187 lines
22 KiB
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187 lines
22 KiB
XML
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<mods:title id="9B4F23970F1E898E9AC2B515B4907D41">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="45240B35059DD6A8EECD3B88DF038E06">Chamorro, William</mods:namePart>
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<mods:namePart id="DCB114B2F6469FA5302381BA2845E66D">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="BA4A9651B2CDAADED581C7E8AC21F29A">senjo, Angelico</mods:namePart>
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<mods:namePart id="5ECB2BE089E027CDA6FE0E05E341540B">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="CE89FEC69225FBD7787D747FDCD943E2">2019</mods:date>
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<mods:number id="574FCAF39DA867D9B0D5DD5A259E0AD5">826</mods:number>
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<mods:classification id="80E6AAE38F2525343B8D1D7198FCC3A2">journal article</mods:classification>
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<mods:identifier id="6D88740BAF02BB51CB31CFEFC62A20CB" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="62E328A98EE507183DF3D0627DB23D04" ID-GBIF-Taxon="154127654" LSID="urn:lsid:plazi:treatment:62E328A98EE507183DF3D0627DB23D04" httpUri="http://treatment.plazi.org/id/62E328A98EE507183DF3D0627DB23D04" lastPageId="92" lastPageNumber="93" pageId="90" pageNumber="91">
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<subSubSection id="AA7A35AFB38C1DEF54CB14493A10500D" pageId="90" pageNumber="91" type="nomenclature">
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<paragraph id="1B60DEF60CB918E7A912E22E7426EBB2" pageId="90" pageNumber="91">
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<taxonomicName id="4E7E58D50710DC1B848930808DDA3C9A" ID-CoL="34KBR" authority="Buquet, 1844" authorityName="Buquet" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (Deltohyboma) aequinoctiale" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="aequinoctiale" subGenus="Deltohyboma">Deltochilum (Deltohyboma) aequinoctiale (Buquet, 1844)</taxonomicName>
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Plate 21D
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</paragraph>
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</subSubSection>
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<subSubSection id="88B35FF295855D8B2FACA38C46699C3C" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="reference_group">
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<paragraph id="90D47A67B54AB0EE5D978CC1C9F8EDA8" pageId="91" pageNumber="92">
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<taxonomicName id="86934E82F0F5A48DEEC449C918ED4FE1" class="Insecta" family="Scarabaeidae" genus="Hyboma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyboma aequinoctialis" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="aequinoctialis">
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<pageBreakToken id="1A5C8C4E51B2A2C3A4607AEE611D86E9" pageId="91" pageNumber="92" start="start">Hyboma</pageBreakToken>
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aequinoctialis
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</taxonomicName>
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Buquet, 1844: 21 (original description. Type locality: Colombie [= Colombia]).
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</paragraph>
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<paragraph id="59838C13859F56941254F38843FBEBEC" pageId="91" pageNumber="92">
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<taxonomicName id="A97E3CD2FA2E0B8AC599E2DDF25A9D6E" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum aequinoctiale" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="aequinoctiale">Deltochilum aequinoctiale</taxonomicName>
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:
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<bibRefCitation id="C5B1FFF3F8BF04B134AD366E520DA928" author="Burmeister, HCC" journalOrPublisher="Zeitung fuer Zoologie, Zootomie und Palaeozoologie" pageId="264" pageNumber="265" pagination="133 - 136" title="Die Entwicklungsgeschichte der Gattung Deltochilum Esch." volume="1" year="1848">Burmeister 1848</bibRefCitation>
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: 135 (redescription);
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<bibRefCitation id="5304AE76D1F6C7B9759315A3723D6F30" pageId="91" pageNumber="92">Gemminger and Harold 1869</bibRefCitation>
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: 995 (list, distribution);
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<bibRefCitation id="6E5E6785077E3C3889603F2DCCF6179D" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
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: 35 (complete list of species);
