402 lines
51 KiB
XML
402 lines
51 KiB
XML
<document id="A7F97B68DD48E13E5DEEE19AF91CC744" ID-CLB-Dataset="4735" ID-DOI="10.1080/713833842" ID-GBIF-Dataset="dbf360e3-f8f0-446e-a50d-c6b9a838063b" ID-ISSN="1464-5262" ID-Zenodo-Dep="5298392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629991316392" checkinUser="felipe" docAuthor="Healy, Brenda & Fend, Steve" docDate="2002" docId="C82087E6FFAD2E59FE01FCD2FE59FBD0" docLanguage="en" docName="JNatHist.36.1.15-23.pdf" docOrigin="Journal of Natural History 36 (1)" docSource="http://www.tandfonline.com/doi/abs/10.1080/713833842" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Mesenchytraeus rhithralis Healy & Fend 2002, sp. n." docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="3419FF9EFFA92E5EFF86FFCBFFB4FFF0" masterDocTitle="The occurrence of Mesenchytraeus (Enchytraeidae: Oligochaeta) in ri ‚ e habitats of north-west American rivers, with description of a new species" masterLastPageNumber="23" masterPageNumber="15" pageNumber="17" updateTime="1699145532973" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A2EE9723D882E80F6AECBA2A181CC165">The occurrence of Mesenchytraeus (Enchytraeidae: Oligochaeta) in ri ‚ e habitats of north-west American rivers, with description of a new species</mods:title>
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<mods:namePart id="69C55063A6679EB6D1B6FA02F583DC32">Healy, Brenda</mods:namePart>
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<mods:namePart id="D238316AEA1E5896550132BC678950B0">Fend, Steve</mods:namePart>
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<mods:title id="528664CB131C631957B814608E88F466">Journal of Natural History</mods:title>
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<mods:part id="B2EF21C6DB823DA1BB73D25153FA82CB">
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<mods:date id="48CC4338D214404C102790B85FE0205A">2002</mods:date>
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<mods:number id="6C5C928F16E19BEC1BAD936A9EEF9043">2002-01-31</mods:number>
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<mods:number id="94E8EFD515D30D176113E8486D26A392">36</mods:number>
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<treatment id="C82087E6FFAD2E59FE01FCD2FE59FBD0" ID-DOI="http://doi.org/10.5281/zenodo.5305293" ID-GBIF-Taxon="185123814" ID-Zenodo-Dep="5305293" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0" httpUri="http://treatment.plazi.org/id/C82087E6FFAD2E59FE01FCD2FE59FBD0" lastPageId="7" lastPageNumber="20" pageId="4" pageNumber="17">
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<subSubSection id="0893657BFFAD2E5AFE01FCD2FDDBFCA1" pageId="4" pageNumber="17" type="nomenclature">
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<heading id="1B7E819CFFAD2E5AFE01FCD2FD5AFCC1" box="[391,750,793,817]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="17" reason="2">
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<taxonomicName id="87894D73FFAD2E5AFE01FCD2FD1CFCC1" ID-CoL="3ZZH7" authority="Healy & Fend, 2002" authorityName="Healy & Fend" authorityYear="2002" box="[391,680,793,817]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="17" phylum="Annelida" rank="species" species="rhithralis" status="sp. nov.">
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<emphasis id="72FDEAE2FFAD2E5AFE01FCD2FD1CFCC1" bold="true" box="[391,680,793,817]" italics="true" pageId="4" pageNumber="17">Mesenchytraeus rhithralis</emphasis>
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<taxonomicNameLabel id="69CE5799FFAD2E5AFD36FCD1FD5AFCC1" box="[688,750,794,817]" pageId="4" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="403636F0FFAD2E5AFD81FCF2FDDBFCA1" blockId="4.[519,623,825,850]" box="[519,623,825,850]" pageId="4" pageNumber="17">(®gure 1)</paragraph>
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<materialsCitation id="F0E13CADFFAD2E5AFF00FCA1FED9FC51" ID-GBIF-Occurrence="3339721302" collectingDate="1988-03-01" collectionCode="USNM" collectorName="S. Fend" pageId="4" pageNumber="17" specimenCode="USNM 186799" specimenCount="1" typeStatus="holotype">
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<typeStatus id="9F328852FFAD2E5AFF00FCA1FF4DFC71" box="[134,249,873,897]" pageId="4" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCode id="102F9E8BFFAD2E5AFE97FCA2FE70FC71" box="[273,452,873,897]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186799</specimenCode>
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, stained, whole mounted specimen, collected by
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<collectorName id="ED7C5326FFAD2E5AFFDBFC42FF02FC51" box="[93,182,905,929]" pageId="4" pageNumber="17">S. Fend</collectorName>
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,
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.
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:
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whole mount from each of the following localities:
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<collectingMunicipality id="A052AC8AFFAD2E5AFC46FC63FF28FC10" pageId="4" pageNumber="17">Yakima River</collectingMunicipality>
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at
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<collectorName id="ED7C5326FFAD2E5AFF48FC03FEA2FC10" box="[206,278,968,992]" pageId="4" pageNumber="17">Thorp</collectorName>
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<collectionCode id="2698AE35FFAD2E5AFEA2FC03FEE0FC10" box="[292,340,968,992]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFEE3FC03FDA2FC10" box="[357,534,968,992]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186800</specimenCode>
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;
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<materialsCitation id="F0E13CADFFAD2E5AFDAAFC03FF1FFBF0" ID-GBIF-Occurrence="3339721303" collectionCode="WA" location="Yakima River" municipality="Cle Elum" pageId="4" pageNumber="17" specimenCode="USNM 186801" specimenCount="1" typeStatus="paratype">
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at
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<collectingMunicipality id="A052AC8AFFAD2E5AFC85FC03FCC0FC10" box="[771,884,968,992]" pageId="4" pageNumber="17">Cle Elum</collectingMunicipality>
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<collectionCode id="2698AE35FFAD2E5AFC04FC03FC06FC10" box="[898,946,968,992]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFC42FC03FF1FFBF0" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186801</specimenCode>
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;
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<materialsCitation id="F0E13CADFFAD2E5AFF38FC23FDC9FBF0" ID-GBIF-Occurrence="3339721304" box="[190,637,1000,1024]" collectionCode="WA" location="Manastash Creek" pageId="4" pageNumber="17" specimenCode="USNM 186802" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFF38FC23FE3CFBF0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFF38FC23FE3CFBF0" box="[190,392,1000,1024]" name="Manastash Creek" pageId="4" pageNumber="17">Manastash Creek</location>
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<collectionCode id="2698AE35FFAD2E5AFE14FC23FE0BFBF0" box="[402,447,1000,1024]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFE56FC23FDC9FBF0" box="[464,637,1000,1024]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186802</specimenCode>
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</materialsCitation>
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;
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<materialsCitation id="F0E13CADFFAD2E5AFD09FC23FBA4FBF0" ID-GBIF-Occurrence="3339721309" box="[655,1040,1000,1024]" collectionCode="WA" location="Ruby Creek" pageId="4" pageNumber="17" specimenCode="USNM 186803" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFD09FC23FCAFFBF0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFD09FC23FCAFFBF0" box="[655,795,1000,1024]" name="Ruby Creek" pageId="4" pageNumber="17">Ruby Creek</location>
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<collectionCode id="2698AE35FFAD2E5AFCA3FC23FCE6FBF0" box="[805,850,1000,1024]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFCE5FC23FBA4FBF0" box="[867,1040,1000,1024]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186803</specimenCode>
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;
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<materialsCitation id="F0E13CADFFAD2E5AFFDBFBC3FE00FBD0" ID-GBIF-Occurrence="3339721310" box="[93,436,1032,1056]" collectionCode="ID" location="Cub River" pageId="4" pageNumber="17" specimenCode="USNM 186804" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFFDBFBC3FF60FBD0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFFDBFBC3FF60FBD0" box="[93,212,1032,1056]" name="Cub River" pageId="4" pageNumber="17">Cub River</location>
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<collectionCode id="2698AE35FFAD2E5AFF5AFBC2FF48FBD0" box="[220,252,1033,1056]" country="USA" lsid="urn:lsid:biocol.org:col:15674" name="University of Idaho" pageId="4" pageNumber="17" type="Herbarium">ID</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFE8CFBC3FE00FBD0" box="[266,436,1032,1056]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186804</specimenCode>
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;
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<materialsCitation id="F0E13CADFFAD2E5AFE42FBC3FC8CFBD0" ID-GBIF-Occurrence="3339721301" box="[452,824,1032,1056]" collectionCode="WY" location="Swift Creek" pageId="4" pageNumber="17" specimenCode="USNM 186805" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFE42FBC3FDFDFBD0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFE42FBC3FDFDFBD0" box="[452,585,1032,1056]" name="Swift Creek" pageId="4" pageNumber="17">Swift Creek</location>
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<collectionCode id="2698AE35FFAD2E5AFDD7FBC2FDCBFBD0" box="[593,639,1033,1056]" pageId="4" pageNumber="17">WY</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFD08FBC3FC8CFBD0" box="[654,824,1032,1056]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="17" type="Museum">USNM 186805</specimenCode>
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</materialsCitation>
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.
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<materialsCitation id="F0E13CADFFAD2E5AFCCEFBC3FCA0FBB0" ID-GBIF-Occurrence="3339721307" collectionCode="WA" collectorName="Thorp" location="Serial" municipality="Yakima River" pageId="4" pageNumber="17" specimenCode="ROMIZ 19980006" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFCCEFBC3FC3CFBD0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFCCEFBC3FC3CFBD0" box="[840,904,1032,1056]" municipality="Yakima River" name="Serial" pageId="4" pageNumber="17">Serial</location>
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longitudinal sections from
|
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<collectingMunicipality id="A052AC8AFFAD2E5AFF7AFBE3FE2FFBB0" box="[252,411,1064,1088]" pageId="4" pageNumber="17">Yakima River</collectingMunicipality>
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at
|
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<collectorName id="ED7C5326FFAD2E5AFE38FBE3FDB2FBB0" box="[446,518,1064,1088]" pageId="4" pageNumber="17">Thorp</collectorName>
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<collectionCode id="2698AE35FFAD2E5AFD8BFBE3FD89FBB0" box="[525,573,1064,1088]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
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,
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<specimenCode id="102F9E8BFFAD2E5AFDCEFBE3FCA0FBB0" box="[584,788,1064,1088]" collectionCode="ROMIZ" pageId="4" pageNumber="17">ROMIZ 19980006</specimenCode>
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</materialsCitation>
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,
|
||
<materialsCitation id="F0E13CADFFAD2E5AFCA0FBE2FE99FB90" ID-GBIF-Occurrence="3339721305" collectionCode="WA" location="Teanaway River" pageId="4" pageNumber="17" specimenCode="ROMIZ 19980004" specimenCount="1" typeStatus="paratype">
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<location id="4556602BFFAD2E5AFCA0FBE2FC6AFBB0" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFCA0FBE2FC6AFBB0" box="[806,990,1064,1089]" name="Teanaway River" pageId="4" pageNumber="17">Teanaway River</location>
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||
<collectionCode id="2698AE35FFAD2E5AFC62FBE3FBA5FBB0" box="[996,1041,1064,1088]" country="Poland" lsid="urn:lsid:biocol.org:col:14722" name="University of Warsaw" pageId="4" pageNumber="17" type="Herbarium">WA</collectionCode>
|
||
,
|
||
<specimenCode id="102F9E8BFFAD2E5AFFDBFB83FE99FB90" box="[93,301,1096,1120]" collectionCode="ROMIZ" pageId="4" pageNumber="17">ROMIZ 19980004</specimenCode>
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||
</materialsCitation>
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,
|
||
<materialsCitation id="F0E13CADFFAD2E5AFEC6FB83FD69FB90" ID-GBIF-Occurrence="3339721306" box="[320,733,1096,1120]" collectionCode="WY" location="Swift Creek" pageId="4" pageNumber="17" specimenCode="ROMIZ 19980009" specimenCount="1" typeStatus="paratype">
|
||
<location id="4556602BFFAD2E5AFEC6FB83FE72FB90" LSID="urn:lsid:plazi:treatment:C82087E6FFAD2E59FE01FCD2FE59FBD0:4556602BFFAD2E5AFEC6FB83FE72FB90" box="[320,454,1096,1120]" name="Swift Creek" pageId="4" pageNumber="17">Swift Creek</location>
|
||
<collectionCode id="2698AE35FFAD2E5AFE49FB83FE49FBAF" box="[463,509,1096,1119]" pageId="4" pageNumber="17">WY</collectionCode>
|
||
,
|
||
<specimenCode id="102F9E8BFFAD2E5AFD88FB83FD69FB90" box="[526,733,1096,1120]" collectionCode="ROMIZ" pageId="4" pageNumber="17">ROMIZ 19980009</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAD2E5AFF00FBA2FD27FB6F" blockId="4.[93,1049,873,1184]" pageId="4" pageNumber="17">
|
||
<emphasis id="72FDEAE2FFAD2E5AFF00FBA2FEAAFB70" box="[134,286,1128,1152]" italics="true" pageId="4" pageNumber="17">Type locality.</emphasis>
|
||
Manastash Creek, Kittitas Co.,
|
||
<collectingRegion id="824DF812FFAD2E5AFD20FBA3FC98FB70" box="[678,812,1128,1152]" country="United States of America" name="Washington" pageId="4" pageNumber="17">Washington</collectingRegion>
|
||
, part of the Yakima River drainage. Cobble ri‚e with gravel patches.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAD2E5AFFDBFB03FEC1FB10" blockId="4.[93,1049,1224,1662]" box="[93,373,1224,1248]" pageId="4" pageNumber="17">
|
||
<emphasis id="72FDEAE2FFAD2E5AFFDBFB03FEC1FB10" box="[93,373,1224,1248]" italics="true" pageId="4" pageNumber="17">Other material examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAD2E5AFF00FB23FCFDFACF" blockId="4.[93,1049,1224,1662]" pageId="4" pageNumber="17">
|
||
Whole-mounted specimens (nine slides),
|
||
<collectionCode id="2698AE35FFAD2E5AFDD7FB23FD04FAF0" box="[593,688,1256,1280]" pageId="4" pageNumber="17">ROMIZ</collectionCode>
|
||
19980004, 19980005, 19980007, 19980009. About 50 whole mounts from the 19 localities listed above in the collection of B. H., spirit material and many whole mounts in that of S. F.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAD2E5AFF00FA8CFCF8FAAF" blockId="4.[93,1049,1224,1662]" box="[134,844,1351,1375]" pageId="4" pageNumber="17">
|
||
<emphasis id="72FDEAE2FFAD2E5AFF00FA8CFEB1FAAF" box="[134,261,1351,1375]" italics="true" pageId="4" pageNumber="17">Etymology.</emphasis>
|
||
From the Greek
|
||
<emphasis id="72FDEAE2FFAD2E5AFE65FA8CFDFAFAAF" box="[483,590,1351,1375]" italics="true" pageId="4" pageNumber="17">`rheithron</emphasis>
|
||
’, a current or stream.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAD2E5BFF00FAA3FCDCF9CE" blockId="4.[93,1049,1224,1662]" lastBlockId="5.[93,1049,1000,1662]" lastPageId="5" lastPageNumber="18" pageId="4" pageNumber="17">
|
||
<emphasis id="72FDEAE2FFAD2E5AFF00FAA3FEBFFA8F" box="[134,267,1384,1407]" italics="true" pageId="4" pageNumber="17">Description.</emphasis>
|
||
Somewhat fragile worms, nearly transparent when alive, length of preserved worms
|
||
<quantity id="87719B15FFAD2E5AFEABFA4CFE32FA6F" box="[301,390,1415,1439]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="4" pageNumber="17" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6±9 mm</quantity>
|
||
, width 0.3±0.4 mm in
|
||
<collectionCode id="2698AE35FFAD2E5AFD12FA43FD05FA6F" box="[660,689,1416,1439]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="17">VI</collectionCode>
|
||
, 0.38±0.5 mm at the clitellum, narrowing in terminal segments. Segments 29±41 (exceptionally 45) (mean 35.1,
|
||
<geoCoordinate id="25BD5037FFAD2E5AFC77FA63FFC3FA2E" degrees="5" direction="north" minutes="39" orientation="latitude" pageId="4" pageNumber="17" precision="925" value="5.65">
|
||
<emphasis id="72FDEAE2FFAD2E5AFC77FA63FC4AFA4F" box="[1009,1022,1448,1471]" italics="true" pageId="4" pageNumber="17">n</emphasis>
|
||
5 39
|
||
</geoCoordinate>
|
||
). Epidermal nuclei scattered, not in transverse rows. In most specimens, epidermal gland cells conspicuous throughout the body, regularly scattered, situated on small papillae, numerous on the prostomium; glands and papillae absent in specimens from localities 6, 10 and 19. Prostomium an elongate cone (®gure 1A), head pore at or near the tip. Clitellum over XII±XIII with small, irregularly scattered gland cells, absent mid ventrally, up to 30 Mm thick dorsally. Chaetae sigmoid, sharply pointed, with weak nodulus (®gure 1B), in fan-like bundles, four to eight, rarely 10, in ventral bundles, absent ventrally in XII, three to six laterally; length 90±140 Mm (70 Mm in specimens from Hat Creek), more or less equal within a bundle. Foregut widening gradually at 6/7, wall distinctly thicker in VII±VIII, occasionally also in IX, due to taller cells of the gut lining. Gut contents predominantly diatoms. Choragocytes forming a distinct layer from
|
||
<collectionCode id="2698AE35FFAC2E5BFDEBFB43FD30FB6F" box="[621,644,1160,1183]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="18" type="Museum">V</collectionCode>
|
||
. Post-pharyngeal bulbs small, not observed in many specimens, other gut diverticula absent. Three pairs of septal glands at 3/4, 4/5 and 5/6, septum at 3/4 often poorly developed, reduced to dorsal and ventral strands of tissue. Septal glands consisting of small dorsal lobes, second and third pairs occasionally united across oesophagus, ventral lobes usually present in
|
||
<collectionCode id="2698AE35FFAC2E5BFFF8FAE3FF25FACF" box="[126,145,1320,1343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="18" type="Museum">V</collectionCode>
|
||
and
|
||
<collectionCode id="2698AE35FFAC2E5BFF56FAE3FF59FACF" box="[208,237,1320,1343]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="18">VI</collectionCode>
|
||
, secondary glands present in mid
|
||
<collectionCode id="2698AE35FFAC2E5BFD07FAE3FD20FACF" box="[641,660,1320,1343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="18" type="Museum">V</collectionCode>
|
||
and
|
||
<collectionCode id="2698AE35FFAC2E5BFD55FAE3FD44FACF" box="[723,752,1320,1343]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="18">VI</collectionCode>
|
||
, sometimes reduced in
|
||
<collectionCode id="2698AE35FFAC2E5BFB80FAE3FBADFACF" box="[1030,1049,1320,1343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="18" type="Museum">V</collectionCode>
|
||
(®gure 1A). Anterior border of brain deeply cleft, posterior border sinuous, length about 100 Mm, maximum width 70±90 Mm (®gure 1A). Dorsal vessel originating in XII or XIII, blood colourless. Coelomocytes oval, 10±28 Mm long, with several small vacuoles (®gure 1C). Nephridia of the
|
||
<taxonomicName id="87894D73FFAC2E5BFDB4FA6CFD57FA4F" authorityName="Eisen" authorityYear="1878" box="[562,739,1447,1471]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="18" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAC2E5BFDB4FA6CFD57FA4F" box="[562,739,1447,1471]" italics="true" pageId="5" pageNumber="18">Mesenchytraeus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="9F328852FFAC2E5BFD77FA63FC95FA30" box="[753,801,1448,1472]" pageId="5" pageNumber="18">type</typeStatus>
|
||
, i.e. lobed and with tubules clearly visible due to poor development of interstitial tissue; three preclitellar pairs at 6/7±8/9, occasionally at 7/8±9/10, anterior pair sometimes absent. Preseptal consisting of nephrostome on a short neck, postseptale with two lobes, eOEerent duct long and partly folded, arising anteriorly between lobes (®gure 1D).
|
||
</paragraph>
|
||
<caption id="14F66678FFAC2E5BFFDBFCC4FE1FFC59" ID-DOI="http://doi.org/10.5281/zenodo.5298394" ID-Zenodo-Dep="5298394" httpUri="https://zenodo.org/record/5298394/files/figure.png" pageId="5" pageNumber="18" startId="5.[93,109,783,804]" targetBox="[94,1049,106,755]" targetPageId="5">
|
||
<paragraph id="403636F0FFAC2E5BFFDBFCC4FE1FFC59" blockId="5.[93,1049,782,937]" pageId="5" pageNumber="18">
|
||
FIG. 1.
|
||
<taxonomicName id="87894D73FFAC2E5BFF3FFCC5FE09FCD4" authority="Healy & Fend, 2002" authorityName="Healy & Fend" authorityYear="2002" box="[185,445,782,804]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="18" phylum="Annelida" rank="species" species="rhithralis" status="sp. nov.">
|
||
<emphasis id="72FDEAE2FFAC2E5BFF3FFCC5FE09FCD4" box="[185,445,782,804]" italics="true" pageId="5" pageNumber="18">Mesenchytraeus rhithralis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69CE5799FFAC2E5BFE40FCC4FDB5FCD4" box="[454,513,783,804]" pageId="5" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
|
||
: (A) dorsal view of anterior segments; (B) chaeta; (C) coelomocytes; (D) preclitellar nephridium; (E) sperm funnel; (F) part of vas deferens; (G, H) two views of the penial bulb. am, spermathecal ampulla; br, brain; co, collar; ed, spermathecal ectal duct; ef, eOEerent duct; ne, nephrostome; pb, penial bulb; sg, septal glands; vd, vas deferens. Scale bars: 0.5 mm (A), 50 Mm (B, D, E, G, H) and 20 Mm (C, F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="403636F0FFAC2E58FF00F98CFC6BFCF1" blockId="5.[93,1049,1000,1662]" lastBlockId="6.[93,1049,107,1662]" lastPageId="6" lastPageNumber="19" pageId="5" pageNumber="18">
|
||
Testes compact, rarely lobed. Seminal vesicle unpaired in XI, extending forward as one or two lobes into X and IX, and back into XII. In a few specimens two lobes extended posteriorly in egg sacs as far as XIV. Sperm funnels about twice as long as wide when extended, but usually bent, of variable size, 120±280
|
||
<emphasis id="72FDEAE2FFAF2E58FC0AFF40FC11FF53" bold="true" box="[908,933,139,163]" italics="true" pageId="6" pageNumber="19">Ö</emphasis>
|
||
40±70 Mm, tapering distally. Collar variable, either narrow and equal in diameter to funnel, ¯ared, or apparently absent (®gure 1E). Vasa deferentia stout, thick-walled (®gure 1F), diameter 24±50 Mm, of more or less equal diameter throughout but slightly narrower near funnel and penial bulb, entering penial bulbs centrally; length about equal to diameter of clitellum. Refringent particles sometimes present in vas wall. Bulbs sac-like, compact when retracted, nearly spherical, 80±140 Mm across, consisting of densely staining glandular tissue surrounding a central chamber (®gure 1G, H) which opens to outside by way of an irregular slit. Everted bulbs irregular, appearing thin-walled. Vas enters bulb through a thick pad lined by a glandular epithelium with goblet cells (seen in some sections). Atrial enlargement of vas, atrial glands, and accessory glands attached to bulb absent. Paired egg sacs extending to XIV, occasionally to XV or XVI when fully distended with eggs and ovarian tissue. Spermathecae free, without diverticula, opening ventro-laterally in intersegmental groove 4/5. Ectal ducts stout, thick-walled, 240±360 Mm long, 18± 22 Mm in diameter, leading to large, cylindrical ampullae which are thick-walled when empty but become thin-walled when distended with sperm (®gure 1A). Spermathecae extend ventro-laterall y through several segments, sometimes reaching to X or even XI where they may overlap seminal vesicle. In a few specimens from Ruby Creek, a third spermatheca was present, opening ventro-laterall y at 5/6.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAF2E58FF00FCC2FE48FB0F" blockId="6.[93,1049,107,1662]" pageId="6" pageNumber="19">
|
||
<emphasis id="72FDEAE2FFAF2E58FF00FCC2FF44FCD1" box="[134,240,777,801]" italics="true" pageId="6" pageNumber="19">Remarks.</emphasis>
|
||
The species shares the following characters with other members of
|
||
<taxonomicName id="87894D73FFAF2E58FFDBFCE2FEBAFCB1" authorityName="Eisen" authorityYear="1878" box="[93,270,809,833]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAF2E58FFDBFCE2FEBAFCB1" box="[93,270,809,833]" italics="true" pageId="6" pageNumber="19">Mesenchytraeus</emphasis>
|
||
</taxonomicName>
|
||
: nodulated, sigmoid setae and lobed nephridia with little interstitial tissue.
