619 lines
81 KiB
XML
619 lines
81 KiB
XML
<document id="56AD7232E08EAE43650EEF82D0B00DB3" ID-DOI="10.11646/zootaxa.5536.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14024191" ID-ZooBank="991D72E2-7948-4433-8434-FE647649EA75" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1730456217362" checkinUser="plazi" docAuthor="Farquhar, Jules E., Mulder, Jordan A., Russell, Wyn, Haines, Margaret L., Rangers, Anangu & Chapple, David G." docDate="2024" docId="114FD27EFA65FFDBD4A8278CFB8ECE09" docLanguage="en" docName="zootaxa.5536.1.6.pdf" docOrigin="Zootaxa 5536 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liopholis aputja Farquhar, Mulder, Russell, Haines & Chapple 2024, sp. nov." docType="treatment" docUuid="21D7BF40-8680-436A-B0DA-4D0E3D07D6EE" docUuidSource="ZooBank" docVersion="3" lastPageNumber="167" masterDocId="ED76AA06FA69FFD4D43F255AFFB4CF6C" masterDocTitle="A new species of rock skink (Scincidae: Liopholis) from the Central Ranges bioregion of arid Australia" masterLastPageNumber="177" masterPageNumber="153" pageNumber="165" updateTime="1731010068860" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="C07AF44EBB37B87A5240C1D4167C10D7">A new species of rock skink (Scincidae: Liopholis) from the Central Ranges bioregion of arid Australia</mods:title>
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<mods:name id="31F180021F2E6DD68602B0D5EE0CB76B" type="personal">
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<mods:namePart id="4ED4D4F62E939A8C5C60B89BA308F9E6">Farquhar, Jules E.</mods:namePart>
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<mods:affiliation id="A551C01B5CF6DED38887F264C53CF6C3">School of Biological Sciences, Monash University, Clayton, Victoria, Australia</mods:affiliation>
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<mods:namePart id="C9ADA777519AE4AA186FF161DB3E7100">Mulder, Jordan A.</mods:namePart>
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<mods:affiliation id="0F6E5FF0A459AC5A539658257B5B6B88">School of Biological Sciences, Monash University, Clayton, Victoria, Australia</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="EFEE8D3064E6072CCA0293453A8B2991">Author</mods:roleTerm>
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<mods:namePart id="28B199DA978C9D11013EC1130CE7E700">Russell, Wyn</mods:namePart>
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<mods:affiliation id="A3A4B0520215730D65883962FF325A35">Biosis Pty Ltd, Melbourne, Victoria, Australia</mods:affiliation>
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<mods:name id="ADCE40B1A2D7459EEFED6B2B1D231C07" type="personal">
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<mods:namePart id="415A59C1BF107B672A4FAC95BA36D4F7">Haines, Margaret L.</mods:namePart>
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<mods:affiliation id="416450ABBD015B57D1C8E8F72DFDE121">Museums Victoria, GPO Box 666, Melbourne VIC 3001</mods:affiliation>
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<mods:namePart id="95AC97DA2A21A382FF6DF6B92B0B0440">Rangers, Anangu</mods:namePart>
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<mods:namePart id="D961D2EA36411835AAD129AE31927995">Chapple, David G.</mods:namePart>
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<mods:affiliation id="9F0F047CEAEB7C695E7569F6EAAC9920">School of Biological Sciences, Monash University, Clayton, Victoria, Australia</mods:affiliation>
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<mods:title id="DC950626F12D428545489AC9E05925C8">Zootaxa</mods:title>
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<mods:date id="11EEDD4BD7F7E8C268A3484B99F76B1B">2024</mods:date>
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<mods:number id="DC4A877097588937D2BA72040019DAB6">2024-11-01</mods:number>
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<mods:url id="1E68A1F58B98E324C8064AEA0770E4F0">http://dx.doi.org/10.11646/zootaxa.5536.1.6</mods:url>
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<treatment id="114FD27EFA65FFDBD4A8278CFB8ECE09" LSID="urn:lsid:zoobank.org:act:21D7BF40-8680-436A-B0DA-4D0E3D07D6EE" httpUri="http://treatment.plazi.org/id/114FD27EFA65FFDBD4A8278CFB8ECE09" lastPageId="15" lastPageNumber="167" pageId="12" pageNumber="165">
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<subSubSection id="D1FC30E3FA65FFD8D4A8278CFBBDCD9D" box="[151,1033,726,753]" pageId="12" pageNumber="165" type="nomenclature">
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<heading id="C211D404FA65FFD8D4A8278CFBBDCD9D" bold="true" box="[151,1033,726,753]" fontSize="11" level="1" pageId="12" pageNumber="165" reason="1">
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<taxonomicName id="5EE618EBFA65FFD8D4A8278CFC11CD9C" authority="Farquhar, Mulder, Russell, Haines & Chapple" authorityName="Farquhar, Mulder, Russell, Haines & Chapple" authorityYear="2024" box="[151,933,726,753]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
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<emphasis id="AB92BF7AFA65FFD8D4A8278CFEEACD9D" bold="true" box="[151,350,726,753]" italics="true" pageId="12" pageNumber="165">Liopholis aputja</emphasis>
|
||
Farquhar, Mulder, Russell, Haines & Chapple
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="B0A10201FA65FFD8D793278DFBBDCD9D" box="[940,1033,727,753]" pageId="12" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="D1FC30E3FA65FFD8D4A82644FE47CC54" box="[151,499,798,825]" pageId="12" pageNumber="165" type="vernacular_names">
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<paragraph id="99596368FA65FFD8D4A82644FE47CC54" blockId="12.[151,980,798,897]" box="[151,499,798,825]" pageId="12" pageNumber="165">
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<heading id="C211D404FA65FFD8D4A82644FE47CC54" bold="true" box="[151,499,798,825]" fontSize="11" level="1" pageId="12" pageNumber="165" reason="1">
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<emphasis id="AB92BF7AFA65FFD8D4A82644FE47CC54" bold="true" box="[151,499,798,825]" pageId="12" pageNumber="165">Central Ranges Rock Skink</emphasis>
|
||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D1FC30E3FA65FFD8D4A82619FED1CCED" pageId="12" pageNumber="165" type="description">
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<paragraph id="99596368FA65FFD8D4A82619FC60CC31" blockId="12.[151,980,798,897]" box="[151,980,835,861]" pageId="12" pageNumber="165">
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<uri id="ED776F6AFA65FFD8D4A82619FC60CC31" box="[151,980,835,861]" pageId="12" pageNumber="165">
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urn:lsid:zoobank.org:act:
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<uuid id="ED4059BDFA65FFD8D5962619FC60CC31" box="[425,980,835,861]" pageId="12" pageNumber="165">21D7BF40-8680-436A-B0DA-4D0E3D07D6EE</uuid>
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</uri>
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</paragraph>
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<paragraph id="99596368FA65FFD8D4A8263DFED1CCED" blockId="12.[151,980,798,897]" box="[151,357,870,897]" pageId="12" pageNumber="165">
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(
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<figureCitation id="01DD7FEDFA65FFD8D4A0263DFEB8CCED" box="[159,268,871,897]" captionStart="FIGURE 4" captionStartId="8.[152,255,1283,1307]" captionTargetBox="[151,1436,179,1257]" captionTargetId="figure-31@8.[151,1436,179,1257]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Liopholis aputja sp. nov. (SAMA R60802) from Alalkanya Gorge, South Australia. Photos: Jules Farquhar." figureDoi="http://doi.org/10.5281/zenodo.14024204" httpUri="https://zenodo.org/record/14024204/files/figure.png" pageId="12" pageNumber="165">Figures 4</figureCitation>
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,
|
||
<figureCitation id="01DD7FEDFA65FFD8D527263CFEEACCED" box="[280,350,870,897]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="12" pageNumber="165">5A–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D1FC30E3FA65FFD8D4A826F5FECBCA5D" pageId="12" pageNumber="165" type="materials_examined">
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<paragraph id="99596368FA65FFD8D4A826F5FB9CCB11" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="165">
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<materialsCitation id="298E6935FA65FFD8D4A826F5FB9CCB11" collectingDate="2005-10-20" collectionCode="SAMA" collectorName="J. van Weenan" county="Alalkanya Gorge" latitude="-26.1633" location="Alalkanya Gorge" longLatPrecision="6" longitude="132.1166" municipality="Musgrave Ranges" pageId="12" pageNumber="165" specimenCode="R60802" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
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<emphasis id="AB92BF7AFA65FFD8D4A826F5FE39CCA4" bold="true" box="[151,397,942,969]" pageId="12" pageNumber="165">
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<typeStatus id="465DDDCAFA65FFD8D4A826F5FEB0CCA5" box="[151,260,943,969]" pageId="12" pageNumber="165" type="holotype">Holotype</typeStatus>
|
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(
|
||
<figureCitation id="01DD7FEDFA65FFD8D52A26F4FECACCA4" box="[277,382,942,968]" captionStart="FIGURE 4" captionStartId="8.[152,255,1283,1307]" captionTargetBox="[151,1436,179,1257]" captionTargetId="figure-31@8.[151,1436,179,1257]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Liopholis aputja sp. nov. (SAMA R60802) from Alalkanya Gorge, South Australia. Photos: Jules Farquhar." figureDoi="http://doi.org/10.5281/zenodo.14024204" httpUri="https://zenodo.org/record/14024204/files/figure.png" pageId="12" pageNumber="165">Figure 4</figureCitation>
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).
