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<document id="8CF83A43068271B0B2013B2B13AC6BD1" ID-DOI="10.5281/zenodo.5319032" ID-ISSN="0374-1036" ID-Zenodo-Dep="5319032" ID-ZooBank="32572CA2-D2CB-4A01-B252-CCECB5320114" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630108184603" checkinUser="marcus" docAuthor="Yin, Zi-Wei, Nomura, Shûhei &amp; Li, Li-Zhen" docDate="2015" docId="022DE650D938FFA106F6FB36FBDBFE21" docLanguage="en" docName="ActaEntMusNatPra.2015.55.1.105-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Tribasodites bedosae Yin &amp; Li 2011" docType="treatment" docVersion="4" lastPageNumber="108" masterDocId="FE149E28D93AFFA2074AFFACFFF5FF92" masterDocTitle="Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera: Staphylinidae)" masterLastPageNumber="127" masterPageNumber="105" pageNumber="107" updateTime="1699154406698" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="04AAD9863CBF13C5CCA526E66C0401A6">Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="9CFF4F3BDF9956ACA10D9B1D6B9E61DB">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="40CC5139F46D0FE3D1AF74F73AC58852">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
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<mods:namePart id="117A75A5826199ECD8CC41C258F11712">Nomura, Shûhei</mods:namePart>
<mods:affiliation id="00D46760F0768D23722789C654805706">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan</mods:affiliation>
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<mods:namePart id="FEEE7B3F12DAEE206B1CB6DE4EBA7FE4">Li, Li-Zhen</mods:namePart>
<mods:affiliation id="9E601678703ED3AAAFA7EE1643F7351D">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China &amp; Corresponding author; e-mail: pselaphinae @ gmail. com</mods:affiliation>
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<mods:date id="FE5690C0B9C22B6D8EEE5B8456BDFE1D">2015</mods:date>
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<treatment id="022DE650D938FFA106F6FB36FBDBFE21" ID-DOI="http://doi.org/10.5281/zenodo.5332463" ID-Zenodo-Dep="5332463" LSID="urn:lsid:plazi:treatment:022DE650D938FFA106F6FB36FBDBFE21" httpUri="http://treatment.plazi.org/id/022DE650D938FFA106F6FB36FBDBFE21" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107">
<subSubSection id="C29E04CDD938FFA006F6FB36FCB7FB23" box="[444,834,1178,1201]" pageId="2" pageNumber="107" type="nomenclature">
<paragraph id="8A3B5746D938FFA006F6FB36FCB7FB23" blockId="2.[444,834,1178,1201]" box="[444,834,1178,1201]" pageId="2" pageNumber="107">
<heading id="D173E02AD938FFA006F6FB36FCB7FB23" bold="true" box="[444,834,1178,1201]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="107" reason="3">
<emphasis id="B8F08B54D938FFA006F6FB36FCB7FB23" bold="true" box="[444,834,1178,1201]" pageId="2" pageNumber="107">
<taxonomicName id="4D842CC5D938FFA006F6FB36FD55FB23" ID-CoL="5864C" authorityName="Yin &amp; Li" authorityYear="2011" box="[444,672,1178,1201]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B8F08B54D938FFA006F6FB36FD55FB23" bold="true" box="[444,672,1178,1201]" italics="true" pageId="2" pageNumber="107">Tribasodites bedosae</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD938FFA107DDFB65FEB4FE81" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107" type="diagnosis">
<paragraph id="8A3B5746D938FFA107DDFB65FEB4FE81" blockId="2.[151,1127,1225,1568]" lastBlockId="3.[113,1089,220,435]" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107">
<emphasis id="B8F08B54D938FFA007DDFB65FEF9FB72" bold="true" box="[151,268,1225,1248]" pageId="2" pageNumber="107">Diagnosis.</emphasis>
Cavernicolous
<taxonomicName id="4D842CC5D938FFA006F8FB65FDC3FB72" authorityName="Jeannel" authorityYear="1960" box="[434,566,1225,1248]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B8F08B54D938FFA006F8FB65FDC3FB72" box="[434,566,1225,1248]" italics="true" pageId="2" pageNumber="107">Tribasodites</emphasis>
</taxonomicName>
with slender habitus and elongate antennae and legs. Males of all, except
<taxonomicName id="4D842CC5D938FFA00621FB45FE0AFA92" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[363,511,1257,1280]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="abnormalis" status="sp. nov.">
<emphasis id="B8F08B54D938FFA00621FB45FE8FFA92" box="[363,378,1257,1280]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA006CCFB45FE0AFA92" box="[390,511,1257,1280]" italics="true" pageId="2" pageNumber="107">abnormalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA0054FFB45FDADFA92" box="[517,600,1257,1280]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
, have strongly modified antennomeres VIIIXI. A few species have strongly reduced eyes. Metaventrite modified, often with a pair of rows of dense setae anterior to metacoxae, or setae tufted instead of arranged in rows, or occasionally only sparsely covered.Aedeagus usually with median lobe split apically, with broad dorsal lobe, parameres attached to base of the median lobe, with distinct basoventral projection. Females lack any modification, and can be properly identified only if associated with the males, or sometimes by a combination of the forms of tergite VIII, sternite VIII, and genital complex.
<emphasis id="B8F08B54D938FFA007DDFA65FEA2FA72" bold="true" box="[151,343,1481,1504]" pageId="2" pageNumber="107">Included species.</emphasis>
Thirteen species from
<collectingCountry id="F29317D6D938FFA00503FA65FD7CFA72" box="[585,649,1481,1504]" name="China" pageId="2" pageNumber="107">China</collectingCountry>
and
<collectingCountry id="F29317D6D938FFA005F0FA65FCECFA72" box="[698,793,1481,1504]" name="Thailand" pageId="2" pageNumber="107">Thailand</collectingCountry>
are included:
<taxonomicName id="4D842CC5D938FFA004E4FA65FBB7FA72" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[942,1090,1481,1504]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="abnormalis" status="sp. nov.">
<emphasis id="B8F08B54D938FFA004E4FA65FC48FA72" box="[942,957,1481,1504]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA00483FA65FBB7FA72" box="[969,1090,1481,1504]" italics="true" pageId="2" pageNumber="107">abnormalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA00302FA65FF32F992" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA00799FA45FED9F992" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[211,300,1513,1536]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="bama" status="sp. nov.">
<emphasis id="B8F08B54D938FFA00799FA45FF17F992" box="[211,226,1513,1536]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA007BBFA45FED9F992" box="[241,300,1513,1536]" italics="true" pageId="2" pageNumber="107">bama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA0067EFA45FE7EF992" box="[308,395,1513,1536]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA006DCFA45FD49F992" authority="Yin &amp; Li, 2011" authorityName="Yin &amp; Li" authorityYear="2011" box="[406,700,1513,1536]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B8F08B54D938FFA006DCFA45FE50F992" box="[406,421,1513,1536]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA006FEFA45FDFEF992" box="[436,523,1513,1536]" italics="true" pageId="2" pageNumber="107">bedosae</emphasis>
<bibRefCitation id="EE152AB7D938FFA00559FA45FD49F992" author="YIN Z. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[531,700,1513,1536]" pageId="2" pageNumber="107" pagination="49 - 59" refId="ref9114" refString="YIN Z. - W., LI L. - Z. &amp; ZHAO M. - J. 2011 a: Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3065: 49 - 59." type="journal article" year="2011">Yin &amp; Li, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4D842CC5D938FFA00582FA45FC93F992" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[712,870,1513,1536]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="cehengensis" status="sp. nov.">
<emphasis id="B8F08B54D938FFA00582FA45FD22F992" box="[712,727,1513,1536]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA005AFFA45FC93F992" box="[741,870,1513,1536]" italics="true" pageId="2" pageNumber="107">cehengensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA00424FA45FC30F992" box="[878,965,1513,1536]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA0049BFA45FECBF9B2" authority="Yin &amp; Li, 2011" authorityName="Yin &amp; Li" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B8F08B54D938FFA0049BFA45FC15F992" box="[977,992,1513,1536]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA004A4FA45FB93F992" box="[1006,1126,1513,1536]" italics="true" pageId="2" pageNumber="107">deharvengi</emphasis>
<bibRefCitation id="EE152AB7D938FFA007DDF9A5FECBF9B2" author="YIN Z. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[151,318,1545,1568]" pageId="2" pageNumber="107" pagination="49 - 59" refId="ref9114" refString="YIN Z. - W., LI L. - Z. &amp; ZHAO M. - J. 2011 a: Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3065: 49 - 59." type="journal article" year="2011">Yin &amp; Li, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4D842CC5D938FFA00600F9A5FE2DF9B2" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[330,472,1545,1568]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="hubeiensis" status="sp. nov.">
<emphasis id="B8F08B54D938FFA00600F9A5FEACF9B2" box="[330,345,1545,1568]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA0062DF9A5FE2DF9B2" box="[359,472,1545,1568]" italics="true" pageId="2" pageNumber="107">hubeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA00695F9A5FDC0F9B2" box="[479,565,1545,1568]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA0050AF9A5FD42F9B2" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[576,695,1545,1568]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="kawadai" status="sp. nov.">
<emphasis id="B8F08B54D938FFA0050AF9A5FDBAF9B2" box="[576,591,1545,1568]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA00517F9A5FD42F9B2" box="[605,695,1545,1568]" italics="true" pageId="2" pageNumber="107">kawadai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA005F5F9A5FCE1F9B2" box="[703,788,1545,1568]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA0046AF9A5FC69F9B2" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[800,924,1545,1568]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="liboensis" status="sp. nov.">
<emphasis id="B8F08B54D938FFA0046AF9A5FCDAF9B2" box="[800,815,1545,1568]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA00477F9A5FC69F9B2" box="[829,924,1545,1568]" italics="true" pageId="2" pageNumber="107">liboensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD938FFA004E9F9A5FC0CF9B2" box="[931,1017,1545,1568]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D938FFA1034EF9A5FF4FFF61" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" lastPageId="3" lastPageNumber="108" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="setosiventris" status="sp. nov.">
<emphasis id="B8F08B54D938FFA0034EF9A5FBE6F9B2" box="[1028,1043,1545,1568]" italics="true" pageId="2" pageNumber="107">T</emphasis>
.
<emphasis id="B8F08B54D938FFA1036BF9A5FF4FFF61" italics="true" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107">setosiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD939FFA10788FF70FEEDFF61" box="[194,280,220,243]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D939FFA1066EFF70FE33FF61" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[292,454,220,243]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="thailandicus" status="sp. nov.">
<emphasis id="B8F08B54D939FFA1066EFF70FEC6FF61" box="[292,307,220,243]" italics="true" pageId="3" pageNumber="108">T</emphasis>
.
<emphasis id="B8F08B54D939FFA10608FF70FE33FF61" box="[322,454,220,243]" italics="true" pageId="3" pageNumber="108">thailandicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD939FFA10687FF70FDD1FF61" box="[461,548,220,243]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4D842CC5D939FFA10565FF70FCDBFF61" authority="Yin &amp; Li, 2011" authorityName="Yin &amp; Li" authorityYear="2011" box="[559,814,220,243]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="tiani">
<emphasis id="B8F08B54D939FFA10565FF70FDCBFF61" box="[559,574,220,243]" italics="true" pageId="3" pageNumber="108">T</emphasis>
.
<emphasis id="B8F08B54D939FFA10507FF70FD8BFF61" box="[589,638,220,243]" italics="true" pageId="3" pageNumber="108">tiani</emphasis>
<bibRefCitation id="EE152AB7D939FFA105CFFF70FCDBFF61" author="YIN Z. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[645,814,220,243]" pageId="3" pageNumber="108" pagination="49 - 59" refId="ref9114" refString="YIN Z. - W., LI L. - Z. &amp; ZHAO M. - J. 2011 a: Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3065: 49 - 59." type="journal article" year="2011">Yin &amp; Li, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4D842CC5D939FFA10470FF70FC67FF61" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[826,914,220,243]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="uenoi" status="sp. nov.">
<emphasis id="B8F08B54D939FFA10470FF70FCBCFF61" box="[826,841,220,243]" italics="true" pageId="3" pageNumber="108">T</emphasis>
.
<emphasis id="B8F08B54D939FFA1041DFF70FC67FF61" box="[855,914,220,243]" italics="true" pageId="3" pageNumber="108">uenoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD939FFA104D0FF70FC04FF61" box="[922,1009,220,243]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4D842CC5D939FFA10361FF70FF1DFE81" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="xingyiensis" status="sp. nov.">
<emphasis id="B8F08B54D939FFA10361FF70FBCFFF61" box="[1067,1082,220,243]" italics="true" pageId="3" pageNumber="108">T</emphasis>
.
<emphasis id="B8F08B54D939FFA1073BFF50FF1DFE81" box="[113,232,252,275]" italics="true" pageId="3" pageNumber="108">xingyiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD939FFA107A4FF50FEB4FE81" box="[238,321,252,275]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD939FFA1073BFEB0FBDBFE21" pageId="3" pageNumber="108" type="discussion">
<paragraph id="8A3B5746D939FFA1073BFEB0FBDBFE21" blockId="3.[113,1089,220,435]" pageId="3" pageNumber="108">
<emphasis id="B8F08B54D939FFA1073BFEB0FF06FEA1" bold="true" box="[113,243,284,307]" pageId="3" pageNumber="108">Comments.</emphasis>
Except for
<taxonomicName id="4D842CC5D939FFA10633FEB0FDE4FEA1" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[377,529,284,307]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="abnormalis" status="sp. nov.">
<emphasis id="B8F08B54D939FFA10633FEB0FE7DFEA1" box="[377,392,284,307]" italics="true" pageId="3" pageNumber="108">T</emphasis>
.
<emphasis id="B8F08B54D939FFA106D2FEB0FDE4FEA1" box="[408,529,284,307]" italics="true" pageId="3" pageNumber="108">abnormalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD939FFA10551FEB0FD86FEA1" box="[539,627,284,307]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
, which has simple antennae and modified vertex, all males of this group are morphologically similar to each other. They can be separated only based on a combination of the following features: (1) shape of the aedeagus, (2) presence/absence of the pronotal lateral spines, (3) setation on metaventrite, (4) shape of the antennomeres VIIIXI, and (5) their distribution. An identification key is provided below.
</paragraph>
</subSubSection>
</treatment>
</document>