147 lines
14 KiB
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147 lines
14 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFBDFFFAFF6D6D1DCCC1F99C" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cimbicidae Curtis 1825" docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="23" updateTime="1683290186394" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:A148D509FFBDFFFAFF6D6D1DCCC1F99C" httpUri="http://treatment.plazi.org/id/A148D509FFBDFFFAFF6D6D1DCCC1F99C" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection pageId="22" pageNumber="23" type="nomenclature">
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<paragraph blockId="22.[151,597,1608,1635]" box="[151,597,1608,1635]" pageId="22" pageNumber="23">
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<heading bold="true" box="[151,597,1608,1635]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis bold="true" box="[151,597,1608,1635]" pageId="22" pageNumber="23">
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<taxonomicName ID-CoL="892" ID-ENA="85775" authorityName="Curtis" authorityYear="1825" box="[151,294,1608,1634]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">Cimbicidae</taxonomicName>
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subfamily
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<emphasis bold="true" box="[433,597,1608,1635]" italics="true" pageId="22" pageNumber="23">incertae sedis</emphasis>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="22.[151,551,1680,1743]" box="[151,551,1680,1707]" pageId="22" pageNumber="23">
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<heading bold="true" box="[151,551,1680,1707]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis bold="true" box="[151,551,1680,1707]" pageId="22" pageNumber="23">
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Genus and species
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<emphasis bold="true" box="[387,551,1680,1707]" italics="true" pageId="22" pageNumber="23">incertae sedis</emphasis>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="description">
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<paragraph blockId="22.[151,551,1680,1743]" box="[151,238,1717,1743]" pageId="22" pageNumber="23">
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<figureCitation box="[151,234,1717,1743]" captionStart="FIGURE 22" captionStartId="23.[151,250,1165,1189]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-18@23.[151,1436,181,1141]" captionTargetPageId="23" captionText="FIGURE 22. GSC 142844, Cimbicidae subfamily, genus, and species incertae sedis: A, photograph; B, drawing.Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894920" httpUri="https://zenodo.org/record/7894920/files/figure.png" pageId="22" pageNumber="23">Fig. 22</figureCitation>
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.
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</paragraph>
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<paragraph blockId="22.[151,1436,1788,1995]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[151,289,1788,1814]" pageId="22" pageNumber="23">Description</emphasis>
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. Forewing (
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<figureCitation box="[431,515,1789,1815]" captionStart="FIGURE 22" captionStartId="23.[151,250,1165,1189]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-18@23.[151,1436,181,1141]" captionTargetPageId="23" captionText="FIGURE 22. GSC 142844, Cimbicidae subfamily, genus, and species incertae sedis: A, photograph; B, drawing.Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894920" httpUri="https://zenodo.org/record/7894920/files/figure.png" pageId="22" pageNumber="23">Fig. 22</figureCitation>
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) length
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<quantity box="[613,717,1788,1815]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="22" pageNumber="23" unit="mm" value="15.5">15.5 mm</quantity>
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, maximum width
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<quantity box="[927,1018,1788,1815]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="22" pageNumber="23" unit="mm" value="5.4">5.4 mm</quantity>
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, length/width 2.9; membrane lightly infuscate throughout, pterostigma dark, narrow, tapering, narrower than C-R before pterostigma; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r (3r length about 1.6 2r); 2r-rs longer than Rs between 2r-m, 2r-rs, joining cell 3rm before its middle; 2r-m subvertical, short, slightly curved, joins Rs scarcely behind 2mcu; 1-M almost straight, sub-parallel to 1m-cu; 3r-m slightly angular toward base; cell 2mcu with M slightly shorter than 1m-cu; 2m-cu slightly curved toward base; cu-a postfurcal; anal cells meet at crossvein.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7894920" ID-Zenodo-Dep="7894920" box="[151,1437,1165,1190]" httpUri="https://zenodo.org/record/7894920/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1165,1189]" targetBox="[151,1436,181,1135]" targetPageId="23" targetType="figure">
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<paragraph blockId="23.[151,1437,1165,1190]" box="[151,1437,1165,1190]" pageId="23" pageNumber="24">
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<emphasis bold="true" box="[151,286,1165,1189]" pageId="23" pageNumber="24">FIGURE 22.</emphasis>
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GSC 142844,
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<taxonomicName authorityName="Curtis" authorityYear="1825" box="[431,548,1165,1189]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Cimbicidae</taxonomicName>
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subfamily, genus, and species
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<emphasis box="[851,990,1165,1189]" italics="true" pageId="23" pageNumber="24">incertae sedis</emphasis>
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: A, photograph; B, drawing. Scale bar 5 mm.
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</paragraph>
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</caption>
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<subSubSection pageId="23" pageNumber="24" type="discussion">
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<paragraph blockId="23.[151,1437,1235,1622]" pageId="23" pageNumber="24">
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<emphasis bold="true" box="[199,308,1236,1262]" pageId="23" pageNumber="24">Remarks</emphasis>
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. The straight, short 2r-m excludes this specimen from the
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<taxonomicName authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[944,1130,1235,1261]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Cenocimbicinae">Cenocimbicinae</taxonomicName>
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. Its other forewing venation is a mixture of character states diagnostic of modern subfamilies: M+Sc-R junction is close to that of Rs+M as in
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<taxonomicName authorityName="Benson" authorityYear="1938" box="[151,335,1307,1333]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
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and Corynidinae, but the anal veins are separate throughout, joined by a short crossvein 1a as in the
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<taxonomicName authorityName="Curtis" authorityYear="1825" box="[151,284,1343,1369]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="28" phylum="Arthropoda" rank="subFamily" subFamily="Cimbicinae">Cimbicinae</taxonomicName>
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. We refrain from assigning this specimen to a subfamily or erecting a new one based on it, or naming it pending more specimens to clarify its position. We only note that this forewing is very different not only from those of all other Ypresian
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<taxonomicName authorityName="Curtis" authorityYear="1825" box="[392,524,1415,1441]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Cimbicidae</taxonomicName>
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, but from all others. Unfortunately, the wings of the only other known Ypresian non-cenocimbicine, the Green River Formation
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<taxonomicName authority="(Cockerell)" authorityName="Cockerell" authorityYear="1925" baseAuthorityName="Cockerell" baseAuthorityYear="1925" box="[705,1113,1451,1477]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="byrami">
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<emphasis box="[705,974,1451,1477]" italics="true" pageId="23" pageNumber="24">Eopachylosticta byrami</emphasis>
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(Cockerell)
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</taxonomicName>
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are too poorly preserved to compare.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="24" type="materials_examined">
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<paragraph blockId="23.[151,1437,1235,1622]" pageId="23" pageNumber="24">
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<emphasis bold="true" box="[199,303,1523,1549]" pageId="23" pageNumber="24">Material</emphasis>
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.
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<materialsCitation collectingDate="2001-05-19" collectionCode="GSC" pageId="23" pageNumber="24" specimenCode="GSC 142844" specimenCount="1">
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<specimenCode box="[316,464,1524,1550]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="23" pageNumber="24">GSC 142844</specimenCode>
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, a forewing collected at McAbee by SBA (
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<collectionCode box="[937,993,1524,1550]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="23" pageNumber="24">GSC</collectionCode>
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locality V-016800),
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<date box="[1222,1376,1524,1550]" pageId="23" pageNumber="24" value="2001-05-19">
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<collectingDate box="[1222,1376,1524,1550]" pageId="23" pageNumber="24" value="2001-05-19">May 19, 2001</collectingDate>
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</date>
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(
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<figureCitation captionStart="FIGURE 22" captionStartId="23.[151,250,1165,1189]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-18@23.[151,1436,181,1141]" captionTargetPageId="23" captionText="FIGURE 22. GSC 142844, Cimbicidae subfamily, genus, and species incertae sedis: A, photograph; B, drawing.Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894920" httpUri="https://zenodo.org/record/7894920/files/figure.png" pageId="23" pageNumber="24">Fig. 22</figureCitation>
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), in the collections of the
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</materialsCitation>
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<collectionCode box="[472,526,1560,1586]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="23" pageNumber="24">GSC</collectionCode>
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(Vancouver).
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</paragraph>
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<paragraph blockId="23.[151,1437,1235,1622]" box="[199,1209,1595,1622]" pageId="23" pageNumber="24">
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<emphasis bold="true" box="[199,404,1595,1622]" pageId="23" pageNumber="24">Locality and age.</emphasis>
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McAbee,
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<collectingRegion box="[521,717,1595,1621]" country="Canada" name="British Columbia" pageId="23" pageNumber="24">British Columbia</collectingRegion>
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,
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<collectingCountry box="[729,816,1595,1621]" name="Canada" pageId="23" pageNumber="24">Canada</collectingCountry>
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, Hoodoo Face beds; mid-Ypresian.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |