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<mods:title>Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae)</mods:title>
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<mods:namePart>Kluge, Nikita</mods:namePart>
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Genus 
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<emphasis bold="true" box="[238,388,402,429]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
<bibRefCitation author="Eaton, A. E." box="[396,538,402,429]" pageId="2" pageNumber="529" pagination="207 - 208" refId="ref10667" refString="Eaton, A. E. (1882) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 18, 207 - 208." type="journal article" year="1882">Eaton 1882</bibRefCitation>
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(
<figureCitation box="[159,286,439,465]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2–7" captionStart-10="FIGURES 61–62" captionStart-11="FIGURES 63–68" captionStart-12="FIGURES 69–74" captionStart-13="FIGURES 75–83" captionStart-14="FIGURES 84–92" captionStart-15="FIGURES 93–94" captionStart-16="FIGURES 95–101" captionStart-2="FIGURES 8–11" captionStart-3="FIGURES 12–18" captionStart-4="FIGURES 19–26" captionStart-5="FIGURES 27–30" captionStart-6="FIGURES 31–39" captionStart-7="FIGURES 40–45" captionStart-8="FIGURES 46–49" captionStart-9="FIGURES 50–60" captionStartId-0="1.[151,250,1404,1428]" captionStartId-1="5.[151,264,1841,1865]" captionStartId-10="17.[151,264,1223,1247]" captionStartId-11="18.[151,264,1827,1851]" captionStartId-12="19.[151,264,1841,1865]" captionStartId-13="20.[151,264,1845,1869]" captionStartId-14="21.[151,264,1841,1865]" captionStartId-15="22.[151,264,1218,1242]" captionStartId-16="23.[151,264,1926,1950]" captionStartId-2="9.[151,264,1800,1824]" captionStartId-3="10.[151,264,1841,1865]" captionStartId-4="11.[151,264,1841,1865]" captionStartId-5="12.[151,264,1697,1721]" captionStartId-6="13.[151,264,1840,1864]" captionStartId-7="14.[151,264,1934,1958]" captionStartId-8="15.[151,264,1758,1782]" captionStartId-9="16.[151,264,1932,1956]" captionTargetBox-0="[151,1436,455,1358]" captionTargetBox-1="[151,1436,181,1817]" captionTargetBox-10="[159,1419,181,1200]" captionTargetBox-11="[151,1436,181,1797]" captionTargetBox-12="[151,1436,181,1817]" captionTargetBox-13="[151,1418,181,1821]" captionTargetBox-14="[151,1431,181,1808]" captionTargetBox-15="[151,1435,186,1189]" captionTargetBox-16="[151,1436,273,1903]" captionTargetBox-2="[159,1429,417,1777]" captionTargetBox-3="[151,1436,181,1817]" captionTargetBox-4="[151,1436,181,1817]" captionTargetBox-5="[151,1428,182,1673]" captionTargetBox-6="[150,1438,181,1817]" captionTargetBox-7="[204,1383,634,1911]" captionTargetBox-8="[156,1432,633,1735]" captionTargetBox-9="[151,1436,290,1909]" captionTargetId-0="figure-141@1.[151,1436,450,1378]" captionTargetId-1="figure-15@5.[151,1436,181,1817]" captionTargetId-10="figure-15@17.[159,1428,181,1200]" captionTargetId-11="figure-23@18.[151,1436,181,1803]" captionTargetId-12="figure-15@19.[151,1436,181,1817]" captionTargetId-13="figure-23@20.[151,1436,181,1821]" captionTargetId-14="figure-15@21.[151,1436,181,1817]" captionTargetId-15="figure-23@22.[151,1436,181,1193]" captionTargetId-16="figure-56@23.[151,1436,273,1903]" captionTargetId-2="figure-112@9.[159,1429,417,1777]" captionTargetId-3="figure-23@10.[151,1436,181,1817]" captionTargetId-4="figure-15@11.[151,1436,181,1817]" captionTargetId-5="figure-23@12.[151,1436,181,1673]" captionTargetId-6="figure-15@13.[150,1438,181,1817]" captionTargetId-7="figure-236@14.[204,1383,633,1911]" captionTargetId-8="figure-209@15.[156,1432,633,1735]" captionTargetId-9="figure-61@16.[151,1436,273,1909]" captionTargetPageId-0="1" captionTargetPageId-1="5" captionTargetPageId-10="17" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-13="20" captionTargetPageId-14="21" captionTargetPageId-15="22" captionTargetPageId-16="23" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="16" captionText-0="FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004)." captionText-1="FIGURES 2–7. Details of Teloganodes s.l. 2–4, Teloganodes (Derlethina) eloisae, mature larvae (Malaysia, Sabah, Kindssang, Ranau, Negeri, 9.IX.1994, coll. S.C. Kang): 2, apical part of fore protopteron; 3, hind protopteron; 4, hind leg (si, outer margin of subimaginal femur developing inside). 5, egg of T. (Teloganodes) insignis (Sri Lanka, south-eastern slope of Sri Pada, river Battulu oya, Fishing Hut, 10.II.2020, coll. N. Kluge &amp; L. Sheyko). 6–7, T. (Dudgeodes) celebensis, dorsal cuticle from labrum: 6, focus on proximal part; 7, focus on distal part." captionText-10="FIGURES 61–62. Teloganodes (Dudgeodes) rasus sp. n. 61, male larva of last instar; 62, younger female larva." captionText-11="FIGURES 63–68. Teloganodes (Dudgeodes) rasus sp. n. 63–64, left and right mandibles; 65, maxilla, 66, apex of maxilla; 67, hypopharynx and superlingua; 68, labium. Abbreviations: in, incisor; kd, kinetodontium; m, mola; prs, prostheca." captionText-12="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." captionText-13="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." captionText-14="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." captionText-15="FIGURES 93–94. Teloganodes (Dudgeodes) rasus sp. n., wings of male imago (holotype). 93, in falling light; 94, in transmitted light (arrow shows constant crossvein cua-cup)." captionText-16="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." captionText-2="FIGURES 8–11. Teloganodes (Dudgeodes) celebensis. 8, young larva in dry condition, to show relief; 9–11, fore, middle and hind legs of young larva to show black hypodermal coloration." captionText-3="FIGURES 12–18. Teloganodes (Dudgeodes) celebensis, larval cuticle. 12, latero-posterior margin of head; 13, left half of pronotum and mesonotum; 14–18, left half of abdomen and its enlarged fragments." captionText-4="FIGURES 19–26. Teloganodes (Dudgeodes) celebensis, exuviae of larval legs. 19–20, fore leg and its fragment; 21, middle leg; 22–26, hind leg and its fragments." captionText-5="FIGURES 27–30. Teloganodes (Dudgeodes) celebensis. 27–28, left half of subimaginal exuviae of mesonotum; 29–30, hind wing in falling and transmitted light. Abbreviations: acz, achromozone; cz.a, anterior chromozone; cz.l-p, latero-posterior chromozone; cz.s, chromozonal suture; LPs, lateroparapsidal suture; MPs, medioparapsidal suture; MS, medioscutum; SMS, submedioscutum." captionText-6="FIGURES 31–39. Teloganodes (Dudgeodes) celebensis. 31, male imago; 32, abdomen of male imago; 33, head and thorax of female imago; 34, fore leg of female imago; 35–36, fore legs of male imagines; 37, middle leg of male imago; 38–39, exuviae of fore and middle legs of male subimago (32–39 with equal magnification)." captionText-7="FIGURES 40–45. Teloganodes (Dudgeodes) celebensis. 40–41, penis of male imago, dorsal view; 42, genitalia, ventral view; 43, cuticle of abdominal sternum IX and protopenis of male larva; 44, developing subimaginal penis extracted from the same larva; 45, position of developing penis in larval cuticle of protopenis." captionText-8="FIGURES 46–49. Teloganodes (Dudgeodes) celebensis, eggs." captionText-9="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.7673039" figureDoi-1="http://doi.org/10.5281/zenodo.7673041" figureDoi-10="http://doi.org/10.5281/zenodo.7673063" figureDoi-11="http://doi.org/10.5281/zenodo.7673065" figureDoi-12="http://doi.org/10.5281/zenodo.7673067" figureDoi-13="http://doi.org/10.5281/zenodo.7673069" figureDoi-14="http://doi.org/10.5281/zenodo.7673071" figureDoi-15="http://doi.org/10.5281/zenodo.7673077" figureDoi-16="http://doi.org/10.5281/zenodo.7673083" figureDoi-2="http://doi.org/10.5281/zenodo.7673045" figureDoi-3="http://doi.org/10.5281/zenodo.7673047" figureDoi-4="http://doi.org/10.5281/zenodo.7673049" figureDoi-5="http://doi.org/10.5281/zenodo.7673051" figureDoi-6="http://doi.org/10.5281/zenodo.7673055" figureDoi-7="http://doi.org/10.5281/zenodo.7673057" figureDoi-8="http://doi.org/10.5281/zenodo.7673059" figureDoi-9="http://doi.org/10.5281/zenodo.7673061" httpUri-0="https://zenodo.org/record/7673039/files/figure.png" httpUri-1="https://zenodo.org/record/7673041/files/figure.png" httpUri-10="https://zenodo.org/record/7673063/files/figure.png" httpUri-11="https://zenodo.org/record/7673065/files/figure.png" httpUri-12="https://zenodo.org/record/7673067/files/figure.png" httpUri-13="https://zenodo.org/record/7673069/files/figure.png" httpUri-14="https://zenodo.org/record/7673071/files/figure.png" httpUri-15="https://zenodo.org/record/7673077/files/figure.png" httpUri-16="https://zenodo.org/record/7673083/files/figure.png" httpUri-2="https://zenodo.org/record/7673045/files/figure.png" httpUri-3="https://zenodo.org/record/7673047/files/figure.png" httpUri-4="https://zenodo.org/record/7673049/files/figure.png" httpUri-5="https://zenodo.org/record/7673051/files/figure.png" httpUri-6="https://zenodo.org/record/7673055/files/figure.png" httpUri-7="https://zenodo.org/record/7673057/files/figure.png" httpUri-8="https://zenodo.org/record/7673059/files/figure.png" httpUri-9="https://zenodo.org/record/7673061/files/figure.png" pageId="2" pageNumber="529">Figs 1–101</figureCitation>
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Genus 
<treatmentCitation author="Eaton, A. E." box="[222,523,507,531]" page="208" pageId="2" pageNumber="529" year="1882">
<taxonomicName ID-CoL="7TKG" authority="Eaton 1882: 208" authorityName="Eaton" authorityPageNumber="208" authorityYear="1882" box="[222,523,507,531]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[222,349,507,531]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
<bibRefCitation author="Eaton, A. E." box="[356,523,507,531]" pageId="2" pageNumber="529" pagination="207 - 208" refId="ref10667" refString="Eaton, A. E. (1882) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 18, 207 - 208." type="journal article" year="1882">Eaton 1882: 208</bibRefCitation>
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(imago); 
<bibRefCitation author="Eaton, A. E." box="[623,739,507,531]" pageId="2" pageNumber="529" pagination="1 - 352" refId="ref10695" refString="Eaton, A. E. (1883 - 1888) A revisional monograph of recent Ephemeridae or mayflies. Transactions of the Linnean Society of London, Series 2, 3, 1 - 352, pls. 1 - 65. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 01550 a. x" type="journal article" year="1883">Eaton 1883</bibRefCitation>
–1888 (imago); 
<treatmentCitation author="Ulmer, G." box="[902,1076,507,531]" page="512" pageId="2" pageNumber="529" year="1939">
<bibRefCitation author="Ulmer, G." box="[902,1076,507,531]" pageId="2" pageNumber="529" pagination="443 - 692" refId="ref11924" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692, figs. 1 - 469, 4 tab." type="journal article" year="1939">Ulmer 1939: 512</bibRefCitation>
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(imago), 627 (larva). Teloganodes/fg1: 
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<bibRefCitation author="Kluge, N. J." box="[333,503,538,562]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 320</bibRefCitation>
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(larva, subimago, imago).
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<taxonomicName authority="Wang" authorityName="Wang" authorityYear="1996" box="[151,353,568,593]" class="Insecta" family="Teloganodidae" genus="Macafertiella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[151,289,568,592]" italics="true" pageId="2" pageNumber="529">Macafertiella</emphasis>
Wang
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in 
<treatmentCitation author="Wang, T. - Q. &amp; McCafferty, W. P." box="[385,689,568,593]" page="15" pageId="2" pageNumber="529" year="1996">
<bibRefCitation author="Wang, T. - Q. &amp; McCafferty, W. P." box="[385,689,568,593]" pageId="2" pageNumber="529" pagination="15 - 18" refId="ref11959" refString="Wang, T. - Q. &amp; McCafferty, W. P. (1996) New genus of Teloganodinae (Ephemeroptera: Pannota: Ephemerellidae) from Sri Lanka. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 132, 15 - 18." type="journal article" year="1996">Wang &amp; McCafferty 1996: 15</bibRefCitation>
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(synonymized by 
<bibRefCitation author="Sartori, M. &amp; Peters, J. G. &amp; Hubbard, M. D." box="[877,1203,568,593]" pageId="2" pageNumber="529" pagination="1 - 51" refId="ref11621" refString="Sartori, M., Peters, J. G. &amp; Hubbard, M. D. (2008) A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea). Zootaxa, 1957 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1957.1.1" type="journal article" year="2008">Sartori, Peters &amp; Hubbard 2008</bibRefCitation>
).
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<taxonomicName ID-CoL="48MR" authority="Sartori" authorityName=": Sartori, Peters &amp; Hubbard" authorityYear="2008" box="[151,340,599,623]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[151,265,599,623]" italics="true" pageId="2" pageNumber="529">Dudgeodes</emphasis>
Sartori
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in 
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, 
<emphasis bold="true" box="[746,815,599,623]" pageId="2" pageNumber="529">
<taxonomicNameLabel box="[746,815,599,623]" pageId="2" pageNumber="529">syn. n.</taxonomicNameLabel>
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<emphasis box="[151,261,630,654]" italics="true" pageId="2" pageNumber="529">Derlethina</emphasis>
Sartori
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in 
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, 
<emphasis bold="true" box="[742,811,630,654]" pageId="2" pageNumber="529">
<taxonomicNameLabel box="[742,811,630,654]" pageId="2" pageNumber="529">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="529" type="type_taxon">
<paragraph blockId="2.[151,1437,700,1699]" box="[151,590,700,727]" pageId="2" pageNumber="529">
<emphasis bold="true" box="[151,309,700,726]" pageId="2" pageNumber="529">
<typeStatus box="[151,210,700,726]" pageId="2" pageNumber="529">Type</typeStatus>
species:
</emphasis>
<taxonomicName authority="Hagen 1858" authorityName="Hagen" authorityYear="1858" box="[316,585,700,727]" class="Insecta" family="Baetidae" genus="Cloe" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="tristis">
<emphasis box="[316,439,700,726]" italics="true" pageId="2" pageNumber="529">Cloe tristis</emphasis>
<bibRefCitation author="Hagen, H." box="[446,585,700,727]" pageId="2" pageNumber="529" pagination="471 - 488" refId="ref10893" refString="Hagen, H. (1858) Synopsis der Neuroptera Ceylons. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 8, 471 - 488." type="journal article" year="1858">Hagen 1858</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[151,1437,700,1699]" pageId="2" pageNumber="529">
<emphasis bold="true" box="[199,441,736,762]" pageId="2" pageNumber="529">Systematic position.</emphasis>
<taxonomicName authorityName="Eaton" authorityYear="1882" box="[451,592,736,762]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[451,592,736,762]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
</taxonomicName>
belongs to the taxon 
<emphasis bold="true" box="[848,1045,736,762]" pageId="2" pageNumber="529">Ephemerella/fg1</emphasis>
, or superfamily 
<taxonomicName box="[1240,1433,736,762]" pageId="2" pageNumber="529" rank="superFamily" superFamily="Ephemerelloidea">Ephemerelloidea</taxonomicName>
, which is characterized by peculiar structure of tergalii with lamellate dorsal lobe and two-branched ventral lobe bearing processes on outer sides of its branches (character 7 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." box="[1131,1329,808,834]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 294</bibRefCitation>
) (
<figureCitation box="[1355,1429,808,834]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1428]" captionTargetBox="[151,1436,455,1358]" captionTargetId="figure-141@1.[151,1436,450,1378]" captionTargetPageId="1" captionText="FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004)." figureDoi="http://doi.org/10.5281/zenodo.7673039" httpUri="https://zenodo.org/record/7673039/files/figure.png" pageId="2" pageNumber="529">Figs 1</figureCitation>
, 
<figureCitation box="[151,225,844,871]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="2" pageNumber="529">50–57</figureCitation>
), presence of crossvein 
<emphasis box="[507,597,845,870]" italics="true" pageId="2" pageNumber="529">cua-cup</emphasis>
near base of fore wing (character 6 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 293</bibRefCitation>
) (
<figureCitation box="[218,301,880,906]" captionStart="FIGURES 93–94" captionStartId="22.[151,264,1218,1242]" captionTargetBox="[151,1435,186,1189]" captionTargetId="figure-23@22.[151,1436,181,1193]" captionTargetPageId="22" captionText="FIGURES 93–94. Teloganodes (Dudgeodes) rasus sp. n., wings of male imago (holotype). 93, in falling light; 94, in transmitted light (arrow shows constant crossvein cua-cup)." figureDoi="http://doi.org/10.5281/zenodo.7673077" httpUri="https://zenodo.org/record/7673077/files/figure.png" pageId="2" pageNumber="529">Fig. 94</figureCitation>
), certain reductions in larval maxillae and labium (characters 1–3 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 293</bibRefCitation>
), peculiarities of larval thorax (characters 4–5 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." box="[975,1168,916,942]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 293</bibRefCitation>
), loss of 4th segment of gonostylus (character 8 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." box="[665,852,952,978]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 294</bibRefCitation>
) (
<figureCitation box="[873,960,952,978]" captionStart="FIGURES 40–45" captionStartId="14.[151,264,1934,1958]" captionTargetBox="[204,1383,634,1911]" captionTargetId="figure-236@14.[204,1383,633,1911]" captionTargetPageId="14" captionText="FIGURES 40–45. Teloganodes (Dudgeodes) celebensis. 40–41, penis of male imago, dorsal view; 42, genitalia, ventral view; 43, cuticle of abdominal sternum IX and protopenis of male larva; 44, developing subimaginal penis extracted from the same larva; 45, position of developing penis in larval cuticle of protopenis." figureDoi="http://doi.org/10.5281/zenodo.7673057" httpUri="https://zenodo.org/record/7673057/files/figure.png" pageId="2" pageNumber="529">Figs 42</figureCitation>
, 
<figureCitation box="[970,1001,952,978]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="2" pageNumber="529">97</figureCitation>
), and some other characters (characters 9–19 of Ephemerella/fg 
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<bibRefCitation author="Kluge, N. J." box="[467,716,988,1015]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: 294–295</bibRefCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,700,1699]" pageId="2" pageNumber="529">
The holophyletic taxon Ephemerella/fg1 is divided into two holophyletic taxa, Ephemerella/fg2 (or family 
<taxonomicName authorityName="Klapalek" authorityYear="1909" box="[151,329,1060,1086]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
) and Pantricorythi Kluge 2014, or 
<taxonomicName authorityName="Eaton" authorityYear="1868" box="[719,853,1060,1086]" class="Insecta" family="Tricorythidae" genus="Tricorythus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">Tricorythus</taxonomicName>
/fg1 (usually treated as two or more families).
</paragraph>
<paragraph blockId="2.[151,1437,700,1699]" pageId="2" pageNumber="529">
The taxon 
<emphasis bold="true" box="[317,479,1096,1122]" pageId="2" pageNumber="529">Pantricorythi</emphasis>
is characterized by retention of fully developed tergalii on the 2nd abdominal segment (
<figureCitation box="[159,231,1132,1158]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1428]" captionTargetBox="[151,1436,455,1358]" captionTargetId="figure-141@1.[151,1436,450,1378]" captionTargetPageId="1" captionText="FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004)." figureDoi="http://doi.org/10.5281/zenodo.7673039" httpUri="https://zenodo.org/record/7673039/files/figure.png" pageId="2" pageNumber="529">Figs 1</figureCitation>
, 
<figureCitation box="[247,278,1132,1158]" captionStart="FIGURES 61–62" captionStartId="17.[151,264,1223,1247]" captionTargetBox="[159,1419,181,1200]" captionTargetId="figure-15@17.[159,1428,181,1200]" captionTargetPageId="17" captionText="FIGURES 61–62. Teloganodes (Dudgeodes) rasus sp. n. 61, male larva of last instar; 62, younger female larva." figureDoi="http://doi.org/10.5281/zenodo.7673063" httpUri="https://zenodo.org/record/7673063/files/figure.png" pageId="2" pageNumber="529">62</figureCitation>
), long 1st segment of the 3-segmented gonostyli (
<figureCitation box="[847,936,1132,1158]" captionStart="FIGURES 40–45" captionStartId="14.[151,264,1934,1958]" captionTargetBox="[204,1383,634,1911]" captionTargetId="figure-236@14.[204,1383,633,1911]" captionTargetPageId="14" captionText="FIGURES 40–45. Teloganodes (Dudgeodes) celebensis. 40–41, penis of male imago, dorsal view; 42, genitalia, ventral view; 43, cuticle of abdominal sternum IX and protopenis of male larva; 44, developing subimaginal penis extracted from the same larva; 45, position of developing penis in larval cuticle of protopenis." figureDoi="http://doi.org/10.5281/zenodo.7673057" httpUri="https://zenodo.org/record/7673057/files/figure.png" pageId="2" pageNumber="529">Figs 42</figureCitation>
, 
<figureCitation box="[949,978,1132,1158]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="2" pageNumber="529">97</figureCitation>
) and peculiar arrangement of setal rows on anterior (dorsal) side of larval fore femur, which have a regular row crossing the femur near its middle and continuing distally along the outer margin of the femur (
<figureCitation box="[796,930,1204,1230]" captionStart="FIGURES 19–26" captionStartId="11.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-15@11.[151,1436,181,1817]" captionTargetPageId="11" captionText="FIGURES 19–26. Teloganodes (Dudgeodes) celebensis, exuviae of larval legs. 19–20, fore leg and its fragment; 21, middle leg; 22–26, hind leg and its fragments." figureDoi="http://doi.org/10.5281/zenodo.7673049" httpUri="https://zenodo.org/record/7673049/files/figure.png" pageId="2" pageNumber="529">Figs 19–20</figureCitation>
). Among 
<taxonomicName authorityName="Eaton" authorityYear="1882" box="[1041,1182,1204,1230]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis box="[1041,1182,1204,1230]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1190,1232,1204,1231]" pageId="2" pageNumber="529" sensu="lato">s. l.</taxonomicNameLabel>
, this arrangement of setal rows occurs in most species; in the known species of subgenus 
<taxonomicName authorityName="Eaton" authorityYear="1882" box="[952,1093,1240,1266]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis box="[952,1093,1240,1266]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1100,1161,1241,1267]" pageId="2" pageNumber="529" sensu="stricto">s. str.</taxonomicNameLabel>
, the transverse setal row of fore femur is either secondarily lost (
<bibRefCitation author="Sartori, M. &amp; Peters, J. G. &amp; Hubbard, M. D." box="[603,814,1276,1302]" pageId="2" pageNumber="529" pagination="1 - 51" refId="ref11621" refString="Sartori, M., Peters, J. G. &amp; Hubbard, M. D. (2008) A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea). Zootaxa, 1957 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1957.1.1" type="journal article" year="2008">
Sartori 
<emphasis box="[689,746,1276,1302]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
2008
</bibRefCitation>
: fig. 59–60) or detached from the row on outer margin (
<bibRefCitation author="Selvakumar, C. &amp; Sivaramakrishnan, K. G. &amp; Jacobus, L. M. &amp; Janarthanan, S. &amp; Arumugam, M." box="[159,424,1312,1338]" pageId="2" pageNumber="529" pagination="87 - 104" refId="ref11678" refString="Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, L. M., Janarthanan, S. &amp; Arumugam, M. (2014) Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India. Zootaxa, 3846 (1), 87 - 104. https: // doi. org / 10.11646 / zootaxa. 3846.1.4" type="journal article" year="2014">
Selvakumar 
<emphasis box="[301,358,1312,1338]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
2014
</bibRefCitation>
: fig. 50).
</paragraph>
<paragraph blockId="2.[151,1437,700,1699]" pageId="2" pageNumber="529">
The taxon Pantricorythi contains 
<taxonomicName authority="Tshernova 1972" authorityName="Tshernova" authorityYear="1972" box="[565,883,1348,1374]" class="Insecta" family="Vietnamellidae" genus="Vietnamella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[565,698,1348,1374]" italics="true" pageId="2" pageNumber="529">Vietnamella</emphasis>
<bibRefCitation author="Tshernova, O. A." box="[703,883,1348,1374]" pageId="2" pageNumber="529" pagination="604 - 614" refId="ref11816" refString="Tshernova, O. A. (1972) Some new species of may-flies from Asia (Ephemeroptera, Heptageniidae, Ephemerellidae). Entomological Review, 51 (3), 364 - 369. [English translation from Entomologicheskoe Obozrenie, 51 (3), 604 - 614]" type="journal article" year="1972">Tshernova 1972</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Riek 1963" authorityName="Riek" authorityYear="1963" box="[892,1171,1348,1374]" class="Insecta" family="Austremerellidae" genus="Austremerella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[892,1049,1348,1374]" italics="true" pageId="2" pageNumber="529">Austremerella</emphasis>
<bibRefCitation author="Riek, E. F." box="[1054,1171,1348,1374]" pageId="2" pageNumber="529" pagination="48 - 50" refId="ref11568" refString="Riek, E. F. (1963) An Australian mayfly of the family Ephemerellidae (Ephemeroptera). Journal of the Entomological Society of Queensland, 2, 48 - 50. https: // doi. org / 10.1111 / j. 1440 - 6055.1963. tb 00387. x" type="journal article" year="1963">Riek 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Lestage 1924" authorityName="Lestage" authorityYear="1924" class="Insecta" family="Teloganodidae" genus="Ephemerellina" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1180,1344,1348,1374]" italics="true" pageId="2" pageNumber="529">Ephemerellina</emphasis>
<bibRefCitation author="Lestage, J. - A." pageId="2" pageNumber="529" pagination="316 - 352" refId="ref11261" refString="Lestage, J. - A. (1924) Les Ephemeres de l'Afrique du Sud. Catalogue critique &amp; systematique des especes connues et description de trois genera nouveaux et de sept especes nouvelles. Revue Zoologique Africaine, 12, 316 - 352." type="journal article" year="1924">Lestage 1924</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Barnard 1932" authorityName="Barnard" authorityYear="1932" box="[221,506,1384,1410]" class="Insecta" family="Teloganodidae" genus="Lithogloea" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[221,342,1384,1410]" italics="true" pageId="2" pageNumber="529">Lithogloea</emphasis>
<bibRefCitation author="Barnard, K. H." box="[350,506,1384,1410]" pageId="2" pageNumber="529" pagination="201 - 259" refId="ref10492" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Demoulin 1970" authorityName="Demoulin" authorityYear="1970" box="[517,819,1384,1410]" class="Insecta" family="Teloganodidae" genus="Lestagella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[517,634,1384,1410]" italics="true" pageId="2" pageNumber="529">Lestagella</emphasis>
<bibRefCitation author="Demoulin, G." box="[641,819,1384,1410]" pageId="2" pageNumber="529" pagination="24 - 170" refId="ref10607" refString="Demoulin, G. (1970) Ephemeroptera des faunes ethiopienne et malgache. In: Hanstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life (Results of the Lunds University Expedition in 1950 - 1951), 14, pp. 24 - 170." type="journal article" year="1970">Demoulin 1970</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="McCafferty &amp; Wang 1998" authorityName="McCafferty &amp; Wang" authorityYear="1998" box="[830,1263,1384,1411]" class="Insecta" family="Teloganodidae" genus="Nadinetella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[830,961,1384,1410]" italics="true" pageId="2" pageNumber="529">Nadinetella</emphasis>
<bibRefCitation author="McCafferty, W. P. &amp; Wang, T. - Q." box="[967,1263,1384,1411]" pageId="2" pageNumber="529" pagination="344" refId="ref11492" refString="McCafferty, W. P. &amp; Wang, T. - Q. (1998) New name for a generic homonym in Teloganodinae (Ephemeroptera). Entomological News, 109 (5), 344." type="journal article" year="1998">McCafferty &amp; Wang 1998</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Allen 1973" authorityName="Allen" authorityYear="1973" class="Insecta" family="Teloganodidae" genus="Manohyphella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1274,1436,1384,1410]" italics="true" pageId="2" pageNumber="529">Manohyphella</emphasis>
<bibRefCitation author="Allen, R. K." box="[151,277,1420,1446]" pageId="2" pageNumber="529" pagination="160 - 164" refId="ref10392" refString="Allen, R. K. (1973) New Ephemerellidae from Madagascar and Afghanistan (Ephemeroptera). Pan-Pacific Entomologist, 49 (2), 160 - 164." type="journal article" year="1973">Allen 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Selvakumar et al. 2014" authorityName="Selvakumar" authorityYear="2014" box="[286,691,1420,1446]" class="Insecta" family="Teloganodidae" genus="Indoganodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[286,431,1420,1446]" italics="true" pageId="2" pageNumber="529">Indoganodes</emphasis>
<bibRefCitation author="Selvakumar, C. &amp; Sivaramakrishnan, K. G. &amp; Jacobus, L. M. &amp; Janarthanan, S. &amp; Arumugam, M." box="[435,691,1420,1446]" pageId="2" pageNumber="529" pagination="87 - 104" refId="ref11678" refString="Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, L. M., Janarthanan, S. &amp; Arumugam, M. (2014) Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India. Zootaxa, 3846 (1), 87 - 104. https: // doi. org / 10.11646 / zootaxa. 3846.1.4" type="journal article" year="2014">
Selvakumar 
<emphasis box="[573,627,1420,1446]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
2014
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Ulmer 1920" authorityName="Ulmer" authorityYear="1920" box="[699,1007,1420,1446]" class="Insecta" family="Melanemerellidae" genus="Melanemerella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[699,867,1420,1446]" italics="true" pageId="2" pageNumber="529">Melanemerella</emphasis>
<bibRefCitation author="Ulmer, G." box="[872,1007,1420,1446]" pageId="2" pageNumber="529" pagination="1 - 80" refId="ref11868" refString="Ulmer, G. (1920) Neue Ephemeropteren. Archiv fur Naturgeschichte, (A), 85 (11), 1 - 80. [1919]" type="journal article" year="1920">Ulmer 1920</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Ulmer 1939" authorityName="Ulmer" authorityYear="1939" box="[1016,1287,1420,1446]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1016,1147,1420,1446]" italics="true" pageId="2" pageNumber="529">Teloganella</emphasis>
<bibRefCitation author="Ulmer, G." box="[1152,1287,1420,1446]" pageId="2" pageNumber="529" pagination="443 - 692" refId="ref11924" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692, figs. 1 - 469, 4 tab." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Eaton 1882" authorityName="Eaton" authorityYear="1882" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1296,1437,1420,1446]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
<bibRefCitation author="Eaton, A. E." box="[151,282,1456,1483]" pageId="2" pageNumber="529" pagination="207 - 208" refId="ref10667" refString="Eaton, A. E. (1882) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 18, 207 - 208." type="journal article" year="1882">Eaton 1882</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Gillies 1960" authorityName="Gillies" authorityYear="1960" box="[292,599,1456,1482]" class="Insecta" family="Ephemerythidae" genus="Ephemerythus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[292,452,1456,1482]" italics="true" pageId="2" pageNumber="529">Ephemerythus</emphasis>
<bibRefCitation author="Gillies, M. T." box="[459,599,1456,1482]" pageId="2" pageNumber="529" pagination="35 - 40" refId="ref10829" refString="Gillies, M. T. (1960) A new genus of Tricorythidae (Ephemeroptera) from East Africa. Proceedings of the Royal Entomological Society of London, (B), 29 (3 - 4), 35 - 40. https: // doi. org / 10.1111 / j. 1365 - 3113.1960. tb 01135. x" type="journal article" year="1960">Gillies 1960</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Demoulin 1954" authorityName="Demoulin" authorityYear="1954" box="[609,953,1456,1483]" class="Insecta" family="Dicercomyzidae" genus="Dicercomyzon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[609,770,1457,1482]" italics="true" pageId="2" pageNumber="529">Dicercomyzon</emphasis>
<bibRefCitation author="Demoulin, G." box="[776,953,1456,1483]" pageId="2" pageNumber="529" pagination="1 - 4" refId="ref10533" refString="Demoulin, G. (1954) Description preliminaire d'un type larvaire nouveau d'Ephemeropteres Tricorythidae du Congo Belge. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 3 (6), 1 - 4." type="journal article" year="1954">Demoulin 1954</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Demoulin 1959" authorityName="Demoulin" authorityYear="1959" box="[964,1323,1456,1483]" class="Insecta" family="Machadorythidae" genus="Machadorythus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[964,1140,1456,1482]" italics="true" pageId="2" pageNumber="529">Machadorythus</emphasis>
<bibRefCitation author="Demoulin, G." box="[1146,1323,1456,1483]" pageId="2" pageNumber="529" pagination="249 - 252" refId="ref10571" refString="Demoulin, G. (1959) Une curieuse larve d'Ephemeropteres de l'Angola portugais. Bulletin et Annales de la Societe Entomologique de Belgique, 95 (7 - 8), 249 - 252." type="journal article" year="1959">Demoulin 1959</bibRefCitation>
</taxonomicName>
, the taxon with the non-typified circumscriptional name Tricorygnatha 
<bibRefCitation author="Kluge, N. J." box="[850,984,1492,1518]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
and the large taxon with the rank-free hierarchical name Leptohyphes/fg1 (
<bibRefCitation author="Kluge, N. J." box="[562,701,1528,1554]" pageId="2" pageNumber="529" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,700,1699]" pageId="2" pageNumber="529">
Various authors either arrange these taxa among the families 
<taxonomicName authority="Lestage 1942" authorityName="Lestage" authorityYear="1942" box="[893,1204,1564,1591]" class="Insecta" family="Tricorythidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="family">
Tricorythidae 
<bibRefCitation author="Lestage, J. - A." box="[1052,1204,1564,1591]" pageId="2" pageNumber="529" pagination="1 - 20" refId="ref11307" refString="Lestage, J. - A. (1942) Contribution a l'etude des Ephemeropteres. XXV. Notes critiques sur les anciens Caenidiens d'Afrique et sur l'independance de l'evolution tricorythido-caenidienne. Bulletin du Musee Roual d'Histoire Naturelle de Belgique, 18 (48), 1 - 20." type="journal article" year="1942">Lestage 1942</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Klapalek 1909" authorityName="Klapalek" authorityYear="1909" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="family">
Ephemerellidae 
<bibRefCitation author="Klapalek, F." box="[151,319,1600,1626]" pageId="2" pageNumber="529" pagination="33 - 95" refId="ref10918" refString="Klapalek, F. (1909) I. Ephemerida, Eintagsfliegen. In: Brauer, A. (Ed.), Die Susswasserfauna Deutschlands. Bd. 8. Verlag von Gustav Fischer, Jena, pp. 1 - 32 + 33 - 95." type="book chapter" year="1909">Klapálek 1909</bibRefCitation>
</taxonomicName>
, or accept greater number of families. In the past, the genus 
<taxonomicName authorityName="Eaton" authorityYear="1882" box="[1009,1150,1600,1626]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1009,1150,1600,1626]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
</taxonomicName>
was placed in the family 
<taxonomicName authorityName="Klapalek" authorityYear="1909" box="[151,330,1636,1662]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
, which was paraphyletic in such sense. Currently, it is used to place 
<taxonomicName authorityName="Eaton" authorityYear="1882" box="[1154,1295,1636,1662]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis box="[1154,1295,1636,1662]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
</taxonomicName>
in a family 
<taxonomicName authority="Allen 1965" authorityName="Allen" authorityYear="1965" box="[151,449,1672,1699]" class="Insecta" family="Teloganodidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="family">
Teloganodidae 
<bibRefCitation author="Allen, R. K." box="[321,449,1672,1699]" pageId="2" pageNumber="529" pagination="262 - 266" refId="ref10356" refString="Allen, R. K. (1965) A review of the subfamilies of Ephemerellidae (Ephemeroptera). Journal of the Kansas Entomological Society, 38 (3), 262 - 266." type="journal article" year="1965">Allen 1965</bibRefCitation>
</taxonomicName>
, which has no generally accepted circumscription and diagnosis.
</paragraph>
</subSubSection>
</treatment>
</document>