|
||
</emphasis>
|
||
<collectionCode id="FFF7FBADFA65FFD8D5A926F5FE51CCA5" box="[406,485,943,969]" collectionName="Australia, South Australia, Adelaide, South Australian Museum" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="12" pageNumber="165" type="Museum">SAMA</collectionCode>
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<specimenCode id="C940CB13FA65FFD8D5D126F5FDFECCA5" box="[494,586,943,969]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="165">R60802</specimenCode>
|
||
, adult, collected at night while active in
|
||
<location id="9C3935B3FA65FFD8D02226F5FB55CCA5" LSID="urn:lsid:plazi:treatment:114FD27EFA65FFDBD4A8278CFB8ECE09:9C3935B3FA65FFD8D02226F5FB55CCA5" box="[1053,1249,943,969]" county="Alalkanya Gorge" latitude="-26.1633" longLatPrecision="6" longitude="132.1166" municipality="Musgrave Ranges" name="Alalkanya Gorge" pageId="12" pageNumber="165" stateProvince="South Australia">Alalkanya Gorge</location>
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,
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<locationDeviation id="88EB7A4AFA65FFD8D0D026F5FF5ACC81" pageId="12" pageNumber="165">
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<quantity id="5E1ECE8DFA65FFD8D0D026F5FA8CCCA5" box="[1263,1336,943,969]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="165" unit="km" value="13.0">13 km</quantity>
|
||
north of Pukatja
|
||
</locationDeviation>
|
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,
|
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<collectingMunicipality id="793DF912FA65FFD8D4C42689FE7DCC81" box="[251,457,979,1005]" pageId="12" pageNumber="165">Musgrave Ranges</collectingMunicipality>
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,
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<collectingRegion id="5B22AD8AFA65FFD8D5E82689FD32CC81" box="[471,646,979,1005]" country="Australia" name="South Australia" pageId="12" pageNumber="165">South Australia</collectingRegion>
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(
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<geoCoordinate id="FCD205AFFA65FFD8D6A82689FCB4CC81" box="[663,768,979,1005]" degrees="26.1633" direction="south" orientation="latitude" pageId="12" pageNumber="165" precision="5" value="-26.1633">-26.1633</geoCoordinate>
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,
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<geoCoordinate id="FCD205AFFA65FFD8D7322689FCCDCC81" box="[781,889,979,1005]" degrees="132.1166" direction="east" orientation="longitude" pageId="12" pageNumber="165" precision="5" value="132.1166">132.1166</geoCoordinate>
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||
), by
|
||
<collectorName id="341306BEFA65FFD8D78C2689FBECCC81" box="[947,1112,979,1005]" pageId="12" pageNumber="165">J. van Weenan</collectorName>
|
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on
|
||
<date id="ED5845A8FA65FFD8D0B92689FAF3CC81" box="[1158,1351,978,1005]" pageId="12" pageNumber="165" value="2005-10-20">
|
||
<collectingDate id="FD1CBC40FA65FFD8D0B92689FAF3CC81" box="[1158,1351,978,1005]" pageId="12" pageNumber="165" value="2005-10-20">20 October 2005</collectingDate>
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||
</date>
|
||
(J. van Weenan pers. comm. 2024). Measurements in millimetres: SVL 92, tail 158 (original, 170% of SVL), HL 20, HW 15.1, HD 9.6, AGL 46.2, BW 18.7, FLL 26.3, HLL 36. Scale counts: MSR 37, SubDig 28, SupLab 9, SupCil 6, Cil 13, Lob 5 (with an additional 6
|
||
<superScript id="6E93CE20FA65FFD8D5C42164FDBCCB20" attach="left" box="[507,520,1086,1100]" fontSize="6" pageId="12" pageNumber="165">th</superScript>
|
||
miniscule uppermost lobule), nuchal scales in two pairs (
|
||
<emphasis id="AB92BF7AFA65FFD8D0A72165FB00CB35" box="[1176,1204,1087,1113]" italics="true" pageId="12" pageNumber="165">cf.</emphasis>
|
||
one pair in all other specimens). Colouration of this specimen is typical of the species in preservative.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="99596368FA65FFD8D4F821DDFBB6CB85" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="165">
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<materialsCitation id="298E6935FA65FFD8D4F821DDFE38CBA9" collectingDate="1998-10-25" collectionCode="R" collectorName="M. Hutchinson" latitude="-26.2558" location="South Australia" longLatPrecision="6" longitude="131.4933" pageId="12" pageNumber="165" specimenCode="R51590" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
|
||
<emphasis id="AB92BF7AFA65FFD8D4F821DDFEFCCBCD" bold="true" box="[199,328,1159,1185]" pageId="12" pageNumber="165">
|
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<typeStatus id="465DDDCAFA65FFD8D4F821DDFEF7CBCD" box="[199,323,1159,1185]" pageId="12" pageNumber="165" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="C940CB13FA65FFD8D57121DDFE1ECBCD" box="[334,426,1158,1185]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="165">R51590</specimenCode>
|
||
,
|
||
<locationDeviation id="88EB7A4AFA65FFD8D58C21DCFD01CBCD" box="[435,693,1158,1185]" pageId="12" pageNumber="165">
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<quantity id="5E1ECE8DFA65FFD8D58C21DCFDBACBCD" box="[435,526,1158,1185]" metricMagnitude="4" metricUnit="m" metricValue="3.65" pageId="12" pageNumber="165" unit="km" value="36.5">36.5 km</quantity>
|
||
ESE of Amata
|
||
</locationDeviation>
|
||
,
|
||
<collectingRegion id="5B22AD8AFA65FFD8D68121DDFCDECBCD" box="[702,874,1159,1185]" country="Australia" name="South Australia" pageId="12" pageNumber="165">South Australia</collectingRegion>
|
||
(
|
||
<geoCoordinate id="FCD205AFFA65FFD8D74821DCFC54CBCD" box="[887,992,1158,1185]" degrees="26.2558" direction="south" orientation="latitude" pageId="12" pageNumber="165" precision="5" value="-26.2558">-26.2558</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FCD205AFFA65FFD8D7D521DDFBE3CBCD" box="[1002,1111,1159,1185]" degrees="131.4933" direction="east" orientation="longitude" pageId="12" pageNumber="165" precision="5" value="131.4933">131.4933</geoCoordinate>
|
||
), collected by
|
||
<collectorName id="341306BEFA65FFD8D0C921DDFA28CBCD" box="[1270,1436,1159,1185]" pageId="12" pageNumber="165">M. Hutchinson</collectorName>
|
||
on
|
||
<date id="ED5845A8FA65FFD8D4FF21F0FE38CBA9" box="[192,396,1194,1221]" pageId="12" pageNumber="165" value="1998-10-25">
|
||
<collectingDate id="FD1CBC40FA65FFD8D4FF21F0FE38CBA9" box="[192,396,1194,1221]" pageId="12" pageNumber="165" value="1998-10-25">25 October 1998</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="298E6935FA65FFD8D5A121F1FE18CB85" collectingDate="1961-07-07" collectionCode="R" latitude="-26.35833" location="Erliwunyawunya Rockhole" longLatPrecision="1" longitude="131.72499" pageId="12" pageNumber="165" specimenCode="R17456" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
|
||
<specimenCode id="C940CB13FA65FFD8D5A121F1FE4DCBA9" box="[414,505,1194,1221]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="165">R17456</specimenCode>
|
||
,
|
||
<specimenCount id="8FE0A8E1FA65FFD8D63521F1FDD3CBA9" box="[522,615,1195,1221]" pageId="12" pageNumber="165" type="juvenile">juvenile</specimenCount>
|
||
,
|
||
<location id="9C3935B3FA65FFD8D64621F1FC07CBA9" LSID="urn:lsid:plazi:treatment:114FD27EFA65FFDBD4A8278CFB8ECE09:9C3935B3FA65FFD8D64621F1FC07CBA9" box="[633,947,1195,1221]" latitude="-26.35833" longLatPrecision="1" longitude="131.72499" name="Erliwunyawunya Rockhole" pageId="12" pageNumber="165" stateProvince="South Australia">Erliwunyawunya Rockhole</location>
|
||
,
|
||
<collectingRegion id="5B22AD8AFA65FFD8D7FA21F1FBCCCBA9" box="[965,1144,1195,1221]" country="Australia" name="South Australia" pageId="12" pageNumber="165">South Australia</collectingRegion>
|
||
(
|
||
<geoCoordinate id="FCD205AFFA65FFD8D0B221F0FAB7CBA9" box="[1165,1283,1194,1221]" degrees="26.35833" direction="south" orientation="latitude" pageId="12" pageNumber="165" precision="1" value="-26.35833">-26.35833</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FCD205AFFA65FFD8D12A21F1FA24CBA9" box="[1301,1424,1195,1221]" degrees="131.72499" direction="east" orientation="longitude" pageId="12" pageNumber="165" precision="1" value="131.72499">131.72499</geoCoordinate>
|
||
), collected on
|
||
<date id="ED5845A8FA65FFD8D5192195FE18CB85" box="[294,428,1231,1257]" pageId="12" pageNumber="165" value="1961-07-07">
|
||
<collectingDate id="FD1CBC40FA65FFD8D5192195FE18CB85" box="[294,428,1231,1257]" pageId="12" pageNumber="165" value="1961-07-07">7 July 1961</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="298E6935FA65FFD8D5872195FC4ACB85" box="[440,1022,1231,1257]" collectionCode="R" latitude="-26.116" location="Piltadi" longLatPrecision="74" longitude="130.283" pageId="12" pageNumber="165" specimenCode="R17271" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
|
||
<specimenCode id="C940CB13FA65FFD8D5872195FDA7CB85" box="[440,531,1231,1257]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="165">R17271</specimenCode>
|
||
,
|
||
<location id="9C3935B3FA65FFD8D6212195FDDECB85" LSID="urn:lsid:plazi:treatment:114FD27EFA65FFDBD4A8278CFB8ECE09:9C3935B3FA65FFD8D6212195FDDECB85" box="[542,618,1231,1257]" latitude="-26.116" longLatPrecision="74" longitude="130.283" name="Piltadi" pageId="12" pageNumber="165" stateProvince="South Australia">Piltadi</location>
|
||
,
|
||
<collectingRegion id="5B22AD8AFA65FFD8D6492195FC97CB85" box="[630,803,1231,1257]" country="Australia" name="South Australia" pageId="12" pageNumber="165">South Australia</collectingRegion>
|
||
(
|
||
<geoCoordinate id="FCD205AFFA65FFD8D70D2195FC3FCB85" box="[818,907,1231,1257]" degrees="26.116" direction="south" orientation="latitude" pageId="12" pageNumber="165" precision="55" value="-26.116">-26.116</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FCD205AFFA65FFD8D7A82195FC43CB85" box="[919,1015,1231,1257]" degrees="130.283" direction="east" orientation="longitude" pageId="12" pageNumber="165" precision="55" value="130.283">130.283</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="99596368FA65FFD8D4F821A8FECBCA5D" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="165">
|
||
<emphasis id="AB92BF7AFA65FFD8D4F821A8FE1FCA60" bold="true" box="[199,427,1266,1292]" pageId="12" pageNumber="165">Referred Material:</emphasis>
|
||
Details of wild specimens from which supplementary data were obtained are provided in
|
||
<tableCitation id="D46456D3FA65FFD8D4A8204DFF4ACA5D" box="[151,254,1303,1329]" captionStart="TABLE 1" captionStartId="5.[152,244,585,609]" captionTargetPageId="5" captionText="TABLE 1. Summary statistics (mean ± SD, range in parentheses) of unadjusted mensural (millimetres) and meristic (count) characters for each species." pageId="12" pageNumber="165">Table A1</tableCitation>
|
||
, Appendix.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA65FFD8D4F82060FF69C919" pageId="12" pageNumber="165" type="diagnosis">
|
||
<paragraph id="99596368FA65FFD8D4F82060FF69C919" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="165">
|
||
<emphasis id="AB92BF7AFA65FFD8D4F82060FEF6CA38" bold="true" box="[199,322,1338,1364]" pageId="12" pageNumber="165">Diagnosis.</emphasis>
|
||
<taxonomicName id="5EE618EBFA65FFD8D5782061FE4BCA38" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[327,511,1339,1365]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA65FFD8D5782061FE4BCA38" box="[327,511,1339,1365]" italics="true" pageId="12" pageNumber="165">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA65FFD8D63B2061FDE9CA39" bold="true" box="[516,605,1339,1365]" pageId="12" pageNumber="165">
|
||
<taxonomicNameLabel id="B0A10201FA65FFD8D63B2061FDE9CA39" box="[516,605,1339,1365]" pageId="12" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from other members of the genus by a combination of the following characters: moderately large size (maximum SVL up to
|
||
<quantity id="5E1ECE8DFA65FFD8D7982004FBBCCA15" box="[935,1032,1374,1401]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" pageId="12" pageNumber="165" unit="mm" value="135.0">135 mm</quantity>
|
||
); snout relatively pointed—distance between outer edges of parietals usually no more than the distance from the snout to the posterior edge of the frontal; dorsal scales smooth; conspicuous black callus along free edge of subdigital lamellae; palmar and plantar scales each with a prominent black tubercle. Usually six supraciliary scales; usually four enlarged lobules projecting from anterior border of ear opening. Dorsal surfaces dull orangish-brown, typically with pair of broad diffuse cream dorsolateral stripes and faint indications of five longitudinally aligned black dorsal stripes; head brigher orangish-brown than body; chin and throat salmon orange and without dark marking; ventral surface uniform creamish-white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA65FFD9D4F82324FE7FCA39" lastPageId="13" lastPageNumber="166" pageId="12" pageNumber="165" type="description">
|
||
<paragraph id="99596368FA65FFD8D4F82324FD86C845" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="165">
|
||
<emphasis id="AB92BF7AFA65FFD8D4F82324FEEDC9F4" bold="true" box="[199,345,1662,1688]" pageId="12" pageNumber="165">Description.</emphasis>
|
||
Full summary statistics of morphology data are presented in
|
||
<tableCitation id="D46456D3FA65FFD8D0302325FB37C9F5" box="[1039,1155,1663,1689]" captionStart="TABLE 1" captionStartId="5.[152,244,585,609]" captionTargetPageId="5" captionText="TABLE 1. Summary statistics (mean ± SD, range in parentheses) of unadjusted mensural (millimetres) and meristic (count) characters for each species." pageId="12" pageNumber="165">Table 1. A</tableCitation>
|
||
moderately large, robust
|
||
<taxonomicName id="5EE618EBFA65FFD8D4A823F9FEEBC9D1" box="[151,351,1699,1725]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="AB92BF7AFA65FFD8D4A823F9FEB5C9D1" box="[151,257,1699,1725]" italics="true" pageId="12" pageNumber="165">Liopholis</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Measurements (as means with range in parentheses) of all specimens: SVL
|
||
<quantity id="5E1ECE8DFA65FFD8D09C23F9FAA0C9D1" box="[1187,1300,1699,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.011" pageId="12" pageNumber="165" unit="mm" value="101.1">101.1 mm</quantity>
|
||
(60.2–135); HL
|
||
<quantity id="5E1ECE8DFA65FFD8D4FB239DFE9DC98D" box="[196,297,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="12" pageNumber="165" unit="mm" value="19.2">19.2 mm</quantity>
|
||
(14.2–20.9); HW
|
||
<quantity id="5E1ECE8DFA65FFD8D5C6239DFDEBC98D" box="[505,607,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="12" pageNumber="165" unit="mm" value="14.2">14.2 mm</quantity>
|
||
(10.7–15.4); HD
|
||
<quantity id="5E1ECE8DFA65FFD8D716239DFC35C98D" box="[809,897,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="12" pageNumber="165" unit="mm" value="9.7">9.7 mm</quantity>
|
||
(7.3–10.7); AGL
|
||
<quantity id="5E1ECE8DFA65FFD8D073239DFB05C98D" box="[1100,1201,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="4.74" pageId="12" pageNumber="165" unit="mm" value="47.4">47.4 mm</quantity>
|
||
(31–62.7); BW
|
||
<quantity id="5E1ECE8DFA65FFD8D154239DFF77C869" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="12" pageNumber="165" unit="mm" value="17.8">17.8 mm</quantity>
|
||
(12.9–19.6); FLL
|
||
<quantity id="5E1ECE8DFA65FFD8D5AC23B0FE4CC869" box="[403,504,1770,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="165" unit="mm" value="25.4">25.4 mm</quantity>
|
||
(18.1–29.7); HLL
|
||
<quantity id="5E1ECE8DFA65FFD8D6F323B1FC85C869" box="[716,817,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="3.41" pageId="12" pageNumber="165" unit="mm" value="34.1">34.1 mm</quantity>
|
||
(25–39.8). The original tail length (TL) of adults (>
|
||
<quantity id="5E1ECE8DFA65FFD8D1BF23B0FF77C845" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="165" unit="mm" value="75.0">75 mm</quantity>
|
||
SVL) is
|
||
<quantity id="5E1ECE8DFA65FFD8D5162254FE2FC845" box="[297,411,1806,1833]" metricMagnitude="-1" metricUnit="m" metricValue="1.5059999999999998" pageId="12" pageNumber="165" unit="mm" value="150.6">150.6 mm</quantity>
|
||
(130.5–158).
|
||
</paragraph>
|
||
<paragraph id="99596368FA65FFD9D4F82269FBC0CF95" blockId="12.[151,1437,942,2013]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="166" pageId="12" pageNumber="165">
|
||
<emphasis id="AB92BF7AFA65FFD8D4F82269FEBFC821" box="[199,267,1843,1869]" italics="true" pageId="12" pageNumber="165">Head.</emphasis>
|
||
Head dorsoventrally depressed (head depth is 48%–52% of head length,
|
||
<emphasis id="AB92BF7AFA65FFD8D05B226EFBC7C823" box="[1124,1139,1844,1871]" italics="true" pageId="12" pageNumber="165">x̄</emphasis>
|
||
= 50%). Snout relatively pointed (head length is 132%–138% of head width,
|
||
<emphasis id="AB92BF7AFA65FFD8D6EB2202FD57C81F" box="[724,739,1880,1907]" italics="true" pageId="12" pageNumber="165">x̄</emphasis>
|
||
= 135%) and relatively shallow. Supralabials 8–10 (usually 9); supraciliaries 6–7 (usually 6); ciliaries 10–13 (usually 12); ear opening large, oblong and oriented vertically, with 3– 5 (usually 4) enlarged lobules on the anterior margin, projecting over opening, often with a single additional tiny ear lobule above the enlarged series, ear lobule shape variable, being either obtuse, rectangular or acute; typically only one pair of enlarged nuchal scales (two pairs in SAMA R60802); nasal scales separated, moderately to widely so; post-narial groove absent; prefrontals in broad contact, occasionally in narrow contact; neck only slightly narrower than widest part of head (above tympani); lower eyelid movable; parietal scales divided.
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4F82459FC30CE2D" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82459FEB3CE71" box="[199,263,259,285]" italics="true" pageId="13" pageNumber="166">Body.</emphasis>
|
||
Moderately robust body (axilla-groin length is 48%–61% of SVL,
|
||
<emphasis id="AB92BF7AFA64FFD9D03A245EFBA0CE73" box="[1029,1044,260,287]" italics="true" pageId="13" pageNumber="166">x̄</emphasis>
|
||
= 52%) with smooth, overlapping scales in 32–39 rows (MSR), but usually 36–38 (rarely 32 and 39).
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4F82411FBECCEBD" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82411FEA7CE09" box="[199,275,331,357]" italics="true" pageId="13" pageNumber="166">Limbs.</emphasis>
|
||
Limbs of moderate length (forelimb is 27%–30% of SVL,
|
||
<emphasis id="AB92BF7AFA64FFD9D78C2416FC76CE0B" box="[947,962,332,359]" italics="true" pageId="13" pageNumber="166">x̄</emphasis>
|
||
= 28%; hindlimb is 34%–42% of SVL,
|
||
<emphasis id="AB92BF7AFA64FFD9D1B22416FA28CE0B" box="[1421,1436,332,359]" italics="true" pageId="13" pageNumber="166">x̄</emphasis>
|
||
= 38%); digits short and well-developed; finger length: 4>3>2>5>1, toe length: 4>3>5>2>1; subdigital lamellae 22–28 (usually 26) under fourth toe, with conspicuous black callus along free edge of lamellae; claws sharp and recurved; palmar and plantar scales each with prominent black callus along free edge.
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4F82481FEA3CD75" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82481FF4DCE99" box="[199,249,475,501]" italics="true" pageId="13" pageNumber="166">Tail.</emphasis>
|
||
Original tail long (128%–176% of SVL,
|
||
<emphasis id="AB92BF7AFA64FFD9D6E92486FD51CE9B" box="[726,741,476,503]" italics="true" pageId="13" pageNumber="166">x̄</emphasis>
|
||
= 157%;
|
||
<emphasis id="AB92BF7AFA64FFD9D7652481FCDCCE98" box="[858,872,475,500]" italics="true" pageId="13" pageNumber="166">n</emphasis>
|
||
= 4) and approximately cylindrical, tapering to a pointed tip.
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4F82779FAF6CCA5" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82779FDDBCD51" box="[199,623,547,573]" italics="true" pageId="13" pageNumber="166">
|
||
Colour pattern in life (
|
||
<figureCitation id="01DD7FEDFA64FFD9D5F92779FDD2CD51" box="[454,614,547,573]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figures 5A–D</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: (
|
||
<emphasis id="AB92BF7AFA64FFD9D6B82779FD21CD50" box="[647,661,547,572]" italics="true" pageId="13" pageNumber="166">n</emphasis>
|
||
= 8) Background colouration of the dorsal and lateral surfaces of the body, limbs and tail typically dull orangish-brown with thin black scale margins; top and sides of head usually more vibrant orangish-brown than body. Ciliary scales bright yellow; ear lobules pale cream. Ventral surfaces uniform creamish-white; chin and throat salmon orange and without dark markings. Patterning of the head, dorsum, limbs and flanks is variable (see ‘
|
||
<emphasis id="AB92BF7AFA64FFD9D5A927E9FE4ACDA0" box="[406,510,691,716]" italics="true" pageId="13" pageNumber="166">Variation</emphasis>
|
||
’ section below). On top of head there is often irregular black markings, typically on scale margins; lateral head markings often include black vertical bars between some of the supralabial scales, and a black streak running from about the loreal region, through eye and extending along the suture of the upper and lower secondary temporals, and terminating on the tertiary temporals. Irregular pale blotches may be present in labial region. Usually there is a pair of broad diffuse cream dorsolateral stripes, and faint indications of five longitudinally aligned black dorsal stripes, commencing on the nape and extending to tail base; these stripes typically are more distinct anteriorly and often become diffuse posteriorly. Lateral patterning consists of scattered white spots, with dark anterior margins. Tail and limb colour patterning is generally similar to that of the dorsum and flanks.
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4F82689FB46CB35" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82689FD1FCC81" box="[199,683,979,1005]" italics="true" pageId="13" pageNumber="166">
|
||
Colour pattern in preservative (
|
||
<figureCitation id="01DD7FEDFA64FFD9D6022689FD10CC81" box="[573,676,979,1005]" captionStart="FIGURE 4" captionStartId="8.[152,255,1283,1307]" captionTargetBox="[151,1436,179,1257]" captionTargetId="figure-31@8.[151,1436,179,1257]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Liopholis aputja sp. nov. (SAMA R60802) from Alalkanya Gorge, South Australia. Photos: Jules Farquhar." figureDoi="http://doi.org/10.5281/zenodo.14024204" httpUri="https://zenodo.org/record/14024204/files/figure.png" pageId="13" pageNumber="166">Figure 4</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: Colour pattern in preservative (
|
||
<emphasis id="AB92BF7AFA64FFD9D00E2689FB8BCC80" box="[1073,1087,979,1004]" italics="true" pageId="13" pageNumber="166">n</emphasis>
|
||
= 4) is largely similar to the description in life but with slightly to significantly faded and less vibrant colouration overall. The salmon orange chin and throat colour (present in live specimens) is absent in all preserved specimens. The characteristic black calli of the subdigital lamellae, plantar and palmar scales is substantially faded or absent in preservative.
|
||
</paragraph>
|
||
<paragraph id="99596368FA64FFD9D4C82139FE7FCA39" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4C82139FD9FCB10" box="[247,555,1123,1149]" italics="true" pageId="13" pageNumber="166">
|
||
Variation:
|
||
<taxonomicName id="5EE618EBFA64FFD9D54E2139FD9FCB10" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[369,555,1123,1149]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">Liopholis aputja</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="AB92BF7AFA64FFD9D60C2139FD3ACB11" bold="true" box="[563,654,1123,1149]" pageId="13" pageNumber="166">
|
||
<taxonomicNameLabel id="B0A10201FA64FFD9D60C2139FD3ACB11" box="[563,654,1123,1149]" pageId="13" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is variable in pattern, ranging from heavily patterned (
|
||
<figureCitation id="01DD7FEDFA64FFD9D0C02139FAC2CB11" box="[1279,1398,1122,1149]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figure 5A</figureCitation>
|
||
) to almost completely patternless (
|
||
<figureCitation id="01DD7FEDFA64FFD9D5C821DDFDDACBCD" box="[503,622,1158,1185]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figure 5C</figureCitation>
|
||
), and this variability occurs within the same location. In well-patterned individuals, there are scattered white spots with dark anterior margins on the dorsum, flanks, limbs and tail (
|
||
<figureCitation id="01DD7FEDFA64FFD9D16E21F1FF69CB85" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figure 5A, D</figureCitation>
|
||
); but for some intermediate (
|
||
<figureCitation id="01DD7FEDFA64FFD9D6232195FD3ACB85" box="[540,654,1230,1257]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figure 5B</figureCitation>
|
||
) to weakly patterned (
|
||
<figureCitation id="01DD7FEDFA64FFD9D7BB2195FC42CB85" box="[900,1014,1230,1257]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="13" pageNumber="166">Figure 5C</figureCitation>
|
||
) specimens, these spots occur only on the flanks, or are absent. In pattern-reduced specimens, head patterning may be absent, or at least consists of a black streak in front of and behind eye. In patterned-reduced specimens, the five thin black dorsal lines may be absent, or perceivable only anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA64FFD9D4F82005FD63C845" pageId="13" pageNumber="166" type="etymology">
|
||
<paragraph id="99596368FA64FFD9D4F82005FD63C845" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82005FEFACA15" bold="true" box="[199,334,1375,1401]" pageId="13" pageNumber="166">Etymology.</emphasis>
|
||
Aputja is a word from the Pitjantjatjara/Yankunytjatjara language spoken by First Nations peoples where this lizard species is found in APY (Aṉangu Pitjantjatjara Yankunytjatjara) Lands of north-western
|
||
<collectingRegion id="5B22AD8AFA64FFD9D16420D9FEB5CAAD" country="Australia" name="South Australia" pageId="13" pageNumber="166">South Australia</collectingRegion>
|
||
. Aputja means ‘of the hills’, in reference to this species occurring in the hills and gorges of the Mann-Musgrave Ranges, unlike other
|
||
<taxonomicName id="5EE618EBFA64FFD9D6352091FD18CA89" box="[522,684,1483,1509]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="AB92BF7AFA64FFD9D6352091FDC0CA89" box="[522,628,1483,1509]" italics="true" pageId="13" pageNumber="166">Liopholis</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the region which occur on sandplains. The specific epithet is constructed as a noun in apposition. In arriving at this name, we spoke to rangers, Traditional Owners and knowledge holders across the APY Lands; in Pukatja, Fregon, Mimili/Amaroona, Indulkana, Amata and Pipalyatjara. Responses to asking for the name of the new
|
||
<taxonomicName id="5EE618EBFA64FFD9D62F236DFD65C93D" box="[528,721,1591,1617]" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D62F236DFDCEC93D" box="[528,634,1591,1617]" italics="true" pageId="13" pageNumber="166">Liopholis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
varied. However, particular clarity in narrowing the name down to Aputja was provided by the following Traditional Owners: Allan Wilson (Indulkana/Pukatja), Johnny Roberts (Fregon), Winmati Roberts (Fregon/Umuwa), Manyiritjanu Lennon (Pukatja), and Hughie Cullinan (Mimili/ Amaroona). The common name Central Ranges Rock Skink is chosen because
|
||
<taxonomicName id="5EE618EBFA64FFD9D01423F9FB27C9D0" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[1067,1171,1699,1724]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA64FFD9D01423F9FB27C9D0" box="[1067,1171,1699,1724]" italics="true" pageId="13" pageNumber="166">L. aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA64FFD9D0A323F9FB4CC9D1" bold="true" box="[1180,1272,1699,1725]" pageId="13" pageNumber="166">
|
||
<taxonomicNameLabel id="B0A10201FA64FFD9D0A323F9FB4CC9D1" box="[1180,1272,1699,1725]" pageId="13" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is endemic to the Central Ranges bioregion (
|
||
<bibRefCitation id="FD771E99FA64FFD9D5C1239DFCE0C98D" author="Thackway, R. & Cresswell, I. D." box="[510,852,1734,1761]" pageId="13" pageNumber="166" refId="ref15114" refString="Thackway, R. & Cresswell, I. D. (1995) An Interim Biogeographic Regionalisation for Australia: a framework for establishing the national system of reserves. Version 4.0. Australian Nature Conservation Agency, Canberra, 88 pp. Available from: https: // www. dcceew. gov. au / sites / default / files / documents / ibra-framework-setting-priorities-nrs-cooperative-program. pdf" type="book" year="1995">Thackway & Cresswell 1995</bibRefCitation>
|
||
). We chose the words ‘rock skink’ for clarity and consistency, given that most other saxicoline members of the
|
||
<taxonomicName id="5EE618EBFA64FFD9D76123B1FC7CC869" authorityName="Fitzinger" authorityYear="1843" box="[862,968,1771,1797]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="genus">
|
||
<emphasis id="AB92BF7AFA64FFD9D76123B1FC7CC869" box="[862,968,1771,1797]" italics="true" pageId="13" pageNumber="166">Liopholis</emphasis>
|
||
</taxonomicName>
|
||
genus are commonly referred to as rock skinks (Australian Society of Herpetologists 2023).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA64FFDAD4F82268FB62CEC1" lastPageId="14" lastPageNumber="167" pageId="13" pageNumber="166" type="description">
|
||
<paragraph id="99596368FA64FFDAD4F82268FB62CEC1" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1437,150,1113]" lastPageId="14" lastPageNumber="167" pageId="13" pageNumber="166">
|
||
<emphasis id="AB92BF7AFA64FFD9D4F82268FDCFC821" bold="true" box="[199,635,1842,1869]" pageId="13" pageNumber="166">
|
||
Comparison with similar
|
||
<taxonomicName id="5EE618EBFA64FFD9D6382268FDC0C821" authorityName="Fitzinger" authorityYear="1843" box="[519,628,1842,1869]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="genus">
|
||
<emphasis id="AB92BF7AFA64FFD9D6382268FDC0C821" bold="true" box="[519,628,1842,1869]" italics="true" pageId="13" pageNumber="166">Liopholis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="5EE618EBFA64FFD9D6B82269FCF1C820" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[647,837,1843,1869]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA64FFD9D6B82269FCF1C820" box="[647,837,1843,1869]" italics="true" pageId="13" pageNumber="166">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA64FFD9D76E2269FC1BC821" bold="true" box="[849,943,1843,1869]" pageId="13" pageNumber="166">
|
||
<taxonomicNameLabel id="B0A10201FA64FFD9D76E2269FC1BC821" box="[849,943,1843,1869]" pageId="13" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most closely related to
|
||
<taxonomicName id="5EE618EBFA64FFD9D0CD2269FA22C820" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[1266,1430,1843,1868]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="AB92BF7AFA64FFD9D0CD2269FA22C820" box="[1266,1430,1843,1868]" italics="true" pageId="13" pageNumber="166">L. margaretae</emphasis>
|
||
</taxonomicName>
|
||
, from which it is allopatric and genetically distinct (net between group mean distance of 12.4% in the ND4 gene).
|
||
<taxonomicName id="5EE618EBFA64FFD9D4A82221FEE4C8F8" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[151,336,1915,1941]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA64FFD9D4A82221FEE4C8F8" box="[151,336,1915,1941]" italics="true" pageId="13" pageNumber="166">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA64FFD9D5682221FE05C8F9" bold="true" box="[343,433,1915,1941]" pageId="13" pageNumber="166">
|
||
<taxonomicNameLabel id="B0A10201FA64FFD9D5682221FE05C8F9" box="[343,433,1915,1941]" pageId="13" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="5EE618EBFA64FFD9D6792221FD51C8F8" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[582,741,1915,1940]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="13" pageNumber="166" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="AB92BF7AFA64FFD9D6792221FD51C8F8" box="[582,741,1915,1940]" italics="true" pageId="13" pageNumber="166">L. margaretae</emphasis>
|
||
</taxonomicName>
|
||
as follows: in having only the head (and sometimes the neck) a brighter orangish-brown than the rest of the body (versus the entire anterior half of the body reddish-brown); in more frequently exhibiting intermediate forms with varying degrees of pattern prominence (versus an almost discrete dichotomous separation between patterned and unpatterned morphs (
|
||
<figureCitation id="01DD7FEDFA67FFDAD02C25CDFBCDCFDD" box="[1043,1145,150,177]" captionStart="FIGURE 5" captionStartId="9.[152,255,1375,1399]" captionTargetBox="[164,1423,179,1349]" captionTargetId="figure-21@9.[164,1423,179,1349]" captionTargetPageId="9" captionText="FIGURE 5. Colour pattern variation in Liopholis aputja sp. nov. from Aliwanyiwanyinya Rockhole, SA (A, B); Apara Springs, SA (C); Alalkanya Gorge, SA (D, GenBank: PQ067254); and comparison with L. margaretae patterned morph at Ormiston Gorge, NT (E, GenBank: PQ067258) and plain morph at Kings Canyon, NT (F, GenBank: PQ067257). Photos: Jordan Mulder (A–C); Jules Farquhar (D–F)." figureDoi="http://doi.org/10.5281/zenodo.14024206" httpUri="https://zenodo.org/record/14024206/files/figure.png" pageId="14" pageNumber="167">Figure 5</figureCitation>
|
||
); in lacking black lateral spots which tend to form oblique rows directed up and back (versus sometimes present); in always lacking dark suture markings under chin (versus sometimes present); in having a pair of broad diffuse cream dorsolateral stripes and faint indications of five longitudinally aligned black dorsal stripes (versus absence of such stripes). For meristic characters,
|
||
<taxonomicName id="5EE618EBFA67FFDAD523247DFE32CE2C" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[284,390,295,320]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD523247DFE32CE2C" box="[284,390,295,320]" italics="true" pageId="14" pageNumber="167">L. aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD5AD247DFE44CE2D" bold="true" box="[402,496,295,321]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD5AD247DFE44CE2D" box="[402,496,295,321]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has on average more midbody rows, subdigital lamellae, supralabials, ciliaries, ear lobules, but less supraciliaries (
|
||
<figureCitation id="01DD7FEDFA67FFDAD6132411FD10CE09" box="[556,676,331,357]" captionStart="FIGURE 3" captionStartId="7.[152,255,832,856]" captionTargetBox="[157,1427,187,798]" captionTargetId="figure-21@7.[151,1436,179,806]" captionTargetPageId="7" captionText="FIGURE 3. Meristic character variation between lineages based on n = 59 specimens.(A) Biplot of non-metric multidimensional scaling (nMDS) performed on meristic characters. Specimens (coloured points) that are ordinated closer to one another are phenotypically more similar than those further apart. Holotype specimens are denoted by ‘H’ above data points. (B) Comparative boxplots of meristic characters showing the mean (white dot), median (thick grey horizontal line) range, and inter-quartile range (coloured box). Asterisks denote characters bearing statistically significant (GLM) differences in expected counts." figureDoi="http://doi.org/10.5281/zenodo.14024200" httpUri="https://zenodo.org/record/14024200/files/figure.png" pageId="14" pageNumber="167">Figure 3B</figureCitation>
|
||
;
|
||
<figureCitation id="01DD7FEDFA67FFDAD68C2411FCA1CE09" box="[691,789,331,357]" captionStart="FIGURE 6" captionStartId="10.[152,255,1085,1109]" captionTargetBox="[155,1431,183,1055]" captionTargetId="figure-31@10.[151,1435,179,1059]" captionTargetPageId="10" captionText="FIGURE 6. Comparison of diagnostic scalation between species. Liopholis aputja sp. nov. tend to have fewer supraciliary scales and more ear lobules than L. margaretae. (A) Holotype (SAMA R60802) of L. aputja sp. nov. with six supraciliary scales and five enlarged ear lobules. (B) Liopholis margaretae (GenBank: PQ067257) with eight supraciliary scales and three enlarged ear lobules. Illustrations by Christina Hunter." figureDoi="http://doi.org/10.5281/zenodo.14024211" httpUri="https://zenodo.org/record/14024211/files/figure.png" pageId="14" pageNumber="167">Figure 6</figureCitation>
|
||
;
|
||
<tableCitation id="D46456D3FA67FFDAD71B2411FCCDCE09" box="[804,889,331,357]" captionStart="TABLE 1" captionStartId="5.[152,244,585,609]" captionTargetPageId="5" captionText="TABLE 1. Summary statistics (mean ± SD, range in parentheses) of unadjusted mensural (millimetres) and meristic (count) characters for each species." pageId="14" pageNumber="167">Table 1</tableCitation>
|
||
). While multiple features should be considered simultaneously for identification, supraciliary counts are particularly reliable to distinguish these two species, given
|
||
<taxonomicName id="5EE618EBFA67FFDAD4A824C9FF49CEC0" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[151,253,403,428]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD4A824C9FF49CEC0" box="[151,253,403,428]" italics="true" pageId="14" pageNumber="167">L. aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD53C24C9FEE9CEC1" bold="true" box="[259,349,403,429]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD53C24C9FEE9CEC1" box="[259,349,403,429]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
usually (84%) have 6, whereas
|
||
<taxonomicName id="5EE618EBFA67FFDAD6FC24C9FCD6CEC0" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[707,866,403,428]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="AB92BF7AFA67FFDAD6FC24C9FCD6CEC0" box="[707,866,403,428]" italics="true" pageId="14" pageNumber="167">L. margaretae</emphasis>
|
||
</taxonomicName>
|
||
usually (74%) have 8 (
|
||
<figureCitation id="01DD7FEDFA67FFDAD05924C9FB7CCEC1" box="[1126,1224,403,429]" captionStart="FIGURE 6" captionStartId="10.[152,255,1085,1109]" captionTargetBox="[155,1431,183,1055]" captionTargetId="figure-31@10.[151,1435,179,1059]" captionTargetPageId="10" captionText="FIGURE 6. Comparison of diagnostic scalation between species. Liopholis aputja sp. nov. tend to have fewer supraciliary scales and more ear lobules than L. margaretae. (A) Holotype (SAMA R60802) of L. aputja sp. nov. with six supraciliary scales and five enlarged ear lobules. (B) Liopholis margaretae (GenBank: PQ067257) with eight supraciliary scales and three enlarged ear lobules. Illustrations by Christina Hunter." figureDoi="http://doi.org/10.5281/zenodo.14024211" httpUri="https://zenodo.org/record/14024211/files/figure.png" pageId="14" pageNumber="167">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA67FFDAD4F824EDFC80CCED" pageId="14" pageNumber="167" type="reference_group">
|
||
<paragraph id="99596368FA67FFDAD4F824EDFDCDCDE9" blockId="14.[151,1437,150,1113]" pageId="14" pageNumber="167">
|
||
While
|
||
<taxonomicName id="5EE618EBFA67FFDAD52D24EDFEC2CEBC" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[274,374,439,464]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD52D24EDFEC2CEBC" box="[274,374,439,464]" italics="true" pageId="14" pageNumber="167">L. aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD54324EDFE61CEBD" bold="true" box="[380,469,439,465]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD54324EDFE61CEBD" box="[380,469,439,465]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the only rock
|
||
<emphasis id="AB92BF7AFA67FFDAD6BA24EDFD3ACEBD" bold="true" box="[645,654,439,465]" pageId="14" pageNumber="167">-</emphasis>
|
||
dwelling
|
||
<taxonomicName id="5EE618EBFA67FFDAD6C824EDFCD5CEBD" authorityName="Fitzinger" authorityYear="1843" box="[759,865,439,465]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="genus">
|
||
<emphasis id="AB92BF7AFA67FFDAD6C824EDFCD5CEBD" box="[759,865,439,465]" italics="true" pageId="14" pageNumber="167">Liopholis</emphasis>
|
||
</taxonomicName>
|
||
occurring in the Mann-Musgrave Ranges, there are several congeners recorded in the desert sand plains adjoining these ranges:
|
||
<taxonomicName id="5EE618EBFA67FFDAD7C72481FAACCE99" authority="(Rosen 1905)" baseAuthorityName="Rosen" baseAuthorityYear="1905" box="[1016,1304,474,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="inornata">
|
||
<emphasis id="AB92BF7AFA67FFDAD7C72481FBBCCE98" box="[1016,1032,475,500]" italics="true" pageId="14" pageNumber="167">L</emphasis>
|
||
.
|
||
<emphasis id="AB92BF7AFA67FFDAD0282481FBC3CE98" box="[1047,1143,475,500]" italics="true" pageId="14" pageNumber="167">inornata</emphasis>
|
||
(Rosén 1905)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5EE618EBFA67FFDAD1182481FA28CE99" baseAuthorityName="Stirling & Zietz" baseAuthorityYear="1893" box="[1319,1436,475,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="kintorei">
|
||
<emphasis id="AB92BF7AFA67FFDAD1182481FA28CE99" box="[1319,1436,475,501]" italics="true" pageId="14" pageNumber="167">L. kintorei</emphasis>
|
||
</taxonomicName>
|
||
(Stirling & Zietz 1893),
|
||
<taxonomicName id="5EE618EBFA67FFDAD58D24A5FD05CD75" authority="(Storr 1968)" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[434,689,511,537]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="slateri">
|
||
<emphasis id="AB92BF7AFA67FFDAD58D24A5FDAECD75" box="[434,538,511,537]" italics="true" pageId="14" pageNumber="167">L. slateri</emphasis>
|
||
(
|
||
<bibRefCitation id="FD771E99FA67FFDAD61424A5FD1ECD75" author="Storr, G. M." box="[555,682,511,537]" pageId="14" pageNumber="167" pagination="51 - 62" refId="ref15040" refString="Storr, G. M. (1968) Revision of the Egernia whitei species-group (Lacertilia, Scincidae). J ournal of the Royal Society of Western Australia, 51 (2), 51 - 62." type="journal article" year="1968">Storr 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="5EE618EBFA67FFDAD6CD24A5FB96CD75" authority="(Sternfeld 1919)" baseAuthorityName="Sternfeld" baseAuthorityYear="1919" box="[754,1058,511,537]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="striata">
|
||
<emphasis id="AB92BF7AFA67FFDAD6CD24A5FCEFCD74" box="[754,859,511,536]" italics="true" pageId="14" pageNumber="167">L. striata</emphasis>
|
||
(Sternfeld 1919)
|
||
</taxonomicName>
|
||
. Unlike
|
||
<taxonomicName id="5EE618EBFA67FFDAD0BA24A5FB59CD74" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[1157,1261,511,536]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD0BA24A5FB59CD74" box="[1157,1261,511,536]" italics="true" pageId="14" pageNumber="167">L. aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD0C924A5FAE6CD75" bold="true" box="[1270,1362,511,537]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD0C924A5FAE6CD75" box="[1270,1362,511,537]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, these species have blunt heads (i.e. distance between outer edges of parietals usually greater than the distance from the snout to the hind edge of frontal scale), and lack the conspicuous black callus along free edge of the subdigital lamellae and the palmar and plantar scales.
