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<paragraph id="C12ED8007F00FFE1FC91FED4FB0EFEC8" blockId="25.[813,1437,235,1891]" box="[813,1260,299,323]" pageId="25" pageNumber="26">
Genus
<taxonomicName id="0691A3837F00FFE1FCC5FED4FB04FEC8" authority="Trautschold, 1879" authorityName="Trautschold" authorityYear="1879" box="[889,1254,299,323]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F00FFE1FCC5FED4FBFEFEC8" box="[889,1052,299,323]" italics="true" pageId="25" pageNumber="26">Bothrophyllum</emphasis>
<bibRefCitation id="A500A5F17F00FFE1FB9EFED4FB04FEC8" author="Trautschold, H." box="[1058,1254,299,323]" pageId="25" pageNumber="26" pagination="1 - 82" refId="ref29577" refString="Trautschold, H. (1879). Die KalkbrUche von Myatschkowa. Eine Monographie des Oberen Bergkalks. Socie´te´ Impe´ riale des Naturalistes de Moscou, Me´m. 14, 1 - 82." type="journal article" year="1879">Trautschold, 1879</bibRefCitation>
</taxonomicName>
.
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<emphasis id="F3E504127F00FFE1FC91FEB4FC2AFEE8" bold="true" box="[813,968,331,355]" pageId="25" pageNumber="26">
<typeStatus id="1E2A66A27F00FFE1FC91FEB4FC84FEE8" box="[813,870,331,355]" pageId="25" pageNumber="26">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F00FFE1FC6DFEB4FC83FE08" ID-CoL="8KQV2" authority="Fischer von Waldheim, 1837" authorityName="Fischer von Waldheim" authorityYear="1837" class="Anthozoa" family="Turbinoliidae" genus="Turbinolia" kingdom="Animalia" order="Scleractinia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="conica">
<emphasis id="F3E504127F00FFE1FC6DFEB4FB76FEE8" box="[977,1172,331,355]" italics="true" pageId="25" pageNumber="26">Turbinolia conica</emphasis>
<bibRefCitation id="A500A5F17F00FFE1FB21FEB4FC83FE08" author="Fischer von Waldheim, G." pageId="25" pageNumber="26" refId="ref25952" refString="Fischer von Waldheim, G. (1837). Oryctographie du Gouvernement de Moscou. Moscou: de l'imprimerie d'Auguste Semen. 202 pp." type="book" year="1837">Fischer von Waldheim, 1837</bibRefCitation>
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.
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<emphasis id="F3E504127F00FFE1FC91FE74FC45FE28" bold="true" box="[813,935,395,419]" pageId="25" pageNumber="26">Diagnosis:</emphasis>
see Kora et al. (2019).
</paragraph>
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<subSubSection id="898B8B8B7F00FFE1FC91FE54FB9BFDE8" pageId="25" pageNumber="26" type="distribution">
<paragraph id="C12ED8007F00FFE1FC91FE54FB9BFDE8" blockId="25.[813,1437,235,1891]" pageId="25" pageNumber="26">
<emphasis id="F3E504127F00FFE1FC91FE54FB35FE48" bold="true" box="[813,1239,427,451]" pageId="25" pageNumber="26">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F00FFE1FB5CFE54FA61FE48" authorityName="Trautschold" authorityYear="1879" box="[1248,1411,427,451]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F00FFE1FB5CFE54FA61FE48" box="[1248,1411,427,451]" italics="true" pageId="25" pageNumber="26">Bothrophyllum</emphasis>
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is known from the Viséan to the lower Permian, being typical of the Pennsylvanian (Chwieduk, 2013). It is recorded from the British Isles, Northern Spain, North Africa, Donets and Moscow basins, Ural Mountains,
<collectingCountry id="B98698907F00FFE1FAE8FDD4FA7AFDC8" box="[1364,1432,555,579]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
,
<collectingCountry id="B98698907F00FFE1FC91FDB4FC70FDE8" box="[813,914,587,611]" name="Australia" pageId="25" pageNumber="26">Australia</collectingCountry>
, and North America.
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<taxonomicName id="0691A3837F00FFE1FC91FD94FB07FD08" authority="Kossovaya, 2001" authorityName="Kossovaya" authorityYear="2001" box="[813,1253,619,644]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="okense">
<emphasis id="F3E504127F00FFE1FC91FD94FBCFFD08" bold="true" box="[813,1069,619,643]" italics="true" pageId="25" pageNumber="26">Bothrophyllum okense</emphasis>
Kossovaya, 2001
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</paragraph>
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<subSubSection id="898B8B8B7F00FFE1FC91FD74FC34FD28" box="[813,982,651,675]" pageId="25" pageNumber="26" type="description">
<paragraph id="C12ED8007F00FFE1FC91FD74FC34FD28" blockId="25.[813,1437,235,1891]" box="[813,982,651,675]" pageId="25" pageNumber="26">
(
<figureCitation id="59AAC4857F00FFE1FC88FD74FC6DFD28" box="[820,911,651,675]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FC33FD74FC58FD28" box="[911,954,651,675]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="25" pageNumber="26">1B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FC07FD74FC28FD28" box="[955,970,651,675]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
).
</paragraph>
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<subSubSection id="898B8B8B7F00FFE1FC91FD54FA6AFC88" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="C12ED8007F00FFE1FC91FD54FA6AFC88" blockId="25.[813,1437,235,1891]" pageId="25" pageNumber="26">
<emphasis id="F3E504127F00FFE1FC91FD54FC75FD48" bold="true" box="[813,919,683,707]" pageId="25" pageNumber="26">Material:</emphasis>
One preserved corallite (RAh 37), collected from the basal shales of lower member of the Aheimer Formation. Three transverse thin-sections are available.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F00FFE1FC91FCF4FAC5F88F" pageId="25" pageNumber="26" type="description">
<paragraph id="C12ED8007F00FFE1FC91FCF4FB1BFC28" blockId="25.[813,1437,235,1891]" pageId="25" pageNumber="26">
<emphasis id="F3E504127F00FFE1FC91FCF4FC5FFCA8" bold="true" box="[813,957,779,803]" pageId="25" pageNumber="26">Description:</emphasis>
External characters: The corallite is ca.
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in length. Calyx and apical end parts of are largely eroded. The external wall is partly damaged, eroded and enclosed in fossiliferous mud (
<figureCitation id="59AAC4857F00FFE1FBC2FC94FB3FFC08" box="[1150,1245,875,899]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FB61FC94FB05FC08" box="[1245,1255,875,899]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="25" pageNumber="26">1</figureCitation>
). The outer wall of the corallite in cross section is irregular.
</paragraph>
<paragraph id="C12ED8007F00FFE1FCFEFC54FAC5F88F" blockId="25.[813,1437,235,1891]" pageId="25" pageNumber="26">
Internal characters: The early preserved growth stage (
<figureCitation id="59AAC4857F00FFE1FC88FC34FC70FC68" box="[820,914,971,995]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FC2EFC34FC42FC68" box="[914,928,971,995]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="25" pageNumber="26">2</figureCitation>
), in a diameter of
<quantity id="066975E57F00FFE1FBCFFC34FB20FC68" box="[1139,1218,971,995]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="25" pageNumber="26" unit="mm" value="14.0">14 mm</quantity>
there are 30 major septa.
<taxonomicName id="0691A3837F00FFE1FCCBFC14FC03FB88" box="[887,993,1003,1027]" genus="All" pageId="25" pageNumber="26" rank="species" species="major">All major</taxonomicName>
septa are long, some reach more than 2/3 of the corallite radius, leaving a small axial area which is filled with a diagenetically altered axial structure formed from the axial ends of the major septa and axial tabellae.
<collectingRegion id="035516E27F00FFE1FC33FB94FC2EFB08" box="[911,972,1131,1155]" country="Spain" name="Ceuta" pageId="25" pageNumber="26">Septa</collectingRegion>
are strongly thickened in the tabularium, but thin in the very narrow dissepimentarium that consists of two rows of dissepiments. In the following higher section (
<figureCitation id="59AAC4857F00FFE1FC31FB34FC0EFB68" box="[909,1004,1227,1251]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FC50FB34FC1AFB68" box="[1004,1016,1227,1251]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="25" pageNumber="26">3</figureCitation>
) cardinal and counter septa are long and meet in the corallite center forming an axial septum that contributes with other septal axial ends in making a loose axial structure. At this stage, there are 31 septa for about
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. In the higher mature thin-section near the eroded calice (
<figureCitation id="59AAC4857F00FFE1FC71FA94FBCBFA08" box="[973,1065,1387,1411]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FB95FA94FBD5FA08" box="[1065,1079,1387,1411]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
), all septa are broken due to the compaction of the calicular part. There are about 34 septa in 13.5 mm compressed diameter. The counter septum is still long and reach the corallite centre, in contrast to the cardinal septum that seems to be shorten (
<figureCitation id="59AAC4857F00FFE1FB45FA14FAB6F988" box="[1273,1364,1515,1539]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FAE8FA14FA80F988" box="[1364,1378,1515,1539]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
). The axial structure that formed of the axial tabellae and inner ends of the longest major sept is still present. In a fairly good-preserved part of the corallite wall, a narrow dissepimentarium composed of 2 rows of simple concentric inter-septal dissepiments is present (
<figureCitation id="59AAC4857F00FFE1FB7CF974FAFFF928" box="[1216,1309,1675,1699]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FAA1F974FAC9F928" box="[1309,1323,1675,1699]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
).
<taxonomicName id="0691A3837F00FFE1FA82F974FA7EF928" box="[1342,1436,1675,1699]" genus="Cardinal" pageId="25" pageNumber="8" rank="genus">Cardinal</taxonomicName>
fossula is poorly developed. Minor septa are very short, confined to the dissepimentarium in the mature section (
<figureCitation id="59AAC4857F00FFE1FC88F914FC6CF888" box="[820,910,1771,1795]" captionStart="Fig" captionStartId="24.[167,198,1215,1235]" captionTargetBox="[287,1297,238,1199]" captionTargetId="figure-298@24.[283,1303,236,1202]" captionTargetPageId="24" captionText="Fig. 13 A Ufimia sp. A1 External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated. A2A3 Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:A1: 5 mm; A2, A3: 2 mm. B Bothrophyllum okense Kossovaya, 2001. B1 External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in B2 up to the more mature section below the calice in B4, the transverse sections show the axial septum and a very narrow dissepimentarium. Scale bars: 5mm" figureDoi="http://doi.org/10.5281/zenodo.12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
<figureCitation id="59AAC4857F00FFE1FC32F914FC7EF888" box="[910,924,1771,1795]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
), enter the tabularium as thick pegs.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F00FFE2FCFEF8F4FD14FE28" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
<paragraph id="C12ED8007F00FFE2FCFEF8F4FD14FE28" blockId="25.[813,1437,235,1891]" lastBlockId="26.[151,775,235,419]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="F3E504127F00FFE1FCFEF8F4FC25F8A8" bold="true" box="[834,967,1803,1827]" pageId="25" pageNumber="26">Discussion:</emphasis>
The specimen shows greatest similarity to
<taxonomicName id="0691A3837F00FFE1FCC9F8D4FACFF8C8" authority="Kossovaya, 2001" authorityName="Kossovaya" authorityYear="2001" box="[885,1325,1835,1860]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="okense">
<emphasis id="F3E504127F00FFE1FCC9F8D4FB8BF8C8" box="[885,1129,1835,1859]" italics="true" pageId="25" pageNumber="26">Bothrophyllum okense</emphasis>
Kossovaya, 2001
</taxonomicName>
from the Moscovian of the
<collectingRegion id="035516E27F00FFE1FC41F8B3FBBBF8E8" box="[1021,1113,1868,1891]" country="Russia" name="Moscow" pageId="25" pageNumber="26">Moscow</collectingRegion>
Basin in
<collectingCountry id="B98698907F00FFE1FB76F8B4FAF0F8E8" box="[1226,1298,1867,1891]" name="Russia" pageId="25" pageNumber="26">Russia</collectingCountry>
, despite the incomplete preservation and the diagenetic alteration of the skeleton of the current species. It was compared with the Egyptian slightly older bothrophyllids (Rod EL Hamal Formation) and with other Moscovian and Kasimovian
<taxonomicName id="0691A3837F03FFE2FF2BFE94FE8CFE08" box="[151,366,363,387]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F03FFE2FF2BFE94FED8FE08" box="[151,314,363,387]" italics="true" pageId="26" pageNumber="27">Bothrophyllum</emphasis>
spp.
</taxonomicName>
from the
<typeStatus id="1E2A66A27F03FFE2FE62FE93FDEFFE0F" box="[478,525,364,388]" pageId="26" pageNumber="27">type</typeStatus>
region of the
<collectingRegion id="035516E27F03FFE2FD17FE93FCE5FE08" box="[683,775,364,387]" country="Russia" name="Moscow" pageId="26" pageNumber="27">Moscow</collectingRegion>
Basin have been discussed by Kora et al. (2019;
<tableCitation id="8C13EDBB7F03FFE2FD25FE74FD04FE28" box="[665,742,395,419]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="26" pageNumber="27">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,279 @@
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<mods:roleTerm id="FB60DE5D7FAC45C4C0DE94B76C88DFDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E83F422F1EC6515913C652EDCA4C68F0">El-Desouky, Heba</mods:namePart>
</mods:name>
<mods:name id="4B507612760656445B62D7734FF09281" type="personal">
<mods:role id="B22EC54DC4A1D115CFF5A54E92B66BF9">
<mods:roleTerm id="3DE3DB34EA8E5928B061121053BEB0E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59C7A5898F237639BE7CB356071DAF3E">Herbig, Hans-Georg</mods:namePart>
</mods:name>
<mods:name id="76CD7BB8E351E137FA4E137752BCDFCC" type="personal">
<mods:role id="BFC9400AA9F32EDBF901781324A84DC2">
<mods:roleTerm id="EDA114E74CED69CB4A7E7E53A09BA60B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C858DB92A035159FEF031FB8CC4A9EE">Kora, Mahmoud</mods:namePart>
</mods:name>
<mods:typeOfResource id="C58EF81CEE92C92583CF0D991640CBF0">text</mods:typeOfResource>
<mods:relatedItem id="4A6E7762CDB78529BE060381BDACF11D" type="host">
<mods:titleInfo id="D3043192987C24FC7D68485EFCCAAC28">
<mods:title id="970DBA714407C05A238B1978BE32CB0C">Swiss Journal of Palaeontology</mods:title>
</mods:titleInfo>
<mods:part id="10383E4236015E349EA5220C1063BB0D">
<mods:date id="FB5BAB38C3A2EED3A729744B95D71A50">2023</mods:date>
<mods:detail id="81923DDCAC1428DA851C43E013D1C970" type="series">
<mods:title id="DC7E61D4FB1F1837806DB10C2A18F2B2">32</mods:title>
</mods:detail>
<mods:detail id="4AFBCDD2AD55454085D4FCC3ADCDC7D3" type="pubDate">
<mods:number id="B5EC1E9DA329198D77EA6F525A0D2A85">2023-11-27</mods:number>
</mods:detail>
<mods:detail id="F70FFA71ECA68F5F1A961475891E0F1A" type="volume">
<mods:number id="4DEAF3303BB7478F84F830F171412BF2">142</mods:number>
</mods:detail>
<mods:detail id="3137F3185739D9628C65FF0BF07873AC" type="issue">
<mods:number id="557CFD09BD3C8EB5A928D07F4AD3EB31">1</mods:number>
</mods:detail>
<mods:extent id="73A16E7A3F30C6686C1C2DC25F1BA80D" unit="page">
<mods:start id="14AFBEB446EBE5F0BA4BEBB6BDDE26BF">1</mods:start>
<mods:end id="6A1FEBF99FB91A7DAA2A7A25BF40B3E7">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="044A649412F0291C0C25DE9F2C1C5680">
<mods:url id="0AEE3F16E3AD34C05E7CE88B4F3F848B">http://dx.doi.org/10.1186/s13358-023-00296-0</mods:url>
</mods:location>
<mods:classification id="75E2023239B9B0212AC52C9A4BDAEED9">journal article</mods:classification>
<mods:identifier id="A65811EFD7BAA8C7917F3EE1667CDBE1" type="DOI">10.1186/s13358-023-00296-0</mods:identifier>
<mods:identifier id="AB5715330D3837CF168174398620DCBF" type="ISSN">1664-2384</mods:identifier>
<mods:identifier id="6E444BD8B1082C2B97502CD0932022D6" type="Zenodo-Dep">12003174</mods:identifier>
</mods:mods>
<treatment id="493869167F08FFEAFF2BFAD2FE64FCE8" LSID="urn:lsid:plazi:treatment:493869167F08FFEAFF2BFAD2FE64FCE8" httpUri="http://treatment.plazi.org/id/493869167F08FFEAFF2BFAD2FE64FCE8" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="898B8B8B7F08FFE9FF2BFAD2FDE1FACD" box="[151,515,1325,1350]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="C12ED8007F08FFE9FF2BFAD2FDE1FACD" blockId="17.[151,515,1325,1350]" box="[151,515,1325,1350]" pageId="17" pageNumber="18">
<heading id="9A666F6C7F08FFE9FF2BFAD2FDE1FACD" bold="true" box="[151,515,1325,1350]" fontSize="10" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="F3E504127F08FFE9FF2BFAD2FDE1FACD" bold="true" box="[151,515,1325,1350]" pageId="17" pageNumber="18">
<taxonomicName id="0691A3837F08FFE9FF2BFAD2FE22FACE" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[151,448,1325,1349]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="galalaensis" status="sp. nov.">
<emphasis id="F3E504127F08FFE9FF2BFAD2FE22FACE" bold="true" box="[151,448,1325,1349]" italics="true" pageId="17" pageNumber="18">Monophyllum galalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F08FFE9FE7AFAD1FDE1FACD" box="[454,515,1326,1350]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F08FFE9FF2BFAB1FDCFFA2D" pageId="17" pageNumber="18" type="description">
<paragraph id="C12ED8007F08FFE9FF2BFAB1FE2CFAED" blockId="17.[151,775,1353,1894]" box="[151,462,1353,1382]" pageId="17" pageNumber="18">
(
<figureCitation id="59AAC4857F08FFE9FF23FAB1FF12FAED" box="[159,240,1358,1382]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FF4CFAB1FEFFFAED" box="[240,285,1358,1382]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FEA2FAB1FED1FAD1" box="[286,307,1353,1382]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="17" pageNumber="18">
<superScript id="36E475487F08FFE9FEA2FAB1FED1FAD1" attach="none" box="[286,307,1353,1382]" fontSize="7" pageId="17" pageNumber="18">82</superScript>
</figureCitation>
, B
<figureCitation id="59AAC4857F08FFE9FEEDFAB1FE9EFAED" box="[337,380,1358,1382]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FEC0FAB1FE68FAED" box="[380,394,1358,1382]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="17" pageNumber="18">6</figureCitation>
,
<figureCitation id="59AAC4857F08FFE9FE29FAB1FE20FAED" box="[405,450,1358,1382]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="17" pageNumber="18">12B</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BFA91FDCFFA2D" blockId="17.[151,775,1353,1894]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FF2BFA91FE4FFA0D" bold="true" box="[151,429,1390,1414]" pageId="17" pageNumber="18">Derivation of the name:</emphasis>
<materialsCitation id="71F9D25D7F08FFE9FE09FA91FDCBFA2D" location="Northern Galala Plateau" pageId="17" pageNumber="18" specimenCount="1">
From the
<typeStatus id="1E2A66A27F08FFE9FD9DFA91FDB2FA0D" box="[545,592,1390,1414]" pageId="17" pageNumber="18">type</typeStatus>
locality (eastern cliffs of the
<location id="C44E8EDB7F08FFE9FEA9FA71FDC0FA2D" LSID="urn:lsid:plazi:treatment:493869167F08FFEAFF2BFAD2FE64FCE8:C44E8EDB7F08FFE9FEA9FA71FDC0FA2D" box="[277,546,1422,1446]" name="Northern Galala Plateau" pageId="17" pageNumber="18">Northern Galala Plateau</location>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F08FFE9FF2BFA51FA7AFEEF" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="C12ED8007F08FFE9FF2BFA51FE82F9AD" blockId="17.[151,775,1353,1894]" pageId="17" pageNumber="18">
<materialsCitation id="71F9D25D7F08FFE9FF2BFA51FE82F9AD" pageId="17" pageNumber="18" specimenCode="RAh 9" specimenCount="1" typeStatus="holotype">
<emphasis id="F3E504127F08FFE9FF2BFA51FEE8FA4D" bold="true" box="[151,266,1454,1478]" pageId="17" pageNumber="18">
<typeStatus id="1E2A66A27F08FFE9FF2BFA51FEE7FA4D" box="[151,261,1454,1478]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="D79713897F08FFE9FEABFA51FE60FA4D" box="[279,386,1454,1478]" pageId="17" pageNumber="18" type="generic">Specimen</specimenCount>
<specimenCode id="9137707B7F08FFE9FE33FA51FE3EFA4D" box="[399,476,1454,1478]" pageId="17" pageNumber="18">RAh 9</specimenCode>
; collected from the basal shales of the lower member of the Aheimer Formation and illustrated in
<figureCitation id="59AAC4857F08FFE9FEE2FA11FE52F98D" box="[350,432,1518,1542]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FE0DFA11FE3CF98D" box="[433,478,1518,1542]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FE62FA11FE11FA71" box="[478,499,1513,1542]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="17" pageNumber="18">
<superScript id="36E475487F08FFE9FE62FA11FE11FA71" attach="none" box="[478,499,1513,1542]" fontSize="7" pageId="17" pageNumber="18">82</superScript>
</figureCitation>
. Six transverse thin-sections are available.
</materialsCitation>
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BF9D1FEC2F8AD" blockId="17.[151,775,1353,1894]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FF2BF9D1FEE0F9CD" bold="true" box="[151,258,1582,1606]" pageId="17" pageNumber="18">Material:</emphasis>
<typeStatus id="1E2A66A27F08FFE9FEB0F9D1FE94F9CD" box="[268,374,1582,1606]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
are represented by five partly preserved corallites (RAh 13, RAh 41, RAh 52, RAh 80 and RAh 92); collected from the lower shales of the Aheimer Formation. The internal structures of these corallites are diagenetically altered. Eighteen transverse thin-sections are available, four of them belonging to RAh (13) are extremely altered by diagenesis and thus excluded from illustrations.
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BF8D1FC51FEAF" blockId="17.[151,775,1353,1894]" lastBlockId="17.[813,1437,236,1924]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FF2BF8D1FEDCF8CD" bold="true" box="[151,318,1838,1862]" pageId="17" pageNumber="18">Type locality:</emphasis>
<materialsCitation id="71F9D25D7F08FFE9FEF1F8D1FCE1F8ED" latitude="29.466946" location="Eastern" longLatPrecision="20" longitude="32.460556" municipality="Northern Galala" pageId="17" pageNumber="18" specimenCount="1">
Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
<emphasis id="F3E504127F08FFE9FEC8F8B3FE9DF8EE" box="[372,383,1868,1893]" italics="true" pageId="17" pageNumber="18"></emphasis>
01
<emphasis id="F3E504127F08FFE9FE18F8B3FE56F8EE" box="[420,436,1868,1893]" italics="true" pageId="17" pageNumber="18"></emphasis>
N and Long. 32° 27
<emphasis id="F3E504127F08FFE9FD1CF8B3FD49F8EE" box="[672,683,1868,1893]" italics="true" pageId="17" pageNumber="18"></emphasis>
38
<emphasis id="F3E504127F08FFE9FD73F8B3FD3DF8EE" box="[719,735,1868,1893]" italics="true" pageId="17" pageNumber="18"></emphasis>
E)
</materialsCitation>
, northern Eastern Desert, western side of the Gulf of
<collectingRegion id="035516E27F08FFE9FC91FEF3FC81FEAF" box="[813,867,268,292]" country="Egypt" name="As Suways" pageId="17" pageNumber="18">Suez</collectingRegion>
,
<collectingCountry id="B98698907F08FFE9FCD0FEF3FC4CFEAF" box="[876,942,268,292]" name="Egypt" pageId="17" pageNumber="18">Egypt</collectingCountry>
.
</paragraph>
<paragraph id="C12ED8007F08FFE9FC91FED3FA7AFEEF" blockId="17.[813,1437,236,1924]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FC91FED3FC32FECF" bold="true" box="[813,976,300,324]" pageId="17" pageNumber="18">Type horizon:</emphasis>
Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F08FFE9FC91FE93FA6DFDAF" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="C12ED8007F08FFE9FC91FE93FA6DFDAF" blockId="17.[813,1437,236,1924]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FC91FE93FC42FE0F" bold="true" box="[813,928,364,388]" pageId="17" pageNumber="18">Diagnosis</emphasis>
:
<taxonomicName id="0691A3837F08FFE9FC0AFE93FBACFE0F" authorityName="Fomichev, Shortened" authorityYear="1953" box="[950,1102,364,388]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FC0AFE93FBACFE0F" box="[950,1102,364,388]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
having maximum n:d ratio 24:11. Major septa meet in the corallite axis forming a stereocolumn; they retreat near the calice floor, forming a small free axial area occupied only by the incipient columella resulting from the elongated counter septum.
<taxonomicName id="0691A3837F08FFE9FAD7FE13FC81FDAF" genus="Cardinal" pageId="17" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum moderately shortened from early maturity.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F08FFEAFC91FDD3FD27FE48" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="C12ED8007F08FFE9FC91FDD3FB32FCCF" blockId="17.[813,1437,236,1924]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FC91FDD3FC5FFDCF" bold="true" box="[813,957,556,580]" pageId="17" pageNumber="18">Description:</emphasis>
External characters: Corallites are ceratoid, about 2.2
<quantity id="066975E57F08FFE9FC1AFDB3FC3EFDE8" box="[934,988,588,612]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="cm" value="3.0">3 cm</quantity>
long. Calyx and apex are not preserved in the
<typeStatus id="1E2A66A27F08FFE9FCC4FD93FC3AFD0F" box="[888,984,620,644]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(
<figureCitation id="59AAC4857F08FFE9FC54FD93FBDCFD0F" box="[1000,1086,620,644]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB82FD93FBA5FD0F" box="[1086,1095,620,644]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1</figureCitation>
) but preserved in most of the
<typeStatus id="1E2A66A27F08FFE9FC91FD73FC6EFD2F" box="[813,908,652,676]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
specimens (
<figureCitation id="59AAC4857F08FFE9FBA9FD73FB85FD2F" box="[1045,1127,652,676]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FBDBFD73FB93FD2F" box="[1127,1137,652,676]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1</figureCitation>
). When preserved, calyx is deep and laterally compressed. A remarkably long counter septum that forms an incipient columella with its end can be detected on its floor (
<figureCitation id="59AAC4857F08FFE9FBC9FD13FB2BFC8F" box="[1141,1225,748,772]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB75FD13FB35FC8F" box="[1225,1239,748,772]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">2</figureCitation>
). Delicate growth lines and distinct septal furrows are visible in the good preserved external wall (
<figureCitation id="59AAC4857F08FFE9FB8AFCD3FB6AFCCF" box="[1078,1160,812,836]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB34FCD3FB55FCCF" box="[1160,1207,812,836]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="17" pageNumber="18">1A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB0AFCD3FB26FCCF" box="[1206,1220,812,836]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">2</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F08FFE9FCFEFCB3FC3FFB4F" blockId="17.[813,1437,236,1924]" pageId="17" pageNumber="18">
Internal characters: The young stages of ontogeny are described from the
<typeStatus id="1E2A66A27F08FFE9FBABFC93FB94FC0F" box="[1047,1142,876,900]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
specimen (
<figureCitation id="59AAC4857F08FFE9FABDFC93FAB4FC0F" box="[1281,1366,876,900]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FAEAFC93FA67FC0F" box="[1366,1413,876,900]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">2B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FA38FC93FA72FC0F" box="[1412,1424,876,900]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="17" pageNumber="18">3</figureCitation>
), where the apical part is well-preserved. In the earliest ontogenetically preserved section (nepionic stage) of the
<typeStatus id="1E2A66A27F08FFE9FCE5FC33FC5AFC6F" box="[857,952,972,996]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
RAh (13) only 3 protosepta can be traced in about
<quantity id="066975E57F08FFE9FC2DFC13FC30FB88" box="[913,978,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
diameter (
<figureCitation id="59AAC4857F08FFE9FBF7FC13FB78FB8F" box="[1099,1178,1004,1028]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB26FC13FB4AFB8F" box="[1178,1192,1004,1028]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">2</figureCitation>
). The subsequent section records the early neanic stage (
<figureCitation id="59AAC4857F08FFE9FB12FBF3FB19FBAF" box="[1198,1275,1036,1060]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FB40FBF3FAEAFBAF" box="[1276,1288,1036,1060]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="17" pageNumber="18">3</figureCitation>
), where there are 14 major septa in 2.5 mm diameter, all meet near the corallite axis with their thick inner ends. The subsequent neanic growth stages of the
<typeStatus id="1E2A66A27F08FFE9FBDBFB93FB24FB0F" box="[1127,1222,1132,1156]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
have n:d of 1922: 56.7 mm. These sections are not suitable for illustration as stated before.
</paragraph>
<paragraph id="C12ED8007F08FFEAFCFEFB33FD27FE48" blockId="17.[813,1437,236,1924]" lastBlockId="18.[151,776,235,867]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
The two earliest preserved ontogenetic sections of the
<typeStatus id="1E2A66A27F08FFE9FCE7FB13FC59FA8F" box="[859,955,1260,1284]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
have 2122 septa in 66.5 mm diameter (
<figureCitation id="59AAC4857F08FFE9FC88FAF3FC64FAAF" box="[820,902,1292,1316]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FC3AFAF3FC54FAAF" box="[902,950,1292,1316]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="17" pageNumber="18">3A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FC09FAF3FC21FAAF" box="[949,963,1292,1316]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="17" pageNumber="18">4</figureCitation>
). In this stage major septa are thick, except for the cardinal septum. The counter septum is slightly thicker than the others and is longer.
<collectingRegion id="035516E27F08FFE9FB49FAB3FAD0FAEF" box="[1269,1330,1356,1380]" country="Spain" name="Ceuta" pageId="17" pageNumber="18">Septa</collectingRegion>
are pinnately to sub-radial arranged and meet in the centre to form a stereocolumn. Minor septa are mostly missing. In the successively youngest sections (
<figureCitation id="59AAC4857F08FFE9FB5EFA53FADAFA4F" box="[1250,1336,1452,1476]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FA84FA53FA83FA4F" box="[1336,1377,1452,1476]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[167,198,1874,1894]" captionStartId-1="9.[167,198,1520,1540]" captionStartId-2="12.[167,198,1342,1362]" captionTargetBox-0="[168,1421,1179,1861]" captionTargetBox-1="[287,1299,246,1507]" captionTargetBox-2="[285,1299,238,1327]" captionTargetId-0="figure-436@6.[166,1421,1179,1861]" captionTargetId-1="figure-67@9.[283,1303,236,1508]" captionTargetId-2="figure-189@12.[283,1303,236,1329]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionTargetPageId-2="12" captionText-0="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" captionText-1="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" captionText-2="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi-0="http://doi.org/10.5281/zenodo.12003232" figureDoi-1="http://doi.org/10.5281/zenodo.12003244" figureDoi-2="http://doi.org/10.5281/zenodo.12003254" httpUri-0="https://zenodo.org/record/12003232/files/figure.png" httpUri-1="https://zenodo.org/record/12003244/files/figure.png" httpUri-2="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">57</figureCitation>
, B
<figureCitation id="59AAC4857F08FFE9FA3EFA53FA72FA4F" box="[1410,1424,1452,1476]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="17" pageNumber="18">4</figureCitation>
), major septa count
<quantity id="066975E57F08FFE9FBBDFA33FB87FA6F" box="[1025,1125,1484,1508]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.096" metricValueMin="5.842" pageId="17" pageNumber="18" unit="in" value="23.5" valueMax="24.0" valueMin="23.0">2324 in</quantity>
810.5 mm diameter. They are joined with their thickened axial ends to form a dense axial column, with a narrower free axial area. The counter septum is the longest and exceeds all the majors in the axial stereocolumn with a rhopaloid thickened end.
<taxonomicName id="0691A3837F08FFE9FAD7F9B3FC81F90F" genus="Cardinal" pageId="17" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum thins towards the axis (
<figureCitation id="59AAC4857F08FFE9FB76F993FAFFF90F" box="[1226,1309,1644,1668]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FAA1F993FAC8F90F" box="[1309,1322,1644,1668]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="17" pageNumber="18">5</figureCitation>
), immediately becoming short, situated within a triangular, adaxial open cardinal fossula, which is located on the concave side of the corallite (
<figureCitation id="59AAC4857F08FFE9FBA6F933FB8DF96F" box="[1050,1135,1740,1764]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<figureCitation id="59AAC4857F08FFE9FBCCF933FB78F96F" box="[1136,1178,1740,1764]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[167,198,1520,1540]" captionStartId-1="12.[167,198,1342,1362]" captionTargetBox-0="[287,1299,246,1507]" captionTargetBox-1="[285,1299,238,1327]" captionTargetId-0="figure-67@9.[283,1303,236,1508]" captionTargetId-1="figure-189@12.[283,1303,236,1329]" captionTargetPageId-0="9" captionTargetPageId-1="12" captionText-0="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" captionText-1="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi-0="http://doi.org/10.5281/zenodo.12003244" figureDoi-1="http://doi.org/10.5281/zenodo.12003254" httpUri-0="https://zenodo.org/record/12003244/files/figure.png" httpUri-1="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">67</figureCitation>
). Alar fossulae are not visible. In the last mature sections, below the calice floor (
<figureCitation id="59AAC4857F08FFE9FC88F8F3FC64F8AF" box="[820,902,1804,1828]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9A</figureCitation>
<superScript id="36E475487F08FFE9FC3BF8F3FC4CF893" attach="none" box="[903,942,1799,1828]" fontSize="7" pageId="17" pageNumber="18">
<figureCitation id="59AAC4857F08FFE9FC3BF8F3FC79F893" box="[903,923,1799,1828]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="17" pageNumber="18">81</figureCitation>
,
<figureCitation id="59AAC4857F08FFE9FC19F8F8FC4CF893" box="[933,942,1799,1816]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">2</figureCitation>
</superScript>
, B
<figureCitation id="59AAC4857F08FFE9FC73F8F3FC3EF8AF" box="[975,988,1804,1828]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="17" pageNumber="18">5</figureCitation>
), and within the calice (
<figureCitation id="59AAC4857F08FFE9FB56F8F3FAD8F8AF" box="[1258,1338,1804,1828]" captionStart="Fig" captionStartId="16.[167,198,1543,1563]" captionTargetBox="[291,1292,238,1527]" captionTargetId="figure-16@16.[283,1303,236,1531]" captionTargetPageId="16" captionText="Fig. 9 A Monophyllum galalaensis n. sp. (Holotype specimen, Rah 9). A1 Lateral external view of the corallite, positions of thin-sections indicated in A1. A2 A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow). A3A5 Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn. A67 Two successive mature sections, showing a short cardinal septum. A81, 2 Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note, A82 is a redrawing of section A81. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines. B2 Nepionic stage.B3 Early neanic stage. B45 Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.B6 Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
<figureCitation id="59AAC4857F08FFE9FA86F8F3FAA5F8AF" box="[1338,1351,1804,1828]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="17" pageNumber="18">6</figureCitation>
), major septa are a little withdrawn from the axis and a narrow free axial area is developed that disappears in the lower, less mature sections. They are radially arranged. Minor septa are underdeveloped and form small triangular spikes at the inner wall. However, the long rhopaloid counter septum, incipient columella, and the short cardinal septum in the triangular fossula are still developed. The arrangement of the major septa changes from radial to pseudo-radial and then pinnate when going down towards the successively older parts of the corallite.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0BFFEAFF11FE34FE64FCE8" pageId="18" pageNumber="19" type="discussion">
<paragraph id="C12ED8007F0BFFEAFF11FE34FE64FCE8" blockId="18.[151,776,235,867]" pageId="18" pageNumber="19">
<emphasis id="F3E504127F0BFFEAFF11FE34FED0FE68" bold="true" box="[173,306,459,483]" pageId="18" pageNumber="19">Discussion:</emphasis>
The species discussed differs from the Upper Moscovian
<typeStatus id="1E2A66A27F0BFFEAFECDFE13FE42FD8F" box="[369,416,492,516]" pageId="18" pageNumber="19">type</typeStatus>
species
<taxonomicName id="0691A3837F0BFFEAFDB7FE14FEA6FDA8" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F0BFFEAFDB7FE14FCE5FD88" box="[523,775,491,515]" italics="true" pageId="18" pageNumber="19">Monophyllum sokolovi</emphasis>
<bibRefCitation id="A500A5F17F0BFFEAFF2BFDF4FEA6FDA8" author="Fomichev, V." box="[151,324,523,547]" pageId="18" pageNumber="19" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">
Fomichev,
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the strong elongation of the counter septum up to the calice floor, forming an incipient columella.
<taxonomicName id="0691A3837F0BFFEAFF58FDB4FD75FDE8" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[228,663,587,611]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F0BFFEAFF58FDB4FE3CFDE8" box="[228,478,587,611]" italics="true" pageId="18" pageNumber="19">Monophyllum parvum</emphasis>
<bibRefCitation id="A500A5F17F0BFFEAFE55FDB4FD75FDE8" author="Fomichev, V." box="[489,663,587,611]" pageId="18" pageNumber="19" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
differs in its smaller n:d ratio (
<figureCitation id="59AAC4857F0BFFEAFEC4FD94FE36FD08" box="[376,468,619,643]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="18" pageNumber="19">Fig. 12B</figureCitation>
). The current species differs from
<taxonomicName id="0691A3837F0BFFEAFF69FD73FDF6FD28" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[213,532,651,675]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F0BFFEAFF69FD73FF0FFD28" box="[213,237,652,675]" italics="true" pageId="18" pageNumber="19">M</emphasis>
.
<emphasis id="F3E504127F0BFFEAFF40FD73FEAEFD28" box="[252,332,652,675]" italics="true" pageId="18" pageNumber="19">cassum</emphasis>
Fedorowski, 1987
</taxonomicName>
by its larger n:d ratio (24:11.5 vs. 22:9.3). In addition,
<taxonomicName id="0691A3837F0BFFEAFDBCFD53FD95FD48" authorityName="Fedorowski" authorityYear="1987" box="[512,631,684,707]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F0BFFEAFDBCFD53FD95FD48" box="[512,631,684,707]" italics="true" pageId="18" pageNumber="19">M. cassum</emphasis>
</taxonomicName>
shows withdrawing of major septa from the corallite axis already in the late neanic/early ephebic stage, while this happens in the late ephebic stage near or at the calice floor in the Egyptian species. The formation of a stereocolumn is missing in
<taxonomicName id="0691A3837F0BFFEAFEB0FCB3FE63FCE8" authorityName="Fedorowski" authorityYear="1987" box="[268,385,844,867]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F0BFFEAFEB0FCB3FE63FCE8" box="[268,385,844,867]" italics="true" pageId="18" pageNumber="19">M. cassum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,186 @@
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<subSubSection id="898B8B8B7F0BFFEAFF2BFC52FDF4FC4F" box="[151,534,940,965]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="C12ED8007F0BFFEAFF2BFC52FDF4FC4F" blockId="18.[151,775,940,1925]" box="[151,534,940,965]" pageId="18" pageNumber="19">
<heading id="9A666F6C7F0BFFEAFF2BFC52FDF4FC4F" box="[151,534,940,965]" fontSize="36" level="2" pageId="18" pageNumber="19" reason="3">
Genus
<taxonomicName id="0691A3837F0BFFEAFF58FC53FDF2FC4E" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" box="[228,528,940,965]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFF58FC53FE91FC4F" bold="true" box="[228,371,940,964]" italics="true" pageId="18" pageNumber="19">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F0BFFEAFEC5FC52FDF2FC4E" author="Hudson, R. G. S." box="[377,528,941,965]" pageId="18" pageNumber="19" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0BFFEAFF2BFC32FE40FB8F" pageId="18" pageNumber="19" type="type_taxon">
<paragraph id="C12ED8007F0BFFEAFF2BFC32FE40FB8F" blockId="18.[151,775,940,1925]" pageId="18" pageNumber="19">
<materialsCitation id="71F9D25D7F0BFFEAFF2BFC32FE7CFB8F" location="Upper Visean of Ireland" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<emphasis id="F3E504127F0BFFEAFF2BFC32FED1FC6F" bold="true" box="[151,307,972,997]" pageId="18" pageNumber="19">
<typeStatus id="1E2A66A27F0BFFEAFF2BFC32FF32FC6E" box="[151,208,973,997]" pageId="18" pageNumber="19">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F0BFFEAFE81FC33FCE0FC6E" authority="Vaughan, 1908" authorityName="Vaughan" authorityYear="1908" box="[317,770,972,997]" class="Anthozoa" family="Streptelasmatidae" genus="Densiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="rushianum">
<emphasis id="F3E504127F0BFFEAFE81FC33FDB7FC6F" box="[317,597,972,996]" italics="true" pageId="18" pageNumber="19">Densiphyllum rushianum</emphasis>
Vaughan, 1908
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;
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.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0BFFEAFF2BFBF3FDA9FACE" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="C12ED8007F0BFFEAFF2BFBF3FDA9FACE" blockId="18.[151,775,940,1925]" pageId="18" pageNumber="19">
<emphasis id="F3E504127F0BFFEAFF2BFBF3FEF3FBAF" bold="true" box="[151,273,1036,1060]" pageId="18" pageNumber="19">Diagnosis:</emphasis>
Antiphylline coral with major septa meeting near corallite axis.
<taxonomicName id="0691A3837F0BFFEAFEC5FBD3FE35FBCF" box="[377,471,1068,1092]" genus="Cardinal" pageId="18" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum reaching corallite axis along cardinal fossula in all growth stages, shortened on calice wall but long on floor; counter septum commonly thicker and slightly longer than adjacent major septa; major septa evenly spaced, with extra thickening in inner third of their length, forming a dense axial structure; minor septa short; tabularium normal, sometimes weakly biform in some septal loculi. (
<bibRefCitation id="A500A5F17F0BFFEAFD82FAF2FCE2FAAE" author="Fedorowski, J." box="[574,768,1293,1317]" pageId="18" pageNumber="19" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
, slightly modified by
<bibRefCitation id="A500A5F17F0BFFEAFECFFAD2FDDFFACE" author="Fedorowski, J." box="[371,573,1325,1349]" pageId="18" pageNumber="19" pagination="1 - 37" refId="ref25654" refString="Fedorowski, J. (2009 a). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin, Ukraine. Part 1. Introductory considerations and the genus Rotiphyllum Hudson, 1942. Acta Geologica Polonica, 59 (1), 1 - 37." type="journal article" year="2009">Fedorowski, 2009a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0BFFEAFF2BFAB2FC8DFD08" pageId="18" pageNumber="19" type="discussion">
<paragraph id="C12ED8007F0BFFEAFF2BFAB2FC8DFD08" blockId="18.[151,775,940,1925]" lastBlockId="18.[813,1437,235,643]" pageId="18" pageNumber="19">
<emphasis id="F3E504127F0BFFEAFF2BFAB2FF1FFAEE" bold="true" box="[151,253,1357,1381]" pageId="18" pageNumber="19">Remarks</emphasis>
: Generic relations of
<taxonomicName id="0691A3837F0BFFEAFE48FAB2FD9BFAEE" authorityName="Hudson" authorityYear="1942" box="[500,633,1357,1381]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFE48FAB2FD9BFAEE" box="[500,633,1357,1381]" italics="true" pageId="18" pageNumber="19">Rotiphyllum</emphasis>
</taxonomicName>
were widely discussed by authors, among them
<bibRefCitation id="A500A5F17F0BFFEAFD9BFA92FD56FA0F" author="Weyer, D." box="[551,692,1388,1413]" pageId="18" pageNumber="19" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975</bibRefCitation>
,
<bibRefCitation id="A500A5F17F0BFFEAFD78FA92FD1AFA0E" author="Weyer, D." box="[708,760,1389,1413]" pageId="18" pageNumber="19" pagination="459 - 468" refId="ref30399" refString="Weyer, D. (1977). Review of the rugose coral faunas of the Lower Namurian Ostrava Formation (Upper Silesian Coal Basin). In V. M. Holub, &amp; R. H. Wagner (Eds.), Symposium on Carboniferous Stratigraphy (IUGS Subcommission on Carboniferous Stratigraphy (SCCS), Field and general meeting, Czechoslovakia, September 17 - 27, 1973), 459 - 468 (Geological Survey, Prague)." type="book chapter" year="1977">1977</bibRefCitation>
),
<bibRefCitation id="A500A5F17F0BFFEAFF2BFA72FE96FA2E" author="Fedorowski, J." box="[151,372,1421,1445]" pageId="18" pageNumber="19" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
, and
<bibRefCitation id="A500A5F17F0BFFEAFE00FA72FDADFA2E" author="Wang, X. D." box="[444,591,1421,1445]" pageId="18" pageNumber="19" pagination="118 - 129" refId="ref29776" refString="Wang, X. D. (1994). Revision of Antiphyllinae Ilina, 1970 with a method of cladistics analysis. Acta Palaeontologica, Sinica, 33 (1), 118 - 129." type="journal article" year="1994">Wang (1994)</bibRefCitation>
who supported Fedorowskis concept of the genus. Fedorowksi (1987a) especially discussed the relation to
<taxonomicName id="0691A3837F0BFFEAFD8BFA33FD30FA6F" authorityName="Grabau" authorityYear="1928" box="[567,722,1484,1508]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFD8BFA33FD30FA6F" box="[567,722,1484,1508]" italics="true" pageId="18" pageNumber="19">Bradyphyllum</emphasis>
</taxonomicName>
and stressed the difference of
<taxonomicName id="0691A3837F0BFFEAFE04FA13FDDFF98F" authorityName="Hudson" authorityYear="1942" box="[440,573,1516,1540]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFE04FA13FDDFF98F" box="[440,573,1516,1540]" italics="true" pageId="18" pageNumber="19">Rotiphyllum</emphasis>
</taxonomicName>
by its long cardinal septum along the fossula floor, though, later,
<bibRefCitation id="A500A5F17F0BFFEAFD03F9F2FEC5F9CE" author="Fedorowski, J." pageId="18" pageNumber="19" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski (2004)</bibRefCitation>
stated the very close relationship of both genera. The synonymy of
<taxonomicName id="0691A3837F0BFFEAFE65F9B2FD9AF9EE" authorityName="Fomichev" authorityYear="1953" box="[473,632,1613,1637]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFE65F9B2FD9AF9EE" box="[473,632,1613,1637]" italics="true" pageId="18" pageNumber="19">Actinophrentis</emphasis>
</taxonomicName>
, unambiguously synonymised by
<bibRefCitation id="A500A5F17F0BFFEAFE2BF992FDCCF90E" author="Weyer, D." box="[407,558,1644,1669]" pageId="18" pageNumber="19" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
and also supposed by
<bibRefCitation id="A500A5F17F0BFFEAFF01F972FED5F92E" author="Hill, D." box="[189,311,1677,1701]" pageId="18" pageNumber="19" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
was rejected by
<bibRefCitation id="A500A5F17F0BFFEAFDBCF972FD31F92E" author="Fedorowski, J." box="[512,723,1677,1701]" pageId="18" pageNumber="19" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a</bibRefCitation>
; see above under
<taxonomicName id="0691A3837F0BFFEAFE95F952FE2AF94E" authorityName="Fomichev" authorityYear="1953" box="[297,456,1709,1733]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFE95F952FE2AF94E" box="[297,456,1709,1733]" italics="true" pageId="18" pageNumber="19">Actinophrentis</emphasis>
</taxonomicName>
), who also rejected the conclusion of de Groot (1963),
<bibRefCitation id="A500A5F17F0BFFEAFE7FF932FDB5F96E" author="Weyer, D." box="[451,599,1740,1765]" pageId="18" pageNumber="19" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
, and
<bibRefCitation id="A500A5F17F0BFFEAFD2EF932FCE5F96E" author="Hill, D." box="[658,775,1741,1765]" pageId="18" pageNumber="19" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
that
<taxonomicName id="0691A3837F0BFFEAFF6CF912FDC4F88F" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" box="[208,550,1772,1797]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFF6CF912FE8AF88E" box="[208,360,1773,1797]" italics="true" pageId="18" pageNumber="19">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F0BFFEAFEC9F912FDC4F88F" author="Fomichev, V." box="[373,550,1772,1797]" pageId="18" pageNumber="19" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
would be a junior synonym of
<taxonomicName id="0691A3837F0BFFEAFEA3F8F2FDA6F8AE" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" box="[287,580,1805,1829]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFEA3F8F2FE46F8AE" box="[287,420,1805,1829]" italics="true" pageId="18" pageNumber="19">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F0BFFEAFE10F8F2FDA6F8AE" author="Hudson, R. G. S." box="[428,580,1805,1829]" pageId="18" pageNumber="19" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
(see above under
<taxonomicName id="0691A3837F0BFFEAFF2BF8D2FECDF8CE" authorityName="Fomichev, Shortened" authorityYear="1953" box="[151,303,1837,1861]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFF2BF8D2FECDF8CE" box="[151,303,1837,1861]" italics="true" pageId="18" pageNumber="19">Monophyllum</emphasis>
</taxonomicName>
), (
<tableCitation id="8C13EDBB7F0BFFEAFEEBF8D2FE56F8CE" box="[343,436,1837,1861]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="18" pageNumber="19">Table 1</tableCitation>
). Finally,
<bibRefCitation id="A500A5F17F0BFFEAFD88F8D2FCE2F8CE" author="Fedorowski, J." box="[564,768,1837,1861]" pageId="18" pageNumber="19" pagination="275 - 325" refId="ref25587" refString="Fedorowski, J. (1989). Redescription of the original collection of Zaphrentis calyculus Miller, 1891, Rugosa. Acta Palaeontologica Polonica, 34 (4), 275 - 325." type="journal article" year="1989">Fedorowski (1989</bibRefCitation>
; updated in
<bibRefCitation id="A500A5F17F0BFFEAFEABF8B2FE06F8EE" author="Fedorowski, J." box="[279,484,1869,1893]" pageId="18" pageNumber="19" pagination="1 - 37" refId="ref25654" refString="Fedorowski, J. (2009 a). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin, Ukraine. Part 1. Introductory considerations and the genus Rotiphyllum Hudson, 1942. Acta Geologica Polonica, 59 (1), 1 - 37." type="journal article" year="2009">Fedorowski, 2009a</bibRefCitation>
) gave an extensive review on the synonymy, the included species and the relations of the genus. According to the extended number of species attributed by him, the very long stratigraphical range from the Tournaisian throughout the Lower Permian— and even the middle Permian in
<collectingCountry id="B98698907F0BFFEAFB26FEB3FB34FEE8" box="[1178,1238,332,355]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
(fide
<bibRefCitation id="A500A5F17F0BFFEAFAA4FEB3FC83FE08" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." pageId="18" pageNumber="19" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
)—and geographical distribution from the British Isles and the westernmost Palaeotethys (northern
<collectingCountry id="B98698907F0BFFEAFAEBFE74FA77FE28" box="[1367,1429,395,419]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
) to northeastern
<collectingCountry id="B98698907F0BFFEAFC5FFE54FBC7FE48" box="[995,1061,427,451]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
and the peri-Gondwana terranes from
<collectingRegion id="035516E27F0BFFEAFCD2FE34FC4AFE68" box="[878,936,459,483]" country="China" name="Xizang" pageId="18" pageNumber="19">Tibet</collectingRegion>
to
<collectingRegion id="035516E27F0BFFEAFC6AFE33FBC8FE68" box="[982,1066,460,483]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
(judged from Fedorowskis synonymy list and complemented by
<bibRefCitation id="A500A5F17F0BFFEAFB02FE13FA6FFD88" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." box="[1214,1421,491,516]" pageId="18" pageNumber="19" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
),
<taxonomicName id="0691A3837F0BFFEAFC91FDF4FC50FDA8" authorityName="Hudson" authorityYear="1942" box="[813,946,523,547]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0BFFEAFC91FDF4FC50FDA8" box="[813,946,523,547]" italics="true" pageId="18" pageNumber="19">Rotiphyllum</emphasis>
</taxonomicName>
might be a waste basket of morphotypes belonging to different taxa. It appears to be absent from the CordilleranArcticUralian realm (
<bibRefCitation id="A500A5F17F0BFFEAFABCFDB4FC83FD08" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." pageId="18" pageNumber="19" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="C12ED8007F0DFFECFF2BFAB2FDFEFAEF" blockId="20.[151,775,1293,1925]" box="[151,540,1356,1381]" pageId="20" pageNumber="21">
<heading id="9A666F6C7F0DFFECFF2BFAB2FDFEFAEF" box="[151,540,1356,1381]" fontSize="36" level="2" pageId="20" pageNumber="21" reason="3">
Genus
<taxonomicName id="0691A3837F0DFFECFF58FAB3FDF4FAEE" authority="Hudson, 1941" authorityName="Hudson" authorityYear="1941" box="[228,534,1356,1381]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFF58FAB3FE9BFAEF" bold="true" box="[228,377,1356,1380]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
<bibRefCitation id="A500A5F17F0DFFECFEC3FAB2FDF4FAEE" author="Hudson, R. G. S." box="[383,534,1357,1381]" pageId="20" pageNumber="21" pagination="290 - 311" refId="ref26607" refString="Hudson, R. G. S. (1941). On the Carboniferous corals: Zaphrentis carruthersi sp. nov. from the Mirk Fell Beds and its relation to the Z. delanouei speciesgroup. Yorkshire Geological Society, 24 (4), 290 - 311." type="journal article" year="1941">Hudson, 1941</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFF2BFA93FD3CFA0F" box="[151,734,1388,1413]" pageId="20" pageNumber="21" type="type_taxon">
<paragraph id="C12ED8007F0DFFECFF2BFA93FD3CFA0F" blockId="20.[151,775,1293,1925]" box="[151,734,1388,1413]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFF2BFA93FECFFA0F" bold="true" box="[151,301,1388,1412]" pageId="20" pageNumber="21">Type-species</emphasis>
:
<taxonomicName id="0691A3837F0DFFECFE86FA93FD3AFA0E" authority="Carruthers, 1910" authorityName="Carruthers" authorityYear="1910" box="[314,728,1388,1413]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0DFFECFE86FA93FDFBFA0F" box="[314,537,1388,1412]" italics="true" pageId="20" pageNumber="21">Zaphrentis parallela</emphasis>
Carruthers, 1910
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFF2BFA73FE90F96F" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="C12ED8007F0DFFECFF2BFA73FE90F96F" blockId="20.[151,775,1293,1925]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFF2BFA73FEEFFA2F" bold="true" box="[151,269,1420,1444]" pageId="20" pageNumber="21">Diagnosis</emphasis>
: Small, conical to slightly curved with well-marked longitudinal ribbing.
<taxonomicName id="0691A3837F0DFFECFE47FA53FDBEFA4F" box="[507,604,1452,1476]" genus="Cardinal" pageId="20" pageNumber="8" rank="genus">Cardinal</taxonomicName>
fossula is on the concave side. In early growth stages it is closed and expands adaxially, and major septa are pinnately arranged. In late growth stages cardinal septum is shortened and septa are withdrawn from the fossula, first in cardinal quadrants, then in counter quadrants and thus a radial arrangement of the septa supersedes pinnate arrangement. Minor septa are very short or immersed in the wall. Tabulae are incomplete (after
<bibRefCitation id="A500A5F17F0DFFECFF2BF933FEE6F96E" author="Hill, D." box="[151,260,1740,1765]" pageId="20" pageNumber="21" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill, 1981</bibRefCitation>
; p. F316).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFF2BF912FB7AFD48" pageId="20" pageNumber="21" type="discussion">
<paragraph id="C12ED8007F0DFFECFF2BF912FB7AFD48" blockId="20.[151,775,1293,1925]" lastBlockId="20.[813,1437,235,1507]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFF2BF912FF1FF88E" bold="true" box="[151,253,1773,1797]" pageId="20" pageNumber="21">Remarks</emphasis>
:
<bibRefCitation id="A500A5F17F0DFFECFEB1F913FE67F88E" author="Hill, D." box="[269,389,1772,1797]" pageId="20" pageNumber="21" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
included
<taxonomicName id="0691A3837F0DFFECFE49F913FD62F88F" authorityName="Hudson" authorityYear="1941" box="[501,640,1772,1796]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFE49F913FD62F88F" box="[501,640,1772,1796]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
into
<taxonomicName id="0691A3837F0DFFECFD02F913FF1EF8AE" authorityName="Grabau" authorityYear="1928" class="Anthozoa" family="Hapsiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="family">Hapsiphyllidae</taxonomicName>
, but
<bibRefCitation id="A500A5F17F0DFFECFE8EF8F2FDE8F8AE" author="Fedorowski, J." box="[306,522,1805,1829]" pageId="20" pageNumber="21" pagination="109 - 161" refId="ref25826" refString="Fedorowski, J. (2012 b). The new Upper Serpukhovian genus Zaphrufimia and homeomorphism in some rugose corals. Palaeontogr. Abteilung, 296, 109 - 161." type="journal article" year="2012">Fedorowski (2012b)</bibRefCitation>
insisted on the validity of the family
<taxonomicName id="0691A3837F0DFFECFE94F8D2FD76F8CF" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[296,660,1836,1861]" family="Stereophrentidae" pageId="20" pageNumber="21" rank="family">
Stereophrentidae
<bibRefCitation id="A500A5F17F0DFFECFE57F8D2FD76F8CF" author="Fomichev, V." box="[491,660,1836,1861]" pageId="20" pageNumber="21" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
according to
<collectionCode id="A78040C57F0DFFECFF0AF8B2FF15F8EF" box="[182,247,1869,1892]" pageId="20" pageNumber="21">IRZN</collectionCode>
, although the genus
<taxonomicName id="0691A3837F0DFFECFE5DF8B2FD9EF8EE" authorityName="Fomitchev" authorityYear="1955" box="[481,636,1869,1893]" class="Anthozoa" genus="Stereophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFE5DF8B2FD9EF8EE" box="[481,636,1869,1893]" italics="true" pageId="20" pageNumber="21">Stereophrentis</emphasis>
</taxonomicName>
is not valid, as it is an objective younger synonym of
<taxonomicName id="0691A3837F0DFFECFDCAF892FCE3F80E" authorityName="Hudson" authorityYear="1941" box="[630,769,1901,1925]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFDCAF892FCE3F80E" box="[630,769,1901,1925]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
. Herein, we did not follow this proposal.
<taxonomicName id="0691A3837F0DFFECFAADFF14FA7EFE88" authorityName="Hudson" authorityYear="1941" box="[1297,1436,235,259]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFAADFF14FA7EFE88" box="[1297,1436,235,259]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
shares its simple morphology with many zaphrentoidlike taxa. The position of the cardinal fossula on the concave side of the corallite distinguishes it from the other zaphrentoid corals, where it is located on the convex side. It differs from
<emphasis id="F3E504127F0DFFECFC6BFE74FB47FE28" box="[983,1189,395,419]" italics="true" pageId="20" pageNumber="21">Ampleoizaphrentis</emphasis>
<bibRefCitation id="A500A5F17F0DFFECFB13FE74FAB1FE28" author="Vaughan, A." box="[1199,1363,395,419]" pageId="20" pageNumber="21" pagination="295 - 322" refId="ref29612" refString="Vaughan, A. (1906). Faunal lists and account of the faunal succession and correlation. In C. A. Matley, &amp; A. Vaughan (Eds.), The Carboniferous rocks at Rush County (Dublin). Quarterly Journal of the Geological Society of London, 62, 295 - 322." type="journal article" year="1906">Vaughan, 1906</bibRefCitation>
by its smaller dimensions, missing minor septa and major septa never withdrawn from the axis. Similarities to
<taxonomicName id="0691A3837F0DFFECFA85FE34FC1FFD88" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFA85FE34FCBAFD88" italics="true" pageId="20" pageNumber="21">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F0DFFECFCDEFE14FC1FFD88" author="Hudson, R. G. S." box="[866,1021,491,515]" pageId="20" pageNumber="21" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
are evident and
<bibRefCitation id="A500A5F17F0DFFECFB7FFE14FA7FFD88" author="Fedorowski, J." box="[1219,1437,491,515]" pageId="20" pageNumber="21" pagination="1 - 37" refId="ref25654" refString="Fedorowski, J. (2009 a). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin, Ukraine. Part 1. Introductory considerations and the genus Rotiphyllum Hudson, 1942. Acta Geologica Polonica, 59 (1), 1 - 37." type="journal article" year="2009">Fedorowski (2009a)</bibRefCitation>
moved several species described under “
<taxonomicName id="0691A3837F0DFFECFB57FDF4FA80FDA8" box="[1259,1378,523,547]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFB57FDF4FA80FDA8" box="[1259,1378,523,547]" italics="true" pageId="20" pageNumber="21">Zaphrentis</emphasis>
</taxonomicName>
” and
<emphasis id="F3E504127F0DFFECFC91FDD4FC4CFDC8" box="[813,942,555,579]" italics="true" pageId="20" pageNumber="21">Zaphentites</emphasis>
to the first genus. The main distinctive feature of
<taxonomicName id="0691A3837F0DFFECFCC1FDB4FBEAFDE8" authorityName="Hudson" authorityYear="1941" box="[893,1032,587,611]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFCC1FDB4FBEAFDE8" box="[893,1032,587,611]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
is the morphology of the axial stereocolumn.
<bibRefCitation id="A500A5F17F0DFFECFC26FD94FB91FD08" author="Fedorowski, J." box="[922,1139,619,643]" pageId="20" pageNumber="21" pagination="109 - 161" refId="ref25826" refString="Fedorowski, J. (2012 b). The new Upper Serpukhovian genus Zaphrufimia and homeomorphism in some rugose corals. Palaeontogr. Abteilung, 296, 109 - 161." type="journal article" year="2012">Fedorowski (2012b)</bibRefCitation>
also moved several species from
<taxonomicName id="0691A3837F0DFFECFCCEFD74FC1FFD28" authorityName="Hudson" authorityYear="1941" box="[882,1021,651,675]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFCCEFD74FC1FFD28" box="[882,1021,651,675]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
to
<taxonomicName id="0691A3837F0DFFECFB8FFD74FA74FD28" authority="Fedorowski, 2012" authorityName="Fedorowski" authorityYear="2012" box="[1075,1430,651,675]" class="Anthozoa" genus="Zaphrufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFB8FFD74FB5DFD28" box="[1075,1215,651,675]" italics="true" pageId="20" pageNumber="21">Zaphrufimia</emphasis>
Fedorowski, 2012
</taxonomicName>
, based on the septal development.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFC91FD34FC4DFA68" pageId="20" pageNumber="21" type="distribution">
<paragraph id="C12ED8007F0DFFECFC91FD34FC4DFA68" blockId="20.[813,1437,235,1507]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFC91FD34FB3AFD68" bold="true" box="[813,1240,715,739]" pageId="20" pageNumber="21">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F0DFFECFB5EFD34FA8FFD68" authorityName="Hudson" authorityYear="1941" box="[1250,1389,715,739]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFB5EFD34FA8FFD68" box="[1250,1389,715,739]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
is a common genus through the Mississippian (Tournaisian ViséanSerpukhovian) of the Palaeotethys and adjoining epicontinental seas from Northwest Europe to
<collectingCountry id="B98698907F0DFFECFA8AFCD3FA7EFCC8" box="[1334,1436,811,835]" name="China" pageId="20" pageNumber="21">East Asia</collectingCountry>
(Denayer and Hoşgör, 2014, cum lit.;
<bibRefCitation id="A500A5F17F0DFFECFB5DFCB4FC83FC08" author="Rodriguez, S. &amp; Said, I. &amp; Somerville, I. D. &amp; Cozar, P. &amp; Coronado, I." pageId="20" pageNumber="21" pagination="29 - 42" refId="ref28812" refString="Rodriguez, S., Said, I., Somerville, I. D., Cozar, P., &amp; Coronado, I. (2016). Serpukhovian coral assemblages from Idmarrach and Tirhela Formations (Adarouch, Morocco). Geologica Belgica, 19 (1 - 2), 29 - 42." type="journal article" year="2016">Rodríguez et al., 2016</bibRefCitation>
;
<bibRefCitation id="A500A5F17F0DFFECFCD3FC93FBCDFC08" author="Wang, X. D. &amp; Yang, S. R. &amp; Yao, L. &amp; Sugiyama, T. &amp; Hu, K. Y." box="[879,1071,875,900]" pageId="20" pageNumber="21" pagination="603 - 63" refId="ref29952" refString="Wang, X. D., Yang, S. R., Yao, L., Sugiyama, T., &amp; Hu, K. Y. (2022). Carboniferous biostratigraphy of rugose corals. In S. G. Lucas, J. W. Schneider, X. Wang, S. Nikolaeva (Eds), The Carboniferous timescale. Geological Society, London, Special Publications. 512 (2022). 603 - 63." type="journal article" year="2022">Wang et al., 2022</bibRefCitation>
). It is also known from the Mississippian of the conterminous
<collectingCountry id="B98698907F0DFFECFB2AFC74FB2BFC28" box="[1174,1225,907,931]" name="United States of America" pageId="20" pageNumber="21">USA</collectingCountry>
(fide
<bibRefCitation id="A500A5F17F0DFFECFAAEFC73FC83FC48" author="Wang, X. D. &amp; Yang, S. R. &amp; Yao, L. &amp; Sugiyama, T. &amp; Hu, K. Y." pageId="20" pageNumber="21" pagination="603 - 63" refId="ref29952" refString="Wang, X. D., Yang, S. R., Yao, L., Sugiyama, T., &amp; Hu, K. Y. (2022). Carboniferous biostratigraphy of rugose corals. In S. G. Lucas, J. W. Schneider, X. Wang, S. Nikolaeva (Eds), The Carboniferous timescale. Geological Society, London, Special Publications. 512 (2022). 603 - 63." type="journal article" year="2022">Wang et al., 2022</bibRefCitation>
). From the
<collectingRegion id="035516E27F0DFFECFC5FFC54FB6EFC48" box="[995,1164,939,963]" country="Egypt" name="Shamal Sina'" pageId="20" pageNumber="21">Sinai Peninsula</collectingRegion>
,
<collectingCountry id="B98698907F0DFFECFB24FC53FB3BFC4F" box="[1176,1241,940,964]" name="Egypt" pageId="20" pageNumber="21">Egypt</collectingCountry>
, it was previously reported from the late Viséan Um Bogma Formation (
<bibRefCitation id="A500A5F17F0DFFECFC88FC14FBE7FB88" author="Kora, M. &amp; Jux, U." box="[820,1029,1003,1027]" pageId="20" pageNumber="21" pagination="85 - 98" refId="ref27066" refString="Kora, M., &amp; Jux, U. (1986). On the early Carboniferous macrofauna from the Um Bogma Formation, Sinai. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, 2, 85 - 98." type="journal article" year="1986">Kora and Jux, 1986</bibRefCitation>
).
<taxonomicName id="0691A3837F0DFFECFBA7FC14FB44FB88" authorityName="Hudson" authorityYear="1941" box="[1051,1190,1003,1027]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFBA7FC14FB44FB88" box="[1051,1190,1003,1027]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
is rare in the Pennsylvanian, reported to our knowledge only from northern
<collectingCountry id="B98698907F0DFFECFC91FBD4FC88FBC8" box="[813,874,1067,1091]" name="Spain" pageId="20" pageNumber="21">Spain</collectingCountry>
(“Westphalian D” = Asturian, late Moscovian, de Groot, 1963;
<bibRefCitation id="A500A5F17F0DFFECFC7FFBB4FAC2FBE8" author="Rodriguez, S. &amp; Kullmann, J." box="[963,1312,1099,1123]" pageId="20" pageNumber="21" pagination="19 - 40" refId="ref28690" refString="Rodriguez, S., &amp; Kullmann, J. (1990). Hornformige Einzelkorallen (Rugosa) aus spatoberkarbonischen Flachwasser-Ablagerungen des Kantabrischen Gebirges (Nordspanien). Palaeontographica Abteilung A, 19 - 40." type="book chapter" year="1990">Rodríguez and Kullmann, 1990</bibRefCitation>
;
<bibRefCitation id="A500A5F17F0DFFECFA8CFBB4FBF8FB08" author="Kullmann, J. &amp; Rodriguez, S." pageId="20" pageNumber="21" pagination="15 - 22" refId="ref27842" refString="Kullmann, J., &amp; Rodriguez, S. (1994). Biostratigraphic range and biogeographic relationships of the undissepimented solitary corals from the Picos de Europa Formation (Moscovian, Cantabrian Mountains, NW Spain). Courier Forschungsinstitut Senckenberg, 172, 15 - 22." type="journal article" year="1994">Kullmann and Rodríguez, 1994</bibRefCitation>
and
<bibRefCitation id="A500A5F17F0DFFECFBE1FB94FABEFB08" author="Rodriguez, S. &amp; Coronado, I. &amp; Rodriguez-Castro, I." box="[1117,1372,1131,1155]" pageId="20" pageNumber="21" pagination="87 - 95" refId="ref28771" refString="Rodriguez, S., Coronado, I., &amp; Rodriguez-Castro, I. (2022). Corals from Asturian substage in Cantabrian Mountains: A review. Episodes, 45 (1), 87 - 95." type="journal article" year="2022">Rodríguez et al., 2022</bibRefCitation>
), but both species known (
<taxonomicName id="0691A3837F0DFFECFBA0FB73FADFFB28" authority="de Groot, 1963" authorityName="de Groot" authorityYear="1963" box="[1052,1341,1163,1187]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="clithria">
<emphasis id="F3E504127F0DFFECFBA0FB73FB6EFB28" box="[1052,1164,1163,1187]" italics="true" pageId="20" pageNumber="21">Z. clithria</emphasis>
de Groot, 1963
</taxonomicName>
,
<taxonomicName id="0691A3837F0DFFECFAF1FB73FBA9FB48" authority="de Groot, 1963" authorityName="de Groot" authorityYear="1963" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="paralleloides">
<emphasis id="F3E504127F0DFFECFAF1FB73FC71FB48" italics="true" pageId="20" pageNumber="21">Z. paralleloides</emphasis>
de Groot, 1963
</taxonomicName>
) were moved to
<taxonomicName id="0691A3837F0DFFECFAABFB54FB87FB68" authority="(de Groot, 1963)" baseAuthorityName="de Groot" baseAuthorityYear="1963" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="paralloides">
<emphasis id="F3E504127F0DFFECFAABFB54FC44FB68" italics="true" pageId="20" pageNumber="21">Rotiphyllum paralloides</emphasis>
(de Groot, 1963)
</taxonomicName>
by
<bibRefCitation id="A500A5F17F0DFFECFB32FB34FAB8FB68" author="Fedorowski, J." box="[1166,1370,1227,1251]" pageId="20" pageNumber="21" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski (2004)</bibRefCitation>
. Rodríguez et al., (2022, tab. 1) listed
<taxonomicName id="0691A3837F0DFFECFB23FB14FAC8FA88" authorityName="Hudson" authorityYear="1941" box="[1183,1322,1259,1283]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFB23FB14FAC8FA88" box="[1183,1322,1259,1283]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
also from the late Moscovian (Asturian) of North America (Ellesmere Island, Arctic
<collectingCountry id="B98698907F0DFFECFBBFFAD4FBB6FAC8" box="[1027,1108,1323,1347]" name="Canada" pageId="20" pageNumber="21">Canada</collectingCountry>
—not supported by our literature review) and from the UralianArctic realm). Herein, we describe
<taxonomicName id="0691A3837F0DFFECFC06FA94FBA7FA08" authorityName="Hudson" authorityYear="1941" box="[954,1093,1387,1411]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0DFFECFC06FA94FBA7FA08" box="[954,1093,1387,1411]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
</taxonomicName>
from the Kasimovian West of the Gulf of
<collectingRegion id="035516E27F0DFFECFC0AFA74FC0EFA28" box="[950,1004,1419,1443]" country="Egypt" name="As Suways" pageId="20" pageNumber="21">Suez</collectingRegion>
,
<collectingCountry id="B98698907F0DFFECFC46FA73FBDEFA2F" box="[1018,1084,1420,1444]" name="Egypt" pageId="20" pageNumber="21">Egypt</collectingCountry>
. The genus was also described from the Lower and Middle Permian of
<collectingCountry id="B98698907F0DFFECFB61FA54FAFDFA48" box="[1245,1311,1451,1475]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
(fide
<bibRefCitation id="A500A5F17F0DFFECFAE0FA53FC42FA68" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." pageId="20" pageNumber="21" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:detail id="7D654C8D6623EE0ACF718A06D2C2A99F" type="pubDate">
<mods:number id="A1C689BD7CE484E29E5B8F3E13271713">2023-11-27</mods:number>
</mods:detail>
<mods:detail id="C81589E60E2C709B5E937033F4F2A6EF" type="volume">
<mods:number id="16B870E06FE8E604E09539F4D43272EB">142</mods:number>
</mods:detail>
<mods:detail id="D31F46E5B3E187AC5E14E2905C408B57" type="issue">
<mods:number id="3520E51E0D009A7E0B0ADA6AAA158661">1</mods:number>
</mods:detail>
<mods:extent id="D241284E0B0CC3D2E05C8E9F2D844944" unit="page">
<mods:start id="B5DC59C711A41CB8E11F564D071289D9">1</mods:start>
<mods:end id="353ECC249366559479C8E5D4C8932A25">39</mods:end>
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<mods:location id="91161E90066E3C447EAAEC746ECC5AE4">
<mods:url id="7FE6767B3BA4366ABDAEF1D622BA651D">http://dx.doi.org/10.1186/s13358-023-00296-0</mods:url>
</mods:location>
<mods:classification id="CF554512BE8F28990D23E269D145164C">journal article</mods:classification>
<mods:identifier id="6476DAE0278ED2744000FA97F07A0DAA" type="DOI">10.1186/s13358-023-00296-0</mods:identifier>
<mods:identifier id="057B215BD6C4DF056E06EA49D9999499" type="ISSN">1664-2384</mods:identifier>
<mods:identifier id="81634F566210C7DF8C9A1B38F7893F48" type="Zenodo-Dep">12003174</mods:identifier>
</mods:mods>
<treatment id="493869167F0DFFEFFC91F9D3FE86FB68" LSID="urn:lsid:plazi:treatment:493869167F0DFFEFFC91F9D3FE86FB68" httpUri="http://treatment.plazi.org/id/493869167F0DFFEFFC91F9D3FE86FB68" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="898B8B8B7F0DFFECFC91F9D3FAC8F9CE" box="[813,1322,1580,1605]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="C12ED8007F0DFFECFC91F9D3FAC8F9CE" blockId="20.[813,1436,1580,1892]" box="[813,1322,1580,1605]" pageId="20" pageNumber="21">
<heading id="9A666F6C7F0DFFECFC91F9D3FAC8F9CE" box="[813,1322,1580,1605]" fontSize="9" level="2" pageId="20" pageNumber="21" reason="8">
<taxonomicName id="0691A3837F0DFFECFC91F9D3FAC4F9CE" authority="(Carruthers, 1910)" baseAuthorityName="Carruthers" baseAuthorityYear="1910" box="[813,1318,1580,1605]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Stauriida" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0DFFECFC91F9D3FBB7F9CF" bold="true" box="[813,1109,1580,1604]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFC91F9D3FC20F9CF" bold="true" box="[813,962,1580,1604]" italics="true" pageId="20" pageNumber="21">Zaphrentites</emphasis>
cf.
<emphasis id="F3E504127F0DFFECFC56F9D3FBB7F9CF" bold="true" box="[1002,1109,1580,1604]" italics="true" pageId="20" pageNumber="21">parallela</emphasis>
</emphasis>
(Carruthers, 1910)
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFC91F9B3FAC4F96E" pageId="20" pageNumber="21" type="description">
<paragraph id="C12ED8007F0DFFECFC91F9B3FB88F9EE" blockId="20.[813,1436,1580,1892]" box="[813,1130,1612,1637]" pageId="20" pageNumber="21">
(
<figureCitation id="59AAC4857F0DFFECFC88F9B3FC71F9EE" box="[820,915,1612,1637]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="20" pageNumber="21">Fig. 11A</figureCitation>
<figureCitation id="59AAC4857F0DFFECFC2FF9B2FC20F9EE" box="[915,962,1613,1637]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="20" pageNumber="21">1A</figureCitation>
<figureCitation id="59AAC4857F0DFFECFC7DF9B2FC2FF9EE" box="[961,973,1613,1637]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="20" pageNumber="21">3</figureCitation>
, B
<figureCitation id="59AAC4857F0DFFECFC56F9B2FBF7F9EE" box="[1002,1045,1613,1637]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="20" pageNumber="21">1B</figureCitation>
<figureCitation id="59AAC4857F0DFFECFBA9F9B2FBC1F9EE" box="[1045,1059,1613,1637]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="20" pageNumber="21">4</figureCitation>
,
<figureCitation id="59AAC4857F0DFFECFB92F9B2FBBCF9EE" box="[1070,1118,1613,1637]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="20" pageNumber="21">12D</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F0DFFECFC91F993FC8CF92E" blockId="20.[813,1436,1580,1892]" pageId="20" pageNumber="21">
cf. 1910
<taxonomicName id="0691A3837F0DFFECFC30F993FB15F90F" authority="Carruthers." authorityName="Carruthers." box="[908,1271,1644,1668]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0DFFECFC30F993FB8FF90F" box="[908,1133,1644,1668]" italics="true" pageId="20" pageNumber="21">Zaphrentis parallela</emphasis>
Carruthers.
</taxonomicName>
- p. 533, Pl. 37,
<figureCitation id="59AAC4857F0DFFECFC91F973FC8BF92E" box="[813,873,1676,1701]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="C12ED8007F0DFFECFC91F953FAC4F96E" blockId="20.[813,1436,1580,1892]" pageId="20" pageNumber="21">
cf. 2014
<taxonomicName id="0691A3837F0DFFECFC29F953FAC7F94F" authority="(Carruthers)" baseAuthorityName="Carruthers" baseAuthorityYear="1910" box="[917,1317,1708,1732]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0DFFECFC29F953FB6CF94F" box="[917,1166,1708,1732]" italics="true" pageId="20" pageNumber="21">Zaphrentites parallela</emphasis>
(Carruthers)
</taxonomicName>
.- Denayer and Hoşgör, p. 350,
<figureCitation id="59AAC4857F0DFFECFBBFF933FBA2F96E" box="[1027,1088,1740,1765]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
: 6ad, 14 (cum syn.).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0DFFECFC91F913FC5BF8EF" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="C12ED8007F0DFFECFC91F913FC5BF8EF" blockId="20.[813,1436,1580,1892]" pageId="20" pageNumber="21">
<emphasis id="F3E504127F0DFFECFC91F913FC75F88F" bold="true" box="[813,919,1772,1796]" pageId="20" pageNumber="21">Material:</emphasis>
Three corallites (RAh 109, RAh 111 and RAh 118); collected from the basal shales of the lower member of the Aheimer Formation. Nine transverse thin-sections are available.
</paragraph>
</subSubSection>
<caption id="95EE88887F0CFFEDFF1BF9ADFE4BF88D" ID-DOI="http://doi.org/10.5281/zenodo.12003314" ID-Zenodo-Dep="12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="21" pageNumber="22" startId="21.[167,198,1618,1638]" targetBox="[287,1301,240,1602]" targetPageId="21" targetType="figure">
<paragraph id="C12ED8007F0CFFEDFF1BF9ADFE4BF88D" blockId="21.[167,1415,1618,1798]" pageId="21" pageNumber="22">
<emphasis id="F3E504127F0CFFEDFF1BF9ADFEF1F9ED" bold="true" box="[167,275,1618,1638]" pageId="21" pageNumber="22">Fig. 11 A, B</emphasis>
<taxonomicName id="0691A3837F0CFFEDFEA4F9ADFD9EF9ED" authority="(Carruthers, 1910)" baseAuthorityName="Carruthers" baseAuthorityYear="1910" box="[280,636,1618,1638]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0CFFEDFEA4F9ADFE9CF9ED" box="[280,382,1618,1638]" italics="true" pageId="21" pageNumber="22">Zaphrentites</emphasis>
cf.
<emphasis id="F3E504127F0CFFEDFE27F9ADFE00F9ED" box="[411,482,1618,1638]" italics="true" pageId="21" pageNumber="22">parallela</emphasis>
(Carruthers,1910)
</taxonomicName>
.
<emphasis id="F3E504127F0CFFEDFD38F9ADFD23F9ED" bold="true" box="[644,705,1618,1638]" pageId="21" pageNumber="22">A1A3</emphasis>
Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in
<emphasis id="F3E504127F0CFFEDFD07F993FD36F90B" bold="true" box="[699,724,1644,1664]" pageId="21" pageNumber="22">A1</emphasis>
to the last mature stage below the calice in
<emphasis id="F3E504127F0CFFEDFBF7F993FB81F90B" bold="true" box="[1099,1123,1644,1664]" pageId="21" pageNumber="22">A3</emphasis>
.
<emphasis id="F3E504127F0CFFEDFBD0F993FB66F90B" bold="true" box="[1132,1156,1644,1664]" pageId="21" pageNumber="22">B1</emphasis>
External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated.
<emphasis id="F3E504127F0CFFEDFB1BF978FB00F910" bold="true" box="[1191,1250,1671,1691]" pageId="21" pageNumber="22">B2B4</emphasis>
Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in
<emphasis id="F3E504127F0CFFEDFB3FF95DFB79F93D" bold="true" box="[1155,1179,1698,1718]" pageId="21" pageNumber="22">B2</emphasis>
to the last mature section below the calice floor in B4.
<emphasis id="F3E504127F0CFFEDFE29F943FE40F95B" bold="true" box="[405,418,1724,1744]" pageId="21" pageNumber="22">C</emphasis>
<taxonomicName id="0691A3837F0CFFEDFE1AF943FDDFF95B" box="[422,573,1724,1744]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Cnidaria" rank="species" species="undefined-C">
<emphasis id="F3E504127F0CFFEDFE1AF943FDEEF95B" box="[422,524,1724,1744]" italics="true" pageId="21" pageNumber="22">Zaphrentites</emphasis>
sp.
<emphasis id="F3E504127F0CFFEDFD91F943FDDFF95B" bold="true" box="[557,573,1724,1744]" pageId="21" pageNumber="22">C</emphasis>
</taxonomicName>
<emphasis id="F3E504127F0CFFEDFD81F943FDA7F95B" bold="true" box="[573,581,1724,1744]" pageId="21" pageNumber="22">1</emphasis>
External view of the specimen RAh (84).
<emphasis id="F3E504127F0CFFEDFC22F943FC3AF95B" bold="true" box="[926,984,1724,1744]" pageId="21" pageNumber="22">C2C4</emphasis>
Successive transverse sections; from the immature section in
<emphasis id="F3E504127F0CFFEDFF42F928FEF4F960" bold="true" box="[254,278,1751,1771]" pageId="21" pageNumber="22">C2</emphasis>
to the last mature section below the calice floor in
<emphasis id="F3E504127F0CFFEDFD75F928FD02F960" bold="true" box="[713,736,1751,1771]" pageId="21" pageNumber="22">C4</emphasis>
. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
</paragraph>
</caption>
<caption id="95EE88887F0FFFEEFF1BFAF8FDABFAE0" ID-DOI="http://doi.org/10.5281/zenodo.12003322" ID-Zenodo-Dep="12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="22" pageNumber="23" startId="22.[167,198,1287,1307]" targetBox="[286,1300,248,1266]" targetPageId="22" targetType="figure">
<paragraph id="C12ED8007F0FFFEEFF1BFAF8FDABFAE0" blockId="22.[167,1398,1287,1387]" pageId="22" pageNumber="23">
<emphasis id="F3E504127F0FFFEEFF1BFAF8FF04FA90" bold="true" box="[167,230,1287,1307]" pageId="22" pageNumber="23">Fig. 12</emphasis>
Relation between number of major septa and coralla alar diameters for:
<emphasis id="F3E504127F0FFFEEFCEDFAF8FCBDFA90" bold="true" box="[849,863,1287,1307]" pageId="22" pageNumber="23">A</emphasis>
<taxonomicName id="0691A3837F0FFFEEFCDFFAF8FBFBFA90" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[867,1049,1287,1307]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F0FFFEEFCDFFAF8FBFBFA90" box="[867,1049,1287,1307]" italics="true" pageId="22" pageNumber="23">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F0FFFEEFBA2FAF8FBAEFA90" box="[1054,1100,1287,1307]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="0691A3837F0FFFEEFBEDFAF8FAC4FA90" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[1105,1318,1287,1307]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F0FFFEEFBEDFAF8FB6CFA90" box="[1105,1166,1287,1307]" italics="true" pageId="22" pageNumber="23">L aucta</emphasis>
Fedorowski,1987
</taxonomicName>
,
<taxonomicName id="0691A3837F0FFFEEFA92FAF8FADFFA90" box="[1326,1341,1287,1307]" pageId="22" pageNumber="23">
<emphasis id="F3E504127F0FFFEEFA92FAF8FADFFA90" box="[1326,1341,1287,1307]" italics="true" pageId="22" pageNumber="23">L.</emphasis>
</taxonomicName>
<emphasis id="F3E504127F0FFFEEFF1BFADDFEE2FABD" box="[167,256,1314,1334]" italics="true" pageId="22" pageNumber="23">asymetrica</emphasis>
<bibRefCitation id="A500A5F17F0FFFEEFEB9FADDFE61FABD" author="Soshkina, E. D." box="[261,387,1314,1334]" pageId="22" pageNumber="23" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
described by Chwieduk (2013),
<taxonomicName id="0691A3837F0FFFEEFD2DFADDFC1CFABD" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" box="[657,1022,1314,1334]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F0FFFEEFD2DFADDFD18FABD" box="[657,762,1314,1334]" italics="true" pageId="22" pageNumber="23">L. canadense</emphasis>
<bibRefCitation id="A500A5F17F0FFFEEFD43FADDFC1CFABD" author="Fedorowski, J. &amp; Bamber, E. W." box="[767,1022,1314,1334]" pageId="22" pageNumber="23" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
</taxonomicName>
and the current studied
<taxonomicName id="0691A3837F0FFFEEFB6CFADDFE44FADA" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F0FFFEEFB6CFADDFB3BFABD" box="[1232,1241,1314,1334]" italics="true" pageId="22" pageNumber="23">L</emphasis>
. cf.
<emphasis id="F3E504127F0FFFEEFB44FADDFAB2FABD" box="[1272,1360,1314,1334]" italics="true" pageId="22" pageNumber="23">canadense</emphasis>
<bibRefCitation id="A500A5F17F0FFFEEFF1BFAC2FE44FADA" author="Fedorowski, J. &amp; Bamber, E. W." box="[167,422,1341,1361]" pageId="22" pageNumber="23" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
</taxonomicName>
.
<emphasis id="F3E504127F0FFFEEFE12FAC2FE58FADA" bold="true" box="[430,442,1341,1361]" pageId="22" pageNumber="23">B</emphasis>
<taxonomicName id="0691A3837F0FFFEEFE03FAC2FD70FADA" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[447,658,1341,1361]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="galalaensis" status="sp. nov.">
<emphasis id="F3E504127F0FFFEEFE03FAC2FD70FADA" box="[447,658,1341,1361]" italics="true" pageId="22" pageNumber="23">Monophyllum galalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F0FFFEEFD2BFAC2FD21FADA" box="[663,707,1341,1361]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="0691A3837F0FFFEEFD50FAC2FC2DFADA" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[748,975,1341,1361]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F0FFFEEFD50FAC2FCA7FADA" box="[748,837,1341,1361]" italics="true" pageId="22" pageNumber="23">M. parvum</emphasis>
<bibRefCitation id="A500A5F17F0FFFEEFCF6FAC2FC2DFADA" author="Fomichev, V." box="[842,975,1341,1361]" pageId="22" pageNumber="23" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
<emphasis id="F3E504127F0FFFEEFC6BFAC2FC06FADA" bold="true" box="[983,996,1341,1361]" pageId="22" pageNumber="23">C</emphasis>
<taxonomicName id="0691A3837F0FFFEEFC54FAC2FB17FADA" authority="de Groot, 1963" authorityName="de Groot" authorityYear="1963" box="[1000,1269,1341,1361]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="exile">
<emphasis id="F3E504127F0FFFEEFC54FAC2FB90FADA" box="[1000,1138,1341,1361]" italics="true" pageId="22" pageNumber="23">Rotiphyllum exile</emphasis>
de Groot, 1963
</taxonomicName>
.
<emphasis id="F3E504127F0FFFEEFB41FAC2FAEEFADA" bold="true" box="[1277,1292,1341,1361]" pageId="22" pageNumber="23">D</emphasis>
<taxonomicName id="0691A3837F0FFFEEFAACFAC2FA94FADA" box="[1296,1398,1341,1361]" pageId="22" pageNumber="23">
<emphasis id="F3E504127F0FFFEEFAACFAC2FA94FADA" box="[1296,1398,1341,1361]" italics="true" pageId="22" pageNumber="23">Zaphrentites</emphasis>
</taxonomicName>
cf.
<emphasis id="F3E504127F0FFFEEFF03FAA8FEE4FAE0" box="[191,262,1367,1387]" italics="true" pageId="22" pageNumber="23">parallela</emphasis>
(Carruthers,1910) and
<taxonomicName id="0691A3837F0FFFEEFE76FAA8FDABFAE0" box="[458,585,1367,1387]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F0FFFEEFE76FAA8FDD2FAE0" box="[458,560,1367,1387]" italics="true" pageId="22" pageNumber="23">Zaphrentites</emphasis>
sp
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="898B8B8B7F0FFFEFFF2BFA43FF28FC88" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="C12ED8007F0FFFEEFF2BFA43FD86F9FF" blockId="22.[151,775,1468,1909]" pageId="22" pageNumber="23">
<emphasis id="F3E504127F0FFFEEFF2BFA43FEC5FA5F" bold="true" box="[151,295,1468,1492]" pageId="22" pageNumber="23">Description:</emphasis>
External characters: Straight, conical corallites, about 1.5 cm long, slightly curved in the immature stage. Calyx is largely eroded, apex not preserved. Septal furrows and delicate growth lines are distinctly marked on the external wall (
<figureCitation id="59AAC4857F0FFFEEFE37F9C3FE0EF9DF" box="[395,492,1596,1620]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="22" pageNumber="23">Fig. 11B</figureCitation>
<figureCitation id="59AAC4857F0FFFEEFE50F9C3FE14F9DF" box="[492,502,1596,1620]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="22" pageNumber="23">1</figureCitation>
). An attachment scar is visible in one corallite near the apical part.
</paragraph>
<paragraph id="C12ED8007F0FFFEEFF11F983FB8FF8FF" blockId="22.[151,775,1468,1909]" lastBlockId="22.[813,1437,1468,1908]" pageId="22" pageNumber="23">
Internal characters: In the earliest ontogenetic preserved stage (
<figureCitation id="59AAC4857F0FFFEEFE8EF963FE74F93F" box="[306,406,1692,1716]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="22" pageNumber="23">Fig. 11A</figureCitation>
<figureCitation id="59AAC4857F0FFFEEFE2BF963FE43F93F" box="[407,417,1692,1716]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="22" pageNumber="23">1</figureCitation>
), septa are very thick, reaching the corallite axis. There are 16 septa at a corallite diameter of about
<quantity id="066975E57F0FFFEEFE8EF923FE94F97F" box="[306,374,1756,1780]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="mm" value="3.0">3 mm</quantity>
; of them the counter septum is the thickest. At diameter of 3.7 mm (
<figureCitation id="59AAC4857F0FFFEEFDB4F903FD86F89F" box="[520,612,1788,1812]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="22" pageNumber="23">Fig. 11B</figureCitation>
<figureCitation id="59AAC4857F0FFFEEFDD8F903FD90F89F" box="[612,626,1788,1812]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="22" pageNumber="23">2</figureCitation>
), major septa are still thick, they join in the centre forming a thick stereocolumn; a parallel sided cardinal fossula can be recognised. In the subsequent early mature growth stage at a diameter of
<quantity id="066975E57F0FFFEEFC08FA43FC1AFA5F" box="[948,1016,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
, there are 20 major septa (
<figureCitation id="59AAC4857F0FFFEEFA9FFA43FA61FA5F" box="[1315,1411,1468,1492]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="22" pageNumber="23">Fig. 11A</figureCitation>
<figureCitation id="59AAC4857F0FFFEEFA3FFA43FA73FA5F" box="[1411,1425,1468,1492]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="22" pageNumber="23">2</figureCitation>
). Minor septa are absent. The axial ends of major septa join to form an axial stereocolumn slightly staggered towards the counter quadrants.
<collectingRegion id="035516E27F0FFFEEFB85F9E3FB94F9BF" box="[1081,1142,1564,1588]" country="Spain" name="Ceuta" pageId="22" pageNumber="23">Septa</collectingRegion>
of the counter quadrants are radially arranged, the counter septum is slightly thicker and has the same length as the adjacent septa. Those in the cardinal quadrants are grouped pinnately around a long parallel sided cardinal fossula. The thin cardinal septum is straight, long and reaches beyond the corallite axis to join the stereocolumn at the upper end of the cardinal fossula in the counter quadrants. It further thins towards the axis. Further mature, at a diameter of 6.5 mm, there are 21 septa (
<figureCitation id="59AAC4857F0FFFEEFBD8F8C3FB23F8DF" box="[1124,1217,1852,1876]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="22" pageNumber="23">Fig. 11B</figureCitation>
<figureCitation id="59AAC4857F0FFFEEFB7EF8C3FB2CF8DF" box="[1218,1230,1852,1876]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="22" pageNumber="23">3</figureCitation>
); the cardinal septum becomes slightly shorter.
</paragraph>
<paragraph id="C12ED8007F0EFFEFFF11FF13FF28FC88" blockId="23.[151,775,235,1251]" pageId="23" pageNumber="24">
The uppermost mature transverse section near the calice floor (
<figureCitation id="59AAC4857F0EFFEFFEBCFEF4FEBDFEA8" box="[256,351,267,291]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="23" pageNumber="24">Fig. 11A</figureCitation>
<figureCitation id="59AAC4857F0EFFEFFEE3FEF4FE89FEA8" box="[351,363,267,291]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="23" pageNumber="24">3</figureCitation>
, B
<figureCitation id="59AAC4857F0EFFEFFE3BFEF4FE76FEA8" box="[391,404,267,291]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="23" pageNumber="24">4</figureCitation>
) is slightly deformed due to compaction and some of the septa are broken. In this stage there are maximum 26 septa in 8.6 mm (
<figureCitation id="59AAC4857F0EFFEFFDEEFEB4FD4CFEE8" box="[594,686,331,355]" captionStart="Fig" captionStartId="21.[167,198,1618,1638]" captionTargetBox="[287,1301,240,1602]" captionTargetId="figure-13@21.[283,1303,236,1605]" captionTargetPageId="21" captionText="Fig. 11 A, B Zaphrentites cf. parallela (Carruthers,1910). A1A3 Successive transverse sections in the specimen RAh (118), starting from the early immature stage with zaphrentoid arrangement of the septa in A1 to the last mature stage below the calice in A3. B1 External lateral view of the specimen RAh (109), showing strong longitudinal ribbing on the corallite wall. Positions of thin-sections indicated. B2B4 Successive transverse sections represent the ontogenetic development, starting from the immature section in the apical part in B2 to the last mature section below the calice floor in B4. C Zaphrentites sp. C1 External view of the specimen RAh (84). C2C4 Successive transverse sections; from the immature section in C2 to the last mature section below the calice floor in C4. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003314" httpUri="https://zenodo.org/record/12003314/files/figure.png" pageId="23" pageNumber="24">Fig. 11B</figureCitation>
<figureCitation id="59AAC4857F0EFFEFFD11FEB4FD59FEE8" box="[685,699,331,355]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="23" pageNumber="24">4</figureCitation>
). Axial ends of the major septa in cardinal quadrants are bent to sides with the two pericardinal septa, curved most distinctly. Such septal morphology makes the cardinal fossula opens towards the free axial area. The cardinal fossula is slightly deformed. The cardinal septum is short, and the counter is slightly longer and thicker than the adjacent septa.
<taxonomicName id="0691A3837F0EFFEFFEFDFDD4FE4BFDC8" box="[321,425,555,579]" genus="All" pageId="23" pageNumber="24" rank="species" species="major">All major</taxonomicName>
septa are thick at the base, join at their rhopaloid ends around the well-developed cardinal fossula, which invades the counterpart, being longer than the radius of the corallite. Pseudoalar fossulae are not markedly developed. Minor septa are absent from the corallite lumen, they are visible in the stereoplasm of the wall.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0EFFEFFF2BFCF4FE86FB68" pageId="23" pageNumber="24" type="discussion">
<paragraph id="C12ED8007F0EFFEFFF2BFCF4FE86FB68" blockId="23.[151,775,235,1251]" pageId="23" pageNumber="24">
<emphasis id="F3E504127F0EFFEFFF2BFCF4FEFFFCA8" bold="true" box="[151,285,779,803]" pageId="23" pageNumber="24">Discussion:</emphasis>
According to the septal arrangement and the axial stereocolumn, the current species clearly belongs to the genus
<taxonomicName id="0691A3837F0EFFEFFEB9FCB4FE72FCE8" authorityName="Hudson" authorityYear="1941" box="[261,400,843,867]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFEB9FCB4FE72FCE8" box="[261,400,843,867]" italics="true" pageId="23" pageNumber="24">Zaphrentites</emphasis>
</taxonomicName>
. It is closely similar to
<taxonomicName id="0691A3837F0EFFEFFD3BFCB3FE84FC08" authority="(Carruthers, 1910)" baseAuthorityName="Carruthers" baseAuthorityYear="1910" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0EFFEFFD3BFCB3FCE4FCE8" box="[647,774,843,867]" italics="true" pageId="23" pageNumber="24">Z. parallela</emphasis>
(Carruthers, 1910)
</taxonomicName>
in the septal arrangement and more or less in the shape of the cardinal fossula, with the cardinal septum withdrawn in late stages. However, our specimens reach larger n:d ratios (
<figureCitation id="59AAC4857F0EFFEFFE64FC34FDD4FC68" box="[472,566,971,995]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="23" pageNumber="24">Fig. 12D</figureCitation>
) in the last mature stage near the calice (26:8.6 vs. 20:5.6).
<taxonomicName id="0691A3837F0EFFEFFDCBFC13FDF8FBA8" authority="(Milne-Edwards and Haime, 1851)" baseAuthorityName="Milne-Edwards and Haime" baseAuthorityYear="1851" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="delanouei">
<emphasis id="F3E504127F0EFFEFFDCBFC13FCE5FB88" box="[631,775,1003,1027]" italics="true" pageId="23" pageNumber="24">Z. delanouei</emphasis>
(
<bibRefCitation id="A500A5F17F0EFFEFFF22FBF4FDF2FBA8" author="Milne-Edwards, H. &amp; Haime, J." box="[158,528,1035,1059]" pageId="23" pageNumber="24" pagination="72 - 146" refId="ref28114" refString="Milne-Edwards, H., &amp; Haime, J. (1851). A monograph of the British fossil corals. Palaeontographical Society London, 72 - 146." type="book chapter" year="1851">Milne-Edwards and Haime, 1851</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0691A3837F0EFFEFFDE9FBF3FEE3FBC8" authority="Hudson, 1944" authorityName="Hudson" authorityYear="1944" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="crassus">
<emphasis id="F3E504127F0EFFEFFDE9FBF3FD21FBA8" box="[597,707,1036,1059]" italics="true" pageId="23" pageNumber="24">Z. crassus</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFD71FBF4FEE3FBC8" author="Hudson, R. G. S." pageId="23" pageNumber="24" pagination="145 - 154" refId="ref26698" refString="Hudson, R. G. S. (1944). On the Lower Carboniferous corals: Zaphrentites crassus and Z. tenuis, sp. n. Journal of Natural History, 11 (75), 145 - 154." type="journal article" year="1944">Hudson, 1944</bibRefCitation>
</taxonomicName>
are both larger and have more septa than the herein described species. It is quite similar to
<taxonomicName id="0691A3837F0EFFEFFD1FFBB3FF5DFB08" baseAuthorityName="Carruthers" baseAuthorityYear="1910" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="parallela">
<emphasis id="F3E504127F0EFFEFFD1FFBB3FD56FBE8" box="[675,692,1100,1123]" italics="true" pageId="23" pageNumber="24">Z</emphasis>
.
<emphasis id="F3E504127F0EFFEFFD7FFBB4FF5DFB08" italics="true" pageId="23" pageNumber="24">parallela</emphasis>
</taxonomicName>
of Denayer and Hoşgör (2014) from the Lower Carboniferous of
<collectingCountry id="B98698907F0EFFEFFE8DFB74FE9DFB28" box="[305,383,1163,1187]" name="Turkey" pageId="23" pageNumber="24">Turkey</collectingCountry>
. It differs in the n:d ratio of the last mature stage (21:8.5) and in general counts more septa at the same diameter.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,312 @@
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<subSubSection id="898B8B8B7F0EFFEFFC91FCD2FB7DFCCF" box="[813,1183,812,837]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="C12ED8007F0EFFEFFC91FCD2FB7DFCCF" blockId="23.[813,1437,716,1925]" box="[813,1183,812,837]" pageId="23" pageNumber="24">
<heading id="9A666F6C7F0EFFEFFC91FCD2FB7DFCCF" box="[813,1183,812,837]" fontSize="36" level="2" pageId="23" pageNumber="24" reason="3">
Genus
<taxonomicName id="0691A3837F0EFFEFFCC5FCD3FB7BFCCF" authority="Stuckenberg, 1895" authorityName="Stuckenberg" authorityYear="1895" box="[889,1177,812,836]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFCC5FCD3FC29FCCF" bold="true" box="[889,971,812,836]" italics="true" pageId="23" pageNumber="24">Ufimia</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFC6DFCD3FB7BFCCF" author="Stuckenberg, A. A." box="[977,1177,812,836]" pageId="23" pageNumber="24" refId="ref29373" refString="Stuckenberg, A. A. (1895). Korallen und Bryozoen der Steinkohlenablagerungen des Ural und des Timan. Memoires Du Comite Geologique, 10 (3), 244. in Russian and German." type="book" year="1895">Stuckenberg, 1895</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0EFFEFFC91FCB3FA8DFCEF" box="[813,1391,844,869]" pageId="23" pageNumber="24" type="type_taxon">
<paragraph id="C12ED8007F0EFFEFFC91FCB3FA8DFCEF" blockId="23.[813,1437,716,1925]" box="[813,1391,844,869]" pageId="23" pageNumber="24">
<emphasis id="F3E504127F0EFFEFFC91FCB3FC21FCEF" bold="true" box="[813,963,844,868]" pageId="23" pageNumber="24">Type-species</emphasis>
:
<taxonomicName id="0691A3837F0EFFEFFC73FCB2FA8AFCEF" authority="Stuckenberg, 1895" authorityName="Stuckenberg" authorityYear="1895" box="[975,1384,844,869]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="carbonaria">
<emphasis id="F3E504127F0EFFEFFC73FCB2FB79FCEF" box="[975,1179,844,869]" italics="true" pageId="23" pageNumber="24">Ufimia carbonaria</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFB1CFCB3FA8AFCEF" author="Stuckenberg, A. A." box="[1184,1384,844,868]" pageId="23" pageNumber="24" refId="ref29373" refString="Stuckenberg, A. A. (1895). Korallen und Bryozoen der Steinkohlenablagerungen des Ural und des Timan. Memoires Du Comite Geologique, 10 (3), 244. in Russian and German." type="book" year="1895">Stuckenberg, 1895</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0EFFEFFC91FC93FC58FB6E" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="C12ED8007F0EFFEFFC91FC93FC58FB6E" blockId="23.[813,1437,716,1925]" pageId="23" pageNumber="24">
<emphasis id="F3E504127F0EFFEFFC91FC93FC42FC0F" bold="true" box="[813,928,876,900]" pageId="23" pageNumber="24">Diagnosis</emphasis>
: Small solitary corals with zaphrentoid early stages. Alar and counter lateral septa dominating in late stages becoming longer, taller, thicker, and more rhopaloid. Rhopaloid axial ends of major septa commonly conjunct in axial region in middle stages of development, withdrawing and becoming free axially in late stages. Longest and strongest metasepta are commonly midquadrant.
<taxonomicName id="0691A3837F0EFFEFFC23FBB2FC1FFBEE" box="[927,1021,1101,1125]" genus="Cardinal" pageId="23" pageNumber="8" rank="genus">Cardinal</taxonomicName>
and counter protosepta shorten progressively. Minor septa rudimentary to short. Tabular floors tall and axially depressed; dissepiments missing.
<collectingRegion id="035516E27F0EFFEFFC91FB52FC88FB4E" box="[813,874,1197,1221]" country="Spain" name="Ceuta" pageId="23" pageNumber="24">Septa</collectingRegion>
trabecular, uniseriate (after
<bibRefCitation id="A500A5F17F0EFFEFFB17FB52FAF5FB4E" author="Hill, D." box="[1195,1303,1197,1221]" pageId="23" pageNumber="24" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill, 1981</bibRefCitation>
and
<bibRefCitation id="A500A5F17F0EFFEFFAE8FB52FC49FB6E" author="Fedorowski, J." pageId="23" pageNumber="24" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F0EFFE0FC91FB12FD00FA58" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="discussion">
<paragraph id="C12ED8007F0EFFEFFC91FB12FB33F80E" blockId="23.[813,1437,716,1925]" pageId="23" pageNumber="24">
<emphasis id="F3E504127F0EFFEFFC91FB12FC78FA8E" bold="true" box="[813,922,1261,1285]" pageId="23" pageNumber="24">Remarks:</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFC19FB12FB9FFA8E" author="Schindewolf, O. H." box="[933,1149,1261,1285]" pageId="23" pageNumber="24" pagination="1 - 324" refId="ref29039" refString="Schindewolf, O. H. (1942). Zur Kenntnis der Polycoelien und Plerophyllen. Abhandlungen Des Reichsamt Fur Bodenforschung, Neue Serie, 204, 1 - 324." type="journal article" year="1942">Schindewolf (1942)</bibRefCitation>
discussed the genus and gave earlier synonymies. Like following authors (e.g., de Groot, 1963;
<bibRefCitation id="A500A5F17F0EFFEFFC50FAD2FB7CFACF" author="Kullmann, J." box="[1004,1182,1324,1349]" pageId="23" pageNumber="24" refId="ref27620" refString="Kullmann, J. (1965). Rugose Korallen der Cephalopodenfazies und ihre Verbreitung im Devon des sudostlichen Kantabrischen Gebirges (Nordspanien). Akademie der. Wissenschaft und der Literatur Mainz. Abhandlungen der Naturwissenschaftlichen Klasse 2, 136 pp." type="book" year="1965">Kullmann, 1965</bibRefCitation>
,
<bibRefCitation id="A500A5F17F0EFFEFFB0CFAD3FB06FACF" author="Kullmann, J." box="[1200,1252,1324,1348]" pageId="23" pageNumber="24" pagination="771 - 776" refId="ref27696" refString="Kullmann, J. (1968). Associations of rugose corals and cephalopods in the Devonian of the Cantabrian Mountains (Northern Spain). In D. H. Oswald (Ed.), International Symposium on the Devonian System, Calgary 1, 771 - 776." type="journal article" year="1968">1968</bibRefCitation>
), he considered
<taxonomicName id="0691A3837F0EFFEFFC91FAB2FC9BFAEE" authorityName="Stuckenberg" authorityYear="1895" box="[813,889,1357,1381]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFC91FAB2FC9BFAEE" box="[813,889,1357,1381]" italics="true" pageId="23" pageNumber="24">Ufimia</emphasis>
</taxonomicName>
as subgenus of
<taxonomicName id="0691A3837F0EFFEFFB8BFAB2FABEFAEE" authority="Hinde, 1890" authorityName="Hinde" authorityYear="1890" box="[1079,1372,1357,1381]" class="Anthozoa" family="Plerophyllidae" genus="Plerophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFB8BFAB2FB2AFAEE" box="[1079,1224,1357,1381]" italics="true" pageId="23" pageNumber="24">Plerophyllum</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFB6FFAB2FABEFAEE" author="Hinde, G. J." box="[1235,1372,1357,1381]" pageId="23" pageNumber="24" pagination="194 - 204" refId="ref26525" refString="Hinde, G. J. (1890). Notes on the geology of Western Australia, 2: Corals and Polyzoa. Geological Magazine, New Series, Decade, 3 (7), 194 - 204." type="journal article" year="1890">Hinde, 1890</bibRefCitation>
</taxonomicName>
, only differing by the shortened cardinal septum in adult stages.
<bibRefCitation id="A500A5F17F0EFFEFFC3CFA73FBADFA2E" author="Fedorowski, J." box="[896,1103,1420,1445]" pageId="23" pageNumber="24" pagination="89 - 133" refId="ref25331" refString="Fedorowski, J. (1973). Rugose corals Polycoelaceae and Tachylasmatina subord. n. from Dalnia in the Holy Cross Mts. Acta Geologica Polonica, 23 (1), 89 - 133." type="journal article" year="1973">Fedorowski (1973)</bibRefCitation>
, like in the original designation of
<bibRefCitation id="A500A5F17F0EFFEFFC3EFA52FBBBFA4E" author="Stuckenberg, A. A." box="[898,1113,1452,1477]" pageId="23" pageNumber="24" refId="ref29373" refString="Stuckenberg, A. A. (1895). Korallen und Bryozoen der Steinkohlenablagerungen des Ural und des Timan. Memoires Du Comite Geologique, 10 (3), 244. in Russian and German." type="book" year="1895">Stuckenberg (1895)</bibRefCitation>
, as well as
<bibRefCitation id="A500A5F17F0EFFEFFB63FA52FAADFA4F" author="Hill, D." box="[1247,1359,1452,1477]" pageId="23" pageNumber="24" refId="ref26422" refString="Hill, D. (1956). Rugosa. In R. C. Moore (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata (pp. F 233 - F 324). New York." type="journal volume" year="1956">Hill (1956</bibRefCitation>
,
<bibRefCitation id="A500A5F17F0EFFEFFAE3FA52FA71FA4E" author="Hill, D." box="[1375,1427,1453,1477]" pageId="23" pageNumber="24" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">1981</bibRefCitation>
) and later authors again considered
<taxonomicName id="0691A3837F0EFFEFFB17FA32FB15FA6E" authorityName="Stuckenberg" authorityYear="1895" box="[1195,1271,1485,1509]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFB17FA32FB15FA6E" box="[1195,1271,1485,1509]" italics="true" pageId="23" pageNumber="24">Ufimia</emphasis>
</taxonomicName>
as valid on the genus level.
<taxonomicName id="0691A3837F0EFFEFFC05FA12FB19F98F" authority="Hudson, 1936" authorityName="Hudson" authorityYear="1936" box="[953,1275,1516,1541]" class="Anthozoa" family="Plerophyllidae" genus="Rhopalolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFC05FA12FBB7F98E" box="[953,1109,1517,1541]" italics="true" pageId="23" pageNumber="24">Rhopalolasma</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFBDCFA12FB19F98F" author="Hudson, R. G. S." box="[1120,1275,1516,1541]" pageId="23" pageNumber="24" pagination="1 - 12" refId="ref26566" refString="Hudson, R. G. S. (1936). On the Lower Carboniferous corals Rhopalolasma, gen nov and Cryptophyllum CARR. Proceedings of the Yorkshire Geological Society, 23 (2), 1 - 12." type="journal article" year="1936">Hudson, 1936</bibRefCitation>
</taxonomicName>
like the most probably misspelled
<emphasis id="F3E504127F0EFFEFFBA8F9F2FB4DF9AE" box="[1044,1199,1549,1573]" italics="true" pageId="23" pageNumber="24">Rhopalelasma</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFB05F9F2FA97F9AE" author="Lang, W. D. &amp; Smith, S. &amp; Thomas, H. D." box="[1209,1397,1549,1573]" pageId="23" pageNumber="24" pagination="1 - 231" refId="ref27892" refString="Lang, W. D., Smith, S., &amp; Thomas, H. D. (1940). Index of Palaeozoic coral genera. British Museum London, pp 1 - 231." type="book chapter" year="1940">Lang et al., 1940</bibRefCitation>
, as well as?
<taxonomicName id="0691A3837F0EFFEFFC22F9D2FC83F9EF" authority="Kullmann, 1966" authorityName="Kullmann" authorityYear="1966" class="Anthozoa" family="Plerophyllidae" genus="Plerophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="subGenus" subGenus="Meniscophylloides">
<emphasis id="F3E504127F0EFFEFFC22F9D2FBCDF9CE" box="[926,1071,1581,1605]" italics="true" pageId="23" pageNumber="24">Plerophyllum</emphasis>
(
<emphasis id="F3E504127F0EFFEFFBF6F9D2FAEDF9CE" box="[1098,1295,1581,1605]" italics="true" pageId="23" pageNumber="24">Meniscophylloides</emphasis>
)
<bibRefCitation id="A500A5F17F0EFFEFFA96F9D2FC83F9EF" author="Kullmann, J." pageId="23" pageNumber="24" pagination="443 - 466" refId="ref27663" refString="Kullmann, J. (1966). Goniatiten-Korallen-Vergesellschaftungen im Karbon des Kantabrischen Gebirges (Nordspanien). Neues Iahrbuch Fur Geologie Und Palaontologie, Abhandlungen, 125, 443 - 466." type="journal article" year="1966">Kullmann, 1966</bibRefCitation>
</taxonomicName>
, are younger synonyms of
<taxonomicName id="0691A3837F0EFFEFFB2AF9B2FB00F9EE" authorityName="Stuckenberg" authorityYear="1895" box="[1174,1250,1613,1637]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFB2AF9B2FB00F9EE" box="[1174,1250,1613,1637]" italics="true" pageId="23" pageNumber="24">Ufimia</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="A500A5F17F0EFFEFFA95F9B2FA74F9EE" author="Hill, D." box="[1321,1430,1613,1637]" pageId="23" pageNumber="24" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill, 1981</bibRefCitation>
;
<bibRefCitation id="A500A5F17F0EFFEFFC91F993FB9FF90E" author="Fedorowski, J. &amp; Bamber, E. W." box="[813,1149,1644,1669]" pageId="23" pageNumber="24" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
). The strong similarity of the genus with
<taxonomicName id="0691A3837F0EFFEFFC6FF973FB0CF92E" authority="Grabau, 1922" authorityName="Grabau" authorityYear="1922" box="[979,1262,1676,1701]" class="Anthozoa" family="Pentaphyllidae" genus="Tachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFC6FF973FBB7F92F" box="[979,1109,1676,1700]" italics="true" pageId="23" pageNumber="24">Tachylasma</emphasis>
<bibRefCitation id="A500A5F17F0EFFEFFBE0F973FB0CF92E" author="Grabau, A. W." box="[1116,1262,1676,1701]" pageId="23" pageNumber="24" pagination="1 - 76" refId="ref26116" refString="Grabau, A. W. (1922). Palaeozoic corals of China, Part 1, Tetraseptata. Palaeontologica Sinica, Series B, 2 (1), 1 - 76." type="journal article" year="1922">Grabau, 1922</bibRefCitation>
</taxonomicName>
was stressed by
<bibRefCitation id="A500A5F17F0EFFEFFC91F953FC1BF94E" author="Rodriguez, S." box="[813,1017,1708,1733]" pageId="23" pageNumber="24" refId="ref28568" refString="Rodriguez, S. (1984 b). Corales rugosos del Carbonifero del Este de Asturias. Doctoral thesis, Departamento de Paleontologia, Facultad de Ciencias Geologicas, Universidad Complutense de Madrid, 528 pp." type="book" year="1984">Rodríguez (1984b)</bibRefCitation>
. In the adult stage, the genera are virtually not distinguishable, but in juvenile stages, the septa in
<taxonomicName id="0691A3837F0EFFEFFCF4F912FC28F88E" authorityName="Grabau" authorityYear="1922" box="[840,970,1773,1797]" class="Anthozoa" family="Pentaphyllidae" genus="Tachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFCF4F912FC28F88E" box="[840,970,1773,1797]" italics="true" pageId="23" pageNumber="24">Tachylasma</emphasis>
</taxonomicName>
are not conjunct axially. Moreover, though in cases difficult to see,
<taxonomicName id="0691A3837F0EFFEFFBFAF8F2FB70F8AE" authorityName="Stuckenberg" authorityYear="1895" box="[1094,1170,1805,1829]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFBFAF8F2FB70F8AE" box="[1094,1170,1805,1829]" italics="true" pageId="23" pageNumber="24">Ufimia</emphasis>
</taxonomicName>
has zaphrentoid septal development in earlier ontogenetic stages, opposed to the pentaphyllid development in
<taxonomicName id="0691A3837F0EFFEFFB28F8B2FAF4F8EE" authorityName="Grabau" authorityYear="1922" box="[1172,1302,1869,1893]" class="Anthozoa" family="Pentaphyllidae" genus="Tachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F0EFFEFFB28F8B2FAF4F8EE" box="[1172,1302,1869,1893]" italics="true" pageId="23" pageNumber="24">Tachylasma</emphasis>
</taxonomicName>
(see discussion in
<bibRefCitation id="A500A5F17F0EFFEFFCC0F892FB20F80E" author="Fedorowski, J. &amp; Bamber, E. W." box="[892,1218,1901,1925]" pageId="23" pageNumber="24" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
).
</paragraph>
<caption id="95EE88887F01FFE0FF1BFB40FED3FAD3" ID-DOI="http://doi.org/10.5281/zenodo.12003332" ID-Zenodo-Dep="12003332" httpUri="https://zenodo.org/record/12003332/files/figure.png" pageId="24" pageNumber="25" startId="24.[167,198,1215,1235]" targetBox="[287,1297,238,1199]" targetPageId="24" targetType="figure">
<paragraph id="C12ED8007F01FFE0FF1BFB40FA8CFAB6" blockId="24.[167,1402,1215,1368]" pageId="24" pageNumber="25">
<emphasis id="F3E504127F01FFE0FF1BFB40FF1CFB58" bold="true" box="[167,254,1215,1235]" pageId="24" pageNumber="25">Fig. 13 A</emphasis>
<taxonomicName id="0691A3837F01FFE0FEBEFB40FE89FB58" box="[258,363,1215,1235]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="undefined-A">
<emphasis id="F3E504127F01FFE0FEBEFB40FEDBFB58" box="[258,313,1215,1235]" italics="true" pageId="24" pageNumber="25">Ufimia</emphasis>
sp.
<emphasis id="F3E504127F01FFE0FEE6FB40FE89FB58" bold="true" box="[346,363,1215,1235]" pageId="24" pageNumber="25">A</emphasis>
</taxonomicName>
<emphasis id="F3E504127F01FFE0FED7FB40FE91FB58" bold="true" box="[363,371,1215,1235]" pageId="24" pageNumber="25">1</emphasis>
External view of the specimen RAh (101), showing a straight conical corallite with eroded calice rim and apex. Positions of thin-sections indicated.
<emphasis id="F3E504127F01FFE0FE3BFB26FE26FB66" bold="true" box="[391,452,1241,1261]" pageId="24" pageNumber="25">A2A3</emphasis>
Two successive transverse thin-sections in the mature part of the corallite show the thick, rhopaloid major septa meet in the corallite centre. Scale bars:
<emphasis id="F3E504127F01FFE0FE4DFB0BFDE8FA83" bold="true" box="[497,522,1268,1288]" pageId="24" pageNumber="25">A1</emphasis>
: 5 mm;
<emphasis id="F3E504127F01FFE0FDF1FB0BFD84FA83" bold="true" box="[589,614,1268,1288]" pageId="24" pageNumber="25">A2</emphasis>
, A3: 2 mm.
<emphasis id="F3E504127F01FFE0FD74FB0BFD36FA83" bold="true" box="[712,724,1268,1288]" pageId="24" pageNumber="25">B</emphasis>
<taxonomicName id="0691A3837F01FFE0FD65FB0BFBFCFA83" authority="Kossovaya, 2001" authorityName="Kossovaya" authorityYear="2001" box="[729,1054,1268,1288]" class="Anthozoa" family="Bothrophyllidae" genus="Bothrophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="okense">
<emphasis id="F3E504127F01FFE0FD65FB0BFC72FA83" box="[729,912,1268,1288]" italics="true" pageId="24" pageNumber="25">Bothrophyllum okense</emphasis>
Kossovaya, 2001
</taxonomicName>
.
<emphasis id="F3E504127F01FFE0FB9AFB0BFBDFFA83" bold="true" box="[1062,1085,1268,1288]" pageId="24" pageNumber="25">B1</emphasis>
External view of the partly preserved corallite (RAh 37), with partly eroded wall. Positions of thin-sections indicated. Successive transverse thin-sections from the lower less mature part in
<emphasis id="F3E504127F01FFE0FF00FAD6FF36FAB6" bold="true" box="[188,212,1321,1341]" pageId="24" pageNumber="25">B2</emphasis>
up to the more mature section below the calice in
<emphasis id="F3E504127F01FFE0FD3AFAD6FD7CFAB6" bold="true" box="[646,670,1321,1341]" pageId="24" pageNumber="25">B4</emphasis>
, the transverse sections show the axial septum and a very narrow dissepimentarium.
</paragraph>
<paragraph id="C12ED8007F01FFE0FF1BFABBFED3FAD3" blockId="24.[167,1402,1215,1368]" box="[167,305,1348,1368]" pageId="24" pageNumber="25">Scale bars: 5mm</paragraph>
</caption>
<paragraph id="C12ED8007F01FFE0FF11FA63FD00FA58" blockId="24.[151,775,1435,1907]" pageId="24" pageNumber="25">
The diagnosis of the
<taxonomicName id="0691A3837F01FFE0FE20FA64FE0AFA38" authorityName="Stuckenberg" authorityYear="1895" box="[412,488,1435,1459]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F01FFE0FE20FA64FE0AFA38" box="[412,488,1435,1459]" italics="true" pageId="24" pageNumber="25">Ufimia</emphasis>
</taxonomicName>
was slightly emended by inclusion of septal microstructure (
<bibRefCitation id="A500A5F17F01FFE0FDA9FA44FD36FA58" author="Fedorowski, J." box="[533,724,1467,1491]" pageId="24" pageNumber="25" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F01FFE1FF2BFA24FDECFD08" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="distribution">
<paragraph id="C12ED8007F01FFE1FF2BFA24FDECFD08" blockId="24.[151,775,1435,1907]" lastBlockId="25.[151,775,235,643]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="F3E504127F01FFE0FF2BFA24FDA3FA78" bold="true" box="[151,577,1499,1523]" pageId="24" pageNumber="25">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F01FFE0FDF6FA24FD74FA78" authorityName="Stuckenberg" authorityYear="1895" box="[586,662,1499,1523]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F01FFE0FDF6FA24FD74FA78" box="[586,662,1499,1523]" italics="true" pageId="24" pageNumber="25">Ufimia</emphasis>
</taxonomicName>
is a complex, wide-spread genus—probably a group of intimately related, mostly homeomorphic genera, as indicated by stratigraphic gaps between occurrences. It is recorded from Lower Devonian (Upper Emsian) to Upper Permian strata throughout the Palaeotethys, adjoining epicontinental seas, and the Ural and Franklinian seaways (fide
<bibRefCitation id="A500A5F17F01FFE0FF6FF944FEA3F958" author="Hill, D." box="[211,321,1723,1747]" pageId="24" pageNumber="25" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill, 1981</bibRefCitation>
, and
<bibRefCitation id="A500A5F17F01FFE0FE3AF944FD3EF958" author="Fedorowski, J. &amp; Bamber, E. W." box="[390,732,1723,1747]" pageId="24" pageNumber="25" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
). It is absent from
<collectingCountry id="B98698907F01FFE0FE80F924FE6DF978" box="[316,399,1755,1779]" name="Canada" pageId="24" pageNumber="25">Canada</collectingCountry>
south of the Arctic realm and the conterminous
<collectingCountry id="B98698907F01FFE0FE8AF904FE8FF898" box="[310,365,1787,1811]" name="United States of America" pageId="24" pageNumber="25">USA</collectingCountry>
. (
<bibRefCitation id="A500A5F17F01FFE0FEC3F904FDDCF898" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." box="[383,574,1787,1811]" pageId="24" pageNumber="25" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
,
<bibRefCitation id="A500A5F17F01FFE0FDF0F904FD62F898" author="Wang, X. D. &amp; Yang, S. R. &amp; Yao, L. &amp; Sugiyama, T. &amp; Hu, K. Y." box="[588,640,1787,1811]" pageId="24" pageNumber="25" pagination="603 - 63" refId="ref29952" refString="Wang, X. D., Yang, S. R., Yao, L., Sugiyama, T., &amp; Hu, K. Y. (2022). Carboniferous biostratigraphy of rugose corals. In S. G. Lucas, J. W. Schneider, X. Wang, S. Nikolaeva (Eds), The Carboniferous timescale. Geological Society, London, Special Publications. 512 (2022). 603 - 63." type="journal article" year="2022">2022</bibRefCitation>
). A detailed review is out of scope from the present paper, but we present the distribution patterns from the Pennsylvanian to the Permian.
<bibRefCitation id="A500A5F17F01FFE0FEC2F8A4FDEFF8F8" author="FlUgel, H. W." box="[382,525,1883,1907]" pageId="24" pageNumber="25" pagination="657 - 688" refId="ref26042" refString="FlUgel, H. W. (1991). Rugosa aus dem Karbon der Ozbak-kuh-Gruppe Ost- Irans (Teil 1). Jahrbuch Der Geologischen Bundesanstalt (wien), 134, 657 - 688." type="journal article" year="1991">Flügel (1991)</bibRefCitation>
described a rich fauna consisting of several species known from lower Carboniferous and the new species
<taxonomicName id="0691A3837F01FFE0FB29FA43FA7EFA58" authority="Flugel, 1991" authorityName="Flugel" authorityYear="1991" box="[1173,1436,1467,1491]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="biforma">
<emphasis id="F3E504127F01FFE0FB29FA43FB4AFA58" box="[1173,1192,1468,1491]" italics="true" pageId="24" pageNumber="25">U</emphasis>
.
<emphasis id="F3E504127F01FFE0FB04FA44FAECFA58" box="[1208,1294,1467,1491]" italics="true" pageId="24" pageNumber="25">biforma</emphasis>
<bibRefCitation id="A500A5F17F01FFE0FAABFA44FA7EFA58" author="FlUgel, H. W." box="[1303,1436,1467,1491]" pageId="24" pageNumber="25" pagination="657 - 688" refId="ref26042" refString="FlUgel, H. W. (1991). Rugosa aus dem Karbon der Ozbak-kuh-Gruppe Ost- Irans (Teil 1). Jahrbuch Der Geologischen Bundesanstalt (wien), 134, 657 - 688." type="journal article" year="1991">Flügel, 1991</bibRefCitation>
</taxonomicName>
from Namurian (SerpukhovianBashkirian) strata of central-eastern
<collectingCountry id="B98698907F01FFE0FC64FA03FBEAF998" box="[984,1032,1532,1555]" name="Iran" pageId="24" pageNumber="25">Iran</collectingCountry>
. From the westernmost Palaeotethys
<taxonomicName id="0691A3837F01FFE0FC91F9E4FC9BF9B8" authorityName="Stuckenberg" authorityYear="1895" box="[813,889,1563,1587]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F01FFE0FC91F9E4FC9BF9B8" box="[813,889,1563,1587]" italics="true" pageId="24" pageNumber="25">Ufimia</emphasis>
</taxonomicName>
was recorded from Asturian strata (= Westphalian D, Kasimovian) of the Cantabrian Mountains, northern
<collectingCountry id="B98698907F01FFE0FCE5F9A4FC74F9F8" box="[857,918,1627,1651]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
(de Groot, 1963;
<bibRefCitation id="A500A5F17F01FFE0FBE4F9A4FAF5F9F8" author="Fedorowski, J." box="[1112,1303,1627,1651]" pageId="24" pageNumber="25" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
;
<bibRefCitation id="A500A5F17F01FFE0FA99F9A4FC8DF918" author="Rodriguez, S." pageId="24" pageNumber="25" refId="ref28568" refString="Rodriguez, S. (1984 b). Corales rugosos del Carbonifero del Este de Asturias. Doctoral thesis, Departamento de Paleontologia, Facultad de Ciencias Geologicas, Universidad Complutense de Madrid, 528 pp." type="book" year="1984">Rodríguez, 1984b</bibRefCitation>
;
<bibRefCitation id="A500A5F17F01FFE0FCC2F984FB35F918" author="Rodriguez, S. &amp; Kullmann, J." box="[894,1239,1659,1683]" pageId="24" pageNumber="25" pagination="19 - 40" refId="ref28690" refString="Rodriguez, S., &amp; Kullmann, J. (1990). Hornformige Einzelkorallen (Rugosa) aus spatoberkarbonischen Flachwasser-Ablagerungen des Kantabrischen Gebirges (Nordspanien). Palaeontographica Abteilung A, 19 - 40." type="book chapter" year="1990">Rodríguez and Kullmann, 1990</bibRefCitation>
;
<bibRefCitation id="A500A5F17F01FFE0FB5BF984FC83F938" author="Rodriguez, S. &amp; Coronado, I. &amp; Rodriguez-Castro, I." pageId="24" pageNumber="25" pagination="87 - 95" refId="ref28771" refString="Rodriguez, S., Coronado, I., &amp; Rodriguez-Castro, I. (2022). Corals from Asturian substage in Cantabrian Mountains: A review. Episodes, 45 (1), 87 - 95." type="journal article" year="2022">Rodríguez et al., 2022</bibRefCitation>
). The genus is known from the Middle and Upper Pennsylvanian of the Donets Basin (
<bibRefCitation id="A500A5F17F01FFE0FB78F944FA91F958" author="Fomichev, V." box="[1220,1395,1723,1747]" pageId="24" pageNumber="25" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
). It is an important faunal element of the Upper Artinskian strata from the central to subpolar Urals; also the
<typeStatus id="1E2A66A27F01FFE0FAD1F904FA7EF898" box="[1389,1436,1787,1811]" pageId="24" pageNumber="25">type</typeStatus>
species,
<taxonomicName id="0691A3837F01FFE0FC3BF8E4FAC2F8B8" authority="Stuckenberg, 1895" authorityName="Stuckenberg" authorityYear="1895" box="[903,1312,1819,1843]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="carbonaria">
<emphasis id="F3E504127F01FFE0FC3BF8E4FBB1F8B8" box="[903,1107,1819,1843]" italics="true" pageId="24" pageNumber="25">Ufimia carbonaria</emphasis>
<bibRefCitation id="A500A5F17F01FFE0FBE4F8E4FAC2F8B8" author="Stuckenberg, A. A." box="[1112,1312,1819,1843]" pageId="24" pageNumber="25" refId="ref29373" refString="Stuckenberg, A. A. (1895). Korallen und Bryozoen der Steinkohlenablagerungen des Ural und des Timan. Memoires Du Comite Geologique, 10 (3), 244. in Russian and German." type="book" year="1895">Stuckenberg, 1895</bibRefCitation>
</taxonomicName>
is from the Late Artinskian of the Urals (
<bibRefCitation id="A500A5F17F01FFE0FBDAF8C4FAB0F8D8" author="Kossovaya, O. L. &amp; Kuseva, E. A. &amp; Lukin, A. E. &amp; Zhuravlev, A. V." box="[1126,1362,1851,1875]" pageId="24" pageNumber="25" pagination="95 - 113" refId="ref27550" refString="Kossovaya, O. L., Kuseva, E. A., Lukin, A. E., &amp; Zhuravlev, A. V. (2001). Middle Artinskian (Early Permian) ecological event: a case study of the Urals and northern Timan. Proceedings of the Estonian Academy of Science. Geology, 50 (2), 95 - 113." type="journal article" year="2001">Kossovaya et al., 2001</bibRefCitation>
;
<bibRefCitation id="A500A5F17F01FFE0FAE3F8C4FC49F8F8" author="Kossovaya, O. L." pageId="24" pageNumber="25" pagination="383 - 405" refId="ref27441" refString="Kossovaya, O. L. (2007). Ecological aspects of upper Carboniferous - lower Permian ' Cyathaxonia Fauna' taxonomical diversity (the Urals). In B. Hubmann, &amp; W. E. Piller (Eds.), Fossil Corals and Sponges. Proceedings of the 9 th International Symposium on Fossil Cnidaria and Porifera. - Osterreichische Akademie der Wissenschaften, Schriftenreihe der Erdwissenschaftlichen Kommission 17, 383 - 405." type="journal article" year="2007">Kossovaya, 2007</bibRefCitation>
; see also
<bibRefCitation id="A500A5F17F01FFE0FBADF8A4FB61F8F8" author="Ilina, T. G." box="[1041,1155,1883,1907]" pageId="24" pageNumber="25" refId="ref26797" refString="Ilina, T. G. (1984). Historical evolution of corals (Suborder Polycoeliina). Transactions of Paleontological lnstitute of RAS 198, 184 pp. Moscow, Nauka (in Russian)." type="book" year="1984">Ilina, 1984</bibRefCitation>
who redescribed the
<typeStatus id="1E2A66A27F01FFE0FAD1F8A3FA7EF8FF" box="[1389,1436,1884,1908]" pageId="24" pageNumber="25">type</typeStatus>
species). In the Kungurian to Capitanian interval,
<taxonomicName id="0691A3837F01FFE0FAECF884FA7EF818" authorityName="Stuckenberg" authorityYear="1895" box="[1360,1436,1915,1939]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F01FFE0FAECF884FA7EF818" box="[1360,1436,1915,1939]" italics="true" pageId="24" pageNumber="25">Ufimia</emphasis>
</taxonomicName>
was a common faunal element in South
<collectingCountry id="B98698907F00FFE1FDEDFF14FD71FE88" box="[593,659,235,259]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
and Indochina, but persisted into the latest Permian; in
<collectingRegion id="035516E27F00FFE1FD77FEF3FEE2FEC8" country="China" name="Nei Mongol" pageId="25" pageNumber="26">Inner Mongolia</collectingRegion>
and North
<collectingCountry id="B98698907F00FFE1FE3AFED4FE2AFEC8" box="[390,456,299,323]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
block as well as in the peri-Gondwana terranes the genus also was widespread from the Asselian to Lopingian, appearing also in the Guadalupian of
<collectingCountry id="B98698907F00FFE1FE9EFE73FEBCFE28" box="[290,350,396,419]" name="Japan" pageId="25" pageNumber="26">Japan</collectingCountry>
(fide
<bibRefCitation id="A500A5F17F00FFE1FE1AFE73FD8DFE28" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." box="[422,623,395,420]" pageId="25" pageNumber="26" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
);
<bibRefCitation id="A500A5F17F00FFE1FD35FE74FE8FFE48" author="Fedorowski, J. &amp; Bamber, E. W." pageId="25" pageNumber="26" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber (2001)</bibRefCitation>
described
<taxonomicName id="0691A3837F00FFE1FE54FE54FDD6FE48" authorityName="Stuckenberg" authorityYear="1895" box="[488,564,427,451]" class="Anthozoa" family="Plerophyllidae" genus="Ufimia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F00FFE1FE54FE54FDD6FE48" box="[488,564,427,451]" italics="true" pageId="25" pageNumber="26">Ufimia</emphasis>
</taxonomicName>
from the Wordian (Middle Guadalupian) of the Sverdrup Basin (Canadian Arctic) and the Upper Permian of eastern
<collectingCountry id="B98698907F00FFE1FD32FE14FCE1FD88" box="[654,771,491,515]" name="Greenland" pageId="25" pageNumber="26">Greenland</collectingCountry>
, where it was first described by
<bibRefCitation id="A500A5F17F00FFE1FE55FDF4FD8CFDA8" author="FlUgel, H. W." box="[489,622,523,547]" pageId="25" pageNumber="26" pagination="1 - 57" refId="ref26012" refString="FlUgel, H. W. (1973). Rugose Korallen aus dem oberen Perm Ost-Gronlands. Verhandlungen Der Geologischen Bundes-Anstalt., 1, 1 - 57." type="journal article" year="1973">Flügel (1973</bibRefCitation>
: “
<taxonomicName id="0691A3837F00FFE1FD38FDF4FF20FDC8" authorityName="Carruthers" authorityYear="1919" class="Anthozoa" family="Polycoeliidae" genus="Cryptophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F00FFE1FD38FDF4FF20FDC8" italics="true" pageId="25" pageNumber="26">Cryptophyllum</emphasis>
</taxonomicName>
(
<taxonomicName id="0691A3837F00FFE1FF6EFDD4FEB6FDC8" authorityName="Grabau" authorityYear="1922" box="[210,340,555,579]" class="Anthozoa" family="Pentaphyllidae" genus="Tachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F00FFE1FF6EFDD4FEB6FDC8" box="[210,340,555,579]" italics="true" pageId="25" pageNumber="26">Tachylasma</emphasis>
</taxonomicName>
?) sp.?”). It is also known from the Kungurian (probably Guadalupian according to
<bibRefCitation id="A500A5F17F00FFE1FD3DFDB3FF29FD08" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." pageId="25" pageNumber="26" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al., 2017</bibRefCitation>
) in Spitsbergen (
<bibRefCitation id="A500A5F17F00FFE1FE3FFD94FDE2FD08" author="Ezaki, Y." box="[387,512,619,643]" pageId="25" pageNumber="26" pagination="381 - 388" refId="ref25246" refString="Ezaki, Y. (1997). Cold-water Permian Rugosa and their extinction in Spitsbergen. Boletin De La Real Sociedad Espanola De Histora Natural, Seccion Geologica, 92 (1 - 4), 381 - 388." type="journal article" year="1997">Ezaki, 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,183 @@
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<paragraph id="C12ED8007F11FFF0FF2BFB71FDA1FB2E" blockId="8.[151,776,1165,1926]" box="[151,579,1165,1190]" pageId="8" pageNumber="9">
<heading id="9A666F6C7F11FFF0FF2BFB71FDA1FB2E" box="[151,579,1165,1190]" fontSize="36" level="2" pageId="8" pageNumber="9" reason="3">
Genus
<taxonomicName id="0691A3837F11FFF0FF58FB72FDDEFB2D" authority="Ivanovski, 1967" authorityName="Ivanovski" authorityYear="1967" box="[228,572,1165,1190]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FF58FB72FE6DFB2E" bold="true" box="[228,399,1165,1189]" italics="true" pageId="8" pageNumber="9">Actinophrentis</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FE29FB71FDDEFB2D" author="Ivanovski, A. B." box="[405,572,1166,1190]" pageId="8" pageNumber="9" refId="ref26871" refString="Ivanovski, A. B. (1967). Studies on Early Carboniferous Rugosa. 92 pp., Nauka, Moscow (in Russian)." type="book" year="1967">Ivanovski, 1967</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFB51FF30FB6E" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="C12ED8007F11FFF0FF2BFB51FF30FB6E" blockId="8.[151,776,1165,1926]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFB51FEDDFB4D" bold="true" box="[151,319,1198,1222]" pageId="8" pageNumber="9">
<typeStatus id="1E2A66A27F11FFF0FF2BFB51FF32FB4D" box="[151,208,1198,1222]" pageId="8" pageNumber="9">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F11FFF0FEE9FB51FF29FB6D" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="donetziana">
<emphasis id="F3E504127F11FFF0FEE9FB51FD61FB4D" box="[341,643,1198,1222]" italics="true" pageId="8" pageNumber="9">Actinophrentis donetziana</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FD24FB51FF29FB6D" author="Fomichev, V." pageId="8" pageNumber="9" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFB11FD4BFA4D" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="C12ED8007F11FFF0FF2BFB11FD4BFA4D" blockId="8.[151,776,1165,1926]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFB11FEF3FA8D" bold="true" box="[151,273,1262,1286]" pageId="8" pageNumber="9">Diagnosis:</emphasis>
Non-dissepimented, small, solitary corals with pinnately arranged major septa up to calice floor, united axially by stereocolumn; cardinal septum shortened from early maturity; counter septum distinctly or moderately permanently elongated; cardinal fossula reaches or almost reaches corallite axis; minor septa underdeveloped (after
<bibRefCitation id="A500A5F17F11FFF0FE33FA51FDBBFA4D" author="Fedorowski, J." box="[399,601,1454,1478]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, p. 36).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFA31FB4FFCE8" pageId="8" pageNumber="9" type="discussion">
<paragraph id="C12ED8007F11FFF0FF2BFA31FB33FDC8" blockId="8.[151,776,1165,1926]" lastBlockId="8.[813,1437,235,867]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFA31FEE7FA6D" bold="true" box="[151,261,1486,1510]" pageId="8" pageNumber="9">Remarks:</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FEA8FA31FE53FA6D" author="Weyer, D." box="[276,433,1486,1510]" pageId="8" pageNumber="9" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
unambiguously synonymised
<taxonomicName id="0691A3837F11FFF0FF2BFA11FE13F98D" authority="Ivanovski, 1967" authorityName="Ivanovski" authorityYear="1967" box="[151,497,1518,1542]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FF2BFA11FED4F98D" box="[151,310,1518,1542]" italics="true" pageId="8" pageNumber="9">Actinophrentis</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FEFFFA11FE13F98D" author="Ivanovski, A. B." box="[323,497,1518,1542]" pageId="8" pageNumber="9" refId="ref26871" refString="Ivanovski, A. B. (1967). Studies on Early Carboniferous Rugosa. 92 pp., Nauka, Moscow (in Russian)." type="book" year="1967">Ivanovski, 1967</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="0691A3837F11FFF0FD87FA11FEE6F9AD" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FD87FA11FD22F98D" box="[571,704,1518,1542]" italics="true" pageId="8" pageNumber="9">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FD71FA11FEE6F9AD" author="Hudson, R. G. S." pageId="8" pageNumber="9" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="A500A5F17F11FFF0FEAAF9F1FE73F9AD" author="Hill, D." box="[278,401,1550,1574]" pageId="8" pageNumber="9" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
accepted
<taxonomicName id="0691A3837F11FFF0FDB4F9F1FD45F9AD" authorityName="Ivanovski" authorityYear="1967" box="[520,679,1550,1574]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FDB4F9F1FD45F9AD" box="[520,679,1550,1574]" italics="true" pageId="8" pageNumber="9">Actinophrentis</emphasis>
</taxonomicName>
, though she did not rule out synonymy with the later genus.
<bibRefCitation id="A500A5F17F11FFF0FF2BF9B1FE8FF9ED" author="Fedorowski, J." box="[151,365,1614,1638]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
stated the validity of the genus, especially based on the development of cardinal and counter septum (see diagnosis). He also discussed questionable synonymies with other genera, which include?p.p.
<taxonomicName id="0691A3837F11FFF0FD61F951FE51F96D" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Metriophyllidae" genus="Stereolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FD61F951FF19F96D" italics="true" pageId="8" pageNumber="9">Stereolasma</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FEBAF931FE51F96D" author="Fomichev, V." box="[262,435,1742,1766]" pageId="8" pageNumber="9" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
non
<bibRefCitation id="A500A5F17F11FFF0FE4EF931FD77F96D" author="Simpson, G. B." box="[498,661,1742,1766]" pageId="8" pageNumber="9" pagination="199 - 222" refId="ref29194" refString="Simpson, G. B. (1900). Preliminary descriptions of new genera of Palaeozoic rugose corals. New York State Museum, Bulletin, 8 (39), 199 - 222." type="journal article" year="1900">Simpson, 1900</bibRefCitation>
and 1962?
<taxonomicName id="0691A3837F11FFF0FF1DF911FE27F88D" authority="Easton, 1962" authorityName="Easton" authorityYear="1962" box="[161,453,1774,1798]" class="Anthozoa" family="Hapsiphyllidae" genus="Longiclava" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FF1DF911FEF5F88D" box="[161,279,1774,1798]" italics="true" pageId="8" pageNumber="9">Longiclava</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FE90F911FE27F88D" author="Easton, W. H." box="[300,453,1774,1798]" pageId="8" pageNumber="9" refId="ref24743" refString="Easton, W. H. (1962). Carboniferous formations and faunas of central Montana. U. S. Geol. Surv. Prof. Pap., Washington, 348, 126 pp." type="book" year="1962">Easton, 1962</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="0691A3837F11FFF0FE5CF911FEB5F8AD" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="columnare">
<emphasis id="F3E504127F11FFF0FE5CF911FCE5F88D" box="[480,775,1774,1798]" italics="true" pageId="8" pageNumber="9">Actinophrentis columnare</emphasis>
Fedorowski, 1987
</taxonomicName>
from the lower Wolfcampian of Texas is known in time-equivalent strata from the northern Timan (
<bibRefCitation id="A500A5F17F11FFF0FF4CF8B1FE45F8ED" author="Kossovaya, O. L." box="[240,423,1870,1894]" pageId="8" pageNumber="9" pagination="71 - 82" refId="ref27287" refString="Kossovaya, O. L. (1996). Correlation of uppermost Carboniferous and Lower Permian Rugose Coral Zones from the Urals to Western North America. Palaios, 11 (1), 71 - 82." type="journal article" year="1996">Kossovaya, 1996</bibRefCitation>
); unfortunately, its presence in the Urals and the Russian Platform cannot be deduced from this publication. However, the
<typeStatus id="1E2A66A27F11FFF0FB74FF13FB15FE8F" box="[1224,1271,236,260]" pageId="8" pageNumber="9">type</typeStatus>
species of the genus,
<taxonomicName id="0691A3837F11FFF0FCC6FEF3FB27FEA8" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[890,1221,267,291]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="donetziana">
<emphasis id="F3E504127F11FFF0FCC6FEF3FBF1FEA8" box="[890,1043,267,291]" italics="true" pageId="8" pageNumber="9">A. donetziana</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FBA7FEF4FB27FEA8" author="Fomichev, V." box="[1051,1221,267,291]" pageId="8" pageNumber="9" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
is from the Donets Basin,
<collectingCountry id="B98698907F11FFF0FCC7FED4FC34FEC8" box="[891,982,299,323]" name="Ukraine" pageId="8" pageNumber="9">Ukraine</collectingCountry>
.
<specimenCount id="D79713897F11FFF0FC5AFED4FB68FEC8" box="[998,1162,299,323]" count="1" pageId="8" pageNumber="9" type="generic">One specimen</specimenCount>
described by de Groot (1963) to belong to
<taxonomicName id="0691A3837F11FFF0FBB6FEB4FC83FE08" authority="de Groot, 1963" authorityName="de Groot" authorityYear="1963" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="paralleloides">
<emphasis id="F3E504127F11FFF0FBB6FEB4FACBFEE8" box="[1034,1321,331,355]" italics="true" pageId="8" pageNumber="9">Zaphrentites paralleloides</emphasis>
de Groot, 1963
</taxonomicName>
from the Sierra Corisa Limestone (Westphalian D = upper Moscovian) of northern Palencia (Cantabrian Mountains,
<collectingCountry id="B98698907F11FFF0FC0AFE54FC16FE48" box="[950,1012,427,451]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
) was included by
<bibRefCitation id="A500A5F17F11FFF0FB71FE54FA7FFE48" author="Fedorowski, J." box="[1229,1437,427,451]" pageId="8" pageNumber="9" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski (2004)</bibRefCitation>
as
<taxonomicName id="0691A3837F11FFF0FCF7FE34FBF2FE68" box="[843,1040,459,483]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F11FFF0FCF7FE34FC08FE68" box="[843,1002,459,483]" italics="true" pageId="8" pageNumber="9">Actinophrentis</emphasis>
sp.
</taxonomicName>
In addition, Rodríguez et al., (2022,
<tableCitation id="8C13EDBB7F11FFF0FC91FE14FC9FFD88" box="[813,893,491,515]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="8" pageNumber="9">Table 1</tableCitation>
) accepted the presence of the genus in the Asturian Substage (upper Moscovian) of the Cantabrian Mountains without further discussion.
</paragraph>
<paragraph id="C12ED8007F11FFF0FC91FDB4FB4FFCE8" blockId="8.[813,1437,235,867]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91FDB4FB05FDE8" bold="true" box="[813,1255,587,611]" pageId="8" pageNumber="9">Geographic and stratigraphic range:</emphasis>
Upper Moscovian of the Donets Basin (
<bibRefCitation id="A500A5F17F11FFF0FBD5FD94FAF9FD08" author="Fomichev, V." box="[1129,1307,619,643]" pageId="8" pageNumber="9" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
;
<bibRefCitation id="A500A5F17F11FFF0FA93FD94FC83FD28" author="Ivanovski, A. B." pageId="8" pageNumber="9" refId="ref26871" refString="Ivanovski, A. B. (1967). Studies on Early Carboniferous Rugosa. 92 pp., Nauka, Moscow (in Russian)." type="book" year="1967">Ivanovski, 1967</bibRefCitation>
) and the Cantabrian Mountains (northern Palencia,
<collectingCountry id="B98698907F11FFF0FCDFFD54FC43FD48" box="[867,929,683,707]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
) (
<bibRefCitation id="A500A5F17F11FFF0FC7CFD54FB68FD48" author="Fedorowski, J." box="[960,1162,683,707]" pageId="8" pageNumber="9" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
); Upper Pennsylvanian (Kasimovian) of
<collectingCountry id="B98698907F11FFF0FC5EFD33FBC3FD6F" box="[994,1057,716,740]" name="Egypt" pageId="8" pageNumber="9">Egypt</collectingCountry>
(
<bibRefCitation id="A500A5F17F11FFF0FB93FD34FADFFD68" author="Kora, M. &amp; Mansour, Y." box="[1071,1341,715,739]" pageId="8" pageNumber="9" pagination="597 - 614" refId="ref27107" refString="Kora, M., &amp; Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour, 1991</bibRefCitation>
and this paper); lowermost Permian (Asselian) of N Timan Ridge (
<bibRefCitation id="A500A5F17F11FFF0FC88FCF4FC0DFCA8" author="Kossovaya, O. L." box="[820,1007,779,803]" pageId="8" pageNumber="9" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
); lowermost Permian (lower part of Wolfcampian Series = Nealian Stage) of Glass Mountains, western Texas (
<bibRefCitation id="A500A5F17F11FFF0FC6FFCB4FB7DFCE8" author="Fedorowski, J." box="[979,1183,843,867]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
).
</paragraph>
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@ -0,0 +1,140 @@
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Subfamily
<taxonomicName id="0691A3837F11FFF0FEB6FDB3FE4BFDEF" authority="Ilina, 1970" box="[266,425,588,612]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="8" pageNumber="9" phylum="Cnidaria" rank="subFamily" subFamily="Antiphyllinae">
<emphasis id="F3E504127F11FFF0FEB6FDB3FE4BFDEF" bold="true" box="[266,425,588,612]" pageId="8" pageNumber="9">Antiphyllinae</emphasis>
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<bibRefCitation id="A500A5F17F11FFF0FE13FDB2FDC3FDEE" author="Ilina, T. G." box="[431,545,589,613]" pageId="8" pageNumber="9" pagination="146 - 151" refId="ref26741" refString="Ilina, T. G. (1970). Some new Permian Rugosa from the South-Eastern Pamir. 146 - 151. In G. G. Astrova &amp; I. I. Chudinova (Eds.), New species of the Palaeozoic bryozoans and corals (p. 178). Nauka." type="journal article" year="1970">Ilina, 1970</bibRefCitation>
.
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</paragraph>
<paragraph id="C12ED8007F11FFF0FF2BFD92FE5CFD0F" blockId="8.[151,775,588,1093]" box="[151,446,621,645]" pageId="8" pageNumber="9">
emend.
<bibRefCitation id="A500A5F17F11FFF0FF52FD92FE55FD0E" author="Fedorowski, J." box="[238,439,621,645]" pageId="8" pageNumber="9" pagination="1 - 33" refId="ref25787" refString="Fedorowski, J. (2012 a). On three rugose coral genera from Serpukhovian strata in the Upper Silesian Coal Basin, Poland. Acta Geologica Polonica, 62 (1), 1 - 33." type="journal article" year="2012">Fedorowski, 2012a</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFD72FD61FD2F" box="[151,643,652,677]" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="C12ED8007F11FFF0FF2BFD72FD61FD2F" blockId="8.[151,775,588,1093]" box="[151,643,652,677]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFD72FEC3FD2E" bold="true" box="[151,289,653,677]" pageId="8" pageNumber="9">
<typeStatus id="1E2A66A27F11FFF0FF2BFD72FF32FD2E" box="[151,208,653,677]" pageId="8" pageNumber="9">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="0691A3837F11FFF0FE9BFD73FD9FFD2F" authority="Schindewolf, 1952" authorityName="Schindewolf" authorityYear="1952" box="[295,637,652,676]" class="Anthozoa" family="Metriophyllidae" genus="Antiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FE9BFD73FE52FD2F" box="[295,432,652,676]" italics="true" pageId="8" pageNumber="9">Antiphyllum</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FE0AFD73FD9FFD2F" author="Schindewolf, O. H." box="[438,637,652,676]" pageId="8" pageNumber="9" pagination="1 - 85" refId="ref29071" refString="Schindewolf, O. H. (1952). Korallen aus dem Oberkarbon (Namur) des oberschlesischen Steinkohlen-Beckens. Akademie der Wissenschaften und der Literatur, Abhandlungen der mathematischnaturwissenschaftlichen Klasse 4, 1 - 85." type="journal article" year="1952">Schindewolf, 1952</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFD53FE30FD4F" box="[151,466,684,709]" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="C12ED8007F11FFF0FF2BFD53FE30FD4F" blockId="8.[151,775,588,1093]" box="[151,466,684,709]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFD53FEF3FD4F" bold="true" box="[151,273,684,708]" pageId="8" pageNumber="9">Diagnosis:</emphasis>
As for the family.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FF2BFD32FE90FBCE" pageId="8" pageNumber="9" type="discussion">
<paragraph id="C12ED8007F11FFF0FF2BFD32FE10FC2E" blockId="8.[151,775,588,1093]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FF2BFD32FEE7FD6E" bold="true" box="[151,261,717,741]" pageId="8" pageNumber="9">Remarks:</emphasis>
The most important diagnostic characters of genera of the subfamily recognised in our material are displayed in
<tableCitation id="8C13EDBB7F11FFF0FE9BFCF3FE9BFCAE" box="[295,377,780,805]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="8" pageNumber="9">Table 1</tableCitation>
. For comparison with morphologically close
<taxonomicName id="0691A3837F11FFF0FEA4FCD2FDA8FCCF" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[280,586,812,837]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FEA4FCD2FE76FCCE" box="[280,404,813,837]" italics="true" pageId="8" pageNumber="9">Lytvolasma</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FE1DFCD2FDA8FCCF" author="Soshkina, E. D." box="[417,586,812,837]" pageId="8" pageNumber="9" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
also
<taxonomicName id="0691A3837F11FFF0FD32FCD2FE84FCEE" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FD32FCD2FF20FCEE" italics="true" pageId="8" pageNumber="9">Bradyphyllum</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FF73FCB2FE84FCEE" author="Grabau, A. W." box="[207,358,845,869]" pageId="8" pageNumber="9" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
is listed. Comparisons with further
<taxonomicName id="0691A3837F11FFF0FF2BFC92FEC9FC0E" box="[151,299,877,901]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="35" phylum="Cnidaria" rank="subFamily" subFamily="Antiphyllinae">Antiphyllinae</taxonomicName>
were listed by
<bibRefCitation id="A500A5F17F11FFF0FE71FC92FD40FC0E" author="Fedorowski, J." box="[461,674,877,901]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
and supplemented by Chwieduk (2013).
</paragraph>
<paragraph id="C12ED8007F11FFF0FF2BFC52FE90FBCE" blockId="8.[151,775,588,1093]" pageId="8" pageNumber="9">
The synonymy of the
<typeStatus id="1E2A66A27F11FFF0FE11FC52FE3EFC4E" box="[429,476,941,965]" pageId="8" pageNumber="9">type</typeStatus>
genus of the subfamily,
<taxonomicName id="0691A3837F11FFF0FF2BFC32FE12FC6F" authority="Schindewolf, 1952" authorityName="Schindewolf" authorityYear="1952" box="[151,496,972,997]" class="Anthozoa" family="Metriophyllidae" genus="Antiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FF2BFC32FEC2FC6E" box="[151,288,973,997]" italics="true" pageId="8" pageNumber="9">Antiphyllum</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FE94FC32FE12FC6F" author="Schindewolf, O. H." box="[296,496,972,997]" pageId="8" pageNumber="9" pagination="1 - 85" refId="ref29071" refString="Schindewolf, O. H. (1952). Korallen aus dem Oberkarbon (Namur) des oberschlesischen Steinkohlen-Beckens. Akademie der Wissenschaften und der Literatur, Abhandlungen der mathematischnaturwissenschaftlichen Klasse 4, 1 - 85." type="journal article" year="1952">Schindewolf, 1952</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="0691A3837F11FFF0FD8DFC32FEE3FB8E" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" class="Anthozoa" family="Antiphyllidae" genus="Claviphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FD8DFC32FD27FC6E" box="[561,709,973,997]" italics="true" pageId="8" pageNumber="9">Claviphyllum</emphasis>
<bibRefCitation id="A500A5F17F11FFF0FD71FC32FEE3FB8E" author="Hudson, R. G. S." pageId="8" pageNumber="9" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
, stressed by
<bibRefCitation id="A500A5F17F11FFF0FE29FC12FDC3FB8E" author="Weyer, D." box="[405,545,1005,1029]" pageId="8" pageNumber="9" pagination="345 - 365" refId="ref30333" refString="Weyer, D. (1974). Das Rugosa-Genus Antiphyllum Schindewolf, 1952 (Unternamur, Oberschlesisches Steinkohlenbecken). Casopis pro Mineralogii a Geologii, 19 (4), 345 - 365." type="journal article" year="1974">Weyer (1974</bibRefCitation>
,
<bibRefCitation id="A500A5F17F11FFF0FD8DFC13FD87FB8F" author="Weyer, D." box="[561,613,1004,1028]" pageId="8" pageNumber="9" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">1975</bibRefCitation>
) and followed by
<bibRefCitation id="A500A5F17F11FFF0FF04FBF2FE6DFBAE" author="Fedorowski, J." box="[184,399,1037,1061]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
was not further recommended by
<bibRefCitation id="A500A5F17F11FFF0FF2BFBD2FE8CFBCE" author="Fedorowski, J." box="[151,366,1069,1093]" pageId="8" pageNumber="9" pagination="1 - 33" refId="ref25787" refString="Fedorowski, J. (2012 a). On three rugose coral genera from Serpukhovian strata in the Upper Silesian Coal Basin, Poland. Acta Geologica Polonica, 62 (1), 1 - 33." type="journal article" year="2012">Fedorowski (2012a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,275 @@
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<subSubSection id="898B8B8B7F11FFF0FC91FC53FB5AFC4F" box="[813,1208,940,965]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="C12ED8007F11FFF0FC91FC53FB5AFC4F" blockId="8.[813,1208,940,965]" box="[813,1208,940,965]" pageId="8" pageNumber="9">
<heading id="9A666F6C7F11FFF0FC91FC53FB5AFC4F" bold="true" box="[813,1208,940,965]" fontSize="10" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="0691A3837F11FFF0FC91FC53FB94FC4F" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[813,1142,940,964]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="crassithecata" status="sp. nov.">Actinophrentis crassithecata</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F11FFF0FBC0FC52FB5AFC4F" box="[1148,1208,941,965]" pageId="8" pageNumber="9" rank="species">
<emphasis id="F3E504127F11FFF0FBC0FC52FB50FC4E" bold="true" box="[1148,1202,941,965]" pageId="8" pageNumber="9">n. sp</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FC91FC33FC5CFBCF" pageId="8" pageNumber="9" type="description">
<paragraph id="C12ED8007F11FFF0FC91FC33FBFFFC6E" blockId="8.[813,1437,972,1925]" box="[813,1053,972,997]" pageId="8" pageNumber="9">
(
<figureCitation id="59AAC4857F11FFF0FC88FC33FC64FC6F" box="[820,902,972,996]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="8" pageNumber="9">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F11FFF0FC3AFC33FC56FC6F" box="[902,948,972,996]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="8" pageNumber="9">1A</figureCitation>
<figureCitation id="59AAC4857F11FFF0FC08FC33FC23FC6F" box="[948,961,972,996]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="8" pageNumber="9">5</figureCitation>
, B
<figureCitation id="59AAC4857F11FFF0FC61FC32FBE5FC6E" box="[989,1031,973,997]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="8" pageNumber="9">1B</figureCitation>
<figureCitation id="59AAC4857F11FFF0FBBBFC32FBF7FC6E" box="[1031,1045,973,997]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="8" pageNumber="9">2</figureCitation>
)
</paragraph>
<paragraph id="C12ED8007F11FFF0FC91FC13FC5CFBCF" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91FC13FBDCFB8E" bold="true" box="[813,1086,1004,1029]" pageId="8" pageNumber="9">Derivation of the name:</emphasis>
From the Latin crassus—means thick, theca—F cover—after the great thickness of the external wall.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FC91FBB2FA7AF92E" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="C12ED8007F11FFF0FC91FBB2FAEBFB0F" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<materialsCitation id="71F9D25D7F11FFF0FC91FBB2FAEBFB0F" pageId="8" pageNumber="9" specimenCode="RAh 57" specimenCount="1" typeStatus="holotype">
<emphasis id="F3E504127F11FFF0FC91FBB2FC7DFBEE" bold="true" box="[813,927,1101,1125]" pageId="8" pageNumber="9">
<typeStatus id="1E2A66A27F11FFF0FC91FBB2FC79FBEE" box="[813,923,1101,1125]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="D79713897F11FFF0FC1AFBB3FBF3FBEF" box="[934,1041,1100,1124]" pageId="8" pageNumber="9" type="generic">Specimen</specimenCount>
<specimenCode id="9137707B7F11FFF0FBA5FBB3FB8CFBEF" box="[1049,1134,1100,1124]" pageId="8" pageNumber="9">RAh 57</specimenCode>
, illustrated in
<figureCitation id="59AAC4857F11FFF0FAB7FBB3FABEFBEE" box="[1291,1372,1100,1125]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="8" pageNumber="9">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F11FFF0FAE1FBB2FA68FBEE" box="[1373,1418,1101,1125]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="8" pageNumber="9">1A</figureCitation>
<figureCitation id="59AAC4857F11FFF0FA36FBB2FA7AFBEE" box="[1418,1432,1101,1125]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
. Three transverse thin-sections are available.
</materialsCitation>
</paragraph>
<paragraph id="C12ED8007F11FFF0FC91FB73FAFCFA6E" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91FB73FC75FB2F" bold="true" box="[813,919,1164,1188]" pageId="8" pageNumber="9">Material:</emphasis>
In addition to the
<typeStatus id="1E2A66A27F11FFF0FBDBFB73FB2BFB2F" box="[1127,1225,1164,1188]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, two moderately to badly preserved corallites (RAh 11 and RAh 42) are
<typeStatus id="1E2A66A27F11FFF0FADFFB52FC8BFB6E" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
, with 11 transverse thin-sections. Although looking good preserved in their external form, keeping a good preserved outer epitheca with septal furrows and concentric growth lines (
<figureCitation id="59AAC4857F11FFF0FBAAFAD2FB85FACE" box="[1046,1127,1325,1349]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="8" pageNumber="9">Fig. 6B</figureCitation>
<figureCitation id="59AAC4857F11FFF0FBDBFAD2FB93FACE" box="[1127,1137,1325,1349]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="8" pageNumber="9">1</figureCitation>
, B
<figureCitation id="59AAC4857F11FFF0FB32FAD2FB7EFACE" box="[1166,1180,1325,1349]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="8" pageNumber="9">2</figureCitation>
), serial thin-sections of the specimens (RAh11, 42) are not suitable for illustrations as they suffer from either compaction or ferrugination (e.g.,
<figureCitation id="59AAC4857F11FFF0FC25FA72FC04FA2E" box="[921,998,1421,1445]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="8" pageNumber="9">Fig. 4E</figureCitation>
), The proximal ends of the corallites are largely eroded. They are collected from the basal shales of the lower member of the Aheimer Formation.
</paragraph>
<paragraph id="C12ED8007F11FFF0FC91FA12FC91F9EE" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91FA12FC2EF98F" bold="true" box="[813,972,1516,1541]" pageId="8" pageNumber="9">Type locality:</emphasis>
<materialsCitation id="71F9D25D7F11FFF0FC68FA12FC8DF9EE" country="Egypt" county="Gulf" latitude="29.4725" location="Northern Galala" longLatPrecision="20" longitude="32.460556" municipality="northern Eastern Desert" pageId="8" pageNumber="9" specimenCount="1" stateProvince="As Suways">
Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
<emphasis id="F3E504127F11FFF0FC61F9F4FC0AF9AF" box="[989,1000,1547,1572]" italics="true" pageId="8" pageNumber="9"></emphasis>
21
<emphasis id="F3E504127F11FFF0FBB5F9F4FBFBF9AF" box="[1033,1049,1547,1572]" italics="true" pageId="8" pageNumber="9"></emphasis>
N and Long. 32° 27
<emphasis id="F3E504127F11FFF0FB4CF9F4FB19F9AF" box="[1264,1275,1547,1572]" italics="true" pageId="8" pageNumber="9"></emphasis>
38
<emphasis id="F3E504127F11FFF0FAA7F9F4FAC9F9AF" box="[1307,1323,1547,1572]" italics="true" pageId="8" pageNumber="9"></emphasis>
E), northern Eastern Desert, western side of the Gulf of Suez, Egypt
</materialsCitation>
.
</paragraph>
<paragraph id="C12ED8007F11FFF0FC91F992FA7AF92E" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91F992FC32F90F" bold="true" box="[813,976,1644,1669]" pageId="8" pageNumber="9">Type horizon:</emphasis>
Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF0FC91F953FB19F8CE" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="C12ED8007F11FFF0FC91F953FB19F8CE" blockId="8.[813,1437,972,1925]" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91F953FC44F94F" bold="true" box="[813,934,1708,1732]" pageId="8" pageNumber="9">Diagnosis:</emphasis>
<taxonomicName id="0691A3837F11FFF0FC10F952FBA9F94E" authorityName="Ivanovski" authorityYear="1967" box="[940,1099,1709,1733]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F11FFF0FC10F952FBA9F94E" box="[940,1099,1709,1733]" italics="true" pageId="8" pageNumber="9">Actinophrentis</emphasis>
</taxonomicName>
with thick external wall (about 1.5 mm thick), having maximum n:d ratio at calice floor 24:11. Major septa are pinnately arranged.
<taxonomicName id="0691A3837F11FFF0FAB4F913FA84F88F" box="[1288,1382,1772,1796]" genus="Cardinal" pageId="8" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum shortened with maturity. Counter septum elongated from the immature stage up to calice floor.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F11FFF2FC91F8B3FF34FA48" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="C12ED8007F11FFF1FC91F8B3FAAAF808" blockId="8.[813,1437,972,1925]" lastBlockId="9.[813,1436,1835,1923]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="F3E504127F11FFF0FC91F8B3FC5FF8EF" bold="true" box="[813,957,1868,1892]" pageId="8" pageNumber="9">Description:</emphasis>
External characters: Corallites only faintly conical (see minimal increasing n:d ratios below) and slightly curved (
<figureCitation id="59AAC4857F10FFF1FEF3F8D4FE45F8C8" box="[335,423,1835,1859]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F10FFF1FE1BF8D4FE1CF8C8" box="[423,510,1835,1859]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="9" pageNumber="10">1, B1B</figureCitation>
<figureCitation id="59AAC4857F10FFF1FE43F8D4FDEFF8C8" box="[511,525,1835,1859]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="9" pageNumber="10">2</figureCitation>
). The preserved parts are 22.5 cm long. Calices are deep. If preserved, they are compressed due to compaction. Apexes are not preserved. The thick external wall (1.5 mm-thick) bears distinct furrows of major and minor septa. The latter could only be detected inside the calyx (
<figureCitation id="59AAC4857F10FFF1FB62F894FAD0F808" box="[1246,1330,1899,1923]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F10FFF1FA8DF894FAD9F808" box="[1329,1339,1899,1923]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="9" pageNumber="10">1</figureCitation>
).
</paragraph>
<caption id="95EE88887F10FFF1FF1BFA0FFB15F951" ID-DOI="http://doi.org/10.5281/zenodo.12003244" ID-Zenodo-Dep="12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="9" pageNumber="10" startId="9.[167,198,1520,1540]" targetBox="[287,1299,246,1507]" targetPageId="9" targetType="figure">
<paragraph id="C12ED8007F10FFF1FF1BFA0FFB15F951" blockId="9.[167,1420,1520,1754]" pageId="9" pageNumber="10">
<emphasis id="F3E504127F10FFF1FF1BFA0FFF10F98F" bold="true" box="[167,242,1520,1540]" pageId="9" pageNumber="10">Fig. 6 A</emphasis>
<taxonomicName id="0691A3837F10FFF1FF4BFA0FFE3EF98F" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[247,476,1520,1540]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Cnidaria" rank="species" species="crassithecata" status="sp. nov.">
<emphasis id="F3E504127F10FFF1FF4BFA0FFE3EF98F" box="[247,476,1520,1540]" italics="true" pageId="9" pageNumber="10">Actinophrentis crassithecata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F10FFF1FE5DFA0FFDEDF98F" box="[481,527,1520,1540]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
: (Holotype specimen RAh (57).
<emphasis id="F3E504127F10FFF1FCA9FA0FFCCDF98F" bold="true" box="[789,815,1520,1540]" pageId="9" pageNumber="10">A1</emphasis>
External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated.
<emphasis id="F3E504127F10FFF1FC3CF9F4FC7BF994" bold="true" box="[896,921,1547,1567]" pageId="9" pageNumber="10">A2</emphasis>
Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum.
<emphasis id="F3E504127F10FFF1FC0FF9D9FC2EF9B1" bold="true" box="[947,972,1574,1594]" pageId="9" pageNumber="10">A3</emphasis>
Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks.
<emphasis id="F3E504127F10FFF1FC2FF9BFFC4EF9DF" bold="true" box="[915,940,1600,1620]" pageId="9" pageNumber="10">A4</emphasis>
Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.
<emphasis id="F3E504127F10FFF1FD86F9A4FDB1F9E4" bold="true" box="[570,595,1627,1647]" pageId="9" pageNumber="10">A5</emphasis>
Details of the external wall of
<emphasis id="F3E504127F10FFF1FCEFF9A4FC89F9E4" bold="true" box="[851,875,1627,1647]" pageId="9" pageNumber="10">A3</emphasis>
.
<emphasis id="F3E504127F10FFF1FCC8F9A4FC4FF9E4" bold="true" box="[884,941,1627,1647]" pageId="9" pageNumber="10">B1, B2</emphasis>
External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall.
<emphasis id="F3E504127F10FFF1FB8AF989FBA1F901" bold="true" box="[1078,1091,1654,1674]" pageId="9" pageNumber="10">C</emphasis>
<taxonomicName id="0691A3837F10FFF1FBFBF989FEA2F92F" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F10FFF1FBFBF989FB46F901" box="[1095,1188,1654,1674]" italics="true" pageId="9" pageNumber="10">Lytvolasma</emphasis>
cf.
<emphasis id="F3E504127F10FFF1FB7CF989FAFAF901" box="[1216,1304,1654,1674]" italics="true" pageId="9" pageNumber="10">canadense</emphasis>
<bibRefCitation id="A500A5F17F10FFF1FAA1F989FEA2F92F" author="Fedorowski, J. &amp; Bamber, E. W." pageId="9" pageNumber="10" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber,2001</bibRefCitation>
</taxonomicName>
, specimen RAh (66).
<emphasis id="F3E504127F10FFF1FE4CF96FFDC8F92F" bold="true" box="[496,554,1680,1700]" pageId="9" pageNumber="10">C1, C2</emphasis>
Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end.
<emphasis id="F3E504127F10FFF1FDC4F954FD72F934" bold="true" box="[632,656,1707,1727]" pageId="9" pageNumber="10">C3</emphasis>
Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except
<emphasis id="F3E504127F10FFF1FF59F939FEC0F951" bold="true" box="[229,290,1734,1754]" pageId="9" pageNumber="10">A2A5</emphasis>
: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
</paragraph>
</caption>
<paragraph id="C12ED8007F13FFF2FF11FF14FF34FA48" blockId="10.[151,776,235,1923]" pageId="10" pageNumber="11">
Internal characters (transverse sections): the ontogenetically earliest preserved part has a n:d ratio of 22:
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(
<figureCitation id="59AAC4857F13FFF2FEB4FED4FEB9FEC8" box="[264,347,299,323]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FEE7FED4FE8BFEC8" box="[347,361,299,323]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="10" pageNumber="11">2</figureCitation>
). Major septa are pinnately arranged.
<taxonomicName id="0691A3837F13FFF2FF2BFEB4FF17FEE8" box="[151,245,331,355]" genus="Cardinal" pageId="10" pageNumber="8" rank="genus">Cardinal</taxonomicName>
fossula is distinguishable. Alar fossulae are also distinct, they are marked by the underdeveloped last pair of major septa in the counter quadrants. All septa meet in the corallite axis. The cardinal septum is thinner than the other major septa, whereas the counter is slightly thicker. External wall about 1.5 mm thick and internally enforced by steroplasmatic layer in the most external septal loculi. The mature morphology seen in the next transverse section (
<figureCitation id="59AAC4857F13FFF2FF6DFDB4FEC1FDE8" box="[209,291,587,611]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FE9FFDB4FECDFDE8" box="[291,303,587,611]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="10" pageNumber="11">3</figureCitation>
) is characterised by a pinnate arrangement of long major septa with n:d ratio of 24:8. Major septa are conspicuously acutely thickened in their peripheries and wedge-like rooted in the thick external wall (
<figureCitation id="59AAC4857F13FFF2FF23FD34FF11FD68" box="[159,243,715,739]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FF4FFD34FEE2FD68" box="[243,256,715,739]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="10" pageNumber="11">5</figureCitation>
); they are also slightly thickened to contiguity in their inner ends.
<taxonomicName id="0691A3837F13FFF2FEC0FD14FE38FC88" box="[380,474,747,771]" genus="Cardinal" pageId="10" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum shortens, whereas the counter is the longest. Pericardinal septa are as short as the cardinal. The cardinal fossula extends to the corallite axis, where it moderately widens. Alar septa are long and meet with the axial end of the long counter septum in the axis. The alar fossulae are still well-marked. Minor septa are absent. Peripheral interseptal loculi are filled by stereoplasma, thus apparently increasing the prominent, about 1.8 mm thick external wall (
<figureCitation id="59AAC4857F13FFF2FDA8FC14FD8AFB88" box="[532,616,1003,1027]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FDD4FC14FD97FB88" box="[616,629,1003,1027]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="10" pageNumber="11">5</figureCitation>
). One row of axially updomed tabulae well-developed in peripheral interseptal loculi of right cardinal and counter quadrant. Opposed, strongly horseshoe-arched, isolated and small peripheral, convex tabulae (i.e., inclined towards the external wall) are mostly confined to the left cardinal and counter quadrants. The last mature section below the calice (
<figureCitation id="59AAC4857F13FFF2FEB2FB34FEBDFB68" box="[270,351,1227,1251]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FEE3FB34FE8EFB68" box="[351,364,1227,1251]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="10" pageNumber="11">4</figureCitation>
) has a n:d ratio of 24:11. Major septa are withdrawn from the corallite axis, leaving a free axial area, invaded by the distinctly elongated counter septum.
<taxonomicName id="0691A3837F13FFF2FF2BFAD4FF17FAC8" box="[151,245,1323,1347]" genus="Cardinal" pageId="10" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum strongly shortened. Minor septa are absent. No tabulae are visible except for one in right alar fossula. Like in previous cross section (
<figureCitation id="59AAC4857F13FFF2FD80FA94FD6FFA08" box="[572,653,1387,1411]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
<figureCitation id="59AAC4857F13FFF2FD31FA94FD7BFA08" box="[653,665,1387,1411]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="10" pageNumber="11">3</figureCitation>
) septa are wedge-like thickened in external wall, which is 1.9 mm thick.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F13FFF2FF2BFA34FB0DFE48" pageId="10" pageNumber="11" type="discussion">
<paragraph id="C12ED8007F13FFF2FF2BFA34FB0DFE48" blockId="10.[151,776,235,1923]" lastBlockId="10.[813,1437,235,451]" pageId="10" pageNumber="11">
<emphasis id="F3E504127F13FFF2FF2BFA34FEFFFA68" bold="true" box="[151,285,1483,1507]" pageId="10" pageNumber="11">Discussion:</emphasis>
<taxonomicName id="0691A3837F13FFF2FE99FA34FCE4FA68" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[293,774,1483,1507]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="columnare">
<emphasis id="F3E504127F13FFF2FE99FA34FDDDFA68" box="[293,575,1483,1507]" italics="true" pageId="10" pageNumber="11">Actinophrentis columnare</emphasis>
Fedorowski, 1987
</taxonomicName>
is of similar size with n:d ratios ranging from 20:
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the
<typeStatus id="1E2A66A27F13FFF2FF78F9F4FEC6F9A8" box="[196,292,1547,1571]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
to 24:
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the largest
<typeStatus id="1E2A66A27F13FFF2FD8DF9F3FD71F9AF" box="[561,659,1548,1572]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
. However, in contrast, it has a remarkable thick counter septum at maturity forming a weak columella in the calice. In the studied species the counter septum thins with maturity and/or has the same thickness as the other major septa or is only slightly thicker. Its wall is conspicuously thicker than the wall of the
<taxonomicName id="0691A3837F13FFF2FE38F934FDF8F968" authorityName="Fedorowski" authorityYear="1987" box="[388,538,1739,1763]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="columnare">
<emphasis id="F3E504127F13FFF2FE38F934FDF8F968" box="[388,538,1739,1763]" italics="true" pageId="10" pageNumber="11">A. columnare</emphasis>
</taxonomicName>
. The current species differs from the smaller
<taxonomicName id="0691A3837F13FFF2FE04F914FEB7F8A8" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="bonespringense">
<emphasis id="F3E504127F13FFF2FE04F914FCE5F888" box="[440,775,1771,1795]" italics="true" pageId="10" pageNumber="11">Actinophrentis bonespringense</emphasis>
Fedorowski,
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</taxonomicName>
the wall thickness and n:d ratios. The
<taxonomicName id="0691A3837F13FFF2FF2BF8D4FEBFF8C9" box="[151,349,1835,1859]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F13FFF2FF2BF8D4FED4F8C8" box="[151,310,1835,1859]" italics="true" pageId="10" pageNumber="11">Actinophrentis</emphasis>
sp.
</taxonomicName>
(
<bibRefCitation id="A500A5F17F13FFF2FED7F8D4FDCEF8C8" author="Fedorowski, J." box="[363,556,1835,1859]" pageId="10" pageNumber="11" pagination="71 - 311" refId="ref25622" refString="Fedorowski, J. (2004). Considerations on most Rugosa and the Dividocorallia from de Groot's (1963) collection. Scripta Geologica, 127, 71 - 311." type="journal article" year="2004">Fedorowski, 2004</bibRefCitation>
), previously named
<taxonomicName id="0691A3837F13FFF2FF2BF8B4FD8EF8E8" authority="de Groot, 1963" authorityName="de Groot" authorityYear="1963" box="[151,620,1867,1891]" class="Anthozoa" family="Hapsiphyllidae" genus="Zaphrentites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="paralleloides">
<emphasis id="F3E504127F13FFF2FF2BF8B4FE55F8E8" box="[151,439,1867,1891]" italics="true" pageId="10" pageNumber="11">Zaphrentites paralleloides</emphasis>
de Groot, 1963
</taxonomicName>
differs in the radial arrangement of the major septa that extends from the earliest growth stages up to the calice, vs. the pinnate arrangement in the current species. Diameter and the number of septa in the Spanish species are much smaller. In addition, the type species of the genus,
<taxonomicName id="0691A3837F13FFF2FABFFEB4FC34FE08" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="donetziana">
<emphasis id="F3E504127F13FFF2FABFFEB4FA7EFEE8" box="[1283,1436,331,355]" italics="true" pageId="10" pageNumber="11">A. donetziana</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FC91FE94FC34FE08" author="Fomichev, V." box="[813,982,363,387]" pageId="10" pageNumber="11" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
appears to be smaller with a higher number of septa (n:d = 23:8.5); its wall is conspicuously thinner (based on fig. 203, 7a, b in
<bibRefCitation id="A500A5F17F13FFF2FBC4FE54FB03FE48" author="Hill, D." box="[1144,1249,427,451]" pageId="10" pageNumber="11" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill, 1981</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,158 @@
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<paragraph id="C12ED8007F12FFF3FF2BF973FF30F94F" blockId="11.[151,775,1676,1925]" pageId="11" pageNumber="12">
<heading id="9A666F6C7F12FFF3FF2BF973FF30F94F" fontSize="10" level="2" pageId="11" pageNumber="12" reason="7">
<taxonomicName id="0691A3837F12FFF3FF2BF973FCE5F92E" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" box="[151,775,1676,1701]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Stauriida" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F12FFF3FF2BF973FE31F92F" bold="true" box="[151,467,1676,1700]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FF2BF973FEF9F92F" bold="true" box="[151,283,1676,1700]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
cf.
<emphasis id="F3E504127F12FFF3FEE6F973FE31F92F" bold="true" box="[346,467,1676,1700]" italics="true" pageId="11" pageNumber="12">canadense</emphasis>
</emphasis>
<bibRefCitation id="A500A5F17F12FFF3FE58F972FCE5F92E" author="Fedorowski, J. &amp; Bamber, E. W." box="[484,775,1677,1701]" pageId="11" pageNumber="12" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber,</bibRefCitation>
</taxonomicName>
2001.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FF2BF933FDB0F8AF" pageId="11" pageNumber="12" type="description">
<paragraph id="C12ED8007F12FFF3FF2BF933FE92F96E" blockId="11.[151,775,1676,1925]" box="[151,368,1740,1765]" pageId="11" pageNumber="12">
(
<figureCitation id="59AAC4857F12FFF3FF23F933FF0CF96E" box="[159,238,1740,1765]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
<figureCitation id="59AAC4857F12FFF3FF53F932FEFFF96E" box="[239,285,1741,1765]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="11" pageNumber="12">1C</figureCitation>
<figureCitation id="59AAC4857F12FFF3FEA1F932FEC8F96E" box="[285,298,1741,1765]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
,
<figureCitation id="59AAC4857F12FFF3FE89F932FE86F96E" box="[309,356,1741,1765]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="11" pageNumber="12">12A</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F12FFF3FF2BF913FDB0F8AF" blockId="11.[151,775,1676,1925]" pageId="11" pageNumber="12">
cf. 2001
<taxonomicName id="0691A3837F12FFF3FEBDF913FF5FF8AE" authority="Fedorowski and Bamber." authorityName="Fedorowski and Bamber." class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F12FFF3FEBDF913FE1EF88F" box="[257,508,1772,1796]" italics="true" pageId="11" pageNumber="12">Lytvolasma canadense</emphasis>
Fedorowski and Bamber.
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—Text-Fig. 8, pl. 3,
<figureCitation id="59AAC4857F12FFF3FE2CF8F2FE2EF8AE" box="[400,460,1805,1829]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
, pl. 5,
<figureCitation id="59AAC4857F12FFF3FDADF8F2FDACF8AF" box="[529,590,1804,1829]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FF2BF8D3FC73FEA8" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="C12ED8007F12FFF3FF2BF8D3FC73FEA8" blockId="11.[151,775,1676,1925]" lastBlockId="11.[813,1437,235,1635]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FF2BF8D3FEE0F8CF" bold="true" box="[151,258,1836,1860]" pageId="11" pageNumber="12">Material:</emphasis>
Only one worn specimen (RAh 66) with calice rims and a large part of the corallite not preserved, collected from the basal shales of the lower member of the Aheimer Formation. Two transverse sections are available.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FC91FED4FAB6FC68" pageId="11" pageNumber="12" type="description">
<paragraph id="C12ED8007F12FFF3FC91FED4FBDBFE68" blockId="11.[813,1437,235,1635]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FC91FED4FC5FFEC8" bold="true" box="[813,957,299,323]" pageId="11" pageNumber="12">Description:</emphasis>
External characters: The preserved mature part of the corallite is
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long. It does not show any furrows or ridges on the outer wall except for very delicate growth lines. The calice is not preserved, and also the apex and a large part of the immature ontogenetic stages are not preserved.
</paragraph>
<paragraph id="C12ED8007F12FFF3FCFEFE14FAB6FC68" blockId="11.[813,1437,235,1635]" pageId="11" pageNumber="12">
Internal characters: Only two transverse sections in the mature part of the corallite are available for study. An ontogenetically earliest section (
<figureCitation id="59AAC4857F12FFF3FB00FDD4FAEFFDC8" box="[1212,1293,555,579]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
<figureCitation id="59AAC4857F12FFF3FAB2FDD4FAD8FDC8" box="[1294,1338,555,579]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="11" pageNumber="12">1C</figureCitation>
<figureCitation id="59AAC4857F12FFF3FA86FDD4FAAAFDC8" box="[1338,1352,555,579]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="11" pageNumber="12">2</figureCitation>
) shows sub-radial arrangement of septa, with short cardinal septum and long counter septum with a rhopaloid axial end. The axial area is free of septa, only the swollen end of the counter septum invading the centre. A typical half aulos is developed in the counter quadrants. At this stage n:d is 26:
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. Minor septa are underdeveloped. Biform morphology is developed. The last ephebic stage (
<figureCitation id="59AAC4857F12FFF3FA8CFCF4FA61FCA8" box="[1328,1411,779,803]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
<figureCitation id="59AAC4857F12FFF3FA3FFCF4FA72FCA8" box="[1411,1424,779,803]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
), more or less at the calice floor, shows all septa withdrawn from the axis and the axial area becomes large. Only the counter septum is still prominently longer, and the cardinal septum is slightly shorter than the others. The n:d ratio is 26:12.8 mm. Minor septa are rudimentary.
<taxonomicName id="0691A3837F12FFF3FAEAFC54FCACFC68" genus="Cardinal" pageId="11" pageNumber="8" rank="genus">Cardinal</taxonomicName>
fossula is slightly triangular, but open adaxially.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FC91FC14FA72F9E8" pageId="11" pageNumber="12" type="discussion">
<paragraph id="C12ED8007F12FFF3FC91FC14FA72F9E8" blockId="11.[813,1437,235,1635]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FC91FC14FC50FB88" bold="true" box="[813,946,1003,1027]" pageId="11" pageNumber="12">Discussion:</emphasis>
The current species closely resembles
<taxonomicName id="0691A3837F12FFF3FC91FBF4FB67FBA8" authorityName="Fedorowski" authorityYear="1987" box="[813,1157,1035,1059]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="counterseptatum">
<emphasis id="F3E504127F12FFF3FC91FBF4FB67FBA8" box="[813,1157,1035,1059]" italics="true" pageId="11" pageNumber="12">Bradyphyllum counterseptatum</emphasis>
</taxonomicName>
described by
<bibRefCitation id="A500A5F17F12FFF3FAA3FBF4FC9DFBC8" author="Fedorowski, J." pageId="11" pageNumber="12" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
. It differs from it in the n:d ratio in the ephebic stage; the current species is larger in dimensions than the former species. It also differs in the length of the counter septum; which is longer and thicker in the currently described species than the former.
<taxonomicName id="0691A3837F12FFF3FB15FB54FA7FFB48" box="[1193,1437,1195,1219]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F12FFF3FB15FB54FA7FFB48" box="[1193,1437,1195,1219]" italics="true" pageId="11" pageNumber="12">Lytvolasma canadense</emphasis>
</taxonomicName>
is distinguished from all other
<taxonomicName id="0691A3837F12FFF3FB38FB34FAB5FB68" box="[1156,1367,1227,1251]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FB38FB34FAE2FB68" box="[1156,1280,1227,1251]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
species
</taxonomicName>
by its greater dimensions (
<figureCitation id="59AAC4857F12FFF3FBAAFB14FB98FA88" box="[1046,1146,1259,1283]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="11" pageNumber="12">Fig. 12A</figureCitation>
), the stronger elongation of its counter septum and in the underdevelopment of its minor septa. The shortening of the last major septa in all quadrants is not expressed herein. However, in the n:d ratio (
<figureCitation id="59AAC4857F12FFF3FC1CFA94FBE0FA08" box="[928,1026,1387,1411]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="11" pageNumber="12">Fig. 12A</figureCitation>
) and further morphological aspects our specimen seems to be identical to
<taxonomicName id="0691A3837F12FFF3FB67FA73FA8BFA28" box="[1243,1385,1419,1443]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F12FFF3FB67FA73FA8BFA28" box="[1243,1385,1419,1443]" italics="true" pageId="11" pageNumber="12">L. canadense</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A500A5F17F12FFF3FC91FA54FB9BFA48" author="Fedorowski, J. &amp; Bamber, E. W." box="[813,1145,1451,1475]" pageId="11" pageNumber="12" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
,
<figureCitation id="59AAC4857F12FFF3FB3BFA54FB0FFA48" box="[1159,1261,1451,1475]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="11" pageNumber="12">Fig. 8c, d</figureCitation>
) from the Guadalupian (Wordian) of Ellesmere Island (Sverdrup Basin, Canadian Arctic). Due to the fragmentary nature of our single Egyptian specimen and considerable geographic and stratigraphic differences we did not include it into the nominate species and retained determination as “cf ”.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,250 @@
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<subSubSection id="898B8B8B7F12FFF3FC91F953FB73F94E" box="[813,1169,1708,1733]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="C12ED8007F12FFF3FC91F953FB73F94E" blockId="11.[813,1169,1708,1733]" box="[813,1169,1708,1733]" pageId="11" pageNumber="12">
<heading id="9A666F6C7F12FFF3FC91F953FB73F94E" bold="true" box="[813,1169,1708,1733]" fontSize="10" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="F3E504127F12FFF3FC91F953FB73F94E" bold="true" box="[813,1169,1708,1733]" pageId="11" pageNumber="12">
<taxonomicName id="0691A3837F12FFF3FC91F953FBACF94F" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[813,1102,1708,1732]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="aheimerensis" status="sp. nov.">
<emphasis id="F3E504127F12FFF3FC91F953FBACF94F" bold="true" box="[813,1102,1708,1732]" italics="true" pageId="11" pageNumber="12">Lytvolasma aheimerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F12FFF3FBE8F952FB73F94E" box="[1108,1169,1709,1733]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FC91F933FBF8F8AF" pageId="11" pageNumber="12" type="description">
<paragraph id="C12ED8007F12FFF3FC91F933FC48F96E" blockId="11.[813,1437,1740,1892]" box="[813,938,1740,1765]" pageId="11" pageNumber="12">
(
<figureCitation id="59AAC4857F12FFF3FC88F933FC7CF96E" box="[820,926,1740,1765]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="11" pageNumber="12">Fig. 7AI</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F12FFF3FC91F913FBF8F8AF" blockId="11.[813,1437,1740,1892]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FC91F913FBA9F88E" bold="true" box="[813,1099,1772,1797]" pageId="11" pageNumber="12">Derivation of the name</emphasis>
: From the bearing rock unit “Aheimer Formation&quot;.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF4FC91F8D2FA7AF9D3" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="C12ED8007F12FFF3FC91F8D2FAE1F8EF" blockId="11.[813,1437,1740,1892]" pageId="11" pageNumber="12">
<materialsCitation id="71F9D25D7F12FFF3FC91F8D2FAE1F8EF" pageId="11" pageNumber="12" specimenCode="RAh 2" specimenCount="1" typeStatus="holotype">
<emphasis id="F3E504127F12FFF3FC91F8D2FC7DF8CE" bold="true" box="[813,927,1837,1861]" pageId="11" pageNumber="12">
<typeStatus id="1E2A66A27F12FFF3FC91F8D2FC79F8CE" box="[813,923,1837,1861]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="D79713897F12FFF3FC11F8D3FBFAF8CF" box="[941,1048,1836,1860]" pageId="11" pageNumber="12" type="generic">Specimen</specimenCount>
<specimenCode id="9137707B7F12FFF3FB9BF8D3FB94F8CE" box="[1063,1142,1836,1861]" pageId="11" pageNumber="12">RAh 2</specimenCode>
, illustrated in
<figureCitation id="59AAC4857F12FFF3FA94F8D3FA7BF8CE" box="[1320,1433,1836,1861]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="11" pageNumber="12">Fig. 7AI</figureCitation>
. Eight transverse thin-sections are available.
</materialsCitation>
</paragraph>
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<paragraph id="C12ED8007F15FFF4FF1BFAC1FA8FFA5C" blockId="12.[167,1399,1342,1495]" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FF1BFAC1FF38FAD9" bold="true" box="[167,218,1342,1362]" pageId="12" pageNumber="13">Fig. 7</emphasis>
<taxonomicName id="0691A3837F15FFF4FF58FAC1FE52FAD9" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[228,432,1342,1362]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="aheimerensis" status="sp. nov.">
<emphasis id="F3E504127F15FFF4FF58FAC1FE52FAD9" box="[228,432,1342,1362]" italics="true" pageId="12" pageNumber="13">Lytvolasma aheimerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F15FFF4FE09FAC1FE03FAD9" box="[437,481,1342,1362]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="F3E504127F15FFF4FE58FAC1FE10FAD9" bold="true" box="[484,498,1342,1362]" pageId="12" pageNumber="13">A</emphasis>
External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.
<emphasis id="F3E504127F15FFF4FD5EFAA7FCEAFAE7" bold="true" box="[738,776,1368,1388]" pageId="12" pageNumber="13">BD</emphasis>
Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in
<emphasis id="F3E504127F15FFF4FC56FA8CFC1BFA0C" bold="true" box="[1002,1017,1395,1415]" pageId="12" pageNumber="13">D</emphasis>
.
<emphasis id="F3E504127F15FFF4FBBDFA8CFBC4FA0C" bold="true" box="[1025,1062,1395,1415]" pageId="12" pageNumber="13">EG</emphasis>
Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end.
<emphasis id="F3E504127F15FFF4FE1BFA57FE24FA37" bold="true" box="[423,454,1448,1468]" pageId="12" pageNumber="13">HI</emphasis>
Two transverse thin-sections below the calice, deformed due to rejuvenation in
<emphasis id="F3E504127F15FFF4FBD1FA57FB99FA37" bold="true" box="[1133,1147,1448,1468]" pageId="12" pageNumber="13">H</emphasis>
and lateral compaction in
<emphasis id="F3E504127F15FFF4FAE2FA57FA86FA37" bold="true" box="[1374,1380,1448,1468]" pageId="12" pageNumber="13">I</emphasis>
. Scale bars: 2.5 mm, except
<emphasis id="F3E504127F15FFF4FE37FA3CFE7BFA5C" bold="true" box="[395,409,1475,1495]" pageId="12" pageNumber="13">A</emphasis>
: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
</paragraph>
</caption>
<paragraph id="C12ED8007F15FFF4FF2BF9DFFE94F953" blockId="12.[151,775,1568,1881]" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FF2BF9DFFEE0F9B3" bold="true" box="[151,258,1568,1592]" pageId="12" pageNumber="13">Material:</emphasis>
In addition to the
<typeStatus id="1E2A66A27F15FFF4FE5AF9DFFDAAF9B3" box="[486,584,1568,1592]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, two moderately preserved
<typeStatus id="1E2A66A27F15FFF4FEACF9BEFE9BF9D2" box="[272,377,1601,1625]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
with compressed to crushed calices and eroded apical ends (RAh 10 and RAh 113). Five transverse thin-sections are available. They are collected from the basal shales of the lower member of the Aheimer Formation.
</paragraph>
<paragraph id="C12ED8007F15FFF4FF2BF91FFF3FF8D2" blockId="12.[151,775,1568,1881]" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FF2BF91FFED5F973" bold="true" box="[151,311,1760,1784]" pageId="12" pageNumber="13">Type locality:</emphasis>
<materialsCitation id="71F9D25D7F15FFF4FE83F91EFF3BF8D2" country="Egypt" county="Gulf" latitude="29.4725" location="Northern Galala" longLatPrecision="20" longitude="32.460556" municipality="northern Eastern Desert" pageId="12" pageNumber="13" specimenCount="1" stateProvince="As Suways">
Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
<emphasis id="F3E504127F15FFF4FEF4F900FEB1F893" box="[328,339,1791,1816]" italics="true" pageId="12" pageNumber="13"></emphasis>
21
<emphasis id="F3E504127F15FFF4FECFF900FE61F893" box="[371,387,1791,1816]" italics="true" pageId="12" pageNumber="13"></emphasis>
N and Long. 32° 27
<emphasis id="F3E504127F15FFF4FDE6F900FD87F893" box="[602,613,1791,1816]" italics="true" pageId="12" pageNumber="13"></emphasis>
38
<emphasis id="F3E504127F15FFF4FD3AF900FD74F893" box="[646,662,1791,1816]" italics="true" pageId="12" pageNumber="13"></emphasis>
E), northern Eastern Desert, western side of the Gulf of Suez, Egypt
</materialsCitation>
.
</paragraph>
<paragraph id="C12ED8007F15FFF4FC91F9DFFA7AF9D3" blockId="12.[813,1437,1568,1912]" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FC91F9DFFC32F9B3" bold="true" box="[813,976,1568,1592]" pageId="12" pageNumber="13">Type horizon:</emphasis>
Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F15FFF4FC91F99FFCBEF973" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="C12ED8007F15FFF4FC91F99FFCBEF973" blockId="12.[813,1437,1568,1912]" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FC91F99FFC44F9F3" bold="true" box="[813,934,1632,1656]" pageId="12" pageNumber="13">Diagnosis:</emphasis>
<taxonomicName id="0691A3837F15FFF4FC10F99FFBCAF9F3" box="[940,1064,1632,1656]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F15FFF4FC10F99FFBCAF9F3" box="[940,1064,1632,1656]" italics="true" pageId="12" pageNumber="13">Lytvolasma</emphasis>
</taxonomicName>
with thick wall, up to
<quantity id="066975E57F15FFF4FAA5F99FFABBF9F3" box="[1305,1369,1632,1656]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
thick.
<taxonomicName id="0691A3837F15FFF4FC91F97FFC69F913" box="[813,907,1664,1688]" genus="Cardinal" pageId="12" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum shortened from early maturity. Counter septum elongated up to late maturity without forming a columella. It enters the free axial area with its rhopaloid end.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F15FFF5FC91F8FFFE55FAA8" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="C12ED8007F15FFF5FC91F8FFFD95FEE8" blockId="12.[813,1437,1568,1912]" lastBlockId="13.[151,775,235,1571]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="F3E504127F15FFF4FC91F8FFFC5FF893" bold="true" box="[813,957,1792,1816]" pageId="12" pageNumber="13">Description:</emphasis>
External characters: Solitary, small, conicocylindrical corallites. The preserved parts of the three corallites ranges in length between 2 and
<quantity id="066975E57F15FFF4FAAEF8BFFAADF8D3" box="[1298,1359,1856,1880]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="cm" value="3.0">3 cm</quantity>
. Moderately preserved; their apexes and much of their early neanic stages are missing. Calices are deep. Growth lines, rugae, septal ridges and furrows are well-developed on the outer wall of the corallites. Rejuvenation is not common, only constrictions are visible (
<figureCitation id="59AAC4857F14FFF5FDA5FEB4FD89FEE8" box="[537,619,331,355]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F14FFF5FF11FE94FE55FAA8" blockId="13.[151,775,235,1571]" pageId="13" pageNumber="14">
Internal characters: The ontogenetically earliest growth stage available has 23 septa in 5.9 mm (
<figureCitation id="59AAC4857F14FFF5FD0DFE74FF4EFE48" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7B, C</figureCitation>
). Major septa are radially arranged. They are very thick and joined together in the centre forming a strong axial structure. The cardinal septum is somewhat thinner and the fossular break is noticeable. Further in the ontogeny, at a diameter of 8.813.3 mm with 2426 septa (
<figureCitation id="59AAC4857F14FFF5FF5EFDB4FEBCFDE8" box="[226,350,587,611]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7DH</figureCitation>
), the septa become thinner gradually with maturity, and are more radially arranged. They are united near the axis with their rhopaloid ends, forming a stereozone with a free axial area. A strange attitude of one of the lateral cardinal septa is shown in
<figureCitation id="59AAC4857F14FFF5FD29FD34FCE0FD68" box="[661,770,715,739]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7GI</figureCitation>
. It started to diminish gradually with maturity. This led to the reduction of the number of septa below the calice (
<figureCitation id="59AAC4857F14FFF5FF23FCD4FF0BFCC8" box="[159,233,811,835]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7I</figureCitation>
) to 25 septa. This attitude might be attributed to the rejuvenation took place in this part of the corallite. The cardinal fossula is slightly triangular and open adaxially. Counter septum is the longest with swollen end, invading the free axial area (
<figureCitation id="59AAC4857F14FFF5FDA1FC54FD8DFC48" box="[541,623,939,963]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7F</figureCitation>
), and irregular in shape. Alar fossulae are not developed. The most adult stage, below the calice and the calice itself are compacted and the septa are somewhat deformed, the stereozone gradually diminishes and the septa retreat a little from the centre (
<figureCitation id="59AAC4857F14FFF5FE04FBB4FDE7FBE8" box="[440,517,1099,1123]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="13" pageNumber="14">Fig. 7I</figureCitation>
), the cardinal septum becomes distinctively shorter than the other major septa, whereas the counter is still long with thick rhopaloid end. The wall is thick (0.9
<quantity id="066975E57F14FFF5FDA7FB54FDBDFB48" box="[539,607,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="2.0">2 mm</quantity>
) in all ontogenetic stages. Minor septa are absent. A white line in the midlines of the septa may indicate their original compound trabecular growth.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FF2BFAD4FABFFDEE" pageId="13" pageNumber="14" type="discussion">
<paragraph id="C12ED8007F14FFF5FF2BFAD4FED3F9A8" blockId="13.[151,775,235,1571]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FF2BFAD4FEC2FAC8" bold="true" box="[151,288,1323,1347]" pageId="13" pageNumber="14">Discussion:</emphasis>
The elongation of the counter septum makes this species similar to
<taxonomicName id="0691A3837F14FFF5FE53FAB4FF2FFA08" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F14FFF5FE53FAB4FD69FAE8" box="[495,651,1355,1379]" italics="true" pageId="13" pageNumber="14">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F14FFF5FD29FAB4FF2FFA08" author="Fomichev, V." pageId="13" pageNumber="14" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
. It can be differentiated from
<taxonomicName id="0691A3837F14FFF5FD9EFA94FD5CFA08" authorityName="Fomichev, Shortened" authorityYear="1953" box="[546,702,1387,1411]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F14FFF5FD9EFA94FD5CFA08" box="[546,702,1387,1411]" italics="true" pageId="13" pageNumber="14">Monophyllum</emphasis>
</taxonomicName>
in the shape of the cardinal fossula and the lack of the incipient columella formation in the calice. The current species differs from
<taxonomicName id="0691A3837F14FFF5FEDDFA33FD74FA68" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[353,662,1483,1507]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F14FFF5FEDDFA33FE23FA68" box="[353,449,1484,1507]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
Fedorowski, 1987
</taxonomicName>
from the PennsylvanianPermian transition of southern
<collectingRegion id="035516E27F14FFF5FD17FA13FD08F988" box="[683,746,1516,1539]" country="United States of America" name="Texas" pageId="13" pageNumber="14">Texas</collectingRegion>
in the following:
</paragraph>
<paragraph id="C12ED8007F14FFF5FF10F9B2FD81F90E" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">• Number of septa are higher and diameter (n:d ratio) is larger in the present species.</paragraph>
<paragraph id="C12ED8007F14FFF5FF10F972FEC2F94F" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">• The wall of the current species is remarkably thicker.</paragraph>
<paragraph id="C12ED8007F14FFF5FF10F932FDD9F8CF" blockId="13.[172,775,1613,1861]" pageId="13" pageNumber="14">
• Minor septa are short and restricted to the corallite wall in the latest mature stages of
<taxonomicName id="0691A3837F14FFF5FDFBF912FD43F88F" authorityName="Fedorowski" authorityYear="1987" box="[583,673,1773,1796]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F14FFF5FDFBF912FD43F88F" box="[583,673,1773,1796]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
</taxonomicName>
, whereas they are absent in the current species even in the last ephebic stage near the calice floor.
</paragraph>
<paragraph id="C12ED8007F14FFF5FCFEFF13FAAAFEC8" blockId="13.[834,1436,235,323]" pageId="13" pageNumber="14">
• The counter septum in the present species is longer, enters the free axial area, whereas it is only slightly longer than the other major septa in
<emphasis id="F3E504127F14FFF5FB56FED3FAAAFEC8" box="[1258,1352,300,323]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="0691A3837F14FFF5FB56FED3FAA1FEC8" authorityName="Fedorowski" authorityYear="1987" box="[1258,1347,300,323]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">L. aucta</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="C12ED8007F14FFF5FCFEFE92FABFFDEE" blockId="13.[813,1437,364,613]" pageId="13" pageNumber="14">
The type species of the genus,
<taxonomicName id="0691A3837F14FFF5FB2BFE92FC83FE2F" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="asymetrica">
<emphasis id="F3E504127F14FFF5FB2BFE92FAC9FE0F" box="[1175,1323,365,388]" italics="true" pageId="13" pageNumber="14">L. asymetrica</emphasis>
<bibRefCitation id="A500A5F17F14FFF5FA8FFE93FC83FE2F" author="Soshkina, E. D." pageId="13" pageNumber="14" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
differs by the early appearance of free axial area in the
<typeStatus id="1E2A66A27F14FFF5FCE7FE53FC59FE4F" box="[859,955,428,452]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="A500A5F17F14FFF5FC72FE53FB42FE4E" author="Fedorowski, J." box="[974,1184,428,453]" pageId="13" pageNumber="14" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, with comparisons to further similar specimens from different taxa) and the appearance of minor septa (Chwieduk, 2013), though n:d ratios are similar.
<taxonomicName id="0691A3837F14FFF5FC4AFDF2FBB3FDAF" authorityName="Fedorowski" authorityYear="1987" box="[1014,1105,525,548]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F14FFF5FC4AFDF2FBB3FDAF" box="[1014,1105,525,548]" italics="true" pageId="13" pageNumber="14">L. aucta</emphasis>
</taxonomicName>
illustrated in
<bibRefCitation id="A500A5F17F14FFF5FB4CFDF2FC4CFDCE" author="Kora, M. &amp; Mansour, Y." pageId="13" pageNumber="14" pagination="597 - 614" refId="ref27107" refString="Kora, M., &amp; Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991)</bibRefCitation>
(=
<taxonomicName id="0691A3837F14FFF5FC6AFDD2FB85FDCF" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[982,1127,557,580]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="paraaucta" status="n. sp.">
<emphasis id="F3E504127F14FFF5FC6AFDD2FB85FDCF" box="[982,1127,557,580]" italics="true" pageId="13" pageNumber="14">L. paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F14FFF5FBCEFDD2FB52FDCE" box="[1138,1200,557,581]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
) has smaller dimensions and a lower n:d ratio than the present species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,317 @@
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<subSubSection id="898B8B8B7F13FFF2FC91FDF2FB4CFDAF" box="[813,1198,524,549]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="C12ED8007F13FFF2FC91FDF2FB4CFDAF" blockId="10.[813,1437,524,1925]" box="[813,1198,524,549]" pageId="10" pageNumber="11">
<heading id="9A666F6C7F13FFF2FC91FDF2FB4CFDAF" box="[813,1198,524,549]" fontSize="36" level="2" pageId="10" pageNumber="11" reason="3">
Genus
<taxonomicName id="0691A3837F13FFF2FCC5FDF3FB45FDAF" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[889,1191,524,549]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FCC5FDF3FC1FFDAF" bold="true" box="[889,1021,524,548]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FBB8FDF2FB45FDAF" author="Soshkina, E. D." box="[1028,1191,524,549]" pageId="10" pageNumber="11" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F13FFF2FC91FDD2FC85FDEE" pageId="10" pageNumber="11" type="type_taxon">
<paragraph id="C12ED8007F13FFF2FC91FDD2FC85FDEE" blockId="10.[813,1437,524,1925]" pageId="10" pageNumber="11">
<emphasis id="F3E504127F13FFF2FC91FDD2FC2DFDCF" bold="true" box="[813,975,556,581]" pageId="10" pageNumber="11">
<typeStatus id="1E2A66A27F13FFF2FC91FDD2FC84FDCE" box="[813,870,557,581]" pageId="10" pageNumber="11">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F13FFF2FC62FDD2FA7EFDCF" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[990,1436,556,581]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="asymetrica">
<emphasis id="F3E504127F13FFF2FC62FDD2FB02FDCF" box="[990,1248,557,581]" italics="true" pageId="10" pageNumber="11">Lytvolasma asymetrica</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FB53FDD2FA7EFDCF" author="Soshkina, E. D." box="[1263,1436,556,581]" pageId="10" pageNumber="11" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
[sic!].
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F13FFF2FC91FD93FC24FCCE" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="C12ED8007F13FFF2FC91FD93FC24FCCE" blockId="10.[813,1437,524,1925]" pageId="10" pageNumber="11">
<emphasis id="F3E504127F13FFF2FC91FD93FC44FD0F" bold="true" box="[813,934,620,644]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
Non dissepimented small solitary corals; cardinal fossula deep, key-hole shaped, often bordered by half aulos in counter quadrants; cardinal septum shortened late in ontogeny; counter septum slightly elongated; major septa semi-radial in arrangement; minor septa very short; microstructure of septa trabecular (
<bibRefCitation id="A500A5F17F13FFF2FAEFFCF2FC5AFCCE" author="Fedorowski, J." pageId="10" pageNumber="11" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F13FFF3FC91FCB2FD8CFCA8" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="C12ED8007F13FFF2FC91FCB2FC29FA4F" blockId="10.[813,1437,524,1925]" pageId="10" pageNumber="11">
<emphasis id="F3E504127F13FFF2FC91FCB2FC78FCEE" bold="true" box="[813,922,845,869]" pageId="10" pageNumber="11">Remarks:</emphasis>
<taxonomicName id="0691A3837F13FFF2FC18FCB2FBC2FCEE" box="[932,1056,845,869]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC18FCB2FBC2FCEE" box="[932,1056,845,869]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
is a complex genus with unclear relations to several other taxa, as extensively discussed by
<bibRefCitation id="A500A5F17F13FFF2FCF1FC72FBC6FC2E" author="Fedorowski, J." box="[845,1060,909,933]" pageId="10" pageNumber="11" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
. He modified the diagnosis by
<bibRefCitation id="A500A5F17F13FFF2FACFFC72FC91FC4E" author="Hill, D." pageId="10" pageNumber="11" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
who questioned the validity of
<taxonomicName id="0691A3837F13FFF2FB57FC52FA85FC4E" box="[1259,1383,941,965]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FB57FC52FA85FC4E" box="[1259,1383,941,965]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
and considered it as a possible junior synonym of
<taxonomicName id="0691A3837F13FFF2FA9FFC32FC15FB8E" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FA9FFC32FCBAFB8E" italics="true" pageId="10" pageNumber="11">Bradyphyllum</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FCDEFC12FC15FB8E" author="Grabau, A. W." box="[866,1015,1005,1029]" pageId="10" pageNumber="11" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
. However, due to morphological differences
<bibRefCitation id="A500A5F17F13FFF2FC2DFBF2FB88FBAE" author="Fedorowski, J." box="[913,1130,1037,1061]" pageId="10" pageNumber="11" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
considered both genera as valid and did not include the presence of long, slightly curved, thickened septa that are laterally contiguous over much of their length and almost reach the axis from Hills diagnosis. However, opposed to Fedorowksi, these characteristics were confirmed by Chwieduk (2013, tab. 3). According to
<bibRefCitation id="A500A5F17F13FFF2FC53FB33FB2BFB6E" author="Fedorowski, J." box="[1007,1225,1228,1253]" pageId="10" pageNumber="11" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
, intimate relations of the genus exist with further
<taxonomicName id="0691A3837F13FFF2FB33FB13FAC7FA8F" box="[1167,1317,1260,1284]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="35" phylum="Cnidaria" rank="subFamily" subFamily="Antiphyllinae">Antiphyllinae</taxonomicName>
, as
<taxonomicName id="0691A3837F13FFF2FAEEFB13FC8AFAAF" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FAEEFB13FC8AFAAF" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
specimens were attributed by authors in part to
<taxonomicName id="0691A3837F13FFF2FC91FAD2FB61FACE" authority="Ivanovski, 1967" authorityName="Ivanovski" authorityYear="1967" box="[813,1155,1325,1349]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC91FAD2FC2EFACE" box="[813,972,1325,1349]" italics="true" pageId="10" pageNumber="11">Actinophrentis</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FC6AFAD2FB61FACE" author="Ivanovski, A. B." box="[982,1155,1325,1349]" pageId="10" pageNumber="11" refId="ref26871" refString="Ivanovski, A. B. (1967). Studies on Early Carboniferous Rugosa. 92 pp., Nauka, Moscow (in Russian)." type="book" year="1967">Ivanovski, 1967</bibRefCitation>
</taxonomicName>
, as well as to species of
<taxonomicName id="0691A3837F13FFF2FC91FAB2FB75FAEE" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" box="[813,1175,1357,1381]" genus="Allotropiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC91FAB2FC16FAEE" box="[813,1012,1357,1381]" italics="true" pageId="10" pageNumber="11">Allotropiophyllum</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FBBCFAB2FB75FAEE" author="Grabau, A. W." box="[1024,1175,1357,1381]" pageId="10" pageNumber="11" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0691A3837F13FFF2FB64FAB2FA91FAEE" authorityName="Grabau" authorityYear="1928" box="[1240,1395,1357,1381]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FB64FAB2FA91FAEE" box="[1240,1395,1357,1381]" italics="true" pageId="10" pageNumber="11">Bradyphyllum</emphasis>
</taxonomicName>
. In addition, the
<typeStatus id="1E2A66A27F13FFF2FC01FA92FC0EFA0E" box="[957,1004,1389,1413]" pageId="10" pageNumber="11">type</typeStatus>
species of the hapsiphyllinid
<taxonomicName id="0691A3837F13FFF2FA90FA92FC67FA2E" class="Anthozoa" genus="Parasterophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FA90FA92FC67FA2E" italics="true" pageId="10" pageNumber="11">Parasterophrentis</emphasis>
</taxonomicName>
,
<taxonomicName id="0691A3837F13FFF2FC2FFA72FB4FFA2F" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[915,1197,1420,1445]" class="Anthozoa" genus="Parasterophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="virgata">
<emphasis id="F3E504127F13FFF2FC2FFA72FC18FA2F" box="[915,1018,1421,1444]" italics="true" pageId="10" pageNumber="11">P. virgata</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FBBEFA72FB4FFA2F" author="Fomichev, V." box="[1026,1197,1420,1445]" pageId="10" pageNumber="11" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
has to be transferred to
<taxonomicName id="0691A3837F13FFF2FCF5FA52FC27FA4E" box="[841,965,1453,1477]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FCF5FA52FC27FA4E" box="[841,965,1453,1477]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C12ED8007F13FFF3FC91FA32FD8CFCA8" blockId="10.[813,1437,524,1925]" lastBlockId="11.[151,775,235,1603]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Opposed,
<taxonomicName id="0691A3837F13FFF2FC19FA32FBC3FA6E" box="[933,1057,1485,1509]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC19FA32FBC3FA6E" box="[933,1057,1485,1509]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
(?) cf.
<taxonomicName id="0691A3837F13FFF2FB3FFA32FB17FA6F" box="[1155,1269,1485,1508]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="geinitzi">
<emphasis id="F3E504127F13FFF2FB3FFA32FB17FA6F" box="[1155,1269,1485,1508]" italics="true" pageId="10" pageNumber="11">L. geinitzi</emphasis>
</taxonomicName>
(
<bibRefCitation id="A500A5F17F13FFF2FAB7FA32FA71FA6F" author="Toula, F." box="[1291,1427,1484,1509]" pageId="10" pageNumber="11" pagination="1 - 77" refId="ref29539" refString="Toula, F. (1875). Eine Kohlenkalk-Fauna von den Barents-Inseln (Novaja Semlja NW.). Sitzungsberichte Der Akademie Der Wissenschaften Wien (mathematisch Naturwissenschaftliche Klasse), 71, 1 - 77." type="journal article" year="1875">Toula, 1875</bibRefCitation>
) described by
<bibRefCitation id="A500A5F17F13FFF2FC7AFA12FBBAF98E" author="FlUgel, H. W." box="[966,1112,1517,1541]" pageId="10" pageNumber="11" pagination="1 - 57" refId="ref26012" refString="FlUgel, H. W. (1973). Rugose Korallen aus dem oberen Perm Ost-Gronlands. Verhandlungen Der Geologischen Bundes-Anstalt., 1, 1 - 57." type="journal article" year="1973">Flügel (1973)</bibRefCitation>
from the Upper Permian of eastern
<collectingCountry id="B98698907F13FFF2FC3FF9F2FC14F9AE" box="[899,1014,1549,1573]" name="Greenland" pageId="10" pageNumber="11">Greenland</collectingCountry>
was included in
<taxonomicName id="0691A3837F13FFF2FB0DF9F2FC31F9CF" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" class="Anthozoa" family="Pentaphyllidae" genus="Tachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="rhizoides">
<emphasis id="F3E504127F13FFF2FB0DF9F2FA7FF9AE" box="[1201,1437,1549,1573]" italics="true" pageId="10" pageNumber="11">Tachylasma rhizoides</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FC91F9D2FC31F9CF" author="Soshkina, E. D." box="[813,979,1580,1605]" pageId="10" pageNumber="11" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="A500A5F17F13FFF2FBBDF9D2FB2DF9CE" author="Fedorowski, J." box="[1025,1231,1581,1605]" pageId="10" pageNumber="11" pagination="71 - 91" refId="ref25483" refString="Fedorowski, J. (1982). Some rugose corals from the Upper Permian of East Greenland. Rapport Gronlands Geologiske Undersogelse, 108, 71 - 91." type="journal article" year="1982">Fedorowski (1982)</bibRefCitation>
. He also excluded
<taxonomicName id="0691A3837F13FFF2FC91F9B3FC4BF9EF" box="[813,937,1612,1636]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC91F9B3FC4BF9EF" box="[813,937,1612,1636]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
from Northeast
<collectingCountry id="B98698907F13FFF2FBE3F9B3FB43F9EF" box="[1119,1185,1612,1636]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
figured by
<bibRefCitation id="A500A5F17F13FFF2FAA7F9B2FA75F9EE" author="Guo, S." box="[1307,1431,1613,1637]" pageId="10" pageNumber="11" pagination="1 - 686" refId="ref26265" refString="Guo, S. (1980). Tetracoralla. In: Paleontological atlas of Northeast China. Paleozoic vol. 1 - 686. Beijing." type="book chapter" year="1980">Guo (1980)</bibRefCitation>
, but opposed,
<bibRefCitation id="A500A5F17F13FFF2FC7DF993FAF4F90E" author="Fedorowski, J. &amp; Bamber, E. W." box="[961,1302,1644,1669]" pageId="10" pageNumber="11" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber (2001)</bibRefCitation>
did not rule out the possibility that some of Guos specimens might be included.
<taxonomicName id="0691A3837F13FFF2FC7BF953FBA1F94F" box="[967,1091,1708,1732]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FC7BF953FBA1F94F" box="[967,1091,1708,1732]" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
</taxonomicName>
described from
<collectingRegion id="035516E27F13FFF2FAB0F953FAA4F94F" box="[1292,1350,1708,1732]" country="China" name="Xizang" pageId="10" pageNumber="11">Tibet</collectingRegion>
(
<bibRefCitation id="A500A5F17F13FFF2FAE6F952FC27F96E" author="Wang, Z. - J. &amp; Liu, S. - K." pageId="10" pageNumber="11" refId="ref30148" refString="Wang, Z. - J., &amp; Liu, S. - K. (1982). Early Lower Permian rugose corals from the Saga, Zhongba and Namuco areas of Xizang. Contribution to the geology" type="book" year="1982">Wang and Liu, 1982</bibRefCitation>
;
<bibRefCitation id="A500A5F17F13FFF2FC69F932FBDDF96F" author="Wu, W. S." box="[981,1087,1740,1764]" pageId="10" pageNumber="11" pagination="1966 - 1968" refId="ref30601" refString="Wu, W. S. (1975). The coral fossils from the Qomolangma Feng region. In: A report of scientific investigation in the Qomolangma Feng Region 1966 - 1968 (Palaontology, part 1), 83 - 128; Beijing (Science Press). (in Chinese)." type="book chapter" year="1975">Wu, 1975</bibRefCitation>
) and Xingjiang (
<bibRefCitation id="A500A5F17F13FFF2FB41F932FC83F88F" author="Wang, Z. - J. &amp; Yu, X. - G." pageId="10" pageNumber="11" pagination="657 - 662" refId="ref30209" refString="Wang, Z. - J., &amp; Yu, X. - G. (1986). Early Late Carboniferous corals from Jinghe of Xinjiang. Acta Palaeontologica Sinica, 25 (6), 657 - 662." type="journal article" year="1986">Wang and Yu, 1986</bibRefCitation>
) were included in
<taxonomicName id="0691A3837F13FFF2FB8DF913FA7AF88E" authority="(Fedorowski, 1982)" baseAuthorityName="Fedorowski" baseAuthorityYear="1982" box="[1073,1432,1772,1797]" class="Anthozoa" family="Hapsiphyllidae" genus="Euryphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FB8DF913FB5CF88F" box="[1073,1214,1772,1796]" italics="true" pageId="10" pageNumber="11">Euryphyllum</emphasis>
(
<bibRefCitation id="A500A5F17F13FFF2FB71F913FA6FF88E" author="Fedorowski, J." box="[1229,1421,1772,1797]" pageId="10" pageNumber="11" pagination="71 - 91" refId="ref25483" refString="Fedorowski, J. (1982). Some rugose corals from the Upper Permian of East Greenland. Rapport Gronlands Geologiske Undersogelse, 108, 71 - 91." type="journal article" year="1982">Fedorowski, 1982</bibRefCitation>
)
</taxonomicName>
. Similarities to further genera were extensively discussed by Fedorowksi (1987a). According to the author,
<taxonomicName id="0691A3837F13FFF2FAE5F8D2FBC1F8EF" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FAE5F8D2FC92F8EE" italics="true" pageId="10" pageNumber="11">Lytvolasma</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FCC6F8B2FBC1F8EF" author="Soshkina, E. D." box="[890,1059,1868,1893]" pageId="10" pageNumber="11" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
shows some similarities to
<taxonomicName id="0691A3837F13FFF2FADAF8B2FB83F80E" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" genus="Allotropiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F13FFF2FADAF8B2FC25F80E" italics="true" pageId="10" pageNumber="11">Allotropiophyllum</emphasis>
<bibRefCitation id="A500A5F17F13FFF2FC73F892FB83F80E" author="Grabau, A. W." box="[975,1121,1901,1925]" pageId="10" pageNumber="11" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
. It is distinguished from the latter by the elongation of counter septum and the alar septa in the early growth stage, a slightly but permanently elongated counter septum at maturity, shortening of the cardinal septum late in ontogeny, and the calice floor deeper in the cardinal quadrants. Based on the
<typeStatus id="1E2A66A27F12FFF3FD1BFE94FCE5FE08" box="[679,775,363,387]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
of the type species of
<taxonomicName id="0691A3837F12FFF3FE3EFE74FCE2FE28" authority="Fedorowski, 1982" authorityName="Fedorowski" authorityYear="1982" box="[386,768,395,419]" class="Anthozoa" family="Hapsiphyllidae" genus="Allotropiochisma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FE3EFE74FDDEFE28" box="[386,572,395,419]" italics="true" pageId="11" pageNumber="12">Allotropiochisma</emphasis>
<bibRefCitation id="A500A5F17F12FFF3FDFFFE74FCE2FE28" author="Fedorowski, J." box="[579,768,395,419]" pageId="11" pageNumber="12" pagination="71 - 91" refId="ref25483" refString="Fedorowski, J. (1982). Some rugose corals from the Upper Permian of East Greenland. Rapport Gronlands Geologiske Undersogelse, 108, 71 - 91." type="journal article" year="1982">Fedorowski, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0691A3837F12FFF3FF2BFE54FD94FE48" authority="(Flugel, 1973)" baseAuthorityName="Flugel" baseAuthorityYear="1973" box="[151,630,427,451]" class="Anthozoa" family="Hapsiphyllidae" genus="Allotropiochisma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="longiseptata">
<emphasis id="F3E504127F12FFF3FF2BFE54FE3EFE48" box="[151,476,427,451]" italics="true" pageId="11" pageNumber="12">Allotropiochisma longiseptata</emphasis>
(
<bibRefCitation id="A500A5F17F12FFF3FE56FE54FD8FFE48" author="FlUgel, H. W." box="[490,621,427,451]" pageId="11" pageNumber="12" pagination="1 - 57" refId="ref26012" refString="FlUgel, H. W. (1973). Rugose Korallen aus dem oberen Perm Ost-Gronlands. Verhandlungen Der Geologischen Bundes-Anstalt., 1, 1 - 57." type="journal article" year="1973">Flügel, 1973</bibRefCitation>
)
</taxonomicName>
, this genus is sufficiently different from
<taxonomicName id="0691A3837F12FFF3FE09FE34FDD3FE68" box="[437,561,459,483]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FE09FE34FDD3FE68" box="[437,561,459,483]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
in having a narrow cardinal fossula, lacking an elongated counter septum, &quot;biform reduction&quot; of minor septa, and biform peripheralmost tabularium. In addition,
<taxonomicName id="0691A3837F12FFF3FD87FDD4FF29FDE8" authority="Hill, 1938" authorityName="Hill" authorityYear="1938" class="Anthozoa" family="Hapsiphyllidae" genus="Euryphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FD87FDD4FD2AFDC8" box="[571,712,555,579]" italics="true" pageId="11" pageNumber="12">Euryphyllum</emphasis>
<bibRefCitation id="A500A5F17F12FFF3FD64FDD4FF29FDE8" author="Hill, D." pageId="11" pageNumber="12" pagination="1 - 78" refId="ref26390" refString="Hill, D. (1938). A monograph on the Carboniferous rugose corals of Scotland. Part 1. Palaeontographical Society, London, 91, 1 - 78." type="journal article" year="1938">Hill, 1938</bibRefCitation>
</taxonomicName>
is similar. In fact,
<taxonomicName id="0691A3837F12FFF3FE19FDB4FDA8FDE8" box="[421,586,587,611]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F12FFF3FE19FDB4FDC3FDE8" box="[421,545,587,611]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
sp.
</taxonomicName>
No. 2 described from the Lower Permian of the Northern Urals (Dobrolyubova, 1936) and at least some specimens described as
<taxonomicName id="0691A3837F12FFF3FF2BFD54FEF1FD48" box="[151,275,683,707]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FF2BFD54FEF1FD48" box="[151,275,683,707]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
by
<bibRefCitation id="A500A5F17F12FFF3FE87FD53FE4DFD48" author="Wu, W. S." box="[315,431,683,707]" pageId="11" pageNumber="12" pagination="1966 - 1968" refId="ref30601" refString="Wu, W. S. (1975). The coral fossils from the Qomolangma Feng region. In: A report of scientific investigation in the Qomolangma Feng Region 1966 - 1968 (Palaontology, part 1), 83 - 128; Beijing (Science Press). (in Chinese)." type="book chapter" year="1975">Wu (1975)</bibRefCitation>
and
<bibRefCitation id="A500A5F17F12FFF3FE58FD54FDB4FD48" author="Lin, B. Y." box="[484,598,683,707]" pageId="11" pageNumber="12" pagination="69 - 181" refId="ref28059" refString="Lin, B. Y. (1983). Lower Permian stratigraphy and coral faunas from both flanks of Yarlung Zangbo River in central-southern Xizang (Tibet) Plateau. Contribution to the Geology of the Qinghai-Xizang (tibet) Plateau, 8, 69 - 181. in Chinese with English summary." type="book chapter" year="1983">Lin (1983)</bibRefCitation>
from the Lower Permian of
<collectingRegion id="035516E27F12FFF3FEA7FD34FEB7FD68" box="[283,341,715,739]" country="China" name="Xizang" pageId="11" pageNumber="12">Tibet</collectingRegion>
might belong to that genus, as well as some specimens described by
<bibRefCitation id="A500A5F17F12FFF3FE4DFD14FD92FC88" author="Guo, S." box="[497,624,747,771]" pageId="11" pageNumber="12" pagination="1 - 686" refId="ref26265" refString="Guo, S. (1980). Tetracoralla. In: Paleontological atlas of Northeast China. Paleozoic vol. 1 - 686. Beijing." type="book chapter" year="1980">Guo (1980)</bibRefCitation>
from Northeast
<collectingCountry id="B98698907F12FFF3FF74FCF4FEE8FCA8" box="[200,266,779,803]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
(
<bibRefCitation id="A500A5F17F12FFF3FEA4FCF4FDBDFCA8" author="Fedorowski, J. &amp; Bamber, E. W." box="[280,607,779,803]" pageId="11" pageNumber="12" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F12FFF3FF2BFCD4FDDEF9C8" pageId="11" pageNumber="12" type="distribution">
<paragraph id="C12ED8007F12FFF3FF2BFCD4FDDEF9C8" blockId="11.[151,775,235,1603]" pageId="11" pageNumber="12">
<emphasis id="F3E504127F12FFF3FF2BFCD4FDA8FCC8" bold="true" box="[151,586,811,835]" pageId="11" pageNumber="12">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F12FFF3FDEAFCD4FD30FCC8" box="[598,722,811,835]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FDEAFCD4FD30FCC8" box="[598,722,811,835]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
is a spatiotemporally widespread genus. It is known from the Bashkirian of the Donets Basin and the Southern Urals/ Bashkirian Mountains (
<bibRefCitation id="A500A5F17F12FFF3FE18FC74FDBCFC28" author="Kossovaya, O. L." box="[420,606,907,931]" pageId="11" pageNumber="12" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
), but was not listed from the Bashkirian of the Donets Basin by
<bibRefCitation id="A500A5F17F12FFF3FD03FC54FEC6FC68" author="Fedorowski, J." pageId="11" pageNumber="12" pagination="247 - 316" refId="ref25858" refString="Fedorowski, J. (2022). Bashkirian Rugosa (Anthozoa) from the Donets Basin (Ukraine). Part 12. Concluding considerations. Acta Geologica Polonica, 72 (3), 247 - 316." type="journal article" year="2022">Fedorowski (2022)</bibRefCitation>
. According to
<bibRefCitation id="A500A5F17F12FFF3FE79FC34FD7EFC68" author="Fedorowski, J." box="[453,668,971,995]" pageId="11" pageNumber="12" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
, it occurs there from Kasimovian to Gzhelian (C
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C
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). The genus is known from the Kasimovian of
<collectingCountry id="B98698907F12FFF3FDB3FBF3FDACFBAF" box="[527,590,1036,1060]" name="Egypt" pageId="11" pageNumber="12">Egypt</collectingCountry>
(
<bibRefCitation id="A500A5F17F12FFF3FDE0FBF4FEE8FBC8" author="Kora, M. &amp; Mansour, Y." pageId="11" pageNumber="12" pagination="597 - 614" refId="ref27107" refString="Kora, M., &amp; Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour, 1991</bibRefCitation>
and this paper) and Donets Basin, from the PennsylvanianPermian transition of the Glass Mountains, western Texas (
<taxonomicName id="0691A3837F12FFF3FE3EFB94FE0DFB08" authorityName="Böse" authorityYear="1917" box="[386,495,1131,1155]" class="Cephalopoda" family="Medlicottiidae" genus="Uddenites" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="F3E504127F12FFF3FE3EFB94FE0DFB08" box="[386,495,1131,1155]" italics="true" pageId="11" pageNumber="12">Uddenites</emphasis>
</taxonomicName>
-bearing shale member of Gaptank Formation,
<bibRefCitation id="A500A5F17F12FFF3FEC1FB74FDABFB28" author="Fedorowski, J." box="[381,585,1163,1187]" pageId="11" pageNumber="12" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
), from the upper Artinskian and lowermost Kungirian (lower Saranian) of the Central Urals (
<bibRefCitation id="A500A5F17F12FFF3FE34FB34FD9DFB68" author="Kossovaya, O. L. &amp; Kuseva, E. A. &amp; Lukin, A. E. &amp; Zhuravlev, A. V." box="[392,639,1227,1251]" pageId="11" pageNumber="12" pagination="95 - 113" refId="ref27550" refString="Kossovaya, O. L., Kuseva, E. A., Lukin, A. E., &amp; Zhuravlev, A. V. (2001). Middle Artinskian (Early Permian) ecological event: a case study of the Urals and northern Timan. Proceedings of the Estonian Academy of Science. Geology, 50 (2), 95 - 113." type="journal article" year="2001">Kossovaya et al., 2001</bibRefCitation>
;
<bibRefCitation id="A500A5F17F12FFF3FD33FB33FF29FA88" author="Kossovaya, O. L." pageId="11" pageNumber="12" pagination="383 - 405" refId="ref27441" refString="Kossovaya, O. L. (2007). Ecological aspects of upper Carboniferous - lower Permian ' Cyathaxonia Fauna' taxonomical diversity (the Urals). In B. Hubmann, &amp; W. E. Piller (Eds.), Fossil Corals and Sponges. Proceedings of the 9 th International Symposium on Fossil Cnidaria and Porifera. - Osterreichische Akademie der Wissenschaften, Schriftenreihe der Erdwissenschaftlichen Kommission 17, 383 - 405." type="journal article" year="2007">Kossovaya, 2007</bibRefCitation>
,
<figureCitation id="59AAC4857F12FFF3FF60FB14FEFEFA88" box="[220,284,1259,1283]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
; Shoshkina, 1925), the Sakmarian of Spitsbergen (Chwieduk, 2013) and from the Wordian (Middle Guadalupian) of the Canadian Arctic Archipelago (
<bibRefCitation id="A500A5F17F12FFF3FF22FAB4FE0CFAE8" author="Fedorowski, J. &amp; Bamber, E. W." box="[158,494,1355,1379]" pageId="11" pageNumber="12" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
). The existence of
<taxonomicName id="0691A3837F12FFF3FD78FAB3FF38FA08" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FD78FAB3FF38FA08" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
in Tibet (
<bibRefCitation id="A500A5F17F12FFF3FEECFA94FE5FFA08" author="Wu, W. S." box="[336,445,1387,1411]" pageId="11" pageNumber="12" pagination="1966 - 1968" refId="ref30601" refString="Wu, W. S. (1975). The coral fossils from the Qomolangma Feng region. In: A report of scientific investigation in the Qomolangma Feng Region 1966 - 1968 (Palaontology, part 1), 83 - 128; Beijing (Science Press). (in Chinese)." type="book chapter" year="1975">Wu, 1975</bibRefCitation>
) was rejected by
<bibRefCitation id="A500A5F17F12FFF3FD35FA94FF0BFA28" author="Fedorowski, J." pageId="11" pageNumber="12" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
, but
<bibRefCitation id="A500A5F17F12FFF3FEA3FA73FE13FA28" author="Fang, R. S. &amp; Fan, J. C." box="[287,497,1419,1444]" pageId="11" pageNumber="12" pagination="189 - 190" refId="ref25287" refString="Fang, R. S., &amp; Fan, J. C. (1994). On the cold water coral Lytvolasma fauna in Baoshan-Pengchong region Yunnan. Geology, 1, 189 - 190. in Chinese with English abstract." type="book chapter" year="1994">Fang and Fan (1994</bibRefCitation>
; not seen) again reported a “
<taxonomicName id="0691A3837F12FFF3FF07FA54FED5FA48" box="[187,311,1451,1475]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FF07FA54FED5FA48" box="[187,311,1451,1475]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
fauna” in SW Yunnan (BaoshanPengchong). To us, the presence of
<taxonomicName id="0691A3837F12FFF3FE4AFA34FD90FA68" box="[502,626,1483,1507]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F12FFF3FE4AFA34FD90FA68" box="[502,626,1483,1507]" italics="true" pageId="11" pageNumber="12">Lytvolasma</emphasis>
</taxonomicName>
in Tibet and the Baoshan Block, southwestern Yunnan, i.e., in terranes originally bordering the northern margin of Gondwana, remains unsure and needs revaluation.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,400 @@
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<paragraph id="C12ED8007F14FFF5FC91FD52FB91FD4D" blockId="13.[813,1139,685,710]" box="[813,1139,685,710]" pageId="13" pageNumber="14">
<heading id="9A666F6C7F14FFF5FC91FD52FB91FD4D" bold="true" box="[813,1139,685,710]" fontSize="10" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="F3E504127F14FFF5FC91FD52FB91FD4D" bold="true" box="[813,1139,685,710]" pageId="13" pageNumber="14">
<taxonomicName id="0691A3837F14FFF5FC91FD52FBD3FD4E" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[813,1073,685,709]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F14FFF5FC91FD52FBD3FD4E" bold="true" box="[813,1073,685,709]" italics="true" pageId="13" pageNumber="14">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F14FFF5FB8BFD51FB91FD4D" box="[1079,1139,686,710]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FC91FD31FBC6FCAD" pageId="13" pageNumber="14" type="description">
<paragraph id="C12ED8007F14FFF5FC91FD31FBBFFD6D" blockId="13.[813,1437,718,1894]" box="[813,1117,718,742]" pageId="13" pageNumber="14">
(
<figureCitation id="59AAC4857F14FFF5FC88FD31FC64FD6D" box="[820,902,718,742]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FC3AFD31FC56FD6D" box="[902,948,718,742]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FC08FD31FC23FD6D" box="[948,961,718,742]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
, B
<figureCitation id="59AAC4857F14FFF5FC62FD31FBEBFD6D" box="[990,1033,718,742]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1B</figureCitation>
<figureCitation id="59AAC4857F14FFF5FBB5FD31FBF4FD6D" box="[1033,1046,718,742]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
,
<figureCitation id="59AAC4857F14FFF5FB9DFD31FBB2FD6D" box="[1057,1104,718,742]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="13" pageNumber="14">12A</figureCitation>
).
</paragraph>
<paragraph id="C12ED8007F14FFF5FC91FD11FBC6FCAD" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
1991
<taxonomicName id="0691A3837F14FFF5FCCCFD11FAF2FC8D" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[880,1296,750,774]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F14FFF5FCCCFD11FBD8FC8D" box="[880,1082,750,774]" italics="true" pageId="13" pageNumber="14">Lytvolasma aucta</emphasis>
Fedorowski, 1987
</taxonomicName>
.- Kora and Mansour, figs. 2 c1, c2.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FC91FCD1FAA1FC0D" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="C12ED8007F14FFF5FC91FCD1FBB4FCED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
1991
<taxonomicName id="0691A3837F14FFF5FCDBFCD1FAB3FCCD" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" box="[871,1361,814,838]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="bellicostatum">
<emphasis id="F3E504127F14FFF5FCDBFCD1FBE0FCCD" box="[871,1026,814,838]" italics="true" pageId="13" pageNumber="14">Bradyphyllum</emphasis>
cf.
<emphasis id="F3E504127F14FFF5FB94FCD1FB58FCCD" box="[1064,1210,814,838]" italics="true" pageId="13" pageNumber="14">bellicostatum</emphasis>
<bibRefCitation id="A500A5F17F14FFF5FB7CFCD1FAB3FCCD" author="Grabau, A. W." box="[1216,1361,814,838]" pageId="13" pageNumber="14" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
.- Kora and Mansour, figs. 2 b1, b2.
</paragraph>
<paragraph id="C12ED8007F14FFF5FC91FC91FAA1FC0D" blockId="13.[813,1437,718,1894]" box="[813,1347,878,902]" pageId="13" pageNumber="14">
2001
<taxonomicName id="0691A3837F14FFF5FCDBFC91FC01FC0D" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[871,995,878,902]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F14FFF5FCDBFC91FC01FC0D" box="[871,995,878,902]" italics="true" pageId="13" pageNumber="14">Lytvolasma</emphasis>
</taxonomicName>
—Fedorowski and Bamber, p 48.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FC91FC71FB4AFC4D" pageId="13" pageNumber="14" type="description">
<paragraph id="C12ED8007F14FFF5FC91FC71FB4AFC4D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91FC71FBB6FC2D" bold="true" box="[813,1108,910,934]" pageId="13" pageNumber="14">Derivation of the name:</emphasis>
From the Greek “
<emphasis id="F3E504127F14FFF5FA8EFC70FA87FC2D" box="[1330,1381,911,934]" italics="true" pageId="13" pageNumber="14">para</emphasis>
” for resemblance to the species
<taxonomicName id="0691A3837F14FFF5FBE9FC51FB7FFC4D" authorityName="Fedorowski" authorityYear="1987" box="[1109,1181,942,966]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F14FFF5FBE3FC50FB7FFC4D" box="[1119,1181,943,966]" italics="true" pageId="13" pageNumber="14">aucta</emphasis>
</taxonomicName>
”.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FC91FC31FA7AFA2D" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="C12ED8007F14FFF5FC91FC31FB1FFB8D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<materialsCitation id="71F9D25D7F14FFF5FC91FC31FB1FFB8D" pageId="13" pageNumber="14" specimenCode="RAh 54" specimenCount="1" typeStatus="holotype">
<emphasis id="F3E504127F14FFF5FC91FC31FC7DFC6D" bold="true" box="[813,927,974,998]" pageId="13" pageNumber="14">
<typeStatus id="1E2A66A27F14FFF5FC91FC31FC79FC6D" box="[813,923,974,998]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="D79713897F14FFF5FC1AFC31FBF3FC6D" box="[934,1041,974,998]" pageId="13" pageNumber="14" type="generic">Specimen</specimenCount>
<specimenCode id="9137707B7F14FFF5FBA5FC31FB8CFC6D" box="[1049,1134,974,998]" pageId="13" pageNumber="14">RAh 54</specimenCode>
, illustrated in
<figureCitation id="59AAC4857F14FFF5FAB7FC31FABFFC6D" box="[1291,1373,974,998]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FAE1FC31FA6EFC6D" box="[1373,1420,974,998]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FA30FC31FA7BFC6D" box="[1420,1433,974,998]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
. Four transverse thin-sections are available.
</materialsCitation>
</paragraph>
<paragraph id="C12ED8007F14FFF5FC91FBF1FAFCFB6D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91FBF1FC75FBAD" bold="true" box="[813,919,1038,1062]" pageId="13" pageNumber="14">Material:</emphasis>
In addition to the
<typeStatus id="1E2A66A27F14FFF5FBC4FBF1FB38FBAD" box="[1144,1242,1038,1062]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, four moderately preserved corallites with compressed calices and occasionally preserved apical part (RAh 19, RAh 74 and RAh 81); collected from the basal shales of the lower member of the Aheimer Formation. Eight transverse thin-sections are available. They are collected from the basal shales of the lower member of the Aheimer Formation.
</paragraph>
<paragraph id="C12ED8007F14FFF5FC91FB11FC91FAED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91FB11FC2EFA8D" bold="true" box="[813,972,1262,1286]" pageId="13" pageNumber="14">Type locality:</emphasis>
<materialsCitation id="71F9D25D7F14FFF5FC68FB11FC8DFAED" country="Egypt" county="Gulf" latitude="29.4725" location="Northern Galala" longLatPrecision="20" longitude="32.460556" municipality="northern Eastern Desert" pageId="13" pageNumber="14" specimenCount="1" stateProvince="As Suways">
Eastern cliffs of the Northern Galala plateau (Lat. 29° 28
<emphasis id="F3E504127F14FFF5FC61FAF2FC0AFAAD" box="[989,1000,1293,1318]" italics="true" pageId="13" pageNumber="14"></emphasis>
21
<emphasis id="F3E504127F14FFF5FBB5FAF2FBFBFAAD" box="[1033,1049,1293,1318]" italics="true" pageId="13" pageNumber="14"></emphasis>
N and Long. 32° 27
<emphasis id="F3E504127F14FFF5FB4CFAF2FB19FAAD" box="[1264,1275,1293,1318]" italics="true" pageId="13" pageNumber="14"></emphasis>
38
<emphasis id="F3E504127F14FFF5FAA7FAF2FAC9FAAD" box="[1307,1323,1293,1318]" italics="true" pageId="13" pageNumber="14"></emphasis>
E), northern Eastern Desert, western side of the Gulf of Suez, Egypt
</materialsCitation>
.
</paragraph>
<paragraph id="C12ED8007F14FFF5FC91FA91FA7AFA2D" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91FA91FC32FA0D" bold="true" box="[813,976,1390,1414]" pageId="13" pageNumber="14">Type horizon:</emphasis>
Basal shales of the lower member of the Aheimer Formation; Upper Pennsylvanian (Kasimovian).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF5FC91FA51FC9DF9ED" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="C12ED8007F14FFF5FC91FA51FC9DF9ED" blockId="13.[813,1437,718,1894]" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91FA51FC44FA4D" bold="true" box="[813,934,1454,1478]" pageId="13" pageNumber="14">Diagnosis:</emphasis>
<taxonomicName id="0691A3837F14FFF5FC00FA51FBDAFA4D" box="[956,1080,1454,1478]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F14FFF5FC00FA51FBDAFA4D" box="[956,1080,1454,1478]" italics="true" pageId="13" pageNumber="14">Lytvolasma</emphasis>
</taxonomicName>
having maximum n:d ratio 26:11.5. Major septa frequently thickened in all quadrants to make a stereocolumn that opens in the cardinal quadrants, that changes to half aulos in the counter quadrants with maturity.
<taxonomicName id="0691A3837F14FFF5FBACF9D1FB8CF9CD" box="[1040,1134,1582,1606]" genus="Cardinal" pageId="13" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum is short within the fossula.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F14FFF7FC91F991FDE5FB88" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="C12ED8007F14FFF6FC91F991FD66F956" blockId="13.[813,1437,718,1894]" lastBlockId="14.[151,775,1702,1918]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="F3E504127F14FFF5FC91F991FC5FF90D" bold="true" box="[813,957,1646,1670]" pageId="13" pageNumber="14">Description:</emphasis>
External characters: Corallites are small, up to
<quantity id="066975E57F14FFF5FCCBF971FC51F92E" box="[887,947,1678,1702]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="cm" value="3.0">3 cm</quantity>
long. The external wall is ornamented by septal furrows and fine concentric growth lines in the
<typeStatus id="1E2A66A27F14FFF5FC91F931FC6FF96D" box="[813,909,1742,1766]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
specimen (
<figureCitation id="59AAC4857F14FFF5FBB5F931FBBEF96D" box="[1033,1116,1742,1766]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FBE1F931FB85F96D" box="[1117,1127,1742,1766]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
), in other
<typeStatus id="1E2A66A27F14FFF5FB6BF931FAA2F96D" box="[1239,1344,1742,1766]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
the corallites are enclosed in mud and do not show any septal furrows (
<figureCitation id="59AAC4857F14FFF5FC2DF8F1FC01F8AD" box="[913,995,1806,1830]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8B</figureCitation>
<figureCitation id="59AAC4857F14FFF5FC5FF8F1FC0FF8AD" box="[995,1005,1806,1830]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
). Calices either eroded or crushed and compressed (
<figureCitation id="59AAC4857F14FFF5FC02F8D1FBF6F8CD" box="[958,1044,1838,1862]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F14FFF5FBA8F8D1FBFCF8CD" box="[1044,1054,1838,1862]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="13" pageNumber="14">1</figureCitation>
). Two of the specimens with their apexes preserved. Two shallow constrictions are visible below the calicinal part in the
<typeStatus id="1E2A66A27F17FFF6FE53F959FDADF935" box="[495,591,1702,1726]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
specimen. Constrictions are also observed in the apical part.
</paragraph>
<caption id="95EE88887F17FFF6FF1BFA48FEBEF9DA" ID-DOI="http://doi.org/10.5281/zenodo.12003275" ID-Zenodo-Dep="12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15" startId="14.[167,198,1463,1483]" targetBox="[297,1298,274,1448]" targetPageId="14" targetType="figure">
<paragraph id="C12ED8007F17FFF6FF1BFA48FEBEF9DA" blockId="14.[167,1413,1463,1617]" pageId="14" pageNumber="15">
<emphasis id="F3E504127F17FFF6FF1BFA48FF10FA40" bold="true" box="[167,242,1463,1483]" pageId="14" pageNumber="15">Fig. 8 A</emphasis>
<taxonomicName id="0691A3837F17FFF6FF4BFA48FE4FFA40" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[247,429,1463,1483]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F17FFF6FF4BFA48FE4FFA40" box="[247,429,1463,1483]" italics="true" pageId="14" pageNumber="15">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F17FFF6FE0EFA48FE3DFA40" box="[434,479,1463,1483]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="F3E504127F17FFF6FE5EFA48FE19FA40" bold="true" box="[482,507,1463,1483]" pageId="14" pageNumber="15">A1</emphasis>
Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.
<emphasis id="F3E504127F17FFF6FE91FA2DFEA4FA6D" bold="true" box="[301,326,1490,1510]" pageId="14" pageNumber="15">A2</emphasis>
Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.
<emphasis id="F3E504127F17FFF6FB60FA2DFB17FA6D" bold="true" box="[1244,1269,1490,1510]" pageId="14" pageNumber="15">A3</emphasis>
Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends.
<emphasis id="F3E504127F17FFF6FBE0FA12FB97F98A" bold="true" box="[1116,1141,1517,1537]" pageId="14" pageNumber="15">A4</emphasis>
Laterally compressed section in the calice base,showing a deformed fossula.
<emphasis id="F3E504127F17FFF6FD86F9F8FDB0F990" bold="true" box="[570,594,1543,1563]" pageId="14" pageNumber="15">A5</emphasis>
Transverse thin-section in the compressed calice, with calcareous mud filling.
<emphasis id="F3E504127F17FFF6FB5BF9F8FB1CF990" bold="true" box="[1255,1278,1543,1563]" pageId="14" pageNumber="15">B1</emphasis>
Lateral external view of the paratype specimen (RAh 19).
<emphasis id="F3E504127F17FFF6FDBFF9DDFDDFF9BD" bold="true" box="[515,573,1570,1590]" pageId="14" pageNumber="15">B2B5</emphasis>
Successive transverse thin-sections from the immature part near the apex (
<emphasis id="F3E504127F17FFF6FB07F9DDFB31F9BD" bold="true" box="[1211,1235,1570,1590]" pageId="14" pageNumber="15">B2</emphasis>
) to the last mature part in the calice (
<emphasis id="F3E504127F17FFF6FE83F9C2FEB5F9DA" bold="true" box="[319,343,1597,1617]" pageId="14" pageNumber="15">B5</emphasis>
)
</paragraph>
</caption>
<paragraph id="C12ED8007F17FFF6FF11F919FEE3F895" blockId="14.[151,775,1702,1918]" pageId="14" pageNumber="15">
The following description refers mostly to the
<typeStatus id="1E2A66A27F17FFF6FD1BF919FCE5F975" box="[679,775,1766,1790]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(RAh 54).
</paragraph>
<paragraph id="C12ED8007F17FFF7FF11F8D9FD9BFE28" blockId="14.[151,775,1702,1918]" lastBlockId="15.[151,776,235,1475]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Internal characters: The ontogenetically earliest preserved part has 18 septa in
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of corallite diameter (
<figureCitation id="59AAC4857F17FFF6FF6EF899FEC7F8F5" box="[210,293,1894,1918]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F17FFF6FE9AF899FED6F8F5" box="[294,308,1894,1918]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="14" pageNumber="15">2</figureCitation>
). At this stage major septa show irregular zaphrentoid arrangement. They are thick, among them only the cardinal is thinner, whereas the counter is thicker with rhopaloid ending. The cardinal fossula is noticeable with long cardinal septum that fuses with the other major septa making a dense stereocolumn in the centre, that opens in the cardinal quadrants. Minor septa are absent in this stage (see also
<figureCitation id="59AAC4857F17FFF6FB99F899FB96F8F5" box="[1061,1140,1894,1918]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="14" pageNumber="15">Fig. 8B</figureCitation>
<figureCitation id="59AAC4857F17FFF6FBC8F899FB60F8F5" box="[1140,1154,1894,1918]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="14" pageNumber="15">2</figureCitation>
). The subsequent section made at a diameter of
<quantity id="066975E57F16FFF7FE26FF13FE3FFE88" box="[410,477,236,259]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="7.0">7 mm</quantity>
shows 23 septa which are radially arranged (
<figureCitation id="59AAC4857F16FFF7FEDDFEF4FE56FEA8" box="[353,436,267,291]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F16FFF7FE09FEF4FE23FEA8" box="[437,449,267,291]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3</figureCitation>
). Their inner ends are rhopaloid and form a stereocolumn in the corallite centre. The cardinal fossula is slightly key-hole to subtriangular in shape and open adaxially (
<figureCitation id="59AAC4857F16FFF7FE09FE94FDE4FE08" box="[437,518,363,387]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F16FFF7FDBBFE94FDBEFE08" box="[519,604,363,387]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3, B3B</figureCitation>
<figureCitation id="59AAC4857F16FFF7FDE7FE94FD8BFE08" box="[603,617,363,387]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
), with the cardinal septum thinner and slightly shortened.
</paragraph>
<paragraph id="C12ED8007F16FFF7FF11FE53FDE5FB88" blockId="15.[151,776,235,1475]" pageId="15" pageNumber="16">
The subsequent mature section is made beneath the calice floor (
<figureCitation id="59AAC4857F16FFF7FEA1FE34FE8CFE68" box="[285,366,459,483]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F16FFF7FED2FE34FE9EFE68" box="[366,380,459,483]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
). The corallite in this part is laterally compressed and almost half of the septa are broken and deformed. At this stage of maturity, the deformed cardinal fossula is bordered by a half aulos in counter quadrants. Major septa are almost radially arranged and are
<quantity id="066975E57F16FFF7FD51FDB4FF4FFD08" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="16" unit="in" value="24.0">24 in</quantity>
number.
<taxonomicName id="0691A3837F16FFF7FEA6FD94FE9AFD08" box="[282,376,619,643]" genus="Cardinal" pageId="15" pageNumber="8" rank="genus">Cardinal</taxonomicName>
septum is short within the fossula. Minor septa are rudimentary. They appear as small triangular spikes along the relatively thin epitheca. The last section within the compressed calice (
<figureCitation id="59AAC4857F16FFF7FDF9FD34FD7AFD68" box="[581,664,715,739]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F16FFF7FD24FD34FD47FD68" box="[664,677,715,739]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
) is filled with mud and shows short major septa and underdeveloped minor septa. At a diameter of 11.5 mm of the
<typeStatus id="1E2A66A27F16FFF7FD71FCF3FF24FCCF" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
specimen (RAh 19), more or less at the base of the calice (
<figureCitation id="59AAC4857F16FFF7FF5BFCB4FEDAFCE8" box="[231,312,843,867]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
<figureCitation id="59AAC4857F16FFF7FE85FCB4FEA7FCE8" box="[313,325,843,867]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
) the septal ends are free leaving a free axial area with a short cardinal septum and a relatively longer counter septum than the adjacent septa. The counter septum, always a little longer and/or slightly thicker than other major septa, a character clearly evident in the
<typeStatus id="1E2A66A27F16FFF7FF2BFC13FF14FB8F" box="[151,246,1004,1028]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
specimen (
<figureCitation id="59AAC4857F16FFF7FECEFC14FE22FB88" box="[370,448,1003,1027]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
<figureCitation id="59AAC4857F16FFF7FE7CFC14FE0CFB88" box="[448,494,1003,1027]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3B</figureCitation>
<figureCitation id="59AAC4857F16FFF7FE52FC14FE19FB88" box="[494,507,1003,1027]" captionStart="Fig" captionStartId="6.[167,198,1874,1894]" captionTargetBox="[168,1421,1179,1861]" captionTargetId="figure-436@6.[166,1421,1179,1861]" captionTargetPageId="6" captionText="Fig. 5 Internal morphological characters of the non-dissepimented rugose corals referred to in the systematic descriptions, after Denayer and Hoşgör (2014). A Transverse section. B Longitudinal section" figureDoi="http://doi.org/10.5281/zenodo.12003232" httpUri="https://zenodo.org/record/12003232/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFF7FF2BFBF4FAFCFCA8" pageId="15" pageNumber="16" type="discussion">
<paragraph id="C12ED8007F16FFF7FF2BFBF4FEEDFA48" blockId="15.[151,776,235,1475]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FF2BFBF4FEFFFBA8" bold="true" box="[151,285,1035,1059]" pageId="15" pageNumber="16">Discussion:</emphasis>
<taxonomicName id="0691A3837F16FFF7FE99FBF4FDF9FBA8" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[293,539,1035,1059]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F16FFF7FE99FBF4FDF9FBA8" box="[293,539,1035,1059]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F16FFF7FD9FFBF3FDBDFBA8" box="[547,607,1036,1059]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
was illustrated by
<bibRefCitation id="A500A5F17F16FFF7FF04FBD3FE2EFBC8" author="Kora, M. &amp; Mansour, Y." box="[184,460,1067,1091]" pageId="15" pageNumber="16" pagination="597 - 614" refId="ref27107" refString="Kora, M., &amp; Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991</bibRefCitation>
;
<figureCitation id="59AAC4857F16FFF7FE66FBD4FDCEFBC8" box="[474,556,1067,1091]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">Fig. 2 c</figureCitation>
<figureCitation id="59AAC4857F16FFF7FD90FBD4FDD5FBC8" box="[556,567,1067,1091]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="15" pageNumber="16">1</figureCitation>
, c
<figureCitation id="59AAC4857F16FFF7FDECFBD4FDBDFBC8" box="[592,607,1067,1091]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">2</figureCitation>
). Based on one transverse section in the mature part of the corallite and a longitudinal section, they assigned their specimen to
<taxonomicName id="0691A3837F16FFF7FD4EFB93FE43FB28" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F16FFF7FD4EFB93FF37FB28" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
Fedorowski, 1987
</taxonomicName>
. Due to the lack of description and additional illustrations of younger ontogenetic stages
<bibRefCitation id="A500A5F17F16FFF7FF2BFB34FE17FB68" author="Fedorowski, J. &amp; Bamber, E. W." box="[151,501,1227,1251]" pageId="15" pageNumber="16" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber (2001)</bibRefCitation>
accepted this identification only at a generic level. We also include
<taxonomicName id="0691A3837F16FFF7FD32FB14FDFAFAA8" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="bellicostatum">
<emphasis id="F3E504127F16FFF7FD32FB14FF20FAA8" italics="true" pageId="15" pageNumber="16">Bradyphyllum</emphasis>
cf.
<emphasis id="F3E504127F16FFF7FF57FAF4FE9FFAA8" box="[235,381,1291,1315]" italics="true" pageId="15" pageNumber="16">bellicostatum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FE39FAF4FDFAFAA8" author="Grabau, A. W." box="[389,536,1291,1315]" pageId="15" pageNumber="16" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
briefly described and figured by
<bibRefCitation id="A500A5F17F16FFF7FEB3FAD3FDC4FAC8" author="Kora, M. &amp; Mansour, Y." box="[271,550,1323,1347]" pageId="15" pageNumber="16" pagination="597 - 614" refId="ref27107" refString="Kora, M., &amp; Mansour, Y. (1991). Late Carboniferous solitary rugose corals from the western side of the Gulf of Suez, Egypt. Neues Jahrbuch Fur Geologie Und Palaontologie Monatshefte, Stuttgart, 10, 597 - 614." type="journal article" year="1991">Kora and Mansour (1991</bibRefCitation>
, p. 599,
<figureCitation id="59AAC4857F16FFF7FD3BFAD4FD3BFAC8" box="[647,729,1323,1347]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">Fig. 2 c</figureCitation>
<figureCitation id="59AAC4857F16FFF7FD65FAD4FD12FAC8" box="[729,752,1323,1347]" captionStart="Fig" captionStartId="1.[167,198,1197,1217]" captionTargetBox="[170,1418,237,1183]" captionTargetId="figure-377@1.[169,1419,236,1184]" captionTargetPageId="1" captionText="Fig. 1 Geological map of the eastern cliffs of the Northern Galala Plateau, modified after Abdallah and El Adindani (1965)" figureDoi="http://doi.org/10.5281/zenodo.12003182" httpUri="https://zenodo.org/record/12003182/files/figure.png" pageId="15" pageNumber="16">1c</figureCitation>
<figureCitation id="59AAC4857F16FFF7FD4CFAD4FD1DFAC8" box="[752,767,1323,1347]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="15" pageNumber="16">2</figureCitation>
) in our new species. Additional transverse thin-sections from the youngest stages clearly confirm the difference of the Egyptian specimens from
<taxonomicName id="0691A3837F16FFF7FE44FA73FDB3FA28" authorityName="Fedorowski" authorityYear="1987" box="[504,593,1420,1443]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F16FFF7FE44FA73FDB3FA28" box="[504,593,1420,1443]" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
</taxonomicName>
by the following characters:
</paragraph>
<paragraph id="C12ED8007F16FFF7FF11FA12FEDFF9EE" blockId="15.[173,775,1517,1796]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FF70FA12FE20F98F" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[204,450,1517,1541]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F16FFF7FF70FA12FE20F98F" box="[204,450,1517,1541]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F16FFF7FE73FA12FDECF98F" box="[463,526,1517,1540]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
has higher n:d ratios than the
<typeStatus id="1E2A66A27F16FFF7FE8EF9F3FE72F9AF" box="[306,400,1548,1572]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0691A3837F16FFF7FE05F9F2FDF1F9AF" authorityName="Fedorowski" authorityYear="1987" box="[441,531,1549,1572]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F16FFF7FE05F9F2FE25F9AF" box="[441,455,1549,1572]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis id="F3E504127F16FFF7FE6AF9F2FDF1F9AF" box="[470,531,1549,1572]" italics="true" pageId="15" pageNumber="16">aucta</emphasis>
</taxonomicName>
(
<figureCitation id="59AAC4857F16FFF7FD98F9F3FD66F9AE" box="[548,644,1548,1573]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
). Nevertheless,
<taxonomicName id="0691A3837F16FFF7FEBEF9D2FEBEF9CF" authorityName="Fedorowski" authorityYear="1987" box="[258,348,1581,1604]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F16FFF7FEBEF9D2FEF2F9CF" box="[258,272,1581,1604]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis id="F3E504127F16FFF7FEA3F9D2FEBEF9CF" box="[287,348,1581,1604]" italics="true" pageId="15" pageNumber="16">aucta</emphasis>
</taxonomicName>
<typeStatus id="1E2A66A27F16FFF7FED9F9D2FE2EF9CE" box="[357,460,1581,1605]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
can reach a higher n:d ratio up to 22:9.
</paragraph>
<paragraph id="C12ED8007F16FFF7FF11F992FE7AF88F" blockId="15.[173,775,1517,1796]" pageId="15" pageNumber="16">
• The septal ends in the Egyptian species are thickened in contiguity to form a stereocolumn that is open towards the cardinal fossula, whereas in
<taxonomicName id="0691A3837F16FFF7FD3CF952FD3AF94F" authorityName="Fedorowski" authorityYear="1987" box="[640,728,1709,1732]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="aucta">
<emphasis id="F3E504127F16FFF7FD3CF952FD3AF94F" box="[640,728,1709,1732]" italics="true" pageId="15" pageNumber="16">L. aucta</emphasis>
</taxonomicName>
, the septal ends are free or form a half aulos only in the counter quadrants.
</paragraph>
<paragraph id="C12ED8007F16FFF7FF2BF8D1FAAAFE28" blockId="15.[151,775,1838,1894]" lastBlockId="15.[813,1437,235,803]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FF2BF8D1FE72F8CD" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[151,400,1838,1862]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="paraaucta" status="sp. nov.">
<emphasis id="F3E504127F16FFF7FF2BF8D1FE72F8CD" box="[151,400,1838,1862]" italics="true" pageId="15" pageNumber="16">Lytvolasma paraaucta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F16FFF7FE20F8D1FE38F8CE" box="[412,474,1838,1861]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="0691A3837F16FFF7FDCAF8D0FE01F8ED" authority="Fedorowski and Bamber, 2001" authorityName="Fedorowski and Bamber" authorityYear="2001" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F16FFF7FDCAF8D0FCE5F8CD" box="[630,775,1838,1862]" italics="true" pageId="15" pageNumber="16">L. canadense</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FF2BF8B1FE01F8ED" author="Fedorowski, J. &amp; Bamber, E. W." box="[151,483,1870,1894]" pageId="15" pageNumber="16" pagination="31 - 79" refId="ref25898" refString="Fedorowski, J., &amp; Bamber, E. W. (2001). Guadalupian (Middle Permian) solitary rugose corals from the Degerbols and Trold Fiord formations, Ellesmere and Melville islands, Canadian Arctic Archipelago. Acta Geologica Polonica, 51 (1), 31 - 79." type="journal article" year="2001">Fedorowski and Bamber, 2001</bibRefCitation>
</taxonomicName>
from the Middle Permian of the Canadian Arctic Archipelago, in having an unremarkable counter septum, which is elongated in
<taxonomicName id="0691A3837F16FFF7FA3BFEF3FC7CFEC8" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F16FFF7FA3BFEF3FC7CFEC8" italics="true" pageId="15" pageNumber="16">L. canadense</emphasis>
</taxonomicName>
in the calice and below its bottom. It also slightly differs by its n:d ratio in the ephebic stage near the calice floor (24:12? vs. 26:12.6;
<figureCitation id="59AAC4857F16FFF7FB06FE94FAFEFE08" box="[1210,1308,363,387]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
). The same case is for the Egyptian
<taxonomicName id="0691A3837F16FFF7FB92FE74FAA6FE28" box="[1070,1348,395,419]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="canadense">
<emphasis id="F3E504127F16FFF7FB92FE74FB48FE28" box="[1070,1194,395,419]" italics="true" pageId="15" pageNumber="16">Lytvolasma</emphasis>
cf.
<emphasis id="F3E504127F16FFF7FB6CFE74FAA6FE28" box="[1232,1348,395,419]" italics="true" pageId="15" pageNumber="16">canadense</emphasis>
</taxonomicName>
<emphasis id="F3E504127F16FFF7FAF8FE74FAAAFE28" box="[1348,1352,395,419]" italics="true" pageId="15" pageNumber="16">.</emphasis>
</paragraph>
<paragraph id="C12ED8007F16FFF7FC91FE53FAFCFCA8" blockId="15.[813,1437,235,803]" pageId="15" pageNumber="16">
In relation to
<taxonomicName id="0691A3837F16FFF7FC6EFE53FAC2FE48" authority="Soshkina, 1925" authorityName="Soshkina" authorityYear="1925" box="[978,1312,427,451]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">
<emphasis id="F3E504127F16FFF7FC6EFE53FB88FE48" box="[978,1130,428,451]" italics="true" pageId="15" pageNumber="16">L. asymetrica</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FBCAFE54FAC2FE48" author="Soshkina, E. D." box="[1142,1312,427,451]" pageId="15" pageNumber="16" pagination="76 - 104" refId="ref29330" refString="Soshkina, E. D. (1925). Les coraux du Permien inferieur (etage d'Artinsk) du versant occidental de l'Oural. Bulletin De La Societe Des Naturalistes De Moscou, Section Geologique, 33, 76 - 104." type="journal article" year="1925">Soshkina, 1925</bibRefCitation>
</taxonomicName>
, described from the Early Permian (Sakmarian) of Spitsbergen by Chwieduk (2013), the current new species has slightly similar n:d ratio in the mature stage below the calice (24:12? vs. 2326:
<quantity id="066975E57F16FFF7FC4DFDD4FBA6FDC8" box="[1009,1092,555,579]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.556" metricValueMin="2.032" pageId="15" pageNumber="16" unit="in" value="11.0" valueMax="14.0" valueMin="8.0">814 in</quantity>
<emphasis id="F3E504127F16FFF7FBF7FDD3FB04FDC8" box="[1099,1254,556,579]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FBF7FDD3FB3CFDC8" authorityName="Soshkina" authorityYear="1925" box="[1099,1246,556,579]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">L. asymetrica</taxonomicName>
;
</emphasis>
<figureCitation id="59AAC4857F16FFF7FB51FDD4FAAEFDC8" box="[1261,1356,555,579]" captionStart="Fig" captionStartId="22.[167,198,1287,1307]" captionTargetBox="[286,1300,248,1266]" captionTargetId="figure-285@22.[283,1303,236,1274]" captionTargetPageId="22" captionText="Fig. 12 Relation between number of major septa and coralla alar diameters for: A Lytvolasma paraaucta n. sp.,L aucta Fedorowski,1987, L. asymetrica Soshkina, 1925 described by Chwieduk (2013), L. canadense Fedorowski and Bamber, 2001 and the current studied L. cf. canadense Fedorowski and Bamber, 2001. B Monophyllum galalaensis n. sp. and M. parvum Fomichev, 1953. C Rotiphyllum exile de Groot, 1963. D Zaphrentites cf. parallela (Carruthers,1910) and Zaphrentites sp" figureDoi="http://doi.org/10.5281/zenodo.12003322" httpUri="https://zenodo.org/record/12003322/files/figure.png" pageId="15" pageNumber="16">Fig. 12A</figureCitation>
). However, in
<taxonomicName id="0691A3837F16FFF7FC39FDB3FBFBFDE8" authorityName="Soshkina" authorityYear="1925" box="[901,1049,588,611]" class="Anthozoa" family="Antiphyllidae" genus="Lytvolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="asymetrica">
<emphasis id="F3E504127F16FFF7FC39FDB3FC71FDE8" box="[901,915,588,611]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis id="F3E504127F16FFF7FC1EFDB3FBFBFDE8" box="[930,1049,588,611]" italics="true" pageId="15" pageNumber="16">asymetrica</emphasis>
</taxonomicName>
the inner ends of major septa join to form a quite irregular, incomplete aulos that opens on the alar septum sides throughout most of ontogeny; whereas in the current species a stereocolumn is developed through most of its ontogeny and in the mature stage, the thickened inner ends of the septa form an aulos that opens on the cardinal side (
<figureCitation id="59AAC4857F16FFF7FB3BFCF4FB3AFCA8" box="[1159,1240,779,803]" captionStart="Fig" captionStartId="14.[167,198,1463,1483]" captionTargetBox="[297,1298,274,1448]" captionTargetId="figure-138@14.[283,1303,236,1450]" captionTargetPageId="14" captionText="Fig. 8 A Lytvolasma paraaucta n. sp. A1 Lateral external view of the holotype specimen (RAh 54). Note that, calice is compressed due to compaction.A2 Transverse thin-section in the immature part, showing major septa with irregular zaphrentoid arrangement.A3 Transverse thin section in the early mature part of the corallite, showing radially arranged major septa with rhopaloid ends. A4 Laterally compressed section in the calice base,showing a deformed fossula. A5 Transverse thin-section in the compressed calice, with calcareous mud filling. B1 Lateral external view of the paratype specimen (RAh 19). B2B5 Successive transverse thin-sections from the immature part near the apex (B2) to the last mature part in the calice (B5)" figureDoi="http://doi.org/10.5281/zenodo.12003275" httpUri="https://zenodo.org/record/12003275/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
<figureCitation id="59AAC4857F16FFF7FB64FCF4FB06FCA8" box="[1240,1252,779,803]" captionStart="Fig" captionStartId="4.[167,198,1651,1671]" captionTargetBox="[166,1421,236,1632]" captionTargetId="figure-16@4.[166,1421,236,1638]" captionTargetPageId="4" captionText="Fig. 3 Field aspects of the Aheimer Formation; A General view of the studied succession of the lower member exposed in a faulted block to the north of Porto Sukhna; the shales mudstones close to the base (rectangle) are interbedded with thin, hard dolomitic bands. B Close up view on of the lower fossiliferous shale mudstone beds of A. C, D Close-up view on of two rugose corals entombed in the shale mudstone beds.E Close-up view on of a rugose coral species with diagenetically compressed calyx. Note strong ferrugination of corallite and surrounding matrix within the fossiliferous shale. F Fossiliferous shales mudstone with thin bands of ferruginous dolomitic limestone and secondary evaporite veinlets; Cr = crinoidal stems. G Crinoid columnals (Cr) and rugose corals (RC) are embedded in the shale mudstone. H Close-up view on of a spiriferid valve shell within the shale (arrow). Diameter of coins in CE, G: 25 mm" figureDoi="http://doi.org/10.5281/zenodo.12003198" httpUri="https://zenodo.org/record/12003198/files/figure.png" pageId="15" pageNumber="16">3</figureCitation>
, A
<figureCitation id="59AAC4857F16FFF7FAB8FCF4FAF0FCA8" box="[1284,1298,779,803]" captionStart="Fig" captionStartId="5.[167,198,1402,1422]" captionTargetBox="[172,1418,240,1387]" captionTargetId="figure-124@5.[166,1421,236,1389]" captionTargetPageId="5" captionText="Fig. 4 Effects of diagenesis on some coral specimens: A Thin-section in the calice of the specimen RAh (44) shows a completely compressed calice filled with fossiliferous mud. B Thin-section below the calice of the Bothrophyllum okense (RAh 37) shows beginning of compression and breakage of septa. C Thin-section below the calice of the specimen RAh (79), shows completely broken and altered septa, probable outer dissepimentarium and axial structure, filled with ferruginous mud. D Thin-section of a completely compressed specimen RAh (200) shows high alteration,breakage and ferrugination of skeletal elements.E Thin-section in the early mature part of the Actinophrentis crassithecata n. sp.(RAh 11), shows ferrugination affects a large part of the interior skeleton. F Mature thin-section of Rotiphyllum exile (RAh 29), shows ferrugination and dolomitization of skeletal elements.G, H Thin-sections of mature part of the specimen (RAh 81) in G and the immature part of the specimen RAh (99) in H show extensive dolomitization and ferrugination of the internal skeletal elements" figureDoi="http://doi.org/10.5281/zenodo.12003222" httpUri="https://zenodo.org/record/12003222/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,344 @@
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<mods:title id="1EA8630DC98980524EA78F2084497DE7">Kasimovian (late Pennsylvanian) cornute rugose corals from Egypt: taxonomy, facies and palaeogeography of a cool-water fauna from northern Gondwana</mods:title>
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<mods:title id="84565ED79B3436D041E8FBC993663118">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="97AC6502BE73BF02971C58A5572D56F8">2023</mods:date>
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<treatment id="493869167F16FFE9FC91FC92FE7AFB6E" LSID="urn:lsid:plazi:treatment:493869167F16FFE9FC91FC92FE7AFB6E" httpUri="http://treatment.plazi.org/id/493869167F16FFE9FC91FC92FE7AFB6E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="898B8B8B7F16FFF7FC91FC92FB32FC0F" box="[813,1232,876,901]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="C12ED8007F16FFF7FC91FC92FB32FC0F" blockId="15.[813,1437,876,1477]" box="[813,1232,876,901]" pageId="15" pageNumber="16">
<heading id="9A666F6C7F16FFF7FC91FC92FB32FC0F" box="[813,1232,876,901]" fontSize="36" level="2" pageId="15" pageNumber="16" reason="3">
Genus
<taxonomicName id="0691A3837F16FFF7FCC5FC93FB28FC0F" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" box="[889,1226,876,901]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCC5FC93FBF8FC0F" bold="true" box="[889,1050,876,900]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB9DFC92FB28FC0F" author="Fomichev, V." box="[1057,1226,876,901]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFF7FC91FC72FB49FC6F" pageId="15" pageNumber="16" type="type_taxon">
<paragraph id="C12ED8007F16FFF7FC91FC72FA94FC2F" blockId="15.[813,1437,876,1477]" box="[813,1398,908,933]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FC72FC24FC2F" bold="true" box="[813,966,908,933]" pageId="15" pageNumber="16">
<typeStatus id="1E2A66A27F16FFF7FC91FC72FC84FC2E" box="[813,870,909,933]" pageId="15" pageNumber="16">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F16FFF7FC70FC72FA8DFC2F" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[972,1391,908,933]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F16FFF7FC70FC72FB22FC2E" box="[972,1216,909,933]" italics="true" pageId="15" pageNumber="16">Monophyllum sokolovi</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB7AFC72FA8DFC2F" author="Fomichev, V." box="[1222,1391,908,933]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C12ED8007F16FFF7FC91FC52FB49FC6F" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FC91FC52FBB5FC4F" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[813,1111,940,965]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F16FFF7FC91FC52FCA7FC4F" box="[813,837,941,964]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="F3E504127F16FFF7FCEDFC52FC45FC4E" box="[849,935,941,965]" italics="true" pageId="15" pageNumber="16">sokolovi</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FC12FC52FBB5FC4F" author="Fomichev, V." box="[942,1111,940,965]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
;?
<bibRefCitation id="A500A5F17F16FFF7FBC9FC53FB4BFC4F" author="Fomichev, V." box="[1141,1193,940,964]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">1953</bibRefCitation>
M.
<taxonomicName id="0691A3837F16FFF7FB69FC52FC83FC6F" authority="Fomichev 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F16FFF7FB69FC52FACEFC4F" box="[1237,1324,941,964]" italics="true" pageId="15" pageNumber="16">parvum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FA8EFC52FC83FC6F" author="Fomichev, V." pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev 1953</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0691A3837F16FFF7FCD1FC32FB47FC6E" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[877,1189,972,997]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F16FFF7FCD1FC32FC67FC6F" box="[877,901,973,996]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="F3E504127F16FFF7FC2DFC32FC03FC6F" box="[913,993,973,996]" italics="true" pageId="15" pageNumber="16">cassum</emphasis>
Fedorowski, 1987
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFF7FC91FC13FB26FB4E" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="C12ED8007F16FFF7FC91FC13FB26FB4E" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FC13FC44FB8F" bold="true" box="[813,934,1004,1028]" pageId="15" pageNumber="16">Diagnosis:</emphasis>
Non-dissepimented, small solitary corals, with major septa radially arranged, may withdraw a little from corallite axis in maturity; cardinal septum shortened from early maturity; counter septum permanently elongated to corallite axis; alar septa slightly elongated at least in some portion of growth; minor septa underdeveloped (after
<bibRefCitation id="A500A5F17F16FFF7FC16FB52FB97FB4E" author="Fedorowski, J." box="[938,1141,1197,1221]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, p. 67).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFE9FC91FB32FE7AFB6E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
<paragraph id="C12ED8007F16FFF7FC91FB32FAADFA4E" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FB32FC78FB6E" bold="true" box="[813,922,1229,1253]" pageId="15" pageNumber="16">Remarks:</emphasis>
In contrast to
<bibRefCitation id="A500A5F17F16FFF7FBE5FB32FB2FFB6F" author="Hill, D." box="[1113,1229,1228,1253]" pageId="15" pageNumber="16" refId="ref26422" refString="Hill, D. (1956). Rugosa. In R. C. Moore (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata (pp. F 233 - F 324). New York." type="journal volume" year="1956">Hill (1956</bibRefCitation>
,
<bibRefCitation id="A500A5F17F16FFF7FB5DFB32FAF7FB6E" author="Hill, D." box="[1249,1301,1229,1253]" pageId="15" pageNumber="16" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">1981</bibRefCitation>
), de Groot (1963) and
<bibRefCitation id="A500A5F17F16FFF7FC17FB12FBA2FA8E" author="Weyer, D." box="[939,1088,1260,1285]" pageId="15" pageNumber="16" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
who listed
<taxonomicName id="0691A3837F16FFF7FB7EFB12FC4AFAAF" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB7EFB12FAB8FA8E" box="[1218,1370,1261,1285]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FADEFB12FC4AFAAF" author="Fomichev, V." pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
as a synonym of
<taxonomicName id="0691A3837F16FFF7FBD3FAF2FA74FAAE" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" box="[1135,1430,1293,1317]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FBD3FAF2FB16FAAE" box="[1135,1268,1293,1317]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB41FAF2FA74FAAE" author="Hudson, R. G. S." box="[1277,1430,1293,1317]" pageId="15" pageNumber="16" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="A500A5F17F16FFF7FC91FAD2FBEAFACE" author="Fedorowski, J." box="[813,1032,1325,1349]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
considered it as a separate genus. This view was followed by
<bibRefCitation id="A500A5F17F16FFF7FBEAFAB2FAEEFAEF" author="Kossovaya, O. L." box="[1110,1292,1356,1381]" pageId="15" pageNumber="16" pagination="71 - 82" refId="ref27287" refString="Kossovaya, O. L. (1996). Correlation of uppermost Carboniferous and Lower Permian Rugose Coral Zones from the Urals to Western North America. Palaios, 11 (1), 71 - 82." type="journal article" year="1996">Kossovaya (1996</bibRefCitation>
, 2001),
<bibRefCitation id="A500A5F17F16FFF7FAD9FAB2FC91FA0E" author="Ogar, V. V." pageId="15" pageNumber="16" pagination="340 - 349" refId="ref28259" refString="Ogar, V. V. (2012). Carboniferous buildups in the Donets Basin (Ukraine). Geologica Belgica, 15 (4), 340 - 349." type="journal article" year="2012">Ogar (2012)</bibRefCitation>
and
<bibRefCitation id="A500A5F17F16FFF7FC17FA92FB95FA0E" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." box="[939,1143,1389,1413]" pageId="15" pageNumber="16" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al. (2017)</bibRefCitation>
.
<bibRefCitation id="A500A5F17F16FFF7FB3FFA92FAB9FA0E" author="Fedorowski, J." box="[1155,1371,1389,1413]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
mentioned the following differences to
<taxonomicName id="0691A3837F16FFF7FB08FA72FADBFA2E" authorityName="Hudson" authorityYear="1942" box="[1204,1337,1421,1445]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB08FA72FADBFA2E" box="[1204,1337,1421,1445]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
and further similar antiphyllinid genera (see also
<tableCitation id="8C13EDBB7F16FFF7FB4EFA52FAA2FA4E" box="[1266,1344,1453,1477]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="15" pageNumber="16">Table 1</tableCitation>
):
</paragraph>
<paragraph id="C12ED8007F16FFF7FCFEFA11FB68F96D" blockId="15.[834,1437,1518,1894]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FCDEFA11FC1AF98D" authorityName="Fomichev, Shortened" authorityYear="1953" box="[866,1016,1518,1542]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCDEFA11FC1AF98D" box="[866,1016,1518,1542]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F16FFF7FB28FA11FAF5F98D" authorityName="Hudson" authorityYear="1942" box="[1172,1303,1518,1542]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB28FA11FAF5F98D" box="[1172,1303,1518,1542]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
by: (1) the strongly and permanently elongated counter septum reaching the corallite axis, even on the calice floor, where it may form an incipient columella, (2) shortening of the cardinal septum not only on the calice wall as in
<taxonomicName id="0691A3837F16FFF7FC77F971FBACF92D" authorityName="Hudson" authorityYear="1942" box="[971,1102,1678,1702]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FC77F971FBACF92D" box="[971,1102,1678,1702]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
, but also on the calice floor, (3) the triangular shape of at least peripheral parts of the cardinal fossula at maturity.
</paragraph>
<paragraph id="C12ED8007F16FFE9FCFEF911FE58FEE8" blockId="15.[834,1437,1518,1894]" lastBlockId="17.[173,775,235,547]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FCDEF911FC1AF88D" authorityName="Fomichev, Shortened" authorityYear="1953" box="[866,1016,1774,1798]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCDEF911FC1AF88D" box="[866,1016,1774,1798]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F16FFF7FB3AF911FAFCF88D" authorityName="Grabau" authorityYear="1928" box="[1158,1310,1774,1798]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB3AF911FAFCF88D" box="[1158,1310,1774,1798]" italics="true" pageId="15" pageNumber="16">Bradyphyllum</emphasis>
</taxonomicName>
by the permanently elongated counter septum that reaches the corallite axis and tends to form an incipient columella. Although
<taxonomicName id="0691A3837F16FFF7FBA0F8B1FB50F8ED" authorityName="Fomichev, Shortened" authorityYear="1953" box="[1052,1202,1870,1894]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FBA0F8B1FB50F8ED" box="[1052,1202,1870,1894]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
develops a tendency to form an axial area free of septa, like most of the genera included in
<taxonomicName id="0691A3837F08FFE9FE12FEF4FDDDFEA8" box="[430,575,267,291]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="35" phylum="Cnidaria" rank="subFamily" subFamily="Antiphyllinae">Antiphyllinae</taxonomicName>
(Illina, 1970), its counter septum remains elongated, which is not the case in
<taxonomicName id="0691A3837F08FFE9FEA0FEB4FE56FEE8" authorityName="Grabau" authorityYear="1928" box="[284,436,331,355]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FEA0FEB4FE56FEE8" box="[284,436,331,355]" italics="true" pageId="17" pageNumber="18">Bradyphyllum</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="95EE88887F09FFE8FF1BF9F8FD24F880" ID-DOI="http://doi.org/10.5281/zenodo.12003288" ID-Zenodo-Dep="12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="16" pageNumber="17" startId="16.[167,198,1543,1563]" targetBox="[291,1292,238,1527]" targetPageId="16" targetType="figure">
<paragraph id="C12ED8007F09FFE8FF1BF9F8FD24F880" blockId="16.[167,1422,1543,1803]" pageId="16" pageNumber="17">
<emphasis id="F3E504127F09FFE8FF1BF9F8FF10F990" bold="true" box="[167,242,1543,1563]" pageId="16" pageNumber="17">Fig. 9 A</emphasis>
<taxonomicName id="0691A3837F09FFE8FF4BF9F8FE28F990" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[247,458,1543,1563]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Cnidaria" rank="species" species="galalaensis" status="sp. nov.">
<emphasis id="F3E504127F09FFE8FF4BF9F8FE28F990" box="[247,458,1543,1563]" italics="true" pageId="16" pageNumber="17">Monophyllum galalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F09FFE8FE72F9F8FE19F990" box="[462,507,1543,1563]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
(Holotype specimen, Rah 9).
<emphasis id="F3E504127F09FFE8FD4CF9F8FCEBF990" bold="true" box="[752,777,1543,1563]" pageId="16" pageNumber="17">A1</emphasis>
Lateral external view of the corallite, positions of thin-sections indicated in A1.
<emphasis id="F3E504127F09FFE8FF67F9DDFF16F9BD" bold="true" box="[219,244,1570,1590]" pageId="16" pageNumber="17">A2</emphasis>
A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow).
<emphasis id="F3E504127F09FFE8FF1BF9C2FF06F9DA" bold="true" box="[167,228,1597,1617]" pageId="16" pageNumber="17">A3A5</emphasis>
Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn.
<emphasis id="F3E504127F09FFE8FC4DF9A8FBC3F9E0" bold="true" box="[1009,1057,1623,1643]" pageId="16" pageNumber="17">A67</emphasis>
Two successive mature sections, showing a short cardinal septum.
<emphasis id="F3E504127F09FFE8FEC4F98DFE4AF9F7" bold="true" box="[376,424,1646,1670]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FE2DF991FE4AF9F7" attach="left" box="[401,424,1646,1660]" fontSize="6" pageId="16" pageNumber="17">1, 2</superScript>
</emphasis>
Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note,
<emphasis id="F3E504127F09FFE8FC49F972FBF4F91C" bold="true" box="[1013,1046,1673,1697]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FBB2F976FBF4F91C" attach="left" box="[1038,1046,1673,1687]" fontSize="6" pageId="16" pageNumber="17">2</superScript>
</emphasis>
is a redrawing of section
<emphasis id="F3E504127F09FFE8FB52F972FAEDF91C" bold="true" box="[1262,1295,1673,1697]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FABBF976FAEDF91C" attach="none" box="[1287,1295,1673,1687]" fontSize="6" pageId="16" pageNumber="17">1</superScript>
</emphasis>
. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines.
<emphasis id="F3E504127F09FFE8FB93F958FBA5F930" bold="true" box="[1071,1095,1703,1723]" pageId="16" pageNumber="17">B2</emphasis>
Nepionic stage.
<emphasis id="F3E504127F09FFE8FB6FF958FB09F930" bold="true" box="[1235,1259,1703,1723]" pageId="16" pageNumber="17">B3</emphasis>
Early neanic stage.
<emphasis id="F3E504127F09FFE8FF1BF93DFF34F95D" bold="true" box="[167,214,1730,1750]" pageId="16" pageNumber="17">B45</emphasis>
Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.
<emphasis id="F3E504127F09FFE8FBD5F93DFB63F95D" bold="true" box="[1129,1153,1730,1750]" pageId="16" pageNumber="17">B6</emphasis>
Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
</paragraph>
</caption>
<paragraph id="C12ED8007F08FFE9FF11FE93FD36FDA8" blockId="17.[173,775,235,547]" pageId="17" pageNumber="18">
<taxonomicName id="0691A3837F08FFE9FF71FE94FE81FE08" authorityName="Fomichev, Shortened" authorityYear="1953" box="[205,355,363,387]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FF71FE94FE81FE08" box="[205,355,363,387]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F08FFE9FE49FE94FEE2FE28" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE49FE94FD73FE08" box="[501,657,363,387]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FD26FE94FEE2FE28" author="Fomichev, V." pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
by (1) the radial arrangement of septa in the mature portions of
<taxonomicName id="0691A3837F08FFE9FE0AFE54FDAEFE48" authorityName="Fomichev, Shortened" authorityYear="1953" box="[438,588,427,451]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE0AFE54FDAEFE48" box="[438,588,427,451]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
vs. the pinnate arrangement of the major septa lasting up to the calice floor in
<taxonomicName id="0691A3837F08FFE9FEE8FE14FE12FD88" authorityName="Fomichev" authorityYear="1953" box="[340,496,491,515]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FEE8FE14FE12FD88" box="[340,496,491,515]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
</taxonomicName>
and (2) the lacking tendency of
<taxonomicName id="0691A3837F08FFE9FE92FDF4FE28FDA8" authorityName="Fomichev" authorityYear="1953" box="[302,458,523,547]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE92FDF4FE28FDA8" box="[302,458,523,547]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
</taxonomicName>
to form a free axial area.
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BFDB2FD6DFD4F" blockId="17.[151,776,588,1253]" pageId="17" pageNumber="18">
Species described by
<bibRefCitation id="A500A5F17F08FFE9FE24FDB2FDB4FDEE" author="Fomichev, V." box="[408,598,588,613]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev (1953)</bibRefCitation>
to be “
<taxonomicName id="0691A3837F08FFE9FD07FDB2FF38FD0E" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Metriophyllidae" genus="Stereolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FD07FDB2FF38FD0E" italics="true" pageId="17" pageNumber="18">Stereolasma</emphasis>
</taxonomicName>
” might belong to
<taxonomicName id="0691A3837F08FFE9FE0EFD92FDD5FD0E" authorityName="Hudson" authorityYear="1942" box="[434,567,621,645]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE0EFD92FDD5FD0E" box="[434,567,621,645]" italics="true" pageId="17" pageNumber="18">Rotiphyllum</emphasis>
</taxonomicName>
(in toto, see
<bibRefCitation id="A500A5F17F08FFE9FD64FD92FF29FD2F" author="Hill, D." pageId="17" pageNumber="18" refId="ref26422" refString="Hill, D. (1956). Rugosa. In R. C. Moore (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata (pp. F 233 - F 324). New York." type="journal volume" year="1956">Hill, 1956</bibRefCitation>
; de Groot, 1963), to
<taxonomicName id="0691A3837F08FFE9FE6CFD72FD8AFD2E" authorityName="Fomichev, Shortened" authorityYear="1953" box="[464,616,653,677]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE6CFD72FD8AFD2E" box="[464,616,653,677]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
and/or other Antiphyllinidae (
<bibRefCitation id="A500A5F17F08FFE9FEF1FD52FDFBFD4E" author="Fedorowski, J." box="[333,537,685,709]" pageId="17" pageNumber="18" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, p. 36, 68).
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BFD33FE7AFB6E" blockId="17.[151,776,588,1253]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FF2BFD33FDDEFD6E" bold="true" box="[151,572,716,741]" pageId="17" pageNumber="18">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F08FFE9FDFFFD32FD39FD6E" authorityName="Fomichev, Shortened" authorityYear="1953" box="[579,731,717,741]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FDFFFD32FD39FD6E" box="[579,731,717,741]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
is a long ranging but relatively rare genus. It is known from the Bashkirian of the Donets Basin (
<bibRefCitation id="A500A5F17F08FFE9FDF2FCF2FCE2FCAF" author="Fomichev, V." box="[590,768,780,805]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FF2BFCD2FEB3FCCF" author="Kossovaya, O. L." box="[151,337,812,837]" pageId="17" pageNumber="18" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FED9FCD3FE4AFCCF" box="[357,424,812,836]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FE06FCD3FD9DFCCE" author="Fedorowski, J." box="[442,639,812,837]" pageId="17" pageNumber="18" pagination="247 - 316" refId="ref25858" refString="Fedorowski, J. (2022). Bashkirian Rugosa (Anthozoa) from the Donets Basin (Ukraine). Part 12. Concluding considerations. Acta Geologica Polonica, 72 (3), 247 - 316." type="journal article" year="2022">Fedorowski, 2022</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FD2FFCD3FD35FCCE" box="[659,727,812,837]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
:
<taxonomicName id="0691A3837F08FFE9FD55FCD2FE4BFCEF" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F08FFE9FD55FCD2FCE3FCCF" box="[745,769,813,836]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FF2BFCB3FF0FFCEF" box="[151,237,844,868]" italics="true" pageId="17" pageNumber="18">sokolovi</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FF45FCB3FE4BFCEF" author="Fomichev, V." box="[249,425,844,868]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0691A3837F08FFE9FE07FCB2FD1AFCEF" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[443,760,844,868]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F08FFE9FE07FCB2FE31FCEF" box="[443,467,845,868]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FE5AFCB2FDDFFCEF" box="[486,573,845,868]" italics="true" pageId="17" pageNumber="18">parvum</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FDF5FCB3FD1AFCEF" author="Fomichev, V." box="[585,760,844,868]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
), the Moscovian of the same region (
<bibRefCitation id="A500A5F17F08FFE9FDA8FC93FD5CFC0F" author="Fomichev, V." box="[532,702,876,900]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
: original designation of the
<typeStatus id="1E2A66A27F08FFE9FE26FC72FE2BFC2E" box="[410,457,909,933]" pageId="17" pageNumber="18">type</typeStatus>
species
<taxonomicName id="0691A3837F08FFE9FD91FC72FD49FC2E" authorityName="Fomichev" authorityYear="1953" box="[557,683,909,933]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F08FFE9FD91FC72FD49FC2E" box="[557,683,909,933]" italics="true" pageId="17" pageNumber="18">M. sokolovi</emphasis>
</taxonomicName>
;
<bibRefCitation id="A500A5F17F08FFE9FD00FC72FEEDFC4E" author="Kossovaya, O. L." pageId="17" pageNumber="18" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FE9EFC52FE86FC4F" box="[290,356,941,965]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FECAFC52FE10FC4E" author="Ogar, V. V." box="[374,498,941,965]" pageId="17" pageNumber="18" pagination="340 - 349" refId="ref28259" refString="Ogar, V. V. (2012). Carboniferous buildups in the Donets Basin (Ukraine). Geologica Belgica, 15 (4), 340 - 349." type="journal article" year="2012">Ogar, 2012</bibRefCitation>
) and the Moscovian of Novaya Zemlya (fide
<bibRefCitation id="A500A5F17F08FFE9FE3BFC32FDA5FC6E" author="Fedorowski, J." box="[391,583,973,997]" pageId="17" pageNumber="18" pagination="87 - 160" refId="ref25455" refString="Fedorowski, J. (1981). Carboniferous corals: Distribution and sequence. Acta Palaeontologica Polonica, 26 (2), 87 - 160." type="journal article" year="1981">Fedorowski, 1981</bibRefCitation>
and
<bibRefCitation id="A500A5F17F08FFE9FD3EFC32FF29FB8E" author="Wang, X. D. &amp; Yang, S. R. &amp; Yao, L. &amp; Sugiyama, T. &amp; Hu, K. Y." pageId="17" pageNumber="18" pagination="603 - 63" refId="ref29952" refString="Wang, X. D., Yang, S. R., Yao, L., Sugiyama, T., &amp; Hu, K. Y. (2022). Carboniferous biostratigraphy of rugose corals. In S. G. Lucas, J. W. Schneider, X. Wang, S. Nikolaeva (Eds), The Carboniferous timescale. Geological Society, London, Special Publications. 512 (2022). 603 - 63." type="journal article" year="2022">Wang et al., 2022</bibRefCitation>
). It is also known from the PennsylvanianPermian transition and the Wolfcampian (Asselianmiddle Artinskian) of southwest Texas (
<bibRefCitation id="A500A5F17F08FFE9FE07FBD2FD64FBCE" author="Fedorowski, J." box="[443,646,1069,1093]" pageId="17" pageNumber="18" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
:
<taxonomicName id="0691A3837F08FFE9FD2FFBD2FEB7FBEE" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F08FFE9FD2FFBD2FD49FBCF" box="[659,683,1069,1092]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FD0BFBD2FCE5FBCF" box="[695,775,1069,1092]" italics="true" pageId="17" pageNumber="18">cassum</emphasis>
Fedorowski, 1987
</taxonomicName>
). Kossovaya et al., (2001,
<figureCitation id="59AAC4857F08FFE9FDD2FBB2FD49FBEE" box="[622,683,1101,1125]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
) figured?
<taxonomicName id="0691A3837F08FFE9FF1DFB92FE83FB0F" box="[161,353,1133,1157]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F08FFE9FF1DFB92FEDBFB0E" box="[161,313,1133,1157]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
sp.
</taxonomicName>
from the late Artinskian of the Urals and/or northern Timan Ridge (not further specified). Herein, we report the genus from the Kasimovian west of the Gulf of
<collectingRegion id="035516E27F08FFE9FEAEFB32FEAAFB6E" box="[274,328,1229,1253]" country="Egypt" name="As Suways" pageId="17" pageNumber="18">Suez</collectingRegion>
,
<collectingCountry id="B98698907F08FFE9FEEEFB32FE76FB6E" box="[338,404,1229,1253]" name="Egypt" pageId="17" pageNumber="18">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,118 @@
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<subSubSection id="898B8B8B7F1FFFFEFC91FCF4FB7BFCA8" box="[813,1177,779,803]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="C12ED8007F1FFFFEFC91FCF4FB7BFCA8" blockId="6.[813,1437,715,1091]" box="[813,1177,779,803]" pageId="6" pageNumber="7">
<heading id="9A666F6C7F1FFFFEFC91FCF4FB7BFCA8" box="[813,1177,779,803]" fontSize="36" level="2" pageId="6" pageNumber="7" reason="3">
Family
<taxonomicName id="0691A3837F1FFFFEFCC0FCF4FBF9FCA8" authority="Ilina, 1970" box="[892,1051,779,803]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="6" pageNumber="7" phylum="Cnidaria" rank="family">
<emphasis id="F3E504127F1FFFFEFCC0FCF4FBF9FCA8" bold="true" box="[892,1051,779,803]" pageId="6" pageNumber="7">Antiphyllidae</emphasis>
</taxonomicName>
<bibRefCitation id="A500A5F17F1FFFFEFB9DFCF4FB70FCA8" author="Ilina, T. G." box="[1057,1170,779,803]" pageId="6" pageNumber="7" pagination="146 - 151" refId="ref26741" refString="Ilina, T. G. (1970). Some new Permian Rugosa from the South-Eastern Pamir. 146 - 151. In G. G. Astrova &amp; I. I. Chudinova (Eds.), New species of the Palaeozoic bryozoans and corals (p. 178). Nauka." type="journal article" year="1970">Ilina, 1970</bibRefCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F1FFFFEFC91FCD4FBFBFB88" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="C12ED8007F1FFFFEFC91FCD4FBFBFB88" blockId="6.[813,1437,715,1091]" pageId="6" pageNumber="7">
<emphasis id="F3E504127F1FFFFEFC91FCD4FC42FCC8" bold="true" box="[813,928,811,835]" pageId="6" pageNumber="7">Diagnosis</emphasis>
(
<bibRefCitation id="A500A5F17F1FFFFEFC13FCD4FB9FFCC8" author="Fedorowski, J." box="[943,1149,811,835]" pageId="6" pageNumber="7" pagination="1 - 33" refId="ref25787" refString="Fedorowski, J. (2012 a). On three rugose coral genera from Serpukhovian strata in the Upper Silesian Coal Basin, Poland. Acta Geologica Polonica, 62 (1), 1 - 33." type="journal article" year="2012">Fedorowski, 2012a</bibRefCitation>
): Small, solitary, non-dissepimented corals with cardinal septum located on convex side of corallite; early ontogeny rotiphylloid; cardinal septum shortened late in ontogeny or only on calice wall; counter septum commonly elongated, never shortened; stereocolumn of early ontogeny often replaced by free axial area in maturity.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F1FFFF0FC91FBF4FF0CFD88" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="discussion">
<paragraph id="C12ED8007F1FFFF0FC91FBF4FF0CFD88" blockId="6.[813,1437,715,1091]" lastBlockId="8.[151,775,235,515]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<emphasis id="F3E504127F1FFFFEFC91FBF4FC78FBA8" bold="true" box="[813,922,1035,1059]" pageId="6" pageNumber="7">Remarks:</emphasis>
Opposed to
<bibRefCitation id="A500A5F17F1FFFFEFB93FBF4FB45FBA8" author="Hill, D." box="[1071,1191,1035,1059]" pageId="6" pageNumber="7" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
,
<bibRefCitation id="A500A5F17F1FFFFEFB0AFBF3FAAEFBA8" author="Weyer, D." box="[1206,1356,1035,1060]" pageId="6" pageNumber="7" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
and in consequence
<bibRefCitation id="A500A5F17F1FFFFEFC74FBD4FB44FBC8" author="Fedorowski, J." box="[968,1190,1067,1091]" pageId="6" pageNumber="7" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
did not recognise a family
<taxonomicName id="0691A3837F11FFF0FF59FF14FE9AFE88" box="[229,376,235,259]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="family">Antiphyllidae</taxonomicName>
<bibRefCitation id="A500A5F17F11FFF0FE3EFF14FE1AFE88" author="Ilina, T. G." box="[386,504,235,259]" pageId="8" pageNumber="9" pagination="146 - 151" refId="ref26741" refString="Ilina, T. G. (1970). Some new Permian Rugosa from the South-Eastern Pamir. 146 - 151. In G. G. Astrova &amp; I. I. Chudinova (Eds.), New species of the Palaeozoic bryozoans and corals (p. 178). Nauka." type="journal article" year="1970">Ilina, 1970</bibRefCitation>
, but separated the subfamilies
<taxonomicName id="0691A3837F11FFF0FF40FEF4FDAAFEA8" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" box="[252,584,267,291]" class="Anthozoa" family="Hapsiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="21" phylum="Cnidaria" rank="subFamily" subFamily="Hapsiphyllinae">
Hapsiphyllinae
<bibRefCitation id="A500A5F17F11FFF0FE11FEF4FDAAFEA8" author="Grabau, A. W." box="[429,584,267,291]" pageId="8" pageNumber="9" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
and Antiphyllinae
<bibRefCitation id="A500A5F17F11FFF0FF7BFED4FEDEFEC8" author="Ilina, T. G." box="[199,316,299,323]" pageId="8" pageNumber="9" pagination="146 - 151" refId="ref26741" refString="Ilina, T. G. (1970). Some new Permian Rugosa from the South-Eastern Pamir. 146 - 151. In G. G. Astrova &amp; I. I. Chudinova (Eds.), New species of the Palaeozoic bryozoans and corals (p. 178). Nauka." type="journal article" year="1970">Ilina, 1970</bibRefCitation>
within the
<taxonomicName id="0691A3837F11FFF0FE7DFED4FCE2FEC8" authority="Grabau, 1928" authorityName="Grabau" authorityYear="1928" box="[449,768,299,323]" class="Anthozoa" family="Hapsiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="family">
Hapsiphyllidae
<bibRefCitation id="A500A5F17F11FFF0FDD0FED4FCE2FEC8" author="Grabau, A. W." box="[620,768,299,323]" pageId="8" pageNumber="9" pagination="1 - 175" refId="ref26151" refString="Grabau, A. W. (1928). Palaeozoic Corals of China: Part I, Tetraseptata II. Palaeontologica Sinica, B, 2, 1 - 175." type="journal article" year="1928">Grabau, 1928</bibRefCitation>
</taxonomicName>
. However,
<bibRefCitation id="A500A5F17F11FFF0FEB8FEB4FE33FEE8" author="Fedorowski, J." box="[260,465,331,355]" pageId="8" pageNumber="9" pagination="1 - 37" refId="ref25654" refString="Fedorowski, J. (2009 a). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin, Ukraine. Part 1. Introductory considerations and the genus Rotiphyllum Hudson, 1942. Acta Geologica Polonica, 59 (1), 1 - 37." type="journal article" year="2009">Fedorowski (2009a</bibRefCitation>
,
<bibRefCitation id="A500A5F17F11FFF0FE62FEB4FDC3FEE8" author="Fedorowski, J." box="[478,545,331,355]" pageId="8" pageNumber="9" pagination="283 - 317" refId="ref25702" refString="Fedorowski, J. (2009 b). Early Bashkirian Rugosa (Anthozoa) from the Donets Basin, Ukraine. Part 2. On the genera Axisvacuus gen. nov. and Falsiamplexus Fedorowski, 1987. Acta Geologica Polonica, 59 (3), 283 - 317." type="journal article" year="2009">2009b</bibRefCitation>
) and in his following papers treated the
<taxonomicName id="0691A3837F11FFF0FEDBFE94FE18FE08" box="[359,506,363,387]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="family">Antiphyllidae</taxonomicName>
as an independent family. An emended diagnosis, differing from
<bibRefCitation id="A500A5F17F11FFF0FDDEFE74FD3AFE28" author="Hill, D." box="[610,728,395,419]" pageId="8" pageNumber="9" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">Hill (1981)</bibRefCitation>
and based on the main characters of the subfamily diagnosis by
<bibRefCitation id="A500A5F17F11FFF0FF00FE34FE75FE68" author="Fedorowski, J." box="[188,407,459,483]" pageId="8" pageNumber="9" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
was finally given in
<bibRefCitation id="A500A5F17F11FFF0FD35FE34FF08FD88" author="Fedorowski, J." pageId="8" pageNumber="9" pagination="1 - 33" refId="ref25787" refString="Fedorowski, J. (2012 a). On three rugose coral genera from Serpukhovian strata in the Upper Silesian Coal Basin, Poland. Acta Geologica Polonica, 62 (1), 1 - 33." type="journal article" year="2012">Fedorowski (2012a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>