|
||
<taxonomicName id="87894D73FFAF2E58FF29FC82FD7CFC91" authority="Nielsen and Christensen 1959" authorityName="Nielsen and Christensen" authorityYear="1959" box="[175,712,841,865]" class="Clitellata" family="Enchytraeidae" genus="Cernosvitoviella" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAF2E58FF29FC82FEEAFC91" box="[175,350,841,865]" italics="true" pageId="6" pageNumber="19">Cernosvitoviella</emphasis>
|
||
<bibRefCitation id="24184B01FFAF2E58FEEAFC82FD7CFC91" author="NIELSEN, C. O. & CHRISTENSEN, B." box="[364,712,841,865]" pageId="6" pageNumber="19" pagination="160" refId="ref5374" refString="NIELSEN, C. O. and CHRISTENSEN, B., 1959, The Enchytraeidae, critical revision and taxonomy of European species, Natura jutlandica, 8 ± 9, 1 ± 160." type="journal article" year="1959">Nielsen and Christensen 1959</bibRefCitation>
|
||
</taxonomicName>
|
||
, which also have nodulated setae, have more compact, unlobed nephridia and a muscular penial bulb is absent.
|
||
<taxonomicName id="87894D73FFAF2E58FFDBFC42FE34FC51" authorityName="Healy & Fend" authorityYear="2002" box="[93,384,905,929]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="rhithralis">
|
||
<emphasis id="72FDEAE2FFAF2E58FFDBFC42FE34FC51" box="[93,384,905,929]" italics="true" pageId="6" pageNumber="19">Mesenchytraeus rhithralis</emphasis>
|
||
</taxonomicName>
|
||
is recognized by its relatively small size for the genus, primary septal glands at 3/4, 4/5 and 5/6 and secondaries in
|
||
<collectionCode id="2698AE35FFAF2E58FCA7FC62FC80FC30" box="[801,820,937,960]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="19" type="Museum">V</collectionCode>
|
||
(usually) and
|
||
<collectionCode id="2698AE35FFAF2E58FC5BFC62FC4EFC30" box="[989,1018,937,960]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="19">VI</collectionCode>
|
||
, a thickening of the gut wall in VII and VIII, intraclitellar origin of the dorsal vessel, a short, stout vas deferens without atrial enlargement or atrial glands, compact penial bulbs without accessory glands, and long, free spermathecae without diverticula. It is recognized at low magni®cation in benthic samples by its long-conical prostomium, many ®ne chaetae per bundle, and generally thin integument that leaves it looking rather wilted compared with other enchytraeids. Dimensions of all parts of the male reproductive system are very variable. The testes can be compact or slightly lobed, the seminal vesicle can be con®ned to two segments or extend from 9/10 to 14/15, whilst size of the sperm funnels, and diameters of vas deferentia and penial bulbs, vary over wide ranges.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAF2E59FF00FACCFCAFFE72" blockId="6.[93,1049,107,1662]" lastBlockId="7.[93,1049,107,1057]" lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19">
|
||
Among 25 or so described species of
|
||
<taxonomicName id="87894D73FFAF2E58FDB6FACCFD55FAEF" authorityName="Eisen" authorityYear="1878" box="[560,737,1287,1311]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAF2E58FDB6FACCFD55FAEF" box="[560,737,1287,1311]" italics="true" pageId="6" pageNumber="19">Mesenchytraeus</emphasis>
|
||
</taxonomicName>
|
||
in which the spermathecae are free, i.e. not communicating with the oesophagus, only seven lack diverticula (external or internal) at the junction between the ectal duct and the ampulla.
|
||
<taxonomicName id="87894D73FFAF2E58FFDBFAA3FE34FA8F" authority="Altman, 1936" authorityName="Altman" authorityYear="1936" box="[93,384,1383,1407]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="magnus">
|
||
<emphasis id="72FDEAE2FFAF2E58FFDBFAA3FFC3FA8F" box="[93,119,1384,1407]" italics="true" pageId="6" pageNumber="19">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAF2E58FF0EFAA3FF6EFA8F" box="[136,218,1384,1407]" italics="true" pageId="6" pageNumber="19">magnus</emphasis>
|
||
<bibRefCitation id="24184B01FFAF2E58FF63FAACFE34FA8F" author="ALTMAN, L. C." box="[229,384,1383,1407]" pageId="6" pageNumber="19" pagination="137" refId="ref4665" refString="ALTMAN, L. C., 1936, Oligochaeta of Washington, University of Washington Publications in Biology, 4, 1 ± 137." type="journal article" year="1936">Altman, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
, recorded from Seattle, Washington, reaches over
|
||
<quantity id="87719B15FFAF2E58FC48FAACFBACFA8F" box="[974,1048,1383,1407]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="19" unit="mm" value="70.0">70 mm</quantity>
|
||
and has short spermathecae;
|
||
<taxonomicName id="87894D73FFAF2E58FE3BFA43FD08FA6F" authority="Welch, 1917" authorityName="Welch" authorityYear="1917" box="[445,700,1415,1439]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="altus">
|
||
<emphasis id="72FDEAE2FFAF2E58FE3BFA43FD94FA6F" box="[445,544,1415,1439]" italics="true" pageId="6" pageNumber="19">M. altus</emphasis>
|
||
<bibRefCitation id="24184B01FFAF2E58FDABFA4CFD08FA6F" author="WELCH, P. S." box="[557,700,1415,1439]" pageId="6" pageNumber="19" pagination="71" refId="ref5587" refString="WELCH, P. S., 1917, Enchytraeidae from the Rocky Mountain Region. Transactions of the American Microscopical Society, 36, 67 ± 71." type="journal article" year="1917">Welch, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Rocky Mountains, Colorado has atrial and penial glands, and a long vas deferens;
|
||
<taxonomicName id="87894D73FFAF2E58FCA9FA63FF26FA2E" authority="Eisen, 1878" authorityName="Eisen" authorityYear="1878" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="falciformis">
|
||
<emphasis id="72FDEAE2FFAF2E58FCA9FA63FCFDFA4F" box="[815,841,1448,1471]" italics="true" pageId="6" pageNumber="19">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAF2E58FCD0FA6CFC7FFA4F" box="[854,971,1447,1471]" italics="true" pageId="6" pageNumber="19">falciformis</emphasis>
|
||
<bibRefCitation id="24184B01FFAF2E58FC52FA6CFF26FA2E" author="EISEN, G." pageId="6" pageNumber="19" pagination="79" refId="ref5041" refString="EISEN, G., 1878, RedogoErelse foEr Oligochaeta, samlade under Svenska expeditionerna till Arktiska trakter, O Efversigt af Kongliga Vetenskaps-Akademiens FoErhandlingar, 3, 63 ± 79." type="journal article" year="1878">Eisen, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Russian Arctic and
|
||
<collectingCountry id="389E7660FFAF2E58FE73FA0DFDD8FA2E" box="[501,620,1478,1502]" name="Greenland" pageId="6" pageNumber="19">Greenland</collectingCountry>
|
||
has pigmented coelomocytes, a very small sperm funnel and short spermathecae;
|
||
<taxonomicName id="87894D73FFAF2E58FDD1FA2CFCCAFA0E" authority="Nurminen, 1977" authorityName="Nurminen" authorityYear="1977" box="[599,894,1510,1534]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="franzi">
|
||
<emphasis id="72FDEAE2FFAF2E58FDD1FA2CFDC4FA0E" box="[599,624,1511,1534]" italics="true" pageId="6" pageNumber="19">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAF2E58FDF9FA2DFD0AFA0E" box="[639,702,1510,1534]" italics="true" pageId="6" pageNumber="19">franzi</emphasis>
|
||
<bibRefCitation id="24184B01FFAF2E58FD40FA2DFCCAFA0E" author="NURMINEN, M." box="[710,894,1510,1534]" pageId="6" pageNumber="19" pagination="227" refId="ref5410" refString="NURMINEN, M., 1977, Enchytraeidae (Oligochaeta) from the Grossglockner region of the Austrian Alps, Annales zoologici fennici, 14, 224 ± 227." type="journal article" year="1977">Nurminen, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="389E7660FFAF2E58FC42FA2DFBADFA0E" box="[964,1049,1510,1534]" name="Austria" pageId="6" pageNumber="19">Austria</collectingCountry>
|
||
has only two chaetae per bundle, and spermathecae with ectal glands and long ectal ducts;
|
||
<taxonomicName id="87894D73FFAF2E58FF29F9ECFCB6F9CE" authority="Piper, McLean and Christensen, 1982" authorityName="Piper, McLean and Christensen" authorityYear="1982" box="[175,770,1574,1598]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="tundrus">
|
||
<emphasis id="72FDEAE2FFAF2E58FF29F9ECFF7CF9CE" box="[175,200,1575,1598]" italics="true" pageId="6" pageNumber="19">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAF2E58FF5BF9EDFE9AF9CE" box="[221,302,1574,1598]" italics="true" pageId="6" pageNumber="19">tundrus</emphasis>
|
||
Piper, McLean and Christensen, 1982
|
||
</taxonomicName>
|
||
from N.E.
|
||
<collectingCountry id="389E7660FFAF2E58FC1FF9EDFC51F9CE" box="[921,997,1574,1598]" name="Russia" pageId="6" pageNumber="19">Russia</collectingCountry>
|
||
has yellow-brown coelomocytes, pink blood and short spermathecae;
|
||
<taxonomicName id="87894D73FFAF2E58FCDCF98CFD94F98E" authority="Piper, McLean and Christensen, 1982" authorityName="Piper, McLean and Christensen" authorityYear="1982" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="19" phylum="Annelida" rank="species" species="diverticulatus">
|
||
<emphasis id="72FDEAE2FFAF2E58FCDCF98CFBADF9AE" box="[858,1049,1606,1630]" italics="true" pageId="6" pageNumber="19">M. diverticulatus</emphasis>
|
||
Piper, McLean and Christensen, 1982
|
||
</taxonomicName>
|
||
from N.E.
|
||
<collectingCountry id="389E7660FFAF2E58FD33F9ADFCB5F98E" box="[693,769,1638,1662]" name="Russia" pageId="6" pageNumber="19">Russia</collectingCountry>
|
||
has a gut enlargement with diverticula in VIII, and short spermathecae; and
|
||
<taxonomicName id="87894D73FFAE2E59FD67FFA7FE93FF53" authority="Nielsen and Christensen, 1959" authorityName="Nielsen and Christensen" authorityYear="1959" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="sanguineus">
|
||
<emphasis id="72FDEAE2FFAE2E59FD67FFA7FD4EFF73" box="[737,762,108,131]" italics="true" pageId="7" pageNumber="20">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAE2E59FC8AFFA7FC35FF73" box="[780,897,108,131]" italics="true" pageId="7" pageNumber="20">sanguineus</emphasis>
|
||
<bibRefCitation id="24184B01FFAE2E59FC09FFA0FE93FF53" author="NIELSEN, C. O. & CHRISTENSEN, B." pageId="7" pageNumber="20" pagination="160" refId="ref5374" refString="NIELSEN, C. O. and CHRISTENSEN, B., 1959, The Enchytraeidae, critical revision and taxonomy of European species, Natura jutlandica, 8 ± 9, 1 ± 160." type="journal article" year="1959">Nielsen and Christensen, 1959</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="389E7660FFAE2E59FEE9FF40FE6EFF53" box="[367,474,139,163]" name="Denmark" pageId="7" pageNumber="20">Denmark</collectingCountry>
|
||
, Britain and
|
||
<collectingCountry id="389E7660FFAE2E59FDF2FF40FD72FF53" box="[628,710,139,163]" name="Ireland" pageId="7" pageNumber="20">Ireland</collectingCountry>
|
||
has red blood, sperm funnels with tubular collars, and penial bulbs with long accessory glands. These species diOEer widely from each other and none show particular a nities with
|
||
<taxonomicName id="87894D73FFAE2E59FC02FF07FBBBFF13" authorityName="Healy & Fend" authorityYear="2002" box="[900,1039,203,227]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="rhithralis">
|
||
<emphasis id="72FDEAE2FFAE2E59FC02FF07FC29FF13" box="[900,925,204,227]" italics="true" pageId="7" pageNumber="20">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAE2E59FC2BFF00FBBBFF13" box="[941,1039,203,227]" italics="true" pageId="7" pageNumber="20">rhithralis</emphasis>
|
||
</taxonomicName>
|
||
. For nearly all of them, and for most species described prior to 1959, descriptions are incomplete, so it cannot be stated with certainty whether any features of
|
||
<taxonomicName id="87894D73FFAE2E59FFDBFEE7FF58FEB3" authorityName="Healy & Fend" authorityYear="2002" box="[93,236,299,323]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="rhithralis">
|
||
<emphasis id="72FDEAE2FFAE2E59FFDBFEE7FFC3FEB3" box="[93,119,300,323]" italics="true" pageId="7" pageNumber="20">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAE2E59FF0CFEE0FF58FEB3" box="[138,236,299,323]" italics="true" pageId="7" pageNumber="20">rhithralis</emphasis>
|
||
</taxonomicName>
|
||
, or combinations of characters, are unique. In several species listed above, the origin of the dorsal vessel is post-clitellar, but an intra-clitellar origin has been reported for a number of other
|
||
<taxonomicName id="87894D73FFAE2E59FD8DFEA1FCAFFE72" box="[523,795,362,386]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="72FDEAE2FFAE2E59FD8DFEA1FD08FE72" box="[523,700,362,386]" italics="true" pageId="7" pageNumber="20">Mesenchytraeus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAE2E59FF00FE41FC1CFD31" blockId="7.[93,1049,107,1057]" pageId="7" pageNumber="20">
|
||
The presence of septal glands at 3/4, known only for
|
||
<taxonomicName id="87894D73FFAE2E59FD69FE40FBACFE52" authority="Healy and Timm" authorityName="Healy and Timm" box="[751,1048,394,418]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="kuril">
|
||
<emphasis id="72FDEAE2FFAE2E59FD69FE40FCBCFE52" box="[751,776,395,418]" italics="true" pageId="7" pageNumber="20">M</emphasis>
|
||
.
|
||
<emphasis id="72FDEAE2FFAE2E59FC9EFE41FCFFFE52" box="[792,843,394,418]" italics="true" pageId="7" pageNumber="20">kuril</emphasis>
|
||
Healy and Timm
|
||
</taxonomicName>
|
||
(in press), is unique for enchytraeids and deserves special comment. It is generally accepted that there are no septa anterior to 4/
|
||
<quantity id="87719B15FFAE2E59FDEFFE01FD20FE12" box="[617,660,458,482]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="20" unit="in" value="5.0">5 in</quantity>
|
||
enchytraeids, although nephridia, normally associated with septa, have been reported at 2/
|
||
<quantity id="87719B15FFAE2E59FD5EFE21FCB6FDF2" box="[728,770,490,514]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="20" unit="in" value="3.0">3 in</quantity>
|
||
two species of
|
||
<taxonomicName id="87894D73FFAE2E59FC2AFE21FBA3FDF2" box="[940,1047,490,514]" class="Clitellata" family="Enchytraeidae" genus="Cognettia" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAE2E59FC2AFE21FBA3FDF2" box="[940,1047,490,514]" italics="true" pageId="7" pageNumber="20">Cognettia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="24184B01FFAE2E59FFEEFDC0FF44FDD2" author="BAUER, R." box="[104,240,522,546]" pageId="7" pageNumber="20" pagination="689" refId="ref4691" refString="BAUER, R., 1993, Cognettia clarae n. sp. Deine neue Enchytraeiden Art aus einem O E sterreichischen Fichtenwald (Oligochaeta, Enchytraeidae), Linzer Biologische Beitrage, 25, 685 ± 689." type="journal article" year="1993">Bauer, 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24184B01FFAE2E59FE87FDC1FD11FDD2" author="CHRISTENSEN, B. & DOAZSA-FARKAS, K." box="[257,677,522,547]" pageId="7" pageNumber="20" pagination="37" refId="ref5007" refString="CHRISTENSEN, B. and DOAZSA-FARKAS, K., 1999, The enchytraeid fauna of the Palaearctic tundra (Oligochaeta, Enchytraeidae), Biologiske Skrifter, 52, 1 ± 37." type="journal article" year="1999">Christensen and DoÂzsa-Farkas, 1999</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="87894D73FFAE2E59FD65FDC0FCC5FDD2" authorityName="Healy & Fend" authorityYear="2002" box="[739,881,522,546]" class="Clitellata" family="Enchytraeidae" genus="Mesenchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="rhithralis">
|
||
<emphasis id="72FDEAE2FFAE2E59FD65FDC0FCC5FDD2" box="[739,881,522,546]" italics="true" pageId="7" pageNumber="20">M. rhithralis</emphasis>
|
||
</taxonomicName>
|
||
the septa may be incomplete but are distinctly present, seen for example in longitudinal sections (e.g.
