400 lines
80 KiB
XML
400 lines
80 KiB
XML
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<mods:title id="0592F6ECB098441642D54923AA050595">Kasimovian (late Pennsylvanian) cornute rugose corals from Egypt: taxonomy, facies and palaeogeography of a cool-water fauna from northern Gondwana</mods:title>
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<mods:namePart id="6D1957D932A0CDFCAC95D22FF98D2948">El-Desouky, Heba</mods:namePart>
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<mods:namePart id="3094D74E91B8B0BB889B84A07101F2CD">Herbig, Hans-Georg</mods:namePart>
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<mods:namePart id="D0007FA61E9566E4112EC459D16B4CFA">Kora, Mahmoud</mods:namePart>
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<mods:title id="64BDC64D4E8D7D365DCAC00831D6B449">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="E517B2CE69EE9D6B9E3D3AAAB590F4A7">2023</mods:date>
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<paragraph id="C12ED8007F14FFF5FC91FD52FB91FD4D" blockId="13.[813,1139,685,710]" box="[813,1139,685,710]" pageId="13" pageNumber="14">
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<heading id="9A666F6C7F14FFF5FC91FD52FB91FD4D" bold="true" box="[813,1139,685,710]" fontSize="10" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="F3E504127F14FFF5FC91FD52FB91FD4D" bold="true" box="[813,1139,685,710]" pageId="13" pageNumber="14">
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<taxonomicName id="0691A3837F14FFF5FC91FD52FBD3FD4E" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[813,1073,685,709]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
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<emphasis id="F3E504127F14FFF5FC91FD52FBD3FD4E" bold="true" box="[813,1073,685,709]" italics="true" pageId="13" pageNumber="14">Lytvolasma paraaucta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E8D6B9697F14FFF5FB8BFD51FB91FD4D" box="[1079,1139,686,710]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="898B8B8B7F14FFF5FC91FD31FBC6FCAD" pageId="13" pageNumber="14" type="description">
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<paragraph id="C12ED8007F14FFF5FC91FD31FBBFFD6D" blockId="13.[813,1437,718,1894]" box="[813,1117,718,742]" pageId="13" pageNumber="14">
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(
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<figureCitation id="59AAC4857F14FFF5FC88FD31FC64FD6D" box="[820,902,718,742]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
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<figureCitation id="59AAC4857F14FFF5FC3AFD31FC56FD6D" box="[902,948,718,742]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1–A</figureCitation>
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<figureCitation id="59AAC4857F14FFF5FC08FD31FC23FD6D" box="[948,961,718,742]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
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, B
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<figureCitation id="59AAC4857F14FFF5FC62FD31FBEBFD6D" box="[990,1033,718,742]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1–B</figureCitation>
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||
<figureCitation id="59AAC4857F14FFF5FBB5FD31FBF4FD6D" box="[1033,1046,718,742]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
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,
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<figureCitation id="59AAC4857F14FFF5FB9DFD31FBB2FD6D" box="[1057,1104,718,742]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="13" pageNumber="14">12A</figureCitation>
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).
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</paragraph>
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||
<paragraph id="C12ED8007F14FFF5FC91FD11FBC6FCAD" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
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||
1991
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||
<taxonomicName id="0691A3837F14FFF5FCCCFD11FAF2FC8D" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[880,1296,750,774]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
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<emphasis id="F3E504127F14FFF5FCCCFD11FBD8FC8D" box="[880,1082,750,774]" italics="true" pageId="13" pageNumber="14">Lytvolasma aucta</emphasis>
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Fedorowski, 1987
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</taxonomicName>
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.- Kora and Mansour, figs. 2 c1, c2.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="898B8B8B7F14FFF5FC91FCD1FAA1FC0D" pageId="13" pageNumber="14" type="reference_group">
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||
<paragraph id="C12ED8007F14FFF5FC91FCD1FBB4FCED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
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||
1991
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||
<taxonomicName id="0691A3837F14FFF5FCDBFCD1FAB3FCCD" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" box="[871,1361,814,838]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="bellicostatum">
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||
<emphasis id="F3E504127F14FFF5FCDBFCD1FBE0FCCD" box="[871,1026,814,838]" italics="true" pageId="13" pageNumber="14">Bradyphyllum</emphasis>
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cf.
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<emphasis id="F3E504127F14FFF5FB94FCD1FB58FCCD" box="[1064,1210,814,838]" italics="true" pageId="13" pageNumber="14">bellicostatum</emphasis>
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||
<bibRefCitation id="A500A5F17F14FFF5FB7CFCD1FAB3FCCD" author="Grabau, A. W." box="[1216,1361,814,838]" pageId="13" pageNumber="14" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
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</taxonomicName>
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.- Kora and Mansour, figs. 2 b1, b2.
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</paragraph>
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||
<paragraph id="C12ED8007F14FFF5FC91FC91FAA1FC0D" blockId="13.[813,1437,718,1894]" box="[813,1347,878,902]" pageId="13" pageNumber="14">
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2001
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<taxonomicName id="0691A3837F14FFF5FCDBFC91FC01FC0D" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[871,995,878,902]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
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<emphasis id="F3E504127F14FFF5FCDBFC91FC01FC0D" box="[871,995,878,902]" italics="true" pageId="13" pageNumber="14">Lytvolasma</emphasis>
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</taxonomicName>
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—Fedorowski and Bamber, p 48.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="898B8B8B7F14FFF5FC91FC71FB4AFC4D" pageId="13" pageNumber="14" type="description">
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||
<paragraph id="C12ED8007F14FFF5FC91FC71FB4AFC4D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
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||
<emphasis id="F3E504127F14FFF5FC91FC71FBB6FC2D" bold="true" box="[813,1108,910,934]" pageId="13" pageNumber="14">Derivation of the name:</emphasis>
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From the Greek “
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<emphasis id="F3E504127F14FFF5FA8EFC70FA87FC2D" box="[1330,1381,911,934]" italics="true" pageId="13" pageNumber="14">para</emphasis>
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” for resemblance to the species
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<taxonomicName id="0691A3837F14FFF5FBE9FC51FB7FFC4D" authorityName="Fedorowski" authorityYear="1987" box="[1109,1181,942,966]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
|
||
“
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||
<emphasis id="F3E504127F14FFF5FBE3FC50FB7FFC4D" box="[1119,1181,943,966]" italics="true" pageId="13" pageNumber="14">aucta</emphasis>
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||
</taxonomicName>
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||
”.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="898B8B8B7F14FFF5FC91FC31FA7AFA2D" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="C12ED8007F14FFF5FC91FC31FB1FFB8D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="71F9D25D7F14FFF5FC91FC31FB1FFB8D" pageId="13" pageNumber="14" specimenCode="RAh 54" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="F3E504127F14FFF5FC91FC31FC7DFC6D" bold="true" box="[813,927,974,998]" pageId="13" pageNumber="14">
|
||
<typeStatus id="1E2A66A27F14FFF5FC91FC31FC79FC6D" box="[813,923,974,998]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="D79713897F14FFF5FC1AFC31FBF3FC6D" box="[934,1041,974,998]" pageId="13" pageNumber="14" type="generic">Specimen</specimenCount>
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<specimenCode id="9137707B7F14FFF5FBA5FC31FB8CFC6D" box="[1049,1134,974,998]" pageId="13" pageNumber="14">RAh 54</specimenCode>
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||
, illustrated in
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||
<figureCitation id="59AAC4857F14FFF5FAB7FC31FABFFC6D" box="[1291,1373,974,998]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
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||
<figureCitation id="59AAC4857F14FFF5FAE1FC31FA6EFC6D" box="[1373,1420,974,998]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1–A</figureCitation>
|
||
<figureCitation id="59AAC4857F14FFF5FA30FC31FA7BFC6D" box="[1420,1433,974,998]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
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. Four transverse thin-sections are available.
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</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FC91FBF1FAFCFB6D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FC91FBF1FC75FBAD" bold="true" box="[813,919,1038,1062]" pageId="13" pageNumber="14">Material:</emphasis>
|
||
In addition to the
|
||
<typeStatus id="1E2A66A27F14FFF5FBC4FBF1FB38FBAD" box="[1144,1242,1038,1062]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
, four moderately preserved corallites with compressed calices and occasionally preserved apical part (RAh 19, RAh 74 and RAh 81); collected from the basal shales of the lower member of the Aheimer Formation. Eight transverse thin-sections are available. They are collected from the basal shales of the lower member of the Aheimer Formation.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FC91FB11FC91FAED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FC91FB11FC2EFA8D" bold="true" box="[813,972,1262,1286]" pageId="13" pageNumber="14">Type locality:</emphasis>
|
||
<materialsCitation id="71F9D25D7F14FFF5FC68FB11FC8DFAED" country="Egypt" county="Gulf" latitude="29.4725" location="Northern Galala" longLatPrecision="20" longitude="32.460556" municipality="northern Eastern Desert" pageId="13" pageNumber="14" specimenCount="1" stateProvince="As Suways">
|
||
Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
|
||
<emphasis id="F3E504127F14FFF5FC61FAF2FC0AFAAD" box="[989,1000,1293,1318]" italics="true" pageId="13" pageNumber="14">′</emphasis>
|
||
21
|
||
<emphasis id="F3E504127F14FFF5FBB5FAF2FBFBFAAD" box="[1033,1049,1293,1318]" italics="true" pageId="13" pageNumber="14">″</emphasis>
|
||
N and Long. 32° 27
|
||
<emphasis id="F3E504127F14FFF5FB4CFAF2FB19FAAD" box="[1264,1275,1293,1318]" italics="true" pageId="13" pageNumber="14">′</emphasis>
|
||
38
|
||
<emphasis id="F3E504127F14FFF5FAA7FAF2FAC9FAAD" box="[1307,1323,1293,1318]" italics="true" pageId="13" pageNumber="14">″</emphasis>
|
||
E), northern Eastern Desert, western side of the Gulf of Suez, Egypt
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F14FFF5FC91FA91FA7AFA2D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FC91FA91FC32FA0D" bold="true" box="[813,976,1390,1414]" pageId="13" pageNumber="14">Type horizon:</emphasis>
|
||
Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="898B8B8B7F14FFF5FC91FA51FC9DF9ED" pageId="13" pageNumber="14" type="diagnosis">
|
||
<paragraph id="C12ED8007F14FFF5FC91FA51FC9DF9ED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FC91FA51FC44FA4D" bold="true" box="[813,934,1454,1478]" pageId="13" pageNumber="14">Diagnosis:</emphasis>
|
||
<taxonomicName id="0691A3837F14FFF5FC00FA51FBDAFA4D" box="[956,1080,1454,1478]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="F3E504127F14FFF5FC00FA51FBDAFA4D" box="[956,1080,1454,1478]" italics="true" pageId="13" pageNumber="14">Lytvolasma</emphasis>
|
||
</taxonomicName>
|
||
having maximum n:d ratio 26:11.5. Major septa frequently thickened in all quadrants to make a stereocolumn that opens in the cardinal quadrants, that changes to half aulos in the counter quadrants with maturity.
|
||
<taxonomicName id="0691A3837F14FFF5FBACF9D1FB8CF9CD" box="[1040,1134,1582,1606]" genus="Cardinal" pageId="13" pageNumber="8" rank="genus">Cardinal</taxonomicName>
|
||
septum is short within the fossula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="898B8B8B7F14FFF7FC91F991FDE5FB88" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="C12ED8007F14FFF6FC91F991FD66F956" blockId="13.[813,1437,718,1894]" lastBlockId="14.[151,775,1702,1918]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis id="F3E504127F14FFF5FC91F991FC5FF90D" bold="true" box="[813,957,1646,1670]" pageId="13" pageNumber="14">Description:</emphasis>
|
||
External characters: Corallites are small, up to
|
||
<quantity id="066975E57F14FFF5FCCBF971FC51F92E" box="[887,947,1678,1702]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="cm" value="3.0">3 cm</quantity>
|
||
long. The external wall is ornamented by septal furrows and fine concentric growth lines in the
|
||
<typeStatus id="1E2A66A27F14FFF5FC91F931FC6FF96D" box="[813,909,1742,1766]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
specimen (
|
||
<figureCitation id="59AAC4857F14FFF5FBB5F931FBBEF96D" box="[1033,1116,1742,1766]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F14FFF5FBE1F931FB85F96D" box="[1117,1127,1742,1766]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
|
||
), in other
|
||
<typeStatus id="1E2A66A27F14FFF5FB6BF931FAA2F96D" box="[1239,1344,1742,1766]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
|
||
the corallites are enclosed in mud and do not show any septal furrows (
|
||
<figureCitation id="59AAC4857F14FFF5FC2DF8F1FC01F8AD" box="[913,995,1806,1830]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8B</figureCitation>
|
||
<figureCitation id="59AAC4857F14FFF5FC5FF8F1FC0FF8AD" box="[995,1005,1806,1830]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
|
||
). Calices either eroded or crushed and compressed (
|
||
<figureCitation id="59AAC4857F14FFF5FC02F8D1FBF6F8CD" box="[958,1044,1838,1862]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F14FFF5FBA8F8D1FBFCF8CD" box="[1044,1054,1838,1862]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
|
||
). Two of the specimens with their apexes preserved. Two shallow constrictions are visible below the calicinal part in the
|
||
<typeStatus id="1E2A66A27F17FFF6FE53F959FDADF935" box="[495,591,1702,1726]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
specimen. Constrictions are also observed in the apical part.
|
||
</paragraph>
|
||
<caption id="95EE88887F17FFF6FF1BFA48FEBEF9DA" ID-DOI="http://doi.org/10.5281/zenodo.12003275" ID-Zenodo-Dep="12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15" startId="14.[167,198,1463,1483]" targetBox="[297,1298,274,1448]" targetPageId="14" targetType="figure">
|
||
<paragraph id="C12ED8007F17FFF6FF1BFA48FEBEF9DA" blockId="14.[167,1413,1463,1617]" pageId="14" pageNumber="15">
|
||
<emphasis id="F3E504127F17FFF6FF1BFA48FF10FA40" bold="true" box="[167,242,1463,1483]" pageId="14" pageNumber="15">Fig. 8 A</emphasis>
|
||
<taxonomicName id="0691A3837F17FFF6FF4BFA48FE4FFA40" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[247,429,1463,1483]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
|
||
<emphasis id="F3E504127F17FFF6FF4BFA48FE4FFA40" box="[247,429,1463,1483]" italics="true" pageId="14" pageNumber="15">Lytvolasma paraaucta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8D6B9697F17FFF6FE0EFA48FE3DFA40" box="[434,479,1463,1483]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
<emphasis id="F3E504127F17FFF6FE5EFA48FE19FA40" bold="true" box="[482,507,1463,1483]" pageId="14" pageNumber="15">A1</emphasis>
|
||
Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.
|
||
<emphasis id="F3E504127F17FFF6FE91FA2DFEA4FA6D" bold="true" box="[301,326,1490,1510]" pageId="14" pageNumber="15">A2</emphasis>
|
||
Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.
|
||
<emphasis id="F3E504127F17FFF6FB60FA2DFB17FA6D" bold="true" box="[1244,1269,1490,1510]" pageId="14" pageNumber="15">A3</emphasis>
|
||
Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends.
|
||
<emphasis id="F3E504127F17FFF6FBE0FA12FB97F98A" bold="true" box="[1116,1141,1517,1537]" pageId="14" pageNumber="15">A4</emphasis>
|
||
Laterally compressed section in the calice base,showing a deformed fossula.
|
||
<emphasis id="F3E504127F17FFF6FD86F9F8FDB0F990" bold="true" box="[570,594,1543,1563]" pageId="14" pageNumber="15">A5</emphasis>
|
||
Transverse thin-section in the compressed calice, with calcareous mud filling.
|
||
<emphasis id="F3E504127F17FFF6FB5BF9F8FB1CF990" bold="true" box="[1255,1278,1543,1563]" pageId="14" pageNumber="15">B1</emphasis>
|
||
Lateral external view of the paratype specimen (RAh 19).
|
||
<emphasis id="F3E504127F17FFF6FDBFF9DDFDDFF9BD" bold="true" box="[515,573,1570,1590]" pageId="14" pageNumber="15">B2–B5</emphasis>
|
||
Successive transverse thin-sections from the immature part near the apex (
|
||
<emphasis id="F3E504127F17FFF6FB07F9DDFB31F9BD" bold="true" box="[1211,1235,1570,1590]" pageId="14" pageNumber="15">B2</emphasis>
|
||
) to the last mature part in the calice (
|
||
<emphasis id="F3E504127F17FFF6FE83F9C2FEB5F9DA" bold="true" box="[319,343,1597,1617]" pageId="14" pageNumber="15">B5</emphasis>
|
||
)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C12ED8007F17FFF6FF11F919FEE3F895" blockId="14.[151,775,1702,1918]" pageId="14" pageNumber="15">
|
||
The following description refers mostly to the
|
||
<typeStatus id="1E2A66A27F17FFF6FD1BF919FCE5F975" box="[679,775,1766,1790]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
(RAh 54).
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F17FFF7FF11F8D9FD9BFE28" blockId="14.[151,775,1702,1918]" lastBlockId="15.[151,776,235,1475]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
Internal characters: The ontogenetically earliest preserved part has 18 septa in
|
||
<quantity id="066975E57F17FFF6FE56F8B9FDD2F8D6" box="[490,560,1862,1886]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="mm" value="4.0">4 mm</quantity>
|
||
of corallite diameter (
|
||
<figureCitation id="59AAC4857F17FFF6FF6EF899FEC7F8F5" box="[210,293,1894,1918]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F17FFF6FE9AF899FED6F8F5" box="[294,308,1894,1918]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="14" pageNumber="15">2</figureCitation>
|
||
). At this stage major septa show irregular zaphrentoid arrangement. They are thick, among them only the cardinal is thinner, whereas the counter is thicker with rhopaloid ending. The cardinal fossula is noticeable with long cardinal septum that fuses with the other major septa making a dense stereocolumn in the centre, that opens in the cardinal quadrants. Minor septa are absent in this stage (see also
|
||
<figureCitation id="59AAC4857F17FFF6FB99F899FB96F8F5" box="[1061,1140,1894,1918]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15">Fig. 8B</figureCitation>
|
||
<figureCitation id="59AAC4857F17FFF6FBC8F899FB60F8F5" box="[1140,1154,1894,1918]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="14" pageNumber="15">2</figureCitation>
|
||
). The subsequent section made at a diameter of
|
||
<quantity id="066975E57F16FFF7FE26FF13FE3FFE88" box="[410,477,236,259]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="7.0">7 mm</quantity>
|
||
shows 23 septa which are radially arranged (
|
||
<figureCitation id="59AAC4857F16FFF7FEDDFEF4FE56FEA8" box="[353,436,267,291]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FE09FEF4FE23FEA8" box="[437,449,267,291]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in C–E, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3</figureCitation>
|
||
). Their inner ends are rhopaloid and form a stereocolumn in the corallite centre. The cardinal fossula is slightly key-hole to subtriangular in shape and open adaxially (
|
||
<figureCitation id="59AAC4857F16FFF7FE09FE94FDE4FE08" box="[437,518,363,387]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FDBBFE94FDBEFE08" box="[519,604,363,387]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in C–E, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3, B3–B</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FDE7FE94FD8BFE08" box="[603,617,363,387]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
|
||
), with the cardinal septum thinner and slightly shortened.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F16FFF7FF11FE53FDE5FB88" blockId="15.[151,776,235,1475]" pageId="15" pageNumber="16">
|
||
The subsequent mature section is made beneath the calice floor (
|
||
<figureCitation id="59AAC4857F16FFF7FEA1FE34FE8CFE68" box="[285,366,459,483]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FED2FE34FE9EFE68" box="[366,380,459,483]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
|
||
). The corallite in this part is laterally compressed and almost half of the septa are broken and deformed. At this stage of maturity, the deformed cardinal fossula is bordered by a half aulos in counter quadrants. Major septa are almost radially arranged and are
|
||
<quantity id="066975E57F16FFF7FD51FDB4FF4FFD08" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="16" unit="in" value="24.0">24 in</quantity>
|
||
number.
|
||
<taxonomicName id="0691A3837F16FFF7FEA6FD94FE9AFD08" box="[282,376,619,643]" genus="Cardinal" pageId="15" pageNumber="8" rank="genus">Cardinal</taxonomicName>
|
||
septum is short within the fossula. Minor septa are rudimentary. They appear as small triangular spikes along the relatively thin epitheca. The last section within the compressed calice (
|
||
<figureCitation id="59AAC4857F16FFF7FDF9FD34FD7AFD68" box="[581,664,715,739]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FD24FD34FD47FD68" box="[664,677,715,739]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
|
||
) is filled with mud and shows short major septa and underdeveloped minor septa. At a diameter of 11.5 mm of the
|
||
<typeStatus id="1E2A66A27F16FFF7FD71FCF3FF24FCCF" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
|
||
specimen (RAh 19), more or less at the base of the calice (
|
||
<figureCitation id="59AAC4857F16FFF7FF5BFCB4FEDAFCE8" box="[231,312,843,867]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FE85FCB4FEA7FCE8" box="[313,325,843,867]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
|
||
) the septal ends are free leaving a free axial area with a short cardinal septum and a relatively longer counter septum than the adjacent septa. The counter septum, always a little longer and/or slightly thicker than other major septa, a character clearly evident in the
|
||
<typeStatus id="1E2A66A27F16FFF7FF2BFC13FF14FB8F" box="[151,246,1004,1028]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
|
||
specimen (
|
||
<figureCitation id="59AAC4857F16FFF7FECEFC14FE22FB88" box="[370,448,1003,1027]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FE7CFC14FE0CFB88" box="[448,494,1003,1027]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in C–E, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3–B</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FE52FC14FE19FB88" box="[494,507,1003,1027]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="898B8B8B7F16FFF7FF2BFBF4FAFCFCA8" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="C12ED8007F16FFF7FF2BFBF4FEEDFA48" blockId="15.[151,776,235,1475]" pageId="15" pageNumber="16">
|
||
<emphasis id="F3E504127F16FFF7FF2BFBF4FEFFFBA8" bold="true" box="[151,285,1035,1059]" pageId="15" pageNumber="16">Discussion:</emphasis>
|
||
<taxonomicName id="0691A3837F16FFF7FE99FBF4FDF9FBA8" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[293,539,1035,1059]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
|
||
<emphasis id="F3E504127F16FFF7FE99FBF4FDF9FBA8" box="[293,539,1035,1059]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8D6B9697F16FFF7FD9FFBF3FDBDFBA8" box="[547,607,1036,1059]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
was illustrated by
|
||
<bibRefCitation id="A500A5F17F16FFF7FF04FBD3FE2EFBC8" author="Kora, M. & Mansour, Y." box="[184,460,1067,1091]" pageId="15" pageNumber="16" pagination="597 - 614" refId="ref27107" refString="Kora, M., & Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991</bibRefCitation>
|
||
;
|
||
<figureCitation id="59AAC4857F16FFF7FE66FBD4FDCEFBC8" box="[474,556,1067,1091]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">Fig. 2 c</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FD90FBD4FDD5FBC8" box="[556,567,1067,1091]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="15" pageNumber="16">1</figureCitation>
|
||
, c
|
||
<figureCitation id="59AAC4857F16FFF7FDECFBD4FDBDFBC8" box="[592,607,1067,1091]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">2</figureCitation>
|
||
). Based on one transverse section in the mature part of the corallite and a longitudinal section, they assigned their specimen to
|
||
<taxonomicName id="0691A3837F16FFF7FD4EFB93FE43FB28" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F16FFF7FD4EFB93FF37FB28" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
|
||
Fedorowski, 1987
|
||
</taxonomicName>
|
||
. Due to the lack of description and additional illustrations of younger ontogenetic stages
|
||
<bibRefCitation id="A500A5F17F16FFF7FF2BFB34FE17FB68" author="Fedorowski, J. & Bamber, E. W." box="[151,501,1227,1251]" pageId="15" pageNumber="16" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., & Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber (2001)</bibRefCitation>
|
||
accepted this identification only at a generic level. We also include
|
||
<taxonomicName id="0691A3837F16FFF7FD32FB14FDFAFAA8" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="bellicostatum">
|
||
<emphasis id="F3E504127F16FFF7FD32FB14FF20FAA8" italics="true" pageId="15" pageNumber="16">Bradyphyllum</emphasis>
|
||
cf.
|
||
<emphasis id="F3E504127F16FFF7FF57FAF4FE9FFAA8" box="[235,381,1291,1315]" italics="true" pageId="15" pageNumber="16">bellicostatum</emphasis>
|
||
<bibRefCitation id="A500A5F17F16FFF7FE39FAF4FDFAFAA8" author="Grabau, A. W." box="[389,536,1291,1315]" pageId="15" pageNumber="16" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
briefly described and figured by
|
||
<bibRefCitation id="A500A5F17F16FFF7FEB3FAD3FDC4FAC8" author="Kora, M. & Mansour, Y." box="[271,550,1323,1347]" pageId="15" pageNumber="16" pagination="597 - 614" refId="ref27107" refString="Kora, M., & Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991</bibRefCitation>
|
||
, p. 599,
|
||
<figureCitation id="59AAC4857F16FFF7FD3BFAD4FD3BFAC8" box="[647,729,1323,1347]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">Fig. 2 c</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FD65FAD4FD12FAC8" box="[729,752,1323,1347]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="15" pageNumber="16">1c</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FD4CFAD4FD1DFAC8" box="[752,767,1323,1347]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">2</figureCitation>
|
||
) in our new species. Additional transverse thin-sections from the youngest stages clearly confirm the difference of the Egyptian specimens from
|
||
<taxonomicName id="0691A3837F16FFF7FE44FA73FDB3FA28" authorityName="Fedorowski" authorityYear="1987" box="[504,593,1420,1443]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F16FFF7FE44FA73FDB3FA28" box="[504,593,1420,1443]" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
|
||
</taxonomicName>
|
||
by the following characters:
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F16FFF7FF11FA12FEDFF9EE" blockId="15.[173,775,1517,1796]" pageId="15" pageNumber="16">
|
||
•
|
||
<taxonomicName id="0691A3837F16FFF7FF70FA12FE20F98F" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[204,450,1517,1541]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
|
||
<emphasis id="F3E504127F16FFF7FF70FA12FE20F98F" box="[204,450,1517,1541]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8D6B9697F16FFF7FE73FA12FDECF98F" box="[463,526,1517,1540]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
has higher n:d ratios than the
|
||
<typeStatus id="1E2A66A27F16FFF7FE8EF9F3FE72F9AF" box="[306,400,1548,1572]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="0691A3837F16FFF7FE05F9F2FDF1F9AF" authorityName="Fedorowski" authorityYear="1987" box="[441,531,1549,1572]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F16FFF7FE05F9F2FE25F9AF" box="[441,455,1549,1572]" italics="true" pageId="15" pageNumber="16">L</emphasis>
|
||
.
|
||
<emphasis id="F3E504127F16FFF7FE6AF9F2FDF1F9AF" box="[470,531,1549,1572]" italics="true" pageId="15" pageNumber="16">aucta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="59AAC4857F16FFF7FD98F9F3FD66F9AE" box="[548,644,1548,1573]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
|
||
). Nevertheless,
|
||
<taxonomicName id="0691A3837F16FFF7FEBEF9D2FEBEF9CF" authorityName="Fedorowski" authorityYear="1987" box="[258,348,1581,1604]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F16FFF7FEBEF9D2FEF2F9CF" box="[258,272,1581,1604]" italics="true" pageId="15" pageNumber="16">L</emphasis>
|
||
.
|
||
<emphasis id="F3E504127F16FFF7FEA3F9D2FEBEF9CF" box="[287,348,1581,1604]" italics="true" pageId="15" pageNumber="16">aucta</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="1E2A66A27F16FFF7FED9F9D2FE2EF9CE" box="[357,460,1581,1605]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
|
||
can reach a higher n:d ratio up to 22:9.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F16FFF7FF11F992FE7AF88F" blockId="15.[173,775,1517,1796]" pageId="15" pageNumber="16">
|
||
• The septal ends in the Egyptian species are thickened in contiguity to form a stereocolumn that is open towards the cardinal fossula, whereas in
|
||
<taxonomicName id="0691A3837F16FFF7FD3CF952FD3AF94F" authorityName="Fedorowski" authorityYear="1987" box="[640,728,1709,1732]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
|
||
<emphasis id="F3E504127F16FFF7FD3CF952FD3AF94F" box="[640,728,1709,1732]" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
|
||
</taxonomicName>
|
||
, the septal ends are free or form a half aulos only in the counter quadrants.
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F16FFF7FF2BF8D1FAAAFE28" blockId="15.[151,775,1838,1894]" lastBlockId="15.[813,1437,235,803]" pageId="15" pageNumber="16">
|
||
<taxonomicName id="0691A3837F16FFF7FF2BF8D1FE72F8CD" authority="El-Desouky & Herbig & Kora, 2023" authorityName="El-Desouky & Herbig & Kora" authorityYear="2023" box="[151,400,1838,1862]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
|
||
<emphasis id="F3E504127F16FFF7FF2BF8D1FE72F8CD" box="[151,400,1838,1862]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8D6B9697F16FFF7FE20F8D1FE38F8CE" box="[412,474,1838,1861]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="0691A3837F16FFF7FDCAF8D0FE01F8ED" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
|
||
<emphasis id="F3E504127F16FFF7FDCAF8D0FCE5F8CD" box="[630,775,1838,1862]" italics="true" pageId="15" pageNumber="16">L. canadense</emphasis>
|
||
<bibRefCitation id="A500A5F17F16FFF7FF2BF8B1FE01F8ED" author="Fedorowski, J. & Bamber, E. W." box="[151,483,1870,1894]" pageId="15" pageNumber="16" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., & Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Middle Permian of the Canadian Arctic Archipelago, in having an unremarkable counter septum, which is elongated in
|
||
<taxonomicName id="0691A3837F16FFF7FA3BFEF3FC7CFEC8" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
|
||
<emphasis id="F3E504127F16FFF7FA3BFEF3FC7CFEC8" italics="true" pageId="15" pageNumber="16">L. canadense</emphasis>
|
||
</taxonomicName>
|
||
in the calice and below its bottom. It also slightly differs by its n:d ratio in the ephebic stage near the calice floor (24:12? vs. 26:12.6;
|
||
<figureCitation id="59AAC4857F16FFF7FB06FE94FAFEFE08" box="[1210,1308,363,387]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
|
||
). The same case is for the Egyptian
|
||
<taxonomicName id="0691A3837F16FFF7FB92FE74FAA6FE28" box="[1070,1348,395,419]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
|
||
<emphasis id="F3E504127F16FFF7FB92FE74FB48FE28" box="[1070,1194,395,419]" italics="true" pageId="15" pageNumber="16">Lytvolasma</emphasis>
|
||
cf.
|
||
<emphasis id="F3E504127F16FFF7FB6CFE74FAA6FE28" box="[1232,1348,395,419]" italics="true" pageId="15" pageNumber="16">canadense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3E504127F16FFF7FAF8FE74FAAAFE28" box="[1348,1352,395,419]" italics="true" pageId="15" pageNumber="16">.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C12ED8007F16FFF7FC91FE53FAFCFCA8" blockId="15.[813,1437,235,803]" pageId="15" pageNumber="16">
|
||
In relation to
|
||
<taxonomicName id="0691A3837F16FFF7FC6EFE53FAC2FE48" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[978,1312,427,451]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">
|
||
<emphasis id="F3E504127F16FFF7FC6EFE53FB88FE48" box="[978,1130,428,451]" italics="true" pageId="15" pageNumber="16">L. asymetrica</emphasis>
|
||
<bibRefCitation id="A500A5F17F16FFF7FBCAFE54FAC2FE48" author="Soshkina, E. D." box="[1142,1312,427,451]" pageId="15" pageNumber="16" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from the Early Permian (Sakmarian) of Spitsbergen by Chwieduk (2013), the current new species has slightly similar n:d ratio in the mature stage below the calice (24:12? vs. 23–26:
|
||
<quantity id="066975E57F16FFF7FC4DFDD4FBA6FDC8" box="[1009,1092,555,579]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.556" metricValueMin="2.032" pageId="15" pageNumber="16" unit="in" value="11.0" valueMax="14.0" valueMin="8.0">8–14 in</quantity>
|
||
<emphasis id="F3E504127F16FFF7FBF7FDD3FB04FDC8" box="[1099,1254,556,579]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="0691A3837F16FFF7FBF7FDD3FB3CFDC8" authorityName="Soshkina" authorityYear="1925" box="[1099,1246,556,579]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">L. asymetrica</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<figureCitation id="59AAC4857F16FFF7FB51FDD4FAAEFDC8" box="[1261,1356,555,579]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
|
||
). However, in
|
||
<taxonomicName id="0691A3837F16FFF7FC39FDB3FBFBFDE8" authorityName="Soshkina" authorityYear="1925" box="[901,1049,588,611]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">
|
||
<emphasis id="F3E504127F16FFF7FC39FDB3FC71FDE8" box="[901,915,588,611]" italics="true" pageId="15" pageNumber="16">L</emphasis>
|
||
.
|
||
<emphasis id="F3E504127F16FFF7FC1EFDB3FBFBFDE8" box="[930,1049,588,611]" italics="true" pageId="15" pageNumber="16">asymetrica</emphasis>
|
||
</taxonomicName>
|
||
the inner ends of major septa join to form a quite irregular, incomplete aulos that opens on the alar septum sides throughout most of ontogeny; whereas in the current species a stereocolumn is developed through most of its ontogeny and in the mature stage, the thickened inner ends of the septa form an aulos that opens on the cardinal side (
|
||
<figureCitation id="59AAC4857F16FFF7FB3BFCF4FB3AFCA8" box="[1159,1240,779,803]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2–B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||
<figureCitation id="59AAC4857F16FFF7FB64FCF4FB06FCA8" box="[1240,1252,779,803]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in C–E, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3</figureCitation>
|
||
, A
|
||
<figureCitation id="59AAC4857F16FFF7FAB8FCF4FAF0FCA8" box="[1284,1298,779,803]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |