439 lines
56 KiB
XML
439 lines
56 KiB
XML
<document id="4DF1D5DBCD56DD607EB0B9A4D284E365" ID-DOI="10.11606/1807-0205/2020.60.special-issue.04" ID-ISSN="1807-0205" ID-Zenodo-Dep="4615078" ID-ZooBank="27886CA4-CD45-4D73-89BB-24D00342C8C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616008509816" checkinUser="carolina" docAuthor="Vaz-de-Mello, Fernando Z., Nunes, Luis Gabriel de Oliveira Albuquerque & Costa-Silva, Vinícius da" docDate="2020" docId="7058F5779760FFCAFEBF56746435F906" docLanguage="en" docName="PapAvZool.60.special-issue.e202060.04.pdf" docOrigin="Papéis Avulsos de Zoologia 60 (4)" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.04" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Canthon cleidecostae Vaz-de-Mello & Nunes & Costa-Silva 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="8C618D0F9762FFCEFF88572E610FFF95" masterDocTitle="A new species of the genus Canthon Hoffmannsegg (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini) from central Brazil" masterLastPageNumber="6" masterPageNumber="1" pageNumber="3" updateTime="1698889337663" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:titleInfo id="DFAD2D0C6689F6F99DBA45D0C4C3849A">
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<mods:title id="59A7B485E10FC012D1AE920BDAABA52F">A new species of the genus Canthon Hoffmannsegg (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini) from central Brazil</mods:title>
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<mods:namePart id="CD4958E18A1F1A51EDBEA404A2D64E98">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="EAE1F4E1B2563D83E5A200FE4216E99E">Nunes, Luis Gabriel de Oliveira Albuquerque</mods:namePart>
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<mods:namePart id="0FE19A4F7CE9DEB4305D96F2F0E58529">Costa-Silva, Vinícius da</mods:namePart>
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<mods:title id="E744CCB7B8F549F05D1DB89F111E898B">Papéis Avulsos de Zoologia</mods:title>
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<mods:part id="16C5EEC65CE22D10B348D3B848C247FF">
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<mods:date id="55CE9E437FE0589CA3BD77ED57E193DD">2020</mods:date>
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<mods:number id="4F5AD461055B82154BAA5EA1756EC372">2020-03-04</mods:number>
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<mods:number id="75EC29D13CEB866C3F5867630DEA6FE4">60</mods:number>
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<mods:number id="EC8DBA5DE995B1A3E1C9A19DEDDACFDE">4</mods:number>
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<mods:identifier id="399899D496A470BEF048BD3A5486D3E4" type="DOI">10.11606/1807-0205/2020.60.special-issue.04</mods:identifier>
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<treatment id="7058F5779760FFCAFEBF56746435F906" LSID="urn:lsid:plazi:treatment:7058F5779760FFCAFEBF56746435F906" httpUri="http://treatment.plazi.org/id/7058F5779760FFCAFEBF56746435F906" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection id="B0EB17EA9760FFCCFEBF56746314FE06" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="F84E44619760FFCCFEBF56746314FE06" blockId="2.[252,712,218,403]" pageId="2" pageNumber="3">
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<emphasis id="CA8598739760FFCCFEBF56746314FE06" bold="true" pageId="2" pageNumber="3">
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<taxonomicName id="3FF13FE29760FFCCFEBF56746321FEE6" ID-CoL="QP6V" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[311,558,346,371]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
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<emphasis id="CA8598739760FFCCFEBF56746321FEE6" bold="true" box="[311,558,346,371]" italics="true" pageId="2" pageNumber="3">Canthon cleidecostae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D1B625089760FFCCFDBB56746383FEE6" box="[563,652,346,371]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="60CA58E49760FFCCFE275654631CFE06" box="[431,531,378,403]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[170,229,2106,2131]" captionStartId-1="3.[94,153,2042,2067]" captionStartId-2="4.[170,229,2074,2099]" captionTargetBox-0="[174,1492,830,2073]" captionTargetBox-1="[94,1413,882,2000]" captionTargetBox-2="[171,790,1554,2043]" captionTargetId-0="figure-286@2.[170,1493,829,2076]" captionTargetId-1="figure-360@3.[94,1417,878,2012]" captionTargetId-2="figure-544@4.[170,793,1553,2044]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." captionText-1="Figure 2. Details of structure in Canthon cleidecostae sp. nov.: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm." captionText-2="Figure 3. Metafemur, ventral view of (A) Canthon cleidecostae; and (B) Canthon ibarragrassoi.Scale bar:1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4615080" figureDoi-1="http://doi.org/10.5281/zenodo.4615084" figureDoi-2="http://doi.org/10.5281/zenodo.4615090" httpUri-0="https://zenodo.org/record/4615080/files/figure.png" httpUri-1="https://zenodo.org/record/4615084/files/figure.png" httpUri-2="https://zenodo.org/record/4615090/files/figure.png" pageId="2" pageNumber="3">Figs. 1-3</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="B0EB17EA9760FFCCFF22569460BAFC86" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="F84E44619760FFCCFF22569460BAFC86" blockId="2.[170,795,442,787]" pageId="2" pageNumber="3">
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<emphasis id="CA8598739760FFCCFF2256946059FE46" bold="true" box="[170,342,442,467]" pageId="2" pageNumber="3">Type material:</emphasis>
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<materialsCitation id="48994E3C9760FFCCFED65694608AFDE6" collectingDate="2013-03-13" collectionCode="CEMT" country="Brazil" latitude="-13.236112" location="First" longLatPrecision="21" longitude="-47.15528" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Goias" typeStatus="holotype">
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<typeStatus id="274AFAC39760FFCCFED6569460D8FE46" box="[350,471,442,467]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
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||
(
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<specimenCount id="EEF78FE89760FFCCFE6F569460F5FE44" box="[487,506,442,465]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
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<collectionCode id="9EE0DCA49760FFCCFD8A56946347FE46" box="[514,584,442,467]" pageId="2" pageNumber="3">CEMT</collectionCode>
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):
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<location id="FD2E12BA9760FFCCFDD056946388FE46" LSID="urn:lsid:plazi:treatment:7058F5779760FFCAFEBF56746435F906:FD2E12BA9760FFCCFDD056946388FE46" box="[600,647,442,467]" country="Brazil" latitude="-13.236112" longLatPrecision="21" longitude="-47.15528" name="First" pageId="2" pageNumber="3" stateProvince="Goias">First</location>
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label [white, typeset]: “
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<collectingCountry id="80E604F19760FFCCFE9556F4607EFE66" box="[285,369,474,499]" name="Brazil" pageId="2" pageNumber="3">BRAZIL</collectingCountry>
