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<document id="24DD095DE3D4B50DC20A29F21A463C5F" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C362C73EE12E750AFE5DF9C6" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella impressicollis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="79" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="77" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="43CB2D7B546D52CF95DD091CA37CFE25">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77" type="key">
<paragraph id="EA1E3692C362C73CE12E750AFC4AFEFD" blockId="76.[264,1008,283,310]" box="[264,1008,283,310]" pageId="76" pageNumber="77">
<heading id="B15681FEC362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="D8D5EA80C362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" pageId="76" pageNumber="77">
<key id="9E3061A3C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77">
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<keyLead id="515095A7C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77">
22.
<taxonomicName id="2DA14D11C362C73CE112750DFD58FEFD" ID-CoL="6YBCW" authority="Gusarov" authorityName="Gusarov" box="[308,738,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C362C73CE112750DFDCAFEFE" bold="true" box="[308,624,284,310]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC362C73CE2C9750AFC8BFEFD" box="[751,817,283,309]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C362C73CE366750AFC52FEFD" box="[832,1000,283,309]" captionStart-0="FIGURES 307 ­ 310" captionStart-1="FIGURES 311 ­ 320" captionStartId-0="76.[264,383,1792,1816]" captionStartId-1="77.[264,383,1623,1647]" captionTargetBox-0="[290,1292,893,1759]" captionTargetBox-1="[286,1302,571,1605]" captionTargetId-0="figure@76.[282,1305,884,1766]" captionTargetId-1="figure@77.[276,1310,569,1613]" captionTargetPageId-0="76" captionTargetPageId-1="77" captionText-0="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri-0="https://zenodo.org/record/156474/files/figure.png" httpUri-1="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 307­320</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C362C73CE12E7577FECFFDE9" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="EA1E3692C362C73CE12E7577FECFFE19" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="76" pageNumber="77">
<typeStatus id="351A8830C362C73CE12E7577FEF9FE48" box="[264,323,358,384]" pageId="76" pageNumber="77">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C362C73CE1F17576FDFDFE49" box="[471,583,359,385]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C362C73CE25D7577FC79FE48" bold="true" box="[635,963,358,384]" pageId="76" pageNumber="77">
<collectingCountry id="92B67602C362C73CE25D7577FC82FE48" box="[635,824,358,384]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Cerro Chirripó, Valle de los Conelos,
<geoCoordinate id="8F955055C362C73CE15C759EFE4FFE61" box="[378,501,399,425]" direction="north" orientation="latitude" pageId="76" pageNumber="77" precision="15" value="9.476944">9°28'37&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C362C73CE1D9759EFD35FE61" box="[511,655,399,425]" direction="west" orientation="longitude" pageId="76" pageNumber="77" precision="15" value="-83.489716">83°29'23&quot;W</geoCoordinate>
,
<quantity id="2D599B77C362C73CE2B9759EFD44FE61" box="[671,766,399,425]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="76" pageNumber="77" unit="m" value="3600.0">3600 m</quantity>
, paramo shrub litter (R.Anderson),
<date id="9E1F1052C362C73CE48B759EFA90FE61" box="[1197,1322,399,425]" pageId="76" pageNumber="77" value="1999-06-26">26.vi.1999</date>
(
<collectionCode id="8CB0AE57C362C73CE13675A6FED2FE19" box="[272,360,439,465]" pageId="76" pageNumber="77">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C362C73CE11E75CEFECFFDE9" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<typeStatus id="351A8830C362C73CE11E75CEFE15FE31" box="[312,431,479,505]" pageId="76" pageNumber="77" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C362C73CE1ED75CFFCA2FE30" bold="true" box="[459,792,478,504]" pageId="76" pageNumber="77">
<collectingCountry id="92B67602C362C73CE1ED75CFFD33FE30" box="[459,649,478,504]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC362C73CE31775CEFC62FE31" box="[817,984,479,505]" pageId="76" pageNumber="77" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C362C73CE4E575CEFA90FE31" box="[1219,1322,479,505]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C362C73CE1367616FED2FDE9" box="[272,360,519,545]" pageId="76" pageNumber="77">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C362C73EE11E763FFE3BFE1E" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="EA1E3692C362C73CE11E763FFD78FCD9" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE11E763FFE0DFD80" bold="true" box="[312,439,558,584]" pageId="76" pageNumber="77">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C362C73CE4DE763EFED7FDB8" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C362C73CE4DE763EFED7FDB8" italics="true" pageId="76" pageNumber="77">Seeversiella</emphasis>
</taxonomicName>
by having brown or dark brown body with lighter pronotum and elytra; temples 1.8­2.2 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of male tergum 7 (
<figureCitation id="729A2A17C362C73CE13776DEFECDFD21" box="[273,375,719,745]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77">Fig. 307</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C362C73CE4D676DEFED0FCD9" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 311­319</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C362C73CE26876E6FD0CFCD9" box="[590,694,759,785]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Fig. 320</figureCitation>
).
</paragraph>
<caption id="BEDE661AC362C73CE12E7311FC2CF895" httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77" targetBox="[290,1292,893,1759]" targetPageId="76">
<paragraph id="EA1E3692C362C73CE12E7311FC2CF895" blockId="76.[264,1323,1792,1885]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7311FE52F8D0" bold="true" box="[264,488,1792,1816]" pageId="76" pageNumber="77">FIGURES 307­310</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C362C73CE3647310FB6BF8D0" authority="Gusarov" authorityName="Gusarov" box="[834,1233,1793,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C362C73CE3647310FBDCF8D0" box="[834,1126,1793,1816]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C362C73CE4C37311FA91F8D0" bold="true" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="C3E657FBC362C73CE4C37311FA91F8D0" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE11E750DFD43FE36" blockId="77.[264,1323,284,510]" pageId="77" pageNumber="78">
<taxonomicName id="2DA14D11C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" italics="true" pageId="77" pageNumber="78">S. sulcicollis</emphasis>
</taxonomicName>
in having smaller body; in males pronotum with broad medial impression, posterior margin of tergum 3 not projecting, tergum 7 without medial carina and posterior margin of tergum 8 without medial denticles (
<figureCitation id="729A2A17C363C73DE1377585FEC2FE66" box="[273,376,404,430]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="77" pageNumber="78">Fig. 307</figureCitation>
); apex of median lobe in parameral view without carinae (
<figureCitation id="729A2A17C363C73DE40D7585FB69FE66" box="[1067,1235,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 311­312</figureCitation>
); copulatory piece of internal sac shorter (
<figureCitation id="729A2A17C363C73DE28975ADFC99FE1E" box="[687,803,444,470]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 317</figureCitation>
; 326, 328); sclerites of lateral diverticula of internal sac more slender (
<figureCitation id="729A2A17C363C73DE21875F5FD15FE36" box="[574,687,484,510]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 318</figureCitation>
; 326).
</paragraph>
<caption id="BEDE661AC363C73DE12E7246FEFAF8F7" httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78" targetBox="[286,1302,571,1605]" targetPageId="77">
<paragraph id="EA1E3692C363C73DE12E7246FEFAF8F7" blockId="77.[264,1323,1623,1891]" pageId="77" pageNumber="78">
<emphasis id="D8D5EA80C363C73DE12E7246FE67F9A7" bold="true" box="[264,477,1623,1647]" pageId="77" pageNumber="78">FIGURES 311­320</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C363C73DE2537249FC4BF9A7" authority="Gusarov" authorityName="Gusarov" box="[629,1009,1624,1647]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C363C73DE2537249FC2AF9A7" box="[629,912,1624,1647]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C363C73DE3DA7246FB82F9A7" bold="true" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="C3E657FBC363C73DE3DA7246FB82F9A7" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312­313, 315, 317, 319­ 320).
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE12E735DFA91F8AB" blockId="77.[264,1323,1623,1891]" box="[264,1323,1868,1891]" pageId="77" pageNumber="78">CP copulatory piece; ML medial lamellae; SLD sclerite of lateral diverticulum of internal sac.</paragraph>
<paragraph id="EA1E3692C360C73EE11E750DFE3BFE1E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<taxonomicName id="2DA14D11C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" italics="true" pageId="78" pageNumber="79">S. microphthalma</emphasis>
</taxonomicName>
in having pronotum with more asperate punctation; in males pronotum with broad medial impression, posterior angles of tergum 3 not projecting as spines; lateral diverticula of internal sac with long spiniform sclerites (
<figureCitation id="729A2A17C360C73EE1DE7585FD11FE66" box="[504,683,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 316, 318</figureCitation>
; 343); spermatheca with smaller distal portion (
<figureCitation id="729A2A17C360C73EE4D77585FE8DFE1E" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 320</figureCitation>
; 341).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E75F2FDC4FA5E" pageId="78" pageNumber="79" type="description">
<paragraph id="EA1E3692C360C73EE11E75F2FD37FDEE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E75F2FE6AFE35" bold="true" box="[312,464,483,509]" pageId="78" pageNumber="79">Description.</emphasis>
Length
<quantity id="2D599B77C360C73EE21A75F5FD73FE36" box="[572,713,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="78" pageNumber="79" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2­2.7 mm</quantity>
. Body brown or dark brown with lighter pronotum and elytra; legs brownish yellow.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7625FCB1FD0E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals 2 times their diameter. Temples 1.8­2.2 times as long as eyes. Antennal article 2 longer than 3, article 4 subquadrate, 5­10 transverse (ratio 1.5­1.8).</paragraph>
<paragraph id="EA1E3692C360C73EE11E76C5FE5FFBCE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C360C73EE40D76C5FB6CFD26" box="[1067,1238,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.699999999999999" pageId="78" pageNumber="79" unit="mm" value="0.5" valueMax="0.53" valueMin="0.47">0.47­0.53 mm</quantity>
, length
<quantity id="2D599B77C360C73EE12E76EDFE0BFCDE" box="[264,433,764,790]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.9" metricValueMin="4.3" pageId="78" pageNumber="79" unit="mm" value="0.45999999999999996" valueMax="0.49" valueMin="0.43">0.43­0.49 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.5), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7005FC18FB6E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C360C73EE2C1709DFC2FFB6E" box="[743,917,1164,1190]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Figs. 307, 309</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E70A5FD9CFAD6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
In males pronotum with wide medial impression; posterior angles of tergum 3 not projecting; tergum 7 without medial carina or tubercle; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C360C73EE1947115FDA0FAD6" box="[434,538,1284,1310]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Fig. 307</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E713DFD48FAA6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Aedeagus as in
<figureCitation id="729A2A17C360C73EE1C9713DFD2EFA8E" box="[495,660,1324,1350]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 311­319</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C360C73EE2587145FD5FFAA6" box="[638,741,1364,1390]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 313</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E716DFDC4FA5E" blockId="78.[264,1324,284,1550]" box="[312,638,1404,1430]" pageId="78" pageNumber="79">
Spermatheca as in
<figureCitation id="729A2A17C360C73EE232716DFDC0FA5E" box="[532,634,1404,1430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 320</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E71B2FE5DF9C6" pageId="78" pageNumber="79" type="distribution">
<paragraph id="EA1E3692C360C73EE11E71B2FBFEFA76" blockId="78.[264,1324,284,1550]" box="[312,1092,1443,1470]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71B2FE62FA75" bold="true" box="[312,472,1443,1469]" pageId="78" pageNumber="79">Distribution.</emphasis>
Known from Cerro Chirripó,
<collectingCountry id="92B67602C360C73EE36671B5FC7AFA76" box="[832,960,1444,1470]" name="Costa Rica" pageId="78" pageNumber="79">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C360C73EE3F671B5FB82FA76" box="[976,1080,1444,1470]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="78" pageNumber="79">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E71DAFE5DF9C6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71DAFDBDFA2D" bold="true" box="[312,519,1483,1509]" pageId="78" pageNumber="79">Natural History.</emphasis>
<taxonomicName id="2DA14D11C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
was collected in paramo shrub litter at altitude of
<quantity id="2D599B77C360C73EE1A171E5FE5EF9C6" box="[391,484,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="78" pageNumber="79" unit="m" value="3600.0">3600 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>