treatments-xml/data/62/08/9C/62089C43D7B3558384831625E704FD8A.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1123.90308" ID-Pensoft-Pub="1313-2970-1123-63" ID-Pensoft-UUID="1BCB0D62026158E7B962D3F2EE741FB4" ID-ZooBank="13556B2ABE8744F69A43FCDAFE1C68CB" ModsDocID="1313-2970-1123-63" checkinTime="1664944213626" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana &amp; Gattolliat, Jean-Luc" docDate="2022" docId="62089C43D7B3558384831625E704FD8A" docLanguage="en" docName="ZooKeys 1123: 63-81" docOrigin="ZooKeys 1123" docPubDate="2022-10-04" docSource="http://dx.doi.org/10.3897/zookeys.1123.90308" docTitle="Labiobaetis kui Kaltenbach &amp; Garces &amp; Gattolliat 2022, sp. nov." docType="treatment" docUuid="03B09E8B-57E2-40AA-8BAD-911A6D969606" docUuidSource="ZooBank" docVersion="1" id="1BCB0D62026158E7B962D3F2EE741FB4" lastPageNumber="63" masterDocId="1BCB0D62026158E7B962D3F2EE741FB4" masterDocTitle="First contribution to Labiobaetis Novikova &amp; Kluge in Cambodia (Ephemeroptera, Baetidae), with description of two new species" masterLastPageNumber="81" masterPageNumber="63" pageNumber="63" updateTime="1664944213626" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>First contribution to Labiobaetis Novikova &amp; Kluge in Cambodia (Ephemeroptera, Baetidae), with description of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garces, Jhoana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5150-0281</mods:nameIdentifier>
<mods:affiliation>Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-10-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>63</mods:start>
<mods:end>81</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1123.90308</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1123.90308</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1123-63</mods:identifier>
<mods:identifier type="ZooBank">13556B2ABE8744F69A43FCDAFE1C68CB</mods:identifier>
<mods:identifier type="Pensoft-UUID">1BCB0D62026158E7B962D3F2EE741FB4</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:03B09E8B-57E2-40AA-8BAD-911A6D969606" httpUri="http://treatment.plazi.org/id/62089C43D7B3558384831625E704FD8A" lastPageNumber="63" pageId="0" pageNumber="63">
<subSubSection pageId="0" pageNumber="63" type="nomenclature">
<paragraph pageId="0" pageNumber="63">
<taxonomicName LSID="https://zoobank.org/03B09E8B-57E2-40AA-8BAD-911A6D969606" authority="Kaltenbach &amp; Garces &amp; Gattolliat, 2022" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">Labiobaetis kui</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">, 5c, d</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Labiobaetis in Cambodia a overview map b Labiobaetis species." figureDoi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63">, 6b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="diagnosis">
<paragraph pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Larva.</emphasis>
Following combination of characters: A) antennal scape without process (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4g</figureCitation>
); B) dorsal surface of labrum with submarginal arc of 16-18 long, clavate setae (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3a</figureCitation>
); C) labial palp segment II with an extended, slightly hooked, thumb-like distomedial protuberance (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3i</figureCitation>
); D) left mandible without setae at apex of mola (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3e</figureCitation>
); E) fore femur rather slender, length ca 4 
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with 10-15 curved, spine-like setae (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4a</figureCitation>
); F) hind protoptera absent; G) six pairs of terga-lii; H) paraproct distally slightly expanded, with 33-38 stout, marginal spines (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4f</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., larva morphology 
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
labrum (left: ventral view, right: dorsal view) 
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
seta of submarginal arc 
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
right mandible 
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
right prostheca 
<emphasis bold="true" pageId="0" pageNumber="63">e</emphasis>
left mandible 
<emphasis bold="true" pageId="0" pageNumber="63">f</emphasis>
left prostheca 
<emphasis bold="true" pageId="0" pageNumber="63">g</emphasis>
hypopharynx and superlinguae 
<emphasis bold="true" pageId="0" pageNumber="63">h</emphasis>
maxilla 
<emphasis bold="true" pageId="0" pageNumber="63">i</emphasis>
labium (left: ventral view, right: dorsal view).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., larva morphology 
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
foreleg 
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
seta of tibia dorsal margin 
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
fore claw 
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
tergum IV 
<emphasis bold="true" pageId="0" pageNumber="63">e</emphasis>
tergalius IV 
<emphasis bold="true" pageId="0" pageNumber="63">f</emphasis>
paraproct 
<emphasis bold="true" pageId="0" pageNumber="63">g</emphasis>
antennal base.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">Description.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Larva</emphasis>
(Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">5c, d</figureCitation>
). Body length ca 4.9 mm. Caudalii broken. Antenna broken.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Colouration</emphasis>
(Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">5c, d</figureCitation>
). Head, thorax, and abdomen dorsally uniform brown. Head, thorax, and abdomen ventrally light brown. Legs light brown; femur with a brown medial spot, darker on ventral margin, dorsal margin and apex brown. Caudalii light brown.
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 5.</emphasis>
Habitus, larvae 
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis brao" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="brao">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis brao</emphasis>
</taxonomicName>
sp. nov., dorsal view 
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis brao" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="brao">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis brao</emphasis>
</taxonomicName>
sp. nov., ventral view 
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., dorsal view 
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Antenna</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4g</figureCitation>
) with scape and pedicel subcylindrical, without distolateral process at scape.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Labrum</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3a, b</figureCitation>
). Rectangular, length 0.7 
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 16-18 long, clavate setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae; ventral surface with ca three short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Right mandible</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3c, d</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles, inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Left mandible</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3e, f</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Tuft of setae at apex of mola absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Hypopharynx and superlinguae</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3g</figureCitation>
). Lingua approx. as long as superlinguae. Lingua longer than broad; medial tuft of stout setae well developed; distal half laterally expanded. Superlinguae distally almost straight; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Maxilla</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3h</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid. Medially with one pectinate, spine-like seta and three or four medium, simple setae. Maxillary palp 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; 2-segmented; palp segment II 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Labium</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca six spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with ca four spine-like setae; ventral surface with fine, simple, scattered setae. Paraglossa subrectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and a row of 2-4 medium, simple setae in anteromedial area; dorsally with a row of four or five long, spine-like, simple setae near inner margin. Labial palp with segment I 0.8 
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with extended, slightly hooked, thumb-like distomedial protuberance; distomedial protuberance 0.7 
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with two long, spine-like, simple setae near outer margin. Segment III slightly pentagonal; apex rounded, inner apical margin slightly concave; length subequal to width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Foreleg</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4a-c</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.4:0.2. 
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Femur</emphasis>
.
</emphasis>
Length ca 4 
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with 10-15 long, curved, spine-like setae; length of setae 0.23 
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded, with a pair of long, curved, spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch absent. 
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Tibia</emphasis>
.
</emphasis>
Dorsal margin with row of short, stout, apically rounded setae, on apex one longer, spine-like seta. Ventral margin with row of short, curved, spine-like setae, on apex some longer setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae near ventral margin. Patellatibial suture present on basal 1/3 area. 
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Tarsus</emphasis>
.
</emphasis>
Dorsal margin with some fine, simple setae. Ventral margin with row of curved, spine-like setae. Claw with one row of nine or ten denticles; distally pointed; with ca. five stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Terga</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4d</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Tergalii</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4e</figureCitation>
). Present on segments II-VII. Margins with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Tergalius IV as long as length of segments V and 1/3 VI combined. Tergalius VII as long as length of segment VIII.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Paraproct</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4f</figureCitation>
). Distally slightly expanded, with 33-38 stout, marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with numerous small, marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="etymology">
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="63">The new species is dedicated to the indigenous Kui people from northeastern Cambodia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="distribution">
<paragraph pageId="0" pageNumber="63">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="63">
Cambodia (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Labiobaetis in Cambodia a overview map b Labiobaetis species." figureDoi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63">6b</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 6.</emphasis>
Distribution of 
<taxonomicName authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis</emphasis>
</taxonomicName>
in Cambodia 
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
overview map 
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
<taxonomicName authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="biological aspects">
<paragraph pageId="0" pageNumber="63">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="63">The specimens were collected from 100 m to 640 m, mostly on littoral gravel.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="type material">
<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
.
</emphasis>
Cambodia • larva; Kampong Speu Province, Chambok River, 1.83 Km from Chambok Community; 240 m; 
<geoCoordinate degrees="11" direction="north" minutes="21" orientation="latitude" precision="15" seconds="58" value="11.366112">11°21'58&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="06" orientation="longitude" precision="15" seconds="17" value="104.10472">104°06'17&quot;E</geoCoordinate>
; 11.07.2018; leg. H. Freitag and J. Garces; on slide; GBIFCH00592702; MZL. 
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Paratypes</emphasis>
.
</emphasis>
Cambodia • 14 larvae; same data as holotype; 1 on slide; GBIFCH00592701; MZL; 13 in alcohol; GenBank ON982737, ON982738; GBIFCH00515681, GBIFCH00829876, GBIFCH00829877, GBIFCH00975577, GBIFCH00975578; MZL • 7 larvae; Kampong Speu Province, waterfall at Kirirom National Park; 640 m; 
<geoCoordinate degrees="11" direction="north" minutes="20" orientation="latitude" precision="15" seconds="26" value="11.340555">11°20'26&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="02" orientation="longitude" precision="15" seconds="14" value="104.037224">104°02'14&quot;E</geoCoordinate>
; 13.07.2018; leg. H. Freitag and J. Garces; 1 on slide; GBIFCH00592698; RUPP; 6 in alcohol; GBIFCH00975579; MZL • 1 larva; Kampong Speu Province, Kokie waterfall, secondary forest remnants; 110 m; 
<geoCoordinate degrees="11" direction="north" minutes="12" orientation="latitude" precision="15" seconds="11" value="11.203055">11°12'11&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="03" orientation="longitude" precision="15" seconds="49" value="104.063614">104°03'49&quot;E</geoCoordinate>
; 12.07.2018; leg. H. Freitag and J. Garces; on slide; GBIFCH00592699; RUPP.
</paragraph>
</subSubSection>
</treatment>
</document>