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<bibRefCitation id="6391A8E1386979B2513BBF99C2D74F16" author="Balthasar, V" journalOrPublisher="Beitrag Fauna Perus, Jena" pageId="262" pageNumber="263" pagination="337 - 358" title="Scarabaeidae laparostictae." volume="1" year="1941">Balthasar 1941</bibRefCitation>
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: 344 (cited for Peru);
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<bibRefCitation id="D1ED6D4D70001A6341BBD82163ED1597" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 202 (list of species from Latin America);
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<bibRefCitation id="909461E8C2F9E54249C9A2A96E17E6B1" author="Balthasar, V" journalOrPublisher="Beitraege zur Fauna Perus, Jena" pageId="262" pageNumber="263" pagination="322 - 344" title="Scarabaeidae laparostictae." volume="2" year="1951">Balthasar 1951</bibRefCitation>
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: 329 (cited for Peru);
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<bibRefCitation id="9F2D7203EA4741BF7C56E6ACE77FADF8" author="Contreras, GC" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Fisicas y Naturales" pageId="265" pageNumber="266" pagination="220 - 229" title="Catalogo de los coleopteros colombianos." volume="8" year="1951">Contreras 1951</bibRefCitation>
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: 221 (cited for Colombia, cited as
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<taxonomicName id="DFE72E6668774A2CD9EF441B821FFCAE" family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="D. aequinoctiale" order="Coleoptera" pageId="91" pageNumber="92" rank="species" species="aequinoctiale">D. aequinoctiale</taxonomicName>
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);
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<bibRefCitation id="05F634AB22F100103E094292DAE3B360" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
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: 136 (cited for Colombia);
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<bibRefCitation id="144AFB8B86B9D944E5B5FCA9B6CBBAEB" author="Medina, CA" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="274" pageNumber="275" pagination="23 - 68" title="Morphological variation and systematics of Canthon and related genera of new world Canthonini dung beetles." url="https://doi.org/10.1002/mmnd.20030500105" volume="50" year="2003">Medina et al. 2003</bibRefCitation>
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: 65 (distribution);
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<bibRefCitation id="06E60B2384C69D472F3EB26C7C6E1534" pageId="91" pageNumber="92">Krajcik 2012</bibRefCitation>
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: 88 (complete list of species);
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<bibRefCitation id="61BCC547207EA7FBF1AB9E7431607588" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
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: 195 (cited for Peru).
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</paragraph>
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<paragraph id="2A8828E718B648CAF3A40AAD81FDB21C" pageId="91" pageNumber="92">
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<taxonomicName id="8174C8F535A9BEB2C79D5ED4C1917756" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum (Deltohyboma) aequinoctiale" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="aequinoctiale" subGenus="Deltohyboma">Deltochilum (Deltohyboma) aequinoctiale</taxonomicName>
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:
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<bibRefCitation id="F5CF197D074404532D52058AE6117748" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="213 - 296" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="107" year="1938">Paulian 1938</bibRefCitation>
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: 270 (characters in key), 278 (redescription);
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<bibRefCitation id="20A090A3C38007F76604D890221890C1" author="Vulcano, MA" journalOrPublisher="Entomologische Arbeiten aus dem Museum G Frey" pageId="279" pageNumber="280" pagination="570 - 685" title="Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere." volume="15" year="1964">Vulcano and Pereira 1964</bibRefCitation>
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: 652 (catalog of species, distribution);
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<bibRefCitation id="FC7BFBD4DC481D2AA78A8A8CEF977A01" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
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: 557 (characters in key);
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<bibRefCitation id="CDE4B771A6430E678775182EBB925570" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 316-317 (cited for Ecuador);
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<bibRefCitation id="C0B5BF6DA44EBF6D7A599E511A1BD92A" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 85 (figure 8H), 86 (figure 9A), 94 (cited for Ecuador).
