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<mods:title id="AF7CCBE8549B7230351F5FDB4208CCE7">New or poorly known species of Sinodrepanus Simonis, 1985 (Coleoptera: Scarabaeidae: Oniticellini), with the first identification key for the genus</mods:title>
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<taxonomicName id="B7EC4D65FFFAFFF430CBFD53F075FCF9" authority="Montanaro" authorityName="Montanaro" authorityYear="2024" box="[151,615,762,789]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="schoenfeldi" status="sp. nov.">
<emphasis id="4298EAF4FFFAFFF430CBFD53F3C3FCF9" bold="true" box="[151,465,762,789]" italics="true" pageId="10" pageNumber="587">Sinodrepanus schoenfeldi</emphasis>
Montanaro
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,
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(
<figureCitation id="E8D72A63FFFAFFF430C3FCEAF305FCB1" box="[159,279,835,861]" captionStart="FIGURE 1" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[204,1386,182,1914]" captionTargetId="figure-23@6.[199,1388,179,1917]" captionTargetPageId="6" captionText="FIGURE 1. Habitus of Sinodrepanus species, dorsal view. S. exsul: male (a) and female (b). S. schoenfeldi new species: male paratype from Laos (c) and female paratype from India (d)." figureDoi="http://doi.org/10.5281/zenodo.14240162" httpUri="https://zenodo.org/record/14240162/files/figure.png" pageId="10" pageNumber="587">Figs. 1cd</figureCitation>
,
<figureCitation id="E8D72A63FFFAFFF4317FFCEAF392FCB1" box="[291,384,835,861]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[182,1401,185,1914]" captionTargetId="figure-20@7.[177,1410,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Genitalia of Sinodrepanus species. S. exsul: aedeagus in lateral (a), posterior (b) and dorsal (c) views; lamella copulatrix (d); female genitalia (i). S. schoenfeldi new species: aedeagus in lateral (e), posterior (f) and dorsal (g) views; lamella copulatrix (h); female genitalia (j)." figureDoi="http://doi.org/10.5281/zenodo.14240164" httpUri="https://zenodo.org/record/14240164/files/figure.png" pageId="10" pageNumber="587">2eh, 2j</figureCitation>
,
<figureCitation id="E8D72A63FFFAFFF431D7FCEAF3EAFCB1" box="[395,504,835,861]" captionStart="FIGURE 3" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[155,1433,182,1919]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Details of Sinodrepanus species. S. exsul: male (a) and female (c) head; posterior pronotal punctuation (e). S. schoenfeldi new species: male (b) and female (d) head; posterior pronotal punctuation (f). S. arrowi, lectotype (g) and its labels (h) (pictures by Jiří Hájek, NMPC)." figureDoi="http://doi.org/10.5281/zenodo.14240166" httpUri="https://zenodo.org/record/14240166/files/figure.png" pageId="10" pageNumber="587">3d, 3b, 3f</figureCitation>
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<taxonomicName id="B7EC4D65FFFAFFF430CBFC21F37BFC73" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[151,361,903,927]" class="Insecta" family="Scarabaeidae" genus="Drepanocerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="4298EAF4FFFAFFF430CBFC21F37BFC73" box="[151,361,903,927]" italics="true" pageId="10" pageNumber="587">Drepanocerus falsus</emphasis>
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:
<treatmentCitation id="F14D10F7FFFAFFF4312AFC21F024FC73" author="Janssens, A." box="[374,566,903,927]" page="36" pageId="10" pageNumber="587" year="1953">
<bibRefCitation id="147D4B17FFFAFFF4312AFC21F024FC73" author="Janssens, A." box="[374,566,903,927]" pageId="10" pageNumber="587" pagination="1 - 118" refId="ref14863" refString="Janssens, A. (1953) Exploration du Parc National de l'Upemba. Mission G. F. De Witte (1946 - 1949). Oniticellini (Coleoptera Lamellicornia). Institut des parcs nationaux du Congo Belge, 11, 1 - 118." type="journal article" year="1953">Janssens, 1953: 36</bibRefCitation>
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, 37;
<treatmentCitation id="F14D10F7FFFAFFF4323BFC2EF1B9FC73" author="Biswas, S. &amp; Chatterjee, S. K." box="[615,939,903,927]" page="151" pageId="10" pageNumber="587" year="1985">
<bibRefCitation id="147D4B17FFFAFFF4323BFC2EF1B9FC73" author="Biswas, S. &amp; Chatterjee, S. K." box="[615,939,903,927]" pageId="10" pageNumber="587" pagination="147 - 177" refId="ref14524" refString="Biswas, S. &amp; Chatterjee, S. K. (1985) Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, 84, 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee, 1985: 151</bibRefCitation>
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(?);
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<bibRefCitation id="147D4B17FFFAFFF43387FC2EF681FC73" author="Sewak, R." box="[987,1171,903,927]" pageId="10" pageNumber="587" pagination="249 - 287" refId="ref15684" refString="Sewak, R. (2003) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of India with special reference to Arunachal Pradesh, Uttar Pradesh, Gujarat and Rajasthan. In: Gupta, R. K. (Ed.), Advancements in Insect Biodiversity. Agrobios, Jodhpur, pp. 249 - 287." type="book chapter" year="2003">Sewak, 2003: 253</bibRefCitation>
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, 283 (?)
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<taxonomicName id="B7EC4D65FFFAFFF430CBFC0EF370FC52" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[151,354,934,958]" class="Insecta" family="Scarabaeidae" genus="Drepanocerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFAFFF430CBFC0EF370FC52" box="[151,354,934,958]" italics="true" pageId="10" pageNumber="587">Drepanocerus exsul</emphasis>
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:
<treatmentCitation id="F14D10F7FFFAFFF43132FC0FF031FC52" author="Arrow, G. J." box="[366,547,934,958]" page="381" pageId="10" pageNumber="587" year="1931">
<bibRefCitation id="147D4B17FFFAFFF43132FC0FF031FC52" author="Arrow, G. J." box="[366,547,934,958]" pageId="10" pageNumber="587" refId="ref13876" refString="Arrow, G. J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III. Coprinae. Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow, 1931: 381</bibRefCitation>
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, 385;
<treatmentCitation id="F14D10F7FFFAFFF4323DFC0FF105FC52" author="Sewak, R." box="[609,791,934,958]" page="253" pageId="10" pageNumber="587" year="2003">
<bibRefCitation id="147D4B17FFFAFFF4323DFC0FF105FC52" author="Sewak, R." box="[609,791,934,958]" pageId="10" pageNumber="587" pagination="249 - 287" refId="ref15684" refString="Sewak, R. (2003) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of India with special reference to Arunachal Pradesh, Uttar Pradesh, Gujarat and Rajasthan. In: Gupta, R. K. (Ed.), Advancements in Insect Biodiversity. Agrobios, Jodhpur, pp. 249 - 287." type="book chapter" year="2003">Sewak, 2003: 253</bibRefCitation>
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(?)
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<taxonomicName id="B7EC4D65FFFAFFF430CBFC6CF372FC30" authority="(Sharp, 1875)" box="[151,352,964,988]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFAFFF430CBFC6CF372FC30" box="[151,352,964,988]" italics="true" pageId="10" pageNumber="587">Drepenocerus exsul</emphasis>
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[
<emphasis id="4298EAF4FFFAFFF43133FC6CF399FC30" box="[367,395,965,988]" italics="true" pageId="10" pageNumber="587">sic</emphasis>
!]:
<treatmentCitation id="F14D10F7FFFAFFF431F5FC6DF072FC31" author="Sewak, R." box="[425,608,964,989]" page="283" pageId="10" pageNumber="587" year="2003">
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(?)
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<typeStatus id="AF578844FFFAFFF430CBFBA2F2C0FBC9" box="[151,210,1035,1061]" pageId="10" pageNumber="587">Type</typeStatus>
locality
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: Patkai Mountains, Assam state,
<collectingCountry id="08FB7676FFFAFFF432F8FBA2F0F3FBC9" box="[676,737,1035,1061]" name="India" pageId="10" pageNumber="587">India</collectingCountry>
.
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<paragraph id="705336E6FFFAFFF4309BFB86F6FFFA5D" blockId="10.[151,1437,1034,2033]" pageId="10" pageNumber="587">
<emphasis id="4298EAF4FFFAFFF4309BFB86F3F8FBA4" bold="true" box="[199,490,1070,1097]" pageId="10" pageNumber="587">Type material examined</emphasis>
.
<materialsCitation id="C0843CBBFFFAFFF431A6FB86F013FB59" collectionCode="BMNH, BMNH(E)" collectorName="Fry" determinerName="G. Montanaro" location="Patkai Mts." pageId="10" pageNumber="587" specimenCount="1" specimenCount-male="1" stateProvince="Assam" typeStatus="holotype">
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<typeStatus id="AF578844FFFAFFF431A6FB86F075FBA5" box="[506,615,1071,1097]" pageId="10" pageNumber="587" type="holotype">Holotype</typeStatus>
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,
<specimenCount id="66EAFD6FFFFAFFF4322AFB86F099FBA5" box="[630,651,1071,1097]" count="1" pageId="10" pageNumber="587" type="male"></specimenCount>
(
<collectionCode id="16FDAE23FFFAFFF432C7FB86F0E4FBA5" box="[667,758,1071,1097]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="10" pageNumber="587">BMNH</collectionCode>
):
<collectingRegion id="B228F804FFFAFFF43354FB86F146FBA5" box="[776,852,1071,1097]" country="India" name="Assam" pageId="10" pageNumber="587">Assam</collectingRegion>
/
<location id="7533603DFFFAFFF43330FB86F1FCFBA5" LSID="urn:lsid:plazi:treatment:F84587F0FFFAFFF530CBFD53F136F855:7533603DFFFAFFF43330FB86F1FCFBA5" box="[876,1006,1071,1097]" name="Patkai Mts." pageId="10" pageNumber="587" stateProvince="Assam">Patkai Mts.</location>
[wc, hb] // 61033 [wc, hb] // Doherty [wc, hb] //
<collectorName id="DD195330FFFAFFF4314DFBFAF32AFB81" box="[273,312,1107,1133]" pageId="10" pageNumber="587">Fry</collectorName>
Coll. / 1905-100. [wc, pb] // Determinated from / description G.J.A. /
<taxonomicName id="B7EC4D65FFFAFFF43416FBFAF6D7FB81" authorityName="Dejean" authorityYear="1821" box="[1098,1221,1107,1133]" class="Insecta" family="Scarabaeidae" genus="Oniticellus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="genus">Oniticellus</taxonomicName>
/
<taxonomicName id="B7EC4D65FFFAFFF43486FBFAF776FB81" authority=", Sharp" authorityName="Sharp" box="[1242,1380,1107,1133]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="exsul">exsul, Sharp</taxonomicName>
[wc, pb except last two lines hb] //
<collectionCode id="16FDAE23FFFAFFF431BBFBDEF04DFB7D" box="[487,607,1143,1169]" pageId="10" pageNumber="587">BMNH(E)</collectionCode>
/ 1237595 [wc, pb] //
<taxonomicName id="B7EC4D65FFFAFFF43304FBDEF1E3FB7D" authorityName="Simonis" authorityYear="1985" box="[856,1009,1143,1169]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFAFFF43304FBDEF1E3FB7D" box="[856,1009,1143,1169]" italics="true" pageId="10" pageNumber="587">Sinodrepanus</emphasis>
</taxonomicName>
/
<taxonomicName id="B7EC4D65FFFAFFF4345AFBDEF694FB7D" authorityName="Montanaro" authorityYear="2024" box="[1030,1158,1143,1169]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="schoenfeldi" status="sp. nov.">
<emphasis id="4298EAF4FFFAFFF4345AFBDEF694FB7D" box="[1030,1158,1143,1169]" italics="true" pageId="10" pageNumber="587">schoenfeldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="59AB578FFFFAFFF434D1FBDEF6DBFB7D" box="[1165,1225,1143,1169]" pageId="10" pageNumber="587" rank="species">n. sp.</taxonomicNameLabel>
/ HOLOTYPE ♂ /
<determinerName id="858FDA22FFFAFFF430CBFB32F327FB59" box="[151,309,1179,1205]" pageId="10" pageNumber="587">G. Montanaro</determinerName>
det. 2024 [rc, hb]
</materialsCitation>
.
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<typeStatus id="AF578844FFFAFFF43250FB33F063FB58" box="[524,625,1178,1204]" pageId="10" pageNumber="587" type="allotype">Allotype</typeStatus>
</emphasis>
: NE
<collectingCountry id="08FB7676FFFAFFF432F1FB32F0FBFB59" box="[685,745,1179,1205]" name="India" pageId="10" pageNumber="587">India</collectingCountry>
,
<collectingRegion id="B228F804FFFAFFF432AFFB32F1D9FB59" box="[755,971,1179,1205]" country="India" name="Arunachal Pradesh" pageId="10" pageNumber="587">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="9037AC9CFFFAFFF4338AFB32F666FB59" box="[982,1140,1179,1205]" pageId="10" pageNumber="587">Hunli vicinity</collectingMunicipality>
,
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<geoCoordinate id="15D85021FFFAFFF43553FB33F78AFB59" box="[1295,1432,1178,1205]" degrees="95" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="587" precision="15" seconds="31" value="95.95861">95°5731”E</geoCoordinate>
, 1300±
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,
<collectingMethod id="A9AD4EF1FFFAFFF43172FB16F066FB35" box="[302,628,1215,1241]" pageId="10" pageNumber="587">FIT (flight interception trap)</collectingMethod>
,
<date id="04521026FFFAFFF432DDFB16F13EFB35" box="[641,812,1215,1241]" pageId="10" pageNumber="587" value="2012-05-26" valueMax="2012-06-01" valueMin="2012-05-26">
<collectingDate id="1416E9CEFFFAFFF432DDFB16F13EFB35" box="[641,812,1215,1241]" pageId="10" pageNumber="587" value="2012-05-26" valueMax="2012-06-01" valueMin="2012-05-26">26.v1.vi.2012</collectingDate>
</date>
,
<collectorName id="DD195330FFFAFFF43365FB16F1DDFB35" box="[825,975,1215,1241]" pageId="10" pageNumber="587">L. Dembický</collectorName>
<emphasis id="4298EAF4FFFAFFF4338BFB16F61BFB35" box="[983,1033,1215,1241]" italics="true" pageId="10" pageNumber="587">legit</emphasis>
[
<collectionCode id="16FDAE23FFFAFFF43444FB16F662FB35" box="[1048,1136,1215,1241]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="587">NMPC</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="C0843CBBFFFAFFF434DFFB16F06DFACD" collectingDate="2012-06-02" collectingDateMax="2012-06-05" collectingDateMin="2012-06-02" collectionCode="JSCS" collectorName="L. Dembicky" country="India" elevation="500" latitude="28.142221" location="Roing" longLatPrecision="816" longitude="95.85" municipality="Roing" pageId="10" pageNumber="587" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="4298EAF4FFFAFFF434DFFB16F6E9FB35" bold="true" box="[1155,1275,1215,1241]" pageId="10" pageNumber="587">
<typeStatus id="AF578844FFFAFFF434DFFB16F6E9FB35" box="[1155,1275,1215,1241]" pageId="10" pageNumber="587" type="paratype">Paratypes</typeStatus>
</emphasis>
(
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,
<specimenCount id="66EAFD6FFFFAFFF43508FB17F79EFB35" box="[1364,1420,1214,1241]" count="5" pageId="10" pageNumber="587" type="female">5♀♀</specimenCount>
).
<emphasis id="4298EAF4FFFAFFF430CBFB4BF2C5FB10" bold="true" box="[151,215,1250,1276]" pageId="10" pageNumber="587">
<collectingCountry id="08FB7676FFFAFFF430CBFB4BF2C5FB10" box="[151,215,1250,1276]" name="India" pageId="10" pageNumber="587">India</collectingCountry>
</emphasis>
: NE India,
<collectingRegion id="B228F804FFFAFFF43105FB4AF023FB11" box="[345,561,1251,1277]" country="India" name="Arunachal Pradesh" pageId="10" pageNumber="587">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="9037AC9CFFFAFFF43260FB4AF092FB11" box="[572,640,1251,1277]" pageId="10" pageNumber="587">Roing</collectingMunicipality>
vicinity,
<geoCoordinate id="15D85021FFFAFFF432B6FB4AF166FB11" box="[746,884,1251,1277]" degrees="28" direction="north" minutes="08" orientation="latitude" pageId="10" pageNumber="587" precision="15" seconds="32" value="28.142221">28°0832”N</geoCoordinate>
<geoCoordinate id="15D85021FFFAFFF43327FB4BF1C9FB11" box="[891,987,1250,1277]" degrees="95" direction="east" minutes="51" orientation="longitude" pageId="10" pageNumber="587" precision="925" value="95.85">95°51E</geoCoordinate>
,
<quantity id="B7149B03FFFAFFF433BBFB4BF638FB11" box="[999,1066,1250,1277]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="587" unit="m" value="500.0">
<elevation id="FBC1D1D5FFFAFFF433BBFB4BF638FB11" box="[999,1066,1250,1277]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="587" unit="m" value="500.0">500m</elevation>
</quantity>
, sifting+Winkler app. extraction,
<date id="04521026FFFAFFF430CBFAAFF332FACD" box="[151,288,1286,1313]" pageId="10" pageNumber="587" value="2012-06-02" valueMax="2012-06-05" valueMin="2012-06-02">
<collectingDate id="1416E9CEFFFAFFF430CBFAAFF332FACD" box="[151,288,1286,1313]" pageId="10" pageNumber="587" value="2012-06-02" valueMax="2012-06-05" valueMin="2012-06-02">25.vi.2012</collectingDate>
</date>
,
<collectorName id="DD195330FFFAFFF43176FAAEF3ACFACD" box="[298,446,1287,1313]" pageId="10" pageNumber="587">L. Dembický</collectorName>
<emphasis id="4298EAF4FFFAFFF43198FAAEF3E4FACD" box="[452,502,1287,1313]" italics="true" pageId="10" pageNumber="587">legit</emphasis>
[
<specimenCount id="66EAFD6FFFFAFFF4325AFAAEF03BFACD" box="[518,553,1287,1313]" count="1" pageId="10" pageNumber="587" type="female">1♀</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF43269FAAEF06AFACD" box="[565,632,1287,1313]" pageId="10" pageNumber="587">JSCS</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="C0843CBBFFFAFFF432D5FAAEF1CDFAA9" collectingDate="2002-05-01" collectingDateMax="2002-05-09" collectingDateMin="2002-05-01" collectionCode="NMPC, GMOC" collectorName="Vit Kuban" country="Laos" elevation="1100" latitude="20.75" location="17 km NEE Oudomxai" longLatPrecision="1266" longitude="102.15" pageId="10" pageNumber="587" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Oudomxai province">
<emphasis id="4298EAF4FFFAFFF432D5FAAEF0D1FACD" bold="true" box="[649,707,1287,1313]" pageId="10" pageNumber="587">
<collectingCountry id="08FB7676FFFAFFF432D5FAAEF0D1FACD" box="[649,707,1287,1313]" name="Laos" pageId="10" pageNumber="587">Laos</collectingCountry>
</emphasis>
:
<collectingRegion id="B228F804FFFAFFF4328CFAAEF1A0FACD" box="[720,946,1287,1313]" country="Laos" name="Oudomxai" pageId="10" pageNumber="587">Oudomxai province</collectingRegion>
,
<location id="7533603DFFFAFFF433E0FAAEF6AAFACD" LSID="urn:lsid:plazi:treatment:F84587F0FFFAFFF530CBFD53F136F855:7533603DFFFAFFF433E0FAAEF6AAFACD" box="[956,1208,1287,1313]" country="Laos" latitude="20.75" longLatPrecision="1266" longitude="102.15" name="17 km NEE Oudomxai" pageId="10" pageNumber="587" stateProvince="Oudomxai province">
<quantity id="B7149B03FFFAFFF433E0FAAEF1E9FACD" box="[956,1019,1287,1313]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="587" unit="km" value="17.0">17km</quantity>
NEE Oudomxai
</location>
,
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<geoCoordinate id="15D85021FFFAFFF43576FAAEF785FACD" box="[1322,1431,1287,1313]" degrees="102" direction="east" minutes="09" orientation="longitude" pageId="10" pageNumber="587" precision="925" value="102.15">102°09E</geoCoordinate>
, ~
<quantity id="B7149B03FFFAFFF430F9FA82F2E4FAA9" box="[165,246,1323,1349]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="587" unit="m" value="1100.0">
<elevation id="FBC1D1D5FFFAFFF430F9FA82F2E4FAA9" box="[165,246,1323,1349]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="587" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<date id="04521026FFFAFFF43158FA82F391FAA9" box="[260,387,1323,1349]" pageId="10" pageNumber="587" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">
<collectingDate id="1416E9CEFFFAFFF43158FA82F391FAA9" box="[260,387,1323,1349]" pageId="10" pageNumber="587" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">19.v.2002</collectingDate>
</date>
,
<collectorName id="DD195330FFFAFFF431CDFA82F01AFAA9" box="[401,520,1323,1349]" pageId="10" pageNumber="587">Vit Kubáň</collectorName>
<emphasis id="4298EAF4FFFAFFF4324EFA82F056FAA9" box="[530,580,1323,1349]" italics="true" pageId="10" pageNumber="587">legit</emphasis>
[
<specimenCount id="66EAFD6FFFFAFFF43209FA82F083FAA9" box="[597,657,1323,1349]" count="2" pageId="10" pageNumber="587" type="male">2♂♂</specimenCount>
,
<specimenCount id="66EAFD6FFFFAFFF432FCFA82F0C8FAA9" box="[672,730,1323,1349]" count="2" pageId="10" pageNumber="587" type="female">2♀♀</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF432B5FA82F12EFAA9" box="[745,828,1323,1349]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="587">NMPC</collectionCode>
;
<specimenCount id="66EAFD6FFFFAFFF43316FA82F17DFAA9" box="[842,879,1323,1349]" count="1" pageId="10" pageNumber="587" type="male">1♂</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF43322FA82F1CAFAA9" box="[894,984,1323,1349]" pageId="10" pageNumber="587">GMOC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="C0843CBBFFFAFFF433B2FA82F178FA85" collectionCode="NMPC" collectorName="Vit Kuban" elevation="1100" latitude="21.633333" location="Ban Bun Neua" longLatPrecision="1263" longitude="101.95" pageId="10" pageNumber="587" specimenCount="1" specimenCount-male="1" stateProvince="Phongsaly province">
<collectingRegion id="B228F804FFFAFFF433B2FA82F6C3FAA9" box="[1006,1233,1323,1349]" country="Laos" name="Phongsali" pageId="10" pageNumber="587">Phongsaly province</collectingRegion>
,
<locationDeviation id="61E12FC4FFFAFFF43483FA82F73EFAA9" bearing="90.0" box="[1247,1324,1323,1349]" distance="4000.0" pageId="10" pageNumber="587">
<quantity id="B7149B03FFFAFFF43483FA82F700FAA9" box="[1247,1298,1323,1349]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="587" unit="km" value="4.0">4km</quantity>
E
</locationDeviation>
<location id="7533603DFFFAFFF4356AFA82F2C7FA85" LSID="urn:lsid:plazi:treatment:F84587F0FFFAFFF530CBFD53F136F855:7533603DFFFAFFF4356AFA82F2C7FA85" latitude="21.633333" longLatPrecision="1263" longitude="101.95" name="Ban Bun Neua" pageId="10" pageNumber="587" stateProvince="Phongsaly province">Ban Bun Neua</location>
,
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<geoCoordinate id="15D85021FFFAFFF43119FAE7F3A1FA85" box="[325,435,1358,1385]" degrees="101" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="587" precision="925" value="101.95">101°57E</geoCoordinate>
, ~
<quantity id="B7149B03FFFAFFF43190FAE6F00EFA85" box="[460,540,1359,1385]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="587" unit="m" value="1100.0">
<elevation id="FBC1D1D5FFFAFFF43190FAE6F00EFA85" box="[460,540,1359,1385]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="587" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<collectorName id="DD195330FFFAFFF4327AFAE6F08BFA85" box="[550,665,1359,1385]" pageId="10" pageNumber="587">Vit Kubáň</collectorName>
<emphasis id="4298EAF4FFFAFFF432C2FAE6F0C2FA85" box="[670,720,1359,1385]" italics="true" pageId="10" pageNumber="587">legit</emphasis>
[
<specimenCount id="66EAFD6FFFFAFFF43282FAE6F111FA85" box="[734,771,1359,1385]" count="1" pageId="10" pageNumber="587" type="male">1♂</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF43353FAE6F177FA85" box="[783,869,1359,1385]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="587">NMPC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="C0843CBBFFFAFFF43329FAE6F1A4FA61" collectingDate="2011-05-04" collectingDateMax="2011-05-18" collectingDateMin="2011-05-04" collectionCode="NMBS" elevation="600" location="Bokeo Nature Reserve" municipality="Ban Toup" pageId="10" pageNumber="587" specimenCount="1" specimenCount-female="1" stateProvince="Bokeo province">
<collectingRegion id="B228F804FFFAFFF43329FAE6F63BFA85" box="[885,1065,1359,1385]" country="Laos" name="Bokeo" pageId="10" pageNumber="587">Bokeo province</collectingRegion>
,
<locationDeviation id="61E12FC4FFFAFFF4346EFAE7F691FA85" bearing="270.0" box="[1074,1155,1358,1385]" distance="5000.0" pageId="10" pageNumber="587">
<quantity id="B7149B03FFFAFFF4346EFAE7F676FA85" box="[1074,1124,1358,1385]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="587" unit="km" value="5.0">5km</quantity>
W
</locationDeviation>
<collectingMunicipality id="9037AC9CFFFAFFF434D5FAE6F6E5FA85" box="[1161,1271,1359,1385]" pageId="10" pageNumber="587">Ban Toup</collectingMunicipality>
,
<location id="7533603DFFFAFFF4355DFAE6F2E6FA61" LSID="urn:lsid:plazi:treatment:F84587F0FFFAFFF530CBFD53F136F855:7533603DFFFAFFF4355DFAE6F2E6FA61" municipality="Ban Toup" name="Bokeo Nature Reserve" pageId="10" pageNumber="587" stateProvince="Bokeo province">Bokeo Nature Reserve</location>
, 20°2728N 100°45E,
<quantity id="B7149B03FFFAFFF43256FADBF096FA61" box="[522,644,1394,1421]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="587" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
<elevation id="FBC1D1D5FFFAFFF43256FADBF096FA61" box="[522,644,1394,1421]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="587" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700m</elevation>
</quantity>
,
<date id="04521026FFFAFFF432CCFADAF10BFA61" box="[656,793,1395,1421]" pageId="10" pageNumber="587" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">
<collectingDate id="1416E9CEFFFAFFF432CCFADAF10BFA61" box="[656,793,1395,1421]" pageId="10" pageNumber="587" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">418.v.2011</collectingDate>
</date>
[
<specimenCount id="66EAFD6FFFFAFFF43376FADAF15EFA61" box="[810,844,1395,1421]" count="1" pageId="10" pageNumber="587" type="female">1♀</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF43305FADAF1BCFA61" box="[857,942,1395,1421]" collectionName="Switzerland, Bern, Naturhistorische Museums" pageId="10" pageNumber="587">NMBS</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="C0843CBBFFFAFFF4339DFADAF6F8FA5D" collectingDate="2011-05-04" collectingDateMax="2011-05-18" collectingDateMin="2011-05-04" collectionCode="NMBS" collectorName="M. Brancucci" elevation="600" location="Bokeo Nature Reserve" municipality="Ban Toup" pageId="10" pageNumber="587" specimenCount="1" specimenCount-female="1" stateProvince="Bokeo province">
<collectingRegion id="B228F804FFFAFFF4339DFADAF664FA61" box="[961,1142,1395,1421]" country="Laos" name="Bokeo" pageId="10" pageNumber="587">Bokeo province</collectingRegion>
,
<locationDeviation id="61E12FC4FFFAFFF434DCFADBF6C0FA61" bearing="270.0" box="[1152,1234,1394,1421]" distance="5000.0" pageId="10" pageNumber="587">
<quantity id="B7149B03FFFAFFF434DCFADBF6A0FA61" box="[1152,1202,1394,1421]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="587" unit="km" value="5.0">5km</quantity>
W
</locationDeviation>
<collectingMunicipality id="9037AC9CFFFAFFF43485FADAF75AFA61" box="[1241,1352,1395,1421]" pageId="10" pageNumber="587">Ban Toup</collectingMunicipality>
,
<location id="7533603DFFFAFFF4350FFADAF355FA5D" LSID="urn:lsid:plazi:treatment:F84587F0FFFAFFF530CBFD53F136F855:7533603DFFFAFFF4350FFADAF355FA5D" municipality="Ban Toup" name="Bokeo Nature Reserve" pageId="10" pageNumber="587" stateProvince="Bokeo province">Bokeo Nature Reserve</location>
, 20°2728N 100°45E,
<quantity id="B7149B03FFFAFFF43202FA3FF0CAFA5D" box="[606,728,1430,1457]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="587" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
<elevation id="FBC1D1D5FFFAFFF43202FA3FF0CAFA5D" box="[606,728,1430,1457]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="587" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700m</elevation>
</quantity>
,
<date id="04521026FFFAFFF432B8FA3EF17DFA5D" box="[740,879,1431,1457]" pageId="10" pageNumber="587" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">
<collectingDate id="1416E9CEFFFAFFF432B8FA3EF17DFA5D" box="[740,879,1431,1457]" pageId="10" pageNumber="587" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">418.v.2011</collectingDate>
</date>
,
<collectorName id="DD195330FFFAFFF43327FA3EF606FA5D" box="[891,1044,1431,1457]" pageId="10" pageNumber="587">M. Brancucci</collectorName>
<emphasis id="4298EAF4FFFAFFF43446FA3EF65EFA5D" box="[1050,1100,1431,1457]" italics="true" pageId="10" pageNumber="587">legit</emphasis>
[
<specimenCount id="66EAFD6FFFFAFFF43401FA3EF692FA5D" box="[1117,1152,1431,1457]" count="1" pageId="10" pageNumber="587" type="female">1♀</specimenCount>
,
<collectionCode id="16FDAE23FFFAFFF434D1FA3EF6F1FA5D" box="[1165,1251,1431,1457]" collectionName="Switzerland, Bern, Naturhistorische Museums" pageId="10" pageNumber="587">NMBS</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="38F6656DFFFAFFF4309BFA13F630F941" pageId="10" pageNumber="587" type="diagnosis">
<paragraph id="705336E6FFFAFFF4309BFA13F630F941" blockId="10.[151,1437,1034,2033]" pageId="10" pageNumber="587">
<emphasis id="4298EAF4FFFAFFF4309BFA13F328FA38" bold="true" box="[199,314,1466,1492]" pageId="10" pageNumber="587">Diagnosis</emphasis>
.
<taxonomicName id="B7EC4D65FFFAFFF43115FA12F07BFA39" authorityName="Montanaro" authorityYear="2024" box="[329,617,1467,1493]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="schoenfeldi" status="sp. nov.">
<emphasis id="4298EAF4FFFAFFF43115FA12F07BFA39" box="[329,617,1467,1493]" italics="true" pageId="10" pageNumber="587">Sinodrepanus schoenfeldi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="59AB578FFFFAFFF4322CFA12F0E8FA38" box="[624,762,1466,1493]" pageId="10" pageNumber="587" rank="species">new species</taxonomicNameLabel>
</emphasis>
can be easily separated from most
<taxonomicName id="B7EC4D65FFFAFFF434D4FA12F76BFA39" box="[1160,1401,1467,1493]" pageId="10" pageNumber="587">
<emphasis id="4298EAF4FFFAFFF434D4FA12F733FA39" box="[1160,1313,1467,1493]" italics="true" pageId="10" pageNumber="587">Sinodrepanus</emphasis>
species
</taxonomicName>
by the combination of the following characters: i) the longest setae on interstria 5 irregularly distributed, not arranged into tufts; ii) three teeth on the outer protibial margin; iii) male clypeus with two teeth plus an additional small stout median tooth (
<figureCitation id="E8D72A63FFFAFFF43167F98EF39DF9AD" box="[315,399,1575,1601]" captionStart="FIGURE 3" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[155,1433,182,1919]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Details of Sinodrepanus species. S. exsul: male (a) and female (c) head; posterior pronotal punctuation (e). S. schoenfeldi new species: male (b) and female (d) head; posterior pronotal punctuation (f). S. arrowi, lectotype (g) and its labels (h) (pictures by Jiří Hájek, NMPC)." figureDoi="http://doi.org/10.5281/zenodo.14240166" httpUri="https://zenodo.org/record/14240166/files/figure.png" pageId="10" pageNumber="587">Fig. 3b</figureCitation>
); iv) females with a sharp small denticle between clypeal teeth (
<figureCitation id="E8D72A63FFFAFFF4343AF98EF6A9F9AD" box="[1126,1211,1575,1601]" captionStart="FIGURE 3" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[155,1433,182,1919]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Details of Sinodrepanus species. S. exsul: male (a) and female (c) head; posterior pronotal punctuation (e). S. schoenfeldi new species: male (b) and female (d) head; posterior pronotal punctuation (f). S. arrowi, lectotype (g) and its labels (h) (pictures by Jiří Hájek, NMPC)." figureDoi="http://doi.org/10.5281/zenodo.14240166" httpUri="https://zenodo.org/record/14240166/files/figure.png" pageId="10" pageNumber="587">Fig. 3d</figureCitation>
); v) parameres with very short basolateral plate (
<figureCitation id="E8D72A63FFFAFFF4318BF9E2F040F989" box="[471,594,1611,1637]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[182,1401,185,1914]" captionTargetId="figure-20@7.[177,1410,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Genitalia of Sinodrepanus species. S. exsul: aedeagus in lateral (a), posterior (b) and dorsal (c) views; lamella copulatrix (d); female genitalia (i). S. schoenfeldi new species: aedeagus in lateral (e), posterior (f) and dorsal (g) views; lamella copulatrix (h); female genitalia (j)." figureDoi="http://doi.org/10.5281/zenodo.14240164" httpUri="https://zenodo.org/record/14240164/files/figure.png" pageId="10" pageNumber="587">Figs. 2eg</figureCitation>
). Except for the third and last characters, all other features are also present in
<taxonomicName id="B7EC4D65FFFAFFF430EAF9C6F31DF965" box="[182,271,1647,1673]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFAFFF430EAF9C6F31DF965" box="[182,271,1647,1673]" italics="true" pageId="10" pageNumber="587">S. exsul</emphasis>
</taxonomicName>
, from which however it can be easily separated also by the punctured (not smooth) lateral surface of sternites 36 and the circular (not elongated) punctures of pronotal disc (
<figureCitation id="E8D72A63FFFAFFF4339AF93AF607F941" box="[966,1045,1683,1709]" captionStart="FIGURE 3" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[155,1433,182,1919]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Details of Sinodrepanus species. S. exsul: male (a) and female (c) head; posterior pronotal punctuation (e). S. schoenfeldi new species: male (b) and female (d) head; posterior pronotal punctuation (f). S. arrowi, lectotype (g) and its labels (h) (pictures by Jiří Hájek, NMPC)." figureDoi="http://doi.org/10.5281/zenodo.14240166" httpUri="https://zenodo.org/record/14240166/files/figure.png" pageId="10" pageNumber="587">Fig. 3f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="38F6656DFFFAFFF5309BF91FF164FA71" lastPageId="11" lastPageNumber="588" pageId="10" pageNumber="587" type="description">
<paragraph id="705336E6FFFAFFF5309BF91FF375FBFD" blockId="10.[151,1437,1034,2033]" lastBlockId="11.[151,1437,151,1977]" lastPageId="11" lastPageNumber="588" pageId="10" pageNumber="587">
<emphasis id="4298EAF4FFFAFFF4309BF91FF074F93D" bold="true" box="[199,614,1718,1745]" pageId="10" pageNumber="587">
Description of the
<typeStatus id="AF578844FFFAFFF431F2F91EF004F93D" box="[430,534,1719,1745]" pageId="10" pageNumber="587" type="holotype">holotype</typeStatus>
, male.
</emphasis>
Flat, elongated, integument microreticulated, dark brown and partially with metallic lustre; antennae reddish-brown; body surface almost wholly covered by simple or ocellate punctures which carry light brown scale-like setae.
<emphasis id="4298EAF4FFFAFFF43202F956F0A6F8F5" bold="true" box="[606,692,1791,1817]" pageId="10" pageNumber="587">Length</emphasis>
:
<quantity id="B7149B03FFFAFFF4329EF956F108F8F5" box="[706,794,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="587" unit="mm" value="8.6">8.6 mm</quantity>
.
<emphasis id="4298EAF4FFFAFFF4337FF956F171F8F5" bold="true" box="[803,867,1791,1817]" pageId="10" pageNumber="587">Head</emphasis>
. Subpentagonal, clypeus covered by dense ocellate setigerous punctures; vertex with two weak, short longitudinal carinae bearing long testaceous bristles; carinae dividing the interocular space into three almost equal segments; frons bulging medially; lateral edge of genae obtusely angular; head margin obtusely angular in correspondence of clypeogenal junctions; anterior clypeal margin with two upturned teeth and a third, very small medial clypeal tooth, margin broadly obtusely angled medially.
<emphasis id="4298EAF4FFFAFFF43540F826F787F845" bold="true" box="[1308,1429,1935,1961]" pageId="10" pageNumber="587">Pronotum</emphasis>
. Subquadrangular, widening anteriorly, lateral margin approximately straight posteriorly; anterolateral angles evenly curved; posterior angles broadly rounded; posterior margin broadly rounded, obtuse medially; covered with ocellate setigerous punctures of uneven size, punctures circular on disc. Pronotal disc with 8 ill-defined longitudinal carinae bearing testaceous bristles (in order, from medial to lateral): 2 carinae on either side of the midline, reaching the anterior pronotal border but expiring before posterior edge, slightly converging at about the anterior 2/5 of their length, where disc is transversely swollen; laterally to these, 2 short carinae connected to the posterior pronotal border and 2 short oblique carinae anteriorly, connecting to the anterior part of the medial carinae; other 2 bulge-like oblique carinae on the posterior and anterior thirds of pronotum, close to the lateral margins.
<emphasis id="4298EAF4FFFBFFF53491FEE2F70BFE89" bold="true" box="[1229,1305,331,357]" pageId="11" pageNumber="588">Elytra</emphasis>
. Flat on the whole, interstria 1 raised, interstria 6 depressed longitudinally; interstriae 57 obliquely elevated next to humeral callus, interstria 6 depressed a little behind callus; striae broad, their borders clearly ridged, punctures marked, each one carrying a scale-like seta; interstriae uniformly covered with setigerous punctures; interstriae 13, 5 and 7 with a tuft of long setae apically; interstriae 1, 5 and 7 carrying longer setae, setae on interstria 5 irregularly distributed; interstria 8 covered with longer and more scattered setae.Scutellar shield approximately parallel-sided, smooth.
<emphasis id="4298EAF4FFFBFFF53501FE56F787FDF5" bold="true" box="[1373,1429,511,537]" pageId="11" pageNumber="588">Legs</emphasis>
. Anterior surface of femur covered by dense small setigerous punctures and by scattered ocellate punctures carrying thicker setae. Protibia with distal margin obliquely truncated; ventral protibial margin strongly curved, slightly produced distally; apical protibial spur with distal half slightly bent; dorsal protibial margin with three small teeth.
<emphasis id="4298EAF4FFFBFFF530CBFD27F313FD44" bold="true" box="[151,257,654,680]" pageId="11" pageNumber="588">Tergite 8</emphasis>
. Densely setigerous; strongly swollen medially, with two broad bulges; strongly depressed along anterior margin; with a longitudinal medial carina connected to the anterior margin.
<emphasis id="4298EAF4FFFBFFF53452FD1AF71CFD20" bold="true" box="[1038,1294,690,717]" pageId="11" pageNumber="588">Ventral body surface</emphasis>
. Abdominal sternites with a tuft of long setae laterally next to the epipleuron, evenly covered with setigerous punctures, setae longer medially. Metaventral disc covered by small ocellate punctures intermixed with few bigger punctures, with a pair of symmetrical shallow depressions next to posteromedian margin; lateral metaventral surface microreticulated, with big ocellate puctures intermixed with many smaller setigerous punctures.
<emphasis id="4298EAF4FFFBFFF5344DFCEBF66DFCB0" bold="true" box="[1041,1151,834,860]" pageId="11" pageNumber="588">Genitalia</emphasis>
(
<figureCitation id="E8D72A63FFFBFFF534CCFCEAF718FCB1" box="[1168,1290,835,861]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[182,1401,185,1914]" captionTargetId="figure-20@7.[177,1410,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Genitalia of Sinodrepanus species. S. exsul: aedeagus in lateral (a), posterior (b) and dorsal (c) views; lamella copulatrix (d); female genitalia (i). S. schoenfeldi new species: aedeagus in lateral (e), posterior (f) and dorsal (g) views; lamella copulatrix (h); female genitalia (j)." figureDoi="http://doi.org/10.5281/zenodo.14240164" httpUri="https://zenodo.org/record/14240164/files/figure.png" pageId="11" pageNumber="588">Figs. 2eh</figureCitation>
). Basolateral plates of parameres relatively short, not reaching parameral apices in lateral view; parameral apices well developed, strongly bent downwards; parameral apices rounded in frontal view. Lamella copulatrix (LC) four-lobed: superior left lobe with a flattened digitiform expansion bent posteriorly; superior right lobe simple, flattened; inferior left lobe relatively thick, arched; inferior right lobe separated from the rest of the LC, associated with an inconspicuous additional sclerite.
</paragraph>
<paragraph id="705336E6FFFBFFF5309BFBB2F68DFB29" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BFBB2F30CFBD9" bold="true" box="[199,286,1051,1077]" pageId="11" pageNumber="588">Female</emphasis>
(
<typeStatus id="AF578844FFFBFFF53173FBB2F39DFBD9" box="[303,399,1051,1077]" pageId="11" pageNumber="588" type="allotype">allotype</typeStatus>
). Similar to the male except for the following characters.
<emphasis id="4298EAF4FFFBFFF53479FBB2F677FBD9" bold="true" box="[1061,1125,1051,1077]" pageId="11" pageNumber="588">Head</emphasis>
. Symmetrical clypeal teeth more developed, space between them broader, median denticle more developed and sharp; clypeal margin slightly notched at clypeogenal margin.
<emphasis id="4298EAF4FFFBFFF531AAFBCAF03CFB91" bold="true" box="[502,558,1123,1149]" pageId="11" pageNumber="588">Legs</emphasis>
. Outer protibial teeth larger than in the male; inner protibial margin not strongly curved.
<emphasis id="4298EAF4FFFBFFF530AEFB2FF349FB4C" bold="true" box="[242,347,1158,1184]" pageId="11" pageNumber="588">Tergite 8</emphasis>
. Shorter than in the male.
<emphasis id="4298EAF4FFFBFFF532DAFB2EF16DFB4C" bold="true" box="[646,895,1158,1185]" pageId="11" pageNumber="588">Ventral body surface</emphasis>
. S8 longer than in the male.
<emphasis id="4298EAF4FFFBFFF53498FB2FF720FB4C" bold="true" box="[1220,1330,1158,1184]" pageId="11" pageNumber="588">Genitalia</emphasis>
(
<figureCitation id="E8D72A63FFFBFFF5351EFB2EF79DFB4D" box="[1346,1423,1159,1185]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[182,1401,185,1914]" captionTargetId="figure-20@7.[177,1410,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Genitalia of Sinodrepanus species. S. exsul: aedeagus in lateral (a), posterior (b) and dorsal (c) views; lamella copulatrix (d); female genitalia (i). S. schoenfeldi new species: aedeagus in lateral (e), posterior (f) and dorsal (g) views; lamella copulatrix (h); female genitalia (j)." figureDoi="http://doi.org/10.5281/zenodo.14240164" httpUri="https://zenodo.org/record/14240164/files/figure.png" pageId="11" pageNumber="588">Fig. 2j</figureCitation>
). Vagina relatively poorly sclerotised, with many folds; infundibulum broadly curved distally.
</paragraph>
<paragraph id="705336E6FFFBFFF5309BFB67F74CFAB9" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BFB67F32AFB04" bold="true" box="[199,312,1230,1256]" pageId="11" pageNumber="588">Variation</emphasis>
. Body size ranges from 8.0 to 9.0 mm. The clypeal margin can be more or less markedly angular at clypeogenal junctions. Punctures on the basal half of disc can sometimes be slightly oval, but never clearly elongated as in
<taxonomicName id="B7EC4D65FFFBFFF53119FABEF389FADD" box="[325,411,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFBFFF53119FABEF389FADD" box="[325,411,1303,1329]" italics="true" pageId="11" pageNumber="588">S. exsul</emphasis>
</taxonomicName>
. In
<specimenCount id="66EAFD6FFFFBFFF5319BFABEF03EFADD" box="[455,556,1303,1329]" count="1" pageId="11" pageNumber="588" type="male">one male</specimenCount>
the median clypeal denticle was completely effaced, probably due to wear. No significant external or genital differences were observed between specimens from
<collectingCountry id="08FB7676FFFBFFF5346DFA92F679FAB9" box="[1073,1131,1339,1365]" name="India" pageId="11" pageNumber="588">India</collectingCountry>
and those from
<collectingCountry id="08FB7676FFFBFFF5357CFA92F748FAB9" box="[1312,1370,1339,1365]" name="Laos" pageId="11" pageNumber="588">Laos</collectingCountry>
.
</paragraph>
<paragraph id="705336E6FFFBFFF5309BFAF7F164FA71" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BFAF7F3D3FA94" bold="true" box="[199,449,1374,1400]" pageId="11" pageNumber="588">Verified distribution</emphasis>
:
<collectingCountry id="08FB7676FFFBFFF53189FAF6F01DFA95" box="[469,527,1375,1401]" name="India" pageId="11" pageNumber="588">India</collectingCountry>
(
<collectingRegion id="B228F804FFFBFFF53278FAF6F067FA95" box="[548,629,1375,1401]" country="India" name="Assam" pageId="11" pageNumber="588">Assam</collectingRegion>
,
<collectingRegion id="B228F804FFFBFFF532D8FAF6F170FA95" box="[644,866,1375,1401]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="588">Arunachal Pradesh</collectingRegion>
),
<collectingCountry id="08FB7676FFFBFFF53327FAF6F1A3FA95" box="[891,945,1375,1401]" name="Laos" pageId="11" pageNumber="588">Laos</collectingCountry>
(
<figureCitation id="E8D72A63FFFBFFF5339AFAF6F603FA95" box="[966,1041,1375,1401]" captionStart="FIGURE 4" captionStartId="12.[152,255,1032,1056]" captionTargetBox="[151,1436,182,1009]" captionTargetId="figure-23@12.[151,1436,181,1009]" captionTargetPageId="12" captionText="FIGURE 4. Distribution of Sinodrepanus species. The position of the holotype of S. exsul is chosen arbitrarily within Thailand due to the lack of precise georeferentiation." figureDoi="http://doi.org/10.5281/zenodo.14240172" httpUri="https://zenodo.org/record/14240172/files/figure.png" pageId="11" pageNumber="588">Fig. 4</figureCitation>
). See the remarks in the “
<taxonomicName id="B7EC4D65FFFBFFF5350EFAF6F78EFA95" box="[1362,1436,1375,1401]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="11" pageNumber="595" phylum="Chordata" rank="genus">Indian</taxonomicName>
<taxonomicName id="B7EC4D65FFFBFFF530CBFA2AF322FA71" authorityName="Simonis" authorityYear="1985" box="[151,304,1411,1437]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF530CBFA2AF322FA71" box="[151,304,1411,1437]" italics="true" pageId="11" pageNumber="588">Sinodrepanus</emphasis>
</taxonomicName>
: an overlooked diversity?” section for more details.
</paragraph>
</subSubSection>
<subSubSection id="38F6656DFFFBFFF5309BFA0EF71FFA2D" box="[199,1293,1447,1473]" pageId="11" pageNumber="588" type="biology_ecology">
<paragraph id="705336E6FFFBFFF5309BFA0EF71FFA2D" blockId="11.[151,1437,151,1977]" box="[199,1293,1447,1473]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BFA0EF334FA2D" bold="true" box="[199,294,1447,1473]" pageId="11" pageNumber="588">Ecology</emphasis>
. In
<collectingCountry id="08FB7676FFFBFFF5310EFA0EF39CFA2D" box="[338,398,1447,1473]" name="India" pageId="11" pageNumber="588">India</collectingCountry>
, this species inhabits evergreen rainforests on the eastern foothills of Himalaya.
</paragraph>
</subSubSection>
<subSubSection id="38F6656DFFFBFFF5309BFA62F306F9C1" pageId="11" pageNumber="588" type="etymology">
<paragraph id="705336E6FFFBFFF5309BFA62F306F9C1" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BFA62F35AFA09" bold="true" box="[199,328,1483,1509]" pageId="11" pageNumber="588">Etymology</emphasis>
. I am pleased to dedicate the new species to Joachim Schönfeld (Sinzig,
<collectingCountry id="08FB7676FFFBFFF534CFFA62F712FA09" box="[1171,1280,1483,1509]" name="Germany" pageId="11" pageNumber="588">Germany</collectingCountry>
), specialist of Old World
<taxonomicName id="B7EC4D65FFFBFFF5314EFA46F3BBF9E5" authorityName="Latreille" authorityYear="1802" box="[274,425,1519,1545]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
, to whom I am grateful for his friendly support and help in gathering important dung beetle specimens.
</paragraph>
</subSubSection>
<subSubSection id="38F6656DFFFBFFF5309BF99EF136F855" pageId="11" pageNumber="588" type="discussion">
<paragraph id="705336E6FFFBFFF5309BF99EF3DBF975" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF5309BF99EF326F9BD" bold="true" box="[199,308,1591,1617]" pageId="11" pageNumber="588">Remarks</emphasis>
. This species was overlooked by several authors due to the similarity with
<taxonomicName id="B7EC4D65FFFBFFF5342AF99EF6C1F9BD" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[1142,1235,1591,1617]" class="Insecta" family="Scarabaeidae" genus="Drepanocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="4298EAF4FFFBFFF5342AF99EF6C1F9BD" box="[1142,1235,1591,1617]" italics="true" pageId="11" pageNumber="588">S. falsus</emphasis>
</taxonomicName>
and
<taxonomicName id="B7EC4D65FFFBFFF53554F99EF74CF9BD" box="[1288,1374,1591,1617]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFBFFF53554F99EF74CF9BD" box="[1288,1374,1591,1617]" italics="true" pageId="11" pageNumber="588">S. exsul</emphasis>
</taxonomicName>
. This error was first made by Arrow, who identified the
<typeStatus id="AF578844FFFBFFF5329AF9F2F13AF999" box="[710,808,1627,1653]" pageId="11" pageNumber="588" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B7EC4D65FFFBFFF53310F9F2F1FAF999" authorityName="Montanaro" authorityYear="2024" box="[844,1000,1627,1653]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="schoenfeldi" status="new species">
<emphasis id="4298EAF4FFFBFFF53310F9F2F1FAF999" box="[844,1000,1627,1653]" italics="true" pageId="11" pageNumber="588">S. schoenfeldi</emphasis>
</taxonomicName>
<emphasis id="4298EAF4FFFBFFF533B2F9F2F665F998" bold="true" box="[1006,1143,1626,1653]" pageId="11" pageNumber="588">
<taxonomicNameLabel id="59AB578FFFFBFFF533B2F9F2F665F998" box="[1006,1143,1626,1653]" pageId="11" pageNumber="588" rank="species">new species</taxonomicNameLabel>
</emphasis>
as
<taxonomicName id="B7EC4D65FFFBFFF534C0F9F2F6E0F999" box="[1180,1266,1627,1653]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFBFFF534C0F9F2F6E0F999" box="[1180,1266,1627,1653]" italics="true" pageId="11" pageNumber="588">S. exsul</emphasis>
</taxonomicName>
and depicted it (
<bibRefCitation id="147D4B17FFFBFFF530C3F9D6F376F975" author="Arrow, G. J." box="[159,356,1662,1689]" pageId="11" pageNumber="588" refId="ref13876" refString="Arrow, G. J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III. Coprinae. Taylor &amp; Francis, London, 428 pp." type="book" year="1931">Arrow 1931: 385</bibRefCitation>
, fig. 55).
</paragraph>
<paragraph id="705336E6FFFBFFF5309BF90AF136F855" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="588">
A remarkable feature of the new species is the presence of a third, tiny medial clypeal tooth in both males and females. This character is rare in
<taxonomicName id="B7EC4D65FFFBFFF53212F96EF0C2F90D" authorityName="Kolbe" authorityYear="1905" box="[590,720,1735,1761]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="tribe" tribe="Oniticellini">Oniticellini</taxonomicName>
, being found—to my knowledge—only in females of
<taxonomicName id="B7EC4D65FFFBFFF53518F96EF78FF90D" box="[1348,1437,1735,1761]" class="Insecta" family="Scarabaeidae" genus="Drepenocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="4298EAF4FFFBFFF53518F96EF78FF90D" box="[1348,1437,1735,1761]" italics="true" pageId="11" pageNumber="588">S. exsul</emphasis>
</taxonomicName>
and
<taxonomicName id="B7EC4D65FFFBFFF53094F942F097F8E9" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[200,645,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Scaptocnemis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="species" species="segregis">
<emphasis id="4298EAF4FFFBFFF53094F942F3D4F8E8" box="[200,454,1771,1797]" italics="true" pageId="11" pageNumber="588">Scaptocnemis segregis</emphasis>
<bibRefCitation id="147D4B17FFFBFFF53193F942F097F8E9" author="Peringuey, L." box="[463,645,1771,1797]" pageId="11" pageNumber="588" pagination="1 - 563" refId="ref15467" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society, 12, 1 - 563. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="147D4B17FFFBFFF532C9F942F15EF8E9" author="Peringuey, L." box="[661,844,1771,1797]" pageId="11" pageNumber="588" pagination="1 - 563" refId="ref15467" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society, 12, 1 - 563. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey 1901</bibRefCitation>
), and in the
<taxonomicName id="B7EC4D65FFFBFFF53386F942F78EF8E9" box="[986,1436,1770,1797]" pageId="11" pageNumber="588">
<emphasis id="4298EAF4FFFBFFF53386F942F692F8E9" box="[986,1152,1771,1797]" italics="true" pageId="11" pageNumber="588">Helictopleurus</emphasis>
dOrbigny, 1915 species
</taxonomicName>
of the
<emphasis id="4298EAF4FFFBFFF530BFF8A6F35FF8C5" box="[227,333,1807,1833]" italics="true" pageId="11" pageNumber="588">fungicola</emphasis>
-group (
<bibRefCitation id="147D4B17FFFBFFF531F6F8A6F095F8C5" author="Rossini, M. &amp; Montreuil, O. &amp; Grebennikov, V. &amp; Tarasov, S." box="[426,647,1807,1833]" pageId="11" pageNumber="588" pagination="63 - 79" refId="ref15614" refString="Rossini, M., Montreuil, O., Grebennikov, V. &amp; Tarasov, S. (2021) Genome sequencing reveals extraordinary cephalic horns in the Madagascan dung beetle genus Helictopleurus (Coleoptera, Scarabaeinae): insight from a revision of fungicola species group. ZooKeys, 1033, 63 - 79. https: // doi. org / 10.3897 / zookeys. 1033.63527" type="journal article" year="2021">
Rossini
<emphasis id="4298EAF4FFFBFFF53254F8A6F029F8C5" box="[520,571,1807,1833]" italics="true" pageId="11" pageNumber="588">et al</emphasis>
. 2021
</bibRefCitation>
). Curiously, evidence from multiple studies showed that the subtribe
<taxonomicName id="B7EC4D65FFFBFFF530CBF89AF32DF8A1" authorityName="Montanaro" authorityYear="2024" box="[151,319,1843,1869]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="subTribe" subTribe="Drepanocerina">Drepanocerina</taxonomicName>
, to which
<taxonomicName id="B7EC4D65FFFBFFF5319EF89AF049F8A1" authorityName="Simonis" authorityYear="1985" box="[450,603,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF5319EF89AF049F8A1" box="[450,603,1843,1869]" italics="true" pageId="11" pageNumber="588">Sinodrepanus</emphasis>
</taxonomicName>
is currently assigned, and the genera
<taxonomicName id="B7EC4D65FFFBFFF53479F89AF77AF8A1" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[1061,1384,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Tiniocellus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF53479F89AF6B2F8A1" box="[1061,1184,1843,1869]" italics="true" pageId="11" pageNumber="588">Tiniocellus</emphasis>
<bibRefCitation id="147D4B17FFFBFFF534F1F89AF77AF8A1" author="Peringuey, L." box="[1197,1384,1843,1869]" pageId="11" pageNumber="588" pagination="1 - 563" refId="ref15467" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society, 12, 1 - 563. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="B7EC4D65FFFBFFF530CBF8FEF3E1F89D" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[151,499,1879,1905]" class="Insecta" family="Scarabaeidae" genus="Scaptocnemis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF530CBF8FEF323F89D" box="[151,305,1879,1905]" italics="true" pageId="11" pageNumber="588">Scaptocnemis</emphasis>
<bibRefCitation id="147D4B17FFFBFFF53166F8FEF3E1F89D" author="Peringuey, L." box="[314,499,1879,1905]" pageId="11" pageNumber="588" pagination="1 - 563" refId="ref15467" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Transactions of the South African philosophical Society, 12, 1 - 563. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
</taxonomicName>
, may be strictly related (
<bibRefCitation id="147D4B17FFFBFFF53352F8FEF1C1F89D" author="Ayivi, S. P. G. &amp; Tong, Y. &amp; Storey, K. B. &amp; Yu, D. N. &amp; Zhang, J. Y." box="[782,979,1879,1905]" pageId="11" pageNumber="588" refId="ref13913" refString="Ayivi, S. P. G., Tong, Y., Storey, K. B., Yu, D. N. &amp; Zhang, J. Y. (2021) The mitochondrial genomes of 18 new Pleurosticti (Coleoptera: Scarabaeidae) exhibit a novel trnQ-NCR-trnI-trnM gene rearrangement and clarify phylogenetic relationships of subfamilies within Scarabaeidae. Insects, 12 (11), 1025. https: // doi. org / 10.3390 / insects 12111025" type="journal volume" year="2021">
Ayivi
<emphasis id="4298EAF4FFFBFFF5330AF8FEF19BF89D" box="[854,905,1879,1905]" italics="true" pageId="11" pageNumber="588">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="147D4B17FFFBFFF533BCF8FEF6E3F89D" author="Breeschoten, T. &amp; Doorenweerd, C. &amp; Tarasov, S. &amp; Vogler, A. P." box="[992,1265,1879,1905]" pageId="11" pageNumber="588" pagination="86 - 95" refId="ref14728" refString="Breeschoten, T., Doorenweerd, C., Tarasov, S. &amp; Vogler, A. P. (2016) Phylogenetics and biogeography of the dung beetle genus Onthophagus inferred from mitochondrial genomes. Molecular Phylogenetics and Evolution, 105, 86 - 95. https: // doi. org / 10.1016 / j. ympev. 2016.08.016" type="journal article" year="2016">
Breeschoten
<emphasis id="4298EAF4FFFBFFF5342FF8FEF6B4F89D" box="[1139,1190,1879,1905]" italics="true" pageId="11" pageNumber="588">et al</emphasis>
. 2016
</bibRefCitation>
;
<bibRefCitation id="147D4B17FFFBFFF534A2F8FEF783F89D" author="Philips, T. K." box="[1278,1425,1879,1905]" pageId="11" pageNumber="588" pagination="9 - 57" refId="ref15507" refString="Philips, T. K. (2016) Phylogeny of the Oniticellini and Onthophagini dung beetles (Scarabaeidae, Scarabaeinae) from morphological evidence. ZooKeys, 579, 9 - 57. https: // doi. org / 10.3897 / zookeys. 579.6183" type="journal article" year="2016">Philips 2016</bibRefCitation>
). If this is the case, one may wonder whether the three-toothed clypeus shared by
<taxonomicName id="B7EC4D65FFFBFFF53445F8D2F6A0F879" authorityName="Simonis" authorityYear="1985" box="[1049,1202,1915,1941]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF53445F8D2F6A0F879" box="[1049,1202,1915,1941]" italics="true" pageId="11" pageNumber="588">Sinodrepanus</emphasis>
</taxonomicName>
and
<taxonomicName id="B7EC4D65FFFBFFF534B5F8D2F791F879" authorityName="Peringuey" authorityYear="1901" box="[1257,1411,1915,1941]" class="Insecta" family="Scarabaeidae" genus="Scaptocnemis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="4298EAF4FFFBFFF534B5F8D2F791F879" box="[1257,1411,1915,1941]" italics="true" pageId="11" pageNumber="588">Scaptocnemis</emphasis>
</taxonomicName>
is due to convergence or, instead, is a homologous character.
</paragraph>
</subSubSection>
</treatment>
</document>