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<document id="27B9587447EE9742E9FF6E54AB3498D6" ID-CLB-Dataset="25938" ID-DOI="10.11646/zootaxa.4658.2.2" ID-GBIF-Dataset="349f1d0e-6dfc-443a-998d-0322ea62b866" ID-ISSN="1175-5326" ID-Zenodo-Dep="3375788" ID-ZooBank="A4D55B3B-9BE9-4918-89C6-8049F5895CF2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566574958471" checkinUser="plazi" docAuthor="Grubbs, Scott A. &amp; Baumann, Richard W." docDate="2019" docId="B66387BEF369B37DFF381604FDBFFD78" docLanguage="en" docName="zootaxa.4658.2.2.pdf" docOrigin="Zootaxa 4658 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Soyedina sheldoni Grubbs &amp; Baumann 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="244" masterDocId="4A5AFFC6F37BB368FFAF144DFFB9FFB7" masterDocTitle="Soyedina Ricker, 1952 (Plecoptera: Nemouridae) in the eastern Nearctic: review of species concepts, proposed morphology-based species groups, and description of a new species from North Carolina" masterLastPageNumber="250" masterPageNumber="223" pageNumber="241" updateTime="1698752216948" updateUser="ExternalLinkService">
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<mods:title id="4CC583DC8623944A427A0EC3CDD09AC5">Soyedina Ricker, 1952 (Plecoptera: Nemouridae) in the eastern Nearctic: review of species concepts, proposed morphology-based species groups, and description of a new species from North Carolina</mods:title>
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<mods:namePart id="1DED6692B4E45B98819D1DE37F08F9B1">Grubbs, Scott A.</mods:namePart>
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<mods:date id="3E8D6102C323B87F1139479E95ED48A1">2019</mods:date>
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<taxonomicName id="F9CA4D2BF369B37AFF381604FECDFDD3" ID-CoL="8NH3S" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[151,372,585,612]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="241" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF369B37AFF381604FECDFDD3" bold="true" box="[151,372,585,612]" italics="true" pageId="18" pageNumber="241">Soyedina sheldoni</emphasis>
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<taxonomicNameLabel id="178D57C1F369B37AFED41604FE60FDD4" box="[379,473,585,611]" pageId="18" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
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http://lsid.speciesfile.org/urn:lsid:
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<subSubSection id="76D06523F369B37AFF3816DCFEE1FD1C" box="[151,344,656,683]" pageId="18" pageNumber="241" type="description">
<paragraph id="3E7536A8F369B37AFF3816DCFEE1FD1C" blockId="18.[151,1052,585,683]" box="[151,344,656,683]" pageId="18" pageNumber="241">
(
<figureCitation id="A6F12A2DF369B37AFF3016DCFE91FD1C" box="[159,296,656,683]" captionStart-0="FIGURES 6568" captionStart-1="FIGURES 6974" captionStart-2="FIGURES 7578" captionStartId-0="19.[151,264,1900,1925]" captionStartId-1="20.[151,264,1621,1646]" captionStartId-2="21.[151,264,1719,1744]" captionTargetBox-0="[155,1428,920,1873]" captionTargetBox-1="[157,1424,175,1601]" captionTargetBox-2="[152,1432,737,1696]" captionTargetId-0="figure@19.[151,1435,913,1880]" captionTargetId-1="figure@20.[156,1430,174,1602]" captionTargetId-2="figure@21.[151,1436,733,1700]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." captionText-1="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." captionText-2="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri-0="https://zenodo.org/record/3375808/files/figure.png" httpUri-1="https://zenodo.org/record/3375810/files/figure.png" httpUri-2="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">Figs. 6578</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFE9C16DCFEE9FD1C" box="[307,336,657,683]" captionStart="FIGURE 81" captionStartId="24.[151,250,1453,1478]" captionTargetBox="[176,1413,196,1392]" captionTargetId="figure@24.[151,1435,174,1434]" captionTargetPageId="24" captionText="FIGURE 81. Distribution maps of eastern Nearctic Soyedina based on material examined during this study plus valid literature data. 81, S. amicalola, S. kondratieffi, and S. sheldoni n. sp." httpUri="https://zenodo.org/record/3375818/files/figure.png" pageId="18" pageNumber="241">81</figureCitation>
)
</paragraph>
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<subSubSection id="76D06523F369B37DFF381695FDBFFD78" lastPageId="21" lastPageNumber="244" pageId="18" pageNumber="241" type="materials_examined">
<paragraph id="3E7536A8F369B37AFF381695FB61FC34" blockId="18.[151,1437,728,2015]" pageId="18" pageNumber="241">
<emphasis id="0CBEEABAF369B37AFF381695FE90FD44" bold="true" box="[151,297,728,755]" pageId="18" pageNumber="241">Description.</emphasis>
Macropterous. Male forewing length 4.56.0 mm, body length
<quantity id="F9329B4DF369B37AFC561695FB39FD44" box="[1017,1152,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="18" pageNumber="241" unit="mm" value="7.5" valueMax="8.5" valueMin="6.5">6.58.5 mm</quantity>
(n=26); female forewing length
<quantity id="F9329B4DF369B37AFF4B16B1FED5FCA0" box="[228,364,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="18" pageNumber="241" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.56.5 mm</quantity>
, body length
<quantity id="F9329B4DF369B37AFDAD16B1FD31FCA0" box="[514,648,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="18" pageNumber="241" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
(n=11). General body color dark brown. Head dark brown except for a lighter medial area extending from the occiput anteriorly to, but not enclosing, the lateral ocelli, and extending into the ocellar triangle (
<figureCitation id="A6F12A2DF369B37AFEDB1708FE71FCE8" box="[372,456,836,863]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 65</figureCitation>
). Legs light brown (
<figureCitation id="A6F12A2DF369B37AFD0B1708FD4EFCE8" box="[676,759,836,863]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 65</figureCitation>
). Wings fumose, venation typical for genus with veins A
<subScript id="A24E34EDF369B37AFAC9171BFAD7FCD3" attach="left" box="[1382,1390,854,868]" fontSize="6" pageId="18" pageNumber="241">1</subScript>
and A
<subScript id="A24E34EDF369B37AFF041737FF0AFC3F" attach="left" box="[171,179,890,904]" fontSize="6" pageId="18" pageNumber="241">2</subScript>
united near posterior margin. Cerci membranous (
<figureCitation id="A6F12A2DF369B37AFD451724FC82FC33" box="[746,827,873,900]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 66</figureCitation>
) and single-segmented. Gills absent.
</paragraph>
<paragraph id="3E7536A8F369B37AFF6817C0FC7DFA50" blockId="18.[151,1437,728,2015]" pageId="18" pageNumber="241">
<emphasis id="0CBEEABAF369B37AFF6817C0FEB1FC10" box="[199,264,909,935]" italics="true" pageId="18" pageNumber="241">Male.</emphasis>
(
<figureCitation id="A6F12A2DF369B37AFEB617C0FE1DFC10" box="[281,420,908,935]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Figs. 6567</figureCitation>
;
<figureCitation id="A6F12A2DF369B37AFE1D17C0FE45FC1F" box="[434,508,909,936]" captionStart-0="FIGURES 6974" captionStart-1="FIGURES 7578" captionStartId-0="20.[151,264,1621,1646]" captionStartId-1="21.[151,264,1719,1744]" captionTargetBox-0="[157,1424,175,1601]" captionTargetBox-1="[152,1432,737,1696]" captionTargetId-0="figure@20.[156,1430,174,1602]" captionTargetId-1="figure@21.[151,1436,733,1700]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." captionText-1="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri-0="https://zenodo.org/record/3375810/files/figure.png" httpUri-1="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">6978</figureCitation>
). 7
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and 8
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abdominal terga unmodified. Hypoproct well-developed, base covering medial ⅓ of 9
<superScript id="C9BF9BE0F369B37AFEF017FDFED5FC09" attach="left" box="[351,364,944,958]" fontSize="6" pageId="18" pageNumber="241">th</superScript>
sternum, constricted in basal half to plane of the base of the paraprocts; apical half tapered and set between base of outer paraproct lobes, distally tapered markedly to a lightly-sclerotized, narrowly-triangular terminus. Vesicle base partially obscured on preserved males by posterior margin of 8
<superScript id="C9BF9BE0F369B37AFBFB17B5FBD8FBB1" attach="left" box="[1108,1121,1016,1030]" fontSize="6" pageId="18" pageNumber="241">th</superScript>
abdominal sternum; narrow basally, widest in medial ½, tapered apically; mean length (L):
<quantity id="F9329B4DF369B37AFCFD1050FC7CFB80" box="[850,965,1053,1079]" metricMagnitude="-5" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="241" unit="mm" value="0.031">0.031 mm</quantity>
, mean maximum width (
<collectionCode id="58DBAE6DF369B37AFB761050FAAAFB80" box="[1241,1299,1053,1079]" country="Netherlands" name="Museum Wasmann" pageId="18" pageNumber="241">MW</collectionCode>
):
<quantity id="F9329B4DF369B37AFA8B1050FA21FB80" box="[1316,1432,1053,1079]" metricMagnitude="-5" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="241" unit="mm" value="0.013">0.013 mm</quantity>
, mean L/
<collectionCode id="58DBAE6DF369B37AFF5A100CFE91FBEC" box="[245,296,1089,1115]" country="Netherlands" name="Museum Wasmann" pageId="18" pageNumber="241">MW</collectionCode>
ratio: 2.45 (n=11). Paraprocts with two lobes, inner lobes difficult to discern with standard light microscopy on uncleared specimens, lightly sclerotized and inconspicuous, extending only slightly to base of recurved portion of outer lobes; outer lobes heavily sclerotized, large and robust for genus, in basal half appear bulbous in ventral view (
<figureCitation id="A6F12A2DF369B37AFE9C10E0FE3CFB7F" box="[307,389,1197,1224]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 67</figureCitation>
) but essentially medially-concave in dorsal view (
<figureCitation id="A6F12A2DF369B37AFC1210E0FBA2FB70" box="[957,1051,1197,1223]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Figs. 66</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFB8810E0FBFEFB7F" box="[1063,1095,1197,1224]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">76</figureCitation>
); broadly rounded basally and dorsally (
<figureCitation id="A6F12A2DF369B37AFEAE109CFEE6FB5C" box="[257,351,1233,1259]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Figs. 66</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFEC4109CFE0CFB5C" box="[363,437,1233,1259]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">7778</figureCitation>
); recurved in anterior 1/3, slightly concave laterally (
<figureCitation id="A6F12A2DF369B37AFBA5109CFBDEFB5C" box="[1034,1127,1233,1259]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Figs. 66</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFBDC109CFB28FB5C" box="[1139,1169,1233,1259]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">69</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFB31109CFB02FB5B" box="[1182,1211,1233,1260]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">76</figureCitation>
) and dorsally (
<figureCitation id="A6F12A2DF369B37AFACB109CFF58FAB8" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">Figs. 7778</figureCitation>
), terminating anteriorly in medially-directed triangular flaps (
<figureCitation id="A6F12A2DF369B37AFC2610B8FBA9FAB8" box="[905,1040,1269,1295]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 6970</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFBB510B8FBDDFAB8" box="[1050,1124,1269,1295]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="18" pageNumber="241">7778</figureCitation>
). Epiproct typical for genus, exhibiting asymmetry with right half longer than left (
<figureCitation id="A6F12A2DF369B37AFCAF1154FC0CFA84" box="[768,949,1305,1331]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 6970,74</figureCitation>
); lateral arms short and lightly sclerotized, extending from near the distal terminus of the 10
<superScript id="C9BF9BE0F369B37AFD151171FD7EFAFD" attach="left" box="[698,711,1340,1354]" fontSize="6" pageId="18" pageNumber="241">th</superScript>
tergal sclerite to the basal cushion; ventral sclerites elongate and asymmetric, right longer than left (
<figureCitation id="A6F12A2DF369B37AFDB1112CFD1EFACB" box="[542,679,1377,1404]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 6970</figureCitation>
), studded with small teeth medially and laterally throughout length, tips typically rounded distally (
<figureCitation id="A6F12A2DF369B37AFE5611C8FD3CFA28" box="[505,645,1413,1439]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 7172</figureCitation>
); dorsal sclerite elongate and asymmetric, right longer than left (
<figureCitation id="A6F12A2DF369B37AFACB11C8FF0CFA74" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Figs. 67</figureCitation>
,
<figureCitation id="A6F12A2DF369B37AFF6F11E4FE8DFA74" box="[192,308,1449,1475]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">69, 7374</figureCitation>
), appearing scaly at higher magnifications (
<figureCitation id="A6F12A2DF369B37AFCB711E4FC19FA74" box="[792,928,1449,1475]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 7172</figureCitation>
), open apically, revealing an sclerotized inner member is either slightly tapered or straight-sided distally (
<figureCitation id="A6F12A2DF369B37AFC811180FC01FA50" box="[814,952,1485,1511]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="18" pageNumber="241">Figs. 7173</figureCitation>
).
</paragraph>
<paragraph id="3E7536A8F369B37AFF6811BCFBD3F9E3" blockId="18.[151,1437,728,2015]" pageId="18" pageNumber="241">
<emphasis id="0CBEEABAF369B37AFF6811BCFE9AF9BC" box="[199,291,1521,1547]" italics="true" pageId="18" pageNumber="241">Female.</emphasis>
(
<figureCitation id="A6F12A2DF369B37AFE9C11BCFE30F9BB" box="[307,393,1521,1548]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 68</figureCitation>
). Typical of genus. Subgenital plate well-developed, subtriangular in shape; base broad, extending from posterior of seventh sternum entirely over eighth sternum and approximately ¼ over ninth sternum; apical portion darker than basal portion, terminating in broadly-rounded apex (
<figureCitation id="A6F12A2DF369B37AFBA61274FBE4F9E3" box="[1033,1117,1593,1620]" captionStart="FIGURES 6568" captionStartId="19.[151,264,1900,1925]" captionTargetBox="[155,1428,920,1873]" captionTargetId="figure@19.[151,1435,913,1880]" captionTargetPageId="19" captionText="FIGURES 6568. Soyedina sheldoni n. sp. 6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="18" pageNumber="241">Fig. 68</figureCitation>
).
</paragraph>
<paragraph id="3E7536A8F369B37AFF681213FE14F9C0" blockId="18.[151,1437,728,2015]" box="[199,429,1629,1655]" pageId="18" pageNumber="241">
<emphasis id="0CBEEABAF369B37AFF681213FEB3F9C0" box="[199,266,1630,1655]" italics="true" pageId="18" pageNumber="241">Larva</emphasis>
. Undescribed.
</paragraph>
<paragraph id="3E7536A8F369B37BFF6812CCFBE3FD88" blockId="18.[151,1437,728,2015]" lastBlockId="19.[151,1437,152,863]" lastPageId="19" lastPageNumber="242" pageId="18" pageNumber="241">
<emphasis id="0CBEEABAF369B37AFF6812CCFEC9F92C" bold="true" box="[199,368,1664,1691]" pageId="18" pageNumber="241">Type material</emphasis>
.
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<typeStatus id="E171880AF369B37AFE2F12CCFE51F92C" box="[384,488,1665,1691]" pageId="18" pageNumber="241" type="holotype">Holotype</typeStatus>
male (
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):
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</emphasis>
,
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,
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,
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</date>
,
<collectorName id="933F537EF369B37AFCD912E8FBAAF908" box="[886,1043,1701,1727]" pageId="18" pageNumber="241">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF369B37AFB9712E8FB74F908" box="[1080,1229,1701,1727]" pageId="18" pageNumber="241">E.T. Metzger</collectorName>
</materialsCitation>
.
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<typeStatus id="E171880AF369B37AFB7712E8FAF0F908" box="[1240,1353,1701,1727]" pageId="18" pageNumber="241" type="paratype">Paratypes</typeStatus>
:
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<collectingRegion id="FC0EF84AF369B37AFAFA12E8FEB9F954" country="United States of America" name="North Carolina" pageId="18" pageNumber="241">North Carolina</collectingRegion>
</emphasis>
,
<collectingCounty id="D7144E24F369B37AFEBE1284FE2BF954" box="[273,402,1737,1763]" pageId="18" pageNumber="241">Macon Co.</collectingCounty>
,
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,
<location id="3B156073F369B37AFDDC1284FC5CF954" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFDDC1284FC5CF954" box="[627,997,1737,1763]" country="United States of America" county="Macon Co." latitude="35.03804" longLatPrecision="1" longitude="-83.45908" municipality="upper Ball Creek" name="Coweeta Hydrologic Laboratory" pageId="18" pageNumber="241" stateProvince="North Carolina">Coweeta Hydrologic Laboratory</location>
,
<location id="3B156073F369B37AFC5D1284FAA6F954" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFC5D1284FAA6F954" box="[1010,1311,1737,1763]" country="United States of America" county="Macon Co." latitude="35.03804" longLatPrecision="1" longitude="-83.45908" municipality="upper Ball Creek" name="Nantahala National Forest" pageId="18" pageNumber="241" stateProvince="North Carolina">Nantahala National Forest</location>
,
<geoCoordinate id="5BFE506FF369B37AFA831285FA21F954" box="[1324,1432,1736,1763]" degrees="35.03804" direction="north" orientation="latitude" pageId="18" pageNumber="241" precision="1" value="35.03804">35.03804</geoCoordinate>
,
<geoCoordinate id="5BFE506FF369B37AFF3812A1FEB4F8B0" box="[151,269,1772,1799]" degrees="83.45908" direction="west" orientation="longitude" pageId="18" pageNumber="241" precision="1" value="-83.45908">-83.45908</geoCoordinate>
,
<date id="4A741068F369B37AFEB612A0FD90F8B0" box="[281,553,1773,1799]" pageId="18" pageNumber="241" value="1984-03-20" valueMax="1984-04-07" valueMin="1984-03-20">
<collectingDate id="5A30E980F369B37AFEB612A0FD90F8B0" box="[281,553,1773,1799]" pageId="18" pageNumber="241" value="1984-03-20" valueMax="1984-04-07" valueMin="1984-03-20">20 March7 April 1984</collectingDate>
</date>
,
<collectorName id="933F537EF369B37AFD9B12A0FD04F8B0" box="[564,701,1773,1799]" pageId="18" pageNumber="241">A.D. Huryn</collectorName>
,
<specimenCount id="28CCFD21F369B37AFD6412A0FCAEF8B0" box="[715,791,1773,1799]" pageId="18" pageNumber="241" type="male">1 male</specimenCount>
(
<specimenCode id="6E6C9ED3F369B37AFC8712A0FC13F8B0" box="[808,938,1773,1799]" collectionCode="BYU" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="18" pageNumber="241" type="University or college">BYU 1933</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37AFC6F12A0FEC8F89C" ID-GBIF-Occurrence="2351014758" collectingDate="1984-04-07" collectingDateMax="1984-04-20" collectingDateMin="1984-04-07" collectionCode="BYU" collectorName="Huryn" country="United States of America" pageId="18" pageNumber="241" specimenCode="BYU 1934" specimenCount="2" specimenCount-male="2" stateProvince="North Carolina" typeStatus="paratype">
same but
<date id="4A741068F369B37AFB8112A0FB54F8B0" box="[1070,1261,1773,1799]" pageId="18" pageNumber="241" value="1984-04-07" valueMax="1984-04-20" valueMin="1984-04-07">
<collectingDate id="5A30E980F369B37AFB8112A0FB54F8B0" box="[1070,1261,1773,1799]" pageId="18" pageNumber="241" value="1984-04-07" valueMax="1984-04-20" valueMin="1984-04-07">720 April 1984</collectingDate>
</date>
, A.D.
<collectorName id="933F537EF369B37AFA9812A0FA38F8B0" box="[1335,1409,1773,1799]" pageId="18" pageNumber="241">Huryn</collectorName>
,
<specimenCount id="28CCFD21F369B37AFA2112A0FF61F89C" pageId="18" pageNumber="241" type="male">2 males</specimenCount>
(
<specimenCode id="6E6C9ED3F369B37AFF47135CFED0F89C" box="[232,361,1809,1835]" collectionCode="BYU" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="18" pageNumber="241" type="University or college">BYU 1934</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37AFED1135CFB5AF89B" ID-GBIF-Occurrence="2351014724" box="[382,1251,1809,1836]" collectingDate="1984-04-20" collectingDateMax="1984-05-20" collectingDateMin="1984-04-20" collectionCode="BYU" collectorName="Huryn" country="United States of America" pageId="18" pageNumber="241" specimenCode="BYU 1936" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
same but same but
<date id="4A741068F369B37AFDF6135CFCE7F89C" box="[601,862,1809,1835]" pageId="18" pageNumber="241" value="1984-04-20" valueMax="1984-05-20" valueMin="1984-04-20">
<collectingDate id="5A30E980F369B37AFDF6135CFCE7F89C" box="[601,862,1809,1835]" pageId="18" pageNumber="241" value="1984-04-20" valueMax="1984-05-20" valueMin="1984-04-20">20 April20 May 1984</collectingDate>
</date>
, A.D.
<collectorName id="933F537EF369B37AFC08135CFC48F89C" box="[935,1009,1809,1835]" pageId="18" pageNumber="241">Huryn</collectorName>
,
<specimenCount id="28CCFD21F369B37AFC51135CFBF3F89C" box="[1022,1098,1809,1835]" pageId="18" pageNumber="241" type="male">1 male</specimenCount>
(
<specimenCode id="6E6C9ED3F369B37AFBF5135CFB62F89B" box="[1114,1243,1809,1836]" collectionCode="BYU" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="18" pageNumber="241" type="University or college">BYU 1936</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37AFB40135CFF51F8C4" ID-GBIF-Occurrence="2351014865" collectingDate="1994-05-25" collectionCode="BYU" collectorName="Szczytko &amp; J. B. Sandberg &amp; B. P. Stark" country="United States of America" county="Yancey Co." location="Lower Creek" municipality="Camp Alice" pageId="18" pageNumber="241" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F369B37AFB40135CFACDF89C" box="[1263,1396,1809,1835]" pageId="18" pageNumber="241">Yancey Co.</collectingCounty>
, on road to
<collectingMunicipality id="DE11ACD2F369B37AFF431378FEC8F8F8" box="[236,369,1845,1871]" pageId="18" pageNumber="241">Camp Alice</collectingMunicipality>
(=
<location id="3B156073F369B37AFE391378FD96F8E7" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFE391378FD96F8E7" box="[406,559,1845,1872]" country="United States of America" county="Yancey Co." municipality="Camp Alice" name="Lower Creek" pageId="18" pageNumber="241" stateProvince="North Carolina">Lower Creek</location>
?),
<date id="4A741068F369B37AFDE31379FD5AF8F8" box="[588,739,1844,1871]" pageId="18" pageNumber="241" value="1994-05-25">
<collectingDate id="5A30E980F369B37AFDE31379FD5AF8F8" box="[588,739,1844,1871]" pageId="18" pageNumber="241" value="1994-05-25">25 May 1994</collectingDate>
</date>
, S.W.
<collectorName id="933F537EF369B37AFC851378FC2AF8F8" box="[810,915,1845,1871]" pageId="18" pageNumber="241">Szczytko</collectorName>
,
<collectorName id="933F537EF369B37AFC311378FB80F8F8" box="[926,1081,1845,1871]" pageId="18" pageNumber="241">J.B. Sandberg</collectorName>
&amp;
<collectorName id="933F537EF369B37AFBF31378FB77F8F8" box="[1116,1230,1845,1871]" pageId="18" pageNumber="241">B.P. Stark</collectorName>
,
<specimenCount id="28CCFD21F369B37AFB771378FA9FF8F8" box="[1240,1318,1845,1871]" pageId="18" pageNumber="241" type="male">1 male</specimenCount>
,
<specimenCount id="28CCFD21F369B37AFA9E1378FA25F8F8" box="[1329,1436,1845,1871]" pageId="18" pageNumber="241" type="female">4 females</specimenCount>
(
<collectionCode id="58DBAE6DF369B37AFF301314FF59F8C4" box="[159,224,1881,1907]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="18" pageNumber="241" type="University or college">BYU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37AFF5A1314FC36F820" ID-GBIF-Occurrence="2351014756" collectingDate="2016-04-28" collectionCode="NCDWR" collectorName="E. Fleek &amp; S. Beaty" country="Netherlands" county="Balsam Spring" latitude="35.76653" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.26403" municipality="Mount Mitchell Summit" pageId="18" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
seep (=
<collectingCounty id="D7144E24F369B37AFEFE1314FE44F8C3" box="[337,509,1881,1908]" pageId="18" pageNumber="241">Balsam Spring</collectingCounty>
), near
<collectingMunicipality id="DE11ACD2F369B37AFDE51314FCD9F8C4" box="[586,864,1881,1907]" pageId="18" pageNumber="241">Mount Mitchell Summit</collectingMunicipality>
,
<location id="3B156073F369B37AFCC21314FB27F8C4" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFCC21314FB27F8C4" box="[877,1182,1881,1907]" country="Netherlands" county="Balsam Spring" latitude="35.76653" longLatPrecision="1" longitude="-82.26403" municipality="Mount Mitchell Summit" name="Mount Mitchell State Park" pageId="18" pageNumber="241" stateProvince="North Carolina">Mount Mitchell State Park</location>
,
<geoCoordinate id="5BFE506FF369B37AFB051315FAAFF8C4" box="[1194,1302,1880,1907]" degrees="35.76653" direction="north" orientation="latitude" pageId="18" pageNumber="241" precision="1" value="35.76653">35.76653</geoCoordinate>
,
<geoCoordinate id="5BFE506FF369B37AFA8C1314FA21F8C4" box="[1315,1432,1881,1907]" degrees="82.26403" direction="west" orientation="longitude" pageId="18" pageNumber="241" precision="1" value="-82.26403">-82.26403</geoCoordinate>
,
<date id="4A741068F369B37AFF381330FE8EF820" box="[151,311,1917,1943]" pageId="18" pageNumber="241" value="2016-04-28">
<collectingDate id="5A30E980F369B37AFF381330FE8EF820" box="[151,311,1917,1943]" pageId="18" pageNumber="241" value="2016-04-28">28 April 2016</collectingDate>
</date>
, V.
<collectingCountry id="46DD7638F369B37AFECD1330FE06F820" box="[354,447,1917,1943]" name="Netherlands" pageId="18" pageNumber="241">Holland</collectingCountry>
,
<collectorName id="933F537EF369B37AFE641330FD90F820" box="[459,553,1917,1943]" pageId="18" pageNumber="241">E. Fleek</collectorName>
&amp;
<collectorName id="933F537EF369B37AFDE01330FD16F820" box="[591,687,1917,1943]" pageId="18" pageNumber="241">S. Beaty</collectorName>
,
<specimenCount id="28CCFD21F369B37AFD131330FCB1F820" box="[700,776,1917,1943]" pageId="18" pageNumber="241" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF369B37AFCB71330FC3EF820" box="[792,903,1917,1943]" pageId="18" pageNumber="241">NCDWR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37AFC351330FEBAF868" ID-GBIF-Occurrence="2351014691" collectingDate="2019-04-25" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Balsam Spring" latitude="35.76653" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.26403" municipality="Mount Mitchell" pageId="18" pageNumber="241" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F369B37AFC351330FBFAF820" box="[922,1091,1917,1943]" pageId="18" pageNumber="241">Balsam Spring</collectingCounty>
, near
<collectingMunicipality id="DE11ACD2F369B37AFB271330FA80F820" box="[1160,1337,1917,1943]" pageId="18" pageNumber="241">Mount Mitchell</collectingMunicipality>
summit,
<location id="3B156073F369B37AFF3813ECFE7AF80C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFF3813ECFE7AF80C" box="[151,451,1953,1979]" country="Netherlands" county="Balsam Spring" latitude="35.76653" longLatPrecision="1" longitude="-82.26403" municipality="Mount Mitchell" name="Mount Mitchell State Park" pageId="18" pageNumber="241" stateProvince="North Carolina">Mount Mitchell State Park</location>
<geoCoordinate id="5BFE506FF369B37AFE6413EDFD8EF80C" box="[459,567,1952,1979]" degrees="35.76653" direction="north" orientation="latitude" pageId="18" pageNumber="241" precision="1" value="35.76653">35.76653</geoCoordinate>
,
<geoCoordinate id="5BFE506FF369B37AFDEC13ECFD00F80C" box="[579,697,1953,1979]" degrees="82.26403" direction="west" orientation="longitude" pageId="18" pageNumber="241" precision="1" value="-82.26403">-82.26403</geoCoordinate>
,
<date id="4A741068F369B37AFD6B13EDFCDCF80C" box="[708,869,1952,1979]" pageId="18" pageNumber="241" value="2019-04-25">
<collectingDate id="5A30E980F369B37AFD6B13EDFCDCF80C" box="[708,869,1952,1979]" pageId="18" pageNumber="241" value="2019-04-25">25 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF369B37AFCDE13ECFBB4F80C" box="[881,1037,1953,1979]" pageId="18" pageNumber="241">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF369B37AFB9C13ECFB71F80C" box="[1075,1224,1953,1979]" pageId="18" pageNumber="241">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F369B37AFB7B13ECFA9AF80C" box="[1236,1315,1953,1979]" pageId="18" pageNumber="241" type="male">1 male</specimenCount>
,
<specimenCount id="28CCFD21F369B37AFA8013ECFA25F80C" box="[1327,1436,1953,1979]" pageId="18" pageNumber="241" type="female">2 females</specimenCount>
(
<collectionCode id="58DBAE6DF369B37AFF301388FF42F868" box="[159,251,1989,2015]" pageId="18" pageNumber="241">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F369B37BFEBE1388FC85FF04" ID-GBIF-Occurrence="2351014843" collectingDate="2019-04-25" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" lastPageId="19" lastPageNumber="242" latitude="35.74395" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.28172" pageId="18" pageNumber="241" specimenCount="5" specimenCount-larva="5" stateProvince="North Carolina" typeStatus="paratype">
spring along old railroad bed,
<location id="3B156073F369B37AFDDE1388FC1FF868" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F369B37AFDDE1388FC1FF868" box="[625,934,1989,2015]" country="Netherlands" latitude="35.74395" longLatPrecision="1" longitude="-82.28172" name="Mount Mitchell State Park" pageId="18" pageNumber="241" stateProvince="North Carolina">Mount Mitchell State Park</location>
,
<geoCoordinate id="5BFE506FF369B37AFC1A1389FB98F868" box="[949,1057,1988,2015]" degrees="35.74395" direction="north" orientation="latitude" pageId="18" pageNumber="241" precision="1" value="35.74395">35.74395</geoCoordinate>
,
<geoCoordinate id="5BFE506FF369B37AFB801388FB1DF868" box="[1071,1188,1989,2015]" degrees="82.28172" direction="west" orientation="longitude" pageId="18" pageNumber="241" precision="1" value="-82.28172">-82.28172</geoCoordinate>
,
<date id="4A741068F369B37AFB1D1389FAEFF868" box="[1202,1366,1988,2015]" pageId="18" pageNumber="241" value="2019-04-25">
<collectingDate id="5A30E980F369B37AFB1D1389FAEFF868" box="[1202,1366,1988,2015]" pageId="18" pageNumber="241" value="2019-04-25">25 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF369B37BFACB1388FF4DFF04" lastPageId="19" lastPageNumber="242" pageId="18" pageNumber="241">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFEB614D4FE17FF04" box="[281,430,153,179]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
, 1 pharate male,
<specimenCount id="28CCFD21F368B37BFDC014D5FD71FF04" box="[623,712,152,179]" pageId="19" pageNumber="242" type="larva">5 larvae</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFD7714D4FC8DFF04" box="[728,820,153,179]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFCE714D4FA21FF60" ID-GBIF-Occurrence="2351014752" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Right Prong South Toe River" latitude="35.72718" location="Pisgah National Forest" longLatPrecision="1" longitude="-82.28379" municipality="Buncombe Trail" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
tributary to
<collectingCounty id="D7144E24F368B37BFC6214D4FAAEFF04" box="[973,1303,153,179]" pageId="19" pageNumber="242">Right Prong South Toe River</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFA8C14D4FF76FF60" pageId="19" pageNumber="242">Buncombe Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFF7514F0FE63FF60" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFF7514F0FE63FF60" box="[218,474,189,215]" country="Netherlands" county="Right Prong South Toe River" latitude="35.72718" longLatPrecision="1" longitude="-82.28379" municipality="Buncombe Trail" name="Pisgah National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Pisgah National Forest</location>
,
<geoCoordinate id="5BFE506FF368B37BFE4B14F1FDE9FF60" box="[484,592,188,215]" degrees="35.72718" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.72718">35.72718</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFDF514F0FD69FF60" box="[602,720,189,215]" degrees="82.28379" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.28379">-82.28379</geoCoordinate>
,
<date id="4A741068F368B37BFD7514F0FCCFFF60" box="[730,886,189,215]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFD7514F0FCCFFF60" box="[730,886,189,215]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFC2E14F0FBA5FF60" box="[897,1052,189,215]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFB9114F0FB68FF60" box="[1086,1233,189,215]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFB7414F0FA9FFF60" box="[1243,1318,189,215]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFA9B14F0FA29FF60" box="[1332,1424,189,215]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFF3814ACFD56FEA8" ID-GBIF-Occurrence="2351014803" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Right Prong South Toe River" latitude="35.729" location="Pisgah National Forest" longLatPrecision="1" longitude="-82.28331" municipality="Buncombe Trail" pageId="19" pageNumber="242" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F368B37BFF3814ACFE58FF4C" box="[151,481,225,251]" pageId="19" pageNumber="242">Right Prong South Toe River</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFE4214ACFD1CFF4C" box="[493,677,225,251]" pageId="19" pageNumber="242">Buncombe Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFD1E14ACFC0CFF4C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFD1E14ACFC0CFF4C" box="[689,949,225,251]" country="Netherlands" county="Right Prong South Toe River" latitude="35.729" longLatPrecision="1" longitude="-82.28331" municipality="Buncombe Trail" name="Pisgah National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Pisgah National Forest</location>
,
<geoCoordinate id="5BFE506FF368B37BFC6F14ADFB95FF4C" box="[960,1068,224,251]" degrees="35.72900" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.729">35.72900</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFB9714ACFB14FF4C" box="[1080,1197,225,251]" degrees="82.28331" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.28331">-82.28331</geoCoordinate>
,
<date id="4A741068F368B37BFB1614ACFAE0FF4C" box="[1209,1369,225,251]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFB1614ACFAE0FF4C" box="[1209,1369,225,251]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFACB14ACFF4DFEA8" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFEB41548FE09FEA8" box="[283,432,261,287]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFE121548FDB5FEA8" box="[445,524,261,287]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFDB61548FDC2FEA8" box="[537,635,261,287]" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFD241548FD5EFEA8" box="[651,743,261,287]" pageId="19" pageNumber="242">WKUC</collectionCode>
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;
<materialsCitation id="8EA23CF5F368B37BFD531548FAF2FEF4" ID-GBIF-Occurrence="2351014879" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="South Fork Upper Creek" latitude="35.73634" location="Pisgah National Forest" longLatPrecision="1" longitude="-82.27946" municipality="Buncombe Trail" pageId="19" pageNumber="242" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F368B37BFD531548FBAEFEA8" box="[764,1047,261,287]" pageId="19" pageNumber="242">South Fork Upper Creek</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFB8C1548FB65FEA8" box="[1059,1244,261,287]" pageId="19" pageNumber="242">Buncombe Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFB461548FF59FEF4" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFB461548FF59FEF4" country="Netherlands" county="South Fork Upper Creek" latitude="35.73634" longLatPrecision="1" longitude="-82.27946" municipality="Buncombe Trail" name="Pisgah National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Pisgah National Forest</location>
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<geoCoordinate id="5BFE506FF368B37BFEC81564FE64FEF4" box="[359,477,297,323]" degrees="82.27946" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.27946">-82.27946</geoCoordinate>
,
<date id="4A741068F368B37BFE451564FD37FEF4" box="[490,654,297,323]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFE451564FD37FEF4" box="[490,654,297,323]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFD331564FC83FEF4" box="[668,826,297,323]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFCCB1564FC43FEF4" box="[868,1018,297,323]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFBA71564FBDDFEF4" box="[1032,1124,297,323]" pageId="19" pageNumber="242" type="male">4 males</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFBDD1564FB6CFEF4" box="[1138,1237,297,323]" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFB481564FAFAFEF4" box="[1255,1347,297,323]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFAF61564FE71FE3C" ID-GBIF-Occurrence="2351014716" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" latitude="35.74279" location="Pisgah National Forest" longLatPrecision="1" longitude="-82.27468" municipality="Buncombe Trail" pageId="19" pageNumber="242" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="North Carolina" typeStatus="paratype">
seeps,
<collectingMunicipality id="DE11ACD2F368B37BFF381500FEEBFED0" box="[151,338,333,359]" pageId="19" pageNumber="242">Buncombe Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFECF1500FDDEFED0" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFECF1500FDDEFED0" box="[352,615,333,359]" country="Netherlands" latitude="35.74279" longLatPrecision="1" longitude="-82.27468" municipality="Buncombe Trail" name="Pisgah National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Pisgah National Forest</location>
,
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,
<geoCoordinate id="5BFE506FF368B37BFD411500FCDAFED0" box="[750,867,333,359]" degrees="82.27468" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.27468">-82.27468</geoCoordinate>
,
<date id="4A741068F368B37BFCDE1500FBADFED0" box="[881,1044,333,359]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFCDE1500FBADFED0" box="[881,1044,333,359]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFB8D1500FB78FED0" box="[1058,1217,333,359]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFB451500FA38FED0" box="[1258,1409,333,359]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFA211500FF62FE3C" pageId="19" pageNumber="242" type="male">4 males</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFF46153CFEF5FE3C" box="[233,332,369,395]" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFEF2153CFE0DFE3C" box="[349,436,369,395]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFE78153CFC6BFE18" ID-GBIF-Occurrence="2351014826" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Lower Creek" latitude="35.75564" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.26803" municipality="Buncombe Trail" pageId="19" pageNumber="242" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F368B37BFE78153CFDC9FE3C" box="[471,624,369,395]" pageId="19" pageNumber="242">Lower Creek</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFDD2153CFC8EFE3C" box="[637,823,369,395]" pageId="19" pageNumber="242">Buncombe Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFCEB153CFBCEFE3C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFCEB153CFBCEFE3C" box="[836,1143,369,395]" country="Netherlands" county="Lower Creek" latitude="35.75564" longLatPrecision="1" longitude="-82.26803" municipality="Buncombe Trail" name="Mount Mitchell State Park" pageId="19" pageNumber="242" stateProvince="North Carolina">Mount Mitchell State Park</location>
,
<geoCoordinate id="5BFE506FF368B37BFB2A153DFB48FE3C" box="[1157,1265,368,395]" degrees="35.75564" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.75564">35.75564</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFB51153CFACAFE3C" box="[1278,1395,369,395]" degrees="82.26803" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.26803">-82.26803</geoCoordinate>
,
<date id="4A741068F368B37BFA2F153CFEAEFE18" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFA2F153CFEAEFE18" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFE8B15D8FE7AFE18" box="[292,451,405,431]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFE4315D8FD3BFE18" box="[492,642,405,431]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFD3F15D8FD52FE18" box="[656,747,405,431]" pageId="19" pageNumber="242" type="male">2 males</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFD5615D8FCE4FE18" box="[761,861,405,431]" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFCC115D8FC73FE18" box="[878,970,405,431]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFC4F15D8FEEFFE40" ID-GBIF-Occurrence="2351014708" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Lower Creek" latitude="35.7563" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.2646" municipality="Commissary Trail" pageId="19" pageNumber="242" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="North Carolina" typeStatus="paratype">
tributary to
<collectingCounty id="D7144E24F368B37BFBC615D8FAB8FE18" box="[1129,1281,405,431]" pageId="19" pageNumber="242">Lower Creek</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFAA015D8FF76FE64" pageId="19" pageNumber="242">Commissary Trail</collectingMunicipality>
,
<location id="3B156073F368B37BFF7315F4FDB4FE64" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFF7315F4FDB4FE64" box="[220,525,441,467]" country="Netherlands" county="Lower Creek" latitude="35.7563" longLatPrecision="1" longitude="-82.2646" municipality="Commissary Trail" name="Mount Mitchell State Park" pageId="19" pageNumber="242" stateProvince="North Carolina">Mount Mitchell State Park</location>
,
<geoCoordinate id="5BFE506FF368B37BFDB615F5FD3CFE64" box="[537,645,440,467]" degrees="35.75630" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.7563">35.75630</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFD3D15F4FCBEFE64" box="[658,775,441,467]" degrees="82.26460" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.2646">-82.26460</geoCoordinate>
,
<date id="4A741068F368B37BFCBC15F4FC0CFE64" box="[787,949,441,467]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFCBC15F4FC0CFE64" box="[787,949,441,467]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFC6E15F4FBE7FE64" box="[961,1118,441,467]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFB2A15F4FAA2FE64" box="[1157,1307,441,467]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFA8815F4FA3BFE64" box="[1319,1410,441,467]" pageId="19" pageNumber="242" type="male">3 males</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFA2115F4FF5AFE40" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFF5D1590FEF7FE40" box="[242,334,477,503]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFECF1590FD5FFDAC" ID-GBIF-Occurrence="2351014687" collectingDate="2019-04-27" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" county="Lower Creek" latitude="35.75953" location="Mount Mitchell State Park" longLatPrecision="1" longitude="-82.26683" municipality="Camp Alice" pageId="19" pageNumber="242" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
<collectingCounty id="D7144E24F368B37BFECF1590FE4CFE40" box="[352,501,477,503]" pageId="19" pageNumber="242">Lower Creek</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFDAF1590FD3FFE40" box="[512,646,477,503]" pageId="19" pageNumber="242">Camp Alice</collectingMunicipality>
,
<location id="3B156073F368B37BFD3E1590FC05FE40" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFD3E1590FC05FE40" box="[657,956,477,503]" country="Netherlands" county="Lower Creek" latitude="35.75953" longLatPrecision="1" longitude="-82.26683" municipality="Camp Alice" name="Mount Mitchell State Park" pageId="19" pageNumber="242" stateProvince="North Carolina">Mount Mitchell State Park</location>
,
<geoCoordinate id="5BFE506FF368B37BFC681591FB8AFE40" box="[967,1075,476,503]" degrees="35.75953" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.75953">35.75953</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFB921590FB0BFE40" box="[1085,1202,477,503]" degrees="82.26683" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.26683">-82.26683</geoCoordinate>
,
<date id="4A741068F368B37BFB121590FAE3FE40" box="[1213,1370,477,503]" pageId="19" pageNumber="242" value="2019-04-27">
<collectingDate id="5A30E980F368B37BFB121590FAE3FE40" box="[1213,1370,477,503]" pageId="19" pageNumber="242" value="2019-04-27">27 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFACB1590FF4DFDAC" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFEB7164CFE14FDAC" box="[280,429,513,539]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFE17164CFDBFFDAC" box="[440,518,513,539]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFDBD164CFDCAFDAC" box="[530,627,513,539]" pageId="19" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFD2D164CFD67FDAC" box="[642,734,513,539]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFD5E164CFBEFFD88" ID-GBIF-Occurrence="2351014754" collectingDate="2019-04-28" collectionCode="WKUC" collectorName="M. L. Metzger &amp; E. T. Metzger" country="Netherlands" latitude="35.71005" location="Pisgah National Forest" longLatPrecision="1" longitude="-82.24959" municipality="Hemphill Creek" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
tributary to
<collectingMunicipality id="DE11ACD2F368B37BFCD9164CFB95FDAC" box="[886,1068,513,539]" pageId="19" pageNumber="242">Hemphill Creek</collectingMunicipality>
, FS 472,
<location id="3B156073F368B37BFB39164CFA21FDAC" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFB39164CFA21FDAC" box="[1174,1432,513,539]" country="Netherlands" latitude="35.71005" longLatPrecision="1" longitude="-82.24959" municipality="Hemphill Creek" name="Pisgah National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Pisgah National Forest</location>
,
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,
<geoCoordinate id="5BFE506FF368B37BFEA11669FE3DFD88" box="[270,388,548,575]" degrees="82.24959" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.24959">-82.24959</geoCoordinate>
,
<date id="4A741068F368B37BFE201668FD94FD88" box="[399,557,549,575]" pageId="19" pageNumber="242" value="2019-04-28">
<collectingDate id="5A30E980F368B37BFE201668FD94FD88" box="[399,557,549,575]" pageId="19" pageNumber="242" value="2019-04-28">28 April 2019</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFD971668FD6DFD88" box="[568,724,549,575]" pageId="19" pageNumber="242">M.L. Metzger</collectorName>
&amp;
<collectorName id="933F537EF368B37BFD571668FC35FD88" box="[760,908,549,575]" pageId="19" pageNumber="242">E.T. Metzger</collectorName>
,
<specimenCount id="28CCFD21F368B37BFC381668FC5BFD88" box="[919,994,549,575]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFC5D1668FBF7FD88" box="[1010,1102,549,575]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3E7536A8F368B37BFF681605FAF2FCE8" blockId="19.[151,1437,152,863]" pageId="19" pageNumber="242">
<emphasis id="0CBEEABAF368B37BFF681605FD89FDD4" bold="true" box="[199,560,584,611]" pageId="19" pageNumber="242">Additional material examined.</emphasis>
<materialsCitation id="8EA23CF5F368B37BFD9A1605FCA8FD30" ID-GBIF-Occurrence="2351014855" collectingDate="2014-05-26" collectionCode="WKUC" collectorName="A. L. Sheldon" country="United States of America" county="Haywood Co." latitude="35.3225" location="Trail" longLatPrecision="1" longitude="-82.83357" municipality="Yellowstone Prong" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<emphasis id="0CBEEABAF368B37BFD9A1605FC97FDD4" bold="true" box="[565,814,584,611]" pageId="19" pageNumber="242">
<collectingCountry id="46DD7638F368B37BFD9A1605FDD6FDD4" box="[565,623,584,611]" name="United States of America" pageId="19" pageNumber="242">USA</collectingCountry>
,
<collectingRegion id="FC0EF84AF368B37BFDD71604FC97FDD4" box="[632,814,584,611]" country="United States of America" name="North Carolina" pageId="19" pageNumber="242">North Carolina</collectingRegion>
</emphasis>
,
<collectingCounty id="D7144E24F368B37BFC951604FC6FFDD4" box="[826,982,585,611]" pageId="19" pageNumber="242">Haywood Co.</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFC711604FB0DFDD4" box="[990,1204,585,611]" pageId="19" pageNumber="242">Yellowstone Prong</collectingMunicipality>
,
<location id="3B156073F368B37BFB131604FB48FDD4" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFB131604FB48FDD4" box="[1212,1265,585,611]" country="United States of America" county="Haywood Co." latitude="35.3225" longLatPrecision="1" longitude="-82.83357" municipality="Yellowstone Prong" name="Trail" pageId="19" pageNumber="242" stateProvince="North Carolina">Trail</location>
440,
<geoCoordinate id="5BFE506FF368B37BFA831605FA21FDD4" box="[1324,1432,584,611]" degrees="35.32250" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.3225">35.32250</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFF381621FEB4FD30" box="[151,269,620,647]" degrees="82.83357" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.83357">-82.83357</geoCoordinate>
,
<date id="4A741068F368B37BFEB91620FE15FD30" box="[278,428,621,647]" pageId="19" pageNumber="242" value="2014-05-26">
<collectingDate id="5A30E980F368B37BFEB91620FE15FD30" box="[278,428,621,647]" pageId="19" pageNumber="242" value="2014-05-26">26 May 2014</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFE1B1620FDF5FD30" box="[436,588,621,647]" pageId="19" pageNumber="242">A.L. Sheldon</collectorName>
,
<specimenCount id="28CCFD21F368B37BFDF91620FD26FD30" box="[598,671,621,647]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFD021620FCB0FD30" box="[685,777,621,647]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFCB41620FA93FD1C" ID-GBIF-Occurrence="2351014773" collectingDate="1984-03-20" collectionCode="WKUC" collectorName="A. D. Huryn" country="United States of America" county="Macon Co." latitude="35.03804" location="Coweeta Hydrologic Laboratory" longLatPrecision="1" longitude="-83.45908" municipality="upper Ball Creek" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<collectingCounty id="D7144E24F368B37BFCB41620FC20FD30" box="[795,921,621,647]" pageId="19" pageNumber="242">Macon Co.</collectingCounty>
,
<collectingMunicipality id="DE11ACD2F368B37BFC0C1620FBDAFD30" box="[931,1123,621,647]" pageId="19" pageNumber="242">upper Ball Creek</collectingMunicipality>
,
<location id="3B156073F368B37BFBC31620FF62FD1C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFBC31620FF62FD1C" country="United States of America" county="Macon Co." latitude="35.03804" longLatPrecision="1" longitude="-83.45908" municipality="upper Ball Creek" name="Coweeta Hydrologic Laboratory" pageId="19" pageNumber="242" stateProvince="North Carolina">Coweeta Hydrologic Laboratory</location>
,
<location id="3B156073F368B37BFF4816DCFDAAFD1C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFF4816DCFDAAFD1C" box="[231,531,657,683]" country="United States of America" county="Macon Co." latitude="35.03804" longLatPrecision="1" longitude="-83.45908" municipality="upper Ball Creek" name="Nantahala National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Nantahala National Forest</location>
,
<geoCoordinate id="5BFE506FF368B37BFDB016DDFD32FD1C" box="[543,651,656,683]" degrees="35.03804" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.03804">35.03804</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFD3816DDFCB4FD1C" box="[663,781,656,683]" degrees="83.45908" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-83.45908">-83.45908</geoCoordinate>
,
<date id="4A741068F368B37BFCB616DCFC70FD1C" box="[793,969,657,683]" pageId="19" pageNumber="242" value="1984-03-20">
<collectingDate id="5A30E980F368B37BFCB616DCFC70FD1C" box="[793,969,657,683]" pageId="19" pageNumber="242" value="1984-03-20">20 March 1984</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFC7B16DCFBE4FD1C" box="[980,1117,657,683]" pageId="19" pageNumber="242">A.D. Huryn</collectorName>
,
<specimenCount id="28CCFD21F368B37BFBC516DCFB0FFD1C" box="[1130,1206,657,683]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFB6916DCFA9BFD1C" box="[1222,1314,657,683]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFA9816DCFD1EFD78" ID-GBIF-Occurrence="2351014728" collectingDate="1984-04-08" collectingDateMax="1984-04-20" collectingDateMin="1984-04-08" collectionCode="ADHC" collectorName="Huryn" country="United States of America" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
same but
<date id="4A741068F368B37BFF3816F8FEEBFD78" box="[151,338,693,719]" pageId="19" pageNumber="242" value="1984-04-08" valueMax="1984-04-20" valueMin="1984-04-08">
<collectingDate id="5A30E980F368B37BFF3816F8FEEBFD78" box="[151,338,693,719]" pageId="19" pageNumber="242" value="1984-04-08" valueMax="1984-04-20" valueMin="1984-04-08">820 April 1984</collectingDate>
</date>
, A.D.
<collectorName id="933F537EF368B37BFE3616F8FE5AFD78" box="[409,483,693,719]" pageId="19" pageNumber="242">Huryn</collectorName>
,
<specimenCount id="28CCFD21F368B37BFE4016F8FD83FD78" box="[495,570,693,719]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFDE616F8FD26FD78" box="[585,671,693,719]" pageId="19" pageNumber="242">ADHC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFD1D16F8FA21FD78" ID-GBIF-Occurrence="2351014722" box="[690,1432,693,719]" collectingDate="20 April- 2019 April 1984" collectionCode="ADHC" collectorName="Huryn" country="United States of America" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
same but
<collectingDate id="5A30E980F368B37BFCB116F8FBFAFD78" box="[798,1091,693,719]" isEnumeration="true" pageId="19" pageNumber="242">
<date id="4A741068F368B37BFCB116F8FC79FD78" box="[798,960,693,719]" pageId="19" pageNumber="242" value="2019-04-20">20 April2019</date>
<date id="4A741068F368B37BFC6916F8FBFAFD78" box="[966,1091,693,719]" pageId="19" pageNumber="242" value="1984-04">April 1984</date>
</collectingDate>
, A.D.
<collectorName id="933F537EF368B37BFB2516F8FB6DFD78" box="[1162,1236,693,719]" pageId="19" pageNumber="242">Huryn</collectorName>
,
<specimenCount id="28CCFD21F368B37BFB4F16F8FA92FD78" box="[1248,1323,693,719]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFA9516F8FA29FD78" box="[1338,1424,693,719]" pageId="19" pageNumber="242">ADHC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFF381694FD16FD44" ID-GBIF-Occurrence="2351014878" box="[151,687,729,755]" collectingDate="1984-04-21" collectionCode="ADHC" collectorName="Huryn" country="United States of America" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
but
<date id="4A741068F368B37BFF6E1694FEE7FD44" box="[193,350,729,755]" pageId="19" pageNumber="242" value="1984-04-21">
<collectingDate id="5A30E980F368B37BFF6E1694FEE7FD44" box="[193,350,729,755]" pageId="19" pageNumber="242" value="1984-04-21">21 April 1984</collectingDate>
</date>
, A.D.
<collectorName id="933F537EF368B37BFE0B1694FE57FD44" box="[420,494,729,755]" pageId="19" pageNumber="242">Huryn</collectorName>
,
<specimenCount id="28CCFD21F368B37BFE561694FDFAFD44" box="[505,579,729,755]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFDFE1694FD1EFD44" box="[593,679,729,755]" pageId="19" pageNumber="242">ADHC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFD151694FC1EFCA0" ID-GBIF-Occurrence="2351014796" collectingDate="2002-03-23" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Rattlesnake Springs" latitude="35.1706" location="Nantahala National Forest" longLatPrecision="1" longitude="-83.57741" municipality="Wayah Bald" pageId="19" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<collectingCounty id="D7144E24F368B37BFD151694FC23FD44" box="[698,922,729,755]" pageId="19" pageNumber="242">Rattlesnake Springs</collectingCounty>
, near
<collectingMunicipality id="DE11ACD2F368B37BFC741694FBDEFD44" box="[987,1127,729,755]" pageId="19" pageNumber="242">Wayah Bald</collectingMunicipality>
,
<location id="3B156073F368B37BFBDE1694FA21FD44" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFBDE1694FA21FD44" box="[1137,1432,729,755]" country="United States of America" county="Rattlesnake Springs" latitude="35.1706" longLatPrecision="1" longitude="-83.57741" municipality="Wayah Bald" name="Nantahala National Forest" pageId="19" pageNumber="242" stateProvince="North Carolina">Nantahala National Forest</location>
,
<geoCoordinate id="5BFE506FF368B37BFF3816B1FEBAFCA0" box="[151,259,764,791]" degrees="35.17060" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="35.1706">35.17060</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFEA016B1FE3CFCA0" box="[271,389,764,791]" degrees="83.57741" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-83.57741">-83.57741</geoCoordinate>
,
<date id="4A741068F368B37BFE3F16B0FDF9FCA0" box="[400,576,765,791]" pageId="19" pageNumber="242" value="2002-03-23">
<collectingDate id="5A30E980F368B37BFE3F16B0FDF9FCA0" box="[400,576,765,791]" pageId="19" pageNumber="242" value="2002-03-23">23 March 2002</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFDE316B0FD63FCA0" box="[588,730,765,791]" pageId="19" pageNumber="242">S.A. Grubbs</collectorName>
,
<specimenCount id="28CCFD21F368B37BFD4816B0FC8AFCA0" box="[743,819,765,791]" pageId="19" pageNumber="242" type="male">1 male</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFCEC16B0FC26FCA0" box="[835,927,765,791]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFC1C16B0FB4AFC8C" ID-GBIF-Occurrence="2351014831" collectingDate="2014-05-19" collectionCode="SEM" country="United States of America" county="Mitchell Co." latitude="36.10178" location="Roan Mountain" longLatPrecision="1" longitude="-82.1265" municipality="Holder Creek" pageId="19" pageNumber="242" specimenCount="3" specimenCount-male="3" stateProvince="North Carolina">
<collectingCounty id="D7144E24F368B37BFC1C16B0FBFAFCA0" box="[947,1091,765,791]" pageId="19" pageNumber="242">Mitchell Co.</collectingCounty>
, tributary to
<collectingMunicipality id="DE11ACD2F368B37BFB7916B0FAC8FCA0" box="[1238,1393,765,791]" pageId="19" pageNumber="242">Holder Creek</collectingMunicipality>
, FS 130A,
<location id="3B156073F368B37BFF4C176CFE2CFC8C" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFF4C176CFE2CFC8C" box="[227,405,801,827]" country="United States of America" county="Mitchell Co." latitude="36.10178" longLatPrecision="1" longitude="-82.1265" municipality="Holder Creek" name="Roan Mountain" pageId="19" pageNumber="242" stateProvince="North Carolina">Roan Mountain</location>
,
<geoCoordinate id="5BFE506FF368B37BFE0F176CFDB5FC8C" box="[416,524,801,827]" degrees="36.10178" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="36.10178">36.10178</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFDB8176DFD35FC8C" box="[535,652,800,827]" degrees="82.12650" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.1265">-82.12650</geoCoordinate>
,
<date id="4A741068F368B37BFD38176CFC96FC8C" box="[663,815,801,827]" pageId="19" pageNumber="242" value="2014-05-19">
<collectingDate id="5A30E980F368B37BFD38176CFC96FC8C" box="[663,815,801,827]" pageId="19" pageNumber="242" value="2014-05-19">19 May 2014</collectingDate>
</date>
,
<specimenCount id="28CCFD21F368B37BFC96176CFC36FC8C" box="[825,911,801,827]" pageId="19" pageNumber="242" type="male">3 males</specimenCount>
(WKUC—sacrificed for
<collectionCode id="58DBAE6DF368B37BFB03176CFB53FC8C" box="[1196,1258,801,827]" pageId="19" pageNumber="242">SEM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EA23CF5F368B37BFB51176CFAFEFCE8" ID-GBIF-Occurrence="2351014791" collectingDate="2014-05-19" collectionCode="WKUC" collectorName="A. L. Sheldon" country="United States of America" latitude="36.1006" location="Roan Mountain" longLatPrecision="1" longitude="-82.12862" municipality="Holder Creek" pageId="19" pageNumber="242" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="North Carolina">
<collectingMunicipality id="DE11ACD2F368B37BFB51176CFA21FC8C" box="[1278,1432,801,827]" pageId="19" pageNumber="242">Holder Creek</collectingMunicipality>
, FS 130A,
<location id="3B156073F368B37BFEA51708FE05FCE8" LSID="urn:lsid:plazi:treatment:B66387BEF369B37DFF381604FDBFFD78:3B156073F368B37BFEA51708FE05FCE8" box="[266,444,837,863]" country="United States of America" latitude="36.1006" longLatPrecision="1" longitude="-82.12862" municipality="Holder Creek" name="Roan Mountain" pageId="19" pageNumber="242" stateProvince="North Carolina">Roan Mountain</location>
,
<geoCoordinate id="5BFE506FF368B37BFE681708FD8AFCE8" box="[455,563,837,863]" degrees="36.10060" direction="north" orientation="latitude" pageId="19" pageNumber="242" precision="1" value="36.1006">36.10060</geoCoordinate>
,
<geoCoordinate id="5BFE506FF368B37BFD911708FD0AFCE8" box="[574,691,837,863]" degrees="82.12862" direction="west" orientation="longitude" pageId="19" pageNumber="242" precision="1" value="-82.12862">-82.12862</geoCoordinate>
,
<date id="4A741068F368B37BFD111708FCEFFCE8" box="[702,854,837,863]" pageId="19" pageNumber="242" value="2014-05-19">
<collectingDate id="5A30E980F368B37BFD111708FCEFFCE8" box="[702,854,837,863]" pageId="19" pageNumber="242" value="2014-05-19">19 May 2014</collectingDate>
</date>
,
<collectorName id="933F537EF368B37BFCCF1708FC40FCE8" box="[864,1017,837,863]" pageId="19" pageNumber="242">A.L. Sheldon</collectorName>
,
<specimenCount id="28CCFD21F368B37BFBAB1709FBE4FCE8" box="[1028,1117,836,863]" pageId="19" pageNumber="242" type="male">5 males</specimenCount>
,
<specimenCount id="28CCFD21F368B37BFBC71708FB6DFCE8" box="[1128,1236,837,863]" pageId="19" pageNumber="242" type="female">3 females</specimenCount>
(
<collectionCode id="58DBAE6DF368B37BFB4C1708FA86FCE8" box="[1251,1343,837,863]" pageId="19" pageNumber="242">WKUC</collectionCode>
)
</materialsCitation>
;
</paragraph>
<caption id="6AB56620F368B37BFF381321FDF1F846" httpUri="https://zenodo.org/record/3375808/files/figure.png" pageId="19" pageNumber="242" startId="19.[151,264,1900,1925]" targetBox="[155,1428,920,1873]" targetPageId="19">
<paragraph id="3E7536A8F368B37BFF381321FDF1F846" blockId="19.[151,1436,1900,2033]" pageId="19" pageNumber="242">
<emphasis id="0CBEEABAF368B37BFF381321FEF6F832" bold="true" box="[151,335,1900,1925]" pageId="19" pageNumber="242">FIGURES 6568</emphasis>
.
<taxonomicName id="F9CA4D2BF368B37BFEF11320FDA0F832" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[350,537,1901,1925]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="242" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF368B37BFEF11320FDA0F832" box="[350,537,1901,1925]" italics="true" pageId="19" pageNumber="242">Soyedina sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF368B37BFD8E1320FDE5F832" bold="true" box="[545,604,1901,1925]" pageId="19" pageNumber="242">
<taxonomicNameLabel id="178D57C1F368B37BFD8E1320FDE5F832" box="[545,604,1901,1925]" pageId="19" pageNumber="242" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
6566, USA, North Carolina, seeps along Buncombe Trail; 6768: USA, North Carolina, Lower Creek. 65, male, head and pronotum, dorsal view, 4X; 66, male, abdominal terminalia, dorsal view, 11.5X; 67, male, abdominal terminalia, ventral view, 11.5X; 68, female, abdominal terminalia, ventral view, 5X. ds = dorsal sclerite, vs = ventral sclerite, opl = outer paraproct lobe.
</paragraph>
</caption>
<caption id="6AB56620F36FB37CFF381218FC27F96D" httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="20" pageNumber="243" startId="20.[151,264,1621,1646]" targetBox="[157,1424,175,1601]" targetPageId="20">
<paragraph id="3E7536A8F36FB37CFF381218FC27F96D" blockId="20.[151,1437,1621,1754]" pageId="20" pageNumber="243">
<emphasis id="0CBEEABAF36FB37CFF381218FEF4F9D9" bold="true" box="[151,333,1621,1646]" pageId="20" pageNumber="243">FIGURES 6974</emphasis>
.
<taxonomicName id="F9CA4D2BF36FB37CFEF4121BFDADF9D9" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[347,532,1622,1646]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="243" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36FB37CFEF4121BFDADF9D9" box="[347,532,1622,1646]" italics="true" pageId="20" pageNumber="243">Soyedina sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36FB37CFDB4121BFDEDF9D9" bold="true" box="[539,596,1622,1646]" pageId="20" pageNumber="243">
<taxonomicNameLabel id="178D57C1F36FB37CFDB4121BFDEDF9D9" box="[539,596,1622,1646]" pageId="20" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
69, 71, 7374, USA, North
<taxonomicName id="F9CA4D2BF36FB37CFCDA1218FBA6F9D9" authority="Holder" authorityName="Holder" box="[885,1055,1621,1646]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="243" phylum="Arthropoda" rank="genus">Carolina, Holder</taxonomicName>
Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe.
</paragraph>
</caption>
<paragraph id="3E7536A8F36FB37CFF681344FB82F8DD" blockId="20.[151,1437,1800,2043]" pageId="20" pageNumber="243">
<emphasis id="0CBEEABAF36FB37CFF681344FEF7F894" bold="true" box="[199,334,1801,1827]" pageId="20" pageNumber="243">Etymology.</emphasis>
This species is named in honor of the late Dr. Andrew Sheldon. Andy was a close working colleague of both authors and was responsible for collecting the first series of specimens of this new species. Mountain Man Forestfly is proposed as the common name for this species (
<bibRefCitation id="5A5B4B59F36FB37CFCDF131DFB96F8DD" author="Stark, B. P. &amp; Stewart, K. W. &amp; Szczytko, S. W. &amp; Baumann, R. W. &amp; Kondratieff, B. C." box="[880,1071,1872,1899]" pageId="20" pageNumber="243" refId="ref18007" refString="Stark, B. P., Stewart, K. W., Szczytko, S. W., Baumann, R. W. &amp; Kondratieff, B. C. (2012) Scientific and common names of stoneflies of Nearctic stoneflies (Plecoptera), with corrections and additions to the list. The Caddis Press, Miscellaneous Contributions, 1, 20 pp." type="book" year="2012">
Stark
<emphasis id="0CBEEABAF36FB37CFC1C131CFC5CF8DD" box="[947,997,1872,1898]" italics="true" pageId="20" pageNumber="243">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
<paragraph id="3E7536A8F36FB37DFF681339FEC0FEF4" blockId="20.[151,1437,1800,2043]" lastBlockId="21.[151,1437,152,719]" lastPageId="21" lastPageNumber="244" pageId="20" pageNumber="243">
<emphasis id="0CBEEABAF36FB37CFF681339FEFBF838" bold="true" box="[199,322,1908,1935]" pageId="20" pageNumber="243">Diagnosis.</emphasis>
The combination of the (a) nearly straight-sided sclerotized inner epiproct member (
<figureCitation id="A6F12A2DF36FB37CFAAA1339FA36F839" box="[1285,1423,1908,1934]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="20" pageNumber="243">Figs. 7172</figureCitation>
), (b) apically-rounded ventral sclerites (
<figureCitation id="A6F12A2DF36FB37CFD9013D5FD71F804" box="[575,712,1944,1971]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="20" pageNumber="243">Figs. 6970</figureCitation>
), and (c) robust outer paraproct lobe that is medially-concave and has a prominent, anteriorly-directed triangular flap (
<figureCitation id="A6F12A2DF36FB37CFD7113F1FCDDF860" box="[734,868,1980,2007]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="20" pageNumber="243">Figs. 7678</figureCitation>
) readily distinguishes
<taxonomicName id="F9CA4D2BF36FB37CFBCE13F0FB65F861" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[1121,1244,1980,2007]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="243" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36FB37CFBCE13F0FB65F861" box="[1121,1244,1980,2007]" italics="true" pageId="20" pageNumber="243">S. sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36FB37CFB4C13F0FA85F860" bold="true" box="[1251,1340,1981,2007]" pageId="20" pageNumber="243">
<taxonomicNameLabel id="178D57C1F36FB37CFB4C13F0FA85F860" box="[1251,1340,1981,2007]" pageId="20" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from all other eastern Nearctic
<taxonomicName id="F9CA4D2BF36FB37CFE3A13ADFE45F84D" baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[405,508,2016,2042]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="0CBEEABAF36FB37CFE3A13ADFE45F84D" box="[405,508,2016,2042]" italics="true" pageId="20" pageNumber="243">Soyedina</emphasis>
</taxonomicName>
with some form of an anterior projection on the outer paraproct lobe. Several spe- cies of eastern
<taxonomicName id="F9CA4D2BF36EB37DFE9314D4FE1AFF04" baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[316,419,153,179]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="0CBEEABAF36EB37DFE9314D4FE1AFF04" box="[316,419,153,179]" italics="true" pageId="21" pageNumber="244">Soyedina</emphasis>
</taxonomicName>
have a spur or flap present on the outer paraproct lobe. These include
<taxonomicName id="F9CA4D2BF36EB37DFB0414D4FA8FFF04" box="[1195,1334,153,179]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="amicalola">
<emphasis id="0CBEEABAF36EB37DFB0414D4FA8FFF04" box="[1195,1334,153,179]" italics="true" pageId="21" pageNumber="244">S. amicalola</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFAEA14D4FA2EFF04" box="[1349,1431,152,179]" captionStart="FIGURES 5156" captionStartId="16.[151,264,1906,1931]" captionTargetBox="[186,1405,172,1874]" captionTargetId="figure@16.[180,1408,166,1886]" captionTargetPageId="16" captionText="FIGURES 5156. Soyedina amicalola. 51-56: USA: Georgia, Frogtown Creek. 51, epiproct, ventral view, 800X; 52, epiproct, lateral view, 800X; 53, epiproct apex, apical view, 2000X; 54, epiproct, caudal view, 2000X; 55, epiproct apex, caudal view, 3500X; 56, outer paraproct lobe, lateral view, 500X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375804/files/figure.png" pageId="21" pageNumber="244">Fig. 56</figureCitation>
;
<bibRefCitation id="5A5B4B59F36EB37DFF3814F0FECEFF60" author="Verdone, C. J. &amp; Kondratieff, B. &amp; South, E. &amp; DeWalt, R. E." box="[151,375,189,215]" pageId="21" pageNumber="244" pagination="30 - 49" refId="ref18229" refString="Verdone, C. J., Kondratieff, B., South, E. &amp; DeWalt, R. E. (2017) Studies on the stoneflies of Georgia with the description of a new species of Soyedina Ricker, new state records and an annotated checklist. Illiesia, 13, 30 - 49. https: // doi. org / 10.25031 / 2017 / 13.0 3" type="journal article" year="2017">
Verdone
<emphasis id="0CBEEABAF36EB37DFF5314F3FE97FF60" box="[252,302,189,215]" italics="true" pageId="21" pageNumber="244">et al</emphasis>
. 2017
</bibRefCitation>
, their figs. 5, 8), two distinct phenotypes of
<taxonomicName id="F9CA4D2BF36EB37DFCC514F0FBB6FF60" authorityName="Stark" authorityYear="2017" box="[874,1039,189,215]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis id="0CBEEABAF36EB37DFCC514F0FBB6FF60" box="[874,1039,189,215]" italics="true" pageId="21" pageNumber="244">S. carolinensis</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFBB114F0FB11FF60" box="[1054,1192,189,215]" captionStart="FIGURES 3134" captionStartId="11.[151,264,1126,1151]" captionTargetBox="[177,1406,181,1100]" captionTargetId="figure@11.[171,1416,174,1107]" captionTargetPageId="11" captionText="FIGURES 3134. Soyedina carolinensis. 31: USA, North Carolina, Rattlesnake Springs, outer paraproct lobe, lateral view, 250X; 32: USA, West Virginia, Marsh Fork of Buckhannon River, outer paraproct lobe, lateral view, 190X; 33: USA, Virginia, seep into Lewis Fork, outer paraproct lobe, lateral view, 230X; 34, USA, Maryland, Fishing Creek, outer paraproct lobe, lateral view, 270X." httpUri="https://zenodo.org/record/3375798/files/figure.png" pageId="21" pageNumber="244">Figs. 3334</figureCitation>
),
<taxonomicName id="F9CA4D2BF36EB37DFB1314F0FAE2FF60" authorityName="Baumann &amp; Grubbs" authorityYear="1996" box="[1212,1371,189,215]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="kondratieffi">
<emphasis id="0CBEEABAF36EB37DFB1314F0FAE2FF60" box="[1212,1371,189,215]" italics="true" pageId="21" pageNumber="244">S. kondratieffi</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFAC514F0FF58FF4B" captionStart="FIGURES 5764" captionStartId="17.[151,264,1899,1924]" captionTargetBox="[225,1358,175,1875]" captionTargetId="figure@17.[224,1363,166,1879]" captionTargetPageId="17" captionText="FIGURES 5764. Soyedina kondratieffi. 57, 5960, 6263: USA, Maryland, Mill Run; 59, 61, 64: USA, North Carolina, Ball Creek. 57, epiproct, ventral view, 270X; 58, epiproct, ventral view, 354X; 59, epiproct apex, ventral view, 650X; 60, epiproct apex, dorsal view, 850X; 61, epiproct, lateral view, 347X; 62, epiproct, caudal view, 370X; 63, outer paraproct lobe, lateral view, 300X; 64, outer paraproct lobe, lateral view, 262X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member." httpUri="https://zenodo.org/record/3375806/files/figure.png" pageId="21" pageNumber="244">Figs 6364</figureCitation>
), the more subtle spur exhibited by
<taxonomicName id="F9CA4D2BF36EB37DFDDA14ACFD51FF4C" authorityName="Baumann &amp; Grubbs" authorityYear="1996" box="[629,744,225,251]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="merritti">
<emphasis id="0CBEEABAF36EB37DFDDA14ACFD51FF4C" box="[629,744,225,251]" italics="true" pageId="21" pageNumber="244">S. merritti</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFD5814ACFCF2FF4C" box="[759,843,225,251]" captionStart="FIGURES 3542" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[239,1343,180,1835]" captionTargetId="figure@12.[235,1351,174,1841]" captionTargetPageId="12" captionText="FIGURES 3542. Soyedina merritti. 3538, 4041: USA, Pennsylvania, seep into Powdermill Run; 39, 42: USA, Pennsylvania, seep into Indian Creek. 35, epiproct, ventral view, 150X; 36, epiproct, ventral view, 250X; 37, epiproct apex, ventral view, 650X; 38, epiproct apex, dorsal view, 550X; 39, epiproct, lateral view, 270X; 40, epiproct, caudal view, 220X; 41, outer paraproct lobe, lateral view, 180X; 42, outer paraproct lobe, lateral view, 250X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member." httpUri="https://zenodo.org/record/3375800/files/figure.png" pageId="21" pageNumber="244">Fig. 41</figureCitation>
), and the geographically-consistent shape present on
<taxonomicName id="F9CA4D2BF36EB37DFF381548FE85FEA8" authorityName="Illies" authorityYear="1966" box="[151,316,261,287]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="washingtoni">
<emphasis id="0CBEEABAF36EB37DFF381548FE85FEA8" box="[151,316,261,287]" italics="true" pageId="21" pageNumber="244">S. washingtoni</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFEE41548FE6CFEA8" box="[331,469,260,287]" captionStart="FIGURES 4350" captionStartId="14.[151,264,1865,1890]" captionTargetBox="[237,1348,170,1839]" captionTargetId="figure@14.[231,1357,166,1845]" captionTargetPageId="14" captionText="FIGURES 4350. Soyedina washingtoni. 4346, 49: USA, Maine, Thorndike; 47: USA, Maryland, spring into Elk Lick Run; 48: USA, Pennsylvania, Fourmile Creek; 50: USA, Maryland, Mill Run. 43, epiproct, ventral view, 105X; 43, epiproct, ventral view, 247X; 45, epiproct apex, ventral view, 637X; 46, epiproct apex, dorsal view, 488X; 47, epiproct, lateral view, 370X; 48, epiproct, caudal view, 250X; 49, outer paraproct lobe, lateral view, 166X; 50, outer paraproct lobe, lateral view, 300X; ds = dorsal sclerite, vs = ventral sclerite, im = inner member." httpUri="https://zenodo.org/record/3375802/files/figure.png" pageId="21" pageNumber="244">Figs. 4950</figureCitation>
). Hence, the presence alone of a spur is not diagnostic for species-level determinations of eastern
<taxonomicName id="F9CA4D2BF36EB37DFEA31564FECAFEF4" baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[268,371,297,323]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="0CBEEABAF36EB37DFEA31564FECAFEF4" box="[268,371,297,323]" italics="true" pageId="21" pageNumber="244">Soyedina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3E7536A8F36EB37DFF681500FBE1FE40" blockId="21.[151,1437,152,719]" pageId="21" pageNumber="244">
The outer paraproct lobe of
<taxonomicName id="F9CA4D2BF36EB37DFE551500FDCAFED0" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[506,627,333,359]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36EB37DFE551500FDCAFED0" box="[506,627,333,359]" italics="true" pageId="21" pageNumber="244">S. sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36EB37DFDD61500FD68FED0" bold="true" box="[633,721,333,359]" pageId="21" pageNumber="244">
<taxonomicNameLabel id="178D57C1F36EB37DFDD61500FD68FED0" box="[633,721,333,359]" pageId="21" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A6F12A2DF36EB37DFD701500FCDDFED0" box="[735,868,333,359]" captionStart="FIGURES 7578" captionStartId="21.[151,264,1719,1744]" captionTargetBox="[152,1432,737,1696]" captionTargetId="figure@21.[151,1436,733,1700]" captionTargetPageId="21" captionText="FIGURES 7578. Soyedina sheldoni n. sp. 7576, 78: USA, North Carolina, tributary to Holder Creek 77; USA, North Carolina, Holder Creek. 75, epiproct apex, caudal view, 500X; 76, epiproct apex, dorsal view, 250X; 77, outer paraproct lobe, lateral view, 230X; 78, outer paraproct lobe, lateral view, 230X. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375812/files/figure.png" pageId="21" pageNumber="244">Figs. 7778</figureCitation>
) is similar only to
<taxonomicName id="F9CA4D2BF36EB37DFB9E1500FB02FED0" box="[1073,1211,333,359]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="amicalola">
<emphasis id="0CBEEABAF36EB37DFB9E1500FB02FED0" box="[1073,1211,333,359]" italics="true" pageId="21" pageNumber="244">S. amicalola</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFB661500FAA2FED0" box="[1225,1307,332,359]" captionStart="FIGURES 5156" captionStartId="16.[151,264,1906,1931]" captionTargetBox="[186,1405,172,1874]" captionTargetId="figure@16.[180,1408,166,1886]" captionTargetPageId="16" captionText="FIGURES 5156. Soyedina amicalola. 51-56: USA: Georgia, Frogtown Creek. 51, epiproct, ventral view, 800X; 52, epiproct, lateral view, 800X; 53, epiproct apex, apical view, 2000X; 54, epiproct, caudal view, 2000X; 55, epiproct apex, caudal view, 3500X; 56, outer paraproct lobe, lateral view, 500X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375804/files/figure.png" pageId="21" pageNumber="244">Fig. 56</figureCitation>
;
<bibRefCitation id="5A5B4B59F36EB37DFA8A1500FF4CFE3C" author="Verdone, C. J. &amp; Kondratieff, B. &amp; South, E. &amp; DeWalt, R. E." pageId="21" pageNumber="244" pagination="30 - 49" refId="ref18229" refString="Verdone, C. J., Kondratieff, B., South, E. &amp; DeWalt, R. E. (2017) Studies on the stoneflies of Georgia with the description of a new species of Soyedina Ricker, new state records and an annotated checklist. Illiesia, 13, 30 - 49. https: // doi. org / 10.25031 / 2017 / 13.0 3" type="journal article" year="2017">
Verdone
<emphasis id="0CBEEABAF36EB37DFA271503FF14FE3C" italics="true" pageId="21" pageNumber="244">et al</emphasis>
. 2017
</bibRefCitation>
, their figs. 5, 8), but two epiproct characteristics distinguish these two species. First, the ventral sclerites of
<taxonomicName id="F9CA4D2BF36EB37DFF3815D8FEA8FE18" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[151,273,405,431]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36EB37DFF3815D8FEA8FE18" box="[151,273,405,431]" italics="true" pageId="21" pageNumber="244">S. sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36EB37DFEB815D8FED6FE18" bold="true" box="[279,367,405,431]" pageId="21" pageNumber="244">
<taxonomicNameLabel id="178D57C1F36EB37DFEB815D8FED6FE18" box="[279,367,405,431]" pageId="21" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are typically rounded apically (
<figureCitation id="A6F12A2DF36EB37DFD6215D8FCEAFE07" box="[717,851,405,432]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="21" pageNumber="244">Figs. 6970</figureCitation>
) yet subtriangular on
<taxonomicName id="F9CA4D2BF36EB37DFBEB15D8FB76FE18" box="[1092,1231,405,431]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="amicalola">
<emphasis id="0CBEEABAF36EB37DFBEB15D8FB76FE18" box="[1092,1231,405,431]" italics="true" pageId="21" pageNumber="244">S. amicalola</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFB7215D8FA89FE18" box="[1245,1328,404,431]" captionStart="FIGURES 5156" captionStartId="16.[151,264,1906,1931]" captionTargetBox="[186,1405,172,1874]" captionTargetId="figure@16.[180,1408,166,1886]" captionTargetPageId="16" captionText="FIGURES 5156. Soyedina amicalola. 51-56: USA: Georgia, Frogtown Creek. 51, epiproct, ventral view, 800X; 52, epiproct, lateral view, 800X; 53, epiproct apex, apical view, 2000X; 54, epiproct, caudal view, 2000X; 55, epiproct apex, caudal view, 3500X; 56, outer paraproct lobe, lateral view, 500X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375804/files/figure.png" pageId="21" pageNumber="244">Fig. 51</figureCitation>
). Second, the nr. straight-sided sclerotized inner epiproct member of
<taxonomicName id="F9CA4D2BF36EB37DFC8615F4FC1DFE64" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[809,932,441,467]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36EB37DFC8615F4FC1DFE64" box="[809,932,441,467]" italics="true" pageId="21" pageNumber="244">S. sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36EB37DFC0415F4FBBCFE64" bold="true" box="[939,1029,441,467]" pageId="21" pageNumber="244">
<taxonomicNameLabel id="178D57C1F36EB37DFC0415F4FBBCFE64" box="[939,1029,441,467]" pageId="21" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A6F12A2DF36EB37DFBBB15F4FB23FE64" box="[1044,1178,441,467]" captionStart="FIGURES 6974" captionStartId="20.[151,264,1621,1646]" captionTargetBox="[157,1424,175,1601]" captionTargetId="figure@20.[156,1430,174,1602]" captionTargetPageId="20" captionText="FIGURES 6974. Soyedina sheldoni n. sp. 69, 71, 7374, USA, North Carolina, Holder Creek; 70, 72: USA, North Carolina, tributary to Holder Creek. 69, epiproct, ventral view, 200X; 70, epiproct, ventral view, 270X; 71, epiproct apex, ventral view, 650X; 72, epiproct apex, ventral view, 800X; 73, epiproct apex, lateral view, 270X; 74, epiproct, caudal view, 180X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375810/files/figure.png" pageId="21" pageNumber="244">Figs. 7173</figureCitation>
) is easily distinct from the apically-triangular and caudally-split inner member of
<taxonomicName id="F9CA4D2BF36EB37DFC881590FC0AFE40" box="[807,947,477,503]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="amicalola">
<emphasis id="0CBEEABAF36EB37DFC881590FC0AFE40" box="[807,947,477,503]" italics="true" pageId="21" pageNumber="244">S. amicalola</emphasis>
</taxonomicName>
(
<figureCitation id="A6F12A2DF36EB37DFC6D1590FBF2FE40" box="[962,1099,476,503]" captionStart="FIGURES 5156" captionStartId="16.[151,264,1906,1931]" captionTargetBox="[186,1405,172,1874]" captionTargetId="figure@16.[180,1408,166,1886]" captionTargetPageId="16" captionText="FIGURES 5156. Soyedina amicalola. 51-56: USA: Georgia, Frogtown Creek. 51, epiproct, ventral view, 800X; 52, epiproct, lateral view, 800X; 53, epiproct apex, apical view, 2000X; 54, epiproct, caudal view, 2000X; 55, epiproct apex, caudal view, 3500X; 56, outer paraproct lobe, lateral view, 500X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member, opl = outer paraproct lobe." httpUri="https://zenodo.org/record/3375804/files/figure.png" pageId="21" pageNumber="244">Figs. 5355</figureCitation>
).
</paragraph>
<paragraph id="3E7536A8F36EB37DFF68164CFDBFFD78" blockId="21.[151,1437,152,719]" pageId="21" pageNumber="244">
<emphasis id="0CBEEABAF36EB37DFF68164CFE82FDAC" bold="true" box="[199,315,513,539]" pageId="21" pageNumber="244">Remarks.</emphasis>
<taxonomicName id="F9CA4D2BF36EB37DFEEA164CFDACFDAC" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" box="[325,533,513,539]" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36EB37DFEEA164CFDACFDAC" box="[325,533,513,539]" italics="true" pageId="21" pageNumber="244">Soyedina sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36EB37DFDB0164CFDC5FDAC" bold="true" box="[543,636,513,539]" pageId="21" pageNumber="244">
<taxonomicNameLabel id="178D57C1F36EB37DFDB0164CFDC5FDAC" box="[543,636,513,539]" pageId="21" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is presently known from the southern Appalachian Highlands region in western
<collectingRegion id="FC0EF84AF36EB37DFEBB1668FE04FD88" box="[276,445,549,575]" country="United States of America" name="North Carolina" pageId="21" pageNumber="244">North Carolina</collectingRegion>
(
<figureCitation id="A6F12A2DF36EB37DFE611668FD9AFD88" box="[462,547,549,575]" captionStart="FIGURE 81" captionStartId="24.[151,250,1453,1478]" captionTargetBox="[176,1413,196,1392]" captionTargetId="figure@24.[151,1435,174,1434]" captionTargetPageId="24" captionText="FIGURE 81. Distribution maps of eastern Nearctic Soyedina based on material examined during this study plus valid literature data. 81, S. amicalola, S. kondratieffi, and S. sheldoni n. sp." httpUri="https://zenodo.org/record/3375818/files/figure.png" pageId="21" pageNumber="244">Fig. 81</figureCitation>
). The
<typeStatus id="E171880AF36EB37DFDC41668FD74FD88" box="[619,717,549,575]" pageId="21" pageNumber="244" type="holotype">holotype</typeStatus>
locality and most
<typeStatus id="E171880AF36EB37DFC0F1668FBB9FD88" box="[928,1024,549,575]" pageId="21" pageNumber="244" type="paratype">paratype</typeStatus>
localities are positioned in the High Mountains Level IV (
<date id="4A741068F36EB37DFE3D1604FE00FDD3" box="[402,441,585,612]" pageId="21" pageNumber="244" value="1966-01">66i</date>
) Ecoregion. This region is characterized by disjunct high-elevation Appalachian landscapes with remnant patches of red spruce (
<taxonomicName id="F9CA4D2BF36EB37DFD2E1623FC0EFD30" authority="Sargent, 1898" authorityName="Sargent" authorityYear="1898" box="[641,951,621,647]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="21" pageNumber="244" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="0CBEEABAF36EB37DFD2E1623FCADFD30" box="[641,788,621,647]" italics="true" pageId="21" pageNumber="244">Picea rubens</emphasis>
Sargent, 1898
</taxonomicName>
) and Frasier fir (
<taxonomicName id="F9CA4D2BF36EB37DFBD91620FA2DFD30" authority="Pursh, 1817" authorityName="Pursh" authorityYear="1817" box="[1142,1428,621,647]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="21" pageNumber="244" phylum="Tracheophyta" rank="species" species="fraseri">
<emphasis id="0CBEEABAF36EB37DFBD91620FABFFD30" box="[1142,1286,621,647]" italics="true" pageId="21" pageNumber="244">Abies fraseri</emphasis>
Pursh, 1817
</taxonomicName>
) (
<bibRefCitation id="5A5B4B59F36EB37DFF3116DCFEC0FD1C" author="Griffith, G. E. &amp; Omernik, J. M. &amp; Comstock, J. A. &amp; Schafale, M. P. &amp; McNab, W. H. &amp; Lenat, D. R. &amp; MacPherson, T. F. &amp; Glover, J. B. &amp; Shelburne, V. B." box="[158,377,657,683]" pageId="21" pageNumber="244" refId="ref16386" refString="Griffith, G. E., Omernik, J. M., Comstock, J. A., Schafale, M. P., McNab, W. H., Lenat, D. R., MacPherson, T. F., Glover, J. B. &amp; Shelburne, V. B. (2002) Ecoregions of North Carolina and South Carolina, (color poster with map, descriptive text, summary tables, and photographs): Reston, Virginia, U. S. Geological Survey (map scale 1: 1,500,000)." type="book" year="2002">
Griffith
<emphasis id="0CBEEABAF36EB37DFF5216DFFE97FD1C" box="[253,302,657,683]" italics="true" pageId="21" pageNumber="244">et al</emphasis>
. 2002
</bibRefCitation>
).
<taxonomicName id="F9CA4D2BF36EB37DFE2416DCFCB6FD1C" authority="(Claassen, 1937)" baseAuthorityName="Claassen" baseAuthorityYear="1937" box="[395,783,657,683]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sara">
<emphasis id="0CBEEABAF36EB37DFE2416DCFDF5FD1C" box="[395,588,657,683]" italics="true" pageId="21" pageNumber="244">Paraleuctra sara</emphasis>
(Claassen, 1937)
</taxonomicName>
was the only other stonefly species collected with
<taxonomicName id="F9CA4D2BF36EB37DFAE516DCFF70FD78" authority="Grubbs &amp; Baumann, 2019" authorityName="Grubbs &amp; Baumann" authorityYear="2019" class="Insecta" family="Nemouridae" genus="Soyedina" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="244" phylum="Arthropoda" rank="species" species="sheldoni" status="sp. nov.">
<emphasis id="0CBEEABAF36EB37DFAE516DCFF70FD78" italics="true" pageId="21" pageNumber="244">S. sheldoni</emphasis>
</taxonomicName>
<emphasis id="0CBEEABAF36EB37DFF7F16F8FE93FD78" bold="true" box="[208,298,693,719]" pageId="21" pageNumber="244">
<taxonomicNameLabel id="178D57C1F36EB37DFF7F16F8FE93FD78" box="[208,298,693,719]" pageId="21" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type locality.
</paragraph>
</subSubSection>
</treatment>
</document>