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treatments-xml/data/CF/16/87/CF1687E1FFF4FB4B269A2C33FBD8D6CD.xml
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<document id="A2BC639DF3BEA394B291DDBFF4D092DA" ID-CLB-Dataset="31883" ID-DOI="10.11646/zootaxa.4329.3.7" ID-GBIF-Dataset="12141a60-1ea5-430f-b4c2-e105fb314388" ID-ISSN="1175-5326" ID-Zenodo-Dep="1002596" ID-ZooBank="848Bafaa-67C8-4105-B12E-F5C8B18Ff689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507186341443" checkinUser="plazi" docAuthor="Park, Kyung-Hwa" docDate="2017" docId="CF1687E1FFF4FB4B269A2C33FBD8D6CD" docLanguage="en" docName="zootaxa.4329.3.7.pdf" docOrigin="Zootaxa 4329 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leeonychiurus analis Park 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="299" masterDocId="332FFF99FFF7FB4C260D293AFFA9D761" masterDocTitle="Two new species of the genus Leeonychiurus Sun &amp; Arbea, 2014 (Collembola: Onychiuridae) from Korea" masterLastPageNumber="300" masterPageNumber="292" pageNumber="295" updateTime="1698469346388" updateUser="ExternalLinkService">
<mods:mods id="331F2CACD46A32AE753CC73F15489994" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FAF1BC0FE119A6B48DEBC96D6A46AF54">
<mods:title id="27EE1269579A862201B82B1E58F92DD4">Two new species of the genus Leeonychiurus Sun &amp; Arbea, 2014 (Collembola: Onychiuridae) from Korea</mods:title>
</mods:titleInfo>
<mods:name id="1E1BDFAF8107C590F285DE3EFFC3DF3D" type="personal">
<mods:role id="7FD6D41D74576FA4E687544E7AAB336A">
<mods:roleTerm id="A43A3117F9DF9BA59E9FB6914E3ECE3E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94F04031D960CB02B5F002C218917322">Park, Kyung-Hwa</mods:namePart>
</mods:name>
<mods:typeOfResource id="E0FE15FFCA55B76D7D2800883A3E16D5">text</mods:typeOfResource>
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<mods:title id="6F613372A341499C3A48CB6B44F665FB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DECF404BBAFA620B022805612B07C631">
<mods:date id="19F523A9F11CC9505ABEC4857137A4E6">2017</mods:date>
<mods:detail id="2B82AD103986F74E9803E6AD17DCDD46" type="pubDate">
<mods:number id="E6EA770CF5A34BC9F7E75655672AAF7D">2017-10-05</mods:number>
</mods:detail>
<mods:detail id="94A08542CC29E0ADA623B1C539EBD75A" type="volume">
<mods:number id="C38BA668A22E139CCE216AB35E5A909A">4329</mods:number>
</mods:detail>
<mods:detail id="0F92901373CD14C458256AF52D9F5E93" type="issue">
<mods:number id="F98BA1B590F2B26EBEEF764C6472AA25">3</mods:number>
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<mods:start id="0BB2DBAA262A7F1824AC4904AB193894">292</mods:start>
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<mods:classification id="500A9E8094691A1BB80606D54738F603">journal article</mods:classification>
<mods:identifier id="5C01186E6B5EAD16FC2947A7AC038A5F" type="CLB-Dataset">31883</mods:identifier>
<mods:identifier id="B0905EF1D50B3D6E00324E198297E9D5" type="DOI">10.11646/zootaxa.4329.3.7</mods:identifier>
<mods:identifier id="FA3CE0764477D24A0B08D679B04AD9A7" type="GBIF-Dataset">12141a60-1ea5-430f-b4c2-e105fb314388</mods:identifier>
<mods:identifier id="3B690264E5D1549E86F0D05520EFB1F7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9242AEDE6A3E25FADC8A71E73C0D6CEC" type="Zenodo-Dep">1002596</mods:identifier>
<mods:identifier id="6B9F4D4FDC999A3C494AE2693951D2AB" type="ZooBank">848Bafaa-67C8-4105-B12E-F5C8B18Ff689</mods:identifier>
</mods:mods>
<treatment id="CF1687E1FFF4FB4B269A2C33FBD8D6CD" ID-GBIF-Taxon="135111985" LSID="urn:lsid:plazi:treatment:CF1687E1FFF4FB4B269A2C33FBD8D6CD" httpUri="http://treatment.plazi.org/id/CF1687E1FFF4FB4B269A2C33FBD8D6CD" lastPageId="7" lastPageNumber="299" pageId="3" pageNumber="295">
<subSubSection id="0FA5657CFFF4FB4F269A2C33FEF4D224" pageId="3" pageNumber="295" type="nomenclature">
<paragraph id="470036F7FFF4FB4F269A2C33FDA8D242" blockId="3.[151,513,1289,1349]" box="[151,513,1289,1315]" pageId="3" pageNumber="295">
<heading id="1C48819BFFF4FB4F269A2C33FDA8D242" bold="true" box="[151,513,1289,1315]" fontSize="11" level="1" pageId="3" pageNumber="295" reason="1">
<emphasis id="75CBEAE5FFF4FB4F269A2C33FDA8D242" bold="true" box="[151,513,1289,1315]" pageId="3" pageNumber="295">
<taxonomicName id="80BF4D74FFF4FB4F269A2C33FE35D242" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[151,412,1289,1315]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF4FB4F269A2C33FE35D242" bold="true" box="[151,412,1289,1315]" italics="true" pageId="3" pageNumber="295">Leeonychiurus analis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EF8579EFFF4FB4F27AE2C33FDA8D242" box="[419,513,1289,1315]" pageId="3" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="470036F7FFF4FB4F269A2C16FEF4D224" blockId="3.[151,513,1289,1349]" box="[151,349,1324,1349]" pageId="3" pageNumber="295">
(
<figureCitation id="DF842A72FFF4FB4F26932C16FE8ED225" box="[158,295,1324,1349]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,249,1925,1947]" captionStartId-1="5.[151,249,1096,1118]" captionTargetBox-0="[511,1075,981,1886]" captionTargetBox-1="[491,1102,192,1087]" captionTargetId-0="figure@4.[511,1075,979,1886]" captionTargetId-1="figure@5.[484,1102,180,1087]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." captionText-1="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri-0="https://zenodo.org/record/1002602/files/figure.png" httpUri-1="https://zenodo.org/record/1002604/files/figure.png" pageId="3" pageNumber="295">Figures 3–4</figureCitation>
,
<figureCitation id="DF842A72FFF4FB4F273E2C16FEFDD224" box="[307,340,1324,1349]" captionStart="FIGURE 5" captionStartId="5.[151,249,1965,1987]" captionTargetBox="[342,1247,1201,1954]" captionTargetId="figure@5.[340,1247,1188,1955]" captionTargetPageId="5" captionText="FIGURE 5. Ventral side of Abd. II – VI: A, Leeonychiurus koreanus sp. nov. B, Leeonychiurus analis sp. nov. scale bars: 0.1 mm (A, B)." httpUri="https://zenodo.org/record/1002606/files/figure.png" pageId="3" pageNumber="295">5B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF4FB4F269A2C4EFEC5D299" pageId="3" pageNumber="295" type="diagnosis">
<paragraph id="470036F7FFF4FB4F269A2C4EFEC5D299" blockId="3.[151,1436,1396,2038]" pageId="3" pageNumber="295">
<emphasis id="75CBEAE5FFF4FB4F269A2C4EFEB8D2EC" bold="true" box="[151,273,1396,1421]" pageId="3" pageNumber="295">Diagnosis.</emphasis>
Dorsal pseudocellar formula 32/12(3)2/33343, ventral 11/000/0100; subcoxa 1 of legs I–III with one pseudocellus each. Postantennal organ with 18–22 compound vesicles. Abd. I–III terga with 3+3 chaetae along axial line. Th. I–III sterna with 0+0, 1+1, 1+1 chaetae respectively. Chaeta c0 on upper anal valve absent, but chaetae a1 present.
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF4FB4F26CA2F3EFDEAD104" pageId="3" pageNumber="295" type="materials_examined">
<paragraph id="470036F7FFF4FB4F26CA2F3EFDEAD104" blockId="3.[151,1436,1396,2038]" pageId="3" pageNumber="295">
<materialsCitation id="F7D73CAAFFF4FB4F26CA2F3EFDEAD104" ID-GBIF-Occurrence="1661822846" collectingDate="2015-03-27" country="South Korea" county="Sinwol-ri" latitude="127.28972" location="Wanju-gun" longLatPrecision="17" longitude="35.944447" municipality="Dongsang-myeon" pageId="3" pageNumber="295" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Jeollabuk-do" typeStatus="holotype">
<emphasis id="75CBEAE5FFF4FB4F26CA2F3EFED0D17C" bold="true" box="[199,377,1540,1565]" pageId="3" pageNumber="295">
<typeStatus id="98048855FFF4FB4F26CA2F3EFEABD17C" box="[199,258,1540,1565]" pageId="3" pageNumber="295">Type</typeStatus>
material.
</emphasis>
<typeStatus id="98048855FFF4FB4F27882F3EFE5BD17C" box="[389,498,1540,1565]" pageId="3" pageNumber="295" type="holotype">Holotype</typeStatus>
: female on slide,
<collectingCountry id="3FA87667FFF4FB4F24C52F3FFCA6D17D" box="[712,783,1541,1564]" name="South Korea" pageId="3" pageNumber="295">Korea</collectingCountry>
,
<collectingCounty id="AE614E7BFFF4FB4F25122F3EFC22D17C" box="[799,907,1540,1565]" pageId="3" pageNumber="295">Sinwol-ri</collectingCounty>
,
<collectingMunicipality id="A764AC8DFFF4FB4F25912F3FFBCFD17C" box="[924,1126,1541,1565]" pageId="3" pageNumber="295">Dongsang-myeon</collectingMunicipality>
,
<location id="4260602CFFF4FB4F227B2F3EFB5CD17C" LSID="urn:lsid:plazi:treatment:CF1687E1FFF4FB4B269A2C33FBD8D6CD:4260602CFFF4FB4F227B2F3EFB5CD17C" box="[1142,1269,1540,1565]" country="South Korea" county="Sinwol-ri" latitude="127.28972" longLatPrecision="17" longitude="35.944447" municipality="Dongsang-myeon" name="Wanju-gun" pageId="3" pageNumber="295" stateProvince="Jeollabuk-do">Wanju-gun</location>
,
<collectingRegion id="857BF815FFF4FB4F23092F3EFA31D17C" box="[1284,1432,1540,1565]" country="South Korea" name="Jeonbuk" pageId="3" pageNumber="295">Jeollabuk-do</collectingRegion>
, (
<geoCoordinate id="228B5030FFF4FB4F26922F12FE97D121" box="[159,318,1576,1600]" direction="north" orientation="latitude" pageId="3" pageNumber="295" precision="15" value="127.28972">N127°17´23´´</geoCoordinate>
<geoCoordinate id="228B5030FFF4FB4F274A2F12FE7ED120" box="[327,471,1576,1601]" direction="east" orientation="longitude" pageId="3" pageNumber="295" precision="15" value="35.944447">E35°56´40´´</geoCoordinate>
),
<date id="33011037FFF4FB4F27E02F12FDD0D120" box="[493,633,1576,1601]" pageId="3" pageNumber="295" value="2015-03-27">
<collectingDate id="2345E9DFFFF4FB4F27E02F12FDD0D120" box="[493,633,1576,1601]" pageId="3" pageNumber="295" value="2015-03-27">
27–III–
<quantity id="80479B12FFF4FB4F244C2F12FDD0D120" box="[577,633,1576,1601]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="3" pageNumber="295" unit="in" value="2015.0">2015</quantity>
</collectingDate>
</date>
in roots of grass, collected by K.H. Park.
<typeStatus id="98048855FFF4FB4F226B2F13FB71D120" box="[1126,1240,1577,1601]" pageId="3" pageNumber="295" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="51B9FD7EFFF4FB4F22E92F12FAFAD120" box="[1252,1363,1576,1601]" pageId="3" pageNumber="295" type="female">4 females</specimenCount>
and
<specimenCount id="51B9FD7EFFF4FB4F23832F12FF64D104" pageId="3" pageNumber="295" type="male">1 male</specimenCount>
on slides, same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF4FB4826CA2F4AFAC8D474" lastPageId="4" lastPageNumber="296" pageId="3" pageNumber="295" type="description">
<paragraph id="470036F7FFF4FB4F26CA2F4AFDF0D07F" blockId="3.[151,1436,1396,2038]" pageId="3" pageNumber="295">
<emphasis id="75CBEAE5FFF4FB4F26CA2F4AFEF0D1E8" bold="true" box="[199,345,1648,1673]" pageId="3" pageNumber="295">Description.</emphasis>
Body length (excluding antennae) of adults: male 1.65mm, females 1.80–2.12mm. Color white in alcohol. Cuticle granulation more or less uniform. Dorsal pseudocellar formula 32/12(3)2/33343 (
<figureCitation id="DF842A72FFF4FB4F22ED2FAEFA93D1CD" box="[1248,1338,1684,1709]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="3" pageNumber="295">Fig. 3A</figureCitation>
), ventral 11/000/0100; subcoxa 1 of legs I–III with one pseudocellus each. Parapseudocellar formula ventrally 0/000/
<quantity id="80479B12FFF4FB4F269A2FE5FF5ED188" box="[151,247,1755,1783]" metricMagnitude="5" metricUnit="m" metricValue="1.01001" pageId="3" pageNumber="295" unit="m" value="101001.0">101001m</quantity>
(
<figureCitation id="DF842A72FFF4FB4F270E2FE4FEF0D196" box="[259,345,1758,1783]" captionStart="FIGURE 5" captionStartId="5.[151,249,1965,1987]" captionTargetBox="[342,1247,1201,1954]" captionTargetId="figure@5.[340,1247,1188,1955]" captionTargetPageId="5" captionText="FIGURE 5. Ventral side of Abd. II – VI: A, Leeonychiurus koreanus sp. nov. B, Leeonychiurus analis sp. nov. scale bars: 0.1 mm (A, B)." httpUri="https://zenodo.org/record/1002606/files/figure.png" pageId="3" pageNumber="295">Fig. 5B</figureCitation>
), dorsal psx absent. Subcoxa 1 of legs I–III with 1 psx each. Psp formulas: 00/011/11110 dorsally and 00/111/0001m0 0 ventrally (
<figureCitation id="DF842A72FFF4FB4F27F92E3FFDE5D07F" box="[500,588,1797,1822]" captionStart="FIGURE 5" captionStartId="5.[151,249,1965,1987]" captionTargetBox="[342,1247,1201,1954]" captionTargetId="figure@5.[340,1247,1188,1955]" captionTargetPageId="5" captionText="FIGURE 5. Ventral side of Abd. II – VI: A, Leeonychiurus koreanus sp. nov. B, Leeonychiurus analis sp. nov. scale bars: 0.1 mm (A, B)." httpUri="https://zenodo.org/record/1002606/files/figure.png" pageId="3" pageNumber="295">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="470036F7FFF4FB4826CA2E13FA21D621" blockId="3.[151,1436,1396,2038]" lastBlockId="4.[151,1437,151,896]" lastPageId="4" lastPageNumber="296" pageId="3" pageNumber="295">
Head: Antennal bases well marked (
<figureCitation id="DF842A72FFF4FB4F246B2E13FD6BD023" box="[614,706,1833,1858]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="3" pageNumber="295">Fig. 3A</figureCitation>
). Antennae slightly shorter than head; ratio antenna/head=0.83– 0.88. Ant. IV subapical organite with globular apex; invaginated apical bulb absent; microsensillum in lateroexternal position approximately one-third length from the base; sensilla not well distinguish from ordinary chaetae (
<figureCitation id="DF842A72FFF4FB4F26922EAFFF52D0CF" box="[159,251,1941,1966]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="3" pageNumber="295">Fig. 3D</figureCitation>
). Ant. III sensory organ composed of five papillae, five guard chaetae, 2 rods and 2 granulated sensory clubs (
<figureCitation id="DF842A72FFF4FB4F26EF2E83FE95D0B3" box="[226,316,1977,2002]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="3" pageNumber="295">Fig. 3C</figureCitation>
); lateral ms just behind sensory organ. Ant. II with 16 chaetae. Ant. I with 9 chaetae. Postantennal organ with 18–22 compound vesicles arranged in two rows along axis of organ (
<figureCitation id="DF842A72FFF4FB4F22492EE7FB0BD097" box="[1092,1186,2013,2038]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="3" pageNumber="295">Fig. 4A</figureCitation>
). Head with unpaired dorsal chaeta d0. 4+4 p-chaetae between two inner posterior pso on head, p1 anterior to others. Mandible with strong molar plate and four apical teeth, maxilla bearing 3 teeth and 6 lamellae. Maxillary palp simple with 1 basal chaeta and 2 sublobal hairs. Labral formula as 4/342 (
<figureCitation id="DF842A72FFF3FB4824F329E5FCF0D799" box="[766,857,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="4" pageNumber="296">Fig. 4C</figureCitation>
). Labial palp of
<typeStatus id="98048855FFF3FB48221B29DAFBEFD799" box="[1046,1094,224,248]" pageId="4" pageNumber="296">type</typeStatus>
AC (
<figureCitation id="DF842A72FFF3FB48228B29E5FB49D799" box="[1158,1248,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="4" pageNumber="296">Fig. 4B</figureCitation>
: After
<bibRefCitation id="232E4B06FFF3FB48233D29E5FF7AD67D" author="Fjellberg" pageId="4" pageNumber="296" refString="Fjellberg, A. (1999) The labial palp in Collembola. Zoologischer Anzeiger, 237, 309 - 330." type="journal article" year="1999">Fjellberg, 1999</bibRefCitation>
), labial papillae A, B, C, D and E with 1, 4, 0, 3, 3 guard chaetae respectively (
<figureCitation id="DF842A72FFF3FB482272283EFB73D67D" box="[1151,1242,260,285]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="4" pageNumber="296">Fig. 4B</figureCitation>
). Labium with 6 proximal, 4 basomedian and 6 basolateral chaetae. 4+4 postlabial chaetae present along ventral groove (
<figureCitation id="DF842A72FFF3FB48232E281DFAD2D621" box="[1315,1403,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="4" pageNumber="296">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="470036F7FFF3FB4826CA2876FBD3D579" blockId="4.[151,1437,151,896]" pageId="4" pageNumber="296">
Body chaetotaxy: s-chaeta formula as 2/012/222120 dorsally. Tiny and blunt ms, present on Th. II and III. Th. I tergum with 8+8 chaetae on each side. Th. II–Abd. III with 3, 3, 3, 3, 3 chaetae respectively on both sides of axial line and without unpaired axial chaetae, sometimes with asymmetrical chaetae. Abd. IV tergum with unpaired axial chaeta p0, sometimes with m0 chaetae; Abd. V tergum with axial chaeta m0; Abd. VI tergum with 2 unpaired axial chaetae: a0, p0 (
<figureCitation id="DF842A72FFF3FB48275628E6FE13D695" box="[347,442,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,249,1925,1947]" captionTargetBox="[511,1075,981,1886]" captionTargetId="figure@4.[511,1075,979,1886]" captionTargetPageId="4" captionText="FIGURE 3. Leeonychiurus analis sp. nov.: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1 mm (A, B), 0.01 mm (C, D)." httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="4" pageNumber="296">Fig. 3A</figureCitation>
). Th. I, II and III sterna with 0+0, 1+1 and 1+1 chaetae between legs, respectively. Subcoxae 1 of legs I–III with 4, 5, 5 chaetae respectively, subcoxa 2 with 1, 4, 4 chaetae.