|
||
<collectionCode id="2698AE35FFAE2E59FF10FD81FF41FD92" box="[150,245,586,610]" pageId="7" pageNumber="20">ROMIZ</collectionCode>
|
||
19980004B). The glands are small but appear identical in structure to those at 4/5 and 5/6, and ducts leading from them to the pharynx are clearly distinguishable. The irregular pattern of epidermal cells as shown by nuclei not arranged in transverse rows as in most enchytraeids is also worth noting.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAE2E59FF00FD01FDFFFC30" blockId="7.[93,1049,107,1057]" pageId="7" pageNumber="20">
|
||
<emphasis id="72FDEAE2FFAE2E59FF00FD01FF56FD12" box="[134,226,714,738]" italics="true" pageId="7" pageNumber="20">Habitat.</emphasis>
|
||
In low to sixth order rhithral stream segments with coarse substratum. Highest densities in lotic, erosional habitats, becoming scarce in marginal, depositional habitats where they are generally accompanied by other oligochaetes including enchytraeids (species of
|
||
<taxonomicName id="87894D73FFAE2E59FE04FCE2FE59FCB1" box="[386,493,809,833]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="20" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="72FDEAE2FFAE2E59FE04FCE2FE59FCB1" box="[386,493,809,833]" italics="true" pageId="7" pageNumber="20">Fridericia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="87894D73FFAE2E59FDB2FCE2FD57FCB1" authorityName="Nielsen and Christensen" authorityYear="1959" box="[564,739,809,833]" class="Clitellata" family="Enchytraeidae" genus="Cernosvitoviella" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="20" phylum="Annelida" rank="genus">
|
||
<emphasis id="72FDEAE2FFAE2E59FDB2FCE2FD57FCB1" box="[564,739,809,833]" italics="true" pageId="7" pageNumber="20">Cernosvitoviella</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="87894D73FFAE2E59FC85FCE1FCC1FCB1" box="[771,885,810,833]" class="Sordariomycetes" family="Halosphaeriaceae" genus="Nais" kingdom="Fungi" order="Microascales" pageId="7" pageNumber="20" phylum="Ascomycota" rank="species" species="undetermined">
|
||
<emphasis id="72FDEAE2FFAE2E59FC85FCE1FC82FCB1" box="[771,822,810,833]" italics="true" pageId="7" pageNumber="20">Nais</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="87894D73FFAE2E59FC0EFCE2FF3AFC91" class="Clitellata" family="Naididae" genus="Rhyacodrilus" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="72FDEAE2FFAE2E59FC0EFCE2FBACFCB1" box="[904,1048,809,833]" italics="true" pageId="7" pageNumber="20">Rhyacodrilus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="87894D73FFAE2E59FF1BFC82FDAEFC91" authority="ClapareAde" authorityName="ClapareAde" box="[157,538,840,865]" class="Clitellata" family="Lumbriculidae" genus="Stylodrilus" kingdom="Animalia" order="Lumbriculida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="heringianus">
|
||
<emphasis id="72FDEAE2FFAE2E59FF1BFC82FE29FC91" box="[157,413,841,865]" italics="true" pageId="7" pageNumber="20">Stylodrilus heringianus</emphasis>
|
||
ClapareÁde
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="87894D73FFAE2E59FDDFFC82FCA9FC91" box="[601,797,841,865]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="7" pageNumber="20" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="72FDEAE2FFAE2E59FDDFFC82FD50FC91" box="[601,740,841,865]" italics="true" pageId="7" pageNumber="20">Rhynchelmis</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
In the Yakima River system (80 sites), the species was present in almost all of the upstream sites draining a montane, forested landscape, and absent from almost all potamal downstream sites within a valley agricultural landscape.
|
||
</paragraph>
|
||
<paragraph id="403636F0FFAE2E59FF00FC03FE59FBD0" blockId="7.[93,1049,107,1057]" pageId="7" pageNumber="20">
|
||
<emphasis id="72FDEAE2FFAE2E59FF00FC03FEA5FC10" box="[134,273,968,992]" italics="true" pageId="7" pageNumber="20">Distribution.</emphasis>
|
||
Western
|
||
<collectingCountry id="389E7660FFAE2E59FE16FC03FE7DFC10" box="[400,457,968,992]" name="United States of America" pageId="7" pageNumber="20">USA</collectingCountry>
|
||
(
|
||
<collectingRegion id="824DF812FFAE2E59FE5BFC03FDD7FC10" box="[477,611,968,992]" country="United States of America" name="Washington" pageId="7" pageNumber="20">Washington</collectingRegion>
|
||
,
|
||
<collectingRegion id="824DF812FFAE2E59FDF2FC03FD54FC10" box="[628,736,968,992]" country="United States of America" name="Wyoming" pageId="7" pageNumber="20">Wyoming</collectingRegion>
|
||
,
|
||
<collectingRegion id="824DF812FFAE2E59FD77FC03FCF1FC10" box="[753,837,968,992]" country="United States of America" name="Oregon" pageId="7" pageNumber="20">Oregon</collectingRegion>
|
||
,
|
||
<collectingRegion id="824DF812FFAE2E59FCD3FC03FC2CFC10" box="[853,920,968,992]" country="United States of America" name="Idaho" pageId="7" pageNumber="20">Idaho</collectingRegion>
|
||
,
|
||
<collectingRegion id="824DF812FFAE2E59FC2EFC02FBA4FC10" box="[936,1040,969,992]" country="United States of America" name="Montana" pageId="7" pageNumber="20">Montana</collectingRegion>
|
||
,
|
||
<collectingRegion id="824DF812FFAE2E59FFDBFC23FF65FBF0" box="[93,209,1000,1024]" country="United States of America" name="California" pageId="7" pageNumber="20">California</collectingRegion>
|
||
). Recorded in this paper from 19 localities, but known to be present in many more throughout the region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |