Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands Author Archibald, S. Bruce 0000-0002-4397-2497 Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada bruce.archibald@ubc.ca Author Rasnitsyn, Alexandr P. A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca text Zootaxa 2023 2023-05-04 5278 1 1 38 http://dx.doi.org/10.11646/zootaxa.5278.1.1 journal article 10.11646/zootaxa.5278.1.1 1175-5326 7894826 BE0A07DB-170A-4B72-8178-9B8144C1FBF6 Cimbicidae subfamily incertae sedis Genus and species incertae sedis Fig. 22 . Description . Forewing ( Fig. 22 ) length 15.5 mm , maximum width 5.4 mm , length/width 2.9; membrane lightly infuscate throughout, pterostigma dark, narrow, tapering, narrower than C-R before pterostigma; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r (3r length about 1.6 2r); 2r-rs longer than Rs between 2r-m, 2r-rs, joining cell 3rm before its middle; 2r-m subvertical, short, slightly curved, joins Rs scarcely behind 2mcu; 1-M almost straight, sub-parallel to 1m-cu; 3r-m slightly angular toward base; cell 2mcu with M slightly shorter than 1m-cu; 2m-cu slightly curved toward base; cu-a postfurcal; anal cells meet at crossvein. FIGURE 22. GSC 142844, Cimbicidae subfamily, genus, and species incertae sedis : A, photograph; B, drawing. Scale bar 5 mm. Remarks . The straight, short 2r-m excludes this specimen from the Cenocimbicinae . Its other forewing venation is a mixture of character states diagnostic of modern subfamilies: M+Sc-R junction is close to that of Rs+M as in Pachylostictinae and Corynidinae, but the anal veins are separate throughout, joined by a short crossvein 1a as in the Cimbicinae . We refrain from assigning this specimen to a subfamily or erecting a new one based on it, or naming it pending more specimens to clarify its position. We only note that this forewing is very different not only from those of all other Ypresian Cimbicidae , but from all others. Unfortunately, the wings of the only other known Ypresian non-cenocimbicine, the Green River Formation Eopachylosticta byrami (Cockerell) are too poorly preserved to compare. Material . GSC 142844 , a forewing collected at McAbee by SBA ( GSC locality V-016800), May 19, 2001 ( Fig. 22 ), in the collections of the GSC (Vancouver). Locality and age. McAbee, British Columbia , Canada , Hoodoo Face beds; mid-Ypresian.