A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae) Author Enghoff, Henrik text European Journal of Taxonomy 2020 2020-04-14 626 1 32 journal article 10.5852/ejt.2020.626 c906d9cd-cfcb-43db-a24c-b57b39d9184b 3754989 D91881D6-55D8-48FC-A383-069BC643A91E Aquattuor major Enghoff, 2015 Figs 1 , 5 A–C, 9 Aquattuor major Enghoff, in Enghoff & Frederiksen, 2015: 10 . Diagnosis Diameter 1.97–2.11 mm , 53–54 podous rings ( Fig. 9 ). Differs from congeners, except A. longipala and A. mollilobus sp. nov. , by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from A. longipala and A. mollilobus sp. nov. by having a triangular tooth on the anterior margin of the gonopod telomere, from A. mollilobus sp. nov. further by having distinct lateral and mesal incisions on the gonopod coxa and by having the tarsal setae of the first pair of male legs of normal length. Material studied (total: 1 ♂) TANZANIA • 1 ♂ , paratype ; Morogoro Region , Udzungwa Mts , Mwanihana FR, above Sanje ; 1650 m a.s.l. ; 18 Aug. 1982 ; M. Stoltze and N. Scharff leg.; pitfall trap ; NHMD 621644 . Descriptive notes First pair of male legs ( Fig. 5 A–C) each with a single, finger-shaped prefemoral process ( pfp ); in addition each prefemur has a smaller, irregular lobe ( dpl ) on the oral side of the distal margin; setae of femur much shorter than corresponding setae on normal walking legs, as in A. longipala (see Fig. 8B ). Aquattuor major is very similar to A. longipala (see Discussion). Distribution Known only from Mwanihana FR in the Udzungwa Mts. Altitudinal range: 1650–1850 m a.s.l.