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<document id="CD8D6F2E6B299E78AA614300B4B59C99" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C36BC738E12E7252FB8CFA78" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella luridicollis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="73" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="70" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="B157CDAB2DC3985E15D37E5E705F3342">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="3A22DD685C35F19374702F0CEFA906D5">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="99CF502CBDC0E787FB7A11416630AFF3">2003</mods:date>
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<treatment id="62088784C36BC738E12E7252FB8CFA78" ID-DOI="http://doi.org/10.5281/zenodo.6274679" ID-Zenodo-Dep="6274679" LSID="urn:lsid:plazi:treatment:62088784C36BC738E12E7252FB8CFA78" httpUri="http://treatment.plazi.org/id/62088784C36BC738E12E7252FB8CFA78" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
<subSubSection id="A2BB6519C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70" type="key">
<paragraph id="EA1E3692C36BC735E12E7252FC77F995" blockId="69.[264,973,1603,1630]" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<heading id="B15681FEC36BC735E12E7252FC77F995" bold="true" box="[264,973,1603,1630]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
<emphasis id="D8D5EA80C36BC735E12E7252FC77F995" bold="true" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<keyStep id="51552E37C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<key id="9E3061A3C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<keyLead id="515095A7C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
20.
<taxonomicName id="2DA14D11C36BC735E1127255FD05F995" ID-CoL="6YBD5" authority="Gusarov" authorityName="Gusarov" box="[308,703,1603,1630]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C36BC735E1127255FDF4F996" bold="true" box="[308,590,1604,1630]" italics="true" pageId="69" pageNumber="70">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36BC735E2EA7252FCB5F995" box="[716,783,1603,1629]" pageId="69" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C36BC735E3387252FC7FF995" box="[798,965,1603,1629]" captionStart-0="FIGURES 278 ­ 281" captionStart-1="FIGURES 282 ­ 285" captionStart-2="FIGURES 286 ­ 290" captionStartId-0="70.[264,383,1351,1375]" captionStartId-1="71.[264,383,1111,1135]" captionStartId-2="72.[264,382,1260,1284]" captionTargetBox-0="[308,1276,580,1326]" captionTargetBox-1="[293,1297,523,1086]" captionTargetBox-2="[297,1290,539,1225]" captionTargetId-0="figure@70.[302,1286,569,1332]" captionTargetId-1="figure@71.[282,1306,513,1098]" captionTargetId-2="figure@72.[291,1295,529,1233]" captionTargetPageId-0="70" captionTargetPageId-1="71" captionTargetPageId-2="72" captionText-0="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-2="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156468/files/figure.png" httpUri-1="https://zenodo.org/record/156469/files/figure.png" httpUri-2="https://zenodo.org/record/156470/files/figure.png" pageId="69" pageNumber="70">Figs. 278­290</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E12E729FFBB1F8B8" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="EA1E3692C36BC735E12E729FFECFF930" blockId="69.[264,1324,1678,1904]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E12E729FFE04F960" bold="true" box="[264,446,1678,1704]" pageId="69" pageNumber="70">
<typeStatus id="351A8830C36BC735E12E729FFEFFF960" box="[264,325,1678,1704]" pageId="69" pageNumber="70">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36BC735E1EA729FFD86F960" box="[460,572,1678,1704]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36BC735E24C729FFBA6F960" bold="true" box="[618,1052,1678,1704]" pageId="69" pageNumber="70">
<collectingCountry id="92B67602C36BC735E24C729FFC99F960" box="[618,803,1678,1704]" name="Costa Rica" pageId="69" pageNumber="70">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89 Int. Amer. Hwy.,
<geoCoordinate id="8F955055C36BC735E1D872A7FDC3F918" box="[510,633,1718,1744]" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36BC735E2A772A7FCA8F918" box="[641,786,1718,1744]" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36BC735E33972A7FCC6F918" box="[799,892,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="69" pageNumber="70" unit="m" value="3200.0">3200 m</quantity>
, leaf litter (R.Anderson),
<date id="9E1F1052C36BC735E48B72A7FA91F918" box="[1197,1323,1718,1744]" pageId="69" pageNumber="70" value="1998-06-18">18.vi.1998</date>
(
<collectionCode id="8CB0AE57C36BC735E13672CFFED2F930" box="[272,360,1758,1784]" pageId="69" pageNumber="70">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7317FBB1F8B8" blockId="69.[264,1324,1678,1904]" pageId="69" pageNumber="70">
<typeStatus id="351A8830C36BC735E11E7317FE15F8E8" box="[312,431,1798,1824]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36BC735E1E27317FCC4F8E8" bold="true" box="[452,894,1798,1824]" pageId="69" pageNumber="70">
<collectingCountry id="92B67602C36BC735E1E27317FD3AF8E8" box="[452,640,1798,1824]" name="Costa Rica" pageId="69" pageNumber="70">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
7, 7, same data as the
<typeStatus id="351A8830C36BC735E4CB7317FE84F880" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
;
<emphasis id="D8D5EA80C36BC735E16C733FFE00F880" bold="true" box="[330,442,1838,1864]" pageId="69" pageNumber="70">Cartago:</emphasis>
13, 4, Cerro de la Muerte, Pan American Highway, km 89,
<quantity id="2D599B77C36BC735E4ED733FFA9DF880" box="[1227,1319,1838,1864]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="69" pageNumber="70" unit="m" value="3300.0">3300 m</quantity>
, elfin bamboo forest litter (R.Anderson),
<date id="9E1F1052C36BC735E2C57347FCE0F8B8" box="[739,858,1878,1904]" pageId="69" pageNumber="70" value="1996-02-10">10.ii.1996</date>
(all ­
<collectionCode id="8CB0AE57C36BC735E38E7347FC45F8B8" box="[936,1023,1878,1904]" pageId="69" pageNumber="70">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C368C736E11E750AFBA0F96B" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="EA1E3692C368C736E11E750AFCB4FE36" blockId="70.[264,1324,283,510]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C368C736E1E3750DFD61FEFD" box="[453,731,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E1E3750DFD61FEFD" box="[453,731,284,309]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C368C736E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C368C736E4F5750DFEFDFE95" italics="true" pageId="70" pageNumber="71">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body with contrasting lighter pronotum and elytra; temples 1.4­2.1 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of the male tergum 7 (
<figureCitation id="729A2A17C368C736E10C75ADFE2EFE1E" box="[298,404,444,470]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="70" pageNumber="71">Fig. 278</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C368C736E13775F5FE0CFE36" box="[273,438,484,510]" captionStart-0="FIGURES 282 ­ 285" captionStart-1="FIGURES 286 ­ 290" captionStartId-0="71.[264,383,1111,1135]" captionStartId-1="72.[264,382,1260,1284]" captionTargetBox-0="[293,1297,523,1086]" captionTargetBox-1="[297,1290,539,1225]" captionTargetId-0="figure@71.[282,1306,513,1098]" captionTargetId-1="figure@72.[291,1295,529,1233]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-1="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156469/files/figure.png" httpUri-1="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Figs. 282­289</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C368C736E2BC75F5FCB8FE36" box="[666,770,484,510]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
<caption id="BEDE661AC368C736E12E7156FC91FA6B" httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="70" pageNumber="71" targetBox="[308,1276,580,1326]" targetPageId="70">
<paragraph id="EA1E3692C368C736E12E7156FC91FA6B" blockId="70.[264,1323,1351,1443]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E12E7156FE65FA97" bold="true" box="[264,479,1351,1375]" pageId="70" pageNumber="71">FIGURES 278­281</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C368C736E32C7156FBD0FA96" authority="Gusarov" authorityName="Gusarov" box="[778,1130,1351,1374]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C368C736E32C7156FBBDFA96" box="[778,1031,1351,1374]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC368C736E4517156FB09FA97" box="[1143,1203,1351,1375]" pageId="70" pageNumber="71" rank="species">
<emphasis id="D8D5EA80C368C736E4517156FB09FA97" bold="true" box="[1143,1203,1351,1375]" pageId="70" pageNumber="71">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C368C736E11E71FBFBA0F96B" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<taxonomicName id="2DA14D11C368C736E11E71FBFDF5F9CB" box="[312,591,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E11E71FBFDF5F9CB" box="[312,591,1514,1539]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C368C736E2CB71FBFC35F9CB" box="[749,911,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C368C736E2CB71FBFC35F9CB" box="[749,911,1514,1539]" italics="true" pageId="70" pageNumber="71">S. curtipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C368C736E3EC71FBFBDAF9CB" box="[970,1120,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C368C736E3EC71FBFBDAF9CB" box="[970,1120,1514,1539]" italics="true" pageId="70" pageNumber="71">S. lativentris</emphasis>
</taxonomicName>
in having glossy pronotum with weak microsculpture; tergum 8 with four pairs of macrosetae; medial lamellae of internal sac short (
<figureCitation id="729A2A17C368C736E2AA722BFD40F99C" box="[652,762,1594,1620]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 288</figureCitation>
), the apices of lamellae not protruding from retracted sac (
<figureCitation id="729A2A17C368C736E1897273FDA2F9B4" box="[431,536,1634,1660]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Fig. 282</figureCitation>
); the apex of median lobe in parameral view broad, without basal constriction (
<figureCitation id="729A2A17C368C736E184729BFDF1F96C" box="[418,587,1674,1700]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Figs. 282­283</figureCitation>
); and S­shaped spermatheca (
<figureCitation id="729A2A17C368C736E381729BFBB1F96C" box="[935,1035,1674,1700]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C368C738E11E72A3FDC4FEFE" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71" type="description">
<paragraph id="EA1E3692C368C736E11E72A3FAA3F93B" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<taxonomicName id="2DA14D11C368C736E11E72A3FDEAF903" box="[312,592,1714,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E11E72A3FDEAF903" box="[312,592,1714,1739]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C368C736E2D772A3FC19F903" box="[753,931,1714,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C368C736E2D772A3FC19F903" box="[753,931,1714,1739]" italics="true" pageId="70" pageNumber="71">S. micralymma</emphasis>
</taxonomicName>
in having wider apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C368C736E21E72CBFD65F93C" box="[568,735,1754,1780]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Figs. 282­283</figureCitation>
; 295­296) and S­shaped spermatheca (
<figureCitation id="729A2A17C368C736E48272CBFAB3F93C" box="[1188,1289,1754,1780]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C368C736E11E7310FE28F8A4" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E11E7310FE6AF8D3" bold="true" box="[312,464,1793,1819]" pageId="70" pageNumber="71">Description.</emphasis>
Length 2.0­
<quantity id="2D599B77C368C736E24E7313FD7DF8D4" box="[616,711,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="70" pageNumber="71" unit="mm" value="2.5">2.5 mm</quantity>
. Head and abdomen dark brown; pronotum reddish brown to brownish orange; elytra brown to brownish yellow; antennae, mouthparts and legs brown.
</paragraph>
<paragraph id="EA1E3692C369C737E11E750DFC62FE66" blockId="71.[264,1325,284,430]" pageId="71" pageNumber="72">Head surface glossy, with fine and weak isodiametric microsculpture, and fine and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Temples 1.4­2.1 times as long as eyes. Antennal article 2 as long as 3, articles 4­5 elongate, 6­7 subquadrate, 8­10 slightly transverse (ratio 1.5­2.0).</paragraph>
<caption id="BEDE661AC369C737E12E7046FEFAFB1F" httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="71" pageNumber="72" targetBox="[293,1297,523,1086]" targetPageId="71">
<paragraph id="EA1E3692C369C737E12E7046FEFAFB1F" blockId="71.[264,1323,1111,1239]" pageId="71" pageNumber="72">
<emphasis id="D8D5EA80C369C737E12E7046FE59FBA7" bold="true" box="[264,483,1111,1135]" pageId="71" pageNumber="72">FIGURES 282­285</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C369C737E2BE7049FBBBFBA7" authority="Gusarov" authorityName="Gusarov" box="[664,1025,1112,1135]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C369C737E2BE7049FC23FBA7" box="[664,921,1112,1135]" italics="true" pageId="71" pageNumber="72">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC369C737E4367046FBEBFBA7" box="[1040,1105,1111,1135]" pageId="71" pageNumber="72" rank="species">
<emphasis id="D8D5EA80C369C737E4367046FBEBFBA7" bold="true" box="[1040,1105,1111,1135]" pageId="71" pageNumber="72">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285).