|
||
</paragraph>
|
||
<paragraph id="99596368FA67FFDAD4F827D5FC80CCED" blockId="14.[151,1437,150,1113]" pageId="14" pageNumber="167">
|
||
<taxonomicName id="5EE618EBFA67FFDAD4F827D5FE36CDC4" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[199,386,655,681]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD4F827D5FE36CDC4" box="[199,386,655,681]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD5B427D5FE53CDC5" bold="true" box="[395,487,655,681]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD5B427D5FE53CDC5" box="[395,487,655,681]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="5EE618EBFA67FFDAD5C727D5FD2DCDC4" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[504,665,655,680]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="AB92BF7AFA67FFDAD5C727D5FD2DCDC4" box="[504,665,655,680]" italics="true" pageId="14" pageNumber="167">L. margaretae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="AB92BF7AFA67FFDAD69627D5FC8BCDC4" box="[681,831,655,680]" italics="true" pageId="14" pageNumber="167">
|
||
<taxonomicName id="5EE618EBFA67FFDAD69627D5FC88CDC4" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[681,828,655,680]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="personata">L. personata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="5EE618EBFA67FFDAD74427D5FC4CCDC5" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[891,1016,655,681]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="modesta">
|
||
<emphasis id="AB92BF7AFA67FFDAD74427D5FC4CCDC5" box="[891,1016,655,681]" italics="true" pageId="14" pageNumber="167">L. modesta</emphasis>
|
||
</taxonomicName>
|
||
are geographically widely separated from one another, but all share rock-dwelling habits.
|
||
<taxonomicName id="5EE618EBFA67FFDAD6D927E9FC2ACDA0" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[742,926,691,717]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD6D927E9FC2ACDA0" box="[742,926,691,717]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD79B27E9FC49CDA1" bold="true" box="[932,1021,691,717]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD79B27E9FC49CDA1" box="[932,1021,691,717]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
usually has 38 or less MSR, whereas
|
||
<taxonomicName id="5EE618EBFA67FFDAD4A8278DFE91CD9C" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[151,293,727,752]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="personata">
|
||
<emphasis id="AB92BF7AFA67FFDAD4A8278DFE91CD9C" box="[151,293,727,752]" italics="true" pageId="14" pageNumber="167">L. personata</emphasis>
|
||
</taxonomicName>
|
||
has 38 or more (
|
||
<bibRefCitation id="FD771E99FA67FFDAD5DB278DFDD5CD9D" author="Storr, G. M." box="[484,609,727,753]" pageId="14" pageNumber="167" pagination="51 - 62" refId="ref15040" refString="Storr, G. M. (1968) Revision of the Egernia whitei species-group (Lacertilia, Scincidae). J ournal of the Royal Society of Western Australia, 51 (2), 51 - 62." type="journal article" year="1968">Storr 1968</bibRefCitation>
|
||
).
|
||
<taxonomicName id="5EE618EBFA67FFDAD64B278DFC9ACD9C" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[628,814,727,753]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD64B278DFC9ACD9C" box="[628,814,727,753]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD70A278DFC3ACD9D" bold="true" box="[821,910,727,753]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD70A278DFC3ACD9D" box="[821,910,727,753]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
almost always has 6 supraciliaries, whereas
|
||
<taxonomicName id="5EE618EBFA67FFDAD1B9278DFF40CC79" baseAuthorityName="Storr" baseAuthorityYear="1968" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="modesta">
|
||
<emphasis id="AB92BF7AFA67FFDAD1B9278DFF40CC79" italics="true" pageId="14" pageNumber="167">L. modesta</emphasis>
|
||
</taxonomicName>
|
||
usually has 7 or 8 (
|
||
<bibRefCitation id="FD771E99FA67FFDAD5F527A1FDF2CC79" author="Storr, G. M." box="[458,582,763,789]" pageId="14" pageNumber="167" pagination="51 - 62" refId="ref15040" refString="Storr, G. M. (1968) Revision of the Egernia whitei species-group (Lacertilia, Scincidae). J ournal of the Royal Society of Western Australia, 51 (2), 51 - 62." type="journal article" year="1968">Storr 1968</bibRefCitation>
|
||
).
|
||
<taxonomicName id="5EE618EBFA67FFDAD66627A1FCA5CC78" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[601,785,763,789]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD66627A1FCA5CC78" box="[601,785,763,789]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD72927A1FCDBCC79" bold="true" box="[790,879,763,789]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD72927A1FCDBCC79" box="[790,879,763,789]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="5EE618EBFA67FFDAD74327A1FBAECC78" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[892,1050,763,788]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="AB92BF7AFA67FFDAD74327A1FBAECC78" box="[892,1050,763,788]" italics="true" pageId="14" pageNumber="167">L. margaretae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="5EE618EBFA67FFDAD06A27A1FB56CC78" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[1109,1250,763,788]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="personata">
|
||
<emphasis id="AB92BF7AFA67FFDAD06A27A1FB56CC78" box="[1109,1250,763,788]" italics="true" pageId="14" pageNumber="167">L. personata</emphasis>
|
||
</taxonomicName>
|
||
are superficially similar in morphology and all share the character of conspicuous black calluses on subdigital lamellae, palmar, and plantar scales, whereas
|
||
<taxonomicName id="5EE618EBFA67FFDAD5A12619FDACCC31" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[414,536,835,861]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="modesta">
|
||
<emphasis id="AB92BF7AFA67FFDAD5A12619FDACCC31" box="[414,536,835,861]" italics="true" pageId="14" pageNumber="167">L. modesta</emphasis>
|
||
</taxonomicName>
|
||
lacks this scalation and has cream facial and axillary spots (shared by the
|
||
<taxonomicName id="5EE618EBFA67FFDAD16C2619FA27CC31" baseAuthorityName="Lacépède" baseAuthorityYear="1804" box="[1363,1427,835,861]" class="Squamata" family="Scincidae" genus="Egernia" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="whitii">
|
||
<emphasis id="AB92BF7AFA67FFDAD16C2619FA27CC31" box="[1363,1427,835,861]" italics="true" pageId="14" pageNumber="167">whitii</emphasis>
|
||
</taxonomicName>
|
||
- complex) that are absent in the other rock-dwelling species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1FC30E3FA67FFDAD4F826D0FDC2CB35" pageId="14" pageNumber="167" type="distribution">
|
||
<paragraph id="99596368FA67FFDAD4F826D0FDC2CB35" blockId="14.[151,1437,150,1113]" pageId="14" pageNumber="167">
|
||
<emphasis id="AB92BF7AFA67FFDAD4F826D0FE58CCC8" bold="true" box="[199,492,906,933]" pageId="14" pageNumber="167">Distribution and habitat.</emphasis>
|
||
The species’ distribution falls entirely within the Central Ranges bioregion (
|
||
<bibRefCitation id="FD771E99FA67FFDAD11726D1FED1CCA5" author="Thackway, R. & Cresswell, I. D." pageId="14" pageNumber="167" refId="ref15114" refString="Thackway, R. & Cresswell, I. D. (1995) An Interim Biogeographic Regionalisation for Australia: a framework for establishing the national system of reserves. Version 4.0. Australian Nature Conservation Agency, Canberra, 88 pp. Available from: https: // www. dcceew. gov. au / sites / default / files / documents / ibra-framework-setting-priorities-nrs-cooperative-program. pdf" type="book" year="1995">Thackway & Cresswell 1995</bibRefCitation>
|
||
) of
|
||
<collectingCountry id="E1F123F8FA67FFDAD5AC26F5FE4ECCA5" box="[403,506,943,969]" name="Australia" pageId="14" pageNumber="167">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="01DD7FEDFA67FFDAD63626F5FDD8CCA5" box="[521,620,943,969]" captionStart="FIGURE 7" captionStartId="11.[152,255,1637,1661]" captionTargetBox="[161,1429,860,1605]" captionTargetId="figure-355@11.[151,1436,846,1611]" captionTargetPageId="11" captionText="FIGURE 7. Distribution of Liopholis aputja sp. nov. (orange) and Liopholis margaretae (green). Coloured symbols are examined specimens, white dots are observations of Liopholis margaretae and hence not examined material. The landscape is shown as a digital elevation model over a hill-shading layer, with white areas corresponding to higher elevations. Note the clear topographic separation of the two species by theAmadeus lowlands: blue areas denote Lake Amadeus and associated playa lakes of the Amadeus Basin—a significant (~180 km wide) biogeographical barrier for rock-dwelling fauna. Photos: Jules Farquhar (George Gill and Musgrave Ranges); Jason Round (Amadeus lowlands)." figureDoi="http://doi.org/10.5281/zenodo.14024215" httpUri="https://zenodo.org/record/14024215/files/figure.png" pageId="14" pageNumber="167">Figure 7</figureCitation>
|
||
). It occurs in rocky hills and gorges of the Mann and Musgrave Ranges (
|
||
<figureCitation id="01DD7FEDFA67FFDAD4A02689FEB6CC81" box="[159,258,979,1005]" captionStart="FIGURE 8" captionStartId="14.[152,255,1586,1610]" captionTargetBox="[152,1436,1134,1560]" captionTargetId="figure-500@14.[152,1436,1134,1560]" captionTargetPageId="14" captionText="FIGURE 8. Habitat at the type locality of Liopholis aputja sp. nov. at Alalkanya Gorge, South Australia, 13 km north of Pukatja, eastern side of Musgrave Ranges, June 2023. (A) View of the gorge and surrounding rocky slopes and ranges, facing east. (B) Boulder pile at the base of the gorge showing typical microhabitat used by the species; an adult specimen (GenBank: PQ067254) had constructed burrows into soil beneath these rocks. Photos: Jules Farquhar." figureDoi="http://doi.org/10.5281/zenodo.14024217" httpUri="https://zenodo.org/record/14024217/files/figure.png" pageId="14" pageNumber="167">Figure 8</figureCitation>
|
||
), where it constructs burrow systems in the soil beneath boulders and into soil-filled rock crevices. It is possible that the species also occurs outside of SA, where sections of the Mann and Musgrave Ranges extend slightly into the southern NT; however, there are no records outside of SA at present. It is primarily diurnal but occasionally active at night in hot weather.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="CD9933E0FA67FFDAD4A72368FB9EC9DB" ID-DOI="http://doi.org/10.5281/zenodo.14024217" ID-Zenodo-Dep="14024217" httpUri="https://zenodo.org/record/14024217/files/figure.png" pageId="14" pageNumber="167" startId="14.[152,255,1586,1610]" targetBox="[152,1436,1134,1560]" targetPageId="14" targetType="figure">
|
||
<paragraph id="99596368FA67FFDAD4A72368FB9EC9DB" blockId="14.[151,1437,1586,1719]" pageId="14" pageNumber="167">
|
||
<emphasis id="AB92BF7AFA67FFDAD4A72368FEAFC926" bold="true" box="[152,283,1586,1610]" pageId="14" pageNumber="167">FIGURE 8.</emphasis>
|
||
Habitat at the type locality of
|
||
<taxonomicName id="5EE618EBFA67FFDAD65B2369FCBAC926" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[612,782,1587,1611]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD65B2369FCBAC926" box="[612,782,1587,1611]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD7282369FCDEC927" bold="true" box="[791,874,1587,1611]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD7282369FCDEC927" box="[791,874,1587,1611]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at Alalkanya Gorge, South Australia, 13 km north of Pukatja, eastern side of Musgrave Ranges, June 2023. (
|
||
<emphasis id="AB92BF7AFA67FFDAD6F1230CFD54C902" bold="true" box="[718,736,1622,1646]" pageId="14" pageNumber="167">A</emphasis>
|
||
) View of the gorge and surrounding rocky slopes and ranges, facing east. (
|
||
<emphasis id="AB92BF7AFA67FFDAD4EB2321FF51C9FF" bold="true" box="[212,229,1659,1683]" pageId="14" pageNumber="167">B</emphasis>
|
||
) Boulder pile at the base of the gorge showing typical microhabitat used by the species; an adult specimen (GenBank: PQ067254) had constructed burrows into soil beneath these rocks. Photos: Jules Farquhar.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D1FC30E3FA67FFDBD4F82253FB8ECE09" lastPageId="15" lastPageNumber="168" pageId="14" pageNumber="167" type="conservation">
|
||
<paragraph id="99596368FA67FFDBD4F82253FB8ECE09" blockId="14.[151,1437,1801,2043]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="168" pageId="14" pageNumber="167">
|
||
<emphasis id="AB92BF7AFA67FFDAD4F82253FE0FC84F" bold="true" box="[199,443,1801,1827]" pageId="14" pageNumber="167">Conservation status.</emphasis>
|
||
There are few recorded locations for
|
||
<taxonomicName id="5EE618EBFA67FFDAD7612253FBA3C84F" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[862,1047,1801,1827]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="14" pageNumber="167" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA67FFDAD7612253FBA3C84F" box="[862,1047,1801,1827]" italics="true" pageId="14" pageNumber="167">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA67FFDAD0222253FBC2C84F" bold="true" box="[1053,1142,1801,1827]" pageId="14" pageNumber="167">
|
||
<taxonomicNameLabel id="B0A10201FA67FFDAD0222253FBC2C84F" box="[1053,1142,1801,1827]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
given the remote region it occurs in; hence, current occurrence records may be insufficient at describing the species’ distribution. We inferred the species’ distribution based on the extent of rugged terrain in the Mann and Musgrave Ranges region (i.e. the two ranges with occurrence records for the species;
|
||
<figureCitation id="01DD7FEDFA67FFDAD6CD222FFCE1C8E3" box="[754,853,1909,1935]" captionStart="FIGURE 9" captionStartId="15.[152,255,1318,1342]" captionTargetBox="[151,1435,378,1292]" captionTargetId="figure-123@15.[151,1435,378,1292]" captionTargetPageId="15" captionText="FIGURE 9. Area of occupancy (AOO; coloured areas) and extent of occurrence (EOO; minimum convex hull around habitat) for Liopholis aputja sp. nov. (orange) and Liopholis margaretae (green)." figureDoi="http://doi.org/10.5281/zenodo.14024221" httpUri="https://zenodo.org/record/14024221/files/figure.png" pageId="14" pageNumber="167">Figure 9</figureCitation>
|
||
). Based on these calculations the AOO is
|
||
<emphasis id="AB92BF7AFA67FFDAD17F222CFAE7C8E3" box="[1344,1363,1910,1935]" italics="true" pageId="14" pageNumber="167">c.</emphasis>
|
||
<quantity id="5E1ECE8DFA67FFDAD162222FFF0FC8DF" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="167" unit="km" value="5000.0">5,000 km</quantity>
|
||
<superScript id="6E93CE20FA67FFDAD48422C3FF77C8CB" attach="left" box="[187,195,1945,1959]" fontSize="6" pageId="14" pageNumber="167">2</superScript>
|
||
(i.e. the area of rugged habitat in the region) and the EOO is
|
||
<emphasis id="AB92BF7AFA67FFDAD74A22C0FC3CC8DF" box="[885,904,1946,1971]" italics="true" pageId="14" pageNumber="167">c.</emphasis>
|
||
<quantity id="5E1ECE8DFA67FFDAD7B022C3FBB3C8DF" box="[911,1031,1945,1972]" metricMagnitude="7" metricUnit="m" metricValue="1.45" pageId="14" pageNumber="167" unit="km" value="14500.0">14,500 km</quantity>
|
||
<superScript id="6E93CE20FA67FFDAD03822C3FBBBC8CB" attach="left" box="[1031,1039,1945,1959]" fontSize="6" pageId="14" pageNumber="167">2</superScript>
|
||
(i.e. the area of a minimum convex polygon encompassing the rugged habitat used for AOO). The EOO is less than a threshold in the IUCN criteria B (subcriterion B1; <
|
||
<quantity id="5E1ECE8DFA67FFDAD54B22BBFE58C897" box="[372,492,2017,2043]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="167" unit="km" value="20000.0">20,000 km</quantity>
|
||
<superScript id="6E93CE20FA67FFDAD5D322BBFE40C883" attach="left" box="[492,500,2017,2031]" fontSize="6" pageId="14" pageNumber="167">2</superScript>
|
||
; Vulnerable), and it likely meets the condition of occurring in ≤ 10 locations, but at present it remains unclear whether it meets the other conditions relating to severe fragmentation, or decline or extreme fluctuations in population or distribution (
|
||
<bibRefCitation id="FD771E99FA66FFDBD64125E1FCBFCFB9" author="IUCN" box="[638,779,187,213]" pageId="15" pageNumber="168" refId="ref12885" refString="IUCN (2022) The Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. July 2022. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 28 April 2024)" type="journal volume" year="2022">IUCN 2022</bibRefCitation>
|
||
). The geographic distribution of this species is sparselypopulated by humans and is extremely remote, thus populations are likely to be rarely impacted by anthropogenic disturbances. The impacts of feral predators and grazers, weed encroachment (e.g. buffel grass
|
||
<taxonomicName id="5EE618EBFA66FFDBD0F52459FA38CE71" box="[1226,1420,259,285]" class="Liliopsida" family="Poaceae" genus="Cenchrus" kingdom="Plantae" order="Poales" pageId="15" pageNumber="168" phylum="Tracheophyta" rank="species" species="ciliaris">
|
||
<emphasis id="AB92BF7AFA66FFDBD0F52459FA38CE71" box="[1226,1420,259,285]" italics="true" pageId="15" pageNumber="168">Cenchrus ciliaris</emphasis>
|
||
</taxonomicName>
|
||
), fire regimes and climate change on the species should be further investigated. However, the conservation status of
|
||
<taxonomicName id="5EE618EBFA66FFDBD4A82411FEE4CE08" authorityName="Farquhar & Mulder & Russell & Haines & Rangers & Chapple" authorityYear="2024" box="[151,336,331,357]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="15" pageNumber="168" phylum="Chordata" rank="species" species="aputja" status="sp. nov.">
|
||
<emphasis id="AB92BF7AFA66FFDBD4A82411FEE4CE08" box="[151,336,331,357]" italics="true" pageId="15" pageNumber="168">Liopholis aputja</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB92BF7AFA66FFDBD5682411FE04CE09" bold="true" box="[343,432,331,357]" pageId="15" pageNumber="168">
|
||
<taxonomicNameLabel id="B0A10201FA66FFDBD5682411FE04CE09" box="[343,432,331,357]" pageId="15" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
will be examined in further detail in a forthcoming paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |