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<document id="7D13C57E51E2A3C6DF9D169732D40875" ID-CLB-Dataset="5420" ID-DOI="10.11646/zootaxa.3840.1.1" ID-GBIF-Dataset="1478a1d3-fcb4-43b3-9418-010043c5c154" ID-ISSN="1175-5326" ID-Zenodo-Dep="4927892" ID-ZooBank="CDF827EB-8A66-438F-AC5C-07A7D3D83BB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623431855578" checkinUser="felipe" docAuthor="Pešić, Vladimir &amp; Smit, Harry" docDate="2014" docId="F535879E7E74FFD1FF0DFDD9D6BCF84C" docLanguage="en" docName="zt03840p072.pdf" docOrigin="Zootaxa 3840 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Torrenticola minuta" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="090CFFE67E64FFC1FF9AFFB5D561FFFE" masterDocTitle="Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo" masterLastPageNumber="72" masterPageNumber="1" pageNumber="17" updateTime="1699010593832" updateUser="ExternalLinkService">
<mods:mods id="19D729BC1801ED6529AD4CAC880C6A85" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0AFE0B9064DE04EB26736235179C0B1">
<mods:title id="67612F4A4D0D065AD34446212DBC35E3">Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo</mods:title>
</mods:titleInfo>
<mods:name id="1880AB12126EA3E0CADBA7F900948AD9" type="personal">
<mods:role id="DAF3823FACBC8E424ACE2E7E4CDEAF2C">
<mods:roleTerm id="5DEE150F3A178CF319995B5A884D6C3A">Author</mods:roleTerm>
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<mods:namePart id="430EF8212D75C8E594A7C3916463D0EF">Pešić, Vladimir</mods:namePart>
<mods:nameIdentifier id="0FB994DCCA1BB6554265D88B3D593991" type="email">vladopesic@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="25FB4368CFD2955F2F2FE7447CF04317" type="personal">
<mods:role id="046DE7F7CB821DD8834BF198C46CB5A5">
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</mods:role>
<mods:namePart id="9981E589A487463BEFB769FE4CB8DB6F">Smit, Harry</mods:namePart>
<mods:nameIdentifier id="D8A0B55E61EDB7BD4369068D89D256A5" type="email">vladopesic@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="6771CF9445E30765E74C198D592A20C0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BF5AFC9C3E6AB202F8DCFD4E49323B56">
<mods:date id="869044F499A7E4F8B41EB23086686083">2014</mods:date>
<mods:detail id="449C86934A8DAA4A1715B460506455B6" type="pubDate">
<mods:number id="7364E08C1A4755FA920CCD00BDEE9890">2014-07-23</mods:number>
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<mods:number id="88C8C267957232654AF5AEC96847E097">3840</mods:number>
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<mods:number id="EC613E96C80B484E5368A80C169CF030">1</mods:number>
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<mods:classification id="40C8819666179AEE31D6E0259076505C">journal article</mods:classification>
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<mods:identifier id="020A2086F3F0B3C4B4C6014B38152B3D" type="DOI">10.11646/zootaxa.3840.1.1</mods:identifier>
<mods:identifier id="B52E6DB686409AD4A3BC1B24CFEBBD1A" type="GBIF-Dataset">1478a1d3-fcb4-43b3-9418-010043c5c154</mods:identifier>
<mods:identifier id="1A02C5C3F1E117BBE5ACC5EA8FBB5526" type="ISSN">1175-5326</mods:identifier>
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<treatment id="F535879E7E74FFD1FF0DFDD9D6BCF84C" ID-DOI="http://doi.org/10.5281/zenodo.4927707" ID-GBIF-Taxon="183550664" ID-Zenodo-Dep="4927707" LSID="urn:lsid:plazi:treatment:F535879E7E74FFD1FF0DFDD9D6BCF84C" httpUri="http://treatment.plazi.org/id/F535879E7E74FFD1FF0DFDD9D6BCF84C" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="358665037E74FFD1FF0DFDD9D709FD78" box="[151,616,620,646]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="7D2336887E74FFD1FF0DFDD9D709FD78" blockId="16.[151,616,620,679]" box="[151,616,620,646]" pageId="16" pageNumber="17">
<heading id="266B81E47E74FFD1FF0DFDD9D709FD78" bold="true" box="[151,616,620,646]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<taxonomicName id="BA9C4D0B7E74FFD1FF0DFDD9D709FD78" authority="(Lundblad, 1941)" baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[151,616,620,646]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="4FE8EA9A7E74FFD1FF0DFDD9D709FD78" bold="true" box="[151,616,620,646]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF0DFDD9D4E8FD78" bold="true" box="[151,393,620,646]" italics="true" pageId="16" pageNumber="17">Torrenticola minuta</emphasis>
(
<bibRefCitation id="190D4B797E74FFD1FE02FDD9D701FD78" author="Lundblad, O." box="[408,608,620,646]" pageId="16" pageNumber="17" pagination="97 - 121" refId="ref37405" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomologisk Tidskrift, 62, 97 - 121." type="journal article" year="1941">Lundblad, 1941</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="358665037E74FFD1FF0DFD3BD704FD59" box="[151,613,654,679]" pageId="16" pageNumber="17" type="description">
<paragraph id="7D2336887E74FFD1FF0DFD3BD704FD59" blockId="16.[151,616,620,679]" box="[151,613,654,679]" pageId="16" pageNumber="17">
(
<figureCitation id="E5A72A0D7E74FFD1FF05FD3BD453FD59" box="[159,306,654,679]" captionStart="FIGURE 11" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[176,1416,205,1917]" captionTargetId="figure-53@17.[157,1429,195,1933]" captionTargetPageId="17" captionText="FIGURE 11A–D. Torrenticola minuta (Lundblad, 1941) (A,C—male from Kemantis stream; B,D—female from Great Lumotok stream): A–B = idiosoma, ventral view; C–D = palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927927" httpUri="https://zenodo.org/record/4927927/files/figure.png" pageId="16" pageNumber="17">Figs. 11A–D</figureCitation>
,
<figureCitation id="E5A72A0D7E74FFD1FEA4FD3BD4F0FD59" box="[318,401,654,679]" captionStart="FIGURE 16" captionStartId="23.[151,250,1299,1321]" captionTargetBox="[151,1436,193,1277]" captionTargetId="figure-343@23.[151,1436,193,1277]" captionTargetPageId="23" captionText="FIGURE 16A–F. Photographs of dorsal shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F. T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927947" httpUri="https://zenodo.org/record/4927947/files/figure.png" pageId="16" pageNumber="17">16A–B</figureCitation>
,
<figureCitation id="E5A72A0D7E74FFD1FE07FD3BD491FD59" box="[413,496,654,679]" captionStart="FIGURE 17" captionStartId="25.[151,250,1373,1395]" captionTargetBox="[151,1436,195,1352]" captionTargetId="figure-185@25.[151,1436,195,1352]" captionTargetPageId="25" captionText="FIGURE 17A–F. Photographs of ventral shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927949" httpUri="https://zenodo.org/record/4927949/files/figure.png" pageId="16" pageNumber="17">17A–B</figureCitation>
,
<figureCitation id="E5A72A0D7E74FFD1FE66FD3BD73CFD59" box="[508,605,654,679]" captionStart="FIGURE 23" captionStartId="33.[151,250,1268,1290]" captionTargetBox="[244,1342,397,1247]" captionTargetId="figure-364@33.[244,1342,397,1247]" captionTargetPageId="33" captionText="FIGURE 23A–H. Photographs of ejaculatory complex. A Torrenticola borneoensis n. sp. B T. kinabaluensis n. sp.. C T. longipalpis Wiles, 1997. D T. sabahensis n. sp. E Torrenticola minuta (Lundblad, 1941). F T. neoindica n. sp. G Neoatractides uniscutatus n. sp. H Pseudotorrenticola borneoensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4927961" httpUri="https://zenodo.org/record/4927961/files/figure.png" pageId="16" pageNumber="17">Fig. 23E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="358665037E74FFD1FF0DFD63D63EFC3C" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="7D2336887E74FFD1FF0DFD63D742FD12" blockId="16.[151,547,726,748]" box="[151,547,726,748]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="5D8C11A67E74FFD1FF0DFD63D742FD12" box="[151,547,726,748]" pageId="16" pageNumber="17">
<treatmentCitation id="FC3D10997E74FFD1FF0DFD63D77EFD12" author="Lundblad, O." box="[151,543,726,748]" page="99" pageId="16" pageNumber="17" year="1941">
<taxonomicName id="BA9C4D0B7E74FFD1FF0DFD63D77EFD12" authority="Lundblad 1941: 99" authorityName="Lundblad" authorityPageNumber="99" authorityYear="1941" box="[151,543,726,748]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="4FE8EA9A7E74FFD1FF0DFD63D434FD12" box="[151,341,726,748]" italics="true" pageId="16" pageNumber="17">Atractides minutus</emphasis>
<bibRefCitation id="190D4B797E74FFD1FEC1FD63D77EFD12" author="Lundblad, O." box="[347,543,726,748]" pageId="16" pageNumber="17" pagination="97 - 121" refId="ref37405" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomologisk Tidskrift, 62, 97 - 121." type="journal article" year="1941">Lundblad 1941: 99</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7D2336887E74FFD1FF0DFCACD63EFC3C" blockId="16.[151,1437,793,1970]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF0DFCACD44CFCCC" bold="true" box="[151,301,793,818]" pageId="16" pageNumber="17">New records</emphasis>
.
<materialsCitation id="CDF43CD57E74FFD1FEA0FCACD154FCA8" ID-GBIF-Occurrence="3323413333" collectingDate="2012-09-16" collectingDateMax="2012-09-21" collectingDateMin="2012-09-16" country="Malaysia" county="Mt Kinabalu" elevation="928" latitude="6.1556" location="Borneo" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" pageId="16" pageNumber="17" specimenCount="1" specimenCount-juvenile="1">
<collectingCountry id="058B76187E74FFD1FEA0FCACD4C2FCCC" box="[314,419,793,818]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
,
<location id="784360537E74FFD1FE34FCAFD762FCCF" LSID="urn:lsid:plazi:treatment:F535879E7E74FFD1FF0DFDD9D6BCF84C:784360537E74FFD1FE34FCAFD762FCCF" box="[430,515,794,817]" country="Malaysia" county="Mt Kinabalu" latitude="6.1556" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" name="Borneo" pageId="16" pageNumber="17">Borneo</location>
, stream
<location id="784360537E74FFD1FDFAFCACD7AFFCCC" LSID="urn:lsid:plazi:treatment:F535879E7E74FFD1FF0DFDD9D6BCF84C:784360537E74FFD1FDFAFCACD7AFFCCC" box="[608,718,793,818]" country="Malaysia" county="Mt Kinabalu" latitude="6.1556" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" name="Kemantis" pageId="16" pageNumber="17">Kemantis</location>
,
<location id="784360537E74FFD1FD43FCACD641FCCC" LSID="urn:lsid:plazi:treatment:F535879E7E74FFD1FF0DFDD9D6BCF84C:784360537E74FFD1FD43FCACD641FCCC" box="[729,800,793,818]" country="Malaysia" county="Mt Kinabalu" latitude="6.1556" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" name="Sayap" pageId="16" pageNumber="17">Sayap</location>
,
<location id="784360537E74FFD1FCB1FCAFD6DCFCCC" LSID="urn:lsid:plazi:treatment:F535879E7E74FFD1FF0DFDD9D6BCF84C:784360537E74FFD1FCB1FCAFD6DCFCCC" box="[811,957,793,818]" country="Malaysia" county="Mt Kinabalu" latitude="6.1556" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" name="Mt Kinabalu" pageId="16" pageNumber="17">Mt Kinabalu</location>
,
<geoCoordinate id="18A8504F7E74FFD1FC5DFCACD129FCCF" box="[967,1096,793,818]" degrees="6" direction="north" minutes="09.841" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="6.1640167">6º09.841 N</geoCoordinate>
,
<geoCoordinate id="18A8504F7E74FFD1FBCEFCACD18DFCCF" box="[1108,1260,793,818]" degrees="116" direction="east" minutes="33.936" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="116.5656">116º33.936 E</geoCoordinate>
, alt.
<elevation id="F6B1D1BB7E74FFD1FAB8FCACD0F2FCCC" box="[1314,1427,793,818]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="16" pageNumber="17" unit="m" value="928.0">
<quantity id="BA649B6D7E74FFD1FAB8FCACD008FCCF" box="[1314,1385,793,818]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="16" pageNumber="17" unit="m" value="928.0">928 m</quantity>
asl
</elevation>
.,
<date id="092210487E74FFD1FF0DFC88D46FFCA8" box="[151,270,829,854]" pageId="16" pageNumber="17" value="2012-09-16">
<collectingDate id="1966E9A07E74FFD1FF0DFC88D46FFCA8" box="[151,270,829,854]" pageId="16" pageNumber="17" value="2012-09-16">16.ix.2012</collectingDate>
</date>
Smit 2[
<specimenCount id="6B9AFD017E74FFD1FEE2FC88D775FCA8" box="[376,532,829,854]" pageId="16" pageNumber="17" type="juvenile">one juvenile</specimenCount>
]1/0 (1/0/0 mounted); Mahua stream, Mahua,
</materialsCitation>
<materialsCitation id="CDF43CD57E74FFD1FBDFFC88D1BCFC84" ID-GBIF-Occurrence="3323413358" collectingDate="2012-09-21" collectingDateMax="2012-09-21" collectingDateMin="2012-09-16" country="Malaysia" county="Mt Kinabalu" elevation="1052" latitude="6.1556" location="Kibamabangan River" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" pageId="16" pageNumber="17" specimenCount="1" specimenCount-juvenile="1">
Crocker Range,
<geoCoordinate id="18A8504F7E74FFD1FA97FC88D0F9FCAB" box="[1293,1432,829,854]" degrees="5" direction="north" minutes="47.838" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="5.7973">5º47.838 N</geoCoordinate>
,
<geoCoordinate id="18A8504F7E74FFD1FF0DFCD4D454FC87" box="[151,309,865,890]" degrees="116" direction="east" minutes="24.510" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="116.4085">116º24.510 E</geoCoordinate>
, alt.
<elevation id="F6B1D1BB7E74FFD1FEE8FCD4D49AFC84" box="[370,507,865,890]" metricMagnitude="3" metricUnit="m" metricValue="1.052" pageId="16" pageNumber="17" unit="m" value="1052.0">
<quantity id="BA649B6D7E74FFD1FEE8FCD4D4ADFC87" box="[370,460,865,890]" metricMagnitude="3" metricUnit="m" metricValue="1.052" pageId="16" pageNumber="17" unit="m" value="1052.0">1052 m</quantity>
asl
</elevation>
.,
<date id="092210487E74FFD1FD95FCD4D7E6FC84" box="[527,647,865,890]" pageId="16" pageNumber="17" value="2012-09-21">
<collectingDate id="1966E9A07E74FFD1FD95FCD4D7E6FC84" box="[527,647,865,890]" pageId="16" pageNumber="17" value="2012-09-21">21.ix.2012</collectingDate>
</date>
Smit 2/0/0 (1/0/0 mounted);
<collectingMunicipality id="9D47ACF27E74FFD1FC7EFCD4D1B8FC84" box="[996,1241,865,890]" pageId="16" pageNumber="17">Kibamabangan River</collectingMunicipality>
,
</materialsCitation>
<materialsCitation id="CDF43CD57E74FFD1FB72FCD4D113FC63" ID-GBIF-Occurrence="3323413369" collectingDate="2012-09-18" collectingDateMax="2012-09-21" collectingDateMin="2012-09-16" country="Malaysia" county="Mt Kinabalu" elevation="433" latitude="6.1556" location="Borneo" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" pageId="16" pageNumber="17" specimenCount="1" specimenCount-juvenile="1">
Crocker Range,
<geoCoordinate id="18A8504F7E74FFD1FF0DFC30D46AFC63" box="[151,267,901,926]" degrees="5" direction="north" minutes="51.28" orientation="latitude" pageId="16" pageNumber="17" precision="9" value="5.8546667">5º51.28 N</geoCoordinate>
,
<geoCoordinate id="18A8504F7E74FFD1FE8CFC30D4CEFC63" box="[278,431,901,926]" degrees="116" direction="east" minutes="08.417" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="116.14028">116º08.417 E</geoCoordinate>
, alt.