</paragraph>
</caption>
<paragraph id="EA1E3692C369C737E11E7133FB52F9E4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Pronotum slightly transverse, 1.1 times as wide as head, width
<quantity id="2D599B77C369C737E40D7133FB6CFAF4" box="[1067,1238,1314,1340]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="71" pageNumber="72" unit="mm" value="0.485" valueMax="0.51" valueMin="0.46">0.46­0.51 mm</quantity>
, length
<quantity id="2D599B77C369C737E12E715BFE0BFAAC" box="[264,433,1354,1380]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="71" pageNumber="72" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.40­0.47 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, and fine and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.6 times wider than long, glossy, with fine and weak isodiametric microsculpture, and fine punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C369C737E11E722BFC18F904" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­3 times their diameter on terga 3­5 and 2­4 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C369C737E2C172A3FC2FF904" box="[743,917,1714,1740]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="71" pageNumber="72">Figs. 278, 280</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C369C737E11E72CBFBD4F8D4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
In males posterior angles of tergum 3 not projecting; tergum 7 without medial carina or tubercle; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C369C737E3DC7313FBD8F8D4" box="[1018,1122,1794,1820]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="71" pageNumber="72">Fig. 278</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C369C737E11E733BFD48F8A4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Aedeagus as in
<figureCitation id="729A2A17C369C737E1C8733BFD2EF88C" box="[494,660,1834,1860]" captionStart-0="FIGURES 282 ­ 285" captionStart-1="FIGURES 286 ­ 290" captionStartId-0="71.[264,383,1111,1135]" captionStartId-1="72.[264,382,1260,1284]" captionTargetBox-0="[293,1297,523,1086]" captionTargetBox-1="[297,1290,539,1225]" captionTargetId-0="figure@71.[282,1306,513,1098]" captionTargetId-1="figure@72.[291,1295,529,1233]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-1="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156469/files/figure.png" httpUri-1="https://zenodo.org/record/156470/files/figure.png" pageId="71" pageNumber="72">Figs. 282­289</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C369C737E2587343FD5FF8A4" box="[638,741,1874,1900]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="71" pageNumber="72">Fig. 289</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C366C738E11E750DFDC4FEFE" blockId="72.[264,1322,284,430]" box="[312,638,284,310]" pageId="72" pageNumber="73">
Spermatheca as in
<figureCitation id="729A2A17C366C738E232750DFDC0FEFE" box="[532,634,284,310]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="72" pageNumber="73">Fig. 290</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C366C738E11E7552FB8CFA78" pageId="72" pageNumber="73" type="distribution">
<paragraph id="EA1E3692C366C738E11E7552FE17FE66" blockId="72.[264,1322,284,430]" pageId="72" pageNumber="73">
<emphasis id="D8D5EA80C366C738E11E7552FE62FE95" bold="true" box="[312,472,323,349]" pageId="72" pageNumber="73">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C366C738E3547555FC48FE96" box="[882,1010,324,350]" name="Costa Rica" pageId="72" pageNumber="73">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C366C738E4247555FBD3FE96" box="[1026,1129,324,350]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="72" pageNumber="73">Fig. 389</figureCitation>
).
<emphasis id="D8D5EA80C366C738E11E757AFDBDFE4D" bold="true" box="[312,519,363,389]" pageId="72" pageNumber="73">Natural History.</emphasis>
<taxonomicName id="2DA14D11C366C738E23C757DFC88FE4D" box="[538,818,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C366C738E23C757DFC88FE4D" box="[538,818,364,389]" italics="true" pageId="72" pageNumber="73">Seeversiella luridicollis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C366C738E12E7585FE12FE66" box="[264,424,404,430]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="72" pageNumber="73" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">3200­3300 m</quantity>
.
</paragraph>
<caption id="BEDE661AC366C738E12E70FDFEFAFA46" httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="72" pageNumber="73" targetBox="[297,1290,539,1225]" targetPageId="72">
<paragraph id="EA1E3692C366C738E12E70FDFEFAFA46" blockId="72.[264,1322,1260,1456]" pageId="72" pageNumber="73">
<emphasis id="D8D5EA80C366C738E12E70FDFE67FACC" bold="true" box="[264,477,1260,1284]" pageId="72" pageNumber="73">FIGURES 286­290</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C366C738E25270FCFC6BFACB" authority="Gusarov" authorityName="Gusarov" box="[628,977,1260,1284]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C366C738E25270FCFCCAFACC" box="[628,880,1261,1284]" italics="true" pageId="72" pageNumber="73">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C366C738E3FD70FDFBADFACC" bold="true" box="[987,1047,1260,1284]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="C3E657FBC366C738E3FD70FDFBADFACC" box="[987,1047,1260,1284]" pageId="72" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(286, 289, holotype; 287­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287­ 290).
</paragraph>
</caption>
<paragraph id="EA1E3692C366C738E12E7188FB8CFA78" blockId="72.[264,1322,1260,1456]" box="[264,1078,1433,1456]" pageId="72" pageNumber="73">ML medial lamellae; SLD sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
</treatment>
</document>