236 lines
34 KiB
XML
236 lines
34 KiB
XML
<document id="930AE4B466594F122911A77494F95AAE" ID-CLB-Dataset="5420" ID-DOI="10.11646/zootaxa.3840.1.1" ID-GBIF-Dataset="1478a1d3-fcb4-43b3-9418-010043c5c154" ID-ISSN="1175-5326" ID-Zenodo-Dep="4927892" ID-ZooBank="CDF827EB-8A66-438F-AC5C-07A7D3D83BB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623431855578" checkinUser="felipe" docAuthor="Pešić, Vladimir & Smit, Harry" docDate="2014" docId="F535879E7E7BFFDEFF0DFF6BD716F8FD" docLanguage="en" docName="zt03840p072.pdf" docOrigin="Zootaxa 3840 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoatractides uniscutatus Pešić & Smit 2014, n. sp." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="090CFFE67E64FFC1FF9AFFB5D561FFFE" masterDocTitle="Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo" masterLastPageNumber="72" masterPageNumber="1" pageNumber="32" updateTime="1699010593832" updateUser="ExternalLinkService">
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<mods:mods id="66DD0E7AC8A20A9A61B1C296116AC5A9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A702F7107F790385F4A36C20CF1B3A34">
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<mods:title id="4FD602591413771945E8A9AB62371158">Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo</mods:title>
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<mods:name id="32380A31438C09C9CF711E18D407FFBF" type="personal">
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<mods:roleTerm id="3FDE37EC5FA4005B6A803062D220DBAC">Author</mods:roleTerm>
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<mods:namePart id="821BEB432CD358148F4C4A8077EAD750">Pešić, Vladimir</mods:namePart>
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<mods:nameIdentifier id="8F21090DCD0D0CD04342853F6FB3BB2A" type="email">vladopesic@gmail.com</mods:nameIdentifier>
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<mods:name id="C9BA1A22A9DC70807CDE6A3A06BA98F6" type="personal">
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<mods:roleTerm id="AC1C454E32EDA822A40EE56E246D4A4A">Author</mods:roleTerm>
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<mods:namePart id="908F6FB0D7BA87DA511FF9E486CF53CC">Smit, Harry</mods:namePart>
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<mods:nameIdentifier id="24FE4A72300CCF8733BDE06D5C83764E" type="email">vladopesic@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="91E28553B1DDB7C03C3E4ABF429C4C09">text</mods:typeOfResource>
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<mods:titleInfo id="2095C757E07D9B4AA0F5AC2AED575E23">
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<mods:title id="7E70048DD088564624B716219F013E28">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7EF4292FFD82519971653278DDB8F63B">
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<mods:date id="6FD66653F585BA99E714C672265F5786">2014</mods:date>
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<mods:number id="F71929578F2863D41470CDD4DB1D1C0C">2014-07-23</mods:number>
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<mods:detail id="FCD55162047567A34BF712071A3617D9" type="volume">
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<mods:number id="AC7247B6A90DDA022A1703B193DFB076">3840</mods:number>
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<mods:detail id="DC59C18B27B5BDC2DDDD13265FB60651" type="issue">
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<mods:start id="1D27C41222B2C921ADCBEAA02A4020A8">1</mods:start>
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<mods:classification id="567E5B446758444392D111EFBE6A6B36">journal article</mods:classification>
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<mods:identifier id="6B9A654A3AA9E5969AF9C625508194C2" type="DOI">10.11646/zootaxa.3840.1.1</mods:identifier>
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<mods:identifier id="A51E878F85B41CB0B81FE7B6BA688EEA" type="GBIF-Dataset">1478a1d3-fcb4-43b3-9418-010043c5c154</mods:identifier>
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<treatment id="F535879E7E7BFFDEFF0DFF6BD716F8FD" ID-DOI="http://doi.org/10.5281/zenodo.5110208" ID-GBIF-Taxon="183550658" ID-Zenodo-Dep="5110208" LSID="urn:lsid:plazi:treatment:F535879E7E7BFFDEFF0DFF6BD716F8FD" httpUri="http://treatment.plazi.org/id/F535879E7E7BFFDEFF0DFF6BD716F8FD" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="358665037E7BFFDEFF0DFF6BD778FF06" box="[151,537,222,248]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="7D2336887E7BFFDEFF0DFF6BD778FF06" blockId="31.[151,537,222,282]" box="[151,537,222,248]" pageId="31" pageNumber="32">
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<heading id="266B81E47E7BFFDEFF0DFF6BD778FF06" bold="true" box="[151,537,222,248]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
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<emphasis id="4FE8EA9A7E7BFFDEFF0DFF6BD778FF06" bold="true" box="[151,537,222,248]" pageId="31" pageNumber="32">
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<taxonomicName id="BA9C4D0B7E7BFFDEFF0DFF6BD4B1FF06" authority="Pešić & Smit, 2014" authorityName="Pešić & Smit" authorityYear="2014" box="[151,464,222,248]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="uniscutatus" status="sp. nov.">
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<emphasis id="4FE8EA9A7E7BFFDEFF0DFF6BD4B1FF06" bold="true" box="[151,464,222,248]" italics="true" pageId="31" pageNumber="32">Neoatractides uniscutatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="54DB57E17E7BFFDEFE4DFF6BD778FF06" box="[471,537,222,248]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="358665037E7BFFDEFF0DFEB4D4B1FEE4" box="[151,464,257,282]" pageId="31" pageNumber="32" type="description">
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<paragraph id="7D2336887E7BFFDEFF0DFEB4D4B1FEE4" blockId="31.[151,537,222,282]" box="[151,464,257,282]" pageId="31" pageNumber="32">
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(
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<figureCitation id="E5A72A0D7E7BFFDEFF05FEB4D44CFEE4" box="[159,301,257,282]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Figs. 20A–F</figureCitation>
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,
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<figureCitation id="E5A72A0D7E7BFFDEFEA2FEB4D4EDFEE4" box="[312,396,257,282]" captionStart="FIGURE 22" captionStartId="32.[151,250,1414,1436]" captionTargetBox="[151,1436,193,1392]" captionTargetId="figure-284@32.[151,1436,193,1392]" captionTargetPageId="32" captionText="FIGURE 22A–F. Photographs of dorsal (A,C,E) and ventral shield (B,D,F). A–B Neoatractides sundaensis n. sp., female holotype, Kemantis stream. C–D Neoatractides uniscutatus n. sp., male holotype, Kemantis stream. E–F Pseudotorrenticola borneoensis n. sp., male holotype, Little Lumotok stream." figureDoi="http://doi.org/10.5281/zenodo.4927959" httpUri="https://zenodo.org/record/4927959/files/figure.png" pageId="31" pageNumber="32">22C–D</figureCitation>
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,
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<figureCitation id="E5A72A0D7E7BFFDEFE0DFEB4D4A9FEE4" box="[407,456,257,282]" captionStart="FIGURE 23" captionStartId="33.[151,250,1268,1290]" captionTargetBox="[244,1342,397,1247]" captionTargetId="figure-364@33.[244,1342,397,1247]" captionTargetPageId="33" captionText="FIGURE 23A–H. Photographs of ejaculatory complex. A Torrenticola borneoensis n. sp. B T. kinabaluensis n. sp.. C T. longipalpis Wiles, 1997. D T. sabahensis n. sp. E Torrenticola minuta (Lundblad, 1941). F T. neoindica n. sp. G Neoatractides uniscutatus n. sp. H Pseudotorrenticola borneoensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4927961" httpUri="https://zenodo.org/record/4927961/files/figure.png" pageId="31" pageNumber="32">23G</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="358665037E7BFFDEFF0DFEFCD6C7FE78" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="7D2336887E7BFFDEFF0DFEFCD6C7FE78" blockId="31.[151,1437,329,1686]" pageId="31" pageNumber="32">
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<emphasis id="4FE8EA9A7E7BFFDEFF0DFEFCD47EFE9C" bold="true" box="[151,287,329,354]" pageId="31" pageNumber="32">Type series</emphasis>
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.
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<materialsCitation id="CDF43CD57E7BFFDEFEAAFEFCD6C7FE78" ID-GBIF-Occurrence="3323413318" collectingDate="2012-09-18" country="Malaysia" county="Crocker Range" latitude="5.8576" location="Borneo" municipality="Inobong" pageId="31" pageNumber="32" specimenCount="1" typeStatus="holotype">
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<typeStatus id="A227882A7E7BFFDEFEAAFEFCD4F9FE9C" box="[304,408,329,354]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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male, dissected and slide mounted,
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<collectingCountry id="058B76187E7BFFDEFCD9FEFCD6CCFE9C" box="[835,941,329,354]" name="Malaysia" pageId="31" pageNumber="32">Malaysia</collectingCountry>
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,
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<location id="784360537E7BFFDEFC21FEFFD171FE9F" LSID="urn:lsid:plazi:treatment:F535879E7E7BFFDEFF0DFF6BD716F8FD:784360537E7BFFDEFC21FEFFD171FE9F" box="[955,1040,330,353]" country="Malaysia" county="Crocker Range" latitude="5.8576" municipality="Inobong" name="Borneo" pageId="31" pageNumber="32">Borneo</location>
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, unnamed stream
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<location id="784360537E7BFFDEFB7CFEFCD0F9FE9C" LSID="urn:lsid:plazi:treatment:F535879E7E7BFFDEFF0DFF6BD716F8FD:784360537E7BFFDEFB7CFEFCD0F9FE9C" box="[1254,1432,329,354]" country="Malaysia" county="Crocker Range" latitude="5.8576" municipality="Inobong" name="Bansadon Trail" pageId="31" pageNumber="32">Bansadon Trail</location>
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,
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<collectingMunicipality id="9D47ACF27E7BFFDEFF0DFED8D596FE78" box="[151,247,365,390]" pageId="31" pageNumber="32">Inobong</collectingMunicipality>
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,
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<collectingCounty id="94424E047E7BFFDEFE98FED8D4CFFE78" box="[258,430,365,390]" pageId="31" pageNumber="32">Crocker Range</collectingCounty>
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,
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<geoCoordinate id="18A8504F7E7BFFDEFE23FED8D75AFE7B" box="[441,571,365,390]" degrees="5" direction="north" minutes="51.456" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="5.8576">5º51.456 N</geoCoordinate>
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,
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<geoCoordinate id="18A8504F7E7BFFDEFDDCFED8D7BEFE7B" box="[582,735,365,390]" pageId="31" pageNumber="32">116º68.403 E</geoCoordinate>
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,
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<date id="092210487E7BFFDEFD71FED8D603FE78" box="[747,866,365,390]" pageId="31" pageNumber="32" value="2012-09-18">
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<collectingDate id="1966E9A07E7BFFDEFD71FED8D603FE78" box="[747,866,365,390]" pageId="31" pageNumber="32" value="2012-09-18">18.ix.2012</collectingDate>
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</date>
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Smit.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="358665037E7BFFDEFF5DFE24D137FE0C" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="7D2336887E7BFFDEFF5DFE24D137FE0C" blockId="31.[151,1437,329,1686]" pageId="31" pageNumber="32">
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<emphasis id="4FE8EA9A7E7BFFDEFF5DFE24D45BFE54" bold="true" box="[199,314,401,426]" pageId="31" pageNumber="32">Diagnosis</emphasis>
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(Female unknown). Frontal and shoulder plates fused with the large dorsal plate, but suture lines well visible; excretory pore and Vgl-1 embedded in the area of primary sclerotization; gnathosomal rostrum broad and upturned, narrower than remainder of the gnathosoma; P-4 stout (dL/H ratio 2.0).
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</paragraph>
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</subSubSection>
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<subSubSection id="358665037E7BFFDEFF0DFE48D4F8FB60" pageId="31" pageNumber="32" type="description">
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<paragraph id="7D2336887E7BFFDEFF0DFE48D440FDE8" blockId="31.[151,1437,329,1686]" box="[151,289,509,534]" pageId="31" pageNumber="32">
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<emphasis id="4FE8EA9A7E7BFFDEFF0DFE48D440FDE8" bold="true" box="[151,289,509,534]" pageId="31" pageNumber="32">Description</emphasis>
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</paragraph>
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<paragraph id="7D2336887E7BFFDEFF5DFD94D6B9FC80" blockId="31.[151,1437,329,1686]" pageId="31" pageNumber="32">
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Male.
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<emphasis id="4FE8EA9A7E7BFFDEFE8BFD94D4AEFDC4" box="[273,463,545,570]" italics="true" pageId="31" pageNumber="32">General features</emphasis>
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—Frontal and shoulder plates fused with the large dorsal plate, but suture lines well visible (
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<figureCitation id="E5A72A0D7E7BFFDEFF68FDF0D405FDA0" box="[242,356,581,606]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Figs. 20A</figureCitation>
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,
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<figureCitation id="E5A72A0D7E7BFFDEFEEBFDF0D4C2FDA0" box="[369,419,581,606]" captionStart="FIGURE 22" captionStartId="32.[151,250,1414,1436]" captionTargetBox="[151,1436,193,1392]" captionTargetId="figure-284@32.[151,1436,193,1392]" captionTargetPageId="32" captionText="FIGURE 22A–F. Photographs of dorsal (A,C,E) and ventral shield (B,D,F). A–B Neoatractides sundaensis n. sp., female holotype, Kemantis stream. C–D Neoatractides uniscutatus n. sp., male holotype, Kemantis stream. E–F Pseudotorrenticola borneoensis n. sp., male holotype, Little Lumotok stream." figureDoi="http://doi.org/10.5281/zenodo.4927959" httpUri="https://zenodo.org/record/4927959/files/figure.png" pageId="31" pageNumber="32">22C</figureCitation>
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); Cxgl–4 posterior to Cxgl–2; medial suture line of Cx-II+III moderately long; suture line of Cx-IV in medial part directed posteriorly, laterally curved; genital field subrectangular in shape; ejaculatory complex as photographed in
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<figureCitation id="E5A72A0D7E7BFFDEFE43FD38D73DFD58" box="[473,604,653,678]" captionStart="FIGURE 23" captionStartId="33.[151,250,1268,1290]" captionTargetBox="[244,1342,397,1247]" captionTargetId="figure-364@33.[244,1342,397,1247]" captionTargetPageId="33" captionText="FIGURE 23A–H. Photographs of ejaculatory complex. A Torrenticola borneoensis n. sp. B T. kinabaluensis n. sp.. C T. longipalpis Wiles, 1997. D T. sabahensis n. sp. E Torrenticola minuta (Lundblad, 1941). F T. neoindica n. sp. G Neoatractides uniscutatus n. sp. H Pseudotorrenticola borneoensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4927961" httpUri="https://zenodo.org/record/4927961/files/figure.png" pageId="31" pageNumber="32">Figure 23G</figureCitation>
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; excretory pore and Vgl–2 embedded in the area of primary sclerotization; P-1 with dorsal seta, P-2 and -3 without ventral projections, P-2 long with relatively short and slender ventral seta, lateral distal margin of P-3 deeply indented, P-4 stout, longer than P-3, dorsal margin strongly bowed, ventral margin concave with one long and one shorter seta (
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<figureCitation id="E5A72A0D7E7BFFDEFD7EFD4CD618FCEC" box="[740,889,761,786]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Figs. 20C–D</figureCitation>
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). Gnathosoma without conspicuous oral papillae and short posterodorsal projections, rostrum broad and upturned, narrower than remainder of the gnathosoma (
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<figureCitation id="E5A72A0D7E7BFFDEFAF5FCA8D5A9FCA4" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Fig. 20E</figureCitation>
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); chelicera long with relatively short cheliceral claw (L basal segment/claw ratio 4.4). Legs: I-L with I-L-6 stout, L/H ratio 2.0, ventral margin distally strongly protruding (
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<figureCitation id="E5A72A0D7E7BFFDEFCFDFCD0D6AAFC80" box="[871,971,869,894]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Fig. 20F</figureCitation>
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).
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</paragraph>
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<paragraph id="7D2336887E7BFFDEFF5DFC3FD495FB84" blockId="31.[151,1437,329,1686]" pageId="31" pageNumber="32">
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<emphasis id="4FE8EA9A7E7BFFDEFF5DFC3FD408FC5F" box="[199,361,906,929]" italics="true" pageId="31" pageNumber="32">Measurements</emphasis>
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—Idiosoma (ventral view:
|
||
<figureCitation id="E5A72A0D7E7BFFDEFD0DFC3CD667FC5C" box="[663,774,905,930]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Figs. 20B</figureCitation>
|
||
,
|
||
<figureCitation id="E5A72A0D7E7BFFDEFC8BFC3CD622FC5C" box="[785,835,905,930]" captionStart="FIGURE 22" captionStartId="32.[151,250,1414,1436]" captionTargetBox="[151,1436,193,1392]" captionTargetId="figure-284@32.[151,1436,193,1392]" captionTargetPageId="32" captionText="FIGURE 22A–F. Photographs of dorsal (A,C,E) and ventral shield (B,D,F). A–B Neoatractides sundaensis n. sp., female holotype, Kemantis stream. C–D Neoatractides uniscutatus n. sp., male holotype, Kemantis stream. E–F Pseudotorrenticola borneoensis n. sp., male holotype, Little Lumotok stream." figureDoi="http://doi.org/10.5281/zenodo.4927959" httpUri="https://zenodo.org/record/4927959/files/figure.png" pageId="31" pageNumber="32">22D</figureCitation>
|
||
) L 556, W 414; dorsal shield (
|
||
<figureCitation id="E5A72A0D7E7BFFDEFB02FC3CD06BFC5C" box="[1176,1290,905,930]" captionStart="FIGURE 20" captionStartId="29.[151,250,1955,1977]" captionTargetBox="[260,1343,193,1930]" captionTargetId="figure-66@29.[221,1366,193,1933]" captionTargetPageId="29" captionText="FIGURE 20A–F. Neoatractides uniscutatus n. sp., male holotype, Kemantis stream: A = dorsal shield; B = idiosoma, ventral view; C = palp, medial view; D = palp, lateral view; E = gnathosoma and chelicera, F = I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927955" httpUri="https://zenodo.org/record/4927955/files/figure.png" pageId="31" pageNumber="32">Figs. 20A</figureCitation>
|
||
,
|
||
<figureCitation id="E5A72A0D7E7BFFDEFA8FFC3CD024FC5C" box="[1301,1349,905,930]" captionStart="FIGURE 22" captionStartId="32.[151,250,1414,1436]" captionTargetBox="[151,1436,193,1392]" captionTargetId="figure-284@32.[151,1436,193,1392]" captionTargetPageId="32" captionText="FIGURE 22A–F. Photographs of dorsal (A,C,E) and ventral shield (B,D,F). A–B Neoatractides sundaensis n. sp., female holotype, Kemantis stream. C–D Neoatractides uniscutatus n. sp., male holotype, Kemantis stream. E–F Pseudotorrenticola borneoensis n. sp., male holotype, Little Lumotok stream." figureDoi="http://doi.org/10.5281/zenodo.4927959" httpUri="https://zenodo.org/record/4927959/files/figure.png" pageId="31" pageNumber="32">22C</figureCitation>
|
||
) L 455, W 328, L/W ratio 1.39. Gnathosomal bay L 131, Cx-I total L 259, Cx-I mL 128, Cx-II+III mL 106; ratio Cx-I L/ Cx-II+III mL 2.4; Cx-I mL/Cx-II+III mL 1.2. Genital field L/W 93/70, ratio 1.32; distance genital field-excretory pore 78, genital field-caudal idiosoma margin 97. Gnathosoma vL 238–239; chelicera total L 263, basal segment L 220, claw L 50, L basal segment/claw ratio 4.4; palp total L 194–195, dL/H, dL/H ratio: P-1, 32/31–32, 1.03; P-2, 75/32, 2.36; P-3, 34/38, 0.9; P-4, 41–42/21, 2.0; P-5, 12/15, 0.8; P-2/P-4 ratio 1.82. Legs: dL of I-L-5–6: 85, 96, I- L-6 H 48, dL/H I-L-6 ratio 2.0.
|
||
</paragraph>
|
||
<paragraph id="7D2336887E7BFFDEFF5DFB30D4F8FB60" blockId="31.[151,1437,329,1686]" box="[199,409,1157,1182]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF5DFB30D47AFB60" box="[199,283,1157,1182]" italics="true" pageId="31" pageNumber="32">Female</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="358665037E7BFFDEFF5DFB1CD1F9FB3C" box="[199,1176,1193,1218]" pageId="31" pageNumber="32" type="etymology">
|
||
<paragraph id="7D2336887E7BFFDEFF5DFB1CD1F9FB3C" blockId="31.[151,1437,329,1686]" box="[199,1176,1193,1218]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF5DFB1CD42FFB3C" bold="true" box="[199,334,1193,1218]" pageId="31" pageNumber="32">Etymology.</emphasis>
|
||
The species name refers to the undivided dorsal shield of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="358665037E7BFFDEFF5DFB78D6CFF98C" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="7D2336887E7BFFDEFF5DFB78D782F9B0" blockId="31.[151,1437,329,1686]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF5DFB78D424FB18" bold="true" box="[199,325,1229,1254]" pageId="31" pageNumber="32">Discussion</emphasis>
|
||
.
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFEC3FB78D7E5FB1B" authority="Pešić & Smit, 2014" authorityName="Pešić & Smit" authorityYear="2014" box="[345,644,1229,1254]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="uniscutatus" status="sp. nov.">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFEC3FB78D7E5FB1B" box="[345,644,1229,1254]" italics="true" pageId="31" pageNumber="32">Neoatractides uniscutatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="54DB57E17E7BFFDEFD0BFB78D7B6FB1B" box="[657,727,1229,1254]" pageId="31" pageNumber="32" rank="species">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFD0BFB78D7AEFB18" bold="true" box="[657,719,1229,1254]" pageId="31" pageNumber="32">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is the third representative of the subgenus
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFB76FB78D42BFAF4" authority="Lundblad, 1941" authorityName="Lundblad" authorityYear="1941" class="Arachnida" family="Torrenticolidae" genus="Heteratractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFB76FB78D0FDFB18" box="[1260,1436,1229,1254]" italics="true" pageId="31" pageNumber="32">Heteratractides</emphasis>
|
||
<bibRefCitation id="190D4B797E7BFFDEFF0DFB44D42BFAF4" author="Lundblad, O." box="[151,330,1265,1290]" pageId="31" pageNumber="32" pagination="97 - 121" refId="ref37405" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomologisk Tidskrift, 62, 97 - 121." type="journal article" year="1941">Lundblad, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
. Synapomorphies of the species included in this subgenus (i.e. subgeneric autapomorphies) include the following features: lateral distal margin of P-3 deeply indented and gnathosomal rostrum broad and upturned (
|
||
<bibRefCitation id="190D4B797E7BFFDEFF04FA8CD44BFAAC" author="Wiles, P. R." box="[158,298,1337,1362]" pageId="31" pageNumber="32" pagination="191 - 236" refId="ref38559" refString="Wiles, P. R. (1997) Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and description of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia. Journal of Natural History, 31, 191 - 236. http: // dx. doi. org / 10.1080 / 00222939700770121" type="journal article" year="1997">Wiles 1997</bibRefCitation>
|
||
). Two species are known so far,
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFD5FFA8CD1BAFAAC" authority="(Lundblad, 1941)" baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[709,1243,1337,1362]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="serratirostris">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFD5FFA8CD162FAAF" box="[709,1027,1337,1362]" italics="true" pageId="31" pageNumber="32">Neoatractides serratirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="190D4B797E7BFFDEFB82FA8CD1B2FAAC" author="Lundblad, O." box="[1048,1235,1337,1362]" pageId="31" pageNumber="32" pagination="97 - 121" refId="ref37405" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomologisk Tidskrift, 62, 97 - 121." type="journal article" year="1941">Lundblad, 1941</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="058B76187E7BFFDEFAB7FA8CD0FAFAAC" box="[1325,1435,1337,1362]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="190D4B797E7BFFDEFF04FAE8D42FFA88" author="Lundblad, O." box="[158,334,1373,1398]" pageId="31" pageNumber="32" pagination="97 - 121" refId="ref37405" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomologisk Tidskrift, 62, 97 - 121." type="journal article" year="1941">Lundblad 1941</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFE08FAEBD7D5FA88" authority="Wiles, 1991" authorityName="Wiles" authorityYear="1991" box="[402,692,1373,1398]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFE08FAEBD740FA88" box="[402,545,1373,1398]" italics="true" pageId="31" pageNumber="32">N. orientalis</emphasis>
|
||
<bibRefCitation id="190D4B797E7BFFDEFDB3FAE8D7D5FA88" author="Wiles, P. R." box="[553,692,1373,1398]" pageId="31" pageNumber="32" pagination="41 - 56" refId="ref38490" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32 (1), 41 - 56." type="journal article" year="1991">Wiles, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, described on the basis of
|
||
<specimenCount id="6B9AFD017E7BFFDEFC71FAEBD10BFA88" box="[1003,1130,1373,1398]" pageId="31" pageNumber="32" type="female">two female</specimenCount>
|
||
specimens from
|
||
<collectingRegion id="BF58F86A7E7BFFDEFAA8FAE8D0F6FA88" box="[1330,1431,1373,1398]" country="Malaysia" name="Selangor" pageId="31" pageNumber="32">Selangor</collectingRegion>
|
||
, Peninsular
|
||
<collectingCountry id="058B76187E7BFFDEFE86FA34D4E2FA64" box="[284,387,1409,1434]" name="Malaysia" pageId="31" pageNumber="32">Malaysia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="190D4B797E7BFFDEFE03FA34D745FA64" author="Wiles, P. R." box="[409,548,1409,1434]" pageId="31" pageNumber="32" pagination="41 - 56" refId="ref38490" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32 (1), 41 - 56." type="journal article" year="1991">Wiles 1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="190D4B797E7BFFDEFDA2FA34D715FA64" author="Wiles, P. R." box="[568,628,1409,1434]" pageId="31" pageNumber="32" pagination="191 - 236" refId="ref38559" refString="Wiles, P. R. (1997) Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and description of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia. Journal of Natural History, 31, 191 - 236. http: // dx. doi. org / 10.1080 / 00222939700770121" type="journal article" year="1997">1997</bibRefCitation>
|
||
). From the latter species,
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFC21FA34D187FA67" authority="Pešić & Smit, 2014" authorityName="Pešić & Smit" authorityYear="2014" box="[955,1254,1409,1434]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="uniscutatus" status="sp. nov.">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFC21FA34D187FA67" box="[955,1254,1409,1434]" italics="true" pageId="31" pageNumber="32">Neoatractides uniscutatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4FE8EA9A7E7BFFDEFB6CFA34D05DFA64" bold="true" box="[1270,1340,1409,1434]" pageId="31" pageNumber="32">
|
||
<taxonomicNameLabel id="54DB57E17E7BFFDEFB6CFA34D05DFA64" box="[1270,1340,1409,1434]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the frontal platelets fused to the large dorsal plate.
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFC15FA10D1A7FA43" baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[911,1222,1445,1470]" class="Arachnida" family="Torrenticolidae" genus="Neoatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="serratirostris">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFC15FA10D1A7FA43" box="[911,1222,1445,1470]" italics="true" pageId="31" pageNumber="32">Neoatractides serratirostris</emphasis>
|
||
</taxonomicName>
|
||
resembles the new species due to the anterolateral platelets fused with the large dorsal plate forming an undivided dorsal shield, but differs from the new species in having a shorter and broader gnathosomal rostrum, a very long ventral seta on P-2 exceeding the tip of P-5, a conventionally shaped ejaculatory complex, relatively larger genital field and a shorter medial suture line of Cx-II+III (see
|
||
<bibRefCitation id="190D4B797E7BFFDEFDB2F980D7B6F9B0" author="Lundblad, O." box="[552,727,1589,1614]" pageId="31" pageNumber="32" pagination="435 - 585" refId="ref37427" refString="Lundblad, O. (1953) Die Hydracarinenfauna von Colombia. Arkiv for zoology, Serie 2, 5 (8), 435 - 585." type="journal article" year="1953">Lundblad 1953</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7D2336887E7BFFDEFF5DF9ECD6CFF98C" blockId="31.[151,1437,329,1686]" box="[199,942,1625,1650]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF5DF9ECD44BF98C" bold="true" box="[199,298,1625,1650]" pageId="31" pageNumber="32">Habitat.</emphasis>
|
||
Sandy/bouldery stream, shaded by rain forest (
|
||
<figureCitation id="E5A72A0D7E7BFFDEFCA1F9ECD6C0F98C" box="[827,929,1625,1650]" captionStart="FIGURE 43" captionStartId="68.[151,250,1550,1572]" captionTargetBox="[298,1289,193,1529]" captionTargetId="figure-283@68.[298,1289,193,1529]" captionTargetPageId="68" captionText="FIGURE 43A–D. Photographs of selected sampling sites: A—Mahua stream (type locality of Torrenticola kinabaluensis n. sp. and sampling site of Torrenticola borneoensis n. sp., T. sabahensis n. sp., T. minuta, T. neoindica n. sp., T. cf. pugionirostris, Monatractides longiventris, M. parviventris and M. morpheus n. sp.), B—Unnamed stream Bansadon Trail (type locality of Neoatractides uniscutatus n. sp. and sampling site of Torrenticola borneoensis n. sp., T. kinabaluensis n. sp., Monatractides cf. macroporus and M. parviventris). C—Kibamabangan River (type locality of Torrenticola schilthuizeni n. sp. and sampling site of Torrenticola borneoensis n. sp., T. kinabaluensis n. sp., T. minuta and T. neoindica n. sp., Monatractides cf. macroporus, M. oxystomus, M. parviventris and M. morpheus n. sp.). D—Kemantis stream (type locality of Torrenticola borneoensis n. sp., Neoatractides sundaensis n. sp. and Monatractides phantasos n. sp. and sampling site of T. minuta, Monatractides longiventris, M. parviventris and M. morpheus n. sp.). Photos. H. Smit." figureDoi="http://doi.org/10.5281/zenodo.4928004" httpUri="https://zenodo.org/record/4928004/files/figure.png" pageId="31" pageNumber="32">Fig. 43B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="358665037E7BFFDEFF5DF9C8D716F8FD" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="7D2336887E7BFFDEFF5DF9C8D632F96B" blockId="31.[151,1437,329,1686]" box="[199,851,1661,1686]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF5DF9C8D401F968" bold="true" box="[199,352,1661,1686]" pageId="31" pageNumber="32">Distribution.</emphasis>
|
||
Borneo; known only from the
|
||
<emphasis id="4FE8EA9A7E7BFFDEFD21F9C8D62DF96B" box="[699,844,1661,1686]" italics="true" pageId="31" pageNumber="32">locus typicus</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7D2336887E7BFFDEFF0DF95CD716F8FD" blockId="31.[151,631,1769,1795]" box="[151,631,1769,1795]" pageId="31" pageNumber="32">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF0DF95CD716F8FD" bold="true" box="[151,631,1769,1795]" pageId="31" pageNumber="32">
|
||
Genus
|
||
<taxonomicName id="BA9C4D0B7E7BFFDEFF74F95CD716F8FD" authority="Walter, 1906" authorityName="Walter" authorityYear="1906" box="[238,631,1769,1795]" class="Arachnida" family="Torrenticolidae" genus="Pseudotorrenticola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FE8EA9A7E7BFFDEFF74F95CD4B5F8FD" bold="true" box="[238,468,1769,1795]" italics="true" pageId="31" pageNumber="32">Pseudotorrenticola</emphasis>
|
||
Walter, 1906
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |