272 lines
27 KiB
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272 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="62033938359C06EFCA33FEE332841ABE" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole mantadia Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="E52BA14D-F9F4-429E-9A36-372006991B18" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777011" LSID="urn:lsid:zoobank.org:act:E52BA14D-F9F4-429E-9A36-372006991B18" httpUri="http://treatment.plazi.org/id/62033938359C06EFCA33FEE332841ABE" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/E52BA14D-F9F4-429E-9A36-372006991B18" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia" status="sp. nov.">Pheidole mantadia</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">Figs 55A-F</figureCitation>
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85G</figureCitation>
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<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina; Corridor Forestier Analamay-Mantadia, Tsaravoniana;
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<geoCoordinate degrees="18.76124" direction="south" orientation="latitude" precision="1" value="-18.76124">-18.76124</geoCoordinate>
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,
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<geoCoordinate degrees="48.42134" direction="east" orientation="longitude" precision="1" value="48.42134">48.42134</geoCoordinate>
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; alt. 939 m; 3 Dec 2012; B.L. Fisher et al. leg.; BLF30003, CASENT0297006 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1 w.; same data as for holotype; CASENT0923169 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full face view rectangular, slightly widening posteriorly; sides of the head with sparse, relatively short, erect pilosity; frons with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose; inner hypostomal teeth distinct, low, thick, bulge-like, with rounded apex; outer hypostomal teeth higher, thick, with rounded tips, triangular; inner and outer teeth closely spaced and connected by indistinct concavity; propodeal spines moderately long, with base slightly wider than top; first gastral tergite shagreened.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Whole head foveolate, frons and malar area with few additional, indistinct, longitudinal rugae, vertex with indistinct irregular rugae, genae with sculpture reduced to absent; promesonotum low, slightly convex, with steep posterior declivity; mesosoma foveolate, katepisternum, and mesonotum smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 1.68; HW: 1.52; SL: 0.67; EL: 0.17; WL: 1.21; PSL: 0.24; MTL: 0.68; PNW: 0.7; PTW: 0.2; PPW: 0.61; CI: 90.5; SI: 43.9; PSLI: 14.5; PPI: 33.9; PNI: 46.1; MTI: 44.7.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, slightly widening posteriorly, anterior of eyes relatively straight, posterior of eyes convex (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Antennal scrobes indistinct and not delimited by carinulae. Sides of the head with sparse, relatively short, erect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely rugulose; frons with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose; lateral sides of head with fine and sparse rugulae, area between rugulae finely foveolate; malar area with dense and thin longitudinal rugae, interspaces smooth; genae with fine, thin, and dense rugulae. Clypeus shiny and smooth, with thin, longitudinal rugulae on lateral sides; median notch present, narrow, and moderately deep; median longitudinal carina indistinct; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55B, D</figureCitation>
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). Inner hypostomal teeth distinct, low, thick, bulge-like, with rounded apex; outer hypostomal teeth higher, thick, with rounded tips, triangular (Fig.
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85G</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with small tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines moderately long, with base slightly wider than top, apex rounded; humeral area with small and flat tubercles (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D</figureCitation>
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). Surface shiny, finely foveolate, katepisternum, anepisternum, and lateral surfaces of propodeum smooth, with few irregular, thick rugae. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny, finely foveolate; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and narrow, with convex apex, in rear view node relatively straight; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny, with fine and sparse foveolae; short and rounded; in dorsal view sides with relatively long, acute, horn-like projections; pilosity long, moderately dense and erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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First gastral tergite shiny and shagreened; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, reddish brown to brown; legs yellow to brown (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923169) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0297006).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.58; HW: 0.52; SL: 0.6; EL: 0.11; WL: 0.7; PSL: 0.1; MTL: 0.48; PNW: 0.37; PTW: 0.07; PPW: 0. 1; CI: 89.2; SI: 115.7; PSLI: 17.0; PPI: 64.4; PNI: 71.4; MTI: 91.7.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55A</figureCitation>
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). Pilosity moderately sparse, long, and erect. Whole head foveolate, frons and malar area with few additional, indistinct, longitudinal rugae, vertex with indistinct irregular rugae, genae with sculpture reduced to absent. Clypeus shiny, foveolate to rugoreticulate; median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55A, C</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum low, slightly convex, with steep posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines short, triangular, apex acute (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C</figureCitation>
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). Sculpture foveolate, katepisternum and mesonotum smooth. Pilosity sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Peduncle moderately long and thin; node low, globular, and small; with few long, erect setae (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Short, low, and convex; with few short, erect setae (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, yellowish brown to brown (Fig.
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|
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
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<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species was collected at 939 m in elevation, in rainforest. Nest was located in rotten log.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
|
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</taxonomicName>
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sp. nov. is most similar to
|
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<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
|
|
</taxonomicName>
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sp. nov.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. curvistriat" pageId="0" pageNumber="1" rank="species" species="curvistriat">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriat</emphasis>
|
|
</taxonomicName>
|
|
a sp. nov. by presence of fine foveolate sculpture on frons and mesosoma, low and thick inner hypostomal teeth, which are wider than high and thick, high outer hypostomal teeth; from
|
|
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by sparser and shorter pilosity on sides of head.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by frons and malar with indistinct, longitudinal rugae, and vertex with indistinct irregular and never arcuate rugae, absence of irregular rugae on mesosoma, and smooth mesonotum; from
|
|
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by frons and malar area with indistinct, longitudinal rugae, and never smooth clypeus.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |