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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.80.64435" ID-GBIF-Dataset="39652af0-b0d7-4f26-8d4a-d2c765741b96" ID-PMC="PMC8121775" ID-Pensoft-Pub="1314-4049-80-19" ID-Pensoft-UUID="8D1DD25499FC58CE8320275F070DC883" ID-PubMed="34025144" ModsDocID="1314-4049-80-19" checkinTime="1620475062936" checkinUser="pensoft" docAuthor="Johnston, Peter R., Park, Duckchul, Smith, Matthew E., Mujic, Alija B. &amp; May, Tom W." docDate="2021" docId="02DE039B39AB5DCDA28B1DCA5A2C26DD" docLanguage="en" docName="MycoKeys 80: 19-43" docOrigin="MycoKeys 80" docPubDate="2021-05-07" docSource="http://dx.doi.org/10.3897/mycokeys.80.64435" docTitle="Chlorociboria metrosideri P. R. Johnst. 2021, sp. nov." docType="treatment" docVersion="4" id="8D1DD25499FC58CE8320275F070DC883" lastPageNumber="19" masterDocId="8D1DD25499FC58CE8320275F070DC883" masterDocTitle="Brahmaculus gen. nov. (Leotiomycetes, Chlorociboriaceae)" masterLastPageNumber="43" masterPageNumber="19" pageNumber="19" updateTime="1668137167928" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Brahmaculus gen. nov. (Leotiomycetes, Chlorociboriaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johnston, Peter R.</mods:namePart>
<mods:affiliation>Manaaki Whenua-Landcare Research, Private Bag 92170, Auckland 1142, (Aotearoa) New Zealand</mods:affiliation>
<mods:nameIdentifier type="email">johnstonp@landcareresearch.co.nz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Park, Duckchul</mods:namePart>
<mods:affiliation>Manaaki Whenua-Landcare Research, Private Bag 92170, Auckland 1142, (Aotearoa) New Zealand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Matthew E.</mods:namePart>
<mods:affiliation>University of Florida, Department of Plant Pathology, Gainesville FL 32611, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mujic, Alija B.</mods:namePart>
<mods:affiliation>University of Florida, Department of Plant Pathology, Gainesville FL 32611, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>May, Tom W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2214-4972</mods:nameIdentifier>
<mods:affiliation>Royal Botanic Gardens Victoria, Melbourne, Victoria 3004, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>43</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.80.64435</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.80.64435</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-80-19</mods:identifier>
<mods:identifier type="Pensoft-UUID">8D1DD25499FC58CE8320275F070DC883</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181939794" LSID="urn:lsid:plazi:treatment:02DE039B39AB5DCDA28B1DCA5A2C26DD" httpUri="http://treatment.plazi.org/id/02DE039B39AB5DCDA28B1DCA5A2C26DD" lastPageNumber="19" pageId="0" pageNumber="19">
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
<paragraph pageId="0" pageNumber="19">
<taxonomicName LSID="02DE039B-39AB-5DCD-A28B-1DCA5A2C26DD" authority="P. R. Johnst." authorityName="P. R. Johnst." authorityYear="2021" class="Leotiomycetes" family="Chlorociboriaceae" genus="Chlorociboria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Chlorociboria metrosideri" order="Helotiales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="metrosideri" status="sp. nov.">Chlorociboria metrosideri P.R.Johnst.</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Chlorociboria metrosideri (PDD 116740) A fresh apothecia B margin of receptacle in vertical section C surface of receptacle in squash mount showing rough-walled hairs D apex of asci and paraphyses E immature ascus with 8 spores, and mature asci with 4 spores. Scale bars: 0.1 mm (A); 20 µm (B, C); 10 µm (D, E)." figureDoi="10.3897/mycokeys.80.64435.figure7" httpUri="https://binary.pensoft.net/fig/540238" pageId="0" pageNumber="19">Figure 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="typification">
<paragraph pageId="0" pageNumber="19">Typification.</paragraph>
<paragraph pageId="0" pageNumber="19">
New Zealand -
<emphasis bold="true" pageId="0" pageNumber="19">Bay of Plenty</emphasis>
• vic. Rotorua, Tarawera Falls (
<geoCoordinate degrees="38.1573" direction="south" orientation="latitude" precision="5" value="-38.1573">-38.1573</geoCoordinate>
,
<geoCoordinate degrees="176.5193" direction="east" orientation="longitude" precision="5" value="176.5193">176.5193</geoCoordinate>
); on fallen leaves
<taxonomicName class="Leotiomycetes" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Metrosideros excelsa" order="Myrtales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="excelsa">
<emphasis italics="true" pageId="0" pageNumber="19">Metrosideros excelsa</emphasis>
</taxonomicName>
; P.R. Johnston (D2565) leg.; 16 May 2019; PDD 116740 -
<emphasis bold="true" italics="true" pageId="0" pageNumber="19">holotype</emphasis>
; ICMP 23410 - ex type culture.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="etymology">
<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="19">Refers to the host substrate of the known specimens.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="19">
Phylogenetically a
<taxonomicName class="Leotiomycetes" family="Chlorociboriaceae" genus="Chlorociboria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Chlorociboria" order="Helotiales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Chlorociboria</emphasis>
</taxonomicName>
, differs in developing on dead leaves rather than wood and in the asci being 4-spored when mature.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
<paragraph pageId="0" pageNumber="19">
Apothecia developing on partly decomposed fallen leaves, not associated with pigmentation of substrate. Apothecia less than 1 mm diam., sessile, with short, matted hairs around the margin, hymenium yellow. Hairs 20-45
<normalizedToken originalValue="×">x</normalizedToken>
4
<normalizedToken originalValue="µm">µm</normalizedToken>
, cylindric, walls thin, roughened. Apothecium in vertical section with ectal excipulum 30-40
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, comprising short, broad-cylindric cells 5-7.5
<normalizedToken originalValue="µm">µm</normalizedToken>
diam., with walls hyaline, slightly thickened, rows of cells arranged at a high angle to the receptacle surface near the base of the cup, more parallel to the surface near edge of cup; medullary excipulum of narrow-cylindric cells with thin walls. Paraphyses 2-3
<normalizedToken originalValue="µm">µm</normalizedToken>
diam., taper slightly and gradually to rounded apex, extending 5-10
<normalizedToken originalValue="µm">µm</normalizedToken>
beyond asci. Asci 40-55
<normalizedToken originalValue="×">x</normalizedToken>
5.5-7
<normalizedToken originalValue="µm">µm</normalizedToken>
, cylindric, tapering slightly to broad, subtruncate apex, wall thickened at apex with amyloid pore extending as two narrow, parallel bands extending through the wall, initially with 8 spores, 4 spores aborting and 4-spored at maturity, crozier present. Ascospores 7.5-9.5
<normalizedToken originalValue="×">x</normalizedToken>
2.5-3.5
<normalizedToken originalValue="µm">µm</normalizedToken>
(average 8.3
<normalizedToken originalValue="×">x</normalizedToken>
3.1
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 20), oblong