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<document id="2187F66E1246E3C4AE6D0127F10BD156" ID-CLB-Dataset="3608" ID-DOI="10.11646/zootaxa.5067.2.7" ID-GBIF-Dataset="59e0113f-90f0-4cc7-a332-7a8c7269435b" ID-ISSN="1175-5326" ID-Zenodo-Dep="5677973" ID-ZooBank="FE8F2CB8-72EC-4A12-95AF-260F3C2941AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636527369303" checkinUser="plazi" docAuthor="Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong &amp; Wang, Yongjie" docDate="2021" docId="F70987CA151DFFB5A5B9321FFC88FE73" docLanguage="en" docName="zootaxa.5067.2.7.pdf" docOrigin="Zootaxa 5067 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fuscopolystoechotes Xu, Zheng, Shih &amp; Wang 2021, gen. nov." docType="treatment" docUuid="504F861B-6DE0-4C99-B962-0E12E3C83164" docUuidSource="ZooBank" docVersion="8" lastPageNumber="260" masterDocId="0B30FFB2151CFFB7A52E3725FF88FFDE" masterDocTitle="New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)" masterLastPageNumber="266" masterPageNumber="259" pageNumber="260" updateTime="1699468132269" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="49B27D14298B725DFBADF7DF754ADE77">New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)</mods:title>
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<mods:namePart id="DBA1122ADE2522A39C9C3B4E6F73F6FD">Xu, Yifan</mods:namePart>
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<mods:namePart id="F09716568D3563A474D63CD0EB74D101">Zheng, Bingyu</mods:namePart>
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<mods:namePart id="C3D16E3E00CC8FCEE06DD351916D5294">Fang, Hui</mods:namePart>
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<mods:namePart id="58C97206E5A3CD0F32E4BA899189AA71">Shih, Chungkun</mods:namePart>
<mods:affiliation id="4B832E523180EE47BCC3DAAA4CA614D5">Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20013 - 7012, USA Corresponding author. wangyongjie @ cnu. edu. cn</mods:affiliation>
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<mods:namePart id="AFB46F4750B8CB921045D97DB505F33A">Ren, Dong</mods:namePart>
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<mods:namePart id="7542CA9979DDB2BDD737413F567BC556">Wang, Yongjie</mods:namePart>
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<treatment id="F70987CA151DFFB5A5B9321FFC88FE73" ID-DOI="http://doi.org/10.5281/zenodo.5698937" ID-GBIF-Taxon="190302570" ID-Zenodo-Dep="5698937" LSID="urn:lsid:zoobank.org:act:504F861B-6DE0-4C99-B962-0E12E3C83164" httpUri="http://treatment.plazi.org/id/F70987CA151DFFB5A5B9321FFC88FE73" lastPageId="2" lastPageNumber="260" pageId="1" pageNumber="260">
<subSubSection id="37BA6557151DFFB6A5B9321FFC30FAA7" pageId="1" pageNumber="260" type="nomenclature">
<paragraph id="7F1F36DC151DFFB6A5B9321FFC2FFA8B" blockId="1.[151,952,1338,1401]" box="[151,935,1338,1365]" pageId="1" pageNumber="260">
<heading id="245781B0151DFFB6A5B9321FFC2FFA8B" bold="true" box="[151,935,1338,1365]" fontSize="11" level="1" pageId="1" pageNumber="260" reason="1">
<emphasis id="4DD4EACE151DFFB6A5B9321FFC2FFA8B" bold="true" box="[151,935,1338,1365]" pageId="1" pageNumber="260">
Genus
<taxonomicName id="B8A04D5F151DFFB6A5C0321FFCA4FA8B" authority="Xu, Zheng, Shih &amp; Wang" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[238,812,1338,1365]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4DD4EACE151DFFB6A5C0321FFE6FFA8B" bold="true" box="[238,487,1338,1365]" italics="true" pageId="1" pageNumber="260">Fuscopolystoechotes</emphasis>
Xu, Zheng, Shih &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="56E757B5151DFFB6A617321EFC2FFA8B" box="[825,935,1339,1365]" pageId="1" pageNumber="260" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F1F36DC151DFFB6A5B9327BFC30FAA7" blockId="1.[151,952,1338,1401]" box="[151,952,1374,1401]" pageId="1" pageNumber="260">
<uri id="0B313ADE151DFFB6A5B9327BFC30FAA7" box="[151,952,1374,1401]" pageId="1" pageNumber="260">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151DFFB6A5B93282FB15FA1F" box="[151,1181,1446,1473]" pageId="1" pageNumber="260" type="type_taxon">
<paragraph id="7F1F36DC151DFFB6A5B93282FB15FA1F" blockId="1.[151,1437,1446,2013]" box="[151,1181,1446,1473]" pageId="1" pageNumber="260">
<emphasis id="4DD4EACE151DFFB6A5B93282FE2BFA1E" bold="true" box="[151,419,1446,1473]" pageId="1" pageNumber="260">
<typeStatus id="A01B887E151DFFB6A5B93282FF5AFA1F" box="[151,210,1447,1473]" pageId="1" pageNumber="260">Type</typeStatus>
and only species:
</emphasis>
<taxonomicName id="B8A04D5F151DFFB6A4843282FBB0FA1F" authority="Xu, Zheng, Shih &amp; Wang" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[426,1080,1447,1473]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="4DD4EACE151DFFB6A4843282FC99FA1F" box="[426,785,1447,1473]" italics="true" pageId="1" pageNumber="260">Fuscopolystoechotes reticulatus</emphasis>
Xu, Zheng, Shih &amp; Wang
</taxonomicName>
,
<emphasis id="4DD4EACE151DFFB6A16D3282FB15FA1F" bold="true" box="[1091,1181,1447,1473]" pageId="1" pageNumber="260">
<taxonomicNameLabel id="56E757B5151DFFB6A16D3282FB15FA1F" box="[1091,1181,1447,1473]" pageId="1" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151DFFB6A5E932EFFD9FF8F7" pageId="1" pageNumber="260" type="diagnosis">
<paragraph id="7F1F36DC151DFFB6A5E932EFFD9FF8F7" blockId="1.[151,1437,1446,2013]" pageId="1" pageNumber="260">
<emphasis id="4DD4EACE151DFFB6A5E932EFFECAFA3A" bold="true" box="[199,322,1482,1508]" pageId="1" pageNumber="260">Diagnosis.</emphasis>
Forewing elongated, slightly broad with length-width ratio about 3; membrane fuscous, with the distinct transparent patches along the anterior and posterior margins; costal space narrow at base and then expanded to the max in proximal 1/5 of forewing, and subsequently narrowed gradually, with a distinctly shrinking region in distal half; humeral veinlet recurrent; presence of ScA; costal crossveins mostly biforked distally; absence of scp-ra crossvein; RP with more than 10 branches, RP1 deeply branched at the proximal half; crossveins in radial sector irregularly arranged except for the outer gradate series; M forked near the separated point of RP1 from RP; MA and MP bearing the similarly distal pectinate branches; Cu forked before the separated point of RP; CuA distinctly longer than CuP, forming the pectinate branches in distal; CuP with 34 pectinate branches, and each forming the secondary sub-branches; A1 dichotomously branched distally, A2 forked close to the base, A3 single branch, with several distal minor sub-branches.
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151DFFB6A5E93017FE98F84B" pageId="1" pageNumber="260" type="etymology">
<paragraph id="7F1F36DC151DFFB6A5E93017FE98F84B" blockId="1.[151,1437,1446,2013]" pageId="1" pageNumber="260">
<emphasis id="4DD4EACE151DFFB6A5E93017FEC1F893" bold="true" box="[199,329,1842,1869]" pageId="1" pageNumber="260">
<emphasis id="4DD4EACE151DFFB6A5E93017FECAF893" bold="true" box="[199,322,1842,1869]" italics="true" pageId="1" pageNumber="260">Etymology</emphasis>
.
</emphasis>
The new generic name is a combination of the Latin prefix
<emphasis id="4DD4EACE151DFFB6A12B3016FBB3F893" box="[1029,1083,1843,1869]" italics="true" pageId="1" pageNumber="260">fusc-</emphasis>
and
<emphasis id="4DD4EACE151DFFB6A1583016FA93F893" box="[1142,1307,1843,1869]" italics="true" pageId="1" pageNumber="260">polystoechotes</emphasis>
(the representative genus of the
<taxonomicName id="B8A04D5F151DFFB6A4B93072FDB7F8AF" authorityName="Burmeister" authorityYear="1839" box="[407,575,1879,1905]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151DFFB6A4B93072FDB7F8AF" box="[407,575,1879,1905]" italics="true" pageId="1" pageNumber="260">Polystoechotes</emphasis>
</taxonomicName>
genus-group), referring to dark brown markings of forewing. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151DFFB5A5E930BAFC88FE73" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260" type="discussion">
<paragraph id="7F1F36DC151DFFB5A5E930BAFC88FE73" blockId="1.[151,1437,1446,2013]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260">
<emphasis id="4DD4EACE151DFFB6A5E930BAFEB3F867" bold="true" box="[199,315,1951,1977]" pageId="1" pageNumber="260">Remarks.</emphasis>
The new genus is clearly assigned to the
<taxonomicName id="B8A04D5F151DFFB6A63130BAFC4FF867" authorityName="Burmeister" authorityYear="1839" box="[799,967,1951,1977]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151DFFB6A63130BAFC4FF867" box="[799,967,1951,1977]" italics="true" pageId="1" pageNumber="260">Polystoechotes</emphasis>
</taxonomicName>
genus-group of
<taxonomicName id="B8A04D5F151DFFB6A1AB30BAFB67F867" authorityName="Newman, 1853 sensu Winterton &amp; Makarkin" authorityYear="2010" box="[1157,1263,1951,1977]" class="Insecta" family="Ithonidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="family">Ithonidae</taxonomicName>
for the combination of following characters: 1) forewing distinctly elongated (
<emphasis id="4DD4EACE151DFFB6A6A630E6FC14F803" box="[904,924,1987,2013]" italics="true" pageId="1" pageNumber="260">cf</emphasis>
. relatively broad in
<taxonomicName id="B8A04D5F151DFFB6A1A230E6FB5CF803" box="[1164,1236,1987,2013]" class="Insecta" family="Ithonidae" genus="Ithone" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151DFFB6A1A230E6FB5CF803" box="[1164,1236,1987,2013]" italics="true" pageId="1" pageNumber="260">Ithone</emphasis>
</taxonomicName>
genus-group); 2) absence of interlinked veinlets in between costal crossveins (
<emphasis id="4DD4EACE151EFFB5A67A37B2FCE0FF6F" box="[852,872,151,177]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
. commonly present in
<taxonomicName id="B8A04D5F151EFFB5A15D37B2FB5DFF6E" authorityName="McLachlan" authorityYear="1866" box="[1139,1237,151,176]" class="Insecta" family="Ithonidae" genus="Rapisma" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A15D37B2FB5DFF6E" box="[1139,1237,151,176]" italics="true" pageId="2" pageNumber="261">Rapisma</emphasis>
</taxonomicName>
genus-group and
<taxonomicName id="B8A04D5F151EFFB5A5B9379EFE9FFF0B" authorityName="Makarkin &amp; Menon" authorityYear="2007" box="[151,279,187,213]" class="Insecta" family="Ithonidae" genus="Principiala" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A5B9379EFE9FFF0B" box="[151,279,187,213]" italics="true" pageId="2" pageNumber="261">Principiala</emphasis>
</taxonomicName>
genus-group); 3) Sc and RA fused distally (
<emphasis id="4DD4EACE151EFFB5A62C379EFC9EFF0B" box="[770,790,187,213]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
. generally separated in
<taxonomicName id="B8A04D5F151EFFB5A10E379EFBE0FF0B" box="[1056,1128,187,213]" class="Insecta" family="Ithonidae" genus="Ithone" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A10E379EFBE0FF0B" box="[1056,1128,187,213]" italics="true" pageId="2" pageNumber="261">Ithone</emphasis>
</taxonomicName>
genus-groups and
<taxonomicName id="B8A04D5F151EFFB5A014379EFA14FF0A" authorityName="McLachlan" authorityYear="1866" box="[1338,1436,187,212]" class="Insecta" family="Ithonidae" genus="Rapisma" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A014379EFA14FF0A" box="[1338,1436,187,212]" italics="true" pageId="2" pageNumber="261">Rapisma</emphasis>
</taxonomicName>
genus-groups); 4) crossveins in radial sector regularly arranged or relatively sparse (
<emphasis id="4DD4EACE151EFFB5A17137FAFBFBFF27" box="[1119,1139,223,249]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
. abundant and commonly irregularly arranged in other groups). The new genus is characterized by its special markings of forewing that is distinctly different from other genera of
<taxonomicName id="B8A04D5F151EFFB5A76E3602FD60FE9F" authorityName="Burmeister" authorityYear="1839" box="[576,744,295,321]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A76E3602FD60FE9F" box="[576,744,295,321]" italics="true" pageId="2" pageNumber="261">Polystoechotes</emphasis>
</taxonomicName>
genus-group except for
<emphasis id="4DD4EACE151EFFB5A6D03602FB76FE9F" box="[1022,1278,295,321]" italics="true" pageId="2" pageNumber="261">
<taxonomicName id="B8A04D5F151EFFB5A6D03602FB73FE9F" authorityName="Fang et al" authorityYear="2020" box="[1022,1275,295,321]" class="Insecta" family="Ithonidae" genus="Lichenipolystoechotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">Lichenipolystoechotes</taxonomicName>
.
</emphasis>
However, the forewing markings in the new genus are distinctly broader and more regular comparing to
<taxonomicName id="B8A04D5F151EFFB5A1B2366EFA1DFEBB" authorityName="Fang et al" authorityYear="2020" box="[1180,1429,331,357]" class="Insecta" family="Ithonidae" genus="Lichenipolystoechotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="4DD4EACE151EFFB5A1B2366EFA1DFEBB" box="[1180,1429,331,357]" italics="true" pageId="2" pageNumber="261">Lichenipolystoechotes</emphasis>
</taxonomicName>
. In addition, their venation are also quite different, e.g., the branched costal crossveins, and the more complicated configuration of CuP, to justify setting up a new genus.
</paragraph>
</subSubSection>
</treatment>
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