901 lines
126 KiB
XML
901 lines
126 KiB
XML
<document id="C6695905B74EDA28A13F9A5B7CF1ED3A" ID-DOI="10.11646/zootaxa.5481.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742724" ID-ZooBank="FEC2B59D-D79F-4F9C-B082-D5BAABE1A453" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721033477194" checkinUser="plazi" docAuthor="Yin, Wen-Qi, Chen, Jia-Heng & Shi, Hong-Liang" docDate="2024" docId="03A6DA14FFF98D73289D7D68FA65F3D3" docLanguage="en" docName="zootaxa.5481.1.3.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tschitscherinea (Tschitscherinea) Berg 1898" docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="FF9FA26CFFF88D77280A7853FFF9F031" masterDocTitle="Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)" masterLastPageNumber="65" masterPageNumber="43" pageNumber="44" updateTime="1721135857569" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="BE544EDCD31A2159CE7A24F630638B58" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A46C85177432C026FFBEBB2C5CBD7696">
|
||
<mods:title id="DC5A39D8C8A8A19E9CC7AA5EB4109DA6">Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E515A85987276CE1F4FE890C881C009E" type="personal">
|
||
<mods:role id="01A9A23927CD6BFE0DE902C40DC167E3">
|
||
<mods:roleTerm id="F60DED417B0B9C997287BA0CD1F1E02A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6C7F848BAF6FECC6801E2B01760C7313">Yin, Wen-Qi</mods:namePart>
|
||
<mods:nameIdentifier id="57B6BB26DFA823D0338EC4F559801A46" type="ORCID">0000-0003-2903-1399</mods:nameIdentifier>
|
||
<mods:affiliation id="A2890CDCC908883AA235988538A7533C">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="0B47B7B001FF1DECD169A9E265180C5A" type="personal">
|
||
<mods:role id="D25AAE5CE471F226822B2ACBB6ED5F71">
|
||
<mods:roleTerm id="05455DA6FFDA85A8F84E93668B8985AE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9B498527539617F2CF369CA89136C770">Chen, Jia-Heng</mods:namePart>
|
||
<mods:nameIdentifier id="0DBA29F593219431F3AA755FF9722D23" type="ORCID">0000-0003-2449-8838</mods:nameIdentifier>
|
||
<mods:affiliation id="E8B24E4E4405ABA318DAC58A4E774C1C">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="31D25DE299C0608A9638DD313511D5E3" type="personal">
|
||
<mods:role id="690A045D9DE5C7D190D83A848E4E2C65">
|
||
<mods:roleTerm id="1B3FD39FB3C1321DD38258BE6C4E836A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A8FB0CA476402B775C26F34652B95A6A">Shi, Hong-Liang</mods:namePart>
|
||
<mods:nameIdentifier id="316987411FAA662021EF7A8D4C71B760" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
|
||
<mods:affiliation id="12BFE90AB701A5A40B5F2144299FD407">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
|
||
<mods:nameIdentifier id="A5E50AF9939564561455810BADD6304D" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="114E06AF8786489DFE2DC328FA32CC0B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CD4AE56D8A7102C346468EFC4BABC142" type="host">
|
||
<mods:titleInfo id="BA4F541C0D6ED72C89A2BD1E3A169771">
|
||
<mods:title id="780910F83A0FEC18EE7EF37400736FCB">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="021C00C93E2B6D6AA6F6DBAFFB8E994D">
|
||
<mods:date id="75EFFAAF6E79BC1122339AE178173316">2024</mods:date>
|
||
<mods:detail id="4AF762F07A5F29FC0BA58E3C1DAA426B" type="pubDate">
|
||
<mods:number id="605A40BAEBEDDCBCCBF86601277A79A2">2024-07-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AE909BCCF5B9401F8B5EE4C1617D5829" type="volume">
|
||
<mods:number id="2F75F9BDDFDFFFE2E8EAA6B2974801E3">5481</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CDC44DD19504809A6A7A75ABAED1B2EC" type="issue">
|
||
<mods:number id="CBA2D95F17129F63F2C25E3FFC8A3611">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9ED3BE31385A1C0FE75674B4058599BF" unit="page">
|
||
<mods:start id="22B4001334193038D26E3C4E9EA0FC17">43</mods:start>
|
||
<mods:end id="AA20C5FD083655E99AF6D55869CE55C3">65</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="1A5D9E31DE9FC39F42498E0EDE414837">
|
||
<mods:url id="72236704D7344D436B1231D1DA53479F">http://dx.doi.org/10.11646/zootaxa.5481.1.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="8F79E4F737C13568A64ECEF54BE8F450">journal article</mods:classification>
|
||
<mods:identifier id="DD7DC7EA67E9F5107D7EB88AA04466C4" type="DOI">10.11646/zootaxa.5481.1.3</mods:identifier>
|
||
<mods:identifier id="0343E09A5A5974F73F6D772FD3ADD2AD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="A179B8B60A1AB25F9816AE265DC0B58D" type="Zenodo-Dep">12742724</mods:identifier>
|
||
<mods:identifier id="EB787218A434B7A2FF17D257D2C874D6" type="ZooBank">FEC2B59D-D79F-4F9C-B082-D5BAABE1A453</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A6DA14FFF98D73289D7D68FA65F3D3" ID-DOI="http://doi.org/10.5281/zenodo.12750635" ID-Zenodo-Dep="12750635" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3" httpUri="http://treatment.plazi.org/id/03A6DA14FFF98D73289D7D68FA65F3D3" lastPageId="4" lastPageNumber="47" pageId="1" pageNumber="44">
|
||
<subSubSection id="C3153889FFF98D76289D7D68FD9BF564" box="[151,610,1339,1366]" pageId="1" pageNumber="44" type="nomenclature">
|
||
<paragraph id="8BB06B02FFF98D76289D7D68FD9BF564" blockId="1.[151,610,1339,1366]" box="[151,610,1339,1366]" pageId="1" pageNumber="44">
|
||
<heading id="D0F8DC6EFFF98D76289D7D68FD9BF564" bold="true" box="[151,610,1339,1366]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
|
||
<emphasis id="B97BB710FFF98D76289D7D68FD9BF564" bold="true" box="[151,610,1339,1366]" pageId="1" pageNumber="44">
|
||
Subgenus
|
||
<taxonomicName id="4C0F1081FFF98D76291D7D68FD9BF564" authority="Berg, 1898" authorityName="Berg" authorityYear="1898" box="[279,610,1339,1366]" class="Insecta" family="Carabidae" genus="Tschitscherinea" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFF98D76291D7D68FE2CF567" bold="true" box="[279,469,1339,1366]" italics="true" pageId="1" pageNumber="44">Tschitscherinea</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D7629D67D68FD9BF564" author="Berg, C." box="[476,610,1339,1365]" pageId="1" pageNumber="44" pagination="16 - 19" refId="ref15240" refString="Berg, C. (1898) Substitucion de nombres genericos. Comunicaciones del Museo Nacional de Buenos Aires, 1, 16 - 19." type="journal article" year="1898">Berg, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFF98D76289D7DD3FA9CF5C5" pageId="1" pageNumber="44" type="reference_group">
|
||
<paragraph id="8BB06B02FFF98D76289D7DD3FDC8F586" blockId="1.[151,1436,1407,1524]" pageId="1" pageNumber="44">
|
||
<treatmentCitationGroup id="AB1F4C2CFFF98D76289D7DD3FE0BF5A9" box="[151,498,1408,1432]" pageId="1" pageNumber="44">
|
||
<treatmentCitation id="0AAE4D13FFF98D76289D7DD3FE0BF5A9" author="Tschitscherine, T." box="[151,498,1408,1432]" page="70" pageId="1" pageNumber="44" year="1897">
|
||
<taxonomicName id="4C0F1081FFF98D76289D7DD3FE0BF5A9" ID-CoL="7P3PW" authority="Tschitscherine, 1897: 70" authorityName="Tschitscherine" authorityPageNumber="70" authorityYear="1897" box="[151,498,1408,1432]" class="Gastropoda" family="Lophospiridae" genus="Eunema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BB710FFF98D76289D7DD3FF13F5A6" box="[151,234,1408,1431]" italics="true" pageId="1" pageNumber="44">Eunema</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D7628F97DD3FE0BF5A9" author="Tschitscherine, T." box="[243,498,1408,1432]" pageId="1" pageNumber="44" pagination="45 - 75" refId="ref15733" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897: 70</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
[junior homonym of
|
||
<taxonomicName id="4C0F1081FFF98D762AD07DD3FC4AF5A6" ID-CoL="7P3PW" authority="Salter, 1859" authorityName="Salter" authorityYear="1859" box="[730,947,1407,1432]" class="Gastropoda" family="Lophospiridae" genus="Eunema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BB710FFF98D762AD07DD3FCD4F5A6" box="[730,813,1408,1431]" italics="true" pageId="1" pageNumber="44">Eunema</emphasis>
|
||
Salter, 1859
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C0F1081FFF98D762BCF7DD3FBA6F5A9" ID-CoL="7NWQ8" authorityName="Wenz" authorityYear="1938" box="[965,1119,1408,1432]" class="Gastropoda" family="Lophospiridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Mollusca" rank="family">Lophospiridae</taxonomicName>
|
||
)].
|
||
<typeStatus id="54B4D5A0FFF98D762C727DD3FB53F5A9" box="[1144,1194,1408,1432]" pageId="1" pageNumber="44">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C0F1081FFF98D762D047DD3FE62F587" ID-CoL="3DZ83" authority="Tschitscherine, 1897" authorityName="Tschitscherine" authorityYear="1897" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="filum">
|
||
<emphasis id="B97BB710FFF98D762D047DD3FA65F5A9" box="[1294,1436,1408,1432]" italics="true" pageId="1" pageNumber="44">Feronia filum</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D7628CD7DCDFE62F587" author="Tschitscherine, T." box="[199,411,1438,1462]" pageId="1" pageNumber="44" pagination="45 - 75" refId="ref15733" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFF98D76289D7DEEFA68F5E4" blockId="1.[151,1436,1407,1524]" box="[151,1425,1469,1493]" pageId="1" pageNumber="44">
|
||
<treatmentCitationGroup id="AB1F4C2CFFF98D76289D7DEEFE20F5E4" box="[151,473,1469,1493]" pageId="1" pageNumber="44">
|
||
<treatmentCitation id="0AAE4D13FFF98D76289D7DEEFE2CF5E4" author="Berg, C." box="[151,469,1469,1493]" page="18" pageId="1" pageNumber="44" year="1898">
|
||
<taxonomicName id="4C0F1081FFF98D76289D7DEEFE2CF5E4" ID-CoL="8MG8W" authority="Berg, 1898: 18" authorityName="Berg" authorityYear="1898" box="[151,469,1469,1493]" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFF98D76289D7DEEFECCF5E4" box="[151,309,1469,1493]" italics="true" pageId="1" pageNumber="44">Tschitscherinea</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D7629367DEEFE2CF5E4" author="Berg, C." box="[316,469,1469,1493]" pageId="1" pageNumber="44" pagination="16 - 19" refId="ref15240" refString="Berg, C. (1898) Substitucion de nombres genericos. Comunicaciones del Museo Nacional de Buenos Aires, 1, 16 - 19." type="journal article" year="1898">Berg, 1898: 18</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
<treatmentCitationGroup id="AB1F4C2CFFF98D7629D57DEEFA68F5E4" box="[479,1425,1469,1493]" pageId="1" pageNumber="44">
|
||
Replacement name for
|
||
<taxonomicName id="4C0F1081FFF98D762AC17DEDFC01F5E4" ID-CoL="7P3PW" authority="Tschitscherine, 1897" authorityName="Tschitscherine" authorityYear="1897" box="[715,1016,1469,1493]" class="Gastropoda" family="Lophospiridae" genus="Eunema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BB710FFF98D762AC17DEDFCE7F5E4" box="[715,798,1470,1493]" italics="true" pageId="1" pageNumber="44">Eunema</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D762B2E7DEEFC01F5E4" author="Tschitscherine, T." box="[804,1016,1469,1493]" pageId="1" pageNumber="44" pagination="45 - 75" refId="ref15733" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<treatmentCitation id="0AAE4D13FFF98D762C087DEEFB33F5E4" author="Jedlicka, A." box="[1026,1226,1469,1493]" page="228" pageId="1" pageNumber="44" year="1962">
|
||
<bibRefCitation id="EF9E16F3FFF98D762C087DEEFB33F5E4" author="Jedlicka, A." box="[1026,1226,1469,1493]" pageId="1" pageNumber="44" pagination="177 - 346" refId="ref15409" refString="Jedlicka, A. (1962) Monographie des Tribus Pterostichini aus Osatasien (Pterostichini, Trigonotomi, Myadi) (Coleoptera- Carabidae). Abhandlungen und Berichte aus dem Staatlichen Museum fur Tierkunde in Dresden, 26, 177 - 346." type="journal article" year="1962">Jedlička, 1962: 228</bibRefCitation>
|
||
</treatmentCitation>
|
||
. Sciaky, 1997: 166.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFF98D76289D7D8FFA9CF5C5" blockId="1.[151,1436,1407,1524]" box="[151,1381,1499,1524]" pageId="1" pageNumber="44">
|
||
<treatmentCitationGroup id="AB1F4C2CFFF98D76289D7D8FFE46F5C2" box="[151,447,1499,1524]" pageId="1" pageNumber="44">
|
||
<taxonomicName id="4C0F1081FFF98D76289D7D8FFE46F5C2" authority="Sciaky, 1997: 162" authorityName="Sciaky" authorityPageNumber="162" authorityYear="1997" box="[151,447,1499,1524]" class="Insecta" family="Carabidae" genus="Jedlickaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFF98D76289D7D8FFF06F5C5" box="[151,255,1500,1524]" italics="true" pageId="1" pageNumber="44">Jedlickaia</emphasis>
|
||
Sciaky, 1997: 162
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
.
|
||
<typeStatus id="54B4D5A0FFF98D7629C27D8FFE03F5C5" box="[456,506,1500,1524]" pageId="1" pageNumber="44">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C0F1081FFF98D762A5D7D8FFC45F5C5" ID-CoL="4Q3TW" authority="Jedlicka, 1934" authorityName="Jedlicka" authorityYear="1934" box="[599,956,1500,1524]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFF98D762A5D7D8FFCD8F5C5" box="[599,801,1500,1524]" italics="true" pageId="1" pageNumber="44">Pterostichus sterbai</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFF98D762B227D8FFC45F5C5" author="Jedlicka, A." box="[808,956,1500,1524]" pageId="1" pageNumber="44" pagination="13 - 19" refId="ref15381" refString="Jedlicka, A. (1934) Carabiden aus Ostasien. (5 Teil.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 31, 13 - 19." type="journal article" year="1934">Jedlička, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
, by original designation.
|
||
<emphasis id="B97BB710FFF98D762CB77D8FFA9CF5C5" bold="true" box="[1213,1381,1500,1524]" pageId="1" pageNumber="44">New synonymy.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFF98D75289D7E72FD44F57F" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="diagnosis">
|
||
<paragraph id="8BB06B02FFF98D76289D7E72FE93F724" blockId="1.[151,1437,1569,2028]" pageId="1" pageNumber="44">
|
||
<emphasis id="B97BB710FFF98D76289D7E72FEEBF60A" bold="true" box="[151,274,1569,1595]" pageId="1" pageNumber="44">Diagnosis.</emphasis>
|
||
Species of the subgenus
|
||
<taxonomicName id="4C0F1081FFF98D762A2E7E71FD2AF60D" box="[548,723,1570,1596]" pageId="1" pageNumber="44" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFF98D762A2E7E71FD2AF60D" box="[548,723,1570,1596]" italics="true" pageId="1" pageNumber="44">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
can be recognized among Chinese
|
||
<taxonomicName id="4C0F1081FFF98D762C5C7E71FB18F60D" box="[1110,1249,1570,1596]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFF98D762C5C7E71FB18F60D" box="[1110,1249,1570,1596]" italics="true" pageId="1" pageNumber="44">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
by the following combinations: medium-sized pterostichid beetles; body form relatively slender, more or less parallel-sided; dorsum reddish brown to dark brown; submentum with two setae on each side; elytra interval 3 with one to three setigerous pores; umbilicate pore series on interval 9 discontinuous in middle; metepisternum at most slightly longer than basal width; mesofemora with two setae on posterior-ventral margin; metacoxae with three setae; median lobe of aedeagus relatively small, apical lamella short with rounded apex; right paramere elongated, often curved at anterior third (
|
||
<figureCitation id="13347787FFF98D7628D17EA9FEA7F724" box="[219,350,1786,1813]" captionStart-0="FIGURES 13–18. 13–14" captionStart-1="FIGURES 19–24" captionStartId-0="15.[152,270,1767,1791]" captionStartId-1="16.[152,270,1691,1715]" captionTargetBox-0="[158,1431,181,1736]" captionTargetBox-1="[158,1429,188,1658]" captionTargetId-0="figure-18@15.[151,1436,181,1743]" captionTargetId-1="figure-36@16.[151,1436,181,1668]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." captionText-1="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12742732" figureDoi-1="http://doi.org/10.5281/zenodo.12742736" httpUri-0="https://zenodo.org/record/12742732/files/figure.png" httpUri-1="https://zenodo.org/record/12742736/files/figure.png" pageId="1" pageNumber="44">Figs 15–24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFF98D7528CD7F4EFD44F57F" blockId="1.[151,1437,1569,2028]" lastBlockId="2.[151,1437,151,2039]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
|
||
<emphasis id="B97BB710FFF98D7628CD7F4EFE23F709" bold="true" box="[199,474,1821,1848]" pageId="1" pageNumber="44">Subgeneric characters.</emphasis>
|
||
Body form relatively slender, more or less parallel-sided; body length 9.0–
|
||
<quantity id="4CF7C6E7FFF98D762D387F4DFA61F709" box="[1330,1432,1822,1849]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="44" unit="mm" value="15.5">15.5 mm</quantity>
|
||
; dorsum reddish-brown to dark brown, elytra without or with very faint metallic luster. Head small to distinctly thickened, always narrower than pronotum; eyes small, two supraorbital setae present; vertex without puncture or wrinkle; frontal grooves shallow, sometimes indistinct, slightly sinuate, reaching mid-point of eyes. Antennae hardly reaching or slightly exceeding elytra base; scape shorter than combined length of following two segments; antennomere 3 with 6–7 primary long setae forming an apical ring, without short accessory seta. Mandibles long, outer margin nearly straight, curved near apical fourth, apex sharply hooked; terminal labial and maxillary palpomere cylindrical, weakly broadened in males; penultimate labial palpomere with two setae along inner margin, without an extra seta near apex; submentum with two setae on each side. Pronotum usually nearly quadrate, lateral sides nearly paralleled or slightly curved, straight or slightly sinuate before posterior angles, but in
|
||
<taxonomicName id="4C0F1081FFFA8D752CBD7950FAD8F12C" box="[1207,1313,259,285]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752CBD7950FAD8F12C" box="[1207,1313,259,285]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
, pronotum nearly ovate, lateral margins evenly arched near middle; posterior margin almost straight, sometimes very shallowly emarginate at middle, usually a little narrower than base of elytra between humerus; anterior angles slightly protruded with blunt apex; posterior angles usually rectangular, apex forming faint denticles, except for
|
||
<taxonomicName id="4C0F1081FFFA8D752CA5793CFAE1F1B9" box="[1199,1304,366,392]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752CA5793CFAE1F1B9" box="[1199,1304,366,392]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
which with rounded-obtuse posterior angles; lateral margins with one mid-lateral seta near maximum width, with one posterior seta near posterior angle; disc flat to slightly convex, often with transverse wrinkles aside median line. Basal foveae composed of one or two shallow grooves, inner groove about one-third length of pronotum, outer groove short or absent; area between outer groove and lateral margin ridged or not; basal foveal area often finely punctate. Elytra parallel-sided or slightly elliptic, width a little greater than half of length, widest a little behind middle, with similar microsculpture in males and females, isodiametric or transverse. Humerus distinct, humeral angles more or less toothed and projected outward. Striae deeply incised, punctate or not, intervals slightly convex; parascutellar pore absent or present; parascutellar stria joined to stria 1, angular base of stria 1 usually short, free or joined to stria 1; interval 3 with one to three setigerous pores: the basal one (if present) adjacent to stria 3, the rest two adjacent to stria 2, the middle pore always present, the apical pore absent only in
|
||
<taxonomicName id="4C0F1081FFFA8D752BB87A87FBF0F2DC" authorityName="Tschitscherine" authorityYear="1897" box="[946,1033,723,749]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="filum">
|
||
<emphasis id="B97BB710FFFA8D752BB87A87FBF0F2DC" box="[946,1033,723,749]" italics="true" pageId="2" pageNumber="45">P. filum</emphasis>
|
||
</taxonomicName>
|
||
; interval 5 without setigerous pore; umbilicate pore series on interval 9 discontinuous in middle, often with 5–6 pores in basal group, 2–3 pores in the middle, 9–10 pores in apical group. Metepisternum slightly longer than basal width; sternite VII with one seta on each side in males, two in females; male abdominal sexual modification absent in most species, present in two species (
|
||
<taxonomicName id="4C0F1081FFFA8D7528AB7B30FE8BF34D" authority="Jedlicka" authorityName="Jedlicka" box="[161,370,866,892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D7528AB7B30FEF4F34D" box="[161,269,866,892]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
Jedlička
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F1081FFFA8D7529A47B30FDBCF34D" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[430,581,866,892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="jiuzhaiicus" status="sp. nov.">
|
||
<emphasis id="B97BB710FFFA8D7529A47B30FDBCF34D" box="[430,581,866,892]" italics="true" pageId="2" pageNumber="45">P. jiuzhaiicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFA8D752A457B31FD55F34D" bold="true" box="[591,684,866,892]" pageId="2" pageNumber="45">
|
||
<taxonomicNameLabel id="A2480A6BFFFA8D752A457B31FD55F34D" box="[591,684,866,892]" pageId="2" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), with simple median depression or depressed and longitudinally ridged on sternite VII (
|
||
<figureCitation id="13347787FFFA8D7529927BD5FDE5F391" box="[408,540,902,928]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="2" pageNumber="45">Figs 48–49</figureCitation>
|
||
). Mesofemora with two setae on posterior ventral margin; metacoxae with three setae; metatrochanters with one seta; metatarsus slender or thickened; fifth tarsomere glabrous ventrally; apical half of inner margin of meso- and metatibiae crenulate in males. Male genitalia relatively small in compare to other
|
||
<taxonomicName id="4C0F1081FFFA8D7528D17BA2FE9FF43A" box="[219,358,1009,1035]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D7528D17BA2FE9FF43A" box="[219,358,1009,1035]" italics="true" pageId="2" pageNumber="45">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
groups; ventral surface simple, almost smooth, without ridge; apical lamella relatively short with rounded apex, often more or less twisted to right side; apical orifice opened to dorsal-left side. Right paramere strongly elongated, often curved before midpoint, apex thickened or sharp (
|
||
<figureCitation id="13347787FFFA8D752BFA7C6AFB8CF465" box="[1008,1141,1081,1108]" captionStart-0="FIGURES 13–18. 13–14" captionStart-1="FIGURES 19–24" captionStartId-0="15.[152,270,1767,1791]" captionStartId-1="16.[152,270,1691,1715]" captionTargetBox-0="[158,1431,181,1736]" captionTargetBox-1="[158,1429,188,1658]" captionTargetId-0="figure-18@15.[151,1436,181,1743]" captionTargetId-1="figure-36@16.[151,1436,181,1668]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." captionText-1="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12742732" figureDoi-1="http://doi.org/10.5281/zenodo.12742736" httpUri-0="https://zenodo.org/record/12742732/files/figure.png" httpUri-1="https://zenodo.org/record/12742736/files/figure.png" pageId="2" pageNumber="45">Figs 15–24</figureCitation>
|
||
). Endophallus simple, left apical lobe (la) larger and spherical, right apical lobe (ra) smaller and sometimes with slightly sclerotized apex, a small sclerotized projection present near gonopore (
|
||
<figureCitation id="13347787FFFA8D752ADD7CD3FCA3F4AA" box="[727,858,1152,1179]" captionStart="FIGURES 25–30" captionStartId="17.[152,270,1537,1561]" captionTargetBox="[158,1430,183,1507]" captionTargetId="figure-18@17.[151,1436,181,1513]" captionTargetPageId="17" captionText="FIGURES 25–30. Endophallus of Pterostichus (Tschitscherinea) spp., left lateral view and right lateral view. (25) P.spissipedes sp. nov., Jianchuan, paratype. (26) P. jiuzhaiicus sp. nov., holotype. (27) P. mulensis Sciaky, Muli. (28) P. farkaci Sciaky, Hongshan, Zhongdian. (29) P. farkaci Sciaky, Tianchi, Zhongdian. (30) P. sterbai Jedlička, Pe-Yen-Tsin. Abbreviations: gp gonopore, la left apical lobe, ra right apical lobe, va ventral apical lobe. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742740" httpUri="https://zenodo.org/record/12742740/files/figure.png" pageId="2" pageNumber="45">Figs 25–30</figureCitation>
|
||
). Gonocoxite 2 of ovipositor almost identical across in species: falcate in ventral view, length about four times as basal width; outer margin with two ensiform setae, inner margin without ensiform seta; apex a little sharp in lateral view, with two nematiform setae in a groove (
|
||
<figureCitation id="13347787FFFA8D752D307C9BFA76F4D3" box="[1338,1423,1223,1250]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="2" pageNumber="45">Fig. 53</figureCitation>
|
||
). Spermatheca with seminal canal and receptaculum well differentiated; receptaculum straight and digitate, surface smooth, base without sclerotized projection; seminal canal five to six times length of receptaculum; spermathecal gland inserted on base of receptaculum (
|
||
<figureCitation id="13347787FFFA8D752A567D60FD48F57F" box="[604,689,1331,1358]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="2" pageNumber="45">Fig. 52</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFA8D7528CD7D05FDF7F5ED" pageId="2" pageNumber="45" type="biology_ecology">
|
||
<paragraph id="8BB06B02FFFA8D7528CD7D05FDF7F5ED" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="45">
|
||
<emphasis id="B97BB710FFFA8D7528CD7D05FED3F541" bold="true" box="[199,298,1366,1392]" pageId="2" pageNumber="45">Habitat.</emphasis>
|
||
Species of the subgenus
|
||
<taxonomicName id="4C0F1081FFFA8D752A527D04FCFEF540" box="[600,775,1367,1393]" pageId="2" pageNumber="45" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D752A527D04FCFEF540" box="[600,775,1367,1393]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
were usually found on the edge of mixed or coniferous forests in median-high altitudes (
|
||
<quantity id="4CF7C6E7FFFA8D752A017D29FD53F5A5" box="[523,682,1402,1428]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="4.2" metricValueMin="2.6" pageId="2" pageNumber="45" unit="m" value="3400.0" valueMax="4200.0" valueMin="2600.0">2600–4200 m</quantity>
|
||
) in southwest
|
||
<collectingCountry id="F3182B92FFFA8D752B457D29FC6FF5A5" box="[847,918,1402,1428]" name="China" pageId="2" pageNumber="45">China</collectingCountry>
|
||
. Adults tend to live under loose bark of fallen or standing dead trees. Different from other groups of
|
||
<taxonomicName id="4C0F1081FFFA8D752AFE7DCDFC86F589" box="[756,895,1438,1464]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D752AFE7DCDFC86F589" box="[756,895,1438,1464]" italics="true" pageId="2" pageNumber="45">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
, as they are more readily found under or in dead wood, rather than by pitfall traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFA8D7428CD7DB6FE43F718" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="discussion">
|
||
<paragraph id="8BB06B02FFFA8D7528CD7DB6FC52F7C7" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="45">
|
||
<emphasis id="B97BB710FFFA8D7528CD7DB6FE55F631" bold="true" box="[199,428,1509,1536]" pageId="2" pageNumber="45">Taxonomical notes.</emphasis>
|
||
The subgenus
|
||
<taxonomicName id="4C0F1081FFFA8D752A597DB6FD3FF5CE" authorityName="Sciaky" authorityYear="1997" box="[595,710,1509,1535]" class="Insecta" family="Carabidae" genus="Jedlickaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D752A597DB6FD3FF5CE" box="[595,710,1509,1535]" italics="true" pageId="2" pageNumber="45">Jedlickaia</emphasis>
|
||
</taxonomicName>
|
||
contains only one species
|
||
<taxonomicName id="4C0F1081FFFA8D752BE57DB5FBA1F5CE" box="[1007,1112,1509,1535]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752BE57DB5FBA1F5CE" box="[1007,1112,1509,1535]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
distributed in North
|
||
<collectingRegion id="49CBA5E0FFFA8D752D4A7DB5FA60F631" box="[1344,1433,1510,1536]" country="China" name="Yunnan" pageId="2" pageNumber="45">Yunnan</collectingRegion>
|
||
, and was supposed to be related to subgenera
|
||
<taxonomicName id="4C0F1081FFFA8D752A807E5AFCC0F612" box="[650,825,1545,1571]" pageId="2" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D752A807E5AFCC0F612" box="[650,825,1545,1571]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C0F1081FFFA8D752B577E5AFB88F612" authority="(Sciaky, 1997)" baseAuthorityName="Sciaky" baseAuthorityYear="1997" box="[861,1137,1545,1571]" class="Insecta" family="Carabidae" genus="Megabea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D752B577E5AFC3CF612" box="[861,965,1545,1571]" italics="true" pageId="2" pageNumber="45">Megabea</emphasis>
|
||
(Sciaky, 1997)
|
||
</taxonomicName>
|
||
. After examining the
|
||
<typeStatus id="54B4D5A0FFFA8D752D6B7E59FA65F615" box="[1377,1436,1546,1572]" pageId="2" pageNumber="45">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0F1081FFFA8D7528B27E7DFEDCF676" box="[184,293,1581,1607]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D7528B27E7DFEDCF676" box="[184,293,1581,1607]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
, it is found that this species is in accord with subgenus
|
||
<taxonomicName id="4C0F1081FFFA8D752BBF7E7EFB9DF676" box="[949,1124,1581,1607]" pageId="2" pageNumber="45" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D752BBF7E7EFB9DF676" box="[949,1124,1581,1607]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
in the following important aspects: submentum with two setae on each side; metacoxae with three setae; elytra with distinct humeral tooth; umbilicate pore series on interval 9 discontinuous in middle; apical lamella of aedeagus very short; right paramere elongated; endophallus simple and strongly bent ventrally, with la and ra small and in similar size. Moreover,
|
||
<taxonomicName id="4C0F1081FFFA8D75289D7EEEFEFAF6E7" box="[151,259,1724,1750]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D75289D7EEEFEFAF6E7" box="[151,259,1724,1750]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
has two uncommon features among Chinese
|
||
<taxonomicName id="4C0F1081FFFA8D752B047EEFFC60F6E7" box="[782,921,1724,1750]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D752B047EEFFC60F6E7" box="[782,921,1724,1750]" italics="true" pageId="2" pageNumber="45">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
: (1) metatarsus distinctly thickened in males (
|
||
<figureCitation id="13347787FFFA8D7528957EB3FF0EF6CB" box="[159,247,1760,1786]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="2" pageNumber="45">Figs 14</figureCitation>
|
||
,
|
||
<figureCitation id="13347787FFFA8D75290F7EB3FEDCF6CB" box="[261,293,1760,1786]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="2" pageNumber="45">43</figureCitation>
|
||
); (2) eyes small with strongly swollen tempora. These two features also present in two other species belonging to the subgenus
|
||
<taxonomicName id="4C0F1081FFFA8D7529CB7F57FD89F72F" box="[449,624,1796,1822]" pageId="2" pageNumber="45" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D7529CB7F57FD89F72F" box="[449,624,1796,1822]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C0F1081FFFA8D752A757F56FCE1F72F" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[639,792,1796,1822]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="spissipedes" status="sp. nov.">
|
||
<emphasis id="B97BB710FFFA8D752A757F56FCE1F72F" box="[639,792,1796,1822]" italics="true" pageId="2" pageNumber="45">P. spissipedes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFA8D752B147F57FC8EF72F" bold="true" box="[798,887,1796,1822]" pageId="2" pageNumber="45">
|
||
<taxonomicNameLabel id="A2480A6BFFFA8D752B147F57FC8EF72F" box="[798,887,1796,1822]" pageId="2" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0F1081FFFA8D752BA67F56FBC6F72F" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[940,1087,1796,1822]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. nov.">
|
||
<emphasis id="B97BB710FFFA8D752BA67F56FBC6F72F" box="[940,1087,1796,1822]" italics="true" pageId="2" pageNumber="45">P. spectabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFA8D752C4F7F57FB67F72F" bold="true" box="[1093,1182,1796,1822]" pageId="2" pageNumber="45">
|
||
<taxonomicNameLabel id="A2480A6BFFFA8D752C4F7F57FB67F72F" box="[1093,1182,1796,1822]" pageId="2" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) distributed in
|
||
<collectingRegion id="49CBA5E0FFFA8D752D4D7F56FF00F773" country="China" name="Yunnan" pageId="2" pageNumber="45">Yunnan province</collectingRegion>
|
||
as well. Thus, these two species could be the closest relatives to
|
||
<taxonomicName id="4C0F1081FFFA8D752BD67F7AFBBEF773" box="[988,1095,1832,1858]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752BD67F7AFBBEF773" box="[988,1095,1832,1858]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13347787FFFA8D752C5C7F7BFB20F773" box="[1110,1241,1832,1858]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="2" pageNumber="45">Figs 13, 14</figureCitation>
|
||
). Additionally,
|
||
<taxonomicName id="4C0F1081FFFA8D752D827F7AFF1CF757" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752D827F7AFF1CF757" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
is different from most other species of
|
||
<taxonomicName id="4C0F1081FFFA8D752A957F1FFCB7F757" box="[671,846,1868,1894]" pageId="2" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D752A957F1FFCB7F757" box="[671,846,1868,1894]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
in having a straight right paramere of male genitalia (
|
||
<figureCitation id="13347787FFFA8D7528AA7F23FF0AF7BA" box="[160,243,1904,1931]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="2" pageNumber="45">Fig. 15</figureCitation>
|
||
) and sexual modification on sternite VII (
|
||
<figureCitation id="13347787FFFA8D752AD07F23FCC8F7BB" box="[730,817,1904,1930]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="2" pageNumber="45">Fig. 49</figureCitation>
|
||
). But
|
||
<taxonomicName id="4C0F1081FFFA8D752B727F22FBF6F7BB" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[888,1039,1904,1930]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="jiuzhaiicus" status="sp. nov.">
|
||
<emphasis id="B97BB710FFFA8D752B727F22FBF6F7BB" box="[888,1039,1904,1930]" italics="true" pageId="2" pageNumber="45">P. jiuzhaiicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFA8D752C137F23FB8CF7BB" bold="true" box="[1049,1141,1904,1930]" pageId="2" pageNumber="45">
|
||
<taxonomicNameLabel id="A2480A6BFFFA8D752C137F23FB8CF7BB" box="[1049,1141,1904,1930]" pageId="2" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C0F1081FFFA8D752D1B7F22FA84F7BB" box="[1297,1405,1904,1930]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D752D1B7F22FA84F7BB" box="[1297,1405,1904,1930]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
in these two aspects (
|
||
<figureCitation id="13347787FFFA8D75296D7FC7FE44F79F" box="[359,445,1940,1966]" captionStart="FIGURES 19–24" captionStartId="16.[152,270,1691,1715]" captionTargetBox="[158,1429,188,1658]" captionTargetId="figure-36@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742736" httpUri="https://zenodo.org/record/12742736/files/figure.png" pageId="2" pageNumber="45">Figs 19</figureCitation>
|
||
,
|
||
<figureCitation id="13347787FFFA8D7529C27FC7FE11F79F" box="[456,488,1940,1966]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="2" pageNumber="45">48</figureCitation>
|
||
). Therefore, despite the circular pronotum which is different from all other species of
|
||
<taxonomicName id="4C0F1081FFFA8D75289D7FEBFEBFF7E3" box="[151,326,1976,2002]" pageId="2" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D75289D7FEBFEBFF7E3" box="[151,326,1976,2002]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFA8D75295E7FEAFE47F7E3" box="[340,446,1976,2002]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFA8D75295E7FEAFE47F7E3" box="[340,446,1976,2002]" italics="true" pageId="2" pageNumber="45">P. sterbai</emphasis>
|
||
</taxonomicName>
|
||
fits all characters of this subgenus. In conclusion, the subgenus
|
||
<taxonomicName id="4C0F1081FFFA8D752C9B7FEBFA65F7E3" authority="Sciaky, 1997" authorityName="Sciaky" authorityYear="1997" box="[1169,1436,1976,2002]" class="Insecta" family="Carabidae" genus="Jedlickaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFA8D752C9B7FEBFAFDF7E3" box="[1169,1284,1976,2002]" italics="true" pageId="2" pageNumber="45">Jedlickaia</emphasis>
|
||
Sciaky, 1997
|
||
</taxonomicName>
|
||
is newly proposed as a junior synonym of
|
||
<taxonomicName id="4C0F1081FFFA8D752A787F8FFC5FF7C7" authority="Berg, 1898" authorityName="Berg" authorityYear="1898" box="[626,934,2012,2039]" pageId="2" pageNumber="46" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFA8D752A787F8FFCD8F7C7" box="[626,801,2012,2038]" italics="true" pageId="2" pageNumber="45">Tschitscherinea</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFA8D752B2D7F8EFC5FF7C7" author="Berg, C." box="[807,934,2012,2039]" pageId="2" pageNumber="45" pagination="16 - 19" refId="ref15240" refString="Berg, C. (1898) Substitucion de nombres genericos. Comunicaciones del Museo Nacional de Buenos Aires, 1, 16 - 19." type="journal article" year="1898">Berg, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D7428CD78C4FC4AF154" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="46">
|
||
For most of the Chinese
|
||
<taxonomicName id="4C0F1081FFFB8D7429EB78C4FD95F080" box="[481,620,151,177]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFB8D7429EB78C4FD95F080" box="[481,620,151,177]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
subgenera, the metacoxa has only two setae close to the outer-posterior angle and the anterior margin respectively. Metacoxae have one additional seta near the inner angle (see
|
||
<bibRefCitation id="EF9E16F3FFFB8D742D0478E8FF2AF0C8" author="Shi, H. L. & Liang, H. B." pageId="3" pageNumber="46" pagination="1 - 92" refId="ref15602" refString="Shi, H. L. & Liang, H. B. (2015) The genus Pterostichus in China II: The subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini). ZooKeys, 536, 1 - 92. https: // doi. org / 10.3897 / zookeys. 536.5982" type="journal article" year="2015">Shi & Liang, 2015</bibRefCitation>
|
||
: Fig. 137) only in the following four Chinese subgenera:
|
||
<taxonomicName id="4C0F1081FFFB8D742B70788CFB4CF0C8" authority="Berg, 1898" authorityName="Berg" authorityYear="1898" box="[890,1205,223,249]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742B70788CFBD0F0C8" box="[890,1065,223,249]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742C39788CFB4CF0C8" author="Berg, C." box="[1075,1205,223,249]" pageId="3" pageNumber="46" pagination="16 - 19" refId="ref15240" refString="Berg, C. (1898) Substitucion de nombres genericos. Comunicaciones del Museo Nacional de Buenos Aires, 1, 16 - 19." type="journal article" year="1898">Berg, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFB8D742CCE788CFA65F0C8" box="[1220,1436,223,249]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742CCE788CFA65F0C8" box="[1220,1436,223,249]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EF9E16F3FFFB8D74289D7950FE7DF12C" author="Tschitscherine, T." box="[151,388,259,285]" pageId="3" pageNumber="46" pagination="1 - 224" refId="ref15754" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des feronien s. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224. [1898 - 1899]" type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFB8D7429987950FE00F12C" box="[402,505,259,285]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Cryobius">
|
||
<emphasis id="B97BB710FFFB8D7429987950FE00F12C" box="[402,505,259,285]" italics="true" pageId="3" pageNumber="46">Cryobius</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742A097950FD4CF12C" author="Chaudoir, M." box="[515,693,259,285]" pageId="3" pageNumber="46" pagination="3 - 32" refId="ref15294" refString="Chaudoir, M. (1838) Tableau d'une nouvelle subdivision du genre Feronia Dejean suibi d'une caracteristique de trois nouveaux genres de carabiques. Bulletin de la societe Imperiale des Naturalistes de Moscou, 11, 3 - 32." type="journal article" year="1838">Chaudoir, 1838</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C0F1081FFFB8D742AFF7950FBBAF12C" authority="Tian & Huang, 2019" authorityName="Tian & Huang" authorityYear="2019" box="[757,1091,259,285]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Huaius">
|
||
<emphasis id="B97BB710FFFB8D742AFF7950FCBFF12D" box="[757,838,259,284]" italics="true" pageId="3" pageNumber="46">Huaius</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742B5A7950FBBAF12C" author="Tian, M. & Huang, S. & Chen, M. & Ding, K." box="[848,1091,259,285]" pageId="3" pageNumber="46" pagination="1 - 16" refId="ref15452" refString="Tian, M., Huang, S., Chen, M. & Ding, K. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae), Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">Tian & Huang, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
. Based on this synapomorphy (character polarity determined by
|
||
<bibRefCitation id="EF9E16F3FFFB8D742A1B7974FD1AF170" author="Bousquet, Y." box="[529,739,294,321]" pageId="3" pageNumber="46" pagination="1 - 292" refId="ref15266" refString="Bousquet, Y. (1999) Supraspecific classification of the Nearctic Pterostichini (Coleoptera: Carabidae). Fabreries Supplement, 9, 1 - 292." type="journal article" year="1999">Bousquet 1999: 35</bibRefCitation>
|
||
) and phenetic similarities, these four subgenera are considered to be closer to each other than to other subgenera of the Chinese fauna.
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D7428CD793CFDDBF250" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="46">
|
||
Compared to the subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D7429E5793CFDB9F1B9" authorityName="Tian & Huang" authorityYear="2019" box="[495,576,367,392]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Huaius">
|
||
<emphasis id="B97BB710FFFB8D7429E5793CFDB9F1B9" box="[495,576,367,392]" italics="true" pageId="3" pageNumber="46">Huaius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFB8D742A46793CFD02F1B8" box="[588,763,367,393]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742A46793CFD02F1B8" box="[588,763,367,393]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
is different in having a parallel-sided body form, subquadrate or ovate pronotum, and discontinuous umbilicate series on interval 9. whereas, In the troglobitic
|
||
<taxonomicName id="4C0F1081FFFB8D742CDB79C0FADBF19D" authorityName="Tian & Huang" authorityYear="2019" box="[1233,1314,403,428]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Huaius">
|
||
<emphasis id="B97BB710FFFB8D742CDB79C0FADBF19D" box="[1233,1314,403,428]" italics="true" pageId="3" pageNumber="46">Huaius</emphasis>
|
||
</taxonomicName>
|
||
, pronotum is more cordate with lateral margins distinctly sinuate before posterior angles and the umbilicate pore series on interval 9 continuous. The subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D742A327988FD66F1C4" box="[568,671,475,501]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Cryobius">
|
||
<emphasis id="B97BB710FFFB8D742A327988FD66F1C4" box="[568,671,475,501]" italics="true" pageId="3" pageNumber="46">Cryobius</emphasis>
|
||
</taxonomicName>
|
||
contains more than a hundred species with a Holarctic distribution. In
|
||
<collectingCountry id="F3182B92FFFB8D7428BD79ACFF04F228" box="[183,253,511,537]" name="China" pageId="3" pageNumber="46">China</collectingCountry>
|
||
, there are only a few species belonging to the subgenus distributed in the Northeast. For all the Chinese species of
|
||
<taxonomicName id="4C0F1081FFFB8D7429047A70FE8CF20C" box="[270,373,547,573]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Cryobius">
|
||
<emphasis id="B97BB710FFFB8D7429047A70FE8CF20C" box="[270,373,547,573]" italics="true" pageId="3" pageNumber="46">Cryobius</emphasis>
|
||
</taxonomicName>
|
||
, they are distinguishable from
|
||
<taxonomicName id="4C0F1081FFFB8D742AD87A70FC78F20C" box="[722,897,547,573]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742AD87A70FC78F20C" box="[722,897,547,573]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
by having a smaller and stouter body form, and the fifth tarsomere setose ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D7428CD7A38FB2BF564" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="46">
|
||
<taxonomicName id="4C0F1081FFFB8D7428CD7A38FE66F2B4" box="[199,415,619,645]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7428CD7A38FE66F2B4" box="[199,415,619,645]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
is distributed in South
|
||
<collectingCountry id="F3182B92FFFB8D742AA47A38FD0DF2B4" box="[686,756,619,645]" name="China" pageId="3" pageNumber="46">China</collectingCountry>
|
||
, North
|
||
<collectingCountry id="F3182B92FFFB8D742B417A38FC54F2B4" box="[843,941,619,645]" name="Vietnam" pageId="3" pageNumber="46">Vietnam</collectingCountry>
|
||
, and North
|
||
<collectingCountry id="F3182B92FFFB8D742C3C7A38FB5AF2B4" box="[1078,1187,619,645]" name="Myanmar" pageId="3" pageNumber="46">Myanmar</collectingCountry>
|
||
and has 36 described species, and probably more than that undescribed. The subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D742B9D7ADCFBBFF298" box="[919,1094,655,681]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742B9D7ADCFBBFF298" box="[919,1094,655,681]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
is suspected to be closest to
|
||
<taxonomicName id="4C0F1081FFFB8D74289D7AE0FE96F2FC" box="[151,367,691,717]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D74289D7AE0FE96F2FC" box="[151,367,691,717]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
. However, it is difficult to infer the systematic relationship between these two subgenera before the phylogenetic relationship within
|
||
<taxonomicName id="4C0F1081FFFB8D742A4F7A84FD29F2C0" box="[581,720,727,753]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97BB710FFFB8D742A4F7A84FD29F2C0" box="[581,720,727,753]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
is better known. In most cases, these two subgenera can be distinguished through the following morphological features but not always: (1) In
|
||
<taxonomicName id="4C0F1081FFFB8D742C497AA8FB0BF324" box="[1091,1266,763,789]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742C497AA8FB0BF324" box="[1091,1266,763,789]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
the body form is generally more elongated and parallel-sided; whereas in
|
||
<taxonomicName id="4C0F1081FFFB8D742B307B4CFBEBF308" box="[826,1042,799,825]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742B307B4CFBEBF308" box="[826,1042,799,825]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, if the body is somewhat parallel-sided then the body size is stouter and smaller (less than
|
||
<quantity id="4CF7C6E7FFFB8D742B057B10FCAAF36C" box="[783,851,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="46" unit="mm" value="9.0">9 mm</quantity>
|
||
). (2) In
|
||
<taxonomicName id="4C0F1081FFFB8D742BA17B10FBA3F36C" box="[939,1114,835,861]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742BA17B10FBA3F36C" box="[939,1114,835,861]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
the fifth tarsomere is always glabrous ventrally; whereas in
|
||
<taxonomicName id="4C0F1081FFFB8D7429E57B34FD3EF3B0" box="[495,711,871,897]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7429E57B34FD3EF3B0" box="[495,711,871,897]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, the fifth tarsomere is usually setose ventrally with the exception of four described species (
|
||
<taxonomicName id="4C0F1081FFFB8D7429B77BD8FCD5F394" authority="Sciaky & Wrase, 1997" authorityName="Sciaky & Wrase" authorityYear="1997" box="[445,812,907,933]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="geberti">
|
||
<emphasis id="B97BB710FFFB8D7429B77BD8FDDEF394" box="[445,551,907,933]" italics="true" pageId="3" pageNumber="46">P. geberti</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742A247BD8FCD5F394" author="Sciaky, R. & Wrase, D. W." box="[558,812,907,933]" pageId="3" pageNumber="46" pagination="1087 - 1139" refId="ref15565" refString="Sciaky, R. & Wrase, D. W. (1997) Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae). Linzer Biologische Beitrage, 29, 1087 - 1139." type="journal article" year="1997">Sciaky & Wrase, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFB8D742B3D7BD8FB86F394" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[823,1151,907,933]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="glabellus">
|
||
<emphasis id="B97BB710FFFB8D742B3D7BD8FC40F394" box="[823,953,907,933]" italics="true" pageId="3" pageNumber="46">P. glabellus</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742BB57BD8FB86F394" author="Fedorenko, D. N." box="[959,1151,907,933]" pageId="3" pageNumber="46" pagination="16 - 39" refId="ref15335" refString="Fedorenko, D. N. (2023) New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam. Russian Entomological Journal, 32 (1), 16 - 39. https: // doi. org / 10.15298 / rusentj. 32.1.03" type="journal article" year="2023">Fedorenko, 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F1081FFFB8D742C807BD8FF2BF3F8" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="primitivus">
|
||
<emphasis id="B97BB710FFFB8D742C807BD8FAEEF395" box="[1162,1303,907,932]" italics="true" pageId="3" pageNumber="46">P. primitivus</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742D177BD8FF2BF3F8" author="Fedorenko, D. N." pageId="3" pageNumber="46" pagination="16 - 39" refId="ref15335" refString="Fedorenko, D. N. (2023) New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam. Russian Entomological Journal, 32 (1), 16 - 39. https: // doi. org / 10.15298 / rusentj. 32.1.03" type="journal article" year="2023">Fedorenko, 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0F1081FFFB8D7429017BFCFD9BF3F8" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[267,610,943,969]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="unisetosus">
|
||
<emphasis id="B97BB710FFFB8D7429017BFCFE62F3F9" box="[267,411,943,968]" italics="true" pageId="3" pageNumber="46">P. unisetosus</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D7429AB7BFCFD9BF3F8" author="Fedorenko, D. N." box="[417,610,943,969]" pageId="3" pageNumber="46" pagination="16 - 39" refId="ref15335" refString="Fedorenko, D. N. (2023) New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam. Russian Entomological Journal, 32 (1), 16 - 39. https: // doi. org / 10.15298 / rusentj. 32.1.03" type="journal article" year="2023">Fedorenko, 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
), and additionally in some undescribed species. (3) In
|
||
<taxonomicName id="4C0F1081FFFB8D742CB47BFCFA94F3F8" box="[1214,1389,943,969]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742CB47BFCFA94F3F8" box="[1214,1389,943,969]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
, the apical lamella of aedeagus is small with a simple rounded apex; whereas in
|
||
<taxonomicName id="4C0F1081FFFB8D742BD17B80FB4AF3DC" box="[987,1203,979,1005]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742BD17B80FB4AF3DC" box="[987,1203,979,1005]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, the apical lamella is generally larger, often toothed or twisted. (4) In
|
||
<taxonomicName id="4C0F1081FFFB8D742ABC7BA4FC9CF420" box="[694,869,1015,1041]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742ABC7BA4FC9CF420" box="[694,869,1015,1041]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
, the right paramere of aedeagus is elongated, with the length of the apical portion (area distal to the articulation) at least fourth as width, curved or nearly straight, apex blunt or gradually attenuate (
|
||
<figureCitation id="13347787FFFB8D7429D37C6CFDA5F468" box="[473,604,1086,1113]" captionStart-0="FIGURES 13–18. 13–14" captionStart-1="FIGURES 19–24" captionStartId-0="15.[152,270,1767,1791]" captionStartId-1="16.[152,270,1691,1715]" captionTargetBox-0="[158,1431,181,1736]" captionTargetBox-1="[158,1429,188,1658]" captionTargetId-0="figure-18@15.[151,1436,181,1743]" captionTargetId-1="figure-36@16.[151,1436,181,1668]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." captionText-1="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12742732" figureDoi-1="http://doi.org/10.5281/zenodo.12742736" httpUri-0="https://zenodo.org/record/12742732/files/figure.png" httpUri-1="https://zenodo.org/record/12742736/files/figure.png" pageId="3" pageNumber="46">Figs 15–24</figureCitation>
|
||
); whereas in
|
||
<taxonomicName id="4C0F1081FFFB8D742AE47C6CFC3FF468" box="[750,966,1087,1113]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742AE47C6CFC3FF468" box="[750,966,1087,1113]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, the right paramere of aedeagus is usually very short, with length of apical portion at most twice as width, apex widely rounded (
|
||
<figureCitation id="13347787FFFB8D742C577C30FB49F44C" box="[1117,1200,1123,1149]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky & Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="3" pageNumber="46">Fig. 33</figureCitation>
|
||
). The only exception of
|
||
<taxonomicName id="4C0F1081FFFB8D7428BF7CD4FE74F490" box="[181,397,1159,1185]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7428BF7CD4FE74F490" box="[181,397,1159,1185]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
is in
|
||
<taxonomicName id="4C0F1081FFFB8D7429C17CD4FD32F490" authority="Sciaky, 1994" authorityName="Sciaky, Houzhenzi" authorityYear="1994" box="[459,715,1159,1185]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="dundai">
|
||
<emphasis id="B97BB710FFFB8D7429C17CD4FDCDF490" box="[459,564,1159,1185]" italics="true" pageId="3" pageNumber="46">P. dundai</emphasis>
|
||
Sciaky, 1994
|
||
</taxonomicName>
|
||
which has the apical portion of right paramere widely triangular with an abruptly narrowed and elongated apical extension. (5) In
|
||
<taxonomicName id="4C0F1081FFFB8D742B6D7CF8FBEFF4F4" box="[871,1046,1195,1221]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D742B6D7CF8FBEFF4F4" box="[871,1046,1195,1221]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
</taxonomicName>
|
||
, the endophallus is relatively small and simple, only with a spherical left apical lobe and a very small right apical lobe; whereas in
|
||
<taxonomicName id="4C0F1081FFFB8D742CB77C9CFA6CF4D8" box="[1213,1429,1231,1257]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742CB77C9CFA6CF4D8" box="[1213,1429,1231,1257]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, the endophallus is diverse in form, usually much larger with several lobes in more complicated shapes (
|
||
<figureCitation id="13347787FFFB8D742D047CA0FA69F53C" box="[1294,1424,1266,1293]" captionStart="FIGURES 54–57. 54" captionStartId="21.[152,270,1525,1549]" captionTargetBox="[151,1435,181,1494]" captionTargetId="figure-18@21.[151,1435,181,1501]" captionTargetPageId="21" captionText="FIGURES 54–57. 54. Confirmed distribution of the Pterostichus (Tschitscherinea) spp. in China: Red P. spissipedes sp. nov. Blue P. spectabilis sp. nov. Black P. jiuzhaiicus sp. nov. Pink P. filum (Tschitschérine). Purple P. mulensis Sciaky. Green P. fakarci Sciaky. Yellow P. krali Sciaky. Cyan P. sterbai Jedlička. 55–57. Endophallus of Pterostichus spp., right lateral view and left lateral view: (55) Pterostichus (Cryobius) jaechi Krischenhofer, Changbaishan, Jilin. (56) Pterostichus (Morphohaptoderus) janatai Sciaky & Wrase, Pingheliang, Shaanxi. (57) Pterostichus (Morphohaptoderus) dundai Sciaky, Houzhenzi, Shaanxi. Scale bars = 0.5 mm, a (55), b (56–57)." figureDoi="http://doi.org/10.5281/zenodo.12742754" httpUri="https://zenodo.org/record/12742754/files/figure.png" pageId="3" pageNumber="46">Figs 56–57</figureCitation>
|
||
). Among these characters mentioned above, the shape of the right paramere of aedeagus can be well-define in these two subgenera, because this character is always reliable for all known species in both subgenera.
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D7428CD7D0CFB9BF5F0" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="46">
|
||
The characters of the subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D742A3D7D0CFD71F549" authorityName="Tian & Huang" authorityYear="2019" box="[567,648,1375,1400]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Huaius">
|
||
<emphasis id="B97BB710FFFB8D742A3D7D0CFD71F549" box="[567,648,1375,1400]" italics="true" pageId="3" pageNumber="46">Huaius</emphasis>
|
||
</taxonomicName>
|
||
were based on
|
||
<bibRefCitation id="EF9E16F3FFFB8D742B4B7D0CFBF6F548" author="Tian, M. & Huang, S. & Chen, M. & Ding, K." box="[833,1039,1375,1401]" pageId="3" pageNumber="46" pagination="1 - 16" refId="ref15452" refString="Tian, M., Huang, S., Chen, M. & Ding, K. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae), Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">
|
||
Tian
|
||
<emphasis id="B97BB710FFFB8D742B777D0CFC41F548" box="[893,952,1375,1401]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
. The characters of the Vietnamese species of subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D7429707DD0FDABF5AC" box="[378,594,1411,1437]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7429707DD0FDABF5AC" box="[378,594,1411,1437]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
were based on
|
||
<bibRefCitation id="EF9E16F3FFFB8D742AF77DD0FC30F5AC" author="Fedorenko, D. N." box="[765,969,1411,1437]" pageId="3" pageNumber="46" pagination="16 - 39" refId="ref15335" refString="Fedorenko, D. N. (2023) New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam. Russian Entomological Journal, 32 (1), 16 - 39. https: // doi. org / 10.15298 / rusentj. 32.1.03" type="journal article" year="2023">Fedorenko (2023)</bibRefCitation>
|
||
. The characters of subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D742D0D7DD0FA97F5AC" box="[1287,1390,1411,1437]" pageId="3" pageNumber="46" rank="subGenus" subGenus="Cryobius">
|
||
<emphasis id="B97BB710FFFB8D742D0D7DD0FA97F5AC" box="[1287,1390,1411,1437]" italics="true" pageId="3" pageNumber="46">Cryobius</emphasis>
|
||
</taxonomicName>
|
||
and other
|
||
<taxonomicName id="4C0F1081FFFB8D7428D27DF4FE49F5F0" box="[216,432,1447,1473]" pageId="3" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7428D27DF4FE49F5F0" box="[216,432,1447,1473]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
species were based on the following specimen examinations:
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D7428CD7D98FE43F718" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="46">
|
||
<taxonomicName id="4C0F1081FFFB8D7428CD7D98FD2EF5D4" authority="Krischenhofer" authorityName="Krischenhofer" box="[199,727,1483,1509]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="jaechi" subGenus="Cryobius">
|
||
<emphasis id="B97BB710FFFB8D7428CD7D98FEABF5D4" box="[199,338,1483,1509]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D74296D7D98FE37F5D4" box="[359,462,1483,1509]" italics="true" pageId="3" pageNumber="46">Cryobius</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D7429E87D98FDDFF5D4" box="[482,550,1483,1509]" italics="true" pageId="3" pageNumber="46">jaechi</emphasis>
|
||
<materialsCitation id="3B67615FFFFB8D742A387D98FD2EF5D4" box="[562,727,1483,1509]" collectingDate="1987-07-20" collectingDateMax="2017-07-09" collectingDateMin="1987-07-20" collectionCode="IZAS, BJFU" collectorName="Peiyu Yu & Hao Huang" county="Ningshan County" elevation="2500" latitude="33.4718" location="Ningshan County" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" pageId="3" pageNumber="46" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Jilin">Krischenhofer</materialsCitation>
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D742AEF7D98FCCDF5D4" box="[741,820,1483,1509]" count="1" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742B7E7D98FC20F5D4" box="[884,985,1483,1509]" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(IZAS), “
|
||
<collectingRegion id="49CBA5E0FFFB8D742C587D98FB7EF5D4" box="[1106,1159,1483,1509]" country="China" name="Jilin" pageId="3" pageNumber="46">Jilin</collectingRegion>
|
||
, Changbaishan,
|
||
<quantity id="4CF7C6E7FFFB8D742D4D7D99FA61F5D4" box="[1351,1432,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="46" unit="m" value="2500.0">
|
||
<elevation id="00228C31FFFB8D742D4D7D99FA61F5D4" box="[1351,1432,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="46" unit="m" value="2500.0">2500m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB14DC2FFFB8D74289D7DBCFEB5F638" box="[151,332,1519,1545]" pageId="3" pageNumber="46" value="1987-07-20" valueMax="1987-07-21" valueMin="1987-07-20">
|
||
<collectingDate id="EFF5B42AFFFB8D74289D7DBCFEB5F638" box="[151,332,1519,1545]" pageId="3" pageNumber="46" value="1987-07-20" valueMax="1987-07-21" valueMin="1987-07-20">1987.VII.20–21</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FA0ED4FFFB8D7429537DBCFE47F638" box="[345,446,1519,1545]" pageId="3" pageNumber="46">Peiyu Yu</collectorName>
|
||
leg.”.
|
||
<taxonomicName id="4C0F1081FFFB8D742A007DBCFB58F638" authority="Sciaky & Wrase" authorityName="Sciaky & Wrase" box="[522,1185,1519,1545]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="janatai" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742A007DBCFD6CF638" box="[522,661,1519,1545]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D742AAC7DBCFC87F638" box="[678,894,1519,1545]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D742B857DBCFC27F639" box="[911,990,1519,1544]" italics="true" pageId="3" pageNumber="46">janatai</emphasis>
|
||
Sciaky & Wrase
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D742CA67DBCFAFAF638" box="[1196,1283,1519,1545]" count="2" pageId="3" pageNumber="46" type="male">2 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742D307DBCFA65F638" box="[1338,1436,1519,1545]" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(BJFU), “
|
||
<collectingRegion id="49CBA5E0FFFB8D7429037E40FE91F61C" box="[265,360,1555,1581]" country="China" name="Shaanxi" pageId="3" pageNumber="46">Shaanxi</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D1138EFFFB8D7429727E40FDBDF61C" box="[376,580,1555,1581]" pageId="3" pageNumber="46">Ningshan County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BD4F178FFFB8D742A5E7E40FD24F61C" box="[596,733,1555,1581]" pageId="3" pageNumber="46">Pingheliang</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFB8D742AE77E40FCA6F61C" box="[749,863,1555,1581]" degrees="33.4718" direction="north" orientation="latitude" pageId="3" pageNumber="46" precision="5" value="33.4718">N33.4718</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFB8D742B7A7E40FC15F61C" box="[880,1004,1555,1581]" degrees="108.4961" direction="east" orientation="longitude" pageId="3" pageNumber="46" precision="5" value="108.4961">E108.4961</geoCoordinate>
|
||
,
|
||
<quantity id="4CF7C6E7FFFB8D742BF77E40FBB5F61C" box="[1021,1100,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="46" unit="m" value="2110.0">
|
||
<elevation id="00228C31FFFB8D742BF77E40FBB5F61C" box="[1021,1100,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="46" unit="m" value="2110.0">2110m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB14DC2FFFB8D742C577E40FB0EF61C" box="[1117,1271,1555,1581]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">
|
||
<collectingDate id="EFF5B42AFFFB8D742C577E40FB0EF61C" box="[1117,1271,1555,1581]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">2017.VII.2–9</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="524E1315FFFB8D742D0D7E40FEEDF660" pageId="3" pageNumber="46">Weifeng Yan pitfall trap</collectingMethod>
|
||
.”.
|
||
<taxonomicName id="4C0F1081FFFB8D7429247E64FC23F660" authority="Sciaky & Wrase" authorityName="Sciaky & Wrase" authorityYear="1997" box="[302,986,1591,1617]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="schuelkei" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D7429247E64FE40F660" box="[302,441,1591,1617]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D7429C37E64FD58F660" box="[457,673,1591,1617]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D742ABB7E64FCE0F660" box="[689,793,1591,1617]" italics="true" pageId="3" pageNumber="46">schuelkei</emphasis>
|
||
Sciaky & Wrase
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D742BEE7E64FBD6F660" box="[996,1071,1591,1617]" count="1" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742C6F7E64FB3FF660" box="[1125,1222,1591,1617]" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(BJFU), “
|
||
<collectingRegion id="49CBA5E0FFFB8D742D337E64FA61F660" box="[1337,1432,1591,1617]" country="China" name="Shaanxi" pageId="3" pageNumber="46">Shaanxi</collectingRegion>
|
||
,
|
||
<location id="8ED03DD9FFFB8D74289D7E08FE98F644" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D74289D7E08FE98F644" box="[151,353,1627,1653]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Ningshan County" pageId="3" pageNumber="46" stateProvince="Jilin">Ningshan County</location>
|
||
,
|
||
<location id="8ED03DD9FFFB8D74297A7E08FE10F644" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D74297A7E08FE10F644" box="[368,489,1627,1653]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Huoditang" pageId="3" pageNumber="46" stateProvince="Jilin">Huoditang</location>
|
||
,
|
||
<date id="FFB14DC2FFFB8D7429F27E08FD68F644" box="[504,657,1627,1653]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">
|
||
<collectingDate id="EFF5B42AFFFB8D7429F27E08FD68F644" box="[504,657,1627,1653]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">2017.VII.2–9</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8ED03DD9FFFB8D742AAA7E08FCCAF644" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742AAA7E08FCCAF644" box="[672,819,1627,1653]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Weifeng Yan" pageId="3" pageNumber="46" stateProvince="Jilin">Weifeng Yan</location>
|
||
<emphasis id="B97BB710FFFB8D742B347E08FC83F644" box="[830,890,1627,1653]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
<collectingMethod id="524E1315FFFB8D742B8E7E08FBFAF644" box="[900,1027,1627,1653]" pageId="3" pageNumber="46">pitfall trap</collectingMethod>
|
||
”.
|
||
<taxonomicName id="4C0F1081FFFB8D742C167E08FE36F6A8" authority="Sciaky & Wrase" authorityName="Sciaky & Wrase" authorityYear="1997" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="lingshanus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742C167E08FB5EF644" box="[1052,1191,1627,1653]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D742CB17E08FA6AF644" box="[1211,1427,1627,1653]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D74289D7E2CFEE9F6A8" box="[151,272,1663,1689]" italics="true" pageId="3" pageNumber="46">lingshanus</emphasis>
|
||
Sciaky & Wrase
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D7429D37E2CFDDAF6A8" box="[473,547,1663,1689]" count="1" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742A527E2CFD41F6A8" box="[600,696,1663,1689]" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(BJFU), “
|
||
<collectingRegion id="49CBA5E0FFFB8D742B207E2CFC70F6A8" box="[810,905,1663,1689]" country="China" name="Shaanxi" pageId="3" pageNumber="46">Shaanxi</collectingRegion>
|
||
,
|
||
<location id="8ED03DD9FFFB8D742B997E2CFBF8F6A8" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742B997E2CFBF8F6A8" box="[915,1025,1663,1689]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Ningshan" pageId="3" pageNumber="46" stateProvince="Jilin">Ningshan</location>
|
||
,
|
||
<location id="8ED03DD9FFFB8D742C067E2CFB6CF6A8" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742C067E2CFB6CF6A8" box="[1036,1173,1663,1689]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Pingheliang" pageId="3" pageNumber="46" stateProvince="Jilin">Pingheliang</location>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFB8D742C957E2CFAE9F6A8" box="[1183,1296,1663,1689]" degrees="33.4718" direction="north" orientation="latitude" pageId="3" pageNumber="46" precision="5" value="33.4718">N33.4718</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFB8D742D117E2CFA61F6A8" box="[1307,1432,1663,1689]" degrees="108.4961" direction="east" orientation="longitude" pageId="3" pageNumber="46" precision="5" value="108.4961">E108.4961</geoCoordinate>
|
||
,
|
||
<quantity id="4CF7C6E7FFFB8D74289D7EF0FF1FF68C" box="[151,230,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="46" unit="m" value="2110.0">
|
||
<elevation id="00228C31FFFB8D74289D7EF0FF1FF68C" box="[151,230,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="46" unit="m" value="2110.0">2110m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB14DC2FFFB8D7428FE7EF0FE74F68C" box="[244,397,1699,1725]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">
|
||
<collectingDate id="EFF5B42AFFFB8D7428FE7EF0FE74F68C" box="[244,397,1699,1725]" pageId="3" pageNumber="46" value="2017-07-02" valueMax="2017-07-09" valueMin="2017-07-02">2017.VII.2–9</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="524E1315FFFB8D7429937EF0FD4BF68C" box="[409,690,1699,1725]" pageId="3" pageNumber="46">Weifeng Yan pitfall trap</collectingMethod>
|
||
.”.
|
||
<taxonomicName id="4C0F1081FFFB8D742AC47EF0FB05F68C" authority="Sciaky" authorityName="Sciaky, Houzhenzi" authorityYear="1994" box="[718,1276,1699,1725]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="dundai" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742AC47EF0FCA0F68C" box="[718,857,1699,1725]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D742B607EF0FBBBF68C" box="[874,1090,1699,1725]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D742C5F7EF0FB5AF68C" box="[1109,1187,1699,1725]" italics="true" pageId="3" pageNumber="46">dundai</emphasis>
|
||
Sciaky
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D742D027EF0FAADF68C" box="[1288,1364,1699,1725]" count="1" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742D847EF0FF1AF6D0" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(IZAS), “
|
||
<collectingRegion id="49CBA5E0FFFB8D7429537E94FE41F6D0" box="[345,440,1735,1761]" country="China" name="Shaanxi" pageId="3" pageNumber="46">Shaanxi</collectingRegion>
|
||
,
|
||
<quantity id="4CF7C6E7FFFB8D7429CC7E94FE0EF6D0" box="[454,503,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="46" unit="km" value="3.0">3km</quantity>
|
||
of
|
||
<location id="8ED03DD9FFFB8D742A287E94FD62F6D0" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742A287E94FD62F6D0" box="[546,667,1735,1761]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Houzhenzi" pageId="3" pageNumber="46" stateProvince="Jilin">Houzhenzi</location>
|
||
to old
|
||
<location id="8ED03DD9FFFB8D742AF87E94FC42F6D0" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742AF87E94FC42F6D0" box="[754,955,1735,1761]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Foping Township" pageId="3" pageNumber="46" stateProvince="Jilin">Foping Township</location>
|
||
,
|
||
<date id="FFB14DC2FFFB8D742BC37E94FBD6F6D0" box="[969,1071,1735,1761]" pageId="3" pageNumber="46" value="2008-05-09">
|
||
<collectingDate id="EFF5B42AFFFB8D742BC37E94FBD6F6D0" box="[969,1071,1735,1761]" pageId="3" pageNumber="46" value="2008-05-09">2008.V.9</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FA0ED4FFFB8D742C347E94FB3BF6D0" box="[1086,1218,1735,1761]" pageId="3" pageNumber="46">Hao Huang</collectorName>
|
||
leg.”.
|
||
<taxonomicName id="4C0F1081FFFB8D742D1B7E94FD65F734" authority="Sciaky & Wrase" authorityName="Sciaky & Wrase" authorityYear="1997" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="geberti" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFFB8D742D1B7E94FA65F6D0" box="[1297,1436,1735,1761]" italics="true" pageId="3" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFB8D7428AB7EB8FE80F734" box="[161,377,1771,1797]" italics="true" pageId="3" pageNumber="46">Morphohaptoderus</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFB8D7429807EB8FE20F734" box="[394,473,1771,1797]" italics="true" pageId="3" pageNumber="46">geberti</emphasis>
|
||
Sciaky & Wrase
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D09A08BFFFB8D742AAD7EB8FD0AF734" box="[679,755,1771,1797]" count="1" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFB8D742B217EB8FC74F734" box="[811,909,1771,1797]" count="1" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
|
||
(IZAS), “
|
||
<collectingRegion id="49CBA5E0FFFB8D742BF47EB8FBA4F734" box="[1022,1117,1771,1797]" country="China" name="Shaanxi" pageId="3" pageNumber="46">Shaanxi</collectingRegion>
|
||
,
|
||
<location id="8ED03DD9FFFB8D742C637EB8FB30F734" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742C637EB8FB30F734" box="[1129,1225,1771,1797]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Zhouzhi" pageId="3" pageNumber="46" stateProvince="Jilin">Zhouzhi</location>
|
||
,
|
||
<location id="8ED03DD9FFFB8D742CDF7EB8FAA8F734" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D742CDF7EB8FAA8F734" box="[1237,1361,1771,1797]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Houzhenzi" pageId="3" pageNumber="46" stateProvince="Jilin">Houzhenzi</location>
|
||
,
|
||
<date id="FFB14DC2FFFB8D742D577EB8FF34F718" pageId="3" pageNumber="46" value="2007-05-25">
|
||
<collectingDate id="EFF5B42AFFFB8D742D577EB8FF34F718" pageId="3" pageNumber="46" value="2007-05-25">2007. V.25</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8ED03DD9FFFB8D7428D37F5CFE82F718" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFB8D7428D37F5CFE82F718" box="[217,379,1807,1833]" county="Ningshan County" latitude="33.4718" longLatPrecision="6" longitude="108.4961" municipality="Pingheliang" name="Hongliang Shi" pageId="3" pageNumber="46" stateProvince="Jilin">Hongliang Shi</location>
|
||
lgt.”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFB8D73289D7F28FA65F3D3" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46" type="key">
|
||
<paragraph id="8BB06B02FFFB8D74289D7F28FCCDF7A5" blockId="3.[151,820,1914,1941]" box="[151,820,1914,1941]" pageId="3" pageNumber="46">
|
||
<emphasis id="B97BB710FFFB8D74289D7F28FCCDF7A5" bold="true" box="[151,820,1914,1941]" pageId="3" pageNumber="46">
|
||
Key to species of subgenus
|
||
<taxonomicName id="4C0F1081FFFB8D7429E37F29FCCDF7A5" authority="Berg, 1898" authorityName="Berg" authorityYear="1898" box="[489,820,1914,1941]" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFB8D7429E37F29FD5EF7A4" bold="true" box="[489,679,1914,1941]" italics="true" pageId="3" pageNumber="46">Tschitscherinea</emphasis>
|
||
<bibRefCitation id="EF9E16F3FFFB8D742AA47F28FCCDF7A5" author="Berg, C." box="[686,820,1914,1941]" pageId="3" pageNumber="46" pagination="16 - 19" refId="ref15240" refString="Berg, C. (1898) Substitucion de nombres genericos. Comunicaciones del Museo Nacional de Buenos Aires, 1, 16 - 19." type="journal article" year="1898">Berg, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<keyStep id="30FB73A7FFFB8D74289D7F93FA65F7C3" pageId="3" pageNumber="46">
|
||
<paragraph id="8BB06B02FFFB8D74289D7F93FA65F7E5" blockId="3.[151,1436,1983,2035]" box="[151,1436,1983,2006]" pageId="3" pageNumber="46">
|
||
<keyLead id="30FEC837FFFB8D74289D7F93FA65F7E5" box="[151,1436,1983,2006]" pageId="3" pageNumber="46">
|
||
1 Elytra with isodiametric microsculpture, length of each mesh less than 1.5 times than width (
|
||
<figureCitation id="13347787FFFB8D742C567F93FB33F7E7" box="[1116,1226,1984,2006]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="3" pageNumber="46">Figs 44, 46</figureCitation>
|
||
)...............
|
||
<emphasis id="B97BB710FFFB8D742D9A7FECFA65F7E5" bold="true" box="[1424,1436,1983,2004]" pageId="3" pageNumber="46">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFB8D74289D7F8EFA65F7C3" blockId="3.[151,1436,1983,2035]" box="[151,1436,2013,2035]" pageId="3" pageNumber="46">
|
||
<keyLead id="30FEC837FFFB8D74289D7F8EFA65F7C3" box="[151,1436,2013,2035]" pageId="3" pageNumber="46">
|
||
- Elytra with transverse microsculpture, length of each mesh more than 2 times than width (
|
||
<figureCitation id="13347787FFFB8D742C487F8EFB49F7C3" box="[1090,1200,2013,2035]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="3" pageNumber="46">Figs 45, 47</figureCitation>
|
||
)..................
|
||
<emphasis id="B97BB710FFFB8D742D9A7F8EFA65F7C3" bold="true" box="[1424,1436,2013,2034]" pageId="3" pageNumber="46">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D78CBFA65F113" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D78CBFA65F0D9" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D78CBFA65F0D9" pageId="4" pageNumber="47">
|
||
2 Elytral interval 3 with three setigerous pores, the first one adjacent to stria 3, the rest two adjacent to stria 2; eyes large, tempora shorter than eyes; male metatarsus slender, tarsomere 1 with length/width = 3.1–3.5 (
|
||
<figureCitation id="13347787FFFC8D732C1878E6FBA3F0FB" box="[1042,1114,181,203]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="4" pageNumber="47">Fig. 41</figureCitation>
|
||
);
|
||
<collectingRegion id="49CBA5E0FFFC8D732C6178E6FB4DF0FB" box="[1131,1204,181,202]" country="China" name="Yunnan" pageId="4" pageNumber="47">Yunnan</collectingRegion>
|
||
(Zhongdian)..............................................................................................
|
||
<taxonomicName id="4C0F1081FFFC8D732CFE7881FA65F0D9" ID-CoL="4Q4HD" authority="Sciaky" authorityName="Sciaky" box="[1268,1436,210,232]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="farkaci">
|
||
<emphasis id="B97BB710FFFC8D732CFE7881FA65F0D9" bold="true" box="[1268,1436,210,232]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97BB710FFFC8D732CFE7881FAAAF0D9" bold="true" box="[1268,1363,210,232]" italics="true" pageId="4" pageNumber="47">P. farkaci</emphasis>
|
||
Sciaky
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D78A3FA65F113" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D78A3FA65F113" pageId="4" pageNumber="47">
|
||
- Elytral interval 3 with two setigerous pores, both adjacent to stria 2; eyes small, tempora swollen, longer than eyes; male metatarsus in a stout form, tarsomere 1 with length/width = 1.7–2.2 (
|
||
<figureCitation id="13347787FFFC8D732B7C795EFBF1F113" box="[886,1032,269,291]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="4" pageNumber="47">Figs 35–38, 43</figureCitation>
|
||
)................................
|
||
<emphasis id="B97BB710FFFC8D732D9A795EFA65F113" bold="true" box="[1424,1436,269,290]" pageId="4" pageNumber="47">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D7979FA65F1A6" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D7979FA65F16C" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7979FA65F16C" pageId="4" pageNumber="47">
|
||
3 Body larger, BL =
|
||
<quantity id="4CF7C6E7FFFC8D7329997979FDD8F171" box="[403,545,298,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="4" pageNumber="47" unit="mm" value="15.15" valueMax="15.5" valueMin="14.8">14.8–15.5 mm</quantity>
|
||
; head very large, HW/PW = 0.78–0.81; basal foveae with inner groove about 1.5 times length as outer groove;
|
||
<collectingRegion id="49CBA5E0FFFC8D732988791BFE32F16C" box="[386,459,328,349]" country="China" name="Yunnan" pageId="4" pageNumber="47">Yunnan</collectingRegion>
|
||
(Ninglang),
|
||
<collectingRegion id="49CBA5E0FFFC8D732A4C7914FD6AF16C" box="[582,659,327,349]" country="China" name="Sichuan" pageId="4" pageNumber="47">Sichuan</collectingRegion>
|
||
(Yanyuan)........................................
|
||
<emphasis id="B97BB710FFFC8D732CE8791BFA65F16C" bold="true" box="[1250,1436,327,349]" pageId="4" pageNumber="47">
|
||
<taxonomicName id="4C0F1081FFFC8D732CE8791BFA98F16C" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[1250,1377,327,349]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. n.">
|
||
<emphasis id="B97BB710FFFC8D732CE8791BFA98F16C" bold="true" box="[1250,1377,327,349]" italics="true" pageId="4" pageNumber="47">P. spectabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2480A6BFFFC8D732D6C791BFA65F16C" box="[1382,1436,328,349]" pageId="4" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D7936FA65F1A6" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7936FA65F1A6" pageId="4" pageNumber="47">
|
||
- Body smaller, BL =
|
||
<quantity id="4CF7C6E7FFFC8D7329AD7936FDCDF14B" box="[423,564,357,379]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.16" metricValueMin="1.03" pageId="4" pageNumber="47" unit="mm" value="10.95" valueMax="11.6" valueMin="10.3">10.3–11.6 mm</quantity>
|
||
; head smaller, HW/PW = 0.64–0.69; basal foveae with inner groove about twice length as outer groove.........................................................................................
|
||
<emphasis id="B97BB710FFFC8D732D9A79D1FA65F1A6" bold="true" box="[1424,1436,386,407]" pageId="4" pageNumber="47">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D79F3FA65F279" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D79F3FA65F1DE" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D79F3FA65F1DE" pageId="4" pageNumber="47">
|
||
4 Pronotum more quadrate, lateral margins subparallel near middle; elytra parascutellar pore present; maxim width of head at outer margin of eyes; right paramere of male genitalia curved with apex truncated (
|
||
<figureCitation id="13347787FFFC8D732C1479EEFB69F1E2" box="[1054,1168,445,467]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="4" pageNumber="47">Figs 16–17</figureCitation>
|
||
); male sternite VII without modification;
|
||
<collectingRegion id="49CBA5E0FFFC8D7329627988FE48F1C1" box="[360,433,475,496]" country="China" name="Yunnan" pageId="4" pageNumber="47">Yunnan</collectingRegion>
|
||
(Lijiang, Dali, Deqin).................................................
|
||
<emphasis id="B97BB710FFFC8D732CD77989FA65F1DE" bold="true" box="[1245,1436,474,496]" pageId="4" pageNumber="47">
|
||
<taxonomicName id="4C0F1081FFFC8D732CD77989FA99F1C1" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[1245,1376,474,496]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="spissipedes" status="sp. n.">
|
||
<emphasis id="B97BB710FFFC8D732CD77989FA99F1C1" bold="true" box="[1245,1376,474,496]" italics="true" pageId="4" pageNumber="47">P. spissipedes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2480A6BFFFC8D732D6C7989FA65F1DE" box="[1382,1436,474,495]" pageId="4" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D79ABFA65F279" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D79ABFA65F279" pageId="4" pageNumber="47">
|
||
- Pronotum more ovate, lateral margins distinctly arched near middle; elytra parascutellar pore absent; maxim width of head at outer margin of tempora; right paramere of male genitalia nearly straight with apex relatively sharp (
|
||
<figureCitation id="13347787FFFC8D732CA87A46FB11F21A" box="[1186,1256,533,555]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="4" pageNumber="47">Fig. 15</figureCitation>
|
||
); male sternite VII with a round depression (
|
||
<figureCitation id="13347787FFFC8D7329DF7A61FDE4F276" box="[469,541,562,584]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="4" pageNumber="47">Fig. 49</figureCitation>
|
||
);
|
||
<collectingRegion id="49CBA5E0FFFC8D732A247A60FD8EF279" box="[558,631,563,584]" country="China" name="Yunnan" pageId="4" pageNumber="47">Yunnan</collectingRegion>
|
||
(Dayao)............................................
|
||
<taxonomicName id="4C0F1081FFFC8D732CED7A61FA65F279" ID-CoL="4Q3TW" authority="Jedlicka" authorityName="Jedlicka" box="[1255,1436,562,584]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="sterbai">
|
||
<emphasis id="B97BB710FFFC8D732CED7A61FA65F279" bold="true" box="[1255,1436,562,584]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97BB710FFFC8D732CED7A61FAB8F279" bold="true" box="[1255,1345,562,584]" italics="true" pageId="4" pageNumber="47">P. sterbai</emphasis>
|
||
Jedlička
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D7A1CFA65F2B3" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D7A1CFA65F255" blockId="4.[151,1437,151,994]" box="[151,1436,591,613]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7A1CFA65F255" box="[151,1436,591,613]" pageId="4" pageNumber="47">
|
||
5 Elytral parascutellar pore present, occasionally absent on one side; pronotum basal foveae almost impunctate............
|
||
<emphasis id="B97BB710FFFC8D732D9A7A1CFA65F255" bold="true" box="[1424,1436,591,612]" pageId="4" pageNumber="47">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D7A3EFA65F2B3" blockId="4.[151,1437,151,994]" box="[151,1436,621,643]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7A3EFA65F2B3" box="[151,1436,621,643]" pageId="4" pageNumber="47">
|
||
- Elytral parascutellar pore absent; pronotum basal foveae distinctly punctate.......................................
|
||
<emphasis id="B97BB710FFFC8D732D9A7A3EFA65F2B3" bold="true" box="[1424,1436,621,642]" pageId="4" pageNumber="47">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D7AD9FA65F302" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D7AD9FA65F2EA" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7AD9FA65F2EA" pageId="4" pageNumber="47">
|
||
6 Elytral microsculpture stronger, transverse-isodiametric, length of each mesh usually 2–3 times as width (
|
||
<figureCitation id="13347787FFFC8D732D0F7AD9FAA9F291" box="[1285,1360,650,672]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="4" pageNumber="47">Fig. 45</figureCitation>
|
||
); elytra humeral tooth smaller (
|
||
<figureCitation id="13347787FFFC8D7329CE7AF4FDE4F28C" box="[452,541,679,701]" captionStart="FIGURES 7–12" captionStartId="14.[152,270,1814,1838]" captionTargetBox="[158,1429,188,1782]" captionTargetId="figure-36@14.[151,1436,181,1790]" captionTargetPageId="14" captionText="FIGURES 7–12. Habitus of Pterostichus (Tschitscherinea) spp. (7) P. mulensis Sciaky, a male from Muli. (8) P. mulensis Sciaky, holotype. (9) P. krali Sciaky, holotype. (10) P. farkaci Sciaky, a male from Tianchi, Zhongdian. (11) P. farkaci Sciaky, a male from Hongshan mountain, Zhongdian. (12) P. farkaci Sciaky, a male paratype from west of Zhongdian. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742728" httpUri="https://zenodo.org/record/12742728/files/figure.png" pageId="4" pageNumber="47">Figs 7–8</figureCitation>
|
||
); right paramere of male genitalia curved with the maximum curvature near apical fourth of apical portion (
|
||
<figureCitation id="13347787FFFC8D73297E7A96FE45F2EB" box="[372,444,709,731]" captionStart="FIGURES 19–24" captionStartId="16.[152,270,1691,1715]" captionTargetBox="[158,1429,188,1658]" captionTargetId="figure-36@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742736" httpUri="https://zenodo.org/record/12742736/files/figure.png" pageId="4" pageNumber="47">Fig. 20</figureCitation>
|
||
);
|
||
<collectingRegion id="49CBA5E0FFFC8D7329C77A96FDE3F2EA" box="[461,538,709,731]" country="China" name="Sichuan" pageId="4" pageNumber="47">Sichuan</collectingRegion>
|
||
(Muli)......................................................
|
||
<taxonomicName id="4C0F1081FFFC8D732CE97A96FA65F2EA" ID-CoL="4Q4HG" authority="Sciaky" authorityName="Sciaky" box="[1251,1436,708,731]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="mulensis">
|
||
<emphasis id="B97BB710FFFC8D732CE97A96FA65F2EA" bold="true" box="[1251,1436,708,731]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97BB710FFFC8D732CE97A96FAABF2EB" bold="true" box="[1251,1362,708,730]" italics="true" pageId="4" pageNumber="47">P. mulensis</emphasis>
|
||
Sciaky
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D7AB0FA65F302" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7AB0FA65F302" pageId="4" pageNumber="47">
|
||
- Elytral microsculpture shallower, transverse-linear, length of each mesh usually more than 4 times as width (
|
||
<figureCitation id="13347787FFFC8D732D037AB1FAAAF2C6" box="[1289,1363,738,760]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="4" pageNumber="47">Fig. 47</figureCitation>
|
||
); elytra humeral tooth larger (
|
||
<figureCitation id="13347787FFFC8D7329B07AACFE01F324" box="[442,504,767,789]" captionStart="FIGURES 7–12" captionStartId="14.[152,270,1814,1838]" captionTargetBox="[158,1429,188,1782]" captionTargetId="figure-36@14.[151,1436,181,1790]" captionTargetPageId="14" captionText="FIGURES 7–12. Habitus of Pterostichus (Tschitscherinea) spp. (7) P. mulensis Sciaky, a male from Muli. (8) P. mulensis Sciaky, holotype. (9) P. krali Sciaky, holotype. (10) P. farkaci Sciaky, a male from Tianchi, Zhongdian. (11) P. farkaci Sciaky, a male from Hongshan mountain, Zhongdian. (12) P. farkaci Sciaky, a male paratype from west of Zhongdian. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742728" httpUri="https://zenodo.org/record/12742728/files/figure.png" pageId="4" pageNumber="47">Fig. 9</figureCitation>
|
||
); right paramere curved with the maximum curvature near middle of apical portion (
|
||
<figureCitation id="13347787FFFC8D732D4C7AACFA68F324" box="[1350,1425,767,789]" captionStart="FIGURES 19–24" captionStartId="16.[152,270,1691,1715]" captionTargetBox="[158,1429,188,1658]" captionTargetId="figure-36@16.[151,1436,181,1668]" captionTargetPageId="16" captionText="FIGURES 19–24. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (19) P. jiuzhaiicus sp. nov., holotype. (20) P. mulensis Sciaky, holotype. (21) P. krali Sciaky, Haba Snow Mountain. (22) P. farkaci Sciaky, west of Zhongdian, paratype. (23) P. farkaci Sciaky, Tianchi, Zhongdian. (24) P. farkaci Sciaky, Hongshan, Zhongdian. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742736" httpUri="https://zenodo.org/record/12742736/files/figure.png" pageId="4" pageNumber="47">Fig. 21</figureCitation>
|
||
);
|
||
<collectingRegion id="49CBA5E0FFFC8D7328E87B4EFED2F303" box="[226,299,797,818]" country="China" name="Yunnan" pageId="4" pageNumber="47">Yunnan</collectingRegion>
|
||
(
|
||
<location id="8ED03DD9FFFC8D7329337B4EFE6FF303" LSID="urn:lsid:plazi:treatment:03A6DA14FFF98D73289D7D68FA65F3D3:8ED03DD9FFFC8D7329337B4EFE6FF303" box="[313,406,797,819]" name="Haba Mt." pageId="4" pageNumber="47" stateProvince="Yunnan">Haba Mt.</location>
|
||
).........................................................................
|
||
<taxonomicName id="4C0F1081FFFC8D732D067B4EFA65F302" ID-CoL="4Q4HF" authority="Sciaky" authorityName="Sciaky" box="[1292,1436,796,819]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="krali">
|
||
<emphasis id="B97BB710FFFC8D732D067B4EFA65F302" bold="true" box="[1292,1436,796,819]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97BB710FFFC8D732D067B4EFAAAF303" bold="true" box="[1292,1363,796,818]" italics="true" pageId="4" pageNumber="47">P. krali</emphasis>
|
||
Sciaky
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30FB73A7FFFC8D73289D7B68FA65F3D3" pageId="4" pageNumber="47">
|
||
<paragraph id="8BB06B02FFFC8D73289D7B68FA65F3BB" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7B68FA65F3BB" pageId="4" pageNumber="47">
|
||
7 Elytral interval 3 with one setigerous pore; eyes small, tempora strongly swollen and distinctly longer than eyes, head maxim width at outer margin of tempora; pronotum basal foveae without outer groove;
|
||
<collectingRegion id="49CBA5E0FFFC8D732BE97B04FBC9F35C" box="[995,1072,855,877]" country="China" name="Sichuan" pageId="4" pageNumber="47">Sichuan</collectingRegion>
|
||
(Baoxing, Wenchuan)............................................................................................
|
||
<taxonomicName id="4C0F1081FFFC8D732CAC7B26FA65F3BB" ID-CoL="3DZ83" authority="(Tschitscherine)" baseAuthorityName="Tschitscherine" box="[1190,1436,884,906]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="filum">
|
||
<emphasis id="B97BB710FFFC8D732CAC7B26FA65F3BB" bold="true" box="[1190,1436,884,906]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97BB710FFFC8D732CAC7B26FB0AF3BB" bold="true" box="[1190,1267,884,906]" italics="true" pageId="4" pageNumber="47">P. filum</emphasis>
|
||
(Tschitschérine)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFC8D73289D7BC0FA65F3D3" blockId="4.[151,1437,151,994]" pageId="4" pageNumber="47">
|
||
<keyLead id="30FEC837FFFC8D73289D7BC0FA65F3D3" pageId="4" pageNumber="47">
|
||
- Elytral interval 3 with three setigerous pores; eyes large, tempora shorter than eyes, head maxim width at outer margin of eyes; pronotum basal foveae with inner and outer grooves well defined;
|
||
<collectingRegion id="49CBA5E0FFFC8D732B6A7BFCFC54F3F4" box="[864,941,943,965]" country="China" name="Sichuan" pageId="4" pageNumber="47">Sichuan</collectingRegion>
|
||
(Jiuzhaigou, Abaco),
|
||
<collectingRegion id="49CBA5E0FFFC8D732C8B7BE3FB47F3F4" box="[1153,1214,944,965]" country="China" name="Gansu" pageId="4" pageNumber="47">Gansu</collectingRegion>
|
||
(Bailongjiang)..........................................................................................
|
||
<emphasis id="B97BB710FFFC8D732CD77B9EFA65F3D3" bold="true" box="[1245,1436,972,994]" pageId="4" pageNumber="47">
|
||
<taxonomicName id="4C0F1081FFFC8D732CD77B9EFA99F3D3" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[1245,1376,972,994]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="jiuzhaiicus" status="sp. n.">
|
||
<emphasis id="B97BB710FFFC8D732CD77B9EFA99F3D3" bold="true" box="[1245,1376,972,994]" italics="true" pageId="4" pageNumber="47">P. jiuzhaiicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2480A6BFFFC8D732D6C7B9EFA65F3D3" box="[1382,1436,973,994]" pageId="4" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |