="true" pageId="7" pageNumber="460">Goniochilus</emphasis>
Sandberger, 1875
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@ -84,7 +84,7 @@ Sandberger, 1875
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species:
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<taxonomicName id="4C0E1DD3F075FFA7FEEEFE31FD5FFE68" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[324,740,473,498]" class="Gastropoda" family="Pleuroceridae" genus="Pleurocera" kingdom="Animalia" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="costulatum">
<emphasis id="B97ABA42F075FFA7FEEEFE31FDFCFE68" box="[324,583,473,498]" italics="true" pageId="7" pageNumber="460">Pleurocera costulatum</emphasis>
Fuchs, 1870
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@ -103,7 +103,7 @@ designation by
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<emphasis id="B97ABA42F075FFA7FF3DFD81FDF7FD18" bold="true" box="[151,588,617,642]" pageId="7" pageNumber="460">
&quot;
<taxonomicName id="4C0E1DD3F075FFA7FF0CFD81FDFBFD18" authority="Pana, 1990" authorityName="Pana" authorityYear="1990" box="[166,576,617,642]" class="Gastropoda" family="Hydrobiidae" genus="Goniochilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cerithiimorpha" pageId="7" pageNumber="460" phylum="Mollusca" rank="species" species="lecrenthey">
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@ -152,11 +152,11 @@ argues that the name
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was made available under ICZN Article 12.1, because it had an indication for a subordinate taxon that was included originally and this article does not demand a description for the nominal taxon. While it is true that the Code states in the glossary that &quot;a taxon encompasses all taxa of lower rank&quot;, we would like to point out that the situation is a bit more complicated. According to the Principle of Coordination (Article 46), all taxa in the species group are deemed to be simultaneously established at every possible rank within the species group. Therefore, the names
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and
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<emphasis id="B97ABA42F075FFA7FD2FFB25FD5AFB7C" box="[645,737,1229,1254]" italics="true" pageId="7" pageNumber="460">nodosus</emphasis>
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, originally proposed as subspecies of

102
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<taxonomicName id="4C0E1DD3F07AFFA8FF3DFF70FD9DFF28" authority="Pana, 1990" authorityName="Pana" authorityYear="1990" box="[151,550,152,178]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="8" pageNumber="461" rank="species" species="nodosus">
<emphasis id="B97ABA42F07AFFA8FF3DFF70FD9DFF28" bold="true" box="[151,550,152,178]" pageId="8" pageNumber="461">
<emphasis id="B97ABA42F07AFFA8FF3DFF70FE2EFF28" bold="true" box="[151,405,152,178]" italics="true" pageId="8" pageNumber="461">Goniochilus nodosus</emphasis>
<bibRefCitation id="EF9F1BA1F07AFFA8FE37FF70FD9DFF28" author="Pana, I." box="[413,550,152,178]" pageId="8" pageNumber="461" pagination="61 - 69" refId="ref10948" refString="Pana, I. (1990) Nouvelles especes des nannogastropodes pontiens. Revue Roumaine de Geologie, Geophysique et Geographie, 34, 61 - 69." type="journal article" year="1990">Pană, 1990</bibRefCitation>
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<emphasis id="B97ABA42F07AFFA8FF3DFF36FE35FF6E" box="[151,398,222,244]" italics="true" pageId="8" pageNumber="461">Goniochilus loeerenthey</emphasis>
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<taxonomicName id="4C0E1DD3F07AFFA8FE3EFF36FD08FF6E" authority="Pana 1990: 68" authorityName="Pana" authorityPageNumber="68" authorityYear="1990" box="[404,691,222,244]" family="Hydrobiidae" genus="Goniochilus" kingdom="Animalia" order="Cerithiimorpha" pageId="8" pageNumber="461" rank="species" species="nodosus">
[sic]
<emphasis id="B97ABA42F07AFFA8FE62FF36FDA0FF6E" box="[456,539,222,244]" italics="true" pageId="8" pageNumber="461">nodosus</emphasis>
<bibRefCitation id="EF9F1BA1F07AFFA8FD8BFF37FD08FF6E" author="Pana, I." box="[545,691,222,244]" pageId="8" pageNumber="461" pagination="61 - 69" refId="ref10948" refString="Pana, I. (1990) Nouvelles especes des nannogastropodes pontiens. Revue Roumaine de Geologie, Geophysique et Geographie, 34, 61 - 69." type="journal article" year="1990">Pană 1990: 68</bibRefCitation>
@ -125,7 +125,7 @@ III g.
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<emphasis id="B97ABA42F07AFFA8FF6DFE79FEF7FE30" bold="true" box="[199,332,401,426]" inLineHeading="true" pageId="8" pageNumber="461" reason="1">Discussion.</emphasis>
To our knowledge,
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@ -167,7 +167,7 @@ ab.
.
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<taxonomicName id="4C6E8BF71550473FBDDF3142FD0EF8CE" authority="Yablokov-Khnzoryan, 1978: 180" authorityName="Yablokov-Khnzoryan" authorityPageNumber="180" authorityYear="1978" box="[151,707,1853,1877]" genus="Lemnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="264" pageNumber="265" rank="species" species="melanoptera">
<emphasis id="B91A2C661550473FBDDF3142FEA6F8CE" box="[151,363,1853,1877]" italics="true" pageId="264" pageNumber="265">Lemnia melanoptera</emphasis>
<bibRefCitation id="EFFF8D851550473FBC3B3141FD0EF8CE" author="Yablokov-Khnzoryan, S. M." box="[371,707,1853,1877]" pageId="264" pageNumber="265" pagination="180 - 183" refId="ref124692" refString="Yablokov-Khnzoryan, S. M. (1978) Two new species of ladybeetles from Eastern Asia (Coleoptera, Coccinellidae). Doklady Akademii Nauk Armyanskoi SSR, 67 (3), 180 - 183." type="journal article" year="1978">Yablokov-Khnzoryan, 1978: 180</bibRefCitation>
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@ -184,7 +184,7 @@ Poorani
2008: 584.
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<taxonomicName id="4C6E8BF71550473FBDDF3107FD11F809" authority="Iablokoff-Khnzorian, 1984: 205" authorityName="Iablokoff-Khnzorian" authorityPageNumber="205" authorityYear="1984" box="[151,732,1914,1939]" class="Insecta" family="Coccinellidae" genus="Lemnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="species" species="martini" subGenus="Synia">
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<emphasis id="B91A2C661550473FBDDF3107FF2EF809" box="[151,227,1915,1938]" italics="true" pageId="264" pageNumber="265">Lemnia</emphasis>
(
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@ -115,7 +115,7 @@ is known only from Cochimbo, Para, central Brazil (Fig. 42).
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The holotype male specimen is in MZSP labeled, &quot;Type [red label with black line border]/ ♂/Brasilien, Para Cochimbo X.1955 Pereira [black line border]/ 31921/ F. Guignot det., 1956
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@ -62,9 +62,9 @@ J.Balfour-Browne, 1953: 56 by original designation);
: 236, 2013: 214.
</paragraph>
<paragraph id="4E6643B6EA2A5CF85D2438002359DD7C" pageId="5" pageNumber="50">
<taxonomicName id="12E3FF159DB9407ED34C9E4305DC4C08" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">Brinckius</taxonomicName>
<taxonomicName id="12E3FF159DB9407ED34C9E4305DC4C08" genus="Brinckius" lsidName="Brinckius" pageId="5" pageNumber="50" rank="genus">Brinckius</taxonomicName>
Guignot, 1957: 38 (type species:
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<taxonomicName id="834BE715B21D96BA05A407C3CA82CDBB" genus="Brinckius" lsidName="Brinckius biguttatus" pageId="5" pageNumber="50" rank="species" species="biguttatus">Brinckius biguttatus</taxonomicName>
Guignot, 1957: 39 by original designation);
<bibRefCitation id="711DF7F86D3254283678CF7033F1B683" author="Bistroem, O" journalOrPublisher="Acta Zoologica Fennica" pageId="72" pageNumber="117" pagination="1 - 41" title="Generic review of the Bidessini (Coleoptera, Dytiscidae)." volume="184" year="1988">
<normalizedToken id="66BF7753F498776EBFB63DC6D6E48118" originalValue="Biström">Bistroem</normalizedToken>
@ -181,7 +181,7 @@ J. Balfour-Browne, was described along with the genus description (
). Subsequent to this,
<bibRefCitation id="D333C7C2B58247B5DAE371BF54A2959F" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="72" pageNumber="117" pagination="33 - 45" title="Contribution a la connaissance des dytiscides sudamericaines [Coleopt.]." volume="24" year="1957">Guignot (1957)</bibRefCitation>
erected the new genus,
<taxonomicName id="CB745B01D23F585C8E120490D2278BA0" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">Brinckius</taxonomicName>
<taxonomicName id="CB745B01D23F585C8E120490D2278BA0" genus="Brinckius" lsidName="Brinckius" pageId="5" pageNumber="50" rank="genus">Brinckius</taxonomicName>
Guignot, with four new species.
<bibRefCitation id="E736B408071EA640C797C228592A5AB2" author="Spangler, PJ" journalOrPublisher="Monographs of the Academy of Natural Science of Philadelphia" pageId="72" pageNumber="117" pagination="377 - 443" title="The Catherwood Foundation Peruvian-Amazon Expedition. XII. Aquatic Coleoptera (Dytiscidae, Noteridae, Gyrinidae, Hydrophilidae, Dascillidae, Helodidae, Psephenidae, Elmidae)." volume="14" year="1966">Spangler (1966)</bibRefCitation>
added two new species from Peru to
@ -191,13 +191,13 @@ added two new species from Peru to
,
<bibRefCitation id="0CB73895DE8C1C070705CB82C38AB446" author="Young, FN" journalOrPublisher="The Coleopterists Bulletin" pageId="72" pageNumber="117" pagination="75 - 84" title="A key to the genera of American bidessine water beetles, with descriptions of three new genera (Coleoptera: Dytiscidae, Hydroporinae)." volume="21" year="1967">Young (1967)</bibRefCitation>
was uncertain whether to synonymize
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<taxonomicName id="74D6140BE22288EB9C096933045BC941" genus="Brinckius" lsidName="Brinckius" pageId="5" pageNumber="50" rank="genus">Brinckius</taxonomicName>
with
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, though he keyed them out together.
<bibRefCitation id="508C7840D0FA60339FDB9E4FE66C608F" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="72" pageNumber="117" title="Dytiscidae. World Catalogue of Insects." year="2001">Nilsson (2001)</bibRefCitation>
regarded the synonymy of
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<taxonomicName id="41AD24AE10E0F48E33CE3795855CC5F8" genus="Brinckius" lsidName="Brinckius" pageId="5" pageNumber="50" rank="genus">Brinckius</taxonomicName>
with
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to date to
@ -214,7 +214,7 @@ whether
[sic] of Guignot should be accorded recognition.&quot; Even so, he soon (
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) did synonymize
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<taxonomicName id="9E83524F5B1BBAA920BCE3B27EA2F8E4" genus="Brinckius" lsidName="Brinckius" pageId="5" pageNumber="50" rank="genus">Brinckius</taxonomicName>
with
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and provided a list of the included species. He then (

54
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@ -45,7 +45,7 @@ Figs 6, 9, 49
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@ -85,7 +85,7 @@ Spangler, 1985: 82;
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<taxonomicName id="6E99F5779A3845483937F5CFC2A51E1B" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius biguttatus" order="Coleoptera" pageId="15" pageNumber="60" phylum="Arthropoda" rank="species" species="biguttatus">Brinckius biguttatus</taxonomicName>
<taxonomicName id="6E99F5779A3845483937F5CFC2A51E1B" genus="Brinckius" lsidName="Brinckius biguttatus" pageId="15" pageNumber="60" rank="species" species="biguttatus">Brinckius biguttatus</taxonomicName>
Guignot: Brazil,
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State, Cachimbo.

58
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@ -45,7 +45,7 @@ Figs 26, 48
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@ -83,7 +83,7 @@ Spangler, 1985: 85;
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@ -161,7 +161,7 @@ the type of
The
<taxonomicName id="286B999D40FE894AF6C12DD7CA108516" class="Insecta" family="Dytiscidae" genus="Hydrodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrodessus octospilus" order="Coleoptera" pageId="52" pageNumber="97" phylum="Arthropoda" rank="species" species="octospilus">Hydrodessus octospilus</taxonomicName>
male holotype in MZSP was examined, labeled, &quot;Type [red label with black line border]/ Brasilien, Para Cochimbo X.1955 Pereira [black line border]/ F. Guignot det., 1956
<taxonomicName id="DCD65B82E29EAB9C68FF4EEB0CB33A5F" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius octospilus" order="Coleoptera" pageId="52" pageNumber="97" phylum="Arthropoda" rank="species" species="octospilus">Brinckius octospilus</taxonomicName>
<taxonomicName id="DCD65B82E29EAB9C68FF4EEB0CB33A5F" genus="Brinckius" lsidName="Brinckius octospilus" pageId="52" pageNumber="97" rank="species" species="octospilus">Brinckius octospilus</taxonomicName>
n.sp. Type ♂ [handwritten]/ 31904.&quot;
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56
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<paragraph id="5F4E6A9008633D0BE8D036DD5B300CBD" pageId="55" pageNumber="100">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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@ -45,7 +45,7 @@ Figs 4, 45
</subSubSection>
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<paragraph id="7BC99E836347DEEB8582F1D3B428BE94" pageId="55" pageNumber="100">
<taxonomicName id="85F57F94A09CA5AB540FDA0EFB84BCED" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius pereirai" order="Coleoptera" pageId="55" pageNumber="100" phylum="Arthropoda" rank="species" species="pereirai">Brinckius pereirai</taxonomicName>
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Guignot, 1957: 41.
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@ -109,7 +109,7 @@ so the usually definitive character system of male genitalia is not available fo
<paragraph id="DFE0640B0D3C6D620A4A2570E6E09223" pageId="55" pageNumber="100">Specimens.</paragraph>
<paragraph id="022D2A222F7089A52A5201A7C2587898" pageId="55" pageNumber="100">
Only the female holotype in MZSP was examined, labeled, &quot;Type [red label with black border]/ Brasilien, Para Cachimbo X.1955 Pereira/ F. Guignot det., 1956
<taxonomicName id="FEB1D75DA153A3CD009B5B5FEB65C988" class="Insecta" family="Dytiscidae" genus="Brinckius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brinckius" order="Coleoptera" pageId="55" pageNumber="100" phylum="Arthropoda" rank="genus">Brinckius</taxonomicName>
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Pereirai Type ♀ [handwritten]/ 31901.&quot;
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