</paragraph>
<paragraph id="470036F7FFF3FB4826CA2B1EFEAFD5AD" blockId="4.[151,1437,151,896]" pageId="4" pageNumber="296">
Appendages: Tibiotarsi I, II and III with 22 (11, 8, 3), 21 (11, 8, 2) and 21 (11, 8, 2) chaetae, respectively. Claw without teeth. Empodial appendage with basal lamella, appendage length shorter than inner edge of unguis. Ventral tube with 8–9+8–9 distal chaetae, without anterior and basal chaetae. Furca reduced to a finely granulated area, with four small dental chaetae in one row posterior to furcal rudiment; three manubrial rows of chaetae present (
<figureCitation id="DF842A72FFF3FB4826922B8EFF53D5AD" box="[159,250,692,717]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="4" pageNumber="296">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="470036F7FFF3FB4826CA2BEDFAC8D474" blockId="4.[151,1437,151,896]" pageId="4" pageNumber="296">
Anal valves chaetotaxy: each lateral valve with chaetae a0, 2a1, b0 and 2b1; upper valve with chaetae a0, 2a1, b0, 2b1, 2b2, 2c1 and 2c2 (
<figureCitation id="DF842A72FFF3FB4827CA2BC6FDB6D475" box="[455,543,764,789]" captionStart="FIGURE 4" captionStartId="5.[151,249,1096,1118]" captionTargetBox="[491,1102,192,1087]" captionTargetId="figure@5.[484,1102,180,1087]" captionTargetPageId="5" captionText="FIGURE 4. Leeonychiurus analis sp. nov.: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05 mm (A, D, E), 0.01 mm (B, C)." httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="4" pageNumber="296">Fig. 4E</figureCitation>
). Anal spines set on distinct papillae, as long as inner edge of hind unguis.
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF3FB4826CA2A25FD95D459" box="[199,572,799,824]" pageId="4" pageNumber="296" type="biology_ecology">
<paragraph id="470036F7FFF3FB4826CA2A25FD95D459" blockId="4.[151,1437,151,896]" box="[199,572,799,824]" pageId="4" pageNumber="296">
<emphasis id="75CBEAE5FFF3FB4826CA2A25FE82D459" bold="true" box="[199,299,799,824]" pageId="4" pageNumber="296">Ecology.</emphasis>
Found in roots of grass.
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF3FB4A26CA2A7EFBB5D100" lastPageId="6" lastPageNumber="298" pageId="4" pageNumber="296" type="etymology">
<paragraph id="470036F7FFF3FB4826CA2A7EFD93D4E1" blockId="4.[151,1437,151,896]" pageId="4" pageNumber="296">
<emphasis id="75CBEAE5FFF3FB4826CA2A7EFEE7D43C" bold="true" box="[199,334,836,861]" pageId="4" pageNumber="296">Etymology.</emphasis>
The name of the species refers to the anal lobe that has specific chaetotaxy. Korean name of the species: Dongsang-lee-eori-tok-to-gi.
</paragraph>
<caption id="13C0667FFFF3FB48269A2EBFFCA8D0DB" httpUri="https://zenodo.org/record/1002602/files/figure.png" pageId="4" pageNumber="296" targetBox="[511,1075,981,1886]" targetPageId="4">
<paragraph id="470036F7FFF3FB48269A2EBFFCA8D0DB" blockId="4.[151,1435,1925,1978]" pageId="4" pageNumber="296">
<emphasis id="75CBEAE5FFF3FB48269A2EBFFEBDD0FB" bold="true" box="[151,276,1925,1947]" pageId="4" pageNumber="296">FIGURE 3.</emphasis>
<taxonomicName id="80BF4D74FFF3FB4827102EBFFE50D0FA" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[285,505,1925,1947]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF3FB4827102EBFFE50D0FA" box="[285,505,1925,1947]" italics="true" pageId="4" pageNumber="296">Leeonychiurus analis</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF3FB48240C2EBFFDFDD0FB" bold="true" box="[513,596,1925,1946]" pageId="4" pageNumber="296">
<taxonomicNameLabel id="6EF8579EFFF3FB48240C2EBFFDFDD0FB" box="[513,596,1925,1946]" pageId="4" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, Dorsal chaetotaxy. B, Ventral chaetotoaxy on head. C, Ant. III with sensory club. D, Antenna. Scale bars: 0.1mm (A, B), 0.01mm (C, D).
</paragraph>
</caption>
<caption id="13C0667FFFF2FB49269A2D72FC8CD31C" httpUri="https://zenodo.org/record/1002604/files/figure.png" pageId="5" pageNumber="297" targetBox="[491,1102,192,1087]" targetPageId="5">
<paragraph id="470036F7FFF2FB49269A2D72FC8CD31C" blockId="5.[151,1435,1096,1149]" pageId="5" pageNumber="297">
<emphasis id="75CBEAE5FFF2FB49269A2D72FEBAD33C" bold="true" box="[151,275,1096,1118]" pageId="5" pageNumber="297">FIGURE 4.</emphasis>
<taxonomicName id="80BF4D74FFF2FB4927172D72FE5CD33F" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[282,501,1096,1118]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF2FB4927172D72FE5CD33F" box="[282,501,1096,1118]" italics="true" pageId="5" pageNumber="297">Leeonychiurus analis</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF2FB4927F12D73FDE4D33F" bold="true" box="[508,589,1097,1118]" pageId="5" pageNumber="297">
<taxonomicNameLabel id="6EF8579EFFF2FB4927F12D73FDE4D33F" box="[508,589,1097,1118]" pageId="5" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, PAO. B, Labial palp. C, Labrum. D, Central part of Abd. IV sternum. E, Anal lobes chaetotaxy. Scale bars: 0.05mm (A, D, E), 0.01mm (B, C).
</paragraph>
</caption>
<caption id="13C0667FFFF2FB49269A2E97FE82D083" httpUri="https://zenodo.org/record/1002606/files/figure.png" pageId="5" pageNumber="297" targetBox="[342,1247,1201,1954]" targetPageId="5">
<paragraph id="470036F7FFF2FB49269A2E97FE82D083" blockId="5.[151,1437,1965,2018]" pageId="5" pageNumber="297">
<emphasis id="75CBEAE5FFF2FB49269A2E97FEBCD0A2" bold="true" box="[151,277,1965,1987]" pageId="5" pageNumber="297">FIGURE 5.</emphasis>
Ventral side of Abd. II–VI: A,
<taxonomicName id="80BF4D74FFF2FB4924642E97FCCAD0A2" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[617,867,1965,1987]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="75CBEAE5FFF2FB4924642E97FCCAD0A2" box="[617,867,1965,1987]" italics="true" pageId="5" pageNumber="297">Leeonychiurus koreanus</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF2FB4925612E94FC69D0A2" bold="true" box="[876,960,1966,1987]" pageId="5" pageNumber="297">
<taxonomicNameLabel id="6EF8579EFFF2FB4925612E94FC69D0A2" box="[876,960,1966,1987]" pageId="5" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
B,
<taxonomicName id="80BF4D74FFF2FB4925E62E97FB61D0A2" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[1003,1224,1965,1987]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF2FB4925E62E97FB61D0A2" box="[1003,1224,1965,1987]" italics="true" pageId="5" pageNumber="297">Leeonychiurus analis</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF2FB4922DC2E94FA8CD0A2" bold="true" box="[1233,1317,1966,1987]" pageId="5" pageNumber="297">
<taxonomicNameLabel id="6EF8579EFFF2FB4922DC2E94FA8CD0A2" box="[1233,1317,1966,1987]" pageId="5" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
scale bars: 0.1mm (A, B).
</paragraph>
</caption>
<paragraph id="470036F7FFF1FB4A26702B02FCF4D52C" blockId="6.[125,861,568,589]" box="[125,861,568,589]" pageId="6" pageNumber="298">
<emphasis id="75CBEAE5FFF1FB4A26702B02FF4BD52C" bold="true" box="[125,226,568,589]" pageId="6" pageNumber="298">TABLE].</emphasis>
Major characters of all known species in the genus
<emphasis id="75CBEAE5FFF1FB4A24C62B02FCF4D52C" box="[715,861,568,589]" italics="true" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A24C62B02FCF0D52C" authorityName="Park" authorityYear="2017" box="[715,857,568,589]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="genus">Leeonychiurus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="470036F7FFF1FB4A27012BA0F8BAD291" pageId="6" pageNumber="298">
<table id="35BFC457FFF104B326702B4CF89AD291" box="[125,1843,630,1520]" gridcols="8" gridrows="9" pageId="6" pageNumber="298">
<tr id="F98F34B5FFF104B326702B4CF89AD37E" box="[125,1843,630,1055]" gridrow="0" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702B4CFE48D37E" box="[125,481,630,1055]" gridcol="0" gridrow="0" pageId="6" pageNumber="298">character Dorsal pso Ventral pso Number of pso on subcoxa 1 of legs I6III Number of psx on subcoxa 1 of legs I6III Number of chaetae on subcoxa1 of legs I6III Number of vesicles on PAΟ</th>
<th id="BA5E5DC9FFF104B3241A2B4CFD33D37E" box="[535,666,630,1055]" gridcol="1" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A24112B41FDDAD5F1" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[540,627,635,656]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="75CBEAE5FFF1FB4A24112B41FDDAD5F1" box="[540,627,635,656]" italics="true" pageId="6" pageNumber="298">koreanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EF8579EFFF1FB4A24742B41FDC7D5CE" pageId="6" pageNumber="298" rank="species">
<emphasis id="75CBEAE5FFF1FB4A24742B41FDCED5CE" bold="true" pageId="6" pageNumber="298">sp. nov</emphasis>
.
</taxonomicNameLabel>
32/O22/33243 11/OOO/O1OO 1/1/1 absent 3,3,3 13615
</th>
<th id="BA5E5DC9FFF104B324C82B4CFCFED37E" box="[709,855,630,1055]" gridcol="2" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A24C52B41FCABD5F1" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[712,770,635,656]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF1FB4A24C52B41FCABD5F1" box="[712,770,635,656]" italics="true" pageId="6" pageNumber="298">analis</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF1FB4A25052B41FCFAD5F1" bold="true" box="[776,851,635,656]" pageId="6" pageNumber="298">
<taxonomicNameLabel id="6EF8579EFFF1FB4A25052B41FCFAD5F1" box="[776,851,635,656]" pageId="6" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
32/12(3)2/3334 3 11/OOO/O1OO 1/1/1 1/1/1 4, 5, 5 18622
</th>
<th id="BA5E5DC9FFF104B3258C2B4CFBB1D37E" box="[897,1048,630,1055]" gridcol="3" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A25812B41FC45D5F1" authorityName="Sun &amp; Zhang" authorityYear="2012" box="[908,1004,635,656]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="gulinensis">
<emphasis id="75CBEAE5FFF1FB4A25812B41FC45D5F1" box="[908,1004,635,656]" italics="true" pageId="6" pageNumber="298">gulinensis</emphasis>
</taxonomicName>
1), 2) (Sun &amp; Zhang, 2O12) 32/122/33343 11/OOO/O1O1 1/1/1 1/1/1 465, 4–5, 4–5 2O621
</th>
<th id="BA5E5DC9FFF104B3225D2B4CFB74D37E" box="[1104,1245,630,1055]" gridcol="4" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A22592B41FB1FD5F1" authorityName="Sun &amp; Zhang" authorityYear="2012" box="[1108,1206,635,656]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="75CBEAE5FFF1FB4A22592B41FB1FD5F1" box="[1108,1206,635,656]" italics="true" pageId="6" pageNumber="298">antennalis</emphasis>
</taxonomicName>
1), 2) (Sun &amp; Zhang, 2O12) 32/122/33343 11/OOO/O1O1 1/1/1 absent 4,4,4 5266O
</th>
<th id="BA5E5DC9FFF104B323042B4CFA09D37E" box="[1289,1440,630,1055]" gridcol="5" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A231A2B41FA2CD5F1" authorityName="Sun &amp; Arbea" authorityYear="2014" box="[1303,1413,635,656]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="fusongensis">
<emphasis id="75CBEAE5FFF1FB4A231A2B41FA2CD5F1" box="[1303,1413,635,656]" italics="true" pageId="6" pageNumber="298">fusongensis</emphasis>
</taxonomicName>
2) Sun &amp; Arbea, 2O14 32/133/33343 11/OOO/O112 1/1/1 1/1/1 5,6,6 13616
</th>
<th id="BA5E5DC9FFF104B323B02B4CF93AD37E" box="[1469,1683,630,1055]" gridcol="6" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A23B02B41F9EAD5F1" baseAuthorityName="Lee &amp; Park" baseAuthorityYear="1986" box="[1469,1603,635,656]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="polychaetosus">
<emphasis id="75CBEAE5FFF1FB4A23B02B41F9EAD5F1" box="[1469,1603,635,656]" italics="true" pageId="6" pageNumber="298">polychaetosus</emphasis>
</taxonomicName>
2), 3) (Lee &amp; Park, 1986) 22/233/33343 O1/OOO/O112 1/1/1? 14618, 1O612, 8612 3O636
</th>
<th id="BA5E5DC9FFF104B320BD2B4CF89AD37E" box="[1712,1843,630,1055]" gridcol="7" gridrow="0" pageId="6" pageNumber="298">
<taxonomicName id="80BF4D74FFF1FB4A20D42B41F955D5F1" authorityName="Wray" authorityYear="1950" box="[1753,1788,635,656]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="mai">
<emphasis id="75CBEAE5FFF1FB4A20D42B41F955D5F1" box="[1753,1788,635,656]" italics="true" pageId="6" pageNumber="298">mai</emphasis>
</taxonomicName>
4) (Wray, 195O) 32/133/33343 11/OOO/OOO1 1/1/1 absent? 15
</th>
</tr>
<tr id="F98F34B5FFF104B326702D78F89AD336" box="[125,1843,1090,1111]" gridrow="1" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702D78FE48D336" box="[125,481,1090,1111]" gridcol="0" gridrow="1" pageId="6" pageNumber="298">Number of chaetae on Th. I</th>
<td id="BA5E5DC9FFF104B3241A2D78FD33D336" box="[535,666,1090,1111]" gridcol="1" gridrow="1" pageId="6" pageNumber="298">668+668</td>
<td id="BA5E5DC9FFF104B324C82D78FCFED336" box="[709,855,1090,1111]" gridcol="2" gridrow="1" pageId="6" pageNumber="298">8+8</td>
<td id="BA5E5DC9FFF104B3258C2D78FBB1D336" box="[897,1048,1090,1111]" gridcol="3" gridrow="1" pageId="6" pageNumber="298">768+768</td>
<td id="BA5E5DC9FFF104B3225D2D78FB74D336" box="[1104,1245,1090,1111]" gridcol="4" gridrow="1" pageId="6" pageNumber="298">768+768</td>
<td id="BA5E5DC9FFF104B323042D78FA09D336" box="[1289,1440,1090,1111]" gridcol="5" gridrow="1" pageId="6" pageNumber="298">861O+861O</td>
<td id="BA5E5DC9FFF104B323B02D78F93AD336" box="[1469,1683,1090,1111]" gridcol="6" gridrow="1" pageId="6" pageNumber="298">13616+13616</td>
<td id="BA5E5DC9FFF104B320BD2D78F89AD336" box="[1712,1843,1090,1111]" gridcol="7" gridrow="1" pageId="6" pageNumber="298">?</td>
</tr>
<tr id="F98F34B5FFF104B326702D43F89AD3EF" box="[125,1843,1145,1166]" gridrow="2" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702D43FE48D3EF" box="[125,481,1145,1166]" gridcol="0" gridrow="2" pageId="6" pageNumber="298">Unpaiređ chaetae on Abđ. IV</th>
<td id="BA5E5DC9FFF104B3241A2D43FD33D3EF" box="[535,666,1145,1166]" gridcol="1" gridrow="2" pageId="6" pageNumber="298">pO</td>
<td id="BA5E5DC9FFF104B324C82D43FCFED3EF" box="[709,855,1145,1166]" gridcol="2" gridrow="2" pageId="6" pageNumber="298">(mO), pO</td>
<td id="BA5E5DC9FFF104B3258C2D43FBB1D3EF" box="[897,1048,1145,1166]" gridcol="3" gridrow="2" pageId="6" pageNumber="298">mO, pO</td>
<td id="BA5E5DC9FFF104B3225D2D43FB74D3EF" box="[1104,1245,1145,1166]" gridcol="4" gridrow="2" pageId="6" pageNumber="298">mO, pO</td>
<td id="BA5E5DC9FFF104B323042D43FA09D3EF" box="[1289,1440,1145,1166]" gridcol="5" gridrow="2" pageId="6" pageNumber="298">pO</td>
<td id="BA5E5DC9FFF104B323B02D43F93AD3EF" box="[1469,1683,1145,1166]" gridcol="6" gridrow="2" pageId="6" pageNumber="298">pO</td>
<td id="BA5E5DC9FFF104B320BD2D43F89AD3EF" box="[1712,1843,1145,1166]" gridcol="7" gridrow="2" pageId="6" pageNumber="298">pO</td>
</tr>
<tr id="F98F34B5FFF104B326702D8BF89AD3A7" box="[125,1843,1201,1222]" gridrow="3" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702D8BFE48D3A7" box="[125,481,1201,1222]" gridcol="0" gridrow="3" pageId="6" pageNumber="298">Unpaiređ chaetae on Abđ. V</th>
<td id="BA5E5DC9FFF104B3241A2D8BFD33D3A7" box="[535,666,1201,1222]" gridcol="1" gridrow="3" pageId="6" pageNumber="298">mO</td>
<td id="BA5E5DC9FFF104B324C82D8BFCFED3A7" box="[709,855,1201,1222]" gridcol="2" gridrow="3" pageId="6" pageNumber="298">mO</td>
<td id="BA5E5DC9FFF104B3258C2D8BFBB1D3A7" box="[897,1048,1201,1222]" gridcol="3" gridrow="3" pageId="6" pageNumber="298">pO</td>
<td id="BA5E5DC9FFF104B3225D2D8BFB74D3A7" box="[1104,1245,1201,1222]" gridcol="4" gridrow="3" pageId="6" pageNumber="298">absent</td>
<td id="BA5E5DC9FFF104B323042D8BFA09D3A7" box="[1289,1440,1201,1222]" gridcol="5" gridrow="3" pageId="6" pageNumber="298">pO</td>
<td id="BA5E5DC9FFF104B323B02D8BF93AD3A7" box="[1469,1683,1201,1222]" gridcol="6" gridrow="3" pageId="6" pageNumber="298">mO, pO</td>
<td id="BA5E5DC9FFF104B320BD2D8BF89AD3A7" box="[1712,1843,1201,1222]" gridcol="7" gridrow="3" pageId="6" pageNumber="298">mO</td>
</tr>
<tr id="F98F34B5FFF104B326702DD2F89AD39C" box="[125,1843,1256,1277]" gridrow="4" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702DD2FE48D39C" box="[125,481,1256,1277]" gridcol="0" gridrow="4" pageId="6" pageNumber="298">Unpaiređ chaetae on Abđ. VI</th>
<td id="BA5E5DC9FFF104B3241A2DD2FD33D39C" box="[535,666,1256,1277]" gridcol="1" gridrow="4" pageId="6" pageNumber="298">aO, pO</td>
<td id="BA5E5DC9FFF104B324C82DD2FCFED39C" box="[709,855,1256,1277]" gridcol="2" gridrow="4" pageId="6" pageNumber="298">aO, pO</td>
<td id="BA5E5DC9FFF104B3258C2DD2FBB1D39C" box="[897,1048,1256,1277]" gridcol="3" gridrow="4" pageId="6" pageNumber="298">aO, mO</td>
<td id="BA5E5DC9FFF104B3225D2DD2FB74D39C" box="[1104,1245,1256,1277]" gridcol="4" gridrow="4" pageId="6" pageNumber="298">aO, mO</td>
<td id="BA5E5DC9FFF104B323042DD2FA09D39C" box="[1289,1440,1256,1277]" gridcol="5" gridrow="4" pageId="6" pageNumber="298">aO, mO</td>
<td id="BA5E5DC9FFF104B323B02DD2F93AD39C" box="[1469,1683,1256,1277]" gridcol="6" gridrow="4" pageId="6" pageNumber="298">aO, mO, pO</td>
<td id="BA5E5DC9FFF104B320BD2DD2F89AD39C" box="[1712,1843,1256,1277]" gridcol="7" gridrow="4" pageId="6" pageNumber="298">aO, pO</td>
</tr>
<tr id="F98F34B5FFF104B326702C1AF89AD254" box="[125,1843,1312,1333]" gridrow="5" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702C1AFE48D254" box="[125,481,1312,1333]" gridcol="0" gridrow="5" pageId="6" pageNumber="298">Number of chaetae on ventral tube</th>
<td id="BA5E5DC9FFF104B3241A2C1AFD33D254" box="[535,666,1312,1333]" gridcol="1" gridrow="5" pageId="6" pageNumber="298">667+667</td>
<td id="BA5E5DC9FFF104B324C82C1AFCFED254" box="[709,855,1312,1333]" gridcol="2" gridrow="5" pageId="6" pageNumber="298">869+869</td>
<td id="BA5E5DC9FFF104B3258C2C1AFBB1D254" box="[897,1048,1312,1333]" gridcol="3" gridrow="5" pageId="6" pageNumber="298">869+869</td>
<td id="BA5E5DC9FFF104B3225D2C1AFB74D254" box="[1104,1245,1312,1333]" gridcol="4" gridrow="5" pageId="6" pageNumber="298">768+768</td>
<td id="BA5E5DC9FFF104B323042C1AFA09D254" box="[1289,1440,1312,1333]" gridcol="5" gridrow="5" pageId="6" pageNumber="298">869+869</td>
<td id="BA5E5DC9FFF104B323B02C1AF93AD254" box="[1469,1683,1312,1333]" gridcol="6" gridrow="5" pageId="6" pageNumber="298">27637+27637</td>
<td id="BA5E5DC9FFF104B320BD2C1AF89AD254" box="[1712,1843,1312,1333]" gridcol="7" gridrow="5" pageId="6" pageNumber="298">7+7</td>
</tr>
<tr id="F98F34B5FFF104B326702C60F89AD20E" box="[125,1843,1370,1391]" gridrow="6" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702C60FE48D20E" box="[125,481,1370,1391]" gridcol="0" gridrow="6" pageId="6" pageNumber="298">Number of chaetae on Th. I6III sterna</th>
<td id="BA5E5DC9FFF104B3241A2C60FD33D20E" box="[535,666,1370,1391]" gridcol="1" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1+1</td>
<td id="BA5E5DC9FFF104B324C82C60FCFED20E" box="[709,855,1370,1391]" gridcol="2" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1+1</td>
<td id="BA5E5DC9FFF104B3258C2C60FBB1D20E" box="[897,1048,1370,1391]" gridcol="3" gridrow="6" pageId="6" pageNumber="298">O+O/2+2/2+2(3)</td>
<td id="BA5E5DC9FFF104B3225D2C60FB74D20E" box="[1104,1245,1370,1391]" gridcol="4" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1+1</td>
<td id="BA5E5DC9FFF104B323042C60FA09D20E" box="[1289,1440,1370,1391]" gridcol="5" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1(2)+1</td>
<td id="BA5E5DC9FFF104B323B02C60F93AD20E" box="[1469,1683,1370,1391]" gridcol="6" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1(4)+1</td>
<td id="BA5E5DC9FFF104B320BD2C60F89AD20E" box="[1712,1843,1370,1391]" gridcol="7" gridrow="6" pageId="6" pageNumber="298">O+O/1+1/1+1</td>
</tr>
<tr id="F98F34B5FFF104B326702CA3F89AD2CF" box="[125,1843,1433,1454]" gridrow="7" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702CA3FE48D2CF" box="[125,481,1433,1454]" gridcol="0" gridrow="7" pageId="6" pageNumber="298">Chaetae a1 on upper anal valve</th>
<td id="BA5E5DC9FFF104B3241A2CA3FD33D2CF" box="[535,666,1433,1454]" gridcol="1" gridrow="7" pageId="6" pageNumber="298">absent</td>
<td id="BA5E5DC9FFF104B324C82CA3FCFED2CF" box="[709,855,1433,1454]" gridcol="2" gridrow="7" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B3258C2CA3FBB1D2CF" box="[897,1048,1433,1454]" gridcol="3" gridrow="7" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B3225D2CA3FB74D2CF" box="[1104,1245,1433,1454]" gridcol="4" gridrow="7" pageId="6" pageNumber="298">absent</td>
<td id="BA5E5DC9FFF104B323042CA3FA09D2CF" box="[1289,1440,1433,1454]" gridcol="5" gridrow="7" pageId="6" pageNumber="298">absent</td>
<td id="BA5E5DC9FFF104B323B02CA3F93AD2CF" box="[1469,1683,1433,1454]" gridcol="6" gridrow="7" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B320BD2CA3F89AD2CF" box="[1712,1843,1433,1454]" gridcol="7" gridrow="7" pageId="6" pageNumber="298">present</td>
</tr>
<tr id="F98F34B5FFF104B326702CE1F89AD291" box="[125,1843,1499,1520]" gridrow="8" pageId="6" pageNumber="298">
<th id="BA5E5DC9FFF104B326702CE1FE48D291" box="[125,481,1499,1520]" gridcol="0" gridrow="8" pageId="6" pageNumber="298">Chaeta cO on upper anal valve</th>
<td id="BA5E5DC9FFF104B3241A2CE1FD33D291" box="[535,666,1499,1520]" gridcol="1" gridrow="8" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B324C82CE1FCFED291" box="[709,855,1499,1520]" gridcol="2" gridrow="8" pageId="6" pageNumber="298">absent</td>
<td id="BA5E5DC9FFF104B3258C2CE1FBB1D291" box="[897,1048,1499,1520]" gridcol="3" gridrow="8" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B3225D2CE1FB74D291" box="[1104,1245,1499,1520]" gridcol="4" gridrow="8" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B323042CE1FA09D291" box="[1289,1440,1499,1520]" gridcol="5" gridrow="8" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B323B02CE1F93AD291" box="[1469,1683,1499,1520]" gridcol="6" gridrow="8" pageId="6" pageNumber="298">present</td>
<td id="BA5E5DC9FFF104B320BD2CE1F89AD291" box="[1712,1843,1499,1520]" gridcol="7" gridrow="8" pageId="6" pageNumber="298">present</td>
</tr>
</table>
</paragraph>
<paragraph id="470036F7FFF1FB4A20DB2F37FBB5D100" blockId="6.[125,1817,1549,1633]" pageId="6" pageNumber="298">
North Distribution Korea Korea China China China Korea (cave) America
<emphasis id="75CBEAE5FFF1FB4A26832F76FEAAD100" bold="true" box="[142,259,1612,1633]" pageId="6" pageNumber="298">References:</emphasis>
1) Sun &amp; Zhang 2O12; 2) Sun &amp; Arbea 2O14; 3)
<bibRefCitation id="232E4B06FFF1FB4A24D92F76FCD1D100" author="Lee" box="[724,888,1612,1633]" pageId="6" pageNumber="298" refString="Lee, B. H. &amp; Park, K. H. (1986) Three new species of Onychiuridae (Collembola) from a Korean cave. The Korean Journal of Systematic Zoology, 2 (1), 11 - 20." type="journal article" year="1986">Lee &amp; Park 1986</bibRefCitation>
; 4) Bernarđ 2O15
</paragraph>
</subSubSection>
<subSubSection id="0FA5657CFFF0FB4B26CA29ADFBD8D6CD" pageId="7" pageNumber="299" type="discussion">
<paragraph id="470036F7FFF0FB4B26CA29ADFBD8D6CD" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="299">
<emphasis id="75CBEAE5FFF0FB4B26CA29ADFEE5D7D1" bold="true" box="[199,332,151,176]" pageId="7" pageNumber="299">Discussion.</emphasis>
This new species is most similar to
<taxonomicName id="80BF4D74FFF0FB4B250529A3FC26D7D1" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[776,911,152,176]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="75CBEAE5FFF0FB4B250529A3FC26D7D1" box="[776,911,152,176]" italics="true" pageId="7" pageNumber="299">L. koreanus</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF0FB4B259129ADFC53D7D1" bold="true" box="[924,1018,151,176]" pageId="7" pageNumber="299">
<taxonomicNameLabel id="6EF8579EFFF0FB4B259129ADFC53D7D1" box="[924,1018,151,176]" pageId="7" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
having identical number of ventral pseudocelli (11/000/0100) and the same set of unpaired axial chaetae on Abd. IV–VI. Nevertheless it can easily be distinguished due to the presence of dorsal pso on Th. I (absent in
<taxonomicName id="80BF4D74FFF0FB4B259B29DBFBB3D799" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[918,1050,224,248]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="75CBEAE5FFF0FB4B259B29DBFBB3D799" box="[918,1050,224,248]" italics="true" pageId="7" pageNumber="299">L. koreanus</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF0FB4B222929E5FBD6D799" bold="true" box="[1060,1151,223,248]" pageId="7" pageNumber="299">
<taxonomicNameLabel id="6EF8579EFFF0FB4B222929E5FBD6D799" box="[1060,1151,223,248]" pageId="7" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and three dorsal pso on Abd. III (only two pso in
<taxonomicName id="80BF4D74FFF0FB4B27B2283FFDEAD67D" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[447,579,260,284]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="75CBEAE5FFF0FB4B27B2283FFDEAD67D" box="[447,579,260,284]" italics="true" pageId="7" pageNumber="299">L. koreanus</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF0FB4B2441283EFD0ED67C" bold="true" box="[588,679,260,285]" pageId="7" pageNumber="299">
<taxonomicNameLabel id="6EF8579EFFF0FB4B2441283EFD0ED67C" box="[588,679,260,285]" pageId="7" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), chaetotaxy of upper anal valve appears to be also rather specific (Table.1). The new species is also close to
<taxonomicName id="80BF4D74FFF0FB4B248A2813FBB1D621" authority="(Sun &amp; Zhang, 2012)" authorityName="Sun &amp; Zhang" authorityYear="2012" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" box="[647,1048,295,320]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="gulinensis">
<emphasis id="75CBEAE5FFF0FB4B248A2813FCB0D621" box="[647,793,296,320]" italics="true" pageId="7" pageNumber="299">L. gulinensis</emphasis>
(
<bibRefCitation id="232E4B06FFF0FB4B25242812FBB9D621" author="Sun" box="[809,1040,295,320]" pageId="7" pageNumber="299" refString="Sun, X. &amp; Zhang, F. (2012) Two new species of Onychiurus (Collembola: Onychiuridae) from Eastern China. Journal of Natural History, 46 (31 - 32), 1895 - 1904. https: // doi. org / 10.1080 / 00222933.2012.707236" type="journal article" year="2012">Sun &amp; Zhang, 2012</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="80BF4D74FFF0FB4B225E2813FF71D605" authority="(Sun &amp; Zhang, 2012)" authorityName="Sun &amp; Zhang" authorityYear="2012" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="75CBEAE5FFF0FB4B225E2813FB4ED621" box="[1107,1255,296,320]" italics="true" pageId="7" pageNumber="299">L. antennalis</emphasis>
(
<bibRefCitation id="232E4B06FFF0FB4B22F52812FF78D605" author="Sun" pageId="7" pageNumber="299" refString="Sun, X. &amp; Zhang, F. (2012) Two new species of Onychiurus (Collembola: Onychiuridae) from Eastern China. Journal of Natural History, 46 (31 - 32), 1895 - 1904. https: // doi. org / 10.1080 / 00222933.2012.707236" type="journal article" year="2012">Sun &amp; Zhang, 2012</bibRefCitation>
)
</taxonomicName>
as sharing same dorsal pseudocellar formula, but it can be distinguished from two latter species by the ventral pseudocellar formula on abdomen (0100 in
<taxonomicName id="80BF4D74FFF0FB4B24D9284BFC9FD6E9" authority="Park, 2017" authorityName="Park" authorityYear="2017" box="[724,822,368,392]" class="Entognatha" family="Onychiuridae" genus="Leeonychiurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="analis" status="sp. nov.">
<emphasis id="75CBEAE5FFF0FB4B24D9284BFC9FD6E9" box="[724,822,368,392]" italics="true" pageId="7" pageNumber="299">L. analis</emphasis>
</taxonomicName>
<emphasis id="75CBEAE5FFF0FB4B25302855FC3ED6E9" bold="true" box="[829,919,367,392]" pageId="7" pageNumber="299">
<taxonomicNameLabel id="6EF8579EFFF0FB4B25302855FC3ED6E9" box="[829,919,367,392]" pageId="7" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="75CBEAE5FFF0FB4B2593284BFC15D6E9" box="[926,956,369,392]" italics="true" pageId="7" pageNumber="299">vs.</emphasis>
0
<quantity id="80479B12FFF0FB4B25DF284AFBB1D6E9" box="[978,1048,367,392]" metricMagnitude="0" metricUnit="m" metricValue="2.5654" pageId="7" pageNumber="299" unit="in" value="101.0">101 in</quantity>
<taxonomicName id="80BF4D74FFF0FB4B2213284BFB07D6E9" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" box="[1054,1198,368,392]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="gulinensis">
<emphasis id="75CBEAE5FFF0FB4B2213284BFB07D6E9" box="[1054,1198,368,392]" italics="true" pageId="7" pageNumber="299">L. gulinensis</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="80BF4D74FFF0FB4B22DC284BFACAD6E9" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" box="[1233,1379,368,392]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="75CBEAE5FFF0FB4B22DC284BFACAD6E9" box="[1233,1379,368,392]" italics="true" pageId="7" pageNumber="299">L. antennalis</emphasis>
</taxonomicName>
) and the absence of chaetae c0 on upper anal valve (present in
<taxonomicName id="80BF4D74FFF0FB4B251628AFFC03D6CD" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" box="[795,938,404,428]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="gulinensis">
<emphasis id="75CBEAE5FFF0FB4B251628AFFC03D6CD" box="[795,938,404,428]" italics="true" pageId="7" pageNumber="299">L. gulinensis</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="80BF4D74FFF0FB4B25C328AFFBC9D6CD" baseAuthorityName="Sun &amp; Zhang" baseAuthorityYear="2012" box="[974,1120,404,428]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="antennalis">
<emphasis id="75CBEAE5FFF0FB4B25C328AFFBC9D6CD" box="[974,1120,404,428]" italics="true" pageId="7" pageNumber="299">L. antennalis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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