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<document id="08C324BFD109B126DCF7B13E156ECB69" ID-CLB-Dataset="56026" ID-DOI="10.11646/megataxa.3.1.1" ID-GBIF-Dataset="1ece557d-068b-465b-809e-0fc1110035b3" ID-ISSN="2703-3090" ID-Zenodo-Dep="6422776" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614076789781" checkinUser="plazi" docAuthor="Nakayama, Naohide" docDate="2020" docId="B711B23FFFF98620D99DC15FFE187B2D" docLanguage="en" docName="megataxa.3.1.1.pdf" docOrigin="Megataxa 3 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Coelorinchus formosanus Okamura 1963" docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="4B28CA47FFD68612D915C228FFE77D35" masterDocTitle="Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph" masterLastPageNumber="383" masterPageNumber="1" pageNumber="48" updateTime="1698930971026" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="67DE367A74ACED746BF787C7FC57E916">Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph</mods:title>
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<mods:namePart id="B13C779EA57256D489DDE317EDF06007">Nakayama, Naohide</mods:namePart>
<mods:nameIdentifier id="53F4F14C1CA7FF8291E5DC0197E594D6" type="ORCID">0000-0001-9556-1537</mods:nameIdentifier>
<mods:affiliation id="52458FDEC643BBF95B9C8B529FD46D62">Department of Marine Biology, School of Marine Science and Technology, Tokai University, 3 - 20 - 1 Orido, Shimizu, Shizuoka 424 - 8610, Japan ® naonakayama @ tsc. u-tokai. ac. jp or ®</mods:affiliation>
<mods:nameIdentifier id="34081EEFF13CBC6FA673B32166E7C233" type="email">gadiformes@gmail.com</mods:nameIdentifier>
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<mods:date id="D67700AEA1FC0D10AFBB39AAB5A3D0DB">2020</mods:date>
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<treatment id="B711B23FFFF98620D99DC15FFE187B2D" ID-GBIF-Taxon="194588064" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D" httpUri="http://treatment.plazi.org/id/B711B23FFFF98620D99DC15FFE187B2D" lastPageId="50" lastPageNumber="50" pageId="47" pageNumber="48">
<subSubSection id="77A250A2FFF9863DD99DC15FFDF77E81" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="3F070329FFF9863DD99DC15FFD6A7EA4" blockId="47.[136,653,887,981]" box="[136,653,887,914]" pageId="47" pageNumber="48">
<heading id="644FB445FFF9863DD99DC15FFD6A7EA4" box="[136,653,887,914]" fontSize="11" level="2" pageId="47" pageNumber="48" reason="2">
<taxonomicName id="F8B878AAFFF9863DD99DC15FFD6A7EA4" ID-CoL="WSSB" authority="Okamura, 1963" authorityName="Okamura" authorityYear="1963" box="[136,653,887,914]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DD99DC15FFD6A7EA4" bold="true" box="[136,653,887,914]" pageId="47" pageNumber="48">
<emphasis id="0DCCDF3BFFF9863DD99DC15FFE267EA4" bold="true" box="[136,449,887,914]" italics="true" pageId="47" pageNumber="48">Coelorinchus formosanus</emphasis>
Okamura, 1963
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3F070329FFF9863DD99DC1B2FDF77E81" blockId="47.[136,653,887,981]" box="[136,528,921,948]" pageId="47" pageNumber="48">[Japanese name: Taiwan-sokodara]</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFF9863DD99DC193FDFE7EE0" box="[136,537,955,981]" pageId="47" pageNumber="48" type="description">
<paragraph id="3F070329FFF9863DD99DC193FDFE7EE0" blockId="47.[136,653,887,981]" box="[136,537,955,981]" pageId="47" pageNumber="48">
(
<figureCitation id="A7831FACFFF9863DD985C193FEFF7EE0" box="[144,280,955,981]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStartId-0="49.[136,235,787,811]" captionStartId-1="49.[136,235,1467,1491]" captionTargetBox-0="[151,1436,159,744]" captionTargetBox-1="[151,1436,907,1430]" captionTargetId-0="figure-223@49.[151,1436,159,744]" captionTargetId-1="figure-266@49.[151,1436,907,1430]" captionTargetPageId-0="49" captionTargetPageId-1="49" captionText-0="FIGURE 32. Fresh specimen of Coelorinchus formosanus. FAKU 204426, 367+ mm TL, Dong-gang fish market, southwestern Taiwan, South China Sea. (A) Lateral view; (B) dorsal and (C) ventral views of the head and trunk. [Photos: N. Nakayama]" captionText-1="FIGURE 33. Holotype of Coelorhynchus formosanus (= Coelorinchus formosanus). FAKU 35856, 72.8 mm HL, 221+ mm TL, Da-xi fish market, northeastern Taiwan, East China Sea. (A) Lateral view; (B) dorsal and (C) ventral views of the head and trunk. Preserved condition. [Photos: N. Nakayama]" figureDoi-0="http://doi.org/10.5281/zenodo.6422852" figureDoi-1="http://doi.org/10.5281/zenodo.6422856" httpUri-0="https://zenodo.org/record/6422852/files/figure.png" httpUri-1="https://zenodo.org/record/6422856/files/figure.png" pageId="47" pageNumber="48">Figs. 32–33</figureCitation>
,
<figureCitation id="A7831FACFFF9863DD831C193FEB07EE0" box="[292,343,955,981]" captionStart="FIGURE 34" captionStartId="50.[136,235,1229,1253]" captionTargetBox="[151,1436,159,1189]" captionTargetId="figure-279@50.[151,1436,159,1189]" captionTargetPageId="50" captionText="FIGURE 34. Scanning electron micrographs showing body scales (from the dorsum below the interdorsal space) of four species of the Coelorinchus argentatus group. (A) C. formosanus, BSKU 99863, ca. 72 mm HL; (B) C. kamoharai, BSKU 93575, 59.6 mm HL; (C) C. longissimus, BSKU 40448, 95.9 mm HL; (D) C. multispinulosus, BSKU 99273, 69.9 mm HL. Views from above. [Photos: N. Nakayama]" figureDoi="http://doi.org/10.5281/zenodo.6422858" httpUri="https://zenodo.org/record/6422858/files/figure.png" pageId="47" pageNumber="48">34A</figureCitation>
; Appendix 3-2A)
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFF9863DD99DC62BFE8A7A9A" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="3F070329FFF9863DD99DC62BFD9679F7" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC62BFDB2792C" ID-CoL="WSN6" authority="Okamura, 1963 b: 37" authorityName="Okamura" authorityPageNumber="37" authorityYear="1963" box="[136,597,1027,1049]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DD99DC62BFE69792D" box="[136,398,1027,1048]" italics="true" pageId="47" pageNumber="48">Coelorhynchus formosanus</emphasis>
<bibRefCitation id="5B297ED8FFF9863DD881C62BFDB2792C" author="Okamura, O." box="[404,597,1027,1049]" pageId="47" pageNumber="48" pagination="37 - 42" refId="ref297236" refString="Okamura, O. (1963 b) A new macrouroid fish found in the adjacent waters of Formosa. Bulletin of the Misaki Marine Biological Institute, Kyoto University, (4), 37 - 42." type="journal article" year="1963">Okamura, 1963b:37</bibRefCitation>
</taxonomicName>
, figs. 1–2 [original description;
<typeStatus id="E003BD8BFFF9863DD82BC60DFE71790E" box="[318,406,1061,1083]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
: FAKU 35836, from “Tashi” (Daxi);
<specimenCount id="29BEC8A0FFF9863DD9F8C66FFE847969" box="[237,355,1095,1116]" pageId="47" pageNumber="48" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
from
<collectingCountry id="47AF43B9FFF9863DD8B8C66EFE1E7969" box="[429,505,1094,1116]" name="Taiwan" pageId="47" pageNumber="48">Taiwan</collectingCountry>
];
<bibRefCitation id="5B297ED8FFF9863DDB05C66FFD387968" author="Okamura, O." box="[528,735,1095,1117]" pageId="47" pageNumber="48" refId="ref297275" refString="Okamura, O. (1970 a) Fauna Japonica, Macrourina (Pisces). Academic Press of Japan, Tokyo, 216 pp., 64 pls." type="book" year="1970">Okamura 1970a:161</bibRefCitation>
, pl. XXXIV, text-figs. 65–66 (description; re-examination of type specimens; new Japanese name: “Taiwan-sokodara”);
<bibRefCitation id="5B297ED8FFF9863DD9A8C685FE3B79F7" author="Tominaga, Y. &amp; Uyeno, T." box="[189,476,1197,1219]" pageId="47" pageNumber="48" pagination="437 - 574" refId="ref301688" refString="Tominaga, Y. &amp; Uyeno, T. (1981) List of Japanese fishes. In: Yasuda, F., Takagi, K., Tominaga, Y., Uyeno, T., Abe, T., Ishiyama, R., Iwai, T., Ochiai, A., Kuronuma, K. &amp; Nakamura, M. (Eds.), Dictionary of Japanese Fish Names and Their Foreign Equivalents. Sanseido Co., Ltd., Tokyo, pp. 437 - 574." type="book chapter" year="1981">Tominaga &amp; Uyeno 1981:488</bibRefCitation>
(listed;
<collectingCountry id="47AF43B9FFF9863DDB3EC685FD8179F7" box="[555,614,1197,1218]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
).
</paragraph>
<paragraph id="3F070329FFF9863DD99DC6E7FEB97833" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC6E7FD2879D0" ID-CoL="WSN6" authority=": Okamura 1970 b" authorityName="Okamura" authorityYear="1970" box="[136,719,1231,1253]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus" subGenus="Quincuncia">
<emphasis id="0DCCDF3BFFF9863DD99DC6E7FDFB79D1" box="[136,540,1231,1252]" italics="true" pageId="47" pageNumber="48">Coelorhynchus (Quincuncia) formosanus</emphasis>
:
<bibRefCitation id="5B297ED8FFF9863DDB39C6E7FD2879D0" author="Okamura, O." box="[556,719,1231,1253]" pageId="47" pageNumber="48" pagination="1 - 179" refId="ref297305" refString="Okamura, O. (1970 b) Studies on the macrouroid fishes of Japan: morphology, ecology and phylogeny. Reports of the Usa Marine Biological Station, 17, 1 - 179, pls. I-V." type="journal article" year="1970">Okamura 1970b</bibRefCitation>
</taxonomicName>
: table 1 (listed;
<collectingCountry id="47AF43B9FFF9863DD80DC6D9FEB47833" box="[280,339,1265,1286]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
).
</paragraph>
<paragraph id="3F070329FFF9863DD99DC73BFEA97858" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC73BFE65781D" ID-CoL="WST9" authorityName="Matsubara" authorityYear="1943" box="[136,386,1299,1320]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="0DCCDF3BFFF9863DD99DC73BFE65781D" box="[136,386,1299,1320]" italics="true" pageId="47" pageNumber="48">Coelorinchus longissimus</emphasis>
</taxonomicName>
(not Matsubara 1943):
<bibRefCitation id="5B297ED8FFF9863DDB7CC73BFCE1781D" author="Yatou, T." box="[617,774,1299,1320]" pageId="47" pageNumber="48" pagination="363 - 370" refId="ref303487" refString="Yatou, T. (1984) Coryphaenoides and Coelorinchus [in part]. In: Okamura, O. &amp; Kitajima, T. (Eds.), Fishes of the Okinawa Trough and the Adjacent Waters I. Japan Fisheries Resource Conservation Association, Tokyo, pp. 218 - 223, 228 - 235, 224 - 245, 363 - 370." type="journal article" year="1984">Yatou 1984:223</bibRefCitation>
, 366, fig. 159 (brief description;
<collectingRegion id="FD7CCDCBFFF9863DDB1CC71DFD87787E" box="[521,608,1333,1355]" country="Japan" name="Okinawa" pageId="47" pageNumber="48">Okinawa</collectingRegion>
Trough; in part, BSKU 26581).
</paragraph>
<paragraph id="3F070329FFF9863DD99DC751FEE47BF4" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC751FE6E78BB" ID-CoL="WSSB" authorityName=": Okamura" authorityYear="1963" box="[136,393,1401,1422]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DD99DC751FE6E78BB" box="[136,393,1401,1422]" italics="true" pageId="47" pageNumber="48">Coelorinchus formosanus</emphasis>
</taxonomicName>
:
<bibRefCitation id="5B297ED8FFF9863DD8B5C751FD8178BA" author="Okamura, O." box="[416,614,1401,1423]" pageId="47" pageNumber="48" pagination="93 - 99" refId="ref297499" refString="Okamura, O. (1984 b) Macrouroidei. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (Eds.), The Fishes of the Japanese Archipelago. 1 st Edition. Tokai University Press, Tokyo, pp. 93 - 99, pls. 79 - 83. [In Japanese.]" type="book chapter" year="1984">Okamura 1984b:96</bibRefCitation>
, pl. 82, fig. J (compiled);
<bibRefCitation id="5B297ED8FFF9863DD82EC7B3FE157884" author="Okamura, O." box="[315,498,1435,1457]" pageId="47" pageNumber="48" pagination="344 - 373" refId="ref297606" refString="Okamura, O. (1988) Macrouroidei. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (Eds.), The Fishes of the Japanese Archipelago. 2 nd Edition. Tokai University Press, Tokyo, pp. 93 - 99, 453, pls. 79 - 83, 344 - 373." type="journal article" year="1988">Okamura 1988:96</bibRefCitation>
, pl. 82, fig. J (compiled);
<bibRefCitation id="5B297ED8FFF9863DD9A8C795FE9E78E6" author="Iwamoto, T." box="[189,377,1469,1491]" pageId="47" pageNumber="48" refId="ref287931" refString="Iwamoto, T. (1990) Family Macrouridae. In: Cohen, D. M., Inada, T., Iwamoto, T. &amp; Scialabba, N. (Eds.), FAO Species Catalogue, Vol. 10. Gadiform Fishes of the World. An Annotated and Illustrated Catalogue of Cods, Hakes, Grenadiers and Other Gadiform Fishes Known to Date. FAO, Rome, pp. 90 - 317." type="book" year="1990">Iwamoto 1990:158</bibRefCitation>
, fig. 372 (synopsis);
<bibRefCitation id="5B297ED8FFF9863DDB44C795FCE178E6" author="Shao, K. - T. &amp; Iwamoto, T. &amp; Ho, H. - C. &amp; Cheng, T. - Y. &amp; Chen, C. - Y." box="[593,774,1469,1491]" pageId="47" pageNumber="48" pagination="17 - 29" refId="ref299606" refString="Shao, K. - T., Iwamoto, T., Ho, H. - C., Cheng, T. - Y. &amp; Chen, C. - Y. (2008 b) Species composition and distribution pattern of grenadiers (family [sic] Bathygadidae, Macrouridae, and Macrourididae [sic]) from Taiwan. In: Orlov, A. M. &amp; Iwamoto, T. (Eds.), Grenadiers of the World Ocean: Biology, Stock Assessment, and Fisheries. American Fisheries Society Symposium 63. American Fisheries Society, Maryland, pp. 17 - 29." type="book chapter" year="2008">
Shao
<emphasis id="0DCCDF3BFFF9863DDB9FC796FD5978E7" box="[650,702,1469,1491]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2008b
</bibRefCitation>
: table 2 (
<specimenCount id="29BEC8A0FFF9863DD801C7F7FEBA78C1" box="[276,349,1503,1524]" pageId="47" pageNumber="48" type="generic">9 spec.</specimenCount>
listed from northeastern and southwestern
<collectingCountry id="47AF43B9FFF9863DD9A8C429FEE17B22" box="[189,262,1537,1559]" name="Taiwan" pageId="47" pageNumber="48">Taiwan</collectingCountry>
, and South
<collectingCountry id="47AF43B9FFF9863DD869C429FE527B22" box="[380,437,1537,1559]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
Sea);
<bibRefCitation id="5B297ED8FFF9863DD8E3C429FD2D7B22" author="Kim, S. - Y. &amp; Iwamoto, T. &amp; Yabe, M." box="[502,714,1537,1559]" pageId="47" pageNumber="48" pagination="106 - 117" refId="ref291511" refString="Kim, S. - Y., Iwamoto, T. &amp; Yabe, M. (2009 b) Four new records of grenadiers (Macrouridae, Gadiformes, Teleostei) from Korea. Korean Journal of Ichthyology, 21, 106 - 117." type="journal article" year="2009">
Kim
<emphasis id="0DCCDF3BFFF9863DDB3DC42AFDBC7B23" box="[552,603,1537,1559]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2009b:112
</bibRefCitation>
, fig. 6 (brief description;
<specimenCount id="29BEC8A0FFF9863DD861C40BFE5E7B0C" box="[372,441,1571,1593]" pageId="47" pageNumber="48" type="generic">2 spec.</specimenCount>
from
<collectingCountry id="47AF43B9FFF9863DD8E3C40CFDD37B0C" box="[502,564,1572,1593]" name="South Korea" pageId="47" pageNumber="48">Korea</collectingCountry>
; new Korean record);
<bibRefCitation id="5B297ED8FFF9863DD9A8C46EFE677B6E" author="Ho, H. - C. &amp; Shao, K. - T." box="[189,384,1605,1627]" pageId="47" pageNumber="48" pagination="1 - 74" refId="ref287169" refString="Ho, H. - C. &amp; Shao, K. - T. (2011) Annotated checklist and type catalog of fish genera and species described from Taiwan. Zootaxa, 2957, 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 2957.1.1" type="journal article" year="2011">Ho &amp; Shao 2011:36</bibRefCitation>
(
<typeStatus id="E003BD8BFFF9863DD89BC46DFE5E7B6F" box="[398,441,1605,1626]" pageId="47" pageNumber="48">type</typeStatus>
catalog);
<bibRefCitation id="5B297ED8FFF9863DDB0FC46DFCED7B6E" author="Nakabo, T. &amp; Kai, Y." box="[538,778,1605,1627]" pageId="47" pageNumber="48" pagination="1872 - 1876" refId="ref295555" refString="Nakabo, T. &amp; Kai, Y. (2013) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 3 rd Edition. Tokai University Press, Hadano, pp. 493 - 512, 1872 - 1876. [In Japanese.]" type="journal article" year="2013">Nakabo &amp; Kai 2013:506</bibRefCitation>
(in key;
<collectingCountry id="47AF43B9FFF9863DD818C440FEAF7B48" box="[269,328,1640,1661]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
); Iwamoto
<emphasis id="0DCCDF3BFFF9863DD8A1C440FE027B49" box="[436,485,1639,1661]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2015:54 (brief description;
<specimenCount id="29BEC8A0FFF9863DDBE7C44FFF087BAA" pageId="47" pageNumber="48" type="generic">38 spec.</specimenCount>
from northeastern Taiwan);
<bibRefCitation id="5B297ED8FFF9863DDB10C4A2FD277BAB" author="Motomura, H." box="[517,704,1673,1695]" pageId="47" pageNumber="48" refId="ref295068" refString="Motomura, H. (2020) List of Japan's All Fish Species. Current Standard Japanese and Scientific Names of All Fish Species Recorded from Japanese Waters. The Kagoshima University Museum, Kagoshima, 560 pp. [In Japanese.]" type="book" year="2020">Motomura 2020:38</bibRefCitation>
(listed;
<collectingCountry id="47AF43B9FFF9863DD9A8C484FF1F7BF4" box="[189,248,1708,1729]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
).
</paragraph>
<paragraph id="3F070329FFF9863DD99DC4E5FDEC7A5E" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC4E5FE6E7BD7" ID-CoL="WSSB" authorityName=": Okamura" authorityYear="1963" box="[136,393,1741,1762]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DD99DC4E5FE6E7BD7" box="[136,393,1741,1762]" italics="true" pageId="47" pageNumber="48">Caelorinchus formosanus</emphasis>
</taxonomicName>
:
<bibRefCitation id="5B297ED8FFF9863DD88BC4E5FDB57BD6" author="Nakabo, T." box="[414,594,1741,1763]" pageId="47" pageNumber="48" pagination="1276 - 1277" refId="ref295357" refString="Nakabo, T. (1993) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 1 st Edition. Tokai University Press, Tokyo, pp. 353 - 371, 1276 - 1277. [In Japanese.]" type="journal article" year="1993">Nakabo 1993:366</bibRefCitation>
(in key;
<collectingCountry id="47AF43B9FFF9863DDBD1C4E6FD187BD6" box="[708,767,1742,1763]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
);
<bibRefCitation id="5B297ED8FFF9863DD9A8C4C7FEB37A30" author="Shao, K. - T." box="[189,340,1775,1797]" pageId="47" pageNumber="48" pagination="165 - 173" refId="ref299442" refString="Shao, K. - T. (1993) Gadiformes. In: Shen, S. - C. (Ed.), Fishes of Taiwan. Department of Zoology, National Taiwan University, Taipei, pp. 165 - 173. [In Chinese.]" type="book chapter" year="1993">Shao 1993:168</bibRefCitation>
(compiled; Taiwan);
<bibRefCitation id="5B297ED8FFF9863DDB26C4C7FD037A31" author="Nakabo, T." box="[563,740,1775,1797]" pageId="47" pageNumber="48" pagination="417 - 435" refId="ref295412" refString="Nakabo, T. (2000) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 2 nd Edition. Tokai University Press, Tokyo, pp. 417 - 435, 1494. [In Japanese.]" type="journal article" year="2000">Nakabo 2000:430</bibRefCitation>
(in key;
<collectingCountry id="47AF43B9FFF9863DD9E5C53AFECC7A12" box="[240,299,1810,1831]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
);
<bibRefCitation id="5B297ED8FFF9863DD82AC539FE097A13" author="Nakabo, T." box="[319,494,1809,1831]" pageId="47" pageNumber="48" pagination="1491 - 1492" refId="ref295465" refString="Nakabo, T. (2002) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. English Edition. Tokai University Press, Tokyo, pp. 417 - 435, 1491 - 1492. [In Japanese.]" type="journal article" year="2002">Nakabo 2002:430</bibRefCitation>
(in key;
<collectingCountry id="47AF43B9FFF9863DDB5BC53AFD6E7A12" box="[590,649,1810,1831]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
);
<bibRefCitation id="5B297ED8FFF9863DDB8EC539FEEC7A7D" author="Yoda, M. &amp; Tokimura, M. &amp; Horikawa, H. &amp; Yamada, U." pageId="47" pageNumber="48" refId="ref303555" refString="Yoda, M., Tokimura, M., Horikawa, H. &amp; Yamada, U. (2002) A Catalogue of Fishes from the East China and Yellow Seas with Their Local Names. Seikai National Fisheries Research Institute, Fisheries Research Agency, Nagasaki, 41 pp. [In Japanese.]" type="book" year="2002">
Yoda
<emphasis id="0DCCDF3BFFF9863DDBC3C53AFCEC7A13" box="[726,779,1809,1831]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2002:11
</bibRefCitation>
(listed; East
<collectingCountry id="47AF43B9FFF9863DD893C51BFE587A7C" box="[390,447,1843,1865]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
and Yellow Seas);
<bibRefCitation id="5B297ED8FFF9863DDB61C51BFEFF7A5E" author="Shinohara, G. &amp; Sato, T. &amp; Aonuma, Y. &amp; Horikawa, H. &amp; Matsuura, K. &amp; Nakabo, T. &amp; Sato, K." pageId="47" pageNumber="48" pagination="385 - 452" refId="ref300310" refString="Shinohara, G., Sato, T., Aonuma, Y., Horikawa, H., Matsuura, K., Nakabo, T. &amp; Sato, K. (2005) Annotated checklist of deep-sea fishes from the waters around the Ryukyu Islands, Japan. National Science Museum Tokyo Monographs, (29), 385 - 452." type="journal article" year="2005">
Shinohara
<emphasis id="0DCCDF3BFFF9863DDBCEC51CFCEC7A7D" box="[731,779,1843,1865]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2005:416
</bibRefCitation>
(listed; Ryukyu Islands).
</paragraph>
<paragraph id="3F070329FFF9863DD99DC55FFE8A7A9A" blockId="47.[136,779,1027,1967]" pageId="47" pageNumber="48">
<taxonomicName id="F8B878AAFFF9863DD99DC55FFDEE7AB9" ID-CoL="WSSB" authorityName=": Okamura" authorityYear="1963" box="[136,521,1911,1933]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus" subGenus="Quincuncia">
<emphasis id="0DCCDF3BFFF9863DD99DC55FFDEE7AB9" box="[136,521,1911,1933]" italics="true" pageId="47" pageNumber="48">Caelorinchus (Quincuncia) formosanus</emphasis>
</taxonomicName>
:
<bibRefCitation id="5B297ED8FFF9863DDB02C55FFD367AB9" author="Chiou, M. - L. &amp; Shao, K. - T. &amp; Iwamoto, T." box="[535,721,1911,1933]" pageId="47" pageNumber="48" pagination="35 - 50" refId="ref282800" refString="Chiou, M. - L., Shao, K. - T. &amp; Iwamoto, T. (2004 a) A new species, Caelorinchus sheni, and 19 new records of grenadiers (Pisces: Gadiformes: Macrouridae) from Taiwan. Zoological Studies, 43, 35 - 50. http: // dx. doi. org / 10.1643 / ci- 03 - 128 r 1" type="journal article" year="2004">
Chiou
<emphasis id="0DCCDF3BFFF9863DDB4CC550FD6C7AB9" box="[601,651,1911,1933]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2004a
</bibRefCitation>
: table 1 (listed;
<collectingRegion id="FD7CCDCBFFF9863DD80DC5B1FE8E7A9A" box="[280,361,1945,1967]" country="China" name="Taiwan)" pageId="47" pageNumber="48">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFF9863DD99DC5F1FBDC7994" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="3F070329FFF9863DD99DC5F1FBDC7994" blockId="47.[136,778,2009,2070]" lastBlockId="47.[810,1452,173,2070]" pageId="47" pageNumber="48">
<emphasis id="0DCCDF3BFFF9863DD99DC5F1FEE77AC6" bold="true" box="[136,256,2009,2035]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
Light organ tubular, completely covered with scales, externally represented by long black streak extending from immediately anterior to anus to chest just posterior to isthmus. Underside of snout completely naked except for overlapping scales along lateral nasal ridges; underside of head posterior to level of midorbit heavily scaled, but anterior portions of mandibular rami mostly naked; top of snout broadly naked along each side of median rostral ridge. Snout long, sharply pointed, length 66–84% PRL, its dorsal profile slightly concave in lateral view; terminal scute short, arrowhead-shaped, dorsoventrally flattened, length 7–18% PRL. Lateral nasal ridge incompletely supported by nasal bone. Anus abutting anal-fin origin. Premaxillary teeth small, slender, conical in moderately wide, long, tapered band, with outer series slightly enlarged; posterior margin of tooth band almost reaching lateral corner of mouth. Body scales covered with short, spike-like, trihedral spinules in irregularly quincunx order to widely divergent rows; buttresses well developed. Second dorsal fin distinctly lower than anal fin. Upper-jaw length 45–51% PRL; pectoral-fin length 69–75% PRL; height of first dorsal fin 74–85% PRL; interdorsal length 35–50% PRL; length of gill slit 21–26% PRL; barbel length 20–26% PRL. Dorsal half of body with faint, dark, irregular blotches; lower half of head and body silvery white when viewed laterally; median nasal bone blackish; underside of head generally pale; black hair-like papillae scarcely developed on naked areas on head; lips dusky; oral cavity pale; gular and branchiostegal membranes heavily peppered with large melanophores; chest dark; first dorsal fin dusky, but second spinous ray prominently blackish; pelvic fin darker distally.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFF98622DA4AC681FE297B95" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="3F070329FFF98622DA4AC681FD477C80" blockId="47.[810,1452,173,2070]" lastBlockId="48.[136,778,173,2071]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="0DCCDF3BFFF9863DDA4AC681FBB679F6" bold="true" box="[863,1105,1193,1219]" pageId="47" pageNumber="48">Material examined.</emphasis>
<materialsCitation id="8FD00974FFF9863DDD74C681FB32781C" ID-GBIF-Occurrence="3750954990" collectingDate="1961-12-06" county="Da-xi" location="northwestern Taiwan" municipality="Yilan" pageId="47" pageNumber="48" specimenCount="23" typeStatus="holotype">
<specimenCount id="29BEC8A0FFF9863DDD74C681FAE379F6" box="[1121,1284,1193,1219]" pageId="47" pageNumber="48" type="generic">23 specimens</specimenCount>
.
<emphasis id="0DCCDF3BFFF9863DDC02C681FA4C79F6" bold="true" box="[1303,1451,1193,1219]" pageId="47" pageNumber="48">
<typeStatus id="E003BD8BFFF9863DDC02C681FA6379F6" box="[1303,1412,1193,1219]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
of
</emphasis>
<taxonomicName id="F8B878AAFFF9863DDA3FC6E3FB3479D0" authority=": FAKU" authorityName="FAKU" box="[810,1235,1227,1254]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DDA3FC6E3FB9D79D0" bold="true" box="[810,1146,1227,1254]" pageId="47" pageNumber="48">
<emphasis id="0DCCDF3BFFF9863DDA3FC6E3FB9679D0" bold="true" box="[810,1137,1227,1254]" italics="true" pageId="47" pageNumber="48">Coelorhynchus formosanus</emphasis>
:
</emphasis>
FAKU
</taxonomicName>
35856 (
<quantity id="F840AECCFFF9863DDC2BC6E3FA4B79D3" box="[1342,1452,1227,1254]" metricMagnitude="-2" metricUnit="m" metricValue="7.279999999999999" pageId="47" pageNumber="48" unit="mm" value="72.8">72.8 mm</quantity>
HL, 221+ mm TL),
<collectingCounty id="D6667BA5FFF9863DDD16C6C5FBA47832" box="[1027,1091,1261,1287]" pageId="47" pageNumber="48">Da-xi</collectingCounty>
fish market,
<collectingMunicipality id="DF639953FFF9863DDDC5C6C5FAF77832" box="[1232,1296,1261,1287]" pageId="47" pageNumber="48">Yilan</collectingMunicipality>
,
<location id="3A6755F2FFF9863DDC0CC6C5FC98781C" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDC0CC6C5FC98781C" county="Da-xi" municipality="Yilan" name="northwestern Taiwan" pageId="47" pageNumber="48">northwestern Taiwan</location>
, bottom trawl,
<date id="4B0625E9FFF9863DDD29C727FB32781C" box="[1084,1237,1295,1322]" pageId="47" pageNumber="48" value="1961-12-06">
<collectingDate id="5B42DC01FFF9863DDD29C727FB32781C" box="[1084,1237,1295,1322]" pageId="47" pageNumber="48" value="1961-12-06">6 Dec. 1961</collectingDate>
</date>
</materialsCitation>
.
<emphasis id="0DCCDF3BFFF9863DDDF2C727FA64781C" bold="true" box="[1255,1411,1295,1321]" pageId="47" pageNumber="48">
<typeStatus id="E003BD8BFFF9863DDDF2C727FAB8781C" box="[1255,1375,1295,1321]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
of
</emphasis>
<taxonomicName id="F8B878AAFFF9863DDC87C727FBF77879" authority=": FAKU" authorityName="FAKU" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFF9863DDC87C727FC5B787E" bold="true" pageId="47" pageNumber="48">
<emphasis id="0DCCDF3BFFF9863DDC87C727FC54787E" bold="true" italics="true" pageId="47" pageNumber="48">C. formosanus</emphasis>
:
</emphasis>
FAKU
</taxonomicName>
35859 (1,
<quantity id="F840AECCFFF9863DDD81C719FB1C7879" box="[1172,1275,1329,1356]" metricMagnitude="-2" metricUnit="m" metricValue="6.2299999999999995" pageId="47" pageNumber="48" unit="mm" value="62.3">62.3 mm</quantity>
HL, 238+ mm TL),
<materialsCitation id="8FD00974FFF9863DDA70C77CFAA178A5" ID-GBIF-Occurrence="3750955293" collectionCode="FAKU" collectorName="Yilan" location="Daxi" pageId="47" pageNumber="48" specimenCode="FAKU 35860" specimenCount="1" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDA70C77CFC19785B" box="[869,1022,1363,1390]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="47" pageNumber="48">FAKU 35860</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDD38C77BFB74785B" box="[1069,1171,1363,1390]" metricMagnitude="-2" metricUnit="m" metricValue="5.779999999999999" pageId="47" pageNumber="48" unit="mm" value="57.8">57.8 mm</quantity>
HL, 212+ mm TL),
<location id="3A6755F2FFF9863DDC94C77CFCA778BA" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDC94C77CFCA778BA" name="Daxi" pageId="47" pageNumber="48">Daxi</location>
fish market,
<collectorName id="924D66FFFFF9863DDAC2C75DFBF078BA" box="[983,1047,1397,1423]" pageId="47" pageNumber="48">Yilan</collectorName>
, bottom trawl, 1958–1961
</materialsCitation>
; *
<materialsCitation id="8FD00974FFF9863DDC4BC75DFB137887" ID-GBIF-Occurrence="3750955294" collectionCode="FAKU" pageId="47" pageNumber="48" specimenCode="FAKU 35857" specimenCount="1" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC4BC75DFC977887" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="47" pageNumber="48">FAKU 35857</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDAB6C7BFFBEC7887" box="[931,1035,1431,1458]" metricMagnitude="-2" metricUnit="m" metricValue="4.67" pageId="47" pageNumber="48" unit="mm" value="46.7">46.7 mm</quantity>
HL, 167+ mm TL)
</materialsCitation>
, *
<materialsCitation id="8FD00974FFF9863DDC18C7B0FAB178C3" ID-GBIF-Occurrence="3750955259" collectionCode="FAKU" collectorName="Non-types" country="Japan" location="South China Sea" municipality="Kaoshiung" pageId="47" pageNumber="48" specimenCode="FAKU 35858" specimenCount="1" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC18C7B0FA4C7887" box="[1293,1451,1431,1458]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="47" pageNumber="48">FAKU 35858</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA5AC791FC5478E1" box="[847,947,1465,1492]" metricMagnitude="-2" metricUnit="m" metricValue="5.54" pageId="47" pageNumber="48" unit="mm" value="55.4">55.4 mm</quantity>
HL, 195+ mm TL),
<collectingMunicipality id="DF639953FFF9863DDD8CC791FAF278E6" box="[1177,1301,1465,1491]" pageId="47" pageNumber="48">Kaoshiung</collectingMunicipality>
,
<location id="3A6755F2FFF9863DDC35C791FCB278C3" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDC35C791FCB278C3" country="Japan" municipality="Kaoshiung" name="South China Sea" pageId="47" pageNumber="48">South China Sea</location>
, bottom trawl, 1958–1961.
<emphasis id="0DCCDF3BFFF9863DDD97C7F4FAB178C3" bold="true" box="[1154,1366,1500,1526]" pageId="47" pageNumber="48">
<collectorName id="924D66FFFFF9863DDD97C7F4FAE778C3" box="[1154,1280,1500,1526]" pageId="47" pageNumber="48">Non-types</collectorName>
:
<collectingCountry id="47AF43B9FFF9863DDC1CC7F4FAB178C3" box="[1289,1366,1500,1526]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
</emphasis>
</materialsCitation>
<emphasis id="0DCCDF3BFFF9863DDC42C7F4FABB78C3" bold="true" box="[1367,1372,1500,1526]" pageId="47" pageNumber="48">:</emphasis>
<materialsCitation id="8FD00974FFF9863DDC74C7F4FB807B0F" ID-GBIF-Occurrence="3750954581" collectingDate="1951-03-21" collectionCode="BSKU" country="Japan" location="Mimase" pageId="47" pageNumber="48" specimenCode="BSKU 159" specimenCount="1" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC74C7F4FCB37B2D" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 159</specimenCode>
(1, 69.0 mm HL, 272+ mm TL),
<location id="3A6755F2FFF9863DDDD3C7D5FAC77B22" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDDD3C7D5FAC77B22" box="[1222,1312,1533,1559]" country="Japan" name="Mimase" pageId="47" pageNumber="48">Mimase</location>
fish market, bottom trawl,
<date id="4B0625E9FFF9863DDADEC408FB807B0F" box="[971,1127,1567,1594]" pageId="47" pageNumber="48" value="1951-03-21">
<collectingDate id="5B42DC01FFF9863DDADEC408FB807B0F" box="[971,1127,1567,1594]" pageId="47" pageNumber="48" value="1951-03-21">21 Mar. 1951</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="8FD00974FFF9863DDD61C437FC727B4B" ID-GBIF-Occurrence="3750954532" collectingDate="2012-07-25" collectionCode="BSKU" country="Taiwan" location="Da-xi" pageId="47" pageNumber="48" specimenCode="BSKU 108963" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<emphasis id="0DCCDF3BFFF9863DDD61C437FB327B0C" bold="true" box="[1140,1237,1567,1593]" pageId="47" pageNumber="48">
<collectingCountry id="47AF43B9FFF9863DDD61C437FB377B0C" box="[1140,1232,1567,1593]" name="Taiwan" pageId="47" pageNumber="48">Taiwan</collectingCountry>
:
</emphasis>
<specimenCode id="6F1EAB52FFF9863DDDC8C408FA637B0C" box="[1245,1412,1567,1594]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 108963</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA3FC46AFC697B69" box="[810,910,1602,1628]" metricMagnitude="-2" metricUnit="m" metricValue="7.99" pageId="47" pageNumber="48" unit="mm" value="79.9">79.9 mm</quantity>
HL, 318+ mm TL),
<location id="3A6755F2FFF9863DDD66C469FB547B6E" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDD66C469FB547B6E" box="[1139,1203,1601,1627]" country="Taiwan" name="Da-xi" pageId="47" pageNumber="48" stateProvince="Yilan">Da-xi</location>
fish market,
<collectingRegion id="FD7CCDCBFFF9863DDC50C469FA627B6E" box="[1349,1413,1601,1627]" country="Taiwan" name="Yilan" pageId="47" pageNumber="48">Yilan</collectingRegion>
,
<date id="4B0625E9FFF9863DDC9AC469FC727B4B" pageId="47" pageNumber="48" value="2012-07-25">
<collectingDate id="5B42DC01FFF9863DDC9AC469FC727B4B" pageId="47" pageNumber="48" value="2012-07-25">25 Jul. 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8FD00974FFF9863DDAB5C44CFA407B95" ID-GBIF-Occurrence="3750955232" collectingDate="2007-11-08" collectionCode="BSKU" collectorName="H. - C. Ho" country="Taiwan" location="Da-xi" pageId="47" pageNumber="48" specimenCode="BSKU 98972" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDAB5C44CFBDF7B4B" box="[928,1080,1636,1662]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 98972</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDD73C44BFB2D7B4B" box="[1126,1226,1635,1662]" metricMagnitude="-2" metricUnit="m" metricValue="5.79" pageId="47" pageNumber="48" unit="mm" value="57.9">57.9 mm</quantity>
HL, 216+ mm TL),
<location id="3A6755F2FFF9863DDA3FC4ADFC8D7BAA" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDA3FC4ADFC8D7BAA" box="[810,874,1669,1695]" country="Taiwan" name="Da-xi" pageId="47" pageNumber="48" stateProvince="Yilan">Da-xi</location>
fish market,
<collectingRegion id="FD7CCDCBFFF9863DDD1CC4ADFBAE7BAA" box="[1033,1097,1669,1695]" country="Taiwan" name="Yilan" pageId="47" pageNumber="48">Yilan</collectingRegion>
, coll.
<collectorName id="924D66FFFFF9863DDD81C4AEFAE57B95" box="[1172,1282,1670,1696]" pageId="47" pageNumber="48">H.-C. Ho</collectorName>
,
<date id="4B0625E9FFF9863DDC07C4AEFA407B95" box="[1298,1447,1670,1696]" pageId="47" pageNumber="48" value="2007-11-08">
<collectingDate id="5B42DC01FFF9863DDC07C4AEFA407B95" box="[1298,1447,1670,1696]" pageId="47" pageNumber="48" value="2007-11-08">8 Nov. 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8FD00974FFF9863DDA3FC480FAA87BF7" ID-GBIF-Occurrence="3750955038" box="[810,1359,1703,1730]" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50363" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDA3FC480FC2F7BF7" box="[810,968,1703,1730]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50363</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDD14C48FFB8C7BF7" box="[1025,1131,1703,1730]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" pageId="47" pageNumber="48" unit="mm" value="51.3">51.3 mm</quantity>
HL,
<quantity id="F840AECCFFF9863DDDA4C48FFAF27BF7" box="[1201,1301,1703,1730]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="47" pageNumber="48" unit="mm" value="215.0">215 mm</quantity>
TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDC74C480FB077BD1" ID-GBIF-Occurrence="3750955345" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50466" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC74C480FC977BD1" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50466</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA8BC4E1FBE57BD1" box="[926,1026,1737,1764]" metricMagnitude="-2" metricUnit="m" metricValue="3.76" pageId="47" pageNumber="48" unit="mm" value="37.6">37.6 mm</quantity>
HL, 128+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDDF8C4E2FB977A33" ID-GBIF-Occurrence="3750955440" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50468" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDDF8C4E2FA627BD1" box="[1261,1413,1737,1764]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50468</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA3FC4C3FC687A33" box="[810,911,1771,1798]" metricMagnitude="-2" metricUnit="m" metricValue="5.32" pageId="47" pageNumber="48" unit="mm" value="53.2">53.2 mm</quantity>
HL, 189+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDD68C4C4FBE77A1D" ID-GBIF-Occurrence="3750955350" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50469" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDD68C4C4FAF07A33" box="[1149,1303,1771,1798]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50469</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDC53C4C4FA4B7A33" box="[1350,1452,1772,1798]" metricMagnitude="-2" metricUnit="m" metricValue="3.8899999999999997" pageId="47" pageNumber="48" unit="mm" value="38.9">38.9 mm</quantity>
HL, 152+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDD19C526FC6C7A7F" ID-GBIF-Occurrence="3750955205" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50471" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDD19C526FB437A1D" box="[1036,1188,1805,1832]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50471</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDDC7C525FAD17A1D" box="[1234,1334,1805,1832]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" pageId="47" pageNumber="48" unit="mm" value="51.8">51.8 mm</quantity>
HL, 210+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDA8DC518FA407A7F" ID-GBIF-Occurrence="3750954678" box="[920,1447,1839,1866]" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50472" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDA8DC518FBD67A7F" box="[920,1073,1839,1866]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50472</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDD74C507FB217A7F" box="[1121,1222,1839,1866]" metricMagnitude="-2" metricUnit="m" metricValue="5.43" pageId="47" pageNumber="48" unit="mm" value="54.3">54.3 mm</quantity>
HL, 200+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDA3FC57AFAB77A59" ID-GBIF-Occurrence="3750955365" box="[810,1360,1873,1900]" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50473" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDA3FC57AFC217A59" box="[810,966,1873,1900]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50473</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDAEEC579FB837A59" box="[1019,1124,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="47" pageNumber="48" unit="mm" value="59.2">59.2 mm</quantity>
HL, 214+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDC74C57AFB077ABB" ID-GBIF-Occurrence="3750954772" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 50474" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC74C57AFC977ABB" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50474</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA8BC55BFBE57ABB" box="[926,1026,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="5.74" pageId="47" pageNumber="48" unit="mm" value="57.4">57.4 mm</quantity>
HL, 217+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF9863DDDF8C55CFB187AE7" ID-GBIF-Occurrence="3750954383" collectingDate="2012-04-23" collectionCode="BSKU" collectorName="T. Yamakawa &amp; R. Asaoka" country="Taiwan" location="Da-xi" pageId="47" pageNumber="48" specimenCode="BSKU 50488" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDDF8C55CFA627ABB" box="[1261,1413,1907,1934]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 50488</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDA3FC5BDFC6A7A85" box="[810,909,1941,1968]" metricMagnitude="-2" metricUnit="m" metricValue="3.86" pageId="47" pageNumber="48" unit="mm" value="38.6">38.6 mm</quantity>
HL,
<quantity id="F840AECCFFF9863DDAD0C5BDFBC57A85" box="[965,1058,1941,1968]" metricMagnitude="-1" metricUnit="m" metricValue="1.58" pageId="47" pageNumber="48" unit="mm" value="158.0">158 mm</quantity>
TL),
<location id="3A6755F2FFF9863DDD75C5BEFB477A85" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDD75C5BEFB477A85" box="[1120,1184,1942,1968]" country="Taiwan" name="Da-xi" pageId="47" pageNumber="48" stateProvince="Yilan">Da-xi</location>
fish market,
<collectingRegion id="FD7CCDCBFFF9863DDC25C5BEFA977A85" box="[1328,1392,1942,1968]" country="Taiwan" name="Yilan" pageId="47" pageNumber="48">Yilan</collectingRegion>
, coll.
<collectorName id="924D66FFFFF9863DDA3FC590FC587AE7" box="[810,959,1976,2002]" pageId="47" pageNumber="48">T. Yamakawa</collectorName>
and
<collectorName id="924D66FFFFF9863DDAE4C590FB827AE7" box="[1009,1125,1976,2002]" pageId="47" pageNumber="48">R.Asaoka</collectorName>
,
<date id="4B0625E9FFF9863DDD7BC590FB187AE7" box="[1134,1279,1976,2002]" pageId="47" pageNumber="48" value="2012-04-23">
<collectingDate id="5B42DC01FFF9863DDD7BC590FB187AE7" box="[1134,1279,1976,2002]" pageId="47" pageNumber="48" value="2012-04-23">23Apr. 2012</collectingDate>
</date>
</materialsCitation>
; *
<materialsCitation id="8FD00974FFF9863DDC06C590FB467AC1" ID-GBIF-Occurrence="3750954663" collectionCode="BSKU" country="Taiwan" pageId="47" pageNumber="48" specimenCode="BSKU 99863" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDC06C590FA4C7AE7" box="[1299,1451,1976,2002]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 99863</specimenCode>
(1, ca.
<quantity id="F840AECCFFF9863DDA6EC5F2FC2B7AC1" box="[891,972,2010,2036]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="47" pageNumber="48" unit="mm" value="72.0">72 mm</quantity>
HL,
<quantity id="F840AECCFFF9863DDD1EC5F2FB8C7AC1" box="[1035,1131,2010,2036]" metricMagnitude="-1" metricUnit="m" metricValue="3.02" pageId="47" pageNumber="48" unit="mm" value="302.0">302 mm</quantity>
TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFF98622DDBAC5F2FE427DF2" ID-GBIF-Occurrence="3750955221" collectingDate="2009-06-30" collectionCode="BSKU" collectorName="M. - Y. Lee" country="Taiwan" lastPageId="48" lastPageNumber="49" location="Da-xi" pageId="47" pageNumber="48" specimenCode="BSKU 99866" specimenCount="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCode id="6F1EAB52FFF9863DDDBAC5F2FAAE7AC1" box="[1199,1353,2010,2036]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="47" pageNumber="48">BSKU 99866</specimenCode>
(1,
<quantity id="F840AECCFFF9863DDC6FC5F2FCB17523" metricMagnitude="-2" metricUnit="m" metricValue="6.609999999999999" pageId="47" pageNumber="48" unit="mm" value="66.1">66.1 mm</quantity>
HL, 216+ mm TL),
<location id="3A6755F2FFF9863DDD5BC5D4FB697523" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFF9863DDD5BC5D4FB697523" box="[1102,1166,2044,2070]" country="Taiwan" name="Da-xi" pageId="47" pageNumber="48" stateProvince="Yilan">Da-xi</location>
fish market,
<collectingRegion id="FD7CCDCBFFF9863DDC39C5D4FA8B7523" box="[1324,1388,2044,2070]" country="Taiwan" name="Yilan" pageId="47" pageNumber="48">Yilan</collectingRegion>
, coll.
<collectorName id="924D66FFFFE68622D99DC285FF187DF2" box="[136,255,173,199]" pageId="48" pageNumber="49">M.-Y. Lee</collectorName>
,
<date id="4B0625E9FFE68622D818C285FE427DF2" box="[269,421,173,199]" pageId="48" pageNumber="49" value="2009-06-30">
<collectingDate id="5B42DC01FFE68622D818C285FE427DF2" box="[269,421,173,199]" pageId="48" pageNumber="49" value="2009-06-30">30 Jun. 2009</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8FD00974FFE68622D8A6C285FD577C3E" ID-GBIF-Occurrence="3750955494" collectingDate="2011-07-01" collectorName="M. Matsunuma &amp; H. - C. Ho" country="Taiwan" elevation="400" location="Kaohsiung" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Kaohsiung" typeStatus="paratype">
KAUM-I. 44655 (1,
<quantity id="F840AECCFFE68622DBB1C285FCED7DF2" box="[676,778,173,199]" metricMagnitude="-2" metricUnit="m" metricValue="6.97" pageId="48" pageNumber="49" unit="mm" value="69.7">69.7 mm</quantity>
HL, 204+ mm TL), off
<collectingRegion id="FD7CCDCBFFE68622D88FC2E7FDF17DDC" box="[410,534,207,233]" country="Taiwan" name="Kaohsiung" pageId="48" pageNumber="49">Kaohsiung</collectingRegion>
,
<collectingRegion id="FD7CCDCBFFE68622DB36C2E7FD847DDC" box="[547,611,207,233]" country="Taiwan" name="Yilan" pageId="48" pageNumber="49">Yilan</collectingRegion>
,
<quantity id="F840AECCFFE68622DB65C2E7FD5C7DDC" box="[624,699,207,233]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="m" value="400.0">
<elevation id="B495E41AFFE68622DB65C2E7FD5C7DDC" box="[624,699,207,233]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="m" value="400.0">400 m</elevation>
</quantity>
, trawl, coll.
<collectorName id="924D66FFFFE68622D9D6C2D9FE957C3E" box="[195,370,241,267]" pageId="48" pageNumber="49">M. Matsunuma</collectorName>
and
<collectorName id="924D66FFFFE68622D8BAC2D9FDFA7C3E" box="[431,541,241,267]" pageId="48" pageNumber="49">H.-C. Ho</collectorName>
,
<date id="4B0625E9FFE68622DB3EC2D9FD577C3E" box="[555,688,241,267]" pageId="48" pageNumber="49" value="2011-07-01">
<collectingDate id="5B42DC01FFE68622DB3EC2D9FD577C3E" box="[555,688,241,267]" pageId="48" pageNumber="49" value="2011-07-01">1 Jul. 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8FD00974FFE68622DBAAC2D9FDD87C18" ID-GBIF-Occurrence="3750955437" collectionCode="BSKU" country="Taiwan" pageId="48" pageNumber="49" specimenCode="BSKU 29993" specimenCount="1" stateProvince="Kaohsiung" typeStatus="paratype">
<specimenCode id="6F1EAB52FFE68622DBAAC2D9FF297C18" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="48" pageNumber="49">BSKU 29993</specimenCode>
(1,
<quantity id="F840AECCFFE68622D9E9C33BFE867C18" box="[252,353,275,302]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="48" pageNumber="49" unit="mm" value="52.5">52.5 mm</quantity>
HL, 184+ mm TL)
</materialsCitation>
,
<materialsCitation id="8FD00974FFE68622DB5EC33BFDAB7C44" ID-GBIF-Occurrence="3750955540" collectingDate="1978-12-06" collectingDateMax="1978-12-07" collectingDateMin="1978-12-06" collectionCode="BSKU" collectorName="T. Kanayama" country="Taiwan" location="Dong-gang" pageId="48" pageNumber="49" specimenCode="BSKU 29994" specimenCount="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCode id="6F1EAB52FFE68622DB5EC33BFD037C18" box="[587,740,275,301]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="48" pageNumber="49">BSKU 29994</specimenCode>
(1,
<quantity id="F840AECCFFE68622D99DC31DFF0A7C7A" box="[136,237,309,335]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="48" pageNumber="49" unit="mm" value="40.7">40.7 mm</quantity>
HL, 155+ mm TL),
<location id="3A6755F2FFE68622D8C3C31DFDB07C7A" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFE68622D8C3C31DFDB07C7A" box="[470,599,309,335]" country="Taiwan" name="Dong-gang" pageId="48" pageNumber="49" stateProvince="Pingtung">Dong-gang</location>
,
<collectingRegion id="FD7CCDCBFFE68622DB76C31DFD2C7C7A" box="[611,715,309,335]" country="Taiwan" name="Pingtung" pageId="48" pageNumber="49">Pingtung</collectingRegion>
, otter trawl, coll.
<collectorName id="924D66FFFFE68622D813C37FFE7A7C44" box="[262,413,343,369]" pageId="48" pageNumber="49">T. Kanayama</collectorName>
,
<date id="4B0625E9FFE68622D8B2C37FFDAB7C44" box="[423,588,343,369]" pageId="48" pageNumber="49" value="1978-12-06" valueMax="1978-12-07" valueMin="1978-12-06">
<collectingDate id="5B42DC01FFE68622D8B2C37FFDAB7C44" box="[423,588,343,369]" pageId="48" pageNumber="49" value="1978-12-06" valueMax="1978-12-07" valueMin="1978-12-06">6–7 Dec. 1978</collectingDate>
</date>
</materialsCitation>
; *
<materialsCitation id="8FD00974FFE68622DB77C37FFD7A7C80" ID-GBIF-Occurrence="3750954945" collectingDate="2017-02-27" collectionCode="FAKU" collectorName="N. Nakayama" country="Taiwan" location="Dong-gang" pageId="48" pageNumber="49" specimenCode="FAKU 204426" specimenCount="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCode id="6F1EAB52FFE68622DB77C37FFCED7C44" box="[610,778,343,369]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="48" pageNumber="49">FAKU 204426</specimenCode>
(1, 367+ mm TL),
<location id="3A6755F2FFE68622D863C351FE137CA6" LSID="urn:lsid:plazi:treatment:B711B23FFFF98620D99DC15FFE187B2D:3A6755F2FFE68622D863C351FE137CA6" box="[374,500,377,403]" country="Taiwan" name="Dong-gang" pageId="48" pageNumber="49" stateProvince="Pingtung">Dong-gang</location>
fish market,
<collectingRegion id="FD7CCDCBFFE68622DB8BC351FCE17CA6" box="[670,774,377,403]" country="Taiwan" name="Pingtung" pageId="48" pageNumber="49">Pingtung</collectingRegion>
, bottom trawl, coll.
<collectorName id="924D66FFFFE68622D84BC3B3FE1C7C80" box="[350,507,411,437]" pageId="48" pageNumber="49">N. Nakayama</collectorName>
,
<date id="4B0625E9FFE68622DB13C3B3FD7A7C80" box="[518,669,411,437]" pageId="48" pageNumber="49" value="2017-02-27">
<collectingDate id="5B42DC01FFE68622DB13C3B3FD7A7C80" box="[518,669,411,437]" pageId="48" pageNumber="49" value="2017-02-27">27 Feb. 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="3F070329FFE68622D9A8C395FD1C7FF3" blockId="48.[136,778,173,2071]" pageId="48" pageNumber="49">
<emphasis id="0DCCDF3BFFE68622D9A8C395FDE77CE2" bold="true" box="[189,512,445,471]" pageId="48" pageNumber="49">Counts and measurements.</emphasis>
Based on
<specimenCount id="29BEC8A0FFE68622DB66C395FCED7CE2" box="[627,778,445,471]" pageId="48" pageNumber="49" type="generic">19 specimens</specimenCount>
(
<quantity id="F840AECCFFE68622D985C3F7FEDF7CCC" box="[144,312,479,505]" metricMagnitude="-2" metricUnit="m" metricValue="5.875" metricValueMax="7.99" metricValueMin="3.76" pageId="48" pageNumber="49" unit="mm" value="58.75" valueMax="79.9" valueMin="37.6">37.6–79.9 mm</quantity>
HL, 128+–318+ mm TL). Counts: first dorsal-fin rays II,8–10; pectoral-fin rays i15–i17; pelvicfin rays 7; gill rakers on first arch (outer/inner) 0/7–9, on second arch 6–7/8–9; longitudinal scales 36–45; transverse scale rows below first dorsal-fin origin 4.5– 6.5, below first dorsal-fin midbase 3.5–5, below second dorsal-fin origin 3.5–5, above anal-fin origin 11.5–14.5.
</paragraph>
<paragraph id="3F070329FFE68622D9A8C0E5FE297B69" blockId="48.[136,778,173,2071]" pageId="48" pageNumber="49">The following measurements are in % of HL, followed by those in % of PRL in parentheses: snout length 40–46 (66–84); orbit diameter 23–27 (41–47); postorbital length 30–36 (54–59); postrostral length 54– 61; orbit–preopercle distance 33–37 (58–66); suborbital width 10–12 (17–20); upper-jaw length 25–30 (45–51); length of rictus 20–24 (36–43); length of premaxillary tooth band 17–22 (30–36); preoral length 33–39 (55–72); length of terminal snout scute 4–10 (7–18); length of lateral nasal ridge 26–34 (43–61); length of suborbital ridge 93–99 (157–180); snout width 22–25 (38–45); internasal width 18–21 (31–38); interorbital width 21–24 (37–43); occipital width 6–9 (10–15); body width over pectoral-fin bases 32–44 (58–81); body depth at first dorsal-fin origin 47–57 (82–104); body depth at anal-fin origin 41–47 (70–86); prepelvic length 100–109 (171– 195); preanus length 141–152 (249–270); preanal length 144–156 (256–273); isthmus–pelvic distance 30–39 (53– 70); isthmus–anus distance 71–83 (126–146); isthmusanal distance 73–87 (130–153); pelvic–anal distance 40–54 (72–94); anus–anal distance 2–4 (4–7); pelvic-fin length 26–35 (47–61); pectoral-fin length 38–44 (69–75); predorsal length 102–110 (178–200); height of first dorsal fin 41–47 (74–85); length of first dorsal-fin base 17–20 (29–36); interdorsal length 20–28 (35–50); length of gill slit 12–14 (21–26); length of posterior nostril 6–8 (9–14); barbel length 11–15 (20–26).</paragraph>
<paragraph id="3F070329FFE68622D9A8C44BFE297B95" blockId="48.[136,778,173,2071]" pageId="48" pageNumber="49">
<emphasis id="0DCCDF3BFFE68622D9A8C44BFF127B48" bold="true" box="[189,245,1635,1661]" pageId="48" pageNumber="49">Size.</emphasis>
To about
<quantity id="F840AECCFFE68622D872C44CFE4A7B4B" box="[359,429,1636,1662]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="48" pageNumber="49" unit="cm" value="32.0">32 cm</quantity>
TL (
<collectionCode id="59A99BECFFE68622D8F0C44CFDD67B4B" box="[485,561,1636,1662]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="48" pageNumber="49">BSKU</collectionCode>
108963, 318+ mm TL, Da-xi, northern
<collectingRegion id="FD7CCDCBFFE68622D87EC4ADFE2C7B95" box="[363,459,1669,1696]" country="China" name="Taiwan)" pageId="48" pageNumber="49">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFE68622D9A8C48FFD2D7AE7" pageId="48" pageNumber="49" type="distribution">
<paragraph id="3F070329FFE68622D9A8C48FFD2D7AE7" blockId="48.[136,778,173,2071]" pageId="48" pageNumber="49">
<emphasis id="0DCCDF3BFFE68622D9A8C48FFEB07BF4" bold="true" box="[189,343,1703,1729]" pageId="48" pageNumber="49">Distribution.</emphasis>
Restricted to the northwestern Pacific off
<collectingCountry id="47AF43B9FFE68622D9A5C4E1FEE27BD6" box="[176,261,1737,1763]" name="Taiwan" pageId="48" pageNumber="49">Taiwan</collectingCountry>
,
<collectingCountry id="47AF43B9FFE68622D807C4E2FEB27BD1" box="[274,341,1738,1764]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
, and
<collectingCountry id="47AF43B9FFE68622D887C4E2FE3D7BD1" box="[402,474,1738,1764]" name="South Korea" pageId="48" pageNumber="49">Korea</collectingCountry>
. Known from the northern South
<collectingCountry id="47AF43B9FFE68622D9C4C4C3FEF37A30" box="[209,276,1771,1797]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
Sea,
<collectingRegion id="FD7CCDCBFFE68622D847C4C3FE507A30" box="[338,439,1771,1797]" country="Japan" name="Okinawa" pageId="48" pageNumber="49">Okinawa</collectingRegion>
Trough, Tosa Bay, and Sea of
<collectingCountry id="47AF43B9FFE68622D99DC526FF2F7A1D" box="[136,200,1806,1832]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
off the Korean Peninsula (Chu &amp; Lo
<emphasis id="0DCCDF3BFFE68622DB6EC526FD767A12" box="[635,657,1806,1831]" italics="true" pageId="48" pageNumber="49">in</emphasis>
<bibRefCitation id="5B297ED8FFE68622DB8CC525FF247A7C" author="Chu, Y. - T. &amp; Chan, C. - L. &amp; Chen, C. - T." pageId="48" pageNumber="49" refId="ref282947" refString="Chu, Y. - T., Chan, C. - L. &amp; Chen, C. - T. (1963) Ichthyofauna of East China Sea. Science Press, Beijing. xxviii + 642 pp. [In Chinese.]" type="book" year="1963">
Chu
<emphasis id="0DCCDF3BFFE68622DBC5C526FCED7A12" box="[720,778,1805,1831]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
1963
</bibRefCitation>
;
<bibRefCitation id="5B297ED8FFE68622D9C7C507FE467A7F" author="Shao, K. - T. &amp; Ho, H. - C. &amp; Lin, P. - L. &amp; Lee, P. - F. &amp; Lee, M. - Y. &amp; Tsai, C. - Y. &amp; Liao, Y. - C. &amp; Lin, Y. - C. &amp; Chen, J. - P. &amp; Yeh, H. - M." box="[210,417,1839,1866]" pageId="48" pageNumber="49" pagination="233 - 271" refId="ref299493" refString="Shao, K. - T., Ho, H. - C., Lin, P. - L., Lee, P. - F., Lee, M. - Y., Tsai, C. - Y., Liao, Y. - C., Lin, Y. - C., Chen, J. - P. &amp; Yeh, H. - M. (2008 a) A checklist of the fishes of southern Taiwan, northern South China Sea. The Raffles Bulletin of Zoology, (suppl. 19), 233 - 271." type="journal article" year="2008">
Shao
<emphasis id="0DCCDF3BFFE68622D801C518FEB77A7C" box="[276,336,1839,1865]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2008a
</bibRefCitation>
,
<bibRefCitation id="5B297ED8FFE68622D8A5C507FE1E7A7C" author="Shao, K. - T. &amp; Iwamoto, T. &amp; Ho, H. - C. &amp; Cheng, T. - Y. &amp; Chen, C. - Y." box="[432,505,1839,1865]" pageId="48" pageNumber="49" pagination="17 - 29" refId="ref299606" refString="Shao, K. - T., Iwamoto, T., Ho, H. - C., Cheng, T. - Y. &amp; Chen, C. - Y. (2008 b) Species composition and distribution pattern of grenadiers (family [sic] Bathygadidae, Macrouridae, and Macrourididae [sic]) from Taiwan. In: Orlov, A. M. &amp; Iwamoto, T. (Eds.), Grenadiers of the World Ocean: Biology, Stock Assessment, and Fisheries. American Fisheries Society Symposium 63. American Fisheries Society, Maryland, pp. 17 - 29." type="book chapter" year="2008">2008b</bibRefCitation>
;
<bibRefCitation id="5B297ED8FFE68622DB1DC507FD357A7C" author="Kim, S. - Y. &amp; Iwamoto, T. &amp; Yabe, M." box="[520,722,1839,1865]" pageId="48" pageNumber="49" pagination="106 - 117" refId="ref291511" refString="Kim, S. - Y., Iwamoto, T. &amp; Yabe, M. (2009 b) Four new records of grenadiers (Macrouridae, Gadiformes, Teleostei) from Korea. Korean Journal of Ichthyology, 21, 106 - 117." type="journal article" year="2009">
Kim
<emphasis id="0DCCDF3BFFE68622DB51C518FD987A7C" box="[580,639,1839,1865]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2009b
</bibRefCitation>
; this study; Appendix 3-2A). Depth range
<quantity id="F840AECCFFE68622DB21C579FD507A59" box="[564,695,1873,1900]" metricMagnitude="2" metricUnit="m" metricValue="2.79" metricValueMax="4.0" metricValueMin="1.58" pageId="48" pageNumber="49" unit="m" value="279.0" valueMax="400.0" valueMin="158.0">158–400 m</quantity>
, plus a single capture in
<quantity id="F840AECCFFE68622D85DC55BFE347ABB" box="[328,467,1907,1934]" metricMagnitude="3" metricUnit="m" metricValue="1.0474999999999999" metricValueMax="1.1099999999999999" metricValueMin="0.985" pageId="48" pageNumber="49" unit="m" value="1047.5" valueMax="1110.0" valueMin="985.0">985–1110 m</quantity>
(ASIZP 65595; Iwamoto
<emphasis id="0DCCDF3BFFE68622DBE0C55CFF427A85" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2015). One of the most common grenadiers found in fish markets in
<collectingCountry id="47AF43B9FFE68622D820C590FE6D7AE7" box="[309,394,1976,2002]" name="Taiwan" pageId="48" pageNumber="49">Taiwan</collectingCountry>
, but rare in Japanese waters.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FFE68620D9A8C5F2FE187B2D" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49" type="discussion">
<paragraph id="3F070329FFE68622D9A8C5F2FC0D7CCC" blockId="48.[136,778,173,2071]" lastBlockId="48.[810,1452,173,2070]" pageId="48" pageNumber="49">
<emphasis id="0DCCDF3BFFE68622D9A8C5F2FED67AC1" bold="true" box="[189,305,2010,2036]" pageId="48" pageNumber="49">Remarks.</emphasis>
<taxonomicName id="F8B878AAFFE68622D82EC5F2FDB87AC1" authorityName=": Okamura" authorityYear="1963" box="[315,607,2010,2036]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE68622D82EC5F2FDB87AC1" box="[315,607,2010,2036]" italics="true" pageId="48" pageNumber="49">Coelorinchus formosanus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="5B297ED8FFE68622D9BBC5D4FE9F7522" author="Okamura, O." box="[174,376,2044,2071]" pageId="48" pageNumber="49" pagination="37 - 42" refId="ref297236" refString="Okamura, O. (1963 b) A new macrouroid fish found in the adjacent waters of Formosa. Bulletin of the Misaki Marine Biological Institute, Kyoto University, (4), 37 - 42." type="journal article" year="1963">Okamura (1963b)</bibRefCitation>
based on five specimens collected from northeastern
<collectingCountry id="47AF43B9FFE68622DD0DC285FB8D7DF2" box="[1048,1130,173,199]" name="Taiwan" pageId="48" pageNumber="49">Taiwan</collectingCountry>
(
<figureCitation id="A7831FACFFE68622DD9FC285FB0A7DF2" box="[1162,1261,173,199]" captionStart="FIGURE 33" captionStartId="49.[136,235,1467,1491]" captionTargetBox="[151,1436,907,1430]" captionTargetId="figure-266@49.[151,1436,907,1430]" captionTargetPageId="49" captionText="FIGURE 33. Holotype of Coelorhynchus formosanus (= Coelorinchus formosanus). FAKU 35856, 72.8 mm HL, 221+ mm TL, Da-xi fish market, northeastern Taiwan, East China Sea. (A) Lateral view; (B) dorsal and (C) ventral views of the head and trunk. Preserved condition. [Photos: N. Nakayama]" figureDoi="http://doi.org/10.5281/zenodo.6422856" httpUri="https://zenodo.org/record/6422856/files/figure.png" pageId="48" pageNumber="49">Fig. 33</figureCitation>
). The original description is excellent and can be referred to for details. In the same year, Chu &amp; Lo (
<emphasis id="0DCCDF3BFFE68622DD7EC2DAFB667C3E" box="[1131,1153,242,267]" italics="true" pageId="48" pageNumber="49">in</emphasis>
<bibRefCitation id="5B297ED8FFE68622DD92C2D9FAD57C39" author="Chu, Y. - T. &amp; Chan, C. - L. &amp; Chen, C. - T." box="[1159,1330,241,268]" pageId="48" pageNumber="49" refId="ref282947" refString="Chu, Y. - T., Chan, C. - L. &amp; Chen, C. - T. (1963) Ichthyofauna of East China Sea. Science Press, Beijing. xxviii + 642 pp. [In Chinese.]" type="book" year="1963">
Chu
<emphasis id="0DCCDF3BFFE68622DDAEC2DAFB147C3E" box="[1211,1267,241,267]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
1963
</bibRefCitation>
) described
<taxonomicName id="F8B878AAFFE68622DA3FC33BFC327C18" box="[810,981,275,301]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="abbreviatus">
<emphasis id="0DCCDF3BFFE68622DA3FC33BFC327C18" box="[810,981,275,301]" italics="true" pageId="48" pageNumber="49">C. abbreviatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F8B878AAFFE68622DD0EC33BFB207C18" box="[1051,1223,275,301]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="0DCCDF3BFFE68622DD0EC33BFB207C18" box="[1051,1223,275,301]" italics="true" pageId="48" pageNumber="49">C. intermedius</emphasis>
</taxonomicName>
from the northern part of the East
<collectingCountry id="47AF43B9FFE68622DAFDC31DFBCC7C7A" box="[1000,1067,309,335]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
Sea. Although
<bibRefCitation id="5B297ED8FFE68622DDF7C31DFA4C7C7A" author="Okamura, O." box="[1250,1451,309,335]" pageId="48" pageNumber="49" refId="ref297275" refString="Okamura, O. (1970 a) Fauna Japonica, Macrourina (Pisces). Academic Press of Japan, Tokyo, 216 pp., 64 pls." type="book" year="1970">Okamura (1970a)</bibRefCitation>
subsequently regarded these species synonymous,
<taxonomicName id="F8B878AAFFE68622DC87C37FFC4A7CA6" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="abbreviatus">
<emphasis id="0DCCDF3BFFE68622DC87C37FFC4A7CA6" italics="true" pageId="48" pageNumber="49">C. abbreviatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F8B878AAFFE68622DAFDC351FB687CA6" box="[1000,1167,377,403]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="0DCCDF3BFFE68622DAFDC351FB687CA6" box="[1000,1167,377,403]" italics="true" pageId="48" pageNumber="49">C. intermedius</emphasis>
</taxonomicName>
are considered as junior synonyms of
<taxonomicName id="F8B878AAFFE68622DAC3C3B3FA637C80" authority="Katayama, 1942" authorityName="Katayama" authorityYear="1942" box="[982,1412,411,437]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="multispinulosus">
<emphasis id="0DCCDF3BFFE68622DAC3C3B3FB567C80" box="[982,1201,411,437]" italics="true" pageId="48" pageNumber="49">C. multispinulosus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DDD7C3B3FA637C80" author="Katayama, M." box="[1218,1412,411,437]" pageId="48" pageNumber="49" pagination="332 - 334" refId="ref291145" refString="Katayama, M. (1942) A new macrouroid fish from the Japan Sea. Dobutsugaku Zasshi [Zoological Magazine, Japan], 54, 332 - 334." type="journal article" year="1942">
Katayama,
<quantity id="F840AECCFFE68622DC59C3B3FA637C80" box="[1356,1412,411,437]" metricMagnitude="1" metricUnit="m" metricValue="4.93268" pageId="48" pageNumber="49" unit="in" value="1942.0">1942</quantity>
</bibRefCitation>
</taxonomicName>
in this study (see the Nomenclatural discussion of
<taxonomicName id="F8B878AAFFE68622DC87C395FC3D7CCC" authorityName="Katayama" authorityYear="1942" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="multispinulosus">
<emphasis id="0DCCDF3BFFE68622DC87C395FC3D7CCC" italics="true" pageId="48" pageNumber="49">C. multispinulosus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3F070329FFE68622DA4AC029FBD77A7F" blockId="48.[810,1452,173,2070]" pageId="48" pageNumber="49">
<emphasis id="0DCCDF3BFFE68622DA4AC029FBEF7F2E" bold="true" box="[863,1032,513,539]" pageId="48" pageNumber="49">Relationships.</emphasis>
<taxonomicName id="F8B878AAFFE68622DD0CC029FAA37F2E" authorityName=": Okamura" authorityYear="1963" box="[1049,1348,513,539]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE68622DD0CC029FAA37F2E" box="[1049,1348,513,539]" italics="true" pageId="48" pageNumber="49">Coelorinchus formosanus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="F8B878AAFFE68622DA6DC00BFBF37F08" box="[888,1044,547,573]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE68622DA6DC00BFBF37F08" box="[888,1044,547,573]" italics="true" pageId="48" pageNumber="49">C. argentatus</emphasis>
</taxonomicName>
group as defined by
<bibRefCitation id="5B297ED8FFE68622DC01C00BFC797F6A" author="Nakayama, N. &amp; Prokofiev, A. M. &amp; Kawai, T." pageId="48" pageNumber="49" refId="ref296469" refString="Nakayama, N., Prokofiev, A. M. &amp; Kawai, T. (2020) Coelorinchus posteromaculatus (Actinopterygii, Gadiformes, Macrouridae), a new species of grenadier from the eastern Indian Ocean. Ichthyological Research, 8 pp. https: // doi. org / 10.1007 / s 10228 - 020 - 00741 - x" type="book" year="2020">
Nakayama
<emphasis id="0DCCDF3BFFE68622DC82C00CFCA07F6A" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
(2020)
</bibRefCitation>
, which includes the following 13 species:
<taxonomicName id="F8B878AAFFE68622DC87C06DFC467FB4" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE68622DC87C06DFC467FB4" italics="true" pageId="48" pageNumber="49">C. argentatus</emphasis>
</taxonomicName>
Smith &amp; Radcliffe
<emphasis id="0DCCDF3BFFE68622DD86C040FB4E7FB4" box="[1171,1193,616,641]" italics="true" pageId="48" pageNumber="49">in</emphasis>
<bibRefCitation id="5B297ED8FFE68622DDA0C04FFA8F7FB4" author="Radcliffe, L." box="[1205,1384,615,641]" pageId="48" pageNumber="49" pagination="105 - 140" refId="ref298180" refString="Radcliffe, L. (1912) Description of a new family, two new genera, and twenty-nine new species of anacanthine fishes from the Philippine Islands and contiguous waters. Proceedings of the United States National Museum, 43, 105 - 140, pls. 22 - 31. http: // dx. doi. org / 10.5479 / si. 00963801.43 - 1924.105" type="journal article" year="1912">Radcliffe, 1912</bibRefCitation>
from the
<collectingCountry id="47AF43B9FFE68622DA42C0A1FC3F7F96" box="[855,984,649,675]" name="Philippines" pageId="48" pageNumber="49">Philippines</collectingCountry>
,
<collectingCountry id="47AF43B9FFE68622DAF3C0A2FB7F7F96" box="[998,1176,649,676]" name="New Caledonia" pageId="48" pageNumber="49">New Caledonia</collectingCountry>
and northern
<collectingCountry id="47AF43B9FFE68622DC28C0A1FA407F96" box="[1341,1447,649,675]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
;
<taxonomicName id="F8B878AAFFE68622DA3FC083FB9B7FF0" authority="Regan, 1921" authorityName="Regan" authorityYear="1921" box="[810,1148,683,710]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="denticulatus">
<emphasis id="0DCCDF3BFFE68622DA3FC083FC3D7FF0" box="[810,986,683,709]" italics="true" pageId="48" pageNumber="49">C. denticulatus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DAFDC084FB9B7FF0" author="Regan, C. T." box="[1000,1148,683,710]" pageId="48" pageNumber="49" pagination="412 - 420" refId="ref298331" refString="Regan, C. T. (1921) New fishes from deep water off the coast of Natal. Annals and Magazine of Natural History, Series 9, 7, 412 - 420. http: // dx. doi. org / 10.1080 / 00222932108632540" type="journal article" year="1921">Regan, 1921</bibRefCitation>
</taxonomicName>
from the western Indian Ocean;
<taxonomicName id="F8B878AAFFE68622DA92C0E5FBD57FD2" authorityName=": Okamura" authorityYear="1963" box="[903,1074,717,743]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE68622DA92C0E5FBD57FD2" box="[903,1074,717,743]" italics="true" pageId="48" pageNumber="49">C. formosanus</emphasis>
</taxonomicName>
;
<taxonomicName id="F8B878AAFFE68622DD52C0E5FC3C7E3C" authority="Iwamoto &amp; Williams, 1999" authorityName="Iwamoto &amp; Williams" authorityYear="1999" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="gaesorhynchus">
<emphasis id="0DCCDF3BFFE68622DD52C0E5FAF17FD2" box="[1095,1302,717,743]" italics="true" pageId="48" pageNumber="49">C. gaesorhynchus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DC31C0E6FC3C7E3C" author="Iwamoto, T. &amp; Williams, A." pageId="48" pageNumber="49" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams, 1999</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="FD7CCDCBFFE68622DD3FC0D8FB1C7E3C" box="[1066,1275,751,778]" country="Australia" name="Western Australia" pageId="48" pageNumber="49">Western Australia</collectingRegion>
;
<taxonomicName id="F8B878AAFFE68622DC19C0C7FC0B7E1E" authority="Matsubara, 1943" authorityName="Matsubara" authorityYear="1943" class="Actinopterygii" family="Macrouridae" genus="Nezumia" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="kamoharai">
<emphasis id="0DCCDF3BFFE68622DC19C0C7FA4C7E3C" box="[1292,1451,751,777]" italics="true" pageId="48" pageNumber="49">C. kamoharai</emphasis>
Matsubara, 1943
</taxonomicName>
and
<taxonomicName id="F8B878AAFFE68622DD25C139FA4C7E1E" authority="Matsubara, 1943" authorityName="Matsubara" authorityYear="1943" box="[1072,1451,785,811]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="0DCCDF3BFFE68622DD25C139FB3C7E1E" box="[1072,1243,785,811]" italics="true" pageId="48" pageNumber="49">C. longissimus</emphasis>
Matsubara, 1943
</taxonomicName>
from
<collectingCountry id="47AF43B9FFE68622DA7FC11CFC4D7E7B" box="[874,938,820,846]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
to
<collectingCountry id="47AF43B9FFE68622DAC7C11BFBCF7E78" box="[978,1064,819,845]" name="Taiwan" pageId="48" pageNumber="49">Taiwan</collectingCountry>
;
<taxonomicName id="F8B878AAFFE68622DD20C11CFC857E5A" authority="Iwamoto &amp; Williams, 1999" authorityName="Iwamoto &amp; Williams" authorityYear="1999" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mayiae">
<emphasis id="0DCCDF3BFFE68622DD20C11CFB4E7E78" box="[1077,1193,820,846]" italics="true" pageId="48" pageNumber="49">C. mayiae</emphasis>
<bibRefCitation id="5B297ED8FFE68622DDA6C11CFC857E5A" author="Iwamoto, T. &amp; Williams, A." pageId="48" pageNumber="49" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams, 1999</bibRefCitation>
</taxonomicName>
from the Arafura Sea;
<taxonomicName id="F8B878AAFFE68622DD98C17EFA817E5A" authorityName="Katayama" authorityYear="1942" box="[1165,1382,853,880]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="multispinulosus">
<emphasis id="0DCCDF3BFFE68622DD98C17EFA817E5A" box="[1165,1382,853,880]" italics="true" pageId="48" pageNumber="49">C. multispinulosus</emphasis>
</taxonomicName>
from
<collectingCountry id="47AF43B9FFE68622DA3FC150FC8D7EA7" box="[810,874,888,914]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
to
<collectingCountry id="47AF43B9FFE68622DA9AC15FFC167EA4" box="[911,1009,887,913]" name="Vietnam" pageId="48" pageNumber="49">Vietnam</collectingCountry>
;
<taxonomicName id="F8B878AAFFE68622DAEBC150FA4C7EA4" authority="Iwamoto &amp; Williams, 1999" authorityName="Iwamoto &amp; Williams" authorityYear="1999" box="[1022,1451,887,914]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="pardus">
<emphasis id="0DCCDF3BFFE68622DAEBC150FB8A7EA4" box="[1022,1133,887,914]" italics="true" pageId="48" pageNumber="49">C. pardus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DD60C150FA4C7EA4" author="Iwamoto, T. &amp; Williams, A." box="[1141,1451,887,914]" pageId="48" pageNumber="49" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams, 1999</bibRefCitation>
</taxonomicName>
from the Arafura Sea;
<taxonomicName id="F8B878AAFFE68622DD23C1B2FB887EE0" authority="Nakayama, Prokofiev &amp; Kawai, 2020" authorityName="Nakayama, Prokofiev &amp; Kawai" authorityYear="2020" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="posteromaculatus">
<emphasis id="0DCCDF3BFFE68622DD23C1B2FAC67E86" box="[1078,1313,921,948]" italics="true" pageId="48" pageNumber="49">C. posteromaculatus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DC38C1B1FB887EE0" author="Nakayama, N. &amp; Prokofiev, A. M. &amp; Kawai, T." pageId="48" pageNumber="49" refId="ref296469" refString="Nakayama, N., Prokofiev, A. M. &amp; Kawai, T. (2020) Coelorinchus posteromaculatus (Actinopterygii, Gadiformes, Macrouridae), a new species of grenadier from the eastern Indian Ocean. Ichthyological Research, 8 pp. https: // doi. org / 10.1007 / s 10228 - 020 - 00741 - x" type="book" year="2020">Nakayama, Prokofiev &amp; Kawai, 2020</bibRefCitation>
</taxonomicName>
from the eastern Indian Ocean;
<taxonomicName id="F8B878AAFFE68622DA95C1F6FAA27EC2" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[896,1349,989,1016]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="quincunciatus">
<emphasis id="0DCCDF3BFFE68622DA95C1F6FBDA7EC2" box="[896,1085,990,1016]" italics="true" pageId="48" pageNumber="49">C. quincunciatus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DD51C1F5FAA27EC2" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[1092,1349,989,1015]" pageId="48" pageNumber="49" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="47AF43B9FFE68622DA3FC1D7FC4B792C" box="[810,940,1023,1049]" name="Philippines" pageId="48" pageNumber="49">Philippines</collectingCountry>
;
<taxonomicName id="F8B878AAFFE68622DAAEC628FA4C792C" authority="Iwamoto &amp; Merrett, 1997" authorityName="Iwamoto &amp; Merrett" authorityYear="1997" box="[955,1451,1023,1050]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="semaphoreus">
<emphasis id="0DCCDF3BFFE68622DAAEC628FB94792C" box="[955,1139,1023,1050]" italics="true" pageId="48" pageNumber="49">C. semaphoreus</emphasis>
<bibRefCitation id="5B297ED8FFE68622DD68C628FA4C792C" author="Iwamoto, T. &amp; Merrett, N. R." box="[1149,1451,1023,1050]" pageId="48" pageNumber="49" pagination="473 - 570" refId="ref288733" refString="Iwamoto, T. &amp; Merrett, N. R. (1997) Pisces Gadiformes: taxonomy of grenadiers of the New Caledonian region, southwest Pacific. Memoires du Museum National d'Histoire Naturelle, 176, 473 - 570." type="journal article" year="1997">Iwamoto &amp; Merrett, 1997</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="47AF43B9FFE68622DA7FC60AFBFD790E" box="[874,1050,1057,1084]" name="New Caledonia" pageId="48" pageNumber="49">New Caledonia</collectingCountry>
and
<collectingRegion id="FD7CCDCBFFE68622DD40C609FB3B790E" box="[1109,1244,1057,1083]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
,
<collectingCountry id="47AF43B9FFE68622DDFDC609FAB5790E" box="[1256,1362,1057,1083]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
; and
<taxonomicName id="F8B878AAFFE68622DC87C60AFB567968" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="0DCCDF3BFFE68622DC87C60AFC787968" italics="true" pageId="48" pageNumber="49">C. thompsoni</emphasis>
<bibRefCitation id="5B297ED8FFE68622DABDC66BFB567968" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[936,1201,1091,1117]" pageId="48" pageNumber="49" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="47AF43B9FFE68622DC33C66BFA4F7968" box="[1318,1448,1091,1117]" name="Philippines" pageId="48" pageNumber="49">Philippines</collectingCountry>
. These species are distinguished from other congeners by having the following combination of features (
<bibRefCitation id="5B297ED8FFE68622DC27C6AFFC4479F6" author="Nakayama, N. &amp; Prokofiev, A. M. &amp; Kawai, T." pageId="48" pageNumber="49" refId="ref296469" refString="Nakayama, N., Prokofiev, A. M. &amp; Kawai, T. (2020) Coelorinchus posteromaculatus (Actinopterygii, Gadiformes, Macrouridae), a new species of grenadier from the eastern Indian Ocean. Ichthyological Research, 8 pp. https: // doi. org / 10.1007 / s 10228 - 020 - 00741 - x" type="book" year="2020">
Nakayama
<emphasis id="0DCCDF3BFFE68622DA3FC682FC8779F6" box="[810,864,1193,1219]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2020
</bibRefCitation>
): light organ long, extending from anus to chest just posterior to isthmus; no interspace between anus and anal-fin origin; lateral nasal ridge incompletely supported by nasal bone; premaxillary teeth in long tapered band, with posterior margin of tooth band almost reaching lateral corner of mouth; body scales covered with short, erect spinules in quincunx order or subparallel to slightly divergent rows; modified scales on head ridges not prominently spinulated; dorsal surfaces of snout broadly naked along each side of median rostral ridge; second dorsal fin poorly developed, its height distinctly lower than that of anal fin; dark prominent markings, such as spots, blotches, stripes, and vermiculations, usually present dorsally on trunk and tail; lower half of body overlain with silvery reflections (often indistinct in preserved specimens). The above definition generally agrees with the original concept of the subgenus
<taxonomicName id="F8B878AAFFE68622DDC4C4E2FC057A33" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" family="Macrouridae" genus="Caelorinchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="subGenus" subGenus="Quincuncia">
<emphasis id="0DCCDF3BFFE68622DDC4C4E2FAB37BD1" box="[1233,1364,1738,1764]" italics="true" pageId="48" pageNumber="49">Quincuncia</emphasis>
<bibRefCitation id="5B297ED8FFE68622DC49C4E1FC057A33" author="Gilbert, C. H. &amp; Hubbs, C. L." pageId="48" pageNumber="49" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</taxonomicName>
that initially comprised
<taxonomicName id="F8B878AAFFE68622DC1BC4C4FA4B7A30" box="[1294,1452,1772,1798]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE68622DC1BC4C4FA4B7A30" box="[1294,1452,1772,1798]" italics="true" pageId="48" pageNumber="49">C. argentatus</emphasis>
</taxonomicName>
(
<typeStatus id="E003BD8BFFE68622DA24C526FC847A1D" box="[817,867,1806,1832]" pageId="48" pageNumber="49">type</typeStatus>
species),
<taxonomicName id="F8B878AAFFE68622DACFC526FB7B7A12" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[986,1180,1806,1832]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="quincunciatus">
<emphasis id="0DCCDF3BFFE68622DACFC526FB7B7A12" box="[986,1180,1806,1832]" italics="true" pageId="48" pageNumber="49">C. quincunciatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F8B878AAFFE68622DDCFC526FA937A12" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[1242,1396,1805,1832]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="0DCCDF3BFFE68622DDCFC526FA937A12" box="[1242,1396,1805,1832]" italics="true" pageId="48" pageNumber="49">C. thompsoni</emphasis>
</taxonomicName>
(see
<bibRefCitation id="5B297ED8FFE68622DA3FC507FBC37A7F" author="Nakayama, N. &amp; Prokofiev, A. M. &amp; Kawai, T." box="[810,1060,1839,1866]" pageId="48" pageNumber="49" refId="ref296469" refString="Nakayama, N., Prokofiev, A. M. &amp; Kawai, T. (2020) Coelorinchus posteromaculatus (Actinopterygii, Gadiformes, Macrouridae), a new species of grenadier from the eastern Indian Ocean. Ichthyological Research, 8 pp. https: // doi. org / 10.1007 / s 10228 - 020 - 00741 - x" type="book" year="2020">
Nakayama
<emphasis id="0DCCDF3BFFE68622DABDC518FC067A7C" box="[936,993,1839,1865]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph id="3F070329FFE68623DA4AC57AFDFE7A7C" blockId="48.[810,1452,173,2070]" lastBlockId="49.[136,778,1635,2070]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<bibRefCitation id="5B297ED8FFE68622DA4AC57AFB0C7A59" author="McCann, C. &amp; McKnight, D. G." box="[863,1259,1874,1900]" pageId="48" pageNumber="49" pagination="1 - 91" refId="ref293949" refString="McCann, C. &amp; McKnight, D. G. (1980) The marine fauna of New Zealand: macrourid fishes (Pisces: Gadida). New Zealand Oceanographic Institute Memoir, 61, 1 - 91." type="journal article" year="1980">McCann &amp; McKnight (1980)</bibRefCitation>
described
<taxonomicName id="F8B878AAFFE68622DC87C57AFC597ABB" authorityName="McCann &amp; McKnight" authorityYear="1980" family="Macrouridae" genus="Caelorinchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="campbellicus">
<emphasis id="0DCCDF3BFFE68622DC87C57AFC597ABB" italics="true" pageId="48" pageNumber="49">C. campbellicus</emphasis>
</taxonomicName>
from the southeast of
<collectingCountry id="47AF43B9FFE68622DC15C55CFA407ABB" box="[1280,1447,1908,1934]" name="New Zealand" pageId="48" pageNumber="49">New Zealand</collectingCountry>
, classifying the species under
<taxonomicName id="F8B878AAFFE68622DD92C5BEFAED7A85" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[1159,1290,1942,1968]" family="Macrouridae" genus="Caelorinchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="subGenus" subGenus="Quincuncia">
<emphasis id="0DCCDF3BFFE68622DD92C5BEFAED7A85" box="[1159,1290,1942,1968]" italics="true" pageId="48" pageNumber="49">Quincuncia</emphasis>
</taxonomicName>
. This species is excluded from the
<taxonomicName id="F8B878AAFFE68622DD08C590FB517AE4" box="[1053,1206,1976,2002]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE68622DD08C590FB517AE4" box="[1053,1206,1976,2002]" italics="true" pageId="48" pageNumber="49">C. argentatus</emphasis>
</taxonomicName>
group, because of its short light organ [43% HL (fide
<bibRefCitation id="5B297ED8FFE68622DDB9C5F2FC617523" author="McCann, C. &amp; McKnight, D. G." pageId="48" pageNumber="49" pagination="1 - 91" refId="ref293949" refString="McCann, C. &amp; McKnight, D. G. (1980) The marine fauna of New Zealand: macrourid fishes (Pisces: Gadida). New Zealand Oceanographic Institute Memoir, 61, 1 - 91." type="journal article" year="1980">McCann &amp; McKnight 1980:57</bibRefCitation>
) vs. about 70% in species of the
<taxonomicName id="F8B878AAFFE68622DC04C5D4FA4B7520" box="[1297,1452,2044,2070]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE68622DC04C5D4FA4B7520" box="[1297,1452,2044,2070]" italics="true" pageId="48" pageNumber="49">C. argentatus</emphasis>
</taxonomicName>
group].
<bibRefCitation id="5B297ED8FFE78623D9F1C44CFDBB7B4B" author="McCann, C. &amp; McKnight, D. G." box="[228,604,1635,1662]" pageId="49" pageNumber="50" pagination="1 - 91" refId="ref293949" refString="McCann, C. &amp; McKnight, D. G. (1980) The marine fauna of New Zealand: macrourid fishes (Pisces: Gadida). New Zealand Oceanographic Institute Memoir, 61, 1 - 91." type="journal article" year="1980">McCann &amp; McKnight (1980:57)</bibRefCitation>
also described
<taxonomicName id="F8B878AAFFE78623D99DC4AEFED87BAA" authorityName="McCann &amp; McKnight" authorityYear="1980" box="[136,319,1669,1696]" family="Macrouridae" genus="Caelorinchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="campbellicus">
<emphasis id="0DCCDF3BFFE78623D99DC4AEFED87BAA" box="[136,319,1669,1696]" italics="true" pageId="49" pageNumber="50">C. campbellicus</emphasis>
</taxonomicName>
as having “anterior photophore covered by scales, posterior ovoid, naked”, but the light organ of the
<taxonomicName id="F8B878AAFFE78623D9A4C4E2FEAE7BD6" box="[177,329,1738,1764]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE78623D9A4C4E2FEAE7BD6" box="[177,329,1738,1764]" italics="true" pageId="49" pageNumber="50">C. argentatus</emphasis>
</taxonomicName>
group is completely covered with sales. The species seems to be closely related to
<taxonomicName id="F8B878AAFFE78623DB65C4C4FE737A1D" authority="Arai &amp; Iwamoto, 1979" authorityName="Arai &amp; Iwamoto" authorityYear="1979" family="Macrouridae" genus="Caelorinchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="kaiyomaru">
<emphasis id="0DCCDF3BFFE78623DB65C4C4FCED7A30" box="[624,778,1771,1798]" italics="true" pageId="49" pageNumber="50">C. kaiyomaru</emphasis>
<bibRefCitation id="5B297ED8FFE78623D99DC525FE737A1D" author="Arai, T. &amp; Iwamoto, T." box="[136,404,1805,1832]" pageId="49" pageNumber="50" pagination="238 - 246" refId="ref281768" refString="Arai, T. &amp; Iwamoto, T. (1979) A new species of the macrourid fish genus Coelorinchus from off Tasmania, New Zealand, and the Falkland Islands. Japanese Journal of Ichthyology, 26, 238 - 246." type="journal article" year="1979">Arai &amp; Iwamoto, 1979</bibRefCitation>
</taxonomicName>
, widely distributed in temperate waters in the Southern Hemisphere.
</paragraph>
<caption id="6BC753A1FFE78623D99DC13BFA8C7E78" ID-DOI="http://doi.org/10.5281/zenodo.6422852" ID-Zenodo-Dep="6422852" httpUri="https://zenodo.org/record/6422852/files/figure.png" pageId="49" pageNumber="50" startId="49.[136,235,787,811]" targetBox="[151,1436,159,744]" targetPageId="49">
<paragraph id="3F070329FFE78623D99DC13BFA8C7E78" blockId="49.[136,1452,787,845]" pageId="49" pageNumber="50">
<emphasis id="0DCCDF3BFFE78623D99DC13BFEF67E1E" bold="true" box="[136,273,787,811]" pageId="49" pageNumber="50">FIGURE 32.</emphasis>
Fresh specimen of
<taxonomicName id="F8B878AAFFE78623D8C9C13BFD057E1E" authorityName=": Okamura" authorityYear="1963" box="[476,738,787,811]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623D8C9C13BFD057E1E" box="[476,738,787,811]" italics="true" pageId="49" pageNumber="50">Coelorinchus formosanus</emphasis>
</taxonomicName>
. FAKU 204426, 367+ mm TL, Dong-gang fish market, southwestern Taiwan, South China Sea. (A) Lateral view; (B) dorsal and (C) ventral views of the head and trunk. [Photos: N. Nakayama]
</paragraph>
</caption>
<caption id="6BC753A1FFE78623D99DC793FDA97B22" ID-DOI="http://doi.org/10.5281/zenodo.6422856" ID-Zenodo-Dep="6422856" httpUri="https://zenodo.org/record/6422856/files/figure.png" pageId="49" pageNumber="50" startId="49.[136,235,1467,1491]" targetBox="[151,1436,907,1430]" targetPageId="49">
<paragraph id="3F070329FFE78623D99DC793FDA97B22" blockId="49.[136,1452,1467,1559]" pageId="49" pageNumber="50">
<emphasis id="0DCCDF3BFFE78623D99DC793FEF578E6" bold="true" box="[136,274,1467,1491]" pageId="49" pageNumber="50">FIGURE 33.</emphasis>
Holotype of
<taxonomicName id="F8B878AAFFE78623D88EC793FD5678E6" authorityName=": Okamura" authorityYear="1963" box="[411,689,1467,1491]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623D88EC793FD5678E6" box="[411,689,1467,1491]" italics="true" pageId="49" pageNumber="50">Coelorhynchus formosanus</emphasis>
</taxonomicName>
(=
<taxonomicName id="F8B878AAFFE78623DBC2C793FC3A78E6" authorityName=": Okamura" authorityYear="1963" box="[727,989,1467,1491]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623DBC2C793FC3A78E6" box="[727,989,1467,1491]" italics="true" pageId="49" pageNumber="50">Coelorinchus formosanus</emphasis>
</taxonomicName>
). FAKU 35856, 72.8 mm HL, 221+ mm TL, Da-xi fish market, northeastern Taiwan, East China Sea. (A) Lateral view; (B) dorsal and (C) ventral views of the head and trunk. Preserved condition. [Photos: N. Nakayama]
</paragraph>
</caption>
<paragraph id="3F070329FFE78623D9A8C579FA407BF7" blockId="49.[136,778,1635,2070]" lastBlockId="49.[810,1452,1635,2070]" pageId="49" pageNumber="50">
<emphasis id="0DCCDF3BFFE78623D9A8C579FE847A5E" bold="true" box="[189,355,1873,1899]" pageId="49" pageNumber="50">Comparisons.</emphasis>
<taxonomicName id="F8B878AAFFE78623D899C57AFD287A59" authorityName=": Okamura" authorityYear="1963" box="[396,719,1874,1900]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623D899C57AFD287A59" box="[396,719,1874,1900]" italics="true" pageId="49" pageNumber="50">Coelorinchus formosanus</emphasis>
</taxonomicName>
is distinctive among the
<taxonomicName id="F8B878AAFFE78623D886C55CFDD67AB8" box="[403,561,1908,1934]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE78623D886C55CFDD67AB8" box="[403,561,1908,1934]" italics="true" pageId="49" pageNumber="50">C. argentatus</emphasis>
</taxonomicName>
group in that the underside of the head is heavily scaled posteriorly, but almost completely naked anteriorly (vs. either almost entirely scaled or naked). It further differs from all except large specimens of
<taxonomicName id="F8B878AAFFE78623D87FC5D4FDE47520" box="[362,515,2044,2070]" class="Actinopterygii" family="Macrouridae" genus="Coelorhynchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="0DCCDF3BFFE78623D87FC5D4FDE47520" box="[362,515,2044,2070]" italics="true" pageId="49" pageNumber="50">C. argentatus</emphasis>
</taxonomicName>
in having body scales covered with short, spike-like, trihedral spinules with welldeveloped buttresses (
<figureCitation id="A7831FACFFE78623DD0AC4ADFB657B95" box="[1055,1154,1669,1696]" captionStart="FIGURE 34" captionStartId="50.[136,235,1229,1253]" captionTargetBox="[151,1436,159,1189]" captionTargetId="figure-279@50.[151,1436,159,1189]" captionTargetPageId="50" captionText="FIGURE 34. Scanning electron micrographs showing body scales (from the dorsum below the interdorsal space) of four species of the Coelorinchus argentatus group. (A) C. formosanus, BSKU 99863, ca. 72 mm HL; (B) C. kamoharai, BSKU 93575, 59.6 mm HL; (C) C. longissimus, BSKU 40448, 95.9 mm HL; (D) C. multispinulosus, BSKU 99273, 69.9 mm HL. Views from above. [Photos: N. Nakayama]" figureDoi="http://doi.org/10.5281/zenodo.6422858" httpUri="https://zenodo.org/record/6422858/files/figure.png" pageId="49" pageNumber="50">Fig. 34A</figureCitation>
vs. spinules needle-like or triangular, and buttresses poorly developed,
<figureCitation id="A7831FACFFE78623DC06C48FFA847BF7" box="[1299,1379,1703,1730]" captionStart="FIGURE 34" captionStartId="50.[136,235,1229,1253]" captionTargetBox="[151,1436,159,1189]" captionTargetId="figure-279@50.[151,1436,159,1189]" captionTargetPageId="50" captionText="FIGURE 34. Scanning electron micrographs showing body scales (from the dorsum below the interdorsal space) of four species of the Coelorinchus argentatus group. (A) C. formosanus, BSKU 99863, ca. 72 mm HL; (B) C. kamoharai, BSKU 93575, 59.6 mm HL; (C) C. longissimus, BSKU 40448, 95.9 mm HL; (D) C. multispinulosus, BSKU 99273, 69.9 mm HL. Views from above. [Photos: N. Nakayama]" figureDoi="http://doi.org/10.5281/zenodo.6422858" httpUri="https://zenodo.org/record/6422858/files/figure.png" pageId="49" pageNumber="50">Fig. 34</figureCitation>
B–D).
</paragraph>
<paragraph id="3F070329FFE78620DA4AC4E2FE187B2D" blockId="49.[810,1452,1635,2070]" lastBlockId="50.[136,778,1431,1560]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
In Japanese waters,
<taxonomicName id="F8B878AAFFE78623DD2CC4E2FB3C7BD1" authorityName=": Okamura" authorityYear="1963" box="[1081,1243,1738,1764]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623DD2CC4E2FB3C7BD1" box="[1081,1243,1738,1764]" italics="true" pageId="49" pageNumber="50">C. formosanus</emphasis>
</taxonomicName>
might be confused with
<taxonomicName id="F8B878AAFFE78623DA70C4C4FBEB7A30" authorityName="Matsubara" authorityYear="1943" box="[869,1036,1771,1798]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="0DCCDF3BFFE78623DA70C4C4FBEB7A30" box="[869,1036,1771,1798]" italics="true" pageId="49" pageNumber="50">C. longissimus</emphasis>
</taxonomicName>
due to their similar appearance and coloration. In addition to the above-mentioned features,
<taxonomicName id="F8B878AAFFE78623DC87C526FC4A7A7F" authorityName=": Okamura" authorityYear="1963" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE78623DC87C526FC4A7A7F" italics="true" pageId="49" pageNumber="50">C. formosanus</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="F8B878AAFFE78623DC10C518FA4C7A7C" authorityName="Matsubara" authorityYear="1943" box="[1285,1451,1839,1866]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="0DCCDF3BFFE78623DC10C518FA4C7A7C" box="[1285,1451,1839,1866]" italics="true" pageId="49" pageNumber="50">C. longissimus</emphasis>
</taxonomicName>
by having a longer barbel (20–26% PRL vs. 11–18%) and pectoral fin (69–75% PRL vs. 55–64%), higher first dorsal fin (74–85% PRL vs. 58–70%), and body-scale spinules arranged in more or less quincunx order (vs. subparallel narrowly divergent rows). Large individuals of
<taxonomicName id="F8B878AAFFE78623DA58C5D4FC107523" authorityName="Matsubara" authorityYear="1943" box="[845,1015,2044,2070]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="0DCCDF3BFFE78623DA58C5D4FC107523" box="[845,1015,2044,2070]" italics="true" pageId="49" pageNumber="50">C. longissimus</emphasis>
</taxonomicName>
occasionally have rudimentary scaly patch(es) on the ventral surfaces of the preopercles, but the mandibular rami are always naked, except the posterior margins where a few scales are rarely present (vs. usually heavily scaled in
<taxonomicName id="F8B878AAFFE48620D85EC7D6FE087B2D" authorityName=": Okamura" authorityYear="1963" box="[331,495,1534,1560]" class="Actinopterygii" family="Macrouridae" genus="Coelorinchus" kingdom="Animalia" order="Gadiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="0DCCDF3BFFE48620D85EC7D6FE087B2D" box="[331,495,1534,1560]" italics="true" pageId="50" pageNumber="51">C. formosanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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