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:
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<collectingRegion id="3A358A839760FFCCFE0956F460CEFE66" box="[385,449,474,499]" country="Brazil" name="Goias" pageId="2" pageNumber="3">Goiás</collectingRegion>
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, Monte / Alegre de
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<collectingRegion id="3A358A839760FFCCFD4856F4620FFE66" box="[704,768,474,499]" country="Brazil" name="Goias" pageId="2" pageNumber="3">Goiás</collectingRegion>
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, /
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<geoCoordinate id="9DC522A69760FFCCFF2256D4602AFD86" box="[170,293,506,532]" degrees="13" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="10" value="-13.236112">13°14′10″S</geoCoordinate>
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,
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<geoCoordinate id="9DC522A69760FFCCFEA756D460BDFD86" box="[303,434,506,532]" degrees="47" direction="west" minutes="09" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="19" value="-47.15528">47°09′19″W</geoCoordinate>
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, /
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<date id="8C4F62A19760FFCCFE4456D46330FD86" box="[460,575,506,531]" pageId="2" pageNumber="3" value="2013-03-13">
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<collectingDate id="9C0B9B499760FFCCFE4456D46330FD86" box="[460,575,506,531]" pageId="2" pageNumber="3" value="2013-03-13">13.iii.2013</collectingDate>
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</date>
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,
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<emphasis id="CA8598739760FFCCFDC156D461EBFDA6" italics="true" pageId="2" pageNumber="3">
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||
<taxonomicName id="3FF13FE29760FFCCFDC156D463ACFD86" box="[585,675,506,531]" class="Mammalia" family="Caviidae" genus="Kerodon" kingdom="Animalia" order="Rodentia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">Kerodon</taxonomicName>
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/
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||
<taxonomicName id="3FF13FE29760FFCCFD3056D46215FD86" authorityName="Moojen, Locks & Langguth" authorityYear="1997" box="[696,794,506,531]" class="Mammalia" family="Caviidae" genus="Kerodon" kingdom="Animalia" order="Rodentia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="acrobata">acrobata</taxonomicName>
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feces,
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||
</emphasis>
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||
Alexandre / leg.”. Second label [red, handwritten]: “
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<taxonomicName id="3FF13FE29760FFCCFF3C55146093FDC6" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[180,412,570,595]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae">
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<emphasis id="CA8598739760FFCCFF3C55146093FDC6" box="[180,412,570,595]" italics="true" pageId="2" pageNumber="3">Canthon cleidecostae</emphasis>
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||
</taxonomicName>
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||
/ Vaz-de-Mello, Nunes / & Costa- Silva /
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||
<typeStatus id="274AFAC39760FFCCFE8B55746072FDE6" box="[259,381,602,627]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
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”
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||
</materialsCitation>
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.
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||
<typeStatus id="274AFAC39760FFCCFE1D55746309FDE6" box="[405,518,602,627]" pageId="2" pageNumber="3" type="paratype">PARATYPE</typeStatus>
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(
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||
<specimenCount id="EEF78FE89760FFCCFD9655746332FDE4" box="[542,573,602,627]" pageId="2" pageNumber="3" type="male">1♂</specimenCount>
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||
<collectionCode id="9EE0DCA49760FFCCFDC25574639FFDE6" box="[586,656,602,627]" pageId="2" pageNumber="3">CEMT</collectionCode>
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): First label [white, typeset]:“
|
||
<collectingCountry id="80E604F19760FFCCFEEE555460B5FD06" box="[358,442,634,659]" name="Brazil" pageId="2" pageNumber="3">BRAZIL</collectingCountry>
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||
:
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<collectingRegion id="3A358A839760FFCCFE4A5554630DFD06" box="[450,514,634,659]" country="Brazil" name="Goias" pageId="2" pageNumber="3">Goiás</collectingRegion>
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, Monte / Alegre de
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<collectingRegion id="3A358A839760FFCCFD5F55546218FD06" box="[727,791,634,659]" country="Brazil" name="Goias" pageId="2" pageNumber="3">Goiás</collectingRegion>
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, /
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<geoCoordinate id="9DC522A69760FFCCFF3655B46036FD26" box="[190,313,666,692]" degrees="13" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="10" value="-13.236112">13°14′10″S</geoCoordinate>
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,
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<geoCoordinate id="9DC522A69760FFCCFECF55B460C5FD26" box="[327,458,666,692]" degrees="47" direction="west" minutes="09" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="19" value="-47.15528">47°09′19″W</geoCoordinate>
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, /
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<date id="8C4F62A19760FFCCFE6455B46351FD26" box="[492,606,666,691]" pageId="2" pageNumber="3" value="2013-03-13">13.iii.2013</date>
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,
|
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<emphasis id="CA8598739760FFCCFDE455B46012FD46" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="3FF13FE29760FFCCFDE455B463C9FD26" box="[620,710,666,691]" class="Mammalia" family="Caviidae" genus="Kerodon" kingdom="Animalia" order="Rodentia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">Kerodon</taxonomicName>
|
||
/
|
||
<taxonomicName id="3FF13FE29760FFCCFD6A55B461D3FD46" authorityName="Moojen, Locks & Langguth" authorityYear="1997" class="Mammalia" family="Caviidae" genus="Kerodon" kingdom="Animalia" order="Rodentia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="acrobata">acrobata</taxonomicName>
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||
feces,
|
||
</emphasis>
|
||
Alexandre / leg.”. Second label [yellow, handwritten]: “
|
||
<taxonomicName id="3FF13FE29760FFCCFE9D55F460F4FD66" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[277,507,730,755]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae">
|
||
<emphasis id="CA8598739760FFCCFE9D55F460F4FD66" box="[277,507,730,755]" italics="true" pageId="2" pageNumber="3">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
/ Vaz-de-Mello, Nunes / & Costa-Silva /
|
||
<typeStatus id="274AFAC39760FFCCFEB055D460A4FC86" box="[312,427,762,787]" pageId="2" pageNumber="3" type="paratype">PARATYPE</typeStatus>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9760FFCCFCED57F464A7FE46" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="F84E44619760FFCCFCED57F464A7FE46" blockId="2.[869,1493,218,467]" pageId="2" pageNumber="3">
|
||
<emphasis id="CA8598739760FFCCFCED57F462EEFF66" bold="true" box="[869,993,218,243]" pageId="2" pageNumber="3">Diagnosis:</emphasis>
|
||
<taxonomicName id="3FF13FE29760FFCCFC6557F465D8FF66" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[1005,1239,218,243]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
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||
<emphasis id="CA8598739760FFCCFC6557F465D8FF66" box="[1005,1239,218,243]" italics="true" pageId="2" pageNumber="3">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089760FFCCFB6A57F46433FF66" box="[1250,1340,218,243]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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||
is most similar to
|
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<emphasis id="CA8598739760FFCCFC3D57D465A1FE86" box="[949,1198,250,275]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="3FF13FE29760FFCCFC3D57D465A5FE86" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[949,1194,250,275]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ibarragrassoi">Canthon ibarragrassoi</taxonomicName>
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||
,
|
||
</emphasis>
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||
with which it shares the metafemora with a sinuous carina close to the anterior margin (
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<figureCitation id="60CA58E49760FFCCFC4C5614650BFEC6" box="[964,1028,314,339]" captionStart="Figure 3" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[171,790,1554,2043]" captionTargetId="figure-544@4.[170,793,1553,2044]" captionTargetPageId="4" captionText="Figure 3. Metafemur, ventral view of (A) Canthon cleidecostae; and (B) Canthon ibarragrassoi.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615090" httpUri="https://zenodo.org/record/4615090/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
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). Furthermore, it can be easily separated by the clypeal teeth, which are U-shaped in
|
||
<taxonomicName id="3FF13FE29760FFCCFADA56746288FE06" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739760FFCCFADA56746288FE06" italics="true" pageId="2" pageNumber="3">C. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089760FFCCFC05565462EFFE06" box="[909,992,378,403]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and V-shaped in
|
||
<emphasis id="CA8598739760FFCCFB1356546448FE06" box="[1179,1351,378,403]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="3FF13FE29760FFCCFB135654644CFE06" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[1179,1347,378,403]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ibarragrassoi">C. ibarragrassoi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The colour is also characteristic while
|
||
<taxonomicName id="3FF13FE29760FFCCFBF856B46403FE26" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[1136,1292,410,435]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739760FFCCFBF856B46403FE26" box="[1136,1292,410,435]" italics="true" pageId="2" pageNumber="3">C. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089760FFCCFA9956B4646CFE26" box="[1297,1379,410,435]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is metallic green and
|
||
<taxonomicName id="3FF13FE29760FFCCFC545694658EFE46" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[988,1153,442,467]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ibarragrassoi">
|
||
<emphasis id="CA8598739760FFCCFC545694658EFE46" box="[988,1153,442,467]" italics="true" pageId="2" pageNumber="3">C. ibarragrassoi</emphasis>
|
||
</taxonomicName>
|
||
present black colouration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9760FFCAFCED56D46084FEC6" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="F84E44619760FFCDFCED56D4644DFCC6" blockId="2.[869,1494,506,787]" lastBlockId="3.[794,1419,218,851]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="CA8598739760FFCCFCED56D465CDFD86" bold="true" box="[869,1218,506,531]" pageId="2" pageNumber="3">
|
||
Description (
|
||
<typeStatus id="274AFAC39760FFCCFC7556D46561FD86" box="[1021,1134,506,531]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
, Male):
|
||
</emphasis>
|
||
Dorsal habitus (
|
||
<figureCitation id="60CA58E49760FFCCFAF356D464C6FD86" box="[1403,1481,506,531]" captionStart="Figure 1" captionStartId="2.[170,229,2106,2131]" captionTargetBox="[174,1492,830,2073]" captionTargetId="figure-286@2.[170,1493,829,2076]" captionTargetPageId="2" captionText="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615080" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
|
||
). Body length (except the head)
|
||
<quantity id="3F09E9849760FFCCFB395534641DFDA6" box="[1201,1298,538,563]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" pageId="2" pageNumber="3" unit="mm" value="7.55">7.55 mm</quantity>
|
||
.
|
||
<emphasis id="CA8598739760FFCCFA9255346450FDA6" bold="true" box="[1306,1375,538,563]" pageId="2" pageNumber="3">Head:</emphasis>
|
||
dorsal surface slightly concave with green alveolar microsculpture; posterior half with finely spaced punctation and anteri- or half with superficial rugosities. Two triangular clypeal teeth, with rounded apex, separated by a U-shaped emargination (
|
||
<figureCitation id="60CA58E49760FFCCFB8D5594655EFD46" box="[1029,1105,698,723]" captionStart="Figure 1" captionStartId="2.[170,229,2106,2131]" captionTargetBox="[174,1492,830,2073]" captionTargetId="figure-286@2.[170,1493,829,2076]" captionTargetPageId="2" captionText="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615080" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="2" pageNumber="3">Fig. 1D</figureCitation>
|
||
). Clypeal margin and lateral margin of head with yellow setae emanating from underneath. Clypeo-genal suture present and oblique until close to eyes.
|
||
<emphasis id="CA8598739761FFCDFF1657F46013FF66" bold="true" box="[158,284,218,243]" pageId="3" pageNumber="4">Pronotum:</emphasis>
|
||
pronotal disc smooth, metallic green, bright with green alveolar microsculpture. Anterior angles acute, pointed forward (
|
||
<figureCitation id="60CA58E49761FFCDFE3156346302FEA6" box="[441,525,282,307]" captionStart="Figure 1" captionStartId="2.[170,229,2106,2131]" captionTargetBox="[174,1492,830,2073]" captionTargetId="figure-286@2.[170,1493,829,2076]" captionTargetPageId="2" captionText="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615080" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
|
||
). Lateral margin smooth with two little teeth and brief emargination between them (more visible in lateral view).
|
||
<emphasis id="CA8598739761FFCDFDB3567463C1FEE6" bold="true" box="[571,718,346,371]" pageId="3" pageNumber="4">Hypomeron:</emphasis>
|
||
dark green, with coriarious microsculpture, sparse yellowish setae anteriorly, and thin transverse carina at centre not touching lateral margin; anterior region slightly concave (
|
||
<figureCitation id="60CA58E49761FFCDFF4F56F4601DFE66" box="[199,274,474,499]" captionStart="Figure 1" captionStartId="2.[170,229,2106,2131]" captionTargetBox="[174,1492,830,2073]" captionTargetId="figure-286@2.[170,1493,829,2076]" captionTargetPageId="2" captionText="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615080" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="3" pageNumber="4">Fig. 1B</figureCitation>
|
||
).
|
||
<emphasis id="CA8598739761FFCDFEAC56F4607FFE66" bold="true" box="[292,368,474,499]" pageId="3" pageNumber="4">Elytra:</emphasis>
|
||
surface with dark green colour, less bright than pronotum; with fine micropuncture. Striae narrow and poorly visible, with punctation; interstriae wide, flat.
|
||
<emphasis id="CA8598739761FFCDFE8755146042FDC6" bold="true" box="[271,333,570,595]" pageId="3" pageNumber="4">Legs:</emphasis>
|
||
protibiae with three main teeth on outer edge and crenulated until the base; protibial spur flattened dorsoventrally, bifid, asymmetrical (
|
||
<figureCitation id="60CA58E49761FFCDFDE6555463CAFD06" box="[622,709,634,659]" captionStart="Figure 2" captionStartId="3.[94,153,2042,2067]" captionTargetBox="[94,1413,882,2000]" captionTargetId="figure-360@3.[94,1417,878,2012]" captionTargetPageId="3" captionText="Figure 2. Details of structure in Canthon cleidecostae sp. nov.: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4615084" httpUri="https://zenodo.org/record/4615084/files/figure.png" pageId="3" pageNumber="4">Fig. 2A</figureCitation>
|
||
). Mesofemora with ventral side lacking anterior and posterior margin. Metafemora subclaviform, with a sinuous carina (
|
||
<figureCitation id="60CA58E49761FFCDFF2655F461F3FD66" box="[174,252,730,755]" captionStart="Figure 3" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[171,790,1554,2043]" captionTargetId="figure-544@4.[170,793,1553,2044]" captionTargetPageId="4" captionText="Figure 3. Metafemur, ventral view of (A) Canthon cleidecostae; and (B) Canthon ibarragrassoi.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615090" httpUri="https://zenodo.org/record/4615090/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||
, white setae); yellow setae on the anterior edge and close to apex; posterior edge with lobe comprising two-thirds apically (
|
||
<figureCitation id="60CA58E49761FFCDFE1C543460E9FCA6" box="[404,486,794,819]" captionStart="Figure 3" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[171,790,1554,2043]" captionTargetId="figure-544@4.[170,793,1553,2044]" captionTargetPageId="4" captionText="Figure 3. Metafemur, ventral view of (A) Canthon cleidecostae; and (B) Canthon ibarragrassoi.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615090" httpUri="https://zenodo.org/record/4615090/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||
).
|
||
<emphasis id="CA8598739761FFCDFE715434637AFCA6" bold="true" box="[505,629,794,819]" pageId="3" pageNumber="4">Abdomen:</emphasis>
|
||
surface with alveolar microsculpture; dark green. Ventrites with tufts of small yellow setae on each side laterally and medially glabrous (
|
||
<figureCitation id="60CA58E49761FFCDFC3B57D462F1FE86" box="[947,1022,250,275]" captionStart="Figure 1" captionStartId="2.[170,229,2106,2131]" captionTargetBox="[174,1492,830,2073]" captionTargetId="figure-286@2.[170,1493,829,2076]" captionTargetPageId="2" captionText="Figure 1. Holotype of Canthon cleidecostae sp.nov.:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.4615080" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="3" pageNumber="4">Fig. 1B</figureCitation>
|
||
).
|
||
<emphasis id="CA8598739761FFCDFB9957D46587FE86" bold="true" box="[1041,1160,250,275]" pageId="3" pageNumber="4">Pygidium:</emphasis>
|
||
as wide as long; bright green without punctation. Basal margin expanded, forming strongV-shaped carina between pygidium and propygidium.
|
||
<emphasis id="CA8598739761FFCDFCFB567462F8FEE6" bold="true" box="[883,1015,346,371]" pageId="3" pageNumber="4">Parameres:</emphasis>
|
||
lateral view: symmetrical; apex truncate (
|
||
<figureCitation id="60CA58E49761FFCDFCDB56546290FE06" box="[851,927,378,403]" captionStart="Figure 2" captionStartId="3.[94,153,2042,2067]" captionTargetBox="[94,1413,882,2000]" captionTargetId="figure-360@3.[94,1417,878,2012]" captionTargetPageId="3" captionText="Figure 2. Details of structure in Canthon cleidecostae sp. nov.: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4615084" httpUri="https://zenodo.org/record/4615084/files/figure.png" pageId="3" pageNumber="4">Fig. 2B</figureCitation>
|
||
). Superior edge straight and inferior concave, never sinuous. Apical-laterally edge with bulge at centre. Dorsal view: membranous portion separating parameres from base to apex.
|
||
<emphasis id="CA8598739761FFCDFC6456F46574FE66" bold="true" box="[1004,1147,474,499]" pageId="3" pageNumber="4">Internal sac:</emphasis>
|
||
peripheral fronto-lateral sclerite (FLP) connected to of the axial sclerite (Ax), both sclerites forming an apical complex, in which Ax forms a central semiduct (
|
||
<figureCitation id="60CA58E49761FFCDFC5755146526FDC6" box="[991,1065,570,595]" captionStart="Figure 2" captionStartId="3.[94,153,2042,2067]" captionTargetBox="[94,1413,882,2000]" captionTargetId="figure-360@3.[94,1417,878,2012]" captionTargetPageId="3" captionText="Figure 2. Details of structure in Canthon cleidecostae sp. nov.: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4615084" httpUri="https://zenodo.org/record/4615084/files/figure.png" pageId="3" pageNumber="4">Fig. 2E</figureCitation>
|
||
, Ax [black arrow]) and FLP completes the circunference in the semiduct (Ax) (like a roof), base and apex of all the complexes of equal size, without tapered region; subaxial sclerite (SA), about the same size as other sclerites, covering ventrally the FLP + Ax complex; peripheral medial sclerite (MP) flat, rolled, with the basal part more acute (in relation to the apex), lateral in relation to FLP + Ax; superior right peripheral sclerite (SRP) with lateral angular prolongation (
|
||
<figureCitation id="60CA58E49761FFCDFB455414641DFCC6" box="[1229,1298,826,851]" captionStart="Figure 2" captionStartId="3.[94,153,2042,2067]" captionTargetBox="[94,1413,882,2000]" captionTargetId="figure-360@3.[94,1417,878,2012]" captionTargetPageId="3" captionText="Figure 2. Details of structure in Canthon cleidecostae sp. nov.: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4615084" httpUri="https://zenodo.org/record/4615084/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
|
||
E-F).
|
||
</paragraph>
|
||
<caption id="AC8E14E99760FFCCFF225F1465E1F7C6" ID-DOI="http://doi.org/10.5281/zenodo.4615080" ID-Zenodo-Dep="4615080" box="[170,1262,2106,2131]" httpUri="https://zenodo.org/record/4615080/files/figure.png" pageId="2" pageNumber="3" startId="2.[170,229,2106,2131]" targetBox="[174,1492,830,2073]" targetPageId="2">
|
||
<paragraph id="F84E44619760FFCCFF225F1465E1F7C6" blockId="2.[170,1262,2106,2131]" box="[170,1262,2106,2131]" pageId="2" pageNumber="3">
|
||
Figure 1. Holotype of
|
||
<taxonomicName id="3FF13FE29760FFCCFEEA5F146306F7C6" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[354,521,2106,2131]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739760FFCCFEEA5F146306F7C6" bold="true" box="[354,521,2106,2131]" italics="true" pageId="2" pageNumber="3">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089760FFCCFD855F146342F7C6" box="[525,589,2106,2131]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
|
||
:(A) dorsal;(B) ventral;and (C) lateral view;(D) head,dorsal view.Scale bar:1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AC8E14E99761FFCDFFD650D4645AF7C6" ID-DOI="http://doi.org/10.5281/zenodo.4615084" ID-Zenodo-Dep="4615084" httpUri="https://zenodo.org/record/4615084/files/figure.png" pageId="3" pageNumber="4" startId="3.[94,153,2042,2067]" targetBox="[94,1413,882,2000]" targetPageId="3">
|
||
<paragraph id="F84E44619761FFCDFFD650D4645AF7C6" blockId="3.[94,1418,2042,2131]" pageId="3" pageNumber="4">
|
||
Figure 2. Details of structure in
|
||
<taxonomicName id="3FF13FE29761FFCDFEF950D46316F786" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[369,537,2042,2067]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739761FFCDFEF950D46316F786" bold="true" box="[369,537,2042,2067]" italics="true" pageId="3" pageNumber="4">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089761FFCDFD9750D4636EF786" box="[543,609,2042,2067]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: (A) male protibial apex, dorsal; Male genitalia: (B) aedeagus, lateral; internal sac: (C) dorsal and (D) ventral views;internal sac apical and parietal sclerites:(E) dorsal and (F) ventral views [Ax= axial sclerite;FLP = fronto-lateral peripheric sclerite;MP = medial peripheric sclerite;SA = subaxial sclerite;SRP = superior peripheric sclerite].Black arrow showing the semiduct.Scale bars:A-D = 1 mm; E and F = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F84E44619766FFCAFF2257F46084FEC6" blockId="4.[170,794,218,339]" pageId="4" pageNumber="5">
|
||
<emphasis id="CA8598739766FFCAFF2257F460D8FF66" bold="true" box="[170,471,218,243]" pageId="4" pageNumber="5">Morphological variation:</emphasis>
|
||
The variation between the two specimens of the type series is centred on the body length, which varied from
|
||
<quantity id="3F09E9849766FFCAFD9356346371FEA6" box="[539,638,282,307]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="4" pageNumber="5" unit="mm" value="6.05">6.05 mm</quantity>
|
||
(
|
||
<typeStatus id="274AFAC39766FFCAFD05563463FBFEA6" box="[653,756,282,307]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
|
||
) to
|
||
<quantity id="3F09E9849766FFCAFF2256146005FEC6" box="[170,266,314,339]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" pageId="4" pageNumber="5" unit="mm" value="7.55">7.55 mm</quantity>
|
||
(
|
||
<typeStatus id="274AFAC39766FFCAFE9F5614608FFEC6" box="[279,384,314,339]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9766FFCAFF225654620EFDA6" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="F84E44619766FFCAFF225654620EFDA6" blockId="4.[170,795,378,563]" pageId="4" pageNumber="5">
|
||
<emphasis id="CA8598739766FFCAFF225654603DFE06" bold="true" box="[170,306,378,403]" pageId="4" pageNumber="5">Etymology:</emphasis>
|
||
This species is named after Dr. Cleide Costa, in acknowledgement of her great contribution to the study of
|
||
<taxonomicName id="3FF13FE29766FFCAFE8756946085FE46" authorityName="Linnaeus" authorityYear="1758" box="[271,394,442,467]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||
and for supervising many Brazilian coleopterists. An important remark is that the larva of this species is still unknown and we hope it will be described in collaboration with Dr. Costa when available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9766FFCAFF2255746215FD06" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="F84E44619766FFCAFF2255746215FD06" blockId="4.[170,794,602,659]" pageId="4" pageNumber="5">
|
||
<emphasis id="CA8598739766FFCAFF225574604EFDE6" bold="true" box="[170,321,602,627]" pageId="4" pageNumber="5">Distribution:</emphasis>
|
||
So far, this species is only known from the
|
||
<typeStatus id="274AFAC39766FFCAFF22555461D3FD06" box="[170,220,634,659]" pageId="4" pageNumber="5">type</typeStatus>
|
||
locality (Monte Alegre de
|
||
<collectingRegion id="3A358A839766FFCAFE7555546332FD06" box="[509,573,634,659]" country="Brazil" name="Goias" pageId="4" pageNumber="5">Goiás</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFDCD555463CFFD06" box="[581,704,634,659]" country="Brazil" name="Goias" pageId="4" pageNumber="5">Goiás state</collectingRegion>
|
||
) (
|
||
<figureCitation id="60CA58E49766FFCAFD5A55546200FD06" box="[722,783,634,659]" captionStart="Figure 4" captionStartId="4.[869,928,1047,1072]" captionTargetBox="[873,1491,207,1014]" captionTargetId="figure-503@4.[869,1493,204,1017]" captionTargetPageId="4" captionText="Figure 4. The distribution record of Canthon cleidecostae sp. nov., central Brazil [TO = Tocantins;MT = Mato Grosso;GO = Goiás;DF = Distrito Federal; BA = Bahia;MG = Minas Gerais;MS = Mato Grosso do Sul]." figureDoi="http://doi.org/10.5281/zenodo.4615086" httpUri="https://zenodo.org/record/4615086/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9766FFCAFF22559460DCFC06" pageId="4" pageNumber="5" type="biology_ecology">
|
||
<paragraph id="F84E44619766FFCAFF22559460DCFC06" blockId="4.[170,794,698,915]" pageId="4" pageNumber="5">
|
||
<emphasis id="CA8598739766FFCAFF225594600AFD46" bold="true" box="[170,261,698,723]" pageId="4" pageNumber="5">Biology</emphasis>
|
||
: Nothing is known about the biology of
|
||
<taxonomicName id="3FF13FE29766FFCAFF2255F46046FD66" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[170,329,730,755]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFF2255F46046FD66" box="[170,329,730,755]" italics="true" pageId="4" pageNumber="5">C. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFED955F460A7FD66" box="[337,424,730,755]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The feeding habits are probably coprophagous, considering related species of the same genus. Besides, the type-series was collected in
|
||
<taxonomicName id="3FF13FE29766FFCAFD4854346386FCC6" authority="Moojen, Locks & Langguth, 1997" authorityName="Moojen, Locks & Langguth" authorityYear="1997" class="Mammalia" family="Caviidae" genus="Kerodon" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acrobata">
|
||
<emphasis id="CA8598739766FFCAFD4854346003FCC6" italics="true" pageId="4" pageNumber="5">Kerodon acrobata</emphasis>
|
||
Moojen, Locks & Langguth, 1997
|
||
</taxonomicName>
|
||
faeces, a rodent endemic of
|
||
<collectingCountry id="80E604F19766FFCAFEFF547460BCFCE6" box="[375,435,858,883]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
and know only to
|
||
<collectingRegion id="3A358A839766FFCAFD1354746215FCE6" box="[667,794,858,883]" country="Brazil" name="Goias" pageId="4" pageNumber="5">Goiás state</collectingRegion>
|
||
(
|
||
<bibRefCitation id="9C6039909766FFCAFF38545460C6FC06" author="Woods, C. A. & Kilpatrick, C. W." box="[176,457,890,915]" pageId="4" pageNumber="5" pagination="1538 - 1600" refId="ref4669" refString="Woods, C. A. & Kilpatrick, C. W. 2005. Infraorder Hystricognathi. In: Wilson, D. E. & Reeder, D. M. (Eds.). Mammal Species of the World: A taxonomic and geographic reference. 3. ed. Baltimore, Johns Hopkins University Press. v. 2, p. 1538 - 1600." type="journal article" year="2005">Woods & Kilpatrick, 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B0EB17EA9766FFCAFF2254946435F906" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="F84E44619766FFCAFF2254946215FBC6" blockId="4.[170,795,954,1491]" pageId="4" pageNumber="5">
|
||
<emphasis id="CA8598739766FFCAFF225494603EFC46" bold="true" box="[170,305,954,979]" pageId="4" pageNumber="5">Comments:</emphasis>
|
||
The placement of this new species within the subgenera of
|
||
<taxonomicName id="3FF13FE29766FFCAFEFD54F460DDFC66" authorityName="Hoffmannsegg" authorityYear="1817" box="[373,466,986,1011]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CA8598739766FFCAFEFD54F460DDFC66" box="[373,466,986,1011]" italics="true" pageId="4" pageNumber="5">Canthon</emphasis>
|
||
</taxonomicName>
|
||
is still uncertain. Some morphological characteristics used to describe this species are similar to
|
||
<taxonomicName id="3FF13FE29766FFCAFEDE53346218FBA6" authority="(Martinez, 1952)" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[342,791,1050,1075]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ibarragrassoi">
|
||
<emphasis id="CA8598739766FFCAFEDE53346344FBA6" box="[342,587,1050,1075]" italics="true" pageId="4" pageNumber="5">Canthon ibarragrassoi</emphasis>
|
||
(
|
||
<bibRefCitation id="9C6039909766FFCAFDD75334621EFBA6" author="Martinez, A." box="[607,785,1050,1075]" pageId="4" pageNumber="5" pagination="53 - 118" refId="ref4011" refString="Martinez, A. 1952. Scarabaeidae nuevos o poco conocidos, III. Mision de Estudios de Patologia regional argentina, 23 (81 - 82): 53 - 118." type="journal article" year="1952">Martínez, 1952</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, mainly considering the sinuous carina in the metafemur.
|
||
</paragraph>
|
||
<caption id="AC8E14E99766FFCAFCED53396445FBE5" ID-DOI="http://doi.org/10.5281/zenodo.4615086" ID-Zenodo-Dep="4615086" httpUri="https://zenodo.org/record/4615086/files/figure.png" pageId="4" pageNumber="5" startId="4.[869,928,1047,1072]" targetBox="[873,1491,207,1014]" targetPageId="4">
|
||
<paragraph id="F84E44619766FFCAFCED53396445FBE5" blockId="4.[869,1494,1047,1136]" pageId="4" pageNumber="5">
|
||
Figure 4. The distribution record of
|
||
<taxonomicName id="3FF13FE29766FFCAFB1753396446FBA5" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[1183,1353,1047,1072]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFB1753396446FBA5" bold="true" box="[1183,1353,1047,1072]" italics="true" pageId="4" pageNumber="5">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFAD85339649CFBA5" box="[1360,1427,1047,1072]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, central Brazil [TO = Tocantins;MT = Mato Grosso;GO = Goiás;DF = Distrito Federal; BA = Bahia;MG = Minas Gerais;MS = Mato Grosso do Sul].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F84E44619766FFCAFF5853746329FAE6" blockId="4.[170,795,954,1491]" pageId="4" pageNumber="5">
|
||
The geographic distribution of both species is distinct.
|
||
<taxonomicName id="3FF13FE29766FFCAFF61535460C1FB06" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[233,462,1146,1171]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFF61535460C1FB06" box="[233,462,1146,1171]" italics="true" pageId="4" pageNumber="5">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFE5C53546326FB06" box="[468,553,1146,1171]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known for the
|
||
<typeStatus id="274AFAC39766FFCAFD6053546215FB06" box="[744,794,1146,1171]" pageId="4" pageNumber="5">type</typeStatus>
|
||
locality as aforementioned, while
|
||
<taxonomicName id="3FF13FE29766FFCAFDA153B46216FB26" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[553,793,1178,1203]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ibarragrassoi">
|
||
<emphasis id="CA8598739766FFCAFDA153B46216FB26" box="[553,793,1178,1203]" italics="true" pageId="4" pageNumber="5">Canthon ibarragrassoi</emphasis>
|
||
</taxonomicName>
|
||
is recorded to
|
||
<collectingCountry id="80E604F19766FFCAFEC6539460BAFB46" box="[334,437,1210,1235]" name="Paraguay" pageId="4" pageNumber="5">Paraguay</collectingCountry>
|
||
(
|
||
<typeStatus id="274AFAC39766FFCAFE4B539460F8FB46" box="[451,503,1210,1235]" pageId="4" pageNumber="5">type</typeStatus>
|
||
locality: Colonia Natalício Talavera),
|
||
<collectingCountry id="80E604F19766FFCAFE9053F46089FB66" box="[280,390,1242,1267]" name="Argentina" pageId="4" pageNumber="5">Argentina</collectingCountry>
|
||
(
|
||
<collectingCountry id="80E604F19766FFCAFE1953F463ADFB66" box="[401,674,1242,1267]" name="Mesopotamia Argentina" pageId="4" pageNumber="5">Mesopotamia Argentina</collectingCountry>
|
||
) and
|
||
<collectingCountry id="80E604F19766FFCAFD5653F46215FB66" box="[734,794,1242,1267]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="3A358A839766FFCAFF3853D46088FA86" box="[176,391,1274,1299]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="5">Rio Grande do Sul</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFE1C53D46333FA86" box="[404,572,1274,1299]" country="Brazil" name="Santa Catarina" pageId="4" pageNumber="5">Santa Catarina</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFDC153D46398FA86" box="[585,663,1274,1299]" country="Brazil" name="Parana" pageId="4" pageNumber="5">Paraná</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFD2C53D46218FA86" box="[676,791,1274,1299]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="5">São Paulo</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFF225234609EFAA6" box="[170,401,1306,1331]" country="Brazil" name="Mato Grosso do Sul" pageId="4" pageNumber="5">Mato Grosso do Sul</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFE175234634AFAA6" box="[415,581,1306,1331]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="5">Rio de Janeiro</collectingRegion>
|
||
,
|
||
<collectingRegion id="3A358A839766FFCAFDDB523463EAFAA6" box="[595,741,1306,1331]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="5">Minas Gerais</collectingRegion>
|
||
and
|
||
<collectingRegion id="3A358A839766FFCAFF225214604CFAC6" box="[170,323,1338,1363]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="5">Espirito Santo</collectingRegion>
|
||
states, the latter state being the northernmost known record of the species.
|
||
</paragraph>
|
||
<paragraph id="F84E44619766FFCAFF58525464DEFAE6" blockId="4.[170,795,954,1491]" lastBlockId="4.[869,1494,1210,1683]" pageId="4" pageNumber="5">
|
||
As currently understood,
|
||
<taxonomicName id="3FF13FE29766FFCAFD8B525463BFFA06" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[515,688,1402,1427]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ibarragrassoi">
|
||
<emphasis id="CA8598739766FFCAFD8B525463BFFA06" box="[515,688,1402,1427]" italics="true" pageId="4" pageNumber="5">C. ibarragrassoi</emphasis>
|
||
</taxonomicName>
|
||
belongs to the subgenus
|
||
<taxonomicName id="3FF13FE29766FFCAFEF752B46333FA26" authorityName="Martinez" authorityYear="1948" box="[383,572,1434,1459]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subGenus" subGenus="Glaphyrocanthon">
|
||
<emphasis id="CA8598739766FFCAFEF752B46333FA26" box="[383,572,1434,1459]" italics="true" pageId="4" pageNumber="5">Glaphyrocanthon</emphasis>
|
||
</taxonomicName>
|
||
considering many characteristics listed by
|
||
<bibRefCitation id="9C6039909766FFCAFE3752946218FA46" author="Halffter, G. & Martinez, A." box="[447,791,1466,1491]" pageId="4" pageNumber="5" pagination="29 - 107" refId="ref3798" refString="Halffter, G. & Martinez, A. 1977. Revision monografica de los Canthonina americanos, IV parte. Clave para generos y subgeneros. Folia EntomolOgica Mexicana, 38: 29 - 107." type="journal article" year="1977">Halffter & Martínez (1977: 79)</bibRefCitation>
|
||
. However, even
|
||
<taxonomicName id="3FF13FE29766FFCAFB96539465CEFB46" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[1054,1217,1210,1235]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFB96539465CEFB46" box="[1054,1217,1210,1235]" italics="true" pageId="4" pageNumber="5">C. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFB4753946424FB46" box="[1231,1323,1210,1235]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
sharing these characteristics with
|
||
<taxonomicName id="3FF13FE29766FFCAFBE853F4641CFB66" baseAuthorityName="Martinez" baseAuthorityYear="1952" box="[1120,1299,1242,1267]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ibarragrassoi">
|
||
<emphasis id="CA8598739766FFCAFBE853F4641CFB66" box="[1120,1299,1242,1267]" italics="true" pageId="4" pageNumber="5">C. ibarragrassoi</emphasis>
|
||
</taxonomicName>
|
||
we concluded that the new species should not be included in the
|
||
<taxonomicName id="3FF13FE29766FFCAFCED5234652DFAA6" authorityName="Martinez" authorityYear="1948" box="[869,1058,1306,1331]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subGenus" subGenus="Glaphyrocanthon">
|
||
<emphasis id="CA8598739766FFCAFCED5234652DFAA6" box="[869,1058,1306,1331]" italics="true" pageId="4" pageNumber="5">Glaphyrocanthon</emphasis>
|
||
</taxonomicName>
|
||
without a detailed taxonomic revision of this subgenus. Thus, to avoid “taxonomic instability”, we regard
|
||
<taxonomicName id="3FF13FE29766FFCAFC52527465CFFAE6" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[986,1216,1370,1395]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFC52527465CFFAE6" box="[986,1216,1370,1395]" italics="true" pageId="4" pageNumber="5">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFB4D52746416FAE6" box="[1221,1305,1370,1395]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
as
|
||
<emphasis id="CA8598739766FFCAFAB3527464DEFAE6" box="[1339,1489,1370,1395]" italics="true" pageId="4" pageNumber="5">incertae sedis.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="F84E44619766FFCAFC0352546435F906" blockId="4.[869,1494,1210,1683]" pageId="4" pageNumber="5">
|
||
In addition, the sclerites of the internal sac are not similar to those found in
|
||
<emphasis id="CA8598739766FFCAFB4F52B4646DFA26" box="[1223,1378,1434,1459]" italics="true" pageId="4" pageNumber="5">Goniocanthon</emphasis>
|
||
<bibRefCitation id="9C6039909766FFCAFAE552B46502FA46" author="Martinez, A. & Pereira, F. S." pageId="4" pageNumber="5" pagination="363 - 388" refId="ref4064" refString="Martinez, A. & Pereira, F. S. 1956. Dois generos novos de Canthonini americanos (Col. Scarabaeoidea, Scarabaeidae). Papeis Avulsos de Zoologia, Sao Paulo, 12: 363 - 388." type="journal article" year="1956">Pereira & Martínez, 1956</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="9C6039909766FFCAFB93529465E5FA46" author="Nunes, L. G. de & Nunes, R. V. & Vaz-de-Mello, F. Z." box="[1051,1258,1466,1491]" pageId="4" pageNumber="5" pagination="1 - 31" refId="ref4105" refString="Nunes, L. G. de O. A.; Nunes, R. V. & Vaz-de-Mello, F. Z. 2018. Taxonomic revision of the South American subgenus Canthon (Pelthecanthon) Pereira, 1953 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). European Journal of Taxonomy, 437: 1 - 31." type="journal article" year="2018">
|
||
Nunes
|
||
<emphasis id="CA8598739766FFCAFBE4529465A8FA46" box="[1132,1191,1466,1491]" italics="true" pageId="4" pageNumber="5">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), nor in
|
||
<emphasis id="CA8598739766FFCAFAC0529464DAFA46" box="[1352,1493,1466,1491]" italics="true" pageId="4" pageNumber="5">Peltecanthon</emphasis>
|
||
(Nunes
|
||
<emphasis id="CA8598739766FFCAFC3752F462F2FA66" box="[959,1021,1498,1523]" italics="true" pageId="4" pageNumber="5">et al.,</emphasis>
|
||
in press), where the Ax + FLP complex gradually tapers to a much thinner apex, or even other
|
||
<taxonomicName id="3FF13FE29766FFCAFCED513462CDF9A6" authorityName="Hoffmannsegg" authorityYear="1817" box="[869,962,1562,1587]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CA8598739766FFCAFCED513462CDF9A6" box="[869,962,1562,1587]" italics="true" pageId="4" pageNumber="5">Canthon</emphasis>
|
||
</taxonomicName>
|
||
handled by one of the authors (Nunes,
|
||
<emphasis id="CA8598739766FFCAFAF0513462B4F9C6" italics="true" pageId="4" pageNumber="5">personal remarks</emphasis>
|
||
). In
|
||
<taxonomicName id="3FF13FE29766FFCAFC62511465C0F9C6" authority="Vaz-de-Mello & Nunes & Costa-Silva, 2020" authorityName="Vaz-de-Mello & Nunes & Costa-Silva" authorityYear="2020" box="[1002,1231,1594,1619]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="CA8598739766FFCAFC62511465C0F9C6" box="[1002,1231,1594,1619]" italics="true" pageId="4" pageNumber="5">Canthon cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D1B625089766FFCAFB5C51146425F9C6" box="[1236,1322,1594,1619]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the apex is not thin, something similar to that observed in
|
||
<taxonomicName id="3FF13FE29766FFCAFADE51746518F906" authority="Blanchard, 1845" authorityName="Blanchard" authorityYear="1845" class="Insecta" family="Scarabaeidae" genus="Tetraechma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CA8598739766FFCAFADE517464DAF9E6" box="[1366,1493,1626,1651]" italics="true" pageId="4" pageNumber="5">Tetraechma</emphasis>
|
||
<bibRefCitation id="9C6039909766FFCAFCED51546518F906" author="Blanchard, C. E." box="[869,1047,1658,1683]" pageId="4" pageNumber="5" pagination="155 - 168" refId="ref3633" refString="Blanchard, C. E. 1845. Insectes de l'Amerique Meridionale. In: d'Orbigny, A.; Blanchard, C. E. & Brulle, A. Voyage dans l'Amerique Meridionale, Insectes Coleopteres. Tribu des Lamellicornes Paris, P. Bertrand. v. 6, pt 2, p. 155 - 168." type="book chapter" year="1845">Blanchard, 1845</bibRefCitation>
|
||
</taxonomicName>
|
||
(Nunes,
|
||
<emphasis id="CA8598739766FFCAFBFD51546422F906" box="[1141,1325,1658,1683]" italics="true" pageId="4" pageNumber="5">personal remarks</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |