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<document id="084AB8774F8EAB65BC4AA2FB75608763" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C313C74FE12E761AFE17FCF6" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella brunnea Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="62" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="5147867AD0F8D8A6E4E8163DB0D7D599">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="EA1E3692C313C74DE12E761AFC08FDED" blockId="61.[264,946,523,550]" box="[264,946,523,550]" pageId="61" pageNumber="62">
<heading id="B15681FEC313C74DE12E761AFC08FDED" bold="true" box="[264,946,523,550]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
<emphasis id="D8D5EA80C313C74DE12E761AFC08FDED" bold="true" box="[264,946,523,550]" pageId="61" pageNumber="62">
<keyStep id="51552E37C313C74DE12E761AFC08FDED" box="[264,946,523,550]" pageId="61" pageNumber="62">
<key id="9E3061A3C313C74DE12E761AFC08FDED" box="[264,946,523,550]" pageId="61" pageNumber="62">
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17.
<taxonomicName id="2DA14D11C313C74DE112761DFD1EFDED" ID-CoL="6YBCR" authority="Gusarov" authorityName="Gusarov" box="[308,676,523,550]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C313C74DE112761DFD88FDEE" bold="true" box="[308,562,524,550]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC313C74DE297761AFD49FDED" box="[689,755,523,549]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C313C74DE324761AFC10FDED" box="[770,938,523,549]" captionStart-0="FIGURES 235 ­ 238" captionStart-1="FIGURES 239 ­ 246" captionStartId-0="61.[264,382,1659,1683]" captionStartId-1="62.[264,383,1630,1654]" captionTargetBox-0="[313,1272,852,1625]" captionTargetBox-1="[313,1277,660,1592]" captionTargetId-0="figure@61.[304,1283,844,1632]" captionTargetId-1="figure@62.[302,1285,649,1603]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURES 235 ­ 238. Abdominal segment 8 of Seeversiella brunnea Gusarov, sp. n. (235 ­ 236, holotype; 237 ­ 238, paratype, P. N. Tapantí, Costa Rica). 235 male tergum 8; 236 male sternum 8; 237 female tergum 8; 238 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri-0="https://zenodo.org/record/156461/files/figure.png" httpUri-1="https://zenodo.org/record/156462/files/figure.png" pageId="61" pageNumber="62">Figs. 235­246</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C313C74DE12E7647FE29FD21" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="EA1E3692C313C74DE12E7647FB3BFD51" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E7647FE78FDB8" bold="true" box="[264,450,598,624]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE12E7647FEF9FDB8" box="[264,323,598,624]" pageId="61" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C313C74DE1C67646FDEAFDB9" box="[480,592,599,625]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C313C74DE2AC7647FC61FDB8" bold="true" box="[650,987,598,624]" pageId="61" pageNumber="62">
<collectingCountry id="92B67602C313C74DE2AC7647FCF0FDB8" box="[650,842,598,624]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
P. N. Tapantí,
<geoCoordinate id="8F955055C313C74DE4897646FA91FDB9" box="[1199,1323,599,625]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="15" value="9.75">9°45'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C313C74DE12E766EFE23FD51" box="[264,409,639,665]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.816666">83°49'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C313C74DE182766EFDBAFD51" box="[420,512,639,665]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="62" unit="m" value="1500.0">1500 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C313C74DE3B8766EFBB7FD51" box="[926,1037,639,665]" pageId="61" pageNumber="62" value="1997-06-04">4.vi.1997</date>
(
<collectionCode id="8CB0AE57C313C74DE43A766EFBCEFD51" box="[1052,1140,639,665]" pageId="61" pageNumber="62">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C313C74DE11E76B6FE29FD21" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE11E76B6FE15FD09" box="[312,431,679,705]" pageId="61" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C313C74DE1E576B7FCB9FD08" bold="true" box="[451,771,678,704]" pageId="61" pageNumber="62">
<collectingCountry id="92B67602C313C74DE1E576B7FDC7FD08" box="[451,637,678,704]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
,, ditto but,
<geoCoordinate id="8F955055C313C74DE3E976B6FBF0FD09" box="[975,1098,679,705]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="15" value="9.741667">9°44'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C313C74DE47476B6FB59FD09" box="[1106,1251,679,705]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.808334">83°48'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C313C74DE4D676B6FEA5FD21" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="61" pageNumber="62" unit="m" value="1740.0">1740 m</quantity>
(
<collectionCode id="8CB0AE57C313C74DE10876DEFE3CFD21" box="[302,390,719,745]" pageId="61" pageNumber="62">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC313C74DE12E726AFC4EF91F" httpUri="https://zenodo.org/record/156461/files/figure.png" pageId="61" pageNumber="62" targetBox="[313,1272,852,1625]" targetPageId="61">
<paragraph id="EA1E3692C313C74DE12E726AFC4EF91F" blockId="61.[264,1323,1659,1751]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E726AFE53F95B" bold="true" box="[264,489,1659,1683]" pageId="61" pageNumber="62">FIGURES 235­238.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C313C74DE338726AFBD2F95A" authority="Gusarov" authorityName="Gusarov" box="[798,1128,1659,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C313C74DE338726AFBB8F95A" box="[798,1026,1659,1682]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C313C74DE451726AFB0DF95B" bold="true" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="C3E657FBC313C74DE451726AFB0DF95B" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(235­236, holotype; 237­238, paratype, P. N. Tapantí, Costa Rica). 235 male tergum 8; 236 male sternum 8; 237 female tergum 8; 238 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74EE11E733CFB24FE1E" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="EA1E3692C313C74EE11E733CFC51FE4E" blockId="61.[264,1323,1837,1904]" lastBlockId="62.[264,1323,284,590]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E733CFE0DF88F" bold="true" box="[312,439,1837,1863]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C313C74DE4F5733FFEFDF8A7" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C313C74DE4F5733FFEFDF8A7" italics="true" pageId="61" pageNumber="62">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 0.8­1.2 times as long as eyes; matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or partially reduced, 2­4 times as long as elytra; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C310C74EE1CB757DFD29FE4E" box="[493,659,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 239­245</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C310C74EE351757DFC64FE4E" box="[887,990,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Fig. 246</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C310C74EE11E7585FB24FE1E" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" italics="true" pageId="62" pageNumber="63">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" italics="true" pageId="62" pageNumber="63">S. furcativentris</emphasis>
</taxonomicName>
in having concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE32075ADFC14FE1E" box="[774,942,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 197­198; 214­215).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C310C74FE11E75F5FDC4FE1E" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
<paragraph id="EA1E3692C310C74EE11E75F5FD68FD86" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" italics="true" pageId="62" pageNumber="63">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; and concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE18D7625FDE9FD86" box="[427,595,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 228­229).
</paragraph>
<caption id="BEDE661AC310C74EE12E724FFD50F8EA" httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63" targetBox="[313,1277,660,1592]" targetPageId="62">
<paragraph id="EA1E3692C310C74EE12E724FFD50F8EA" blockId="62.[264,1324,1630,1863]" pageId="62" pageNumber="63">
<emphasis id="D8D5EA80C310C74EE12E724FFE5AF9BE" bold="true" box="[264,480,1630,1654]" pageId="62" pageNumber="63">FIGURES 239­246</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C310C74EE2A4724FFC72F9BD" authority="Gusarov" authorityName="Gusarov" box="[642,968,1630,1653]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C310C74EE2A4724FFCDEF9BD" box="[642,868,1630,1653]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C310C74EE3F3724FFBA8F9BE" bold="true" box="[981,1042,1630,1654]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="C3E657FBC310C74EE3F3724FFBA8F9BE" box="[981,1042,1630,1654]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(239­244, holotype; 245­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246).
</paragraph>
</caption>
<paragraph id="EA1E3692C310C74EE12E7321FE47F88F" blockId="62.[264,1324,1630,1863]" box="[264,509,1840,1863]" pageId="62" pageNumber="63">CB connecting band.</paragraph>
<paragraph id="EA1E3692C311C74FE11E750AFB5BFEFE" blockId="63.[264,1323,283,830]" box="[312,1249,283,310]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="63" pageNumber="64">Description.</emphasis>
Length
<quantity id="2D599B77C311C74FE21F750DFD7EFEFE" box="[569,708,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="63" pageNumber="64" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Pronotal width
<quantity id="2D599B77C311C74FE3AA750DFB8FFEFE" box="[908,1077,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.0" metricValueMin="4.4" pageId="63" pageNumber="64" unit="mm" value="0.47" valueMax="0.5" valueMin="0.44">0.44­0.50 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7555FBEFFE4E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
In all character states
<taxonomicName id="2DA14D11C311C74FE2117555FD0CFE95" box="[567,694,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2117555FD0CFE95" box="[567,694,324,349]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7585FD26FE66" blockId="63.[264,1323,283,830]" box="[312,668,404,430]" pageId="63" pageNumber="64">
Aedeagus as in
<figureCitation id="729A2A17C311C74FE1D77585FD2DFE66" box="[497,663,404,430]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­245</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E75ADFDC4FE1E" blockId="63.[264,1323,283,830]" box="[312,638,444,470]" pageId="63" pageNumber="64">
Spermatheca as in
<figureCitation id="729A2A17C311C74FE23375ADFDC1FE1E" box="[533,635,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Fig. 246</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E75F2FBBFFD0E" pageId="63" pageNumber="64" type="discussion">
<paragraph id="EA1E3692C311C74FE11E75F2FBBFFD0E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E75F2FE79FE35" bold="true" box="[312,451,483,509]" pageId="63" pageNumber="64">Discussion.</emphasis>
Despite the similarity of
<taxonomicName id="2DA14D11C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
to
<taxonomicName id="2DA14D11C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
in external characters and the shape of genitalia, the difference between
<taxonomicName id="2DA14D11C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
and the two species in the shape of the apex of median lobe (
<figureCitation id="729A2A17C311C74FE35B7625FB9FFD86" box="[893,1061,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­242</figureCitation>
; 194­198; 212­215) is of the same scale as the difference between
<taxonomicName id="2DA14D11C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
. Since no intermediate forms have been found,
<taxonomicName id="2DA14D11C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is considered to represent a separate species, possibly a local endemic of the area near Tapantí (
<figureCitation id="729A2A17C311C74FE3B776BDFC42FD0E" box="[913,1016,684,710]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E76C2FE17FCF6" pageId="63" pageNumber="64" type="distribution">
<paragraph id="EA1E3692C311C74FE11E76C2FB29FD26" blockId="63.[264,1323,283,830]" box="[312,1171,723,750]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76C2FE62FD25" bold="true" box="[312,472,723,749]" pageId="63" pageNumber="64">Distribution.</emphasis>
Known from the area near Tapantí,
<collectingCountry id="92B67602C311C74FE3A976C5FBB5FD26" box="[911,1039,724,750]" name="Costa Rica" pageId="63" pageNumber="64">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C311C74FE43976C5FB3DFD26" box="[1055,1159,724,750]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C311C74FE11E76EAFE17FCF6" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76EAFDB0FCDD" bold="true" box="[312,522,763,789]" pageId="63" pageNumber="64">Natural History.</emphasis>
<taxonomicName id="2DA14D11C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" italics="true" pageId="63" pageNumber="64">Seeversiella brunnea</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C311C74FE12E7735FE12FCF6" box="[264,424,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="1.7399999999999998" metricValueMin="1.5" pageId="63" pageNumber="64" unit="m" value="1620.0" valueMax="1740.0" valueMin="1500.0">1500­1740 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>