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<mods:title id="4C797A6F91EA5265BC3C94A043C95508">Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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<mods:namePart id="E163352C58DC48920300E89DED39F680">Pall-Gergely, Barna</mods:namePart>
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<mods:namePart id="E7B7AB21CD6E216E1951DE928B566D32">Sajan, Sheikh</mods:namePart>
<mods:affiliation id="F490B23F1F1E965DF2999069E0F2D392">Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
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<mods:namePart id="AF80283E9A44EAADB326C4C2558EBF02">Tripathy, Basudev</mods:namePart>
<mods:affiliation id="9DE82FC5F7A74FD9870F26FCABF9B65D">Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:namePart id="7300C87F858960FD703409CB4E01ECC9">Meng, Kaibaryer</mods:namePart>
<mods:affiliation id="144C92E120E8A6063EB32C5FB74DADA2">National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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<mods:affiliation id="1AB0E2198924E79C17FA44F3BE6516C1">Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
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<mods:namePart id="891D9D8F8ED73F59C5B6E51277BC8D5A">Ablett, Jonathan D.</mods:namePart>
<mods:affiliation id="A842ADBCECA84735F7F32159231D407D">Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
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<treatment id="6BF61B61D6F359F3863DE41A069DC49A" ID-GBIF-Taxon="168715069" LSID="urn:lsid:plazi:treatment:6BF61B61D6F359F3863DE41A069DC49A" httpUri="http://treatment.plazi.org/id/6BF61B61D6F359F3863DE41A069DC49A" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="875A74DE7CB84CCC27DCBF2F2D28EB2A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="24BDFD3AF5A21AEB9CF8D1138C4DFAFD" pageId="0" pageNumber="1">
Genus
<taxonomicName id="824CE696461A179565C5049D8AF92169" LSID="6BF61B61-D6F3-59F3-863D-E41A069DC49A" authority="Moellendorff, 1897" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
Chamalycaeus
<normalizedToken id="50DCEAA1AA1565809664F9CF30FCB8BB" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897
</taxonomicName>
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<subSubSection id="12D378F102586FAF5C739FEC03A8736E" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="31D9B9A40C419D9E45724B4C923C532A" pageId="0" pageNumber="1">
<taxonomicName id="4DA7B92423A3FFE937CCDC25A73ECBE7" authority="" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus)" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus)</taxonomicName>
<normalizedToken id="A6503D2AC740F89DE38AEE912516E191" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897b: 93.
</paragraph>
<paragraph id="DF14FAA8945860291343CB03315E957D" pageId="0" pageNumber="1">
<taxonomicName id="630B4463A9FDFFBB701DE88968A42FD0" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Chamalycaeus</taxonomicName>
-
<bibRefCitation id="A82399BAEC5EC15A4FCD71EEF925B01F" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken id="7EFCA12864F55441A97DD1616D1494BD" originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 148;
<bibRefCitation id="2982374BECCEE426FD9D0EE3F39763DB" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="A5AE116E625FBD2A03606317590CEAE8" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 5-7.
</paragraph>
<paragraph id="677E7D0423E6051FCB96E48D322398B9" pageId="0" pageNumber="1">
<taxonomicName id="051D035B2D6882BEBFE00AF315D95D7B" authority="(Chamalycaeus)" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Chamalycaeus)" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Chamalycaeus">Chamalycaeus (Chamalycaeus)</taxonomicName>
-
<bibRefCitation id="4663704828653C27395F8682AEA46024" author="Thiele, J" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="1" refId="B213" refString="Thiele, J, 1929. Handbuch der systematischen Weichtierkunde, Teil 1. Verlag von Gustav Fischer, Jena" title="Handbuch der systematischen Weichtierkunde, Teil 1." year="1929">Thiele 1929</bibRefCitation>
: 107-108; Wenz 1938: 477-478;
<bibRefCitation id="ADC3F3AE3A5FB4DA597D249F087E364B" author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov 2013</bibRefCitation>
: 35.
</paragraph>
</subSubSection>
<subSubSection id="42F1D0C4671A360789642C4C277F4322" pageId="0" pageNumber="1" type="type species">
<paragraph id="3CD30C30A089A24D8ECFB37031D9DE81" pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph id="5A8EC9B0D357B2CDFBFFBD0D0D28A979" pageId="0" pageNumber="1">
<taxonomicName id="C2F5F15EC431BB854C3028C2F3563273" authorityName="Mollendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus) fruhstorferi</taxonomicName>
(Fig.
<figureCitation id="424C30C6DC1A3F7C9A0A016F5093D4DF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Type species of alycaeid genus-group taxa A Alycaeus eydouxi Venmans, 1956 (SMF 109290; type species of Alycaeus) B Chamalycaeus fruhstorferi (Moellendorff, 1897), lectotype (SMF 109481; type species of Chamalycaeus). Photographs: Barna Pall-Gergely (A) and Frank Walther (B)." figureDoi="10.3897/zookeys.981.53583.figure6" httpUri="https://binary.pensoft.net/fig/468651" pageId="0" pageNumber="1">6B</figureCitation>
) by monotypy, see also Remarks.
</paragraph>
</subSubSection>
<subSubSection id="E6B30899647EEE7E7A3A75047CB65203" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="4CAAFC2B07AE48031BA5D710015FD6AE" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="11E44A340F4B4350B14C66D0C1A6469A" pageId="0" pageNumber="1">Shell very small to medium sized (D: 2-5 mm), usually flattened, discoid or low triangular, protoconch smooth (or very finely pitted), elevated even if the spire is low; R1 usually roughly reticulated due to spiral striation and radial ribs (sometimes prominent); R2 from short to very long, with widely spaced, sharp, elevated ribs; R3 normally developed. Operculum usually thin, without notable outer structures. Radula is known for a single species (central tooth with five cusps, broad, central cusp pointed).</paragraph>
</subSubSection>
<subSubSection id="E8C2D40F31FFE90668D43CDEF2FBF185" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="6E9FA84741E5A9D20E4158F5AE8D3507" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="2D418B8311E80EC9B47663F71D1A8822" pageId="0" pageNumber="1">
See under
<taxonomicName id="0834D2EF0E070284663D6F52BF26A71B" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" superFamily="Cyclophoroidea">
<emphasis id="5777D0200A5D428CA92EF89ADCF60EAD" italics="true" pageId="0" pageNumber="1">Alycaeus</emphasis>
</taxonomicName>
and Table
<tableCitation id="B96467C5462F573076EB29325C10B135" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Important traits of Alycaeid genera." httpUri="http://table.plazi.org/id/19332E0DEA2DC936E472DDFDDAA5DCCC" pageId="0" pageNumber="1" tableUuid="19332E0DEA2DC936E472DDFDDAA5DCCC">3</tableCitation>
.
<taxonomicName id="E9F3EB6B1F32CC0388FB2DF589B219C8" authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="30B355D3C5B3B1C99F22745ED1E6DD17" italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
</taxonomicName>
species are identical, with the exception of the spirally striated protoconch.
</paragraph>
</subSubSection>
<subSubSection id="162072388DFB1DC3C2C1AD48A0CAB61C" pageId="0" pageNumber="1" type="distribution">
<paragraph id="31EA22A7609D9955325152D57DA7D662" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="3E491B9D9EFBFFCFAEFD22BF9A826ED6" pageId="0" pageNumber="1">
<taxonomicName id="141F4475F6EBB5F3A8E2A6D9A8935A62" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="835783C575389D225D295136A694A29D" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
is distributed from the southeastern Himalaya Region, the Malay Peninsula, Sumatra, Java, Borneo, Sulawesi, and the Philippine Palawan Island (Fig.
<figureCitation id="1906247049E808039893B95754F78963" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Chamalycaeus Moellendorff, 1897 (dark shaded area) and Metalycaeus Pilsbry, 1900 (light shaded area)." figureDoi="10.3897/zookeys.981.53583.figure9" httpUri="https://binary.pensoft.net/fig/468654" pageId="0" pageNumber="1">9</figureCitation>
).
</paragraph>
<caption id="62B8E7AA05D9882D050EA60836300E24" doi="10.3897/zookeys.981.53583.figure9" httpUri="https://binary.pensoft.net/fig/468654" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph id="896FA4B04F5F54818B1B6135B82BAA49" pageId="0" pageNumber="1">
<emphasis id="74A9481F3C75FF17813853F76D1EF10C" bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
Distribution of
<taxonomicName id="4B57CBED9462E2D3E080552067CFA336" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="FB099565AD2E651FC094E4F085270814" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
<normalizedToken id="6CA3A0C99212E67D86A6D56BC033AB7F" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897 (dark shaded area) and
<taxonomicName id="68CAE1564943C597918F7F35D21C03C8" authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="A1D189FA1021365ABB3B353206EB1617" italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
</taxonomicName>
Pilsbry, 1900 (light shaded area).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D59D3E2E02E5F0B469A21C8CCCB5997B" pageId="0" pageNumber="1" type="remarks">
<paragraph id="A9C35B6AC08EE7AA0FE2488122ABD343" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="8DE3FC36097A4E6377EFC020D5090F47" pageId="0" pageNumber="1">
<bibRefCitation id="0D053F00245D5FD5381270C9E790B9ED" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken id="7CB03BE63C41BED68DAD2617C4236AE4" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897)
</bibRefCitation>
listed the species of
<normalizedToken id="0B92F5A4740E030483297A438A903E5B" originalValue="“Pneumonopoma”">&quot;Pneumonopoma&quot;</normalizedToken>
, which included all members of the genus
<taxonomicName id="768BC057484672B2E0CCCA8796223961" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" superFamily="Cyclophoroidea">
<emphasis id="B341A11460D39D5A9CACC6137D1574CE" italics="true" pageId="0" pageNumber="1">Alycaeus</emphasis>
</taxonomicName>
. They introduced &quot;Subgenus
<emphasis id="567241075680586C29B875B2D03408B3" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
n.&quot; (
<bibRefCitation id="5543A1A67D92C3DD19571B5C1AAA087C" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken id="A3B243526381ADA43BC2EF1998C3B226" originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 148), indicating it as a new subgenus, and included 44 species within their new group. The subgenus
<taxonomicName id="E3CD1C7F57875B879A09B52D5BED4809" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Chamalycaeus)" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Chamalycaeus</taxonomicName>
, however, was previously mentioned in another paper in the same volume of the Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, in which
<bibRefCitation id="8B8C58C7C2884064739A68235FEAD595" author="Moellendorff, O von" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="89 - 97" publicationUrl="https://biodiversitylibrary.org/page/28228249" refId="B141" refString="Moellendorff, O von, 1897b. Neue Landschnecken von Java. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 89 - 97, https://biodiversitylibrary.org/page/28228249" title="Neue Landschnecken von Java." url="https://biodiversitylibrary.org/page/28228249" volume="29" year="1897 b">
<normalizedToken id="3729E3DD9CA16F3D8287A7692E715DB9" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897b
</bibRefCitation>
: 93) described
<taxonomicName id="C8DF2ED9EFB64B96A98183C4D381CC54" authorityName="Mollendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus) fruhstorferi</taxonomicName>
.
<bibRefCitation id="FD3BEFC270A76A445CDA05712480C8DE" author="Moellendorff, O von" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="89 - 97" publicationUrl="https://biodiversitylibrary.org/page/28228249" refId="B141" refString="Moellendorff, O von, 1897b. Neue Landschnecken von Java. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 89 - 97, https://biodiversitylibrary.org/page/28228249" title="Neue Landschnecken von Java." url="https://biodiversitylibrary.org/page/28228249" volume="29" year="1897 b">
<normalizedToken id="DF3C32814AD6804ADEBF86079D993E5D" originalValue="Möllendorffs">Moellendorff's</normalizedToken>
(1897b)
</bibRefCitation>
paper was published in July-August, whereas that of
<bibRefCitation id="B72390AD6F3B32066A62BB2F4C0E0F01" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken id="5A90E420F69DE5A6282821BBABFAE3F7" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897)
</bibRefCitation>
was published in September-October. Accordingly, the genus
<taxonomicName id="0C2ADC705050F829E71FAEAAFE4E4B36" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="97C0A916806810C01E54F8F6492340E6" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
was described by
<normalizedToken id="DDEA665A43A73663153683266A337A1B" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897) and its type species is
<taxonomicName id="F32A5AEC7DD550E584066F7565A6EF79" authorityName="Mollendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus) fruhstorferi</taxonomicName>
by monotypy. On the other hand,
<taxonomicName id="7328E2DA73FC8BA0E7DFBC740B08BAB2" authorityName="Mollendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus) fruhstorferi</taxonomicName>
was not listed in
<normalizedToken id="961CC54F7B25FF9B5D80E4ED60217D8D" originalValue="Kobelts">Kobelt's</normalizedToken>
and
<normalizedToken id="0671EF298F0CCE8F0DEAEA988A36103F" originalValue="Möllendorffs">Moellendorff's</normalizedToken>
(1897) paper, which indicates that they aimed to describe
<taxonomicName id="17A17DD1C4CAA517AAE6166C3408559A" lsidName="A. fruhstorferi" pageId="0" pageNumber="1" rank="species" species="fruhstorferi">
<emphasis id="18279CFEA2B6A24142E03BF941B46EE7" italics="true" pageId="0" pageNumber="1">A. fruhstorferi</emphasis>
</taxonomicName>
after their revision of
<taxonomicName id="797D16A4FAA756F07DD484C85F215D29" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" superFamily="Cyclophoroidea">
<emphasis id="9B8883C00BD98C2E62EA3FE644DFCECA" italics="true" pageId="0" pageNumber="1">Alycaeus</emphasis>
</taxonomicName>
. Moreover,
<bibRefCitation id="71DE957DD93897A739447FE06B328C00" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">
<normalizedToken id="E4EC7DDAA1A292773A6D3CC04EC72028" originalValue="Kobelts">Kobelt's</normalizedToken>
(1902)
</bibRefCitation>
monograph referred to
<taxonomicName id="8738D88E54ED58A6120E421CB808088F" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="085237EB0562E176BAB238D1C46DCDF3" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
as it was introduced by
<bibRefCitation id="81D82B281F0B4BC02093A4FB824B271F" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken id="34D5FDD56DFC1F790CBFEB78B78EF1BC" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897)
</bibRefCitation>
. Almost all subsequent treatments erroneously attributed the name
<taxonomicName id="C26FE33229B4A3D239B4F33C9717F8FF" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="A1758A3091D5552922AC15A828E81E58" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
to Kobelt &amp;
<normalizedToken id="2763FECA27377CB62A2E720AC8C842EC" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897 (
<bibRefCitation id="12F965632BDD2A05898B3AC8F2452962" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
;
<bibRefCitation id="B519EDA8ED92C0A556F81C535C805EB8" DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
;
<bibRefCitation id="B4D2F0A5479168D98A923406D054083C" author="Yen, T-C" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="1" refId="B229" refString="Yen, T-C, 1939. Die Chinesischen Land-und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" title="Die Chinesischen Land-und Suesswasser-Gastropoden des Natur-Museums Senckenberg." year="1939">Yen 1939</bibRefCitation>
;
<bibRefCitation id="D4DF2A44AA6B34C57FF513657C5FC591" author="Zilch, A" journalOrPublisher="Ltd., Beijing" pageId="0" pageNumber="1" refId="B232" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fuer Molluskenkunde 86(4/6): 141-150." title="Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fuer Molluskenkunde 86 (4 / 6): 141 - 150." year="1957">Zilch 1957</bibRefCitation>
;
<bibRefCitation id="60FD41940F8CE5E95E4BDF8D40DE4C27" author="Azuma, M" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="139 - 141" refId="B6" refString="Azuma, M, 1980. Description of a New Species of the Genus Awalycaeus Kuroda, 1915, (Family Cyclophoridae) from Mt. Sarumasa, Shimane Pref., Japan. Venus 39 (3): 139 - 141" title="Description of a New Species of the Genus Awalycaeus Kuroda, 1915, (Family Cyclophoridae) from Mt. Sarumasa, Shimane Pref., Japan." volume="39" year="1980">Azuma 1980</bibRefCitation>
;
<bibRefCitation id="DCE75353D1313FEBB247BB5699016A11" author="Minato, H" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="124 - 140" refId="B121" refString="Minato, H, 1982b. Land shell fauna of Ujigunto and Kusakakigunto Islets, the southwestern Kyushu, Japan, with the descriptions of a new genus and six new species. Venus 41 (2): 124 - 140" title="Land shell fauna of Ujigunto and Kusakakigunto Islets, the southwestern Kyushu, Japan, with the descriptions of a new genus and six new species." volume="41" year="1982 b">Minato 1982b</bibRefCitation>
,
<bibRefCitation id="2FAA5F2D692CA380CC4A3B4386A6544B" author="Minato, H" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="222 - 225" refId="B122" refString="Minato, H, 1987a. The new alycaeid species from Japan. Venus 45 (4): 222 - 225" title="The new alycaeid species from Japan." volume="45" year="1987 a">1987a</bibRefCitation>
,
<bibRefCitation id="5BC19E7F43144B1031D7542F9CCF6113" author="Minato, H" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="39 - 44" refId="B126" refString="Minato, H, 2005. Two new species of Chamalycaeus (Gastropoda: Alycaeidae) from Miyazaki Prefecture, southeastern Kyushu, Japan. Venus 64: 39 - 44" title="Two new species of Chamalycaeus (Gastropoda: Alycaeidae) from Miyazaki Prefecture, southeastern Kyushu, Japan." volume="64" year="2005">2005</bibRefCitation>
;
<bibRefCitation id="9BDF219307B462E66CC2AFD1D236F349" author="Minato, H" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="33 - 36" refId="B129" refString="Minato, H, Yano, Sh, 1988. Addition to the subgenus Sigmacharax (Alycaeidae) from Chūgoku District, Japan. Venus 47 (1): 33 - 36" title="Addition to the subgenus Sigmacharax (Alycaeidae) from Chūgoku District, Japan." volume="47" year="1988">Minato and Yano 1988</bibRefCitation>
;
<bibRefCitation id="2B52EE9FE8DADEF0A5CAD472BACE4813" author="Egorov, RV" journalOrPublisher="The Thailand Natural History Museum Journal" pageId="0" pageNumber="1" refId="B43" refString="Egorov, RV, 2005. Preliminary classification of the recent terrestrial pectinibranch mollusks. Club Conchylia Informationen 37(1/2): 21-32." title="Preliminary classification of the recent terrestrial pectinibranch mollusks. Club Conchylia Informationen 37 (1 / 2): 21 - 32." year="2005">Egorov 2005</bibRefCitation>
;
<bibRefCitation id="186644057DE0607AC37D178B24130C29" author="Zhang, W-H" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="1" pagination="745 - 747" refId="B230" refString="Zhang, W-H, Chen, D-N, Zhou, W-Ch, 2008. A new species of the genus Chamalycaeus from China (Gastropoda, Prosobranchia, Mesogastropoda, Cyclophoridae). Acta Zootaxonomica Sinica 33 (4): 745 - 747" title="A new species of the genus Chamalycaeus from China (Gastropoda, Prosobranchia, Mesogastropoda, Cyclophoridae)." volume="33" year="2008">Zhang et al. 2008</bibRefCitation>
) and referred to
<taxonomicName id="EAE312ED22D2828E50507FB563878A4F" authorityName="Benson" authorityYear="1861" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus andamaniae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="andamaniae" superFamily="Cyclophoroidea">
<emphasis id="3E3A8D99B27EB660399AE513D5811A80" italics="true" pageId="0" pageNumber="1">Alycaeus andamaniae</emphasis>
</taxonomicName>
Benson, 1861 as the type species as subsequent designated by
<bibRefCitation id="464717C294E48FC07F72344E343DB295" DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude (1921)</bibRefCitation>
. The ICZN Code 70.2 states the following: &quot;If it is found that an earlier type species fixation has been overlooked, the overlooked fixation is to be accepted and any later fixations are invalid. If this is considered to cause instability or confusion the case is to be referred to the Commission for a ruling&quot;. Therefore, we must examine whether the correction of the type fixation would cause instability. In our view, confusion or instability would be caused only if the majority of authors who have described species within
<taxonomicName id="6C79D9F265E862B9851C4AFBE1A283E9" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="BCA4DF24785863A278786D606ADE62A1" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
were unaware of the shell morphology of
<taxonomicName id="E41864F675CA921DCC834C59DBD992B8" authorityName="Benson" authorityYear="1861" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus andamaniae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="andamaniae" superFamily="Cyclophoroidea">
<emphasis id="BCC9AE3DF45753DFB0BA574FA6B6F83E" italics="true" pageId="0" pageNumber="1">Alycaeus andamaniae</emphasis>
</taxonomicName>
(incorrectly selected as the type species for
<taxonomicName id="029EBA9AC0D6A84C6C2E69EDD15B5650" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="044E775EA991FBDC7D5C6CDD4EFF5C82" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
). No detailed description of
<taxonomicName id="60DBDB8F31D4E71ED23EB8319D07677D" authorityName="Benson" authorityYear="1861" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus andamaniae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="andamaniae" superFamily="Cyclophoroidea">
<emphasis id="B62D30CEFC6A069AA000059E12F2399F" italics="true" pageId="0" pageNumber="1">Alycaeus andamaniae</emphasis>
</taxonomicName>
has ever been published, and our revision suggests that most authors who described
<taxonomicName id="6CB9BB4F79C4162090AAF521AD128E4D" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="4AAA35FEBB0A829B929F5B0239141552" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
species did not examine samples of
<taxonomicName id="78DDBB4B12FE86D6CEBB305A1A899FB6" authorityName="Benson" authorityYear="1861" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus andamaniae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="andamaniae" superFamily="Cyclophoroidea">
<emphasis id="F446D73D6904360CCBD45F3F44AFAF25" italics="true" pageId="0" pageNumber="1">Alycaeus andamaniae</emphasis>
</taxonomicName>
. Thus, we find no reason to present this issue to the Commission. Instead, we follow
<bibRefCitation id="D4A905B83B1899621F36E74C36E69C3A" author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov (2013)</bibRefCitation>
in accepting
<bibRefCitation id="5651D7D3394A468CC08F57995D7C35F0" author="Moellendorff, O von" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="89 - 97" publicationUrl="https://biodiversitylibrary.org/page/28228249" refId="B141" refString="Moellendorff, O von, 1897b. Neue Landschnecken von Java. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 89 - 97, https://biodiversitylibrary.org/page/28228249" title="Neue Landschnecken von Java." url="https://biodiversitylibrary.org/page/28228249" volume="29" year="1897 b">
<normalizedToken id="BC9C818630499A29D6C82817ED3F152D" originalValue="Möllendorff">Moellendorff</normalizedToken>
(1897b)
</bibRefCitation>
as the author of
<taxonomicName id="6D41F3F3EE5060D1BAE42BBF922749C2" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="2282AAF49C33FEB94CBC15E9FBD04F9F" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
. Thus, in accordance with Art. 70.2 of the Code we clarify that
<taxonomicName id="450233B7D06F167C3EA6E590E505CFA7" authorityName="Mollendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi" subGenus="Chamalycaeus" superFamily="Cyclophoroidea">Alycaeus (Chamalycaeus) fruhstorferi</taxonomicName>
is the type species of
<taxonomicName id="9DFC475490DAF984D07C9DCE95326E96" authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="34F0EBDF2E0A6B7B1A074FD1966214BC" italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
</taxonomicName>
<normalizedToken id="07A437B206AE4F1FF87D6A755E073A76" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897 by monotypy.
</paragraph>
</subSubSection>
</treatment>
</document>