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<document id="E6B4FAEC02EC2B0115F4DF130B6FE1D0" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFB9FFD5FD9DEFF5FAD21272" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor major Enghoff, in Enghoff &amp; Frederiksen 2015" docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="16" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1FFD8D0D3E9FE6370DCE33DDE23B9E94">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:name id="469736CE64B37D2C245D941BA0FC2800" type="personal">
<mods:role id="292F0C1D5561673B8CF71F1A1894B77F">
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<mods:namePart id="CBCFB71E395C544F5D51A226377DC633">Enghoff, Henrik</mods:namePart>
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<mods:title id="14465484782A988EE9BE61B48F1E05EE">European Journal of Taxonomy</mods:title>
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<mods:part id="FB601EA36C3A29014520A03C21DE7DD7">
<mods:date id="E43B315CA57F097AFA2401001B12F502">2020</mods:date>
<mods:detail id="E07076A92F8DFBA76FCA9C108D70999A" type="pubDate">
<mods:number id="6C18090AA0948609D0DA065EBF60BD0D">2020-04-14</mods:number>
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<mods:number id="5B94B965712DE0E2BD798E6BD3225C54">626</mods:number>
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<mods:start id="1B5F546E375B0FD960E2DF77ADF395DE">1</mods:start>
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<mods:classification id="30265C3CF89CFD9DC9BBEDAF6B469C7D">journal article</mods:classification>
<mods:identifier id="61BECA461E625A894A278F19419F4B3E" type="DOI">10.5852/ejt.2020.626</mods:identifier>
<mods:identifier id="A870132F25E2533CF50673C381FFCFFB" type="GBIF-Dataset">c906d9cd-cfcb-43db-a24c-b57b39d9184b</mods:identifier>
<mods:identifier id="BCB4CB4BF997D8794AF2030841FA6712" type="Zenodo-Dep">3754989</mods:identifier>
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<treatment id="5C11879EFFB9FFD5FD9DEFF5FAD21272" ID-DOI="http://doi.org/10.5281/zenodo.3808350" ID-GBIF-Taxon="163642162" ID-Zenodo-Dep="3808350" LSID="urn:lsid:plazi:treatment:5C11879EFFB9FFD5FD9DEFF5FAD21272" httpUri="http://treatment.plazi.org/id/5C11879EFFB9FFD5FD9DEFF5FAD21272" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="9CA26503FFB9FFD5FD9DEFF5FC1F16F3" box="[604,984,692,719]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="D4073688FFB9FFD5FD9DEFF5FC1F16F3" blockId="15.[604,984,692,755]" box="[604,984,692,719]" pageId="15" pageNumber="16">
<heading id="8F4F81E4FFB9FFD5FD9DEFF5FC1F16F3" box="[604,984,692,719]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="13B84D0BFFB9FFD5FD9DEFF5FC1F16F3" ID-CoL="FZL3" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[604,984,692,719]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="major">
<emphasis id="E6CCEA9AFFB9FFD5FD9DEFF5FCE016F3" bold="true" box="[604,807,693,719]" italics="true" pageId="15" pageNumber="16">Aquattuor major</emphasis>
<bibRefCitation id="B0294B79FFB9FFD5FCEFEFF4FC1F16F3" author="Enghoff H. &amp; Frederiksen S. B." box="[814,984,692,719]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFB9FFD5FD7DEF98FCB016CE" box="[700,887,728,755]" pageId="15" pageNumber="16" type="description">
<paragraph id="D4073688FFB9FFD5FD7DEF98FCB016CE" blockId="15.[604,984,692,755]" box="[700,887,728,755]" pageId="15" pageNumber="16">
<figureCitation id="4C832A0DFFB9FFD5FD7DEF98FCC216CE" box="[700,773,728,754]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="15" pageNumber="16">Figs 1</figureCitation>
,
<figureCitation id="4C832A0DFFB9FFD5FCD5EF98FCE616CF" box="[788,801,728,755]" captionStart="Fig" captionStartId="7.[189,232,1887,1913]" captionTargetBox="[234,1343,270,1842]" captionTargetId="figure@7.[229,1354,262,1848]" captionTargetPageId="7" captionText="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754999" httpUri="https://zenodo.org/record/3754999/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
A–C, 9
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFB9FFD5FF7CEE5AFC65170F" box="[189,930,793,820]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="D4073688FFB9FFD5FF7CEE5AFC65170F" blockId="15.[189,930,793,820]" box="[189,930,793,820]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="F4A811A6FFB9FFD5FF7CEE5AFC65170F" box="[189,930,793,820]" pageId="15" pageNumber="16">
<treatmentCitation id="55191099FFB9FFD5FF7CEE5AFC59170F" ID-DOI="http://doi.org/10.5281/zenodo.3793779" ID-Zenodo-Dep="3793779" LSID="urn:lsid:zoobank.org:act:7DE65287-B22D-4470-A2AF-70556B77DEE9" author="Enghoff H. &amp; Frederiksen S. B." box="[189,926,793,820]" httpUri="http://treatment.plazi.org/id/03A487F2FFF0FFB7750F6B2F961DB558" page="10" pageId="15" pageNumber="16" year="2015">
<taxonomicName id="13B84D0BFFB9FFD5FF7CEE5AFC59170F" ID-CoL="FZL3" authority="Enghoff, in Enghoff &amp; Frederiksen, 2015: 10" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityPageNumber="10" authorityYear="2015" box="[189,926,793,820]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="major">
<emphasis id="E6CCEA9AFFB9FFD5FF7CEE5AFE46170F" box="[189,385,794,820]" italics="true" pageId="15" pageNumber="16">Aquattuor major</emphasis>
Enghoff,
<emphasis id="E6CCEA9AFFB9FFD5FE36EE5AFDC9170F" box="[503,526,794,819]" italics="true" pageId="15" pageNumber="16">in</emphasis>
<bibRefCitation id="B0294B79FFB9FFD5FDD4EE59FC59170F" author="Enghoff H. &amp; Frederiksen S. B." box="[533,926,793,820]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015: 10</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFB9FFD5FF7CEE1AFC7111E2" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="D4073688FFB9FFD5FF7CEE1AFEF11748" blockId="15.[189,310,858,884]" box="[189,310,858,884]" pageId="15" pageNumber="16">
<heading id="8F4F81E4FFB9FFD5FF7CEE1AFEF11748" bold="true" box="[189,310,858,884]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="E6CCEA9AFFB9FFD5FF7CEE1AFEF11748" bold="true" box="[189,310,858,884]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CEEC9FACA106F" blockId="15.[189,1399,905,1108]" pageId="15" pageNumber="16">
Diameter
<quantity id="13409B6DFFB9FFD5FEF3EEC9FE191798" box="[306,478,905,932]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" metricValueMax="2.11" metricValueMin="1.97" pageId="15" pageNumber="16" unit="mm" value="2.04" valueMax="2.11" valueMin="1.97">1.97–2.11 mm</quantity>
, 53–54 podous rings (
<figureCitation id="4C832A0DFFB9FFD5FD20EEC9FCEE179F" box="[737,809,905,931]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff &amp; Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
). Differs from congeners, except
<taxonomicName id="13B84D0BFFB9FFD5FB75EECAFA83179F" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1204,1348,905,932]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFB9FFD5FB75EECAFA83179F" box="[1204,1348,905,932]" italics="true" pageId="15" pageNumber="16">A. longipala</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFB9FFD5FF7CEEEDFE9E17FA" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[189,345,940,967]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
<emphasis id="E6CCEA9AFFB9FFD5FF7CEEEDFE9E17FA" box="[189,345,940,967]" italics="true" pageId="15" pageNumber="16">A. mollilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFB9FFD5FEA1EEEDFE7B17FB" box="[352,444,941,967]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from
<taxonomicName id="13B84D0BFFB9FFD5FF3FEEB3FE481031" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[254,399,1011,1037]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFB9FFD5FF3FEEB3FE481031" box="[254,399,1011,1037]" italics="true" pageId="15" pageNumber="16">A. longipala</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFB9FFD5FE0BEEB3FDA01031" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[458,615,1011,1037]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
<emphasis id="E6CCEA9AFFB9FFD5FE0BEEB3FDA01031" box="[458,615,1011,1037]" italics="true" pageId="15" pageNumber="16">A. mollilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFB9FFD5FDAEEEB3FD0C1031" box="[623,715,1011,1037]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
by having a triangular tooth on the anterior margin of the gonopod telomere, from
<taxonomicName id="13B84D0BFFB9FFD5FE23E956FDB9100C" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[482,638,1046,1072]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
<emphasis id="E6CCEA9AFFB9FFD5FE23E956FDB9100C" box="[482,638,1046,1072]" italics="true" pageId="15" pageNumber="16">A. mollilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFB9FFD5FD47E957FD26100D" box="[646,737,1047,1073]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
further by having distinct lateral and mesal incisions on the gonopod coxa and by having the tarsal setae of the first pair of male legs of normal length.
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CE93AFDD010A9" blockId="15.[189,1399,1146,1255]" box="[189,535,1146,1173]" pageId="15" pageNumber="16">
<emphasis id="E6CCEA9AFFB9FFD5FF7CE93AFE4B10A8" bold="true" box="[189,396,1146,1172]" pageId="15" pageNumber="16">Material studied</emphasis>
(total: 1 ♂)
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CE9E9FBAB10DA" blockId="15.[189,1399,1146,1255]" pageId="15" pageNumber="16">
<materialsCitation id="64D03CD5FFB9FFD5FF7CE9E9FBAF10DA" ID-GBIF-Occurrence="2598688674" collectingDate="1982-08-18" collectingMethod="pitfall trap" collectionCode="FR" collectorName="M. Stoltze &amp; N. Scharff" country="Tanzania" county="Udzungwa Mts" elevation="1650" location="above Sanje" municipality="Mwanihana" pageId="15" pageNumber="16" specimenCode="NHMD 621644" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="paratype">
<collectingCountry id="ACAF7618FFB9FFD5FF7CE9E9FE9610FF" box="[189,337,1193,1219]" name="Tanzania" pageId="15" pageNumber="16">TANZANIA</collectingCountry>
•
<specimenCount id="C2BEFD01FFB9FFD5FEA8E9E9FE5F10FF" box="[361,408,1191,1219]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<typeStatus id="0B03882AFFB9FFD5FE65E9EAFDCA10F8" box="[420,525,1194,1220]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
;
<collectingRegion id="167CF86AFFB9FFD5FDD9E9EAFD2810FF" box="[536,751,1193,1220]" country="Tanzania" name="Morogoro" pageId="15" pageNumber="16">Morogoro Region</collectingRegion>
,
<collectingCounty id="3D664E04FFB9FFD5FD3BE9E9FC7710F8" box="[762,944,1193,1220]" pageId="15" pageNumber="16">Udzungwa Mts</collectingCounty>
,
<collectingMunicipality id="3463ACF2FFB9FFD5FC7DE9E9FB8110FF" box="[956,1094,1193,1219]" pageId="15" pageNumber="16">Mwanihana</collectingMunicipality>
FR,
<location id="D1676053FFB9FFD5FB41E9E9FAD510FF" LSID="urn:lsid:plazi:treatment:5C11879EFFB9FFD5FD9DEFF5FAD21272:D1676053FFB9FFD5FB41E9E9FAD510FF" box="[1152,1298,1193,1219]" country="Tanzania" county="Udzungwa Mts" municipality="Mwanihana" name="above Sanje" pageId="15" pageNumber="16" stateProvince="Morogoro Region">above Sanje</location>
;
<elevation id="5F95D1BBFFB9FFD5FADFE9E9FF3110DA" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="15" pageNumber="16" unit="m" value="1650.0">
<quantity id="13409B6DFFB9FFD5FADFE9E9FAB010F8" box="[1310,1399,1193,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="15" pageNumber="16" unit="m" value="1650.0">1650 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFB9FFD5FEC2E98CFE6210DA" box="[259,421,1228,1255]" pageId="15" pageNumber="16" value="1982-08-18">
<collectingDate id="B042E9A0FFB9FFD5FEC2E98CFE6210DA" box="[259,421,1228,1255]" pageId="15" pageNumber="16" value="1982-08-18">18 Aug. 1982</collectingDate>
</date>
;
<collectorName id="794D535EFFB9FFD5FE70E98DFDEC10DA" box="[433,555,1228,1255]" pageId="15" pageNumber="16">M. Stoltze</collectorName>
and
<collectorName id="794D535EFFB9FFD5FDA4E98DFD1810DA" box="[613,735,1228,1255]" pageId="15" pageNumber="16">N. Scharff</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFB9FFD5FCE1E98CFC5810DB" box="[800,927,1228,1255]" pageId="15" pageNumber="16">pitfall trap</collectingMethod>
;
<specimenCode id="841E9EF3FFB9FFD5FC6AE98DFBAF10DA" box="[939,1128,1228,1255]" collectionCode="NHMD" pageId="15" pageNumber="16">NHMD 621644</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CE84EFE541114" blockId="15.[189,403,1294,1320]" box="[189,403,1294,1320]" pageId="15" pageNumber="16">
<heading id="8F4F81E4FFB9FFD5FF7CE84EFE541114" bold="true" box="[189,403,1294,1320]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="E6CCEA9AFFB9FFD5FF7CE84EFE541114" bold="true" box="[189,403,1294,1320]" pageId="15" pageNumber="16">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CE87CFAE511A1" blockId="15.[189,1398,1340,1437]" pageId="15" pageNumber="16">
First pair of male legs (
<figureCitation id="4C832A0DFFB9FFD5FE07E87CFDCF116B" box="[454,520,1340,1367]" captionStart="Fig" captionStartId="7.[189,232,1887,1913]" captionTargetBox="[234,1343,270,1842]" captionTargetId="figure@7.[229,1354,262,1848]" captionTargetPageId="7" captionText="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754999" httpUri="https://zenodo.org/record/3754999/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
A–C) each with a single, finger-shaped prefemoral process (
<emphasis id="E6CCEA9AFFB9FFD5FB7CE87CFB25116A" box="[1213,1250,1340,1366]" italics="true" pageId="15" pageNumber="16">pfp</emphasis>
); in addition each prefemur has a smaller, irregular lobe (
<emphasis id="E6CCEA9AFFB9FFD5FD13E820FD301146" box="[722,759,1376,1402]" italics="true" pageId="15" pageNumber="16">dpl</emphasis>
) on the oral side of the distal margin; setae of femur much shorter than corresponding setae on normal walking legs, as in
<taxonomicName id="13B84D0BFFB9FFD5FC2BE8C3FBBC11A1" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1002,1147,1411,1437]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFB9FFD5FC2BE8C3FBBC11A1" box="[1002,1147,1411,1437]" italics="true" pageId="15" pageNumber="16">A. longipala</emphasis>
</taxonomicName>
(see
<figureCitation id="4C832A0DFFB9FFD5FB78E8C3FAD311A1" box="[1209,1300,1411,1437]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CE885FC7111E2" blockId="15.[189,950,1476,1503]" box="[189,950,1476,1503]" pageId="15" pageNumber="16">
<taxonomicName id="13B84D0BFFB9FFD5FF7CE885FE4611E2" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[189,385,1477,1503]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="major">
<emphasis id="E6CCEA9AFFB9FFD5FF7CE885FE4611E2" box="[189,385,1477,1503]" italics="true" pageId="15" pageNumber="16">Aquattuor major</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="13B84D0BFFB9FFD5FD97E885FD2011E2" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[598,743,1476,1503]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFB9FFD5FD97E885FD2011E2" box="[598,743,1476,1503]" italics="true" pageId="15" pageNumber="16">A. longipala</emphasis>
</taxonomicName>
(see Discussion).
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFB9FFD5FF7CEB45FAD21272" box="[189,1301,1541,1615]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="D4073688FFB9FFD5FF7CEB45FE911223" blockId="15.[189,342,1541,1567]" box="[189,342,1541,1567]" pageId="15" pageNumber="16">
<heading id="8F4F81E4FFB9FFD5FF7CEB45FE911223" bold="true" box="[189,342,1541,1567]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="E6CCEA9AFFB9FFD5FF7CEB45FE911223" bold="true" box="[189,342,1541,1567]" pageId="15" pageNumber="16">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFB9FFD5FF7CEB75FAD21272" blockId="15.[189,1301,1588,1615]" box="[189,1301,1588,1615]" pageId="15" pageNumber="16">
Known only from Mwanihana FR in the Udzungwa Mts. Altitudinal range:
<quantity id="13409B6DFFB9FFD5FBF4EB74FB101273" box="[1077,1239,1588,1615]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.85" metricValueMin="1.65" pageId="15" pageNumber="16" unit="m" value="1750.0" valueMax="1850.0" valueMin="1650.0">1650–1850 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>
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