122 lines
19 KiB
XML
122 lines
19 KiB
XML
<document id="AE5A67BECAE468A003BFAACA8388495B" ID-DOI="10.5281/zenodo.210778" ID-GBIF-Dataset="b5ccb71c-4f14-426e-a93a-0745da86c76b" ID-ISSN="1175-5326" ID-Zenodo-Dep="210778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460554943001" checkinUser="plazi" docAuthor="Maruyama, Munetoshi" docDate="2012" docId="9F698793A83CFFA1FF7CFE206B428085" docLanguage="en" docName="zt03555p088.pdf" docOrigin="Zootaxa 3555" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eocorythoderus Maruyama, 2012, new genus" docType="treatment" docVersion="7" lastPageNumber="84" masterDocId="6350FFEBA83DFFA0FFEBFFD36A598543" masterDocTitle="A new genus and species of flightless, microphthalmic Corythoderini (Coleoptera: Scarabaeidae: Aphodiinae) from Cambodia, associated with Macrotermes termites" masterLastPageNumber="88" masterPageNumber="83" pageNumber="84" updateTime="1698319025298" updateUser="plazi">
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<mods:titleInfo id="B93FBB2923BF961A67F35835764F94E6">
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<mods:title id="77BF210F9C1DF3EC39972D4BB12536E8">A new genus and species of flightless, microphthalmic Corythoderini (Coleoptera: Scarabaeidae: Aphodiinae) from Cambodia, associated with Macrotermes termites</mods:title>
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<mods:namePart id="073FF10F8555196A5127166A3111DB25">Maruyama, Munetoshi</mods:namePart>
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<mods:title id="3FAD5FCA5D90786A9054FDC0AE813BF5">Zootaxa</mods:title>
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<mods:part id="BDEF5CB2804168D3183FACEECBEDF367">
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<mods:date id="DE8F4BA0598090BC5C87E75BD86115F0">2012</mods:date>
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<mods:detail id="73D3CD1EE8289D0E8A904B75DEAE11C0" type="volume">
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<mods:number id="18C7BA08DE2D3389D8C9A24091B5DF3B">3555</mods:number>
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<mods:start id="48BF060EA4012A1D2699A0EE6315A515">83</mods:start>
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<mods:identifier id="62792F100E17FCE467C67BAFDFE02710" type="DOI">10.5281/zenodo.210778</mods:identifier>
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<mods:identifier id="43E721DF4C70E835FD32A5F0DEB7A1EE" type="GBIF-Dataset">b5ccb71c-4f14-426e-a93a-0745da86c76b</mods:identifier>
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<mods:identifier id="FCFB765754DFE033222D1585DCFE83AE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9F698793A83CFFA1FF7CFE206B428085" ID-DOI="http://doi.org/10.5281/zenodo.6173518" ID-GBIF-Taxon="119555580" ID-Zenodo-Dep="6173518" LSID="urn:lsid:plazi:treatment:9F698793A83CFFA1FF7CFE206B428085" httpUri="http://treatment.plazi.org/id/9F698793A83CFFA1FF7CFE206B428085" lastPageNumber="84" pageId="1" pageNumber="84">
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<subSubSection id="5FDA650EA83CFFA1FF7CFE206B4B876C" pageId="1" pageNumber="84" type="nomenclature">
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<paragraph id="177F3685A83CFFA1FF7CFE206B89874F" blockId="1.[151,464,499,559]" box="[151,464,499,524]" pageId="1" pageNumber="84">
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<heading id="4C3781E9A83CFFA1FF7CFE206B89874F" bold="true" box="[151,464,499,524]" fontSize="10" level="2" pageId="1" pageNumber="84" reason="2">
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<emphasis id="25B4EA97A83CFFA1FF7CFE206B89874F" bold="true" box="[151,464,499,524]" pageId="1" pageNumber="84">
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<taxonomicName id="D0C04D06A83CFFA1FF7CFE206B16874F" box="[151,335,499,524]" class="Insecta" family="Aphodiidae" genus="Eocorythoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="25B4EA97A83CFFA1FF7CFE206B16874F" bold="true" box="[151,335,499,524]" italics="true" pageId="1" pageNumber="84">Eocorythoderus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3E8757ECA83CFFA1FEBEFE276B89874F" box="[341,464,500,524]" pageId="1" pageNumber="84" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="177F3685A83CFFA1FF7CFDC46B4B876C" blockId="1.[151,464,499,559]" box="[151,274,535,559]" pageId="1" pageNumber="84">
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(
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<figureCitation id="8FFB2A00A83CFFA1FF74FDC46B50876C" box="[159,265,535,559]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="2.[151,264,1971,1994]" captionStartId-1="3.[151,264,1875,1898]" captionTargetBox-0="[192,1411,176,1953]" captionTargetBox-1="[151,1435,193,1853]" captionTargetId-0="figure@2.[192,1416,173,1954]" captionTargetId-1="figure@3.[151,1436,193,1854]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." captionText-1="FIGURES 5 – 7. Eocorythoderus incredibilis new genus and new species. 5, Adult beetle being carried by a worker termite. 6, 7, Adult beetle (holotype) walking on a fungus garden." httpUri-0="https://zenodo.org/record/210779/files/figure.png" httpUri-1="https://zenodo.org/record/210780/files/figure.png" pageId="1" pageNumber="84">Figs. 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="5FDA650EA83CFFA1FF7CFD88699A8731" box="[151,963,602,627]" pageId="1" pageNumber="84" type="materials_examined">
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<paragraph id="177F3685A83CFFA1FF7CFD88699A8731" blockId="1.[151,1437,602,1478]" box="[151,963,602,627]" pageId="1" pageNumber="84">
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<emphasis id="25B4EA97A83CFFA1FF7CFD886B6B8730" bold="true" box="[151,306,602,627]" pageId="1" pageNumber="84">
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<typeStatus id="C87B8827A83CFFA1FF7CFD886A8B8730" box="[151,210,603,627]" pageId="1" pageNumber="84">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="D0C04D06A83CFFA1FED2FD88682B8730" box="[313,626,603,627]" class="Insecta" family="Aphodiidae" genus="Eocorythoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="incredibilis" status="sp. nov.">
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<emphasis id="25B4EA97A83CFFA1FED2FD88682B8730" box="[313,626,603,627]" italics="true" pageId="1" pageNumber="84">Eocorythoderus incredibilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3E8757ECA83CFFA1FD93FD88695B8731" box="[632,770,602,626]" pageId="1" pageNumber="84" rank="species">new species</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="5FDA650EA83CFFA1FF2CFDAD6B49879E" pageId="1" pageNumber="84" type="etymology">
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<paragraph id="177F3685A83CFFA1FF2CFDAD6B49879E" blockId="1.[151,1437,602,1478]" pageId="1" pageNumber="84">
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<emphasis id="25B4EA97A83CFFA1FF2CFDAD6B1787D5" bold="true" box="[199,334,638,662]" pageId="1" pageNumber="84">Etymology.</emphasis>
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A combination of
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<taxonomicName id="D0C04D06A83CFFA1FDD9FDAC680587D5" box="[562,604,639,662]" class="Aves" family="Psittacidae" genus="Eos" kingdom="Animalia" order="Psittaciformes" pageId="1" pageNumber="84" phylum="Chordata" rank="genus">
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<emphasis id="25B4EA97A83CFFA1FDD9FDAC680587D5" box="[562,604,639,662]" italics="true" pageId="1" pageNumber="84">Eos</emphasis>
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</taxonomicName>
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, the Greek goddess of the dawn, and
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<taxonomicName id="D0C04D06A83CFFA1FBC8FDAD6EE487D5" box="[1059,1213,638,662]" class="Insecta" family="Aphodiidae" genus="Corythoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
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<emphasis id="25B4EA97A83CFFA1FBC8FDAD6EE487D5" box="[1059,1213,638,662]" italics="true" pageId="1" pageNumber="84">Corythoderus</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="C87B8827A83CFFA1FB17FDAC6F7587D4" box="[1276,1324,639,663]" pageId="1" pageNumber="84">type</typeStatus>
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genus of Corythoderini, in reference to the new taxon having the eastern-most distribution of the genera in the tribe. Gender: masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="5FDA650EA83CFFA1FF2CFD3B6BBB8696" pageId="1" pageNumber="84" type="diagnosis">
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<paragraph id="177F3685A83CFFA1FF2CFD3B6BBB8696" blockId="1.[151,1437,602,1478]" pageId="1" pageNumber="84">
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<emphasis id="25B4EA97A83CFFA1FF2CFD3B6B188642" bold="true" box="[199,321,744,769]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
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The combination of the following character states diagnose
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<taxonomicName id="D0C04D06A83CFFA1FBE9FD3B6EEC8643" box="[1026,1205,744,768]" class="Insecta" family="Aphodiidae" genus="Eocorythoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
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<emphasis id="25B4EA97A83CFFA1FBE9FD3B6EEC8643" box="[1026,1205,744,768]" italics="true" pageId="1" pageNumber="84">Eocorythoderus</emphasis>
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</taxonomicName>
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: (1) body distinctly panduriform; (2) clypeal margin rounded; (3) eyes small, dorsally invisible; (4) pronotum with a pair of shallow depressions medially, (5) pronotum without median sulcus; (6) pronotum without trichome; (7) elytron with base of first and second intervals strongly elevated, to form a large median dorsal projection (
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<figureCitation id="8FFB2A00A83CFFA1FB99FC806EE08628" box="[1138,1209,851,875]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 2</figureCitation>
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: arrow); (8) elytron with basal knob-shaped protrusion; (9) elytron with 5 striae; (10) mesocoxal cavities widely separated; (11) mesolegs distinctly longer than others; (12, 13) mesofemora and metafemora broad, oval; (14, 15) mesotibiae and metatibiae narrowed apically.
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</paragraph>
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</subSubSection>
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<subSubSection id="5FDA650EA83CFFA1FF2CFC336B428085" pageId="1" pageNumber="84" type="description">
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<paragraph id="177F3685A83CFFA1FF2CFC336B428085" blockId="1.[151,1437,602,1478]" pageId="1" pageNumber="84">
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<emphasis id="25B4EA97A83CFFA1FF2CFC336B0086BA" bold="true" box="[199,345,992,1017]" pageId="1" pageNumber="84">Description.</emphasis>
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Body (
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<figureCitation id="8FFB2A00A83CFFA1FE47FC33684E86BB" box="[428,535,992,1016]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="2.[151,264,1971,1994]" captionStartId-1="3.[151,264,1875,1898]" captionTargetBox-0="[192,1411,176,1953]" captionTargetBox-1="[151,1435,193,1853]" captionTargetId-0="figure@2.[192,1416,173,1954]" captionTargetId-1="figure@3.[151,1436,193,1854]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." captionText-1="FIGURES 5 – 7. Eocorythoderus incredibilis new genus and new species. 5, Adult beetle being carried by a worker termite. 6, 7, Adult beetle (holotype) walking on a fungus garden." httpUri-0="https://zenodo.org/record/210779/files/figure.png" httpUri-1="https://zenodo.org/record/210780/files/figure.png" pageId="1" pageNumber="84">Figs. 1–7</figureCitation>
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) short, distinctly panduriform. Head (
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<figureCitation id="8FFB2A00A83CFFA1FC2FFC336E7686BB" box="[964,1071,992,1016]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Figs. 1–3</figureCitation>
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) slightly convex above; clypeus with gently broadened lateral margins; anterior margin rounded, slightly reflexed along anterior margin; eyes (
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<figureCitation id="8FFB2A00A83CFFA1FA85FBD76AFF8103" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) small, dorsally invisible. Pronotum (
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<figureCitation id="8FFB2A00A83CFFA1FD8DFBFB688C8103" box="[614,725,1064,1088]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Figs. 1–3</figureCitation>
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) strongly convex above, much higher than length; posterior impression shallow, short, extending from middle to base of median lobe; with a pair or shallow depressions medially; median lobe triangular, with dense setiferous punctures except for mesal area. Elytra (
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<figureCitation id="8FFB2A00A83CFFA1FB34FBBD6F1581C5" box="[1247,1356,1134,1158]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Figs. 1, 2</figureCitation>
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) short, oval, strongly convex dorsally, but lower than pronotum; disc with 5 striae; each elytron with large basal elevation on intervals I and II to form median projection, which is densely covered with small trichomes at apex; base of intervals III-V with two rather low tubercles to form basal knob-shaped protrusion. Mesoventrite (
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<figureCitation id="8FFB2A00A83CFFA1FAFBFB0A6F0E81B2" box="[1296,1367,1241,1265]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) with intercoxal area broad, distinctly margined laterally. Metaventrite (
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<figureCitation id="8FFB2A00A83CFFA1FC61FB2F69898057" box="[906,976,1276,1300]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) short, much shorter than mesoventrite. Proleg (
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<figureCitation id="8FFB2A00A83CFFA1FF1AFAF36B6E807B" box="[241,311,1312,1336]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) short; femur oblong oval; tibia short, bidentate near apex; tarsus slightly dilated apicad. Mesoleg (
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<figureCitation id="8FFB2A00A83CFFA1FF74FA906ABF8018" box="[159,230,1347,1371]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) stout, long, longer than metaleg; femur oval, broad, flattened, with posterior sulcus to hold tibia; tibia dilated apicad; tarsus narrowed apicad. Metaleg (
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<figureCitation id="8FFB2A00A83CFFA1FD27FAB46948803C" box="[716,785,1383,1407]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) similar in shape to mesoleg, but shorter; femur slightly constricted near base. Pygidium (
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<figureCitation id="8FFB2A00A83CFFA1FDCBFA59683080E1" box="[544,617,1418,1442]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[192,1411,176,1953]" captionTargetId="figure@2.[192,1416,173,1954]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Eocorythoderus incredibilis new genus and new species. 1, Holotype, dorsal view. 2, Paratype, lateral view. 3, Paratype, anterior view. 4, Paratype, ventral view." httpUri="https://zenodo.org/record/210779/files/figure.png" pageId="1" pageNumber="84">Fig. 4</figureCitation>
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) with a pair of shallow depressions (perhaps this character is sexual dimorphic).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |