166 lines
17 KiB
XML
166 lines
17 KiB
XML
<document id="0E93889604F0A0A1EF3A252A3F2F761C" ID-DOI="10.1080/00222930050020122" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747979" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620672552581" checkinUser="carolina" docAuthor="Gussmann S. M. & Scholtz, C. H." docDate="2000" docId="7B60F030FFB0FFE0011CFACFFE4FDB98" docLanguage="en" docName="JNATHIST.34.1045-1123.pdf.imf" docOrigin="Journal of Natural History 34" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Meridiobolbus natalensis Gussmann S. M. & Scholtz 2000, spec. nov." docType="treatment" docVersion="3" lastPageNumber="1093" masterDocId="87598848FF81FFD20079FFE2FFB0DF5E" masterDocTitle="Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)" masterLastPageNumber="1123" masterPageNumber="1045" pageNumber="1092" updateTime="1698914725550" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="2D535D39E367229BF7D489096BBA2973" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D374A85FB88872D5F74472A0890DE485">
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<mods:title id="CEBBE5B60BDB3182E70EB347F43992C1">Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B360D098D51EFC1E33201294FD46784F" type="personal">
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<mods:roleTerm id="C8B49B9A8EAAB3A0A0C64AD09A4D5E55">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F025AC6D1D05C5F7BEC115F41824D176">Gussmann S. M.</mods:namePart>
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</mods:name>
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<mods:name id="69205FFB59BB01653B5A4858181AB463" type="personal">
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<mods:role id="AB0F8D3BC2E33E9A61B27FAD03CE4348">
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<mods:roleTerm id="4722AF399768C1EFD43BA683B234ED19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0547086C228AFF1FDFFDE87F305AF601">Scholtz, C. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2FBB0E815A980A8397912E75CC503CD5">text</mods:typeOfResource>
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<mods:titleInfo id="D957BAA14391E7470143B0073E8A58AA">
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<mods:title id="84D4022211C2002A5BD494ED519D59E5">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="78E3881EAEE70C1B476615065627104C">
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<mods:date id="BB43EEB8EF585DAFE8B3294A3B40FDD3">2000</mods:date>
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<mods:detail id="121F306AB4BCB1327F250FDFD7F38C0D" type="pubDate">
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<mods:number id="61151087107F54955A97E3D3F79A465F">2000-12-31</mods:number>
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</mods:detail>
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<mods:detail id="B81F9D40B550147D011506AB913237D2" type="volume">
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<mods:number id="B584508328D6727EEC5F3B43525FDFA2">34</mods:number>
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</mods:detail>
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<mods:extent id="32FF03256FE858CBD2011B9F4421A41B" unit="page">
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<mods:start id="35B7142720E5F7FD1C3E657D5468675C">1045</mods:start>
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<mods:end id="BFD2DC0CE10BBBB8152773E3FD11E600">1123</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="71A1C28C52E14B3E48981DF715F5D3DB">journal article</mods:classification>
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<mods:identifier id="DFBD131D650F5B56CCF11DCC1D123F3B" type="DOI">10.1080/00222930050020122</mods:identifier>
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<mods:identifier id="2298FB20C245AEF675CF1406E5009F64" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="6B49F2A3DBFD04A76F163775A7301734" type="Zenodo-Dep">4747979</mods:identifier>
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</mods:mods>
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<treatment id="7B60F030FFB0FFE0011CFACFFE4FDB98" LSID="urn:lsid:plazi:treatment:7B60F030FFB0FFE0011CFACFFE4FDB98" httpUri="http://treatment.plazi.org/id/7B60F030FFB0FFE0011CFACFFE4FDB98" lastPageId="50" lastPageNumber="1093" pageId="49" pageNumber="1092">
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<subSubSection id="BBD312ADFFB0FFE3011CFACFFD4BDA3B" pageId="49" pageNumber="1092" type="nomenclature">
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<paragraph id="F3764126FFB0FFE3011CFACFFD45DA1B" blockId="49.[351,763,1325,1382]" box="[357,757,1325,1349]" pageId="49" pageNumber="1092">
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<heading id="A83EF64AFFB0FFE3011CFACFFD45DA1B" box="[357,757,1325,1349]" centered="true" fontSize="10" level="2" pageId="49" pageNumber="1092" reason="3">
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<taxonomicName id="34C93AA5FFB0FFE3011CFACFFDCADA1B" authority="Gussmann S. M. & Scholtz, 2000" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[357,634,1325,1349]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1092" phylum="Arthropoda" rank="species" species="natalensis" status="sp. nov.">
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<emphasis id="C1BD9D34FFB0FFE3011CFACFFDCADA1B" bold="true" box="[357,634,1325,1349]" italics="true" pageId="49" pageNumber="1092">Meridiobolbus natalensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DA8E204FFFB0FFE302FAFACCFD45DA1B" box="[643,757,1326,1349]" pageId="49" pageNumber="1092" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="F3764126FFB0FFE30126FAAFFD4BDA3B" blockId="49.[351,763,1325,1382]" box="[351,763,1357,1382]" pageId="49" pageNumber="1092">
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(®gures 9a±e, 15e, f,
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<quantity id="3431ECC3FFB0FFE30236FAAFFDC6DA3B" box="[591,630,1357,1381]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="49" pageNumber="1092" unit="g" value="16.0">16g</quantity>
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±i, 18b, 20)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="BBD312ADFFB0FFE00036FA6CFE4FDB98" lastPageId="50" lastPageNumber="1093" pageId="49" pageNumber="1092" type="description">
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<paragraph id="F3764126FFB0FFE30036FA6CFF7DDAFB" blockId="49.[79,1035,1422,1700]" box="[79,205,1422,1445]" pageId="49" pageNumber="1092">
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<emphasis id="C1BD9D34FFB0FFE30036FA6CFF7DDAFB" box="[79,205,1422,1445]" italics="true" pageId="49" pageNumber="1092">Description</emphasis>
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</paragraph>
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<paragraph id="F3764126FFB0FFE30001FA4FFE48DA9B" blockId="49.[79,1035,1422,1700]" box="[120,504,1453,1477]" pageId="49" pageNumber="1092">
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<emphasis id="C1BD9D34FFB0FFE30001FA4FFF4DDA9B" box="[120,253,1453,1477]" italics="true" pageId="49" pageNumber="1092">Body length</emphasis>
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.
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<typeStatus id="2C72FF84FFB0FFE30166FA4FFE3ADA9B" box="[287,394,1453,1477]" pageId="49" pageNumber="1092" type="holotype">Holotype</typeStatus>
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<quantity id="3431ECC3FFB0FFE301EAFA4FFE41DA9B" box="[403,497,1453,1477]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="49" pageNumber="1092" unit="mm" value="10.6">10.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="F3764126FFB0FFE30001FA2FFECBDABB" blockId="49.[79,1035,1422,1700]" box="[120,379,1485,1509]" pageId="49" pageNumber="1092">
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<emphasis id="C1BD9D34FFB0FFE30001FA2FFF73DABB" box="[120,195,1485,1509]" italics="true" pageId="49" pageNumber="1092">Colour</emphasis>
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. Dark brown.
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</paragraph>
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<paragraph id="F3764126FFB0FFE00001FA0FFE6EDE77" blockId="49.[79,1035,1422,1700]" lastBlockId="50.[79,1035,146,1222]" lastPageId="50" lastPageNumber="1093" pageId="49" pageNumber="1092">
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<emphasis id="C1BD9D34FFB0FFE30001FA0FFF03D95B" box="[120,179,1517,1541]" italics="true" pageId="49" pageNumber="1092">Head</emphasis>
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.
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<emphasis id="C1BD9D34FFB0FFE300ACFA0FFDBED95B" box="[213,526,1517,1541]" italics="true" pageId="49" pageNumber="1092">Outer margin of mandible</emphasis>
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rounded (®gure 15e).
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<emphasis id="C1BD9D34FFB0FFE3035AFA0FFCCAD95B" box="[803,890,1517,1541]" italics="true" pageId="49" pageNumber="1092">Labrum</emphasis>
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(®gure 15e) declivous anteriad and posteriad from obsolete transverse carina; anterior margin distinctly sinuate.
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<emphasis id="C1BD9D34FFB0FFE30163F9CFFEC2D91B" box="[282,370,1581,1605]" italics="true" pageId="49" pageNumber="1092">Clypeus</emphasis>
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(®gures 15e, 18b) with strongly elevated anterior transverse carina, straight in dorsal view; with well-de®ned, short diagonal, posteriorly steeply inclined carina, the latter connecting the antero-lateral marginal angle and the respective endpoint of the transverse anterior carina; with distinct lateral carina from endpoint of anterior carina extending to above antennal insertion, there forming a distinct angular elevation, from which extending posteriorly in a slightly curved line past eye-canthus on frons to about middle of the eye.
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<emphasis id="C1BD9D34FFB3FFE00283FF31FC88DFB4" box="[762,824,211,234]" italics="true" pageId="50" pageNumber="1093">Frons</emphasis>
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(®gures 15e, 18b) with strongly elevated, feebly trituberculate transverse carina, positioned in line with postero-lateral angle of eye-canthi.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FED0FF38DD56" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FED0FF55DE17" box="[120,229,306,329]" italics="true" pageId="50" pageNumber="1093">Pronotum</emphasis>
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(®gure 18b). With four very feebly tuberculate swellings in anterior third; area anterior to protuberances strongly declivous and feebly concave; with feebly indented midline, not reaching anterior declivous part or posterior pronotal margin; with ®ne punctures in posterior half, interspersed by some large punctures, especially along midline; ®ne punctation increasingly replaced by large, partly con- ¯uent punctures anteriorly and towards sides; protuberances with ®ne punctures only.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FDF2FD4DDD16" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FDF2FF54DD76" box="[120,228,528,552]" italics="true" pageId="50" pageNumber="1093">Scutellum</emphasis>
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(®gure 15f). Basal width about 1.25Ölength; with some large, shallow punctures in the middle, and scattered small punctures.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FDB2FC36DDF9" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FDB2FF7BDD36" box="[120,203,592,616]" italics="true" pageId="50" pageNumber="1093">Elytron</emphasis>
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(®gure 15f). Striae moderately impressed, their punctures small and separated by two to four puncture diameters; intervals with double punctation (approximately three ®ne and 10 minute punctures between two striae).
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FD53FDECDC79" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FD53FF1DDD96" box="[120,173,689,712]" italics="true" pageId="50" pageNumber="1093">Legs</emphasis>
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.
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<emphasis id="C1BD9D34FFB3FFE000B7FD52FE99DD96" box="[206,297,688,712]" italics="true" pageId="50" pageNumber="1093">Protibia</emphasis>
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six-dentate.
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<emphasis id="C1BD9D34FFB3FFE001C7FD52FD89DD96" box="[446,569,688,712]" italics="true" pageId="50" pageNumber="1093">Metafemur</emphasis>
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with punctation and setation of ventral side as in
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<taxonomicName id="34C93AA5FFB3FFE000BAFD32FEE4DDB6" box="[195,340,720,744]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="1093" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FFB3FFE000BAFD32FEE4DDB6" box="[195,340,720,744]" italics="true" pageId="50" pageNumber="1093">Bolboceratex</emphasis>
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</taxonomicName>
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, ®gure 24e.
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<emphasis id="C1BD9D34FFB3FFE0019BFD32FDFFDDB6" box="[482,591,720,744]" italics="true" pageId="50" pageNumber="1093">Metatibia</emphasis>
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in lateral view only feebly broadening towards apex; with subapical carina distinctly bilobed; with toothlike projections arranged in two rows above subapical carina.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FCCDFDCCDC19" blockId="50.[79,1035,146,1222]" box="[120,636,815,840]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FCCDFEAADC19" box="[120,282,815,839]" italics="true" pageId="50" pageNumber="1093">Male genitalia</emphasis>
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. Aedeagus as in ®gure
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<quantity id="3431ECC3FFB3FFE00241FCCDFDEFDC19" box="[568,607,815,839]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="50" pageNumber="1093" unit="g" value="16.0">16g</quantity>
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±i.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FCADFDFBDCD9" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FCADFF4CDC39" box="[120,252,847,871]" italics="true" pageId="50" pageNumber="1093">Distribution</emphasis>
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(®gure 19). The species is known only from the
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<typeStatus id="2C72FF84FFB3FFE00320FCB2FC3ADC36" box="[857,906,848,872]" pageId="50" pageNumber="1093">type</typeStatus>
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locality in north-eastern
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<collectingRegion id="310D8FC4FFB3FFE00088FC8DFE2FDCD9" box="[241,415,879,903]" country="South Africa" name="KwaZulu-Natal" pageId="50" pageNumber="1093">Kwazulu-Natal</collectingRegion>
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,
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<collectingCountry id="8BDE01B6FFB3FFE001C9FC8DFDF4DCD9" box="[432,580,879,903]" name="South Africa" pageId="50" pageNumber="1093">South Africa</collectingCountry>
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.
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FC72FCD1DCB8" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FC72FE32DCF9" box="[120,386,911,935]" italics="true" pageId="50" pageNumber="1093">Type material examined</emphasis>
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.
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<materialsCitation id="43A14B7BFFB3FFE001DDFC72FCD1DCB8" collectionCode="SAMC" latitude="-27.39" longLatPrecision="742" longitude="32.14" pageId="50" pageNumber="1093" specimenCount="1" typeStatus="holotype">
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<typeStatus id="2C72FF84FFB3FFE001DDFC72FDA7DCF9" box="[420,535,911,935]" pageId="50" pageNumber="1093" type="holotype">HOLOTYPE</typeStatus>
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: l`Mkuzi [
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<geoCoordinate id="96FD27E1FFB3FFE002D8FC6DFD5BDCF9" box="[673,747,911,935]" degrees="27.39" direction="south" orientation="latitude" pageId="50" pageNumber="1093" precision="555" value="-27.39">27.39S</geoCoordinate>
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||
<geoCoordinate id="96FD27E1FFB3FFE0028AFC6DFCF1DCF9" box="[755,833,911,935]" degrees="32.14" direction="east" orientation="longitude" pageId="50" pageNumber="1093" precision="555" value="32.14">32.14E</geoCoordinate>
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], Zululand, Dec., 1945, Light-Trap, DDT No. 492’`near indubium’ [handwriting Andreae, curator ex
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<collectionCode id="95D8D9E3FFB3FFE00036FC2CFF2FDCB8" box="[79,159,974,998]" country="South Africa" name="Iziko Museums of Cape Town" pageId="50" pageNumber="1093" type="Museum">SAMC</collectionCode>
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]/`m’ [incorrect]/`SAM ’ [black print on white label] (
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<collectionCode id="95D8D9E3FFB3FFE00283FC2CFCFDDCB8" box="[762,845,974,998]" country="South Africa" name="Iziko Museums of Cape Town" pageId="50" pageNumber="1093" type="Museum">SAMC</collectionCode>
|
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="F3764126FFB3FFE00001FC0CFE4FDB98" blockId="50.[79,1035,146,1222]" pageId="50" pageNumber="1093">
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<emphasis id="C1BD9D34FFB3FFE00001FC0CFF5CDB58" box="[120,236,1006,1030]" italics="true" pageId="50" pageNumber="1093">Comments</emphasis>
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. Although no other specimens are known besides the
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<typeStatus id="2C72FF84FFB3FFE0030FFC0DFC16DB59" box="[886,934,1007,1031]" pageId="50" pageNumber="1093">type</typeStatus>
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||
, there is no doubt that
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||
<taxonomicName id="34C93AA5FFB3FFE0008DFBECFDB9DB78" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[244,521,1038,1062]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="1093" phylum="Arthropoda" rank="species" species="natalensis">
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<emphasis id="C1BD9D34FFB3FFE0008DFBECFDB9DB78" box="[244,521,1038,1062]" italics="true" pageId="50" pageNumber="1093">Meridiobolbus natalensis</emphasis>
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</taxonomicName>
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is a new species. The aedeagus is distinct but the apical projections of the parameres in dorsal view link it to
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<taxonomicName id="34C93AA5FFB3FFE00356FBCCFC7DDB18" authorityName="Krikken" authorityYear="1984" box="[815,973,1070,1094]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="1093" phylum="Arthropoda" rank="genus">
|
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<emphasis id="C1BD9D34FFB3FFE00356FBCCFC7DDB18" box="[815,973,1070,1094]" italics="true" pageId="50" pageNumber="1093">Meridiobolbus</emphasis>
|
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</taxonomicName>
|
||
. The four pronotal protrusions also place the species with the latter genus rather than with
|
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<taxonomicName id="34C93AA5FFB3FFE000F1FB8CFE9CDBD8" authorityName="Krikken" authorityYear="1984" box="[136,300,1134,1158]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="1093" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FFB3FFE000F1FB8CFE9CDBD8" box="[136,300,1134,1158]" italics="true" pageId="50" pageNumber="1093">Namibiobolbus</emphasis>
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</taxonomicName>
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. In the absence of any additional material it can only be assumed that external sexual dimorphism exists with
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<taxonomicName id="34C93AA5FFB3FFE00225FB6CFCC5DBF8" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[604,885,1166,1190]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="1093" phylum="Arthropoda" rank="species" species="natalensis">
|
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<emphasis id="C1BD9D34FFB3FFE00225FB6CFCC5DBF8" box="[604,885,1166,1190]" italics="true" pageId="50" pageNumber="1093">Meridiobolbus natalensis</emphasis>
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</taxonomicName>
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to the same degree as with the congeneric species.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |