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<document ID-CLB-Dataset="297097" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="f9373c90-73c5-4ee4-b0f8-4c371f527d91" ID-ISBN="0-8018-8221-4" checkinTime="1670905209323" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="96F0463750B7A3BA1068BA6839AE18A7" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Muridae.1189-1531.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Thallomys nigricauda" docType="treatment" docVersion="4" id="A2267A57FF8A65DA07E1D2E970F9422A" lastPageNumber="1509" masterDocId="A2267A57FF8A65DA07E1D2E970F9422A" masterDocTitle="Order Rodentia - Family Muridae" masterLastPageNumber="1531" masterPageNumber="1189" pageNumber="1509" updateTime="1716781534040" updateUser="ExternalLinkService">
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<mods:title id="CAF8DE162087D06D2DF5B4623F2E968E">Order Rodentia - Family Muridae</mods:title>
</mods:titleInfo>
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<mods:namePart id="7DD12DF8D8E8402BC86A1B2ECD2A9862">Wilson, Don E.</mods:namePart>
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<mods:namePart id="B50C7794B7503B4C8BDC6760E5D31592">Reeder, DeeAnn</mods:namePart>
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<mods:title id="E362A5A215B3E74324CE515488E46696">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<treatment id="96F0463750B7A3BA1068BA6839AE18A7" ID-DOI="http://doi.org/10.5281/zenodo.11335668" ID-GBIF-Taxon="231540184" ID-Zenodo-Dep="11335668" LSID="urn:lsid:plazi:treatment:96F0463750B7A3BA1068BA6839AE18A7" httpUri="http://treatment.plazi.org/id/96F0463750B7A3BA1068BA6839AE18A7" lastPageNumber="1509" pageNumber="1509" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13001841">
<subSubSection id="9D90D58A70A8673CD934F36BE98C8001" pageNumber="1509" type="nomenclature">
<paragraph id="60E787E4A4AE3A44E883B88D690D0F87" pageNumber="1509">
<taxonomicName id="BAD98E8999879A8CAB02A95310CC3358" ID-CoL="562SD" authority="(Thomas 1882)" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="A27D4FEE9A39F83380C68DE7A8122B57" italics="true" pageNumber="1509">Thallomys nigricauda</emphasis>
(Thomas 1882)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="95864D4EB1FD52B370D3D127F47759CA" pageNumber="1509" type="reference_group">
<paragraph id="6B09380C84C60F7C3330E60643D2D0C3" pageNumber="1509">
<treatmentCitation id="2BADD998707D5FA53C519AFF9BDE0CBE" author="Thomas" authority="Thomas 1882" authorityName="Thomas" authorityYear="1882" class="Mammalia" family="Muridae" genus="Mus" journal="Proc. Zool. Soc. Lond." kingdom="Animalia" order="Rodentia" page="266" pageNumber="1509" phylum="Chordata" rank="species" species="nigricauda" title="Mus nigricauda" volume="1882" year="1882">
<taxonomicName id="B744EB3E11BF175230CCE09FDA26DDCC" authority="Thomas 1882" authorityName="Thomas" authorityYear="1882" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="91813984338F89FE91561507FDF98EB8" italics="true" pageNumber="1509">[Mus] nigricauda</emphasis>
<bibRefCitation id="C5E105BABAE86F3A9B4E0747B56D7699" author="Thomas, O." refId="ref351527" refString="Thomas, O. 1882. On African mongooses. Proceedings of the Zoological Society of London, 1882: 59 - 93." year="1882">Thomas 1882</bibRefCitation>
</taxonomicName>
,
<bibCitation id="EC338C4F64A3D0C15BBC17D16E3AD961" author="Thomas" journal="Proc. Zool. Soc. Lond." pageNumber="1509" pagination="266" title="Mus nigricauda" volume="1882" year="1882">Proc. Zool. Soc. Lond., 1882: 266</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="41BC483B3F8E950C536D1441FA9BBDCF" pageNumber="1509" type="materials_examined">
<paragraph id="61F131C7F20F3A8C5E25AE27067CBAE5" pageNumber="1509">
<emphasis id="A7AD08B15302C10B0DB820F49CC0826C" inLineHeading="true" pageNumber="1509">Type Locality:</emphasis>
<materialsCitation id="AB9D5BD7243C63DB48306DBD48BD3E90" ID-GBIF-Occurrence="4867324433" country="Namibia" pageNumber="1509">
<collectingCountry id="A5C25CD6BBD7DCB7020C54102A799915" name="Namibia">Namibia</collectingCountry>
, Great Namaqualand, Hountop (= Hudup or Hutop) River, west of Gibeon (
<bibRefCitation id="DAD058D2104EE7F5D35DA660A34D4061" author="Meester, J. A. J. &amp; I. L. Rautenbach &amp; N. J. Dippenaar &amp; C. M. Baker" refId="ref236076" refString="Meester, J. A. J., I. L. Rautenbach, N. J. Dippenaar, and C. M. Baker. 1986. Classification of southern African mammals. Transvaal Museum Monograph, 5: 1 - 359." year="1986">Meester et al., 1986</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E531686BCDA54E6A5045A056CF1C7875" pageNumber="1509" type="vernacular_names">
<paragraph id="612009B580792836FCD9192881A60BC5" pageNumber="1509">
<emphasis id="557A79B2EBD48795526DF8EC87427445" inLineHeading="true" pageNumber="1509">Vernacular Names:</emphasis>
<vernacularName id="C3328655EDE210125DC5A28997A68BD9" language="eng" pageNumber="1509">Black-tailed Thallomys</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="8593EA41F75F1E58232CC695587B8080" pageNumber="1509" type="synonym_list">
<paragraph id="2820AB2E294268DFB203935ABFFA6334" pageNumber="1509">
<emphasis id="9BADBB648EBCE9DCF09073519F3C31B8" inLineHeading="true" pageNumber="1509">Synonyms:</emphasis>
<taxonomicName id="AD5480DC64B8F444F4E754768E3D9F05" authority="Roberts 1933" authorityName="Roberts" authorityYear="1933" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="662B808A63A6522E2F17C9BAB63C6CE3" italics="true" pageNumber="1509">Thallomys bradfieldi</emphasis>
Roberts 1933
</taxonomicName>
;
<taxonomicName id="7166F9EB29059B179E26787BB3F8CAD6" authority="(De Winton 1897)" baseAuthorityName="De Winton" baseAuthorityYear="1897" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="damarensis">
<emphasis id="8B74A94E1211F662AA1D562DE638FDAB" italics="true" pageNumber="1509">Thallomys damarensis</emphasis>
(De Winton 1897)
</taxonomicName>
;
<taxonomicName id="BE29327F15792C1759D86A95EB480642" authority="Lundholm 1955" authorityName="Lundholm" authorityYear="1955" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="davisi">
<emphasis id="938E9BF3F0935A44C64DABB0014F998E" italics="true" pageNumber="1509">Thallomys davisi</emphasis>
Lundholm 1955
</taxonomicName>
;
<taxonomicName id="121FAAA9B190D59D5A77E13892C59886" authority="Thomas 1926" authorityName="Thomas" authorityYear="1926" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="herero">
<emphasis id="C474A550FB26A8A27F17E17FDB0D2CD2" italics="true" pageNumber="1509">Thallomys herero</emphasis>
Thomas 1926
</taxonomicName>
;
<taxonomicName id="CE295AFF21FF4098830A754A787BAE01" authority="(Dollman 1911)" baseAuthorityName="Dollman" baseAuthorityYear="1911" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="kalaharicus">
<emphasis id="0A3803353950D727E821DE5C0E128E4D" italics="true" pageNumber="1509">Thallomys kalaharicus</emphasis>
(
<bibRefCitation id="BD97C2AA07ABB1D3F6D063470EA0E739" author="Dollman, G." refId="ref87523" refString="Dollman, G. 1911. On Arvicanthis abyssinicus and allied East-African species, with descriptions of four new forms. Annals and Magazine of Natural History, ser. 8, 8: 334 - 353." year="1911">Dollman 1911</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="E579CA25370AA42102A3EB3933A604A0" authority="Thomas 1926" authorityName="Thomas" authorityYear="1926" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="leuconoe">
<emphasis id="63E08E4540B1B8CC3BC74685C062F705" italics="true" pageNumber="1509">Thallomys leuconoe</emphasis>
Thomas 1926
</taxonomicName>
;
<taxonomicName id="6767757E34929759DFA84600FB7012A9" authority="Roberts 1933" authorityName="Roberts" authorityYear="1933" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="molopensis">
<emphasis id="8E15C83CB4F6CBD8BF9A21675754DA9C" italics="true" pageNumber="1509">Thallomys molopensis</emphasis>
Roberts 1933
</taxonomicName>
;
<taxonomicName id="EC10FA7325E5A602F6E83E388CDB39F7" authority="Thomas and Hinton 1923" authorityName="Thomas and Hinton" authorityYear="1923" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="nitela">
<emphasis id="E48653B4E62CD07E07E48C044C2EFD9B" italics="true" pageNumber="1509">Thallomys nitela</emphasis>
Thomas and
<bibRefCitation id="28BFCF206488CA68ED13A9FD3A972677" author="Hinton, M. A. C." refId="ref153971" refString="Hinton, M. A. C. 1923. On the voles collected by Mr. G. Forrest in Yunnan; with remarks upon the genera Eothenomys and Neodon and upon their allies. Annals and Magazine of Natural History, ser. 9, 11: 145 - 162." year="1923">Hinton 1923</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="74C5F0F3F1CD0194BE9A2E6BF8710307" authority="F. Petter and Beaufort 1960" authorityName="F. Petter and Beaufort" authorityYear="1960" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="quissamae">
<emphasis id="B31C1A53EE767502632D0D079257E684" italics="true" pageNumber="1509">Thallomys quissamae</emphasis>
F.
<bibRefCitation id="9E64F124598B5A8A0FA4DC083972D379" author="Petter, F. &amp; F. de Beaufort" refId="ref284508" refString="Petter, F., and F. de Beaufort. 1960. Description d'une forme nouvelle de rongeur d'Angola, Thallomys damarensis quissamae. Bulletin du Museum National d'Histoire Naturelle, Paris, ser. 2, 32: 269 - 271." year="1960">Petter and Beaufort 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="A1A01321CD6D65CA31C7A53FFE66AC6D" authority="(Ellerman, Morrison-Scott, and Hayman 1953)" baseAuthorityName="Ellerman, Morrison-Scott, and Hayman" baseAuthorityYear="1953" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" pageNumber="1509" phylum="Chordata" rank="species" species="robertsi">
<emphasis id="6FE6A676F033A47F76C95F94403C06C5" italics="true" pageNumber="1509">Thallomys robertsi</emphasis>
(
<bibRefCitation id="604594394F4DAB9C6A45F87DBF793064" author="Ellerman, J. R. &amp; T. C. S. Morrison-Scott &amp; R. W. Hayman" refId="ref93354" refString="Ellerman, J. R., T. C. S. Morrison-Scott, and R. W. Hayman. 1953. Southern African mammals 1758 to 1951: A reclassification. British Museum (Natural History), London, 363 pp." year="1953">Ellerman, Morrison-Scott, and Hayman 1953</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FC8C66B33206DB1B80EEB9CC4AABF2A2" pageNumber="1509" type="distribution">
<paragraph id="96D8754B557545365569C9C86BD03876" pageNumber="1509">
<emphasis id="4B8AB325B6BA5C2B7FAFD1397CD030DD" inLineHeading="true" pageNumber="1509">Distribution:</emphasis>
W and S
<collectingCountry id="5C10CA13926ACBD357A91E6B70DB4D6B" name="Angola">Angola</collectingCountry>
,
<collectingCountry id="B1F097706DD3BC0C158C2F772C669E74" name="Namibia">Namibia</collectingCountry>
, N
<collectingCountry id="835785A6FC7A2483A4425D43AEA42C9F" name="South Africa">South Africa</collectingCountry>
,
<collectingCountry id="559402DACFD05D8285636E8797F24F32" name="Zimbabwe">Zimbabwe</collectingCountry>
, N
<collectingCountry id="4AFD510FFF3A00A39A1A9E33BE4F5BA6" name="Botswana">Botswana</collectingCountry>
, and SE
<collectingCountry id="104D0448A256CAEEFE92F746232CF7F5" name="Zambia">Zambia</collectingCountry>
; northern and eastern limits unknown. The range described here is mapped by
<bibRefCitation id="B01FF35FBD8F8A97D7DE5CC9AE0BD949" author="Taylor, P. J. &amp; I. L. Rautenbach &amp; D. Gordon &amp; K. Sink &amp; P. Lotter" refId="ref349317" refString="Taylor, P. J., I. L. Rautenbach, D. Gordon, K. Sink, and P. Lotter. 1995. Diagnostic morphometrics and southern African distribution of two sibling species of tree rat, Thallomys paedulcus and Thallomys nigricauda (Rodentia: Muridae). Durban Museum Novitates, 20: 49 - 62." year="1995">Taylor et al. (1995)</bibRefCitation>
, who also identified a skull from
<collectingCountry id="CD32177AC3708D2A4AFB50D085314DA6" name="Kenya">Kenya</collectingCountry>
as possibly an example of
<taxonomicName id="B0DC8C03A571B54E73DE03D57ED8EF9E" authorityName="Thomas" authorityYear="1882" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="2040466131EC737898A953EC6BAF6DEA" italics="true" pageNumber="1509">T. nigricauda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CBD59298BB25C5118A7215D63DB01230" pageNumber="1509" type="conservation">
<paragraph id="CB04ADAB282F862246AE755BEDEBB021" pageNumber="1509">
<emphasis id="80FC3E55E29FEE6AC62D77A82B0BC5A5" inLineHeading="true" pageNumber="1509">Conservation:</emphasis>
<collectionCode id="51CC1BB186F7C5F8C3713F968EF34432">IUCN</collectionCode>
Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="B3A76CF4736669AD8CBD0C24957CC8FB" pageNumber="1509" type="discussion">
<paragraph id="1BC472151B60DFFF9877E347458E8546" pageNumber="1509">
<emphasis id="F11259D2F65E1ED197DDB4580F4D3651" inLineHeading="true" pageNumber="1509">Discussion:</emphasis>
<taxonomicName id="8C0CCD1A88FC913F85D7150BCCCE96AF" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="5334C60A78A3F2107E8CF4E8300E6C7C" italics="true" pageNumber="1509">Thallomys nigricauda</emphasis>
</taxonomicName>
and
<taxonomicName id="A3A75E11A93308C4B03F938572614999" baseAuthorityName="Sundevall" baseAuthorityYear="1846" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="paedulcus">
<emphasis id="3016D4FD70FA69BB1478D40BCFF642D3" italics="true" pageNumber="1509">T. paedulcus</emphasis>
</taxonomicName>
are now recognized as occurring in the Southern African Subregion by
<bibRefCitation id="C63823E055D9866B350E49472611D6C1" author="Skinner, J. D. &amp; R. H. N. Smithers" refId="ref329484" refString="Skinner, J. D., and R. H. N. Smithers. 1990. The mammals of the southern African subregion. Second ed. University of Pretoria, Republic of South Africa, 771 pp." year="1990">Skinner and Smithers (1990)</bibRefCitation>
, who also summarized some of the chromosomal, morphological, and ecological distinctions between the two species. They also suggested that
<taxonomicName id="B66478C09F3C6A0ADBDD6F6290A61D13" authorityName="Thomas" authorityYear="1926" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="herero">
<emphasis id="D0DDD4D51BAB1DB9E7CEC29B07F8DABB" italics="true" pageNumber="1509">herero</emphasis>
</taxonomicName>
and
<taxonomicName id="231257A7204DD42C7295E371A0A4DAD4" authorityName="Thomas" authorityYear="1926" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="leuconoe">
<emphasis id="86C518283D13FEDE1E07F110EB1561C6" italics="true" pageNumber="1509">leuconoe</emphasis>
</taxonomicName>
may represent samples of
<taxonomicName id="CC9F82BC83C4475CF6A0FDEF69DAE988" baseAuthorityName="Sundevall" baseAuthorityYear="1846" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="paedulcus">
<emphasis id="096F1278ACAD6965FCCFF2BC67D61F87" italics="true" pageNumber="1509">T. paedulcus</emphasis>
</taxonomicName>
, but we examined the
<typeStatus id="671B8D90CA7A6284C46DB1F4CF778010">holotypes</typeStatus>
and they are examples of
<taxonomicName id="96FC17F6850CEF002815516CC9C78499" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="F87DDF4533AEBA4DA7B82565B54A2A8D" italics="true" pageNumber="1509">T. nigricauda</emphasis>
</taxonomicName>
.
<bibRefCitation id="1422659FC102EC564D9A6BE2D759EA65" author="Musser, G. G. &amp; M. D. Carleton" refId="ref252333" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. Pp. 501 - 755, in: Mammal species of the world, a taxonomic and geographic reference, Second ed. (D. E. Wilson and D. M. Reeder, eds.). Smithsonian Institution Press, Washington D. C., xviii + 1206 pp." year="1993">Musser and Carleton (1993:669)</bibRefCitation>
wrote that &quot;Morphological and geographic definitions of
<taxonomicName id="CF43BAA89DDB4D6113F261EE5F3175A5" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="9A88793224A39E5A7BE4939E65036CC8" italics="true" pageNumber="1509">T. nigricauda</emphasis>
</taxonomicName>
are unsatisfactory. Appreciable geographic variation in body size, length of molar row, pelage coloration, and tail pilosity exists among samples and its significance will have to be assessed by critical systematic revision.&quot; Results of such a study are presented by
<bibRefCitation id="5A1195D53B264240DE27F25FB08B9AB7" author="Taylor, P. J. &amp; I. L. Rautenbach &amp; D. Gordon &amp; K. Sink &amp; P. Lotter" refId="ref349317" refString="Taylor, P. J., I. L. Rautenbach, D. Gordon, K. Sink, and P. Lotter. 1995. Diagnostic morphometrics and southern African distribution of two sibling species of tree rat, Thallomys paedulcus and Thallomys nigricauda (Rodentia: Muridae). Durban Museum Novitates, 20: 49 - 62." year="1995">Taylor et al. (1995)</bibRefCitation>
, who employed multivariate analyses of southern African samples already identified by chromosomal traits as either
<taxonomicName id="41ABF1FAADC1198BAAFB2B34CF481074" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="nigricauda">
<emphasis id="63C232449BA1AEB4A724C75221DD910D" italics="true" pageNumber="1509">T. nigricauda</emphasis>
</taxonomicName>
(2n = 47-50) or
<taxonomicName id="1ABF84F0703F84D1B4785AE489782D2A" baseAuthorityName="Sundevall" baseAuthorityYear="1846" class="Mammalia" family="Muridae" genus="Thallomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="paedulcus">
<emphasis id="06D47158E8FDB4561430696099824C78" italics="true" pageNumber="1509">T. paedulcus</emphasis>
</taxonomicName>
(2n = 43-47) and documented morphometric traits distinguishing samples of the two species, provided clearer resolution of their geographic distributions, and demonstrated broad sympatry in ranges.
<bibRefCitation id="CD46946C6C044446FFF28D0009CADD56" author="Crawford-Cabral, J." refId="ref70924" refString="Crawford-Cabral, J. 1998. The Angolan rodents of the superfamily Muroidea: An account of their distribution. Instituto de Investigacao Cientifica Tropical, Estudos, Ensaios e Documentos, 161: 223 pp." year="1998">Crawford-Cabral (1998)</bibRefCitation>
reviewed and mapped Angolan records, and Taylor (1998) discussed specimens from
<collectingRegion id="2DDFEBF7BD5DB5C9A3912B7E1BBE5471" country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>