<elevation id="F6B1D1BB7E74FFD1FE7CFC30D739FC60" box="[486,600,901,926]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="16" pageNumber="17" unit="m" value="433.0">
<quantity id="BA649B6D7E74FFD1FE7CFC30D74CFC63" box="[486,557,901,926]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="16" pageNumber="17" unit="m" value="433.0">433 m</quantity>
asl
</elevation>
.,
<date id="092210487E74FFD1FDF3FC30D781FC60" box="[617,736,901,926]" pageId="16" pageNumber="17" value="2012-09-18">
<collectingDate id="1966E9A07E74FFD1FDF3FC30D781FC60" box="[617,736,901,926]" pageId="16" pageNumber="17" value="2012-09-18">18.ix.2012</collectingDate>
</date>
Smit 2/1/0; Great Lumotok stream,
</materialsCitation>
<materialsCitation id="CDF43CD57E74FFD1FBE3FC33D63AFC3C" ID-GBIF-Occurrence="3323413375" collectingDate="2012-09-18" collectingDateMax="2012-09-21" collectingDateMin="2012-09-16" country="Malaysia" county="Mt Kinabalu" elevation="433" latitude="6.1556" location="Mt Kinabalu" longLatPrecision="1" longitude="116.14028" municipality="Kibamabangan River" pageId="16" pageNumber="17" specimenCount="1" specimenCount-juvenile="1">
<collectingCounty id="94424E047E74FFD1FBE3FC33D06AFC60" box="[1145,1291,901,926]" pageId="16" pageNumber="17">Mt Kinabalu</collectingCounty>
,
<geoCoordinate id="18A8504F7E74FFD1FA8CFC30D0F9FC63" box="[1302,1432,901,926]" degrees="6" direction="north" minutes="09.336" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="6.1556">6º09.336 N</geoCoordinate>
,
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, alt.
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asl
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.,
<date id="092210487E74FFD1FE7DFC1CD73EFC3C" box="[487,607,937,962]" pageId="16" pageNumber="17" value="2012-09-18">
<collectingDate id="1966E9A07E74FFD1FE7DFC1CD73EFC3C" box="[487,607,937,962]" pageId="16" pageNumber="17" value="2012-09-18">18.ix.2012</collectingDate>
</date>
Smit 0/1/0 (mounted)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="358665037E74FFD1FF0DFC78D06CF8DC" pageId="16" pageNumber="17" type="description">
<paragraph id="7D2336887E74FFD1FF0DFC78D44AFC18" blockId="16.[151,1437,793,1970]" box="[151,299,973,998]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF0DFC78D44AFC18" bold="true" box="[151,299,973,998]" pageId="16" pageNumber="17">Morphology</emphasis>
</paragraph>
<paragraph id="7D2336887E74FFD1FF5DFC44D631FAD4" blockId="16.[151,1437,793,1970]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF5DFC44D4E3FBF4" box="[199,386,1009,1034]" italics="true" pageId="16" pageNumber="17">General features</emphasis>
—Idiosoma roundish (dorsal shield L/W ratio 1.2); shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in
<figureCitation id="E5A72A0D7E74FFD1FCDFFBA0D6BEFBD0" box="[837,991,1045,1070]" captionStart="FIGURE 16" captionStartId="23.[151,250,1299,1321]" captionTargetBox="[151,1436,193,1277]" captionTargetId="figure-343@23.[151,1436,193,1277]" captionTargetPageId="23" captionText="FIGURE 16A–F. Photographs of dorsal shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F. T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927947" httpUri="https://zenodo.org/record/4927947/files/figure.png" pageId="16" pageNumber="17">Figs. 16A–B</figureCitation>
; gnathosomal bay U-shaped; Cxgl–4 subapical, only slightly posterior of Cx-I tips; suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field; excretory and Vgl–2 pore away from the line of primary sclerotization, excretory pore on the level with Vgl–2; gnathosoma with curved ventral margin; P-2 longer than P-4, P-2 ventral margin slightly convex, P-2 ventrodistal protrusion, bluntly pointed, curved towards distal; P-3 ventrodistal protrusion bluntly pointed; P-4 with well developed ventral protuberances, ending in two tips separated by a concavity (
<figureCitation id="E5A72A0D7E74FFD1FB67FB7CD0F1FB1C" box="[1277,1424,1225,1250]" captionStart="FIGURE 11" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[176,1416,205,1917]" captionTargetId="figure-53@17.[157,1429,195,1933]" captionTargetPageId="17" captionText="FIGURE 11A–D. Torrenticola minuta (Lundblad, 1941) (A,C—male from Kemantis stream; B,D—female from Great Lumotok stream): A–B = idiosoma, ventral view; C–D = palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927927" httpUri="https://zenodo.org/record/4927927/files/figure.png" pageId="16" pageNumber="17">Figs. 11C–D</figureCitation>
). Male: medial suture line of Cx-II+III short; genital field subrectangular in shape; ejaculatory complex conventional in shape (
<figureCitation id="E5A72A0D7E74FFD1FE99FAA4D409FAD4" box="[259,360,1297,1322]" captionStart="FIGURE 23" captionStartId="33.[151,250,1268,1290]" captionTargetBox="[244,1342,397,1247]" captionTargetId="figure-364@33.[244,1342,397,1247]" captionTargetPageId="33" captionText="FIGURE 23A–H. Photographs of ejaculatory complex. A Torrenticola borneoensis n. sp. B T. kinabaluensis n. sp.. C T. longipalpis Wiles, 1997. D T. sabahensis n. sp. E Torrenticola minuta (Lundblad, 1941). F T. neoindica n. sp. G Neoatractides uniscutatus n. sp. H Pseudotorrenticola borneoensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4927961" httpUri="https://zenodo.org/record/4927961/files/figure.png" pageId="16" pageNumber="17">Fig. 23E</figureCitation>
). Female: genital field pentagonal in shape.
</paragraph>
<paragraph id="7D2336887E74FFD1FF0DFA83D458FAB3" blockId="16.[151,1437,793,1970]" box="[151,313,1334,1357]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF0DFA83D458FAB3" box="[151,313,1334,1357]" italics="true" pageId="16" pageNumber="17">Measurements</emphasis>
</paragraph>
<paragraph id="7D2336887E74FFD1FF5DFAECD4C7F9B4" blockId="16.[151,1437,793,1970]" pageId="16" pageNumber="17">
Male (from stream Kemantis)—Idiosoma (ventral view:
<figureCitation id="E5A72A0D7E74FFD1FCC9FAECD6A4FA8C" box="[851,965,1369,1394]" captionStart="FIGURE 11" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[176,1416,205,1917]" captionTargetId="figure-53@17.[157,1429,195,1933]" captionTargetPageId="17" captionText="FIGURE 11A–D. Torrenticola minuta (Lundblad, 1941) (A,C—male from Kemantis stream; B,D—female from Great Lumotok stream): A–B = idiosoma, ventral view; C–D = palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927927" httpUri="https://zenodo.org/record/4927927/files/figure.png" pageId="16" pageNumber="17">Figs. 11A</figureCitation>
,
<figureCitation id="E5A72A0D7E74FFD1FC4EFAECD164FA8C" box="[980,1029,1369,1394]" captionStart="FIGURE 17" captionStartId="25.[151,250,1373,1395]" captionTargetBox="[151,1436,195,1352]" captionTargetId="figure-185@25.[151,1436,195,1352]" captionTargetPageId="25" captionText="FIGURE 17A–F. Photographs of ventral shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927949" httpUri="https://zenodo.org/record/4927949/files/figure.png" pageId="16" pageNumber="17">17A</figureCitation>
) L 628, W 459; dorsal shield (
<figureCitation id="E5A72A0D7E74FFD1FAF5FAECD5A9FA68" captionStart="FIGURE 16" captionStartId="23.[151,250,1299,1321]" captionTargetBox="[151,1436,193,1277]" captionTargetId="figure-343@23.[151,1436,193,1277]" captionTargetPageId="23" captionText="FIGURE 16A–F. Photographs of dorsal shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F. T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927947" httpUri="https://zenodo.org/record/4927947/files/figure.png" pageId="16" pageNumber="17">Fig. 16A</figureCitation>
) L 511, W 413, L/W ratio 1.24; dorsal plate L 481; frontal plate L 127–128, W 44, L/W ratio 2.9. Gnathosomal bay L 125, Cx-I total L 266, Cx-I mL 142, Cx-II+III mL 58; ratio Cx-I L/Cx-II+III mL 4.6; Cx-I mL/ Cx-II+III mL 2.5. Genital field L/W 119/100, ratio 1.19; ejaculatory complex L; distance genital field-excretory pore 114, genital field-caudal idiosoma margin 183. Gnathosoma vL 293; chelicera total L 339; palp total L 289–291, dL/H, dL/H ratio: P-1, 31/31, 1.0; P-2, 101-102/55, 1.86; P-3, 52/46, 1.13; P-4, 88-89/28, 3.2; P-5, 17/9, 1.85; P-2/P-4 ratio 1.15.
</paragraph>
<paragraph id="7D2336887E74FFD1FF5DF9E0D06CF8DC" blockId="16.[151,1437,793,1970]" pageId="16" pageNumber="17">
Female (from Great Lumotok stream)—Idiosoma (ventral view:
<figureCitation id="E5A72A0D7E74FFD1FC39F9E0D173F990" box="[931,1042,1621,1646]" captionStart="FIGURE 11" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[176,1416,205,1917]" captionTargetId="figure-53@17.[157,1429,195,1933]" captionTargetPageId="17" captionText="FIGURE 11A–D. Torrenticola minuta (Lundblad, 1941) (A,C—male from Kemantis stream; B,D—female from Great Lumotok stream): A–B = idiosoma, ventral view; C–D = palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4927927" httpUri="https://zenodo.org/record/4927927/files/figure.png" pageId="16" pageNumber="17">Figs. 11B</figureCitation>
,
<figureCitation id="E5A72A0D7E74FFD1FB85F9E0D12FF990" box="[1055,1102,1621,1646]" captionStart="FIGURE 17" captionStartId="25.[151,250,1373,1395]" captionTargetBox="[151,1436,195,1352]" captionTargetId="figure-185@25.[151,1436,195,1352]" captionTargetPageId="25" captionText="FIGURE 17A–F. Photographs of ventral shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927949" httpUri="https://zenodo.org/record/4927949/files/figure.png" pageId="16" pageNumber="17">17B</figureCitation>
) L 625, W 459; dorsal shield (
<figureCitation id="E5A72A0D7E74FFD1FF05F9CCD462F96C" box="[159,259,1657,1682]" captionStart="FIGURE 16" captionStartId="23.[151,250,1299,1321]" captionTargetBox="[151,1436,193,1277]" captionTargetId="figure-343@23.[151,1436,193,1277]" captionTargetPageId="23" captionText="FIGURE 16A–F. Photographs of dorsal shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F. T. schilthuizeni n. sp., female holotype, Kibamabangan River." figureDoi="http://doi.org/10.5281/zenodo.4927947" httpUri="https://zenodo.org/record/4927947/files/figure.png" pageId="16" pageNumber="17">Fig. 16B</figureCitation>
) L 497, W 412, L/W ratio 1.2; dorsal plate L 469; frontal plate L 128, W 41, L/W ratio 3.2. Gnathosomal bay L 130, Cx-I total L 266, Cx-I mL 136, Cx-II+III mL 37.5; ratio Cx-I L/Cx-II+III mL 7.1; Cx-I mL/Cx-II+III mL 3.6. Genital field L/W 129/114, ratio 1.13; distance genital field-excretory pore 113, genital field-caudal idiosoma margin 191. Gnathosoma vL 314; chelicera total L 358; palp total L 296–297, dL/H, dL/H ratio: P-1, 34/ 29, 1.16; P-2, 101-102/57, 1.78; P-3, 54/48, 1.13; P-4, 90/28, 3.25; P-5, 17/10, 1.7; P-2/P-4 ratio 1.13.
</paragraph>
</subSubSection>
<subSubSection id="358665037E74FFD1FF5DF898D061F870" pageId="16" pageNumber="17" type="discussion">
<paragraph id="7D2336887E74FFD1FF5DF898D061F870" blockId="16.[151,1437,793,1970]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF5DF898D455F8B8" bold="true" box="[199,308,1837,1862]" pageId="16" pageNumber="17">Remarks</emphasis>
.
<bibRefCitation id="190D4B797E74FFD1FED8F898D4B2F8B8" author="Wiles, P. R." box="[322,467,1837,1862]" pageId="16" pageNumber="17" pagination="409 - 439" refId="ref38627" refString="Wiles, P. R. (1999) The water mites (Acari: Hydrachnidia) of Borneo and additional new species from Thailand and Sulawesi. The Raffles Bulletin of Zoology, 47, 409 - 439." type="journal article" year="1999">Wiles (1999)</bibRefCitation>
reported
<taxonomicName id="BA9C4D0B7E74FFD1FDA5F89BD7CBF8BB" baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[575,682,1838,1861]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="4FE8EA9A7E74FFD1FDA5F89BD7CBF8BB" box="[575,682,1838,1861]" italics="true" pageId="16" pageNumber="17">T. minuta</emphasis>
</taxonomicName>
from several sites in the catchment area of the River
<collectingRegion id="BF58F86A7E74FFD1FB65F898D0E1F8B8" box="[1279,1408,1837,1862]" country="Brunei" name="Temburong" pageId="16" pageNumber="17">Temburong</collectingRegion>
in North
<collectingCountry id="058B76187E74FFD1FF45F8E4D44EF894" box="[223,303,1873,1898]" name="Brunei" pageId="16" pageNumber="17">Brunei</collectingCountry>
. He mentioned that his specimens differ from those from Peninsular
<collectingCountry id="058B76187E74FFD1FBABF8E4D1F9F894" box="[1073,1176,1873,1898]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
in having Cxgl-4 more posteriorly located, near I-L insertion, instead of being subapical (located just posterior of Cx-I tips).
</paragraph>
</subSubSection>
<subSubSection id="358665037E74FFD1FF5DF82CD6BCF84C" box="[199,989,1945,1970]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="7D2336887E74FFD1FF5DF82CD6BCF84C" blockId="16.[151,1437,793,1970]" box="[199,989,1945,1970]" pageId="16" pageNumber="17">
<emphasis id="4FE8EA9A7E74FFD1FF5DF82CD438F84C" bold="true" box="[199,345,1945,1970]" pageId="16" pageNumber="17">Distribution</emphasis>
.
<collectingRegion id="BF58F86A7E74FFD1FEFDF82CD4ADF84C" box="[359,460,1945,1970]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="17">Sulawesi</collectingRegion>
(
<bibRefCitation id="190D4B797E74FFD1FE41F82CD701F84C" author="Wiles, P. R." box="[475,608,1945,1970]" pageId="16" pageNumber="17" pagination="191 - 236" refId="ref38559" refString="Wiles, P. R. (1997) Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and description of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia. Journal of Natural History, 31, 191 - 236. http: // dx. doi. org / 10.1080 / 00222939700770121" type="journal article" year="1997">Wiles 1997</bibRefCitation>
), Borneo (
<bibRefCitation id="190D4B797E74FFD1FD4EF82CD639F84C" author="Wiles, P. R." box="[724,856,1945,1970]" pageId="16" pageNumber="17" pagination="409 - 439" refId="ref38627" refString="Wiles, P. R. (1999) The water mites (Acari: Hydrachnidia) of Borneo and additional new species from Thailand and Sulawesi. The Raffles Bulletin of Zoology, 47, 409 - 439." type="journal article" year="1999">Wiles 1999</bibRefCitation>
, our study).
</paragraph>
</subSubSection>
</treatment>
</document>
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