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<document id="29C0A92F85DA65EA26F103CAEA9CB593" ID="10.11646/zootaxa.4196.4.1" ID-DOI="10.11646/zootaxa.4196.4.1" ID-GBIF-Dataset="921bb78e-4a75-4fa5-82b6-5c7a75c7ad1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="168273" ID-ZooBank="80863DAE-C9E4-466D-9E27-AD938E826D4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479971428066" checkinUser="plazi" docAuthor="Samyn, Yves &amp; Vandenspiegel, Didier" docDate="2016" docId="6C2D24149073A039FF79FC88B3B4FE5B" docLanguage="en" docName="zootaxa.4196.4.1.pdf" docOrigin="Zootaxa 4196 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudostichopus mollis Theel 1886" docType="treatment" docVersion="13" lastPageNumber="480" masterDocId="90145C6C906FA024FFEEFFFBB052FFF6" masterDocTitle="Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species" masterLastPageNumber="497" masterPageNumber="451" pageNumber="479" updateTime="1698394447745" updateUser="plazi">
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<mods:title id="2EE95619AE175F9239119ADFBA5C3583">Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species</mods:title>
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<mods:namePart id="B6E3FCC97417FE2AD822D3FC42CD099F">Samyn, Yves</mods:namePart>
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<treatment id="6C2D24149073A039FF79FC88B3B4FE5B" ID-DOI="http://doi.org/10.5281/zenodo.3503266" ID-GBIF-Taxon="124831659" ID-Zenodo-Dep="3503266" LSID="urn:lsid:plazi:treatment:6C2D24149073A039FF79FC88B3B4FE5B" httpUri="http://treatment.plazi.org/id/6C2D24149073A039FF79FC88B3B4FE5B" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
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<paragraph id="E43B95029073A038FF79FC88B21AFC7B" blockId="28.[151,584,883,943]" box="[151,584,883,909]" pageId="28" pageNumber="479">
<heading id="BF73226E9073A038FF79FC88B21AFC7B" bold="true" box="[151,584,883,909]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
<taxonomicName id="2384EE819073A038FF79FC88B21AFC7B" authority="Theel, 1886" authorityName="Theel" authorityYear="1886" box="[151,584,883,909]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="mollis">
<emphasis id="D6F049109073A038FF79FC88B21AFC7B" bold="true" box="[151,584,883,909]" pageId="28" pageNumber="479">
<emphasis id="D6F049109073A038FF79FC88B1E3FC7B" bold="true" box="[151,433,883,909]" italics="true" pageId="28" pageNumber="479">Pseudostichopus mollis</emphasis>
<bibRefCitation id="8015E8F39073A038FE57FC88B21AFC7B" author="Theel" box="[441,584,883,909]" pageId="28" pageNumber="479" refString="Theel, H. (1886) Holothuroidea. Part 2. Report of the Scientific Results of the Voyage of the &quot; Challenger &quot; (Zoology), 39, 1 - 290, 16 plates." type="book chapter" year="1886">Théel, 1886</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="E43B95029073A038FF79FC6DB17CFC59" blockId="28.[151,584,883,943]" box="[151,302,918,943]" pageId="28" pageNumber="479">
(
<figureCitation id="7CBF89879073A038FF71FC6DB0A2FC59" box="[159,240,918,943]" captionStart="FIGURE 20" captionStartId="28.[151,250,1918,1940]" captionTargetBox="[151,1436,1164,1896]" captionTargetId="figure@28.[151,1436,1164,1896]" captionTargetPageId="28" captionText="FIGURE 20. Pseudostichopus mollis Théel, 1886. A: different view of specimens, B: ossicles from tentacles Scale bar: A = 1 cm, B = 20 µm." httpUri="https://zenodo.org/record/258674/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
A
<emphasis id="D6F049109073A038FEEAFC6DB140FC59" bold="true" box="[260,274,918,943]" pageId="28" pageNumber="479"></emphasis>
B)
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899073A038FF79FC25B216FBE4" pageId="28" pageNumber="479" type="reference_group">
<paragraph id="E43B95029073A038FF79FC25B216FBE4" blockId="28.[151,1435,990,1042]" pageId="28" pageNumber="479">
<treatmentCitationGroup id="C494B22C9073A038FF79FC25B216FBE4" infered="true" pageId="28" pageNumber="479">
<taxonomicName id="2384EE819073A038FF79FC25B265FC02" authority="Theel, 1886: 169" authorityName="Theel" authorityPageNumber="169" authorityYear="1886" box="[151,567,990,1012]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="mollis">
<emphasis id="D6F049109073A038FF79FC25B1D7FC02" box="[151,389,990,1012]" italics="true" pageId="28" pageNumber="479">Pseudostichopus mollis</emphasis>
<treatmentCitation id="6525B3139073A038FE65FC25B265FC02" author="Theel" box="[395,567,990,1012]" page="169" pageId="28" pageNumber="479" year="1886">
<bibRefCitation id="8015E8F39073A038FE65FC25B255FC02" author="Theel" box="[395,519,990,1012]" pageId="28" pageNumber="479" refString="Theel, H. (1886) Holothuroidea. Part 2. Report of the Scientific Results of the Voyage of the &quot; Challenger &quot; (Zoology), 39, 1 - 290, 16 plates." type="book chapter" year="1886">Théel, 1886</bibRefCitation>
: 169
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170, pl. 10 figs 5, 6; OLoughlin &amp;
<treatmentCitation id="6525B3139073A038FC45FC25B438FC02" author="Ahearn" box="[939,1130,990,1012]" page="171" pageId="28" pageNumber="479" year="2005">
<bibRefCitation id="8015E8F39073A038FC45FC25B46AFC02" author="Ahearn" box="[939,1080,990,1012]" pageId="28" pageNumber="479" refString="O'Louglin, P. M. &amp; Ahearn, C. (2005) A review of pygal-furrowed Synallactidae (Echinodermata: Holothuroidea), with new species from the Antarctic, Atlantic and Pacific oceans. Memoirs of Museum Victoria, 62, 147 - 179." type="journal article" year="2005">Ahearn, 2005</bibRefCitation>
: 171
</treatmentCitation>
, figs 1b, c, 9a, e, 10b, c, 11e, f (synonymy and records before 2005).
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="AC9EC6899073A038FF79FBC4B54FFBAE" box="[151,1309,1087,1112]" pageId="28" pageNumber="479" type="materials_examined">
<paragraph id="E43B95029073A038FF79FBC4B54FFBAE" blockId="28.[151,1309,1087,1112]" box="[151,1309,1087,1112]" pageId="28" pageNumber="479">
<emphasis id="D6F049109073A038FF79FBC4B12BFBAE" bold="true" box="[151,377,1087,1112]" pageId="28" pageNumber="479">Material examined</emphasis>
.
<emphasis id="D6F049109073A038FE69FBBBB206FBAE" box="[391,596,1087,1112]" italics="true" pageId="28" pageNumber="479">Non-type material</emphasis>
:
<emphasis id="D6F049109073A038FD8CFBC4B34EFBAE" bold="true" box="[610,796,1087,1112]" pageId="28" pageNumber="479">IE-2007-778 (3)</emphasis>
(6 specimens, sampled in front of Majumga).
</paragraph>
</subSubSection>
<caption id="B0FBC58A9073A038FF79F885B16CF845" httpUri="https://zenodo.org/record/258674/files/figure.png" pageId="28" pageNumber="479" targetBox="[151,1436,1164,1896]" targetPageId="28">
<paragraph id="E43B95029073A038FF79F885B16CF845" blockId="28.[151,1436,1918,1971]" pageId="28" pageNumber="479">
<emphasis id="D6F049109073A038FF79F885B173F865" bold="true" box="[151,289,1918,1940]" pageId="28" pageNumber="479">FIGURE 20.</emphasis>
<taxonomicName id="2384EE819073A038FEC6F885B2CCF862" authority="Theel, 1886" authorityName="Theel" authorityYear="1886" box="[296,670,1918,1940]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="mollis">
<emphasis id="D6F049109073A038FEC6F885B24AF862" box="[296,536,1918,1940]" italics="true" pageId="28" pageNumber="479">Pseudostichopus mollis</emphasis>
Théel, 1886
</taxonomicName>
. A: different view of specimens, B: ossicles from tentacles Scale bar: A = 1cm, B = 20µm.
</paragraph>
</caption>
<subSubSection id="AC9EC6899072A039FF29FF6CB3B4FE5B" pageId="29" pageNumber="480" type="discussion">
<paragraph id="E43B95029072A039FF29FF6CB3B4FE5B" blockId="29.[151,1436,151,429]" pageId="29" pageNumber="480">
<materialsCitation id="54EC9F5F9072A039FF29FF6CB3B4FE5B" ID-GBIF-Occurrence="1343648503" country="Iceland" location="Southern" pageId="29" pageNumber="480" specimenCount="1" stateProvince="Southern">
<emphasis id="D6F049109072A039FF29FF6CB166FF46" bold="true" box="[199,308,151,176]" pageId="29" pageNumber="480">Remarks</emphasis>
. The species in the genus
<taxonomicName id="2384EE819072A039FD9DFF6CB37DFF46" box="[627,815,151,176]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="genus">
<emphasis id="D6F049109072A039FD9DFF6CB37DFF46" box="[627,815,151,176]" italics="true" pageId="29" pageNumber="480">Pseudostichopus</emphasis>
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are notoriously difficult to identify because they are characterized by lacking ossicles in the body wall. From the region, three species have been documented:
<taxonomicName id="2384EE819072A039FA69FF46B190FF0E" authority="Thandar, 1992" authorityName="Thandar" authorityYear="1992" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="species" species="echinatus">
<emphasis id="D6F049109072A039FA69FF46B151FF0E" italics="true" pageId="29" pageNumber="480">P. echinatus</emphasis>
<bibRefCitation id="8015E8F39072A039FEFAFF24B190FF0E" author="Thandar" box="[276,450,223,248]" pageId="29" pageNumber="480" refString="Thandar, A. S. (1992) The South African Museum's Meiring Naude Cruises. Part 18. Holothuroidea. Annals of the South African Museum, 10, 159 - 180." type="journal article" year="1992">Thandar, 1992</bibRefCitation>
</taxonomicName>
characterized by having very irregular knobbed rod ossicles in the tentacles;
<taxonomicName id="2384EE819072A039FF79FEFFB2D5FEEA" authority="Sluiter, 1901" authorityName="Sluiter" authorityYear="1901" box="[151,647,260,285]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="species" species="hyalegerus">
<emphasis id="D6F049109072A039FF79FEFFB18AFEEB" box="[151,472,260,285]" italics="true" pageId="29" pageNumber="480">Pseudostichopus hyalegerus</emphasis>
(
<bibRefCitation id="8015E8F39072A039FE04FEFFB22DFEEA" author="Sluiter" box="[490,639,260,285]" pageId="29" pageNumber="480" refString="Sluiter C. P. (1901) Die Holothurien der Siboga-Expedition. Siboga - Expeditie. Monograph, 44, 1 - 142,11 plates." type="journal volume" year="1901">Sluiter, 1901</bibRefCitation>
)
</taxonomicName>
(see above) and
<taxonomicName id="2384EE819072A039FCBCFEFEB42EFEEB" authority="Thandar, 2006" authorityName="Thandar" authorityYear="2006" box="[850,1148,260,285]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="species" species="langeae">
<emphasis id="D6F049109072A039FCBCFEFEB398FEEB" box="[850,970,260,285]" italics="true" pageId="29" pageNumber="480">P. langeae</emphasis>
<bibRefCitation id="8015E8F39072A039FC3AFEFFB42EFEEB" author="Thandar" box="[980,1148,260,285]" pageId="29" pageNumber="480" refString="Thandar, A. S. (2006) New species and new records of dendrochirotid and dactylochirotid holothuroids (Echinodermata: Holothuroidea) from off the east coast of South Africa. Zootaxa 1245, 1 - 51." type="journal article" year="2006">Thandar, 2006</bibRefCitation>
</taxonomicName>
, characterized by having branched rods in the gonad.
<taxonomicName id="2384EE819072A039FE3DFED2B260FEB6" box="[467,562,295,320]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="species" species="mollis">
<emphasis id="D6F049109072A039FE3DFED2B260FEB6" box="[467,562,295,320]" italics="true" pageId="29" pageNumber="480">P. mollis</emphasis>
</taxonomicName>
had already been reported from the
<collectingRegion id="26405BE09072A039FC26FEDCB47FFEB6" box="[968,1069,295,320]" country="Sri Lanka" name="Southern" pageId="29" pageNumber="480">Southern</collectingRegion>
Indian Ocean, off Marion
<collectingCountry id="9C93D5929072A039FAB6FEDCB5CEFEB6" box="[1368,1436,295,320]" name="Iceland" pageId="29" pageNumber="480">Island</collectingCountry>
(Oloughlin &amp;
<bibRefCitation id="8015E8F39072A039FEAEFEB7B185FE93" author="Ahearn" box="[320,471,332,357]" pageId="29" pageNumber="480" refString="O'Louglin, P. M. &amp; Ahearn, C. (2005) A review of pygal-furrowed Synallactidae (Echinodermata: Holothuroidea), with new species from the Antarctic, Atlantic and Pacific oceans. Memoirs of Museum Victoria, 62, 147 - 179." type="journal article" year="2005">Ahearn 2005</bibRefCitation>
). The six specimens of
<taxonomicName id="2384EE819072A039FD31FEB6B313FE93" box="[735,833,332,357]" class="Holothuroidea" family="Synallactidae" genus="Pseudostichopus" kingdom="Animalia" order="Aspidochirotida" pageId="29" pageNumber="480" phylum="Echinodermata" rank="species" species="mollis">
<emphasis id="D6F049109072A039FD31FEB6B313FE93" box="[735,833,332,357]" italics="true" pageId="29" pageNumber="480">P. mollis</emphasis>
</taxonomicName>
observed were eviscerated but pieces of gonads were found in two specimens; no ossicles was observed in the gonads but spiny rods (60
<emphasis id="D6F049109072A039FBB2FE94B438FE7E" bold="true" box="[1116,1130,367,392]" pageId="29" pageNumber="480"></emphasis>
240µm long), occasionally branched and with some perforations, were found in the tentacles (
<figureCitation id="7CBF89879072A039FC6FFE6FB399FE5B" box="[897,971,404,429]" captionStart="FIGURE 20" captionStartId="28.[151,250,1918,1940]" captionTargetBox="[151,1436,1164,1896]" captionTargetId="figure@28.[151,1436,1164,1896]" captionTargetPageId="28" captionText="FIGURE 20. Pseudostichopus mollis Théel, 1886. A: different view of specimens, B: ossicles from tentacles Scale bar: A = 1 cm, B = 20 µm." httpUri="https://zenodo.org/record/258674/files/figure.png" pageId="29" pageNumber="480">Fig.20</figureCitation>
B)
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
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