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</paragraph>
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<paragraph id="35B3663E65B256A58F719714A92F0614" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
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<taxonomicName id="E377015798ECBF329611C6E0A6AD1A2E" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum erodioides" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="erodioides">Deltochilum erodioides</taxonomicName>
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<pageBreakToken id="0F620111E1112A9BA7C7CB677556D036" pageId="92" pageNumber="93" start="start">Harold</pageBreakToken>
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, 1867d: 77 (original description);
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<bibRefCitation id="3569E7A872347AB759425AF184F42B20" pageId="92" pageNumber="93">Gemminger and Harold 1869</bibRefCitation>
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: 995 (list, distribution);
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<bibRefCitation id="FBCB413289651ADD9F698CF4B3E19EA6" author="Harold, E" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="269" pageNumber="270" pagination="13 - 46" title="Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien." volume="41" year="1880 a">Harold 1880a</bibRefCitation>
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: 18 (cited for Colombia);
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<bibRefCitation id="0FC6C17201197BA3C92408C09E69412B" author="Kirsch, T" journalOrPublisher="Berliner entomologische Zeitschrift" pageId="270" pageNumber="271" pagination="207 - 224" title="Neue suedamerikanische Kaefer." url="https://doi.org/10.1002/mmnd.18850290206" volume="29" year="1885">Kirsch 1885</bibRefCitation>
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: 212 (cited for Ecuador);
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<bibRefCitation id="FC08D2DCDA9F789782D71DDF74836403" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
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: 36 (complete list of species);
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<bibRefCitation id="783CA2A9C98693F95E7764C47C63D35E" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="213 - 296" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="107" year="1938">Paulian 1938</bibRefCitation>
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: 278 (synonym of
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<taxonomicName id="F0E78F57A89CC81ECA721C619D202E69" family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="D. (D.) aequinoctiale" order="Coleoptera" pageId="92" pageNumber="93" rank="species" species="aequinoctiale" subGenus="D.">D. (D.) aequinoctiale</taxonomicName>
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Buq.);
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<bibRefCitation id="F0DB93EE8FE88FE81BB4237C785E6863" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 202 (list of species of Latin America);
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<bibRefCitation id="57D71DE0ED12010C509576026BFB874D" author="Contreras, GC" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Fisicas y Naturales" pageId="265" pageNumber="266" pagination="220 - 229" title="Catalogo de los coleopteros colombianos." volume="8" year="1951">Contreras 1951</bibRefCitation>
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: 222 (cited for Colombia);
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<bibRefCitation id="1A71142E6D36EC365DA8C2372FBC78F2" author="Vulcano, MA" journalOrPublisher="Entomologische Arbeiten aus dem Museum G Frey" pageId="279" pageNumber="280" pagination="570 - 685" title="Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere." volume="15" year="1964">Vulcano and Pereira 1964</bibRefCitation>
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: 652 (cited as synonym of
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<taxonomicName id="F919ACAB17EC54D561B76BE8CB9E3043" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum aequinotiale" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinotiale">Deltochilum aequinotiale</taxonomicName>
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Buquet);
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<bibRefCitation id="8572D8C686EC0DACD415E01DD167D469" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
|
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: 316-317 (cited for Ecuador);
|
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<bibRefCitation id="0D94D992078BD4D0E996A72AE21C8223" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
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: 195 (cited for Peru).
|
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</paragraph>
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</subSubSection>
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<subSubSection id="1BC8CF9C46EE50B4BD1E7AF4EC528942" pageId="92" pageNumber="93" type="type specimens">
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<paragraph id="612F6AC6459263D5DAD637AD9325647A" pageId="92" pageNumber="93">Type specimens.</paragraph>
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<paragraph id="3C66E8F9CEAB334E1111EEB76FD710B7" pageId="92" pageNumber="93">
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<taxonomicName id="2658A178B4CD95CE7DA1AE8F2DA7A842" class="Insecta" family="Scarabaeidae" genus="Hyboma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyboma aequinotialis" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinotialis">Hyboma aequinotialis</taxonomicName>
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Buquet, 1844. One syntype examined deposited at the MNHN (ex coll. D Sharp). Lectotype to be designated in a future work on this species group.
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</paragraph>
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<paragraph id="7DB067982B9F1BB0FAF9C134FE36BAD5" pageId="92" pageNumber="93">
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<taxonomicName id="C7670DDFA255C3715DD13183567920D2" class="Insecta" family="Scarabaeidae" genus="Deltochilum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltochilum erodioides" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="erodioides">Deltochilum erodioides</taxonomicName>
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Harold, 1867. Two syntypes examined deposited at the MNHN (ex coll. E Steinheil, ex coll. E Harold, ex coll. R Oberthur). Lectotype to be designated in a future work on this species group.
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</paragraph>
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</subSubSection>
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<subSubSection id="417334369189831DB0D304AF01A9251D" pageId="92" pageNumber="93" type="distribution">
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<paragraph id="21579C5FAF2D63637F980AAF109A45B5" pageId="92" pageNumber="93">Distribution.</paragraph>
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<paragraph id="E01414EB01FF3C4655BF76BE23B2E1B2" pageId="92" pageNumber="93">Colombia, Ecuador, and Peru.</paragraph>
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</subSubSection>
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<subSubSection id="F82F673FCC85EC641A6CDA33483657CD" pageId="92" pageNumber="93" type="records examined">
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<paragraph id="FCD3806BB7FECCD7758E3D6FF093F884" pageId="92" pageNumber="93">Records examined.</paragraph>
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<paragraph id="248FA3E21E00CE97C9220D35CB1FADBC" pageId="92" pageNumber="93">
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COTOPAXI: Bosque Integral Otonga, 2080 m (78 specimens CEMT: 7 specimens MQCAZ; 4 specimens MUTPL). PICHINCHA: Chespi, Bellavista, 1380 m (1 specimen MUTPL); Yunguilla, Loma La Liberia, 2400 m (2 specimens MUTPL); Reserva
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<normalizedToken id="5E3F06B217DD5E5727045A6208D74880" originalValue="Orquideológica">Orquideologica</normalizedToken>
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El Pahuma, 1975 m (1 specimen MUTPL).
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</paragraph>
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</subSubSection>
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<subSubSection id="38650CB527A1F994455150F70CEC068F" pageId="92" pageNumber="93" type="literature records">
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<paragraph id="A42782354F37228274554F1A5DB41C57" pageId="92" pageNumber="93">Literature records.</paragraph>
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<paragraph id="A536A51111ABEC58DB420425D1D605E2" pageId="92" pageNumber="93">
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BOLIVAR: au dessus de Chimbo, 1900 m (
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<bibRefCitation id="D488F9E4DC636A878C9411F49F69CDE8" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="213 - 296" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="107" year="1938">Paulian 1938</bibRefCitation>
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: 280). MORONA SANTIAGO: Macas (
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<bibRefCitation id="8908F63A84BDF8DFC99B0BCC7B6221A7" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="213 - 296" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="107" year="1938">Paulian 1938</bibRefCitation>
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: 280).
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</paragraph>
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</subSubSection>
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<subSubSection id="CF469504D61C1E2E0FBA4472CBC92CBC" pageId="92" pageNumber="93" type="temporal data">
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<paragraph id="F384378603908E902E71832E1C0F572E" pageId="92" pageNumber="93">Temporal data.</paragraph>
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<paragraph id="DB0BD7B80A0DA271434B44C48069DAA9" pageId="92" pageNumber="93">Collected in January, February, March, April, June, July, September, and December.</paragraph>
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</subSubSection>
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<subSubSection id="A9DDD1135034D749B88D6D65D54863F6" pageId="92" pageNumber="93" type="remarks">
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<paragraph id="71B6C4DF10B52F04A3CC908B07E4359C" pageId="92" pageNumber="93">Remarks.</paragraph>
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<paragraph id="E7645EEC5C448327214733D0D4D980BB" pageId="92" pageNumber="93">
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Inhabits the evergreen lower montane forests and montane cloud forests of the Andean region from 1300-2400 m a.s.l. Collected with pitfall traps baited with carrion and human feces. It is possible that the records cited by
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<bibRefCitation id="5B72FFBFC01C209F891FB1BCA9D9E91E" author="Paulian, R" journalOrPublisher=") Annales de la Societe de France" pageId="274" pageNumber="275" pagination="213 - 296" title="Contribution a l'etude des Canthonides americains (Coleopt. Lamellic." volume="107" year="1938">Paulian (1938)</bibRefCitation>
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from Macas, Morona Santiago province, in the Ecuadorian Amazon, is a different species belonging to the same subgenus. A review of these species is needed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |