Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae) Author Kluge, Nikita text Zootaxa 2023 2023-02-23 5244 6 527 552 http://dx.doi.org/10.11646/zootaxa.5244.6.2 journal article 10.11646/zootaxa.5244.6.2 57e07918-7ed2-4f62-a052-82d911391af7 1175-5326 7673035 8182831B-25FC-4B92-B319-281659660565 Genus Teloganodes Eaton 1882 ( Figs 1–101 ) Genus Teloganodes Eaton 1882: 208 (imago); Eaton 1883 –1888 (imago); Ulmer 1939: 512 (imago), 627 (larva). Teloganodes/fg1: Kluge 2004: 320 (larva, subimago, imago). Macafertiella Wang in Wang & McCafferty 1996: 15 (synonymized by Sartori, Peters & Hubbard 2008 ). Dudgeodes Sartori in Sartori, Peters & Hubbard 2008: 25 , syn. n. Derlethina Sartori in Sartori, Peters & Hubbard 2008: 44 , syn. n. Type species: Cloe tristis Hagen 1858 . Systematic position. Teloganodes belongs to the taxon Ephemerella/fg1 , or superfamily Ephemerelloidea , which is characterized by peculiar structure of tergalii with lamellate dorsal lobe and two-branched ventral lobe bearing processes on outer sides of its branches (character 7 of Ephemerella/fg 1 in Kluge 2004: 294 ) ( Figs 1 , 50–57 ), presence of crossvein cua-cup near base of fore wing (character 6 of Ephemerella/fg 1 in Kluge 2004: 293 ) ( Fig. 94 ), certain reductions in larval maxillae and labium (characters 1–3 of Ephemerella/fg 1 in Kluge 2004: 293 ), peculiarities of larval thorax (characters 4–5 of Ephemerella/fg 1 in Kluge 2004: 293 ), loss of 4th segment of gonostylus (character 8 of Ephemerella/fg 1 in Kluge 2004: 294 ) ( Figs 42 , 97 ), and some other characters (characters 9–19 of Ephemerella/fg 1 in Kluge 2004: 294–295 ). The holophyletic taxon Ephemerella/fg1 is divided into two holophyletic taxa, Ephemerella/fg2 (or family Ephemerellidae ) and Pantricorythi Kluge 2014, or Tricorythus /fg1 (usually treated as two or more families). The taxon Pantricorythi is characterized by retention of fully developed tergalii on the 2nd abdominal segment ( Figs 1 , 62 ), long 1st segment of the 3-segmented gonostyli ( Figs 42 , 97 ) and peculiar arrangement of setal rows on anterior (dorsal) side of larval fore femur, which have a regular row crossing the femur near its middle and continuing distally along the outer margin of the femur ( Figs 19–20 ). Among Teloganodes s. l. , this arrangement of setal rows occurs in most species; in the known species of subgenus Teloganodes s. str. , the transverse setal row of fore femur is either secondarily lost ( Sartori et al. 2008 : fig. 59–60) or detached from the row on outer margin ( Selvakumar et al. 2014 : fig. 50). The taxon Pantricorythi contains Vietnamella Tshernova 1972 , Austremerella Riek 1963 , Ephemerellina Lestage 1924 , Lithogloea Barnard 1932 , Lestagella Demoulin 1970 , Nadinetella McCafferty & Wang 1998 , Manohyphella Allen 1973 , Indoganodes Selvakumar et al. 2014 , Melanemerella Ulmer 1920 , Teloganella Ulmer 1939 , Teloganodes Eaton 1882 , Ephemerythus Gillies 1960 , Dicercomyzon Demoulin 1954 , Machadorythus Demoulin 1959 , the taxon with the non-typified circumscriptional name Tricorygnatha Kluge 2004 and the large taxon with the rank-free hierarchical name Leptohyphes/fg1 ( Kluge 2004 ). Various authors either arrange these taxa among the families Tricorythidae Lestage 1942 and Ephemerellidae Klapálek 1909 , or accept greater number of families. In the past, the genus Teloganodes was placed in the family Ephemerellidae , which was paraphyletic in such sense. Currently, it is used to place Teloganodes in a family Teloganodidae Allen 1965 , which has no generally accepted circumscription and diagnosis.