188 lines
18 KiB
XML
188 lines
18 KiB
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<document id="2187F66E1246E3C4AE6D0127F10BD156" ID-CLB-Dataset="3608" ID-DOI="10.11646/zootaxa.5067.2.7" ID-GBIF-Dataset="59e0113f-90f0-4cc7-a332-7a8c7269435b" ID-ISSN="1175-5326" ID-Zenodo-Dep="5677973" ID-ZooBank="FE8F2CB8-72EC-4A12-95AF-260F3C2941AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636527369303" checkinUser="plazi" docAuthor="Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong & Wang, Yongjie" docDate="2021" docId="F70987CA151DFFB5A5B9321FFC88FE73" docLanguage="en" docName="zootaxa.5067.2.7.pdf" docOrigin="Zootaxa 5067 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fuscopolystoechotes Xu, Zheng, Shih & Wang 2021, gen. nov." docType="treatment" docUuid="504F861B-6DE0-4C99-B962-0E12E3C83164" docUuidSource="ZooBank" docVersion="8" lastPageNumber="260" masterDocId="0B30FFB2151CFFB7A52E3725FF88FFDE" masterDocTitle="New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)" masterLastPageNumber="266" masterPageNumber="259" pageNumber="260" updateTime="1699468132269" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="D8B8096510E113223414D1FBDE8E464B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8E1F5DA16DA6B3AEF5A9492AC90B0B03">
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<mods:title id="49B27D14298B725DFBADF7DF754ADE77">New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="D9F0A7A7111C28698F86F4A4434D57D2" type="personal">
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<mods:roleTerm id="D3A07589DC45CBD10FC313BFFC92712F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DBA1122ADE2522A39C9C3B4E6F73F6FD">Xu, Yifan</mods:namePart>
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</mods:name>
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<mods:name id="73B017A832EE37C624DBF3E0780E3004" type="personal">
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<mods:roleTerm id="2C662F7C744EC977BDBB23481E6BB355">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F09716568D3563A474D63CD0EB74D101">Zheng, Bingyu</mods:namePart>
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</mods:name>
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<mods:name id="79F69BE0C0E46357F884E71B772302C0" type="personal">
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<mods:role id="A62E64812E2421F343DE93F81450FC70">
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<mods:roleTerm id="11514930399AE91776277E7FAB609639">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C3D16E3E00CC8FCEE06DD351916D5294">Fang, Hui</mods:namePart>
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</mods:name>
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<mods:name id="9E98823A9132DA8513E99FC3BB5B77ED" type="personal">
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<mods:role id="DD2C69DD102D5F33875F006634D0EBDF">
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<mods:roleTerm id="44329F0008EDB2664BC6DCA70F43F564">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58C97206E5A3CD0F32E4BA899189AA71">Shih, Chungkun</mods:namePart>
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<mods:affiliation id="4B832E523180EE47BCC3DAAA4CA614D5">Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20013 - 7012, USA Corresponding author. wangyongjie @ cnu. edu. cn</mods:affiliation>
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</mods:name>
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<mods:name id="17FBDB9AA163B76A1B1BDA12DEBC0964" type="personal">
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<mods:roleTerm id="31F109EB54AACBFBD8179F34F4B197CB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AFB46F4750B8CB921045D97DB505F33A">Ren, Dong</mods:namePart>
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</mods:name>
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<mods:name id="D2E6FE4896BBD440463D7D62ECEB746A" type="personal">
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<mods:roleTerm id="907B1FDA8D9C63FA8E17D04E2E3A5F19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7542CA9979DDB2BDD737413F567BC556">Wang, Yongjie</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C434F8D8E39A6F26F0780777EDDD34E3">text</mods:typeOfResource>
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<mods:title id="A0593C049FDA2BF01320BDC2A203A6E9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D972F5B42DEFE8FEF16A2EE886EE6391">
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<mods:date id="911AFDAD379CA2C40534F89733491A46">2021</mods:date>
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<mods:detail id="D9BC72A6D48E32E0CA31295837219F63" type="pubDate">
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<mods:number id="1742EA58FA57B88AC460EA24AF7C3B44">2021-11-10</mods:number>
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<mods:detail id="FDABD8DBB33DFEC2B900017F58833F51" type="volume">
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<mods:number id="2997CB9984E2D31D16FD84C5D823DE2B">5067</mods:number>
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</mods:detail>
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<mods:detail id="BC5E3044611C6E2CCE7A5B87AE974CAB" type="issue">
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<mods:number id="53615154B387818E6957FE43B6A09853">2</mods:number>
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<mods:start id="292E7C7743E1506DCEBDBF00F547CDB4">259</mods:start>
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<mods:classification id="BB3A607EE6FD830AB685C140E8F47EA5">journal article</mods:classification>
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<mods:identifier id="375ED974D037C587B6914889EE3BB2B5" type="CLB-Dataset">3608</mods:identifier>
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<mods:identifier id="B68198D45CBBD6A115E28D82742C4AD5" type="DOI">10.11646/zootaxa.5067.2.7</mods:identifier>
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<mods:identifier id="256F0F01717BB48407AE40EF2CBB5E75" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0CD3EBAA1482AD038F59BE08EE263C32" type="Zenodo-Dep">5677973</mods:identifier>
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<mods:identifier id="0136EEAEF7097831230D4ADA6CDD3E95" type="ZooBank">FE8F2CB8-72EC-4A12-95AF-260F3C2941AB</mods:identifier>
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</mods:mods>
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<treatment id="F70987CA151DFFB5A5B9321FFC88FE73" ID-DOI="http://doi.org/10.5281/zenodo.5698937" ID-GBIF-Taxon="190302570" ID-Zenodo-Dep="5698937" LSID="urn:lsid:zoobank.org:act:504F861B-6DE0-4C99-B962-0E12E3C83164" httpUri="http://treatment.plazi.org/id/F70987CA151DFFB5A5B9321FFC88FE73" lastPageId="2" lastPageNumber="260" pageId="1" pageNumber="260">
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<subSubSection id="37BA6557151DFFB6A5B9321FFC30FAA7" pageId="1" pageNumber="260" type="nomenclature">
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<paragraph id="7F1F36DC151DFFB6A5B9321FFC2FFA8B" blockId="1.[151,952,1338,1401]" box="[151,935,1338,1365]" pageId="1" pageNumber="260">
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<heading id="245781B0151DFFB6A5B9321FFC2FFA8B" bold="true" box="[151,935,1338,1365]" fontSize="11" level="1" pageId="1" pageNumber="260" reason="1">
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<emphasis id="4DD4EACE151DFFB6A5B9321FFC2FFA8B" bold="true" box="[151,935,1338,1365]" pageId="1" pageNumber="260">
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Genus
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<taxonomicName id="B8A04D5F151DFFB6A5C0321FFCA4FA8B" authority="Xu, Zheng, Shih & Wang" authorityName="Xu, Zheng, Shih & Wang" authorityYear="2021" box="[238,812,1338,1365]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="4DD4EACE151DFFB6A5C0321FFE6FFA8B" bold="true" box="[238,487,1338,1365]" italics="true" pageId="1" pageNumber="260">Fuscopolystoechotes</emphasis>
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Xu, Zheng, Shih & Wang
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</taxonomicName>
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,
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<taxonomicNameLabel id="56E757B5151DFFB6A617321EFC2FFA8B" box="[825,935,1339,1365]" pageId="1" pageNumber="260" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7F1F36DC151DFFB6A5B9327BFC30FAA7" blockId="1.[151,952,1338,1401]" box="[151,952,1374,1401]" pageId="1" pageNumber="260">
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<uri id="0B313ADE151DFFB6A5B9327BFC30FAA7" box="[151,952,1374,1401]" pageId="1" pageNumber="260">
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urn:lsid:zoobank.org:act:
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<uuid id="0B060C09151DFFB6A485327BFC30FAA7" box="[427,952,1374,1401]" pageId="1" pageNumber="260">504F861B-6DE0-4C99-B962-0E12E3C83164</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="37BA6557151DFFB6A5B93282FB15FA1F" box="[151,1181,1446,1473]" pageId="1" pageNumber="260" type="type_taxon">
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<paragraph id="7F1F36DC151DFFB6A5B93282FB15FA1F" blockId="1.[151,1437,1446,2013]" box="[151,1181,1446,1473]" pageId="1" pageNumber="260">
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<emphasis id="4DD4EACE151DFFB6A5B93282FE2BFA1E" bold="true" box="[151,419,1446,1473]" pageId="1" pageNumber="260">
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<typeStatus id="A01B887E151DFFB6A5B93282FF5AFA1F" box="[151,210,1447,1473]" pageId="1" pageNumber="260">Type</typeStatus>
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and only species:
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</emphasis>
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<taxonomicName id="B8A04D5F151DFFB6A4843282FBB0FA1F" authority="Xu, Zheng, Shih & Wang" authorityName="Xu, Zheng, Shih & Wang" authorityYear="2021" box="[426,1080,1447,1473]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
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<emphasis id="4DD4EACE151DFFB6A4843282FC99FA1F" box="[426,785,1447,1473]" italics="true" pageId="1" pageNumber="260">Fuscopolystoechotes reticulatus</emphasis>
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Xu, Zheng, Shih & Wang
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</taxonomicName>
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,
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<emphasis id="4DD4EACE151DFFB6A16D3282FB15FA1F" bold="true" box="[1091,1181,1447,1473]" pageId="1" pageNumber="260">
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<taxonomicNameLabel id="56E757B5151DFFB6A16D3282FB15FA1F" box="[1091,1181,1447,1473]" pageId="1" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="37BA6557151DFFB6A5E932EFFD9FF8F7" pageId="1" pageNumber="260" type="diagnosis">
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<paragraph id="7F1F36DC151DFFB6A5E932EFFD9FF8F7" blockId="1.[151,1437,1446,2013]" pageId="1" pageNumber="260">
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<emphasis id="4DD4EACE151DFFB6A5E932EFFECAFA3A" bold="true" box="[199,322,1482,1508]" pageId="1" pageNumber="260">Diagnosis.</emphasis>
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Forewing elongated, slightly broad with length-width ratio about 3; membrane fuscous, with the distinct transparent patches along the anterior and posterior margins; costal space narrow at base and then expanded to the max in proximal 1/5 of forewing, and subsequently narrowed gradually, with a distinctly shrinking region in distal half; humeral veinlet recurrent; presence of ScA; costal crossveins mostly biforked distally; absence of scp-ra crossvein; RP with more than 10 branches, RP1 deeply branched at the proximal half; crossveins in radial sector irregularly arranged except for the outer gradate series; M forked near the separated point of RP1 from RP; MA and MP bearing the similarly distal pectinate branches; Cu forked before the separated point of RP; CuA distinctly longer than CuP, forming the pectinate branches in distal; CuP with 3–4 pectinate branches, and each forming the secondary sub-branches; A1 dichotomously branched distally, A2 forked close to the base, A3 single branch, with several distal minor sub-branches.
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</paragraph>
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</subSubSection>
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<subSubSection id="37BA6557151DFFB6A5E93017FE98F84B" pageId="1" pageNumber="260" type="etymology">
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<paragraph id="7F1F36DC151DFFB6A5E93017FE98F84B" blockId="1.[151,1437,1446,2013]" pageId="1" pageNumber="260">
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<emphasis id="4DD4EACE151DFFB6A5E93017FEC1F893" bold="true" box="[199,329,1842,1869]" pageId="1" pageNumber="260">
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<emphasis id="4DD4EACE151DFFB6A5E93017FECAF893" bold="true" box="[199,322,1842,1869]" italics="true" pageId="1" pageNumber="260">Etymology</emphasis>
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.
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</emphasis>
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The new generic name is a combination of the Latin prefix
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<emphasis id="4DD4EACE151DFFB6A12B3016FBB3F893" box="[1029,1083,1843,1869]" italics="true" pageId="1" pageNumber="260">fusc-</emphasis>
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and
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<emphasis id="4DD4EACE151DFFB6A1583016FA93F893" box="[1142,1307,1843,1869]" italics="true" pageId="1" pageNumber="260">polystoechotes</emphasis>
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(the representative genus of the
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<taxonomicName id="B8A04D5F151DFFB6A4B93072FDB7F8AF" authorityName="Burmeister" authorityYear="1839" box="[407,575,1879,1905]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
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<emphasis id="4DD4EACE151DFFB6A4B93072FDB7F8AF" box="[407,575,1879,1905]" italics="true" pageId="1" pageNumber="260">Polystoechotes</emphasis>
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</taxonomicName>
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genus-group), referring to dark brown markings of forewing. The gender is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="37BA6557151DFFB5A5E930BAFC88FE73" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260" type="discussion">
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<paragraph id="7F1F36DC151DFFB5A5E930BAFC88FE73" blockId="1.[151,1437,1446,2013]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260">
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<emphasis id="4DD4EACE151DFFB6A5E930BAFEB3F867" bold="true" box="[199,315,1951,1977]" pageId="1" pageNumber="260">Remarks.</emphasis>
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The new genus is clearly assigned to the
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<taxonomicName id="B8A04D5F151DFFB6A63130BAFC4FF867" authorityName="Burmeister" authorityYear="1839" box="[799,967,1951,1977]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
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<emphasis id="4DD4EACE151DFFB6A63130BAFC4FF867" box="[799,967,1951,1977]" italics="true" pageId="1" pageNumber="260">Polystoechotes</emphasis>
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</taxonomicName>
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genus-group of
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<taxonomicName id="B8A04D5F151DFFB6A1AB30BAFB67F867" authorityName="Newman, 1853 sensu Winterton & Makarkin" authorityYear="2010" box="[1157,1263,1951,1977]" class="Insecta" family="Ithonidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="family">Ithonidae</taxonomicName>
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for the combination of following characters: 1) forewing distinctly elongated (
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<emphasis id="4DD4EACE151DFFB6A6A630E6FC14F803" box="[904,924,1987,2013]" italics="true" pageId="1" pageNumber="260">cf</emphasis>
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. relatively broad in
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<taxonomicName id="B8A04D5F151DFFB6A1A230E6FB5CF803" box="[1164,1236,1987,2013]" class="Insecta" family="Ithonidae" genus="Ithone" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="genus">
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<emphasis id="4DD4EACE151DFFB6A1A230E6FB5CF803" box="[1164,1236,1987,2013]" italics="true" pageId="1" pageNumber="260">Ithone</emphasis>
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</taxonomicName>
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genus-group); 2) absence of interlinked veinlets in between costal crossveins (
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<emphasis id="4DD4EACE151EFFB5A67A37B2FCE0FF6F" box="[852,872,151,177]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
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. commonly present in
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<taxonomicName id="B8A04D5F151EFFB5A15D37B2FB5DFF6E" authorityName="McLachlan" authorityYear="1866" box="[1139,1237,151,176]" class="Insecta" family="Ithonidae" genus="Rapisma" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A15D37B2FB5DFF6E" box="[1139,1237,151,176]" italics="true" pageId="2" pageNumber="261">Rapisma</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
genus-group and
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A5B9379EFE9FFF0B" authorityName="Makarkin & Menon" authorityYear="2007" box="[151,279,187,213]" class="Insecta" family="Ithonidae" genus="Principiala" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A5B9379EFE9FFF0B" box="[151,279,187,213]" italics="true" pageId="2" pageNumber="261">Principiala</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
genus-group); 3) Sc and RA fused distally (
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A62C379EFC9EFF0B" box="[770,790,187,213]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
|
|||
|
|
. generally separated in
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A10E379EFBE0FF0B" box="[1056,1128,187,213]" class="Insecta" family="Ithonidae" genus="Ithone" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A10E379EFBE0FF0B" box="[1056,1128,187,213]" italics="true" pageId="2" pageNumber="261">Ithone</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
genus-groups and
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A014379EFA14FF0A" authorityName="McLachlan" authorityYear="1866" box="[1338,1436,187,212]" class="Insecta" family="Ithonidae" genus="Rapisma" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A014379EFA14FF0A" box="[1338,1436,187,212]" italics="true" pageId="2" pageNumber="261">Rapisma</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
genus-groups); 4) crossveins in radial sector regularly arranged or relatively sparse (
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A17137FAFBFBFF27" box="[1119,1139,223,249]" italics="true" pageId="2" pageNumber="261">cf</emphasis>
|
|||
|
|
. abundant and commonly irregularly arranged in other groups). The new genus is characterized by its special markings of forewing that is distinctly different from other genera of
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A76E3602FD60FE9F" authorityName="Burmeister" authorityYear="1839" box="[576,744,295,321]" class="Insecta" family="Ithonidae" genus="Polystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A76E3602FD60FE9F" box="[576,744,295,321]" italics="true" pageId="2" pageNumber="261">Polystoechotes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
genus-group except for
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A6D03602FB76FE9F" box="[1022,1278,295,321]" italics="true" pageId="2" pageNumber="261">
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A6D03602FB73FE9F" authorityName="Fang et al" authorityYear="2020" box="[1022,1275,295,321]" class="Insecta" family="Ithonidae" genus="Lichenipolystoechotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">Lichenipolystoechotes</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
However, the forewing markings in the new genus are distinctly broader and more regular comparing to
|
|||
|
|
<taxonomicName id="B8A04D5F151EFFB5A1B2366EFA1DFEBB" authorityName="Fang et al" authorityYear="2020" box="[1180,1429,331,357]" class="Insecta" family="Ithonidae" genus="Lichenipolystoechotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="4DD4EACE151EFFB5A1B2366EFA1DFEBB" box="[1180,1429,331,357]" italics="true" pageId="2" pageNumber="261">Lichenipolystoechotes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. In addition, their venation are also quite different, e.g., the branched costal crossveins, and the more complicated configuration of CuP, to justify setting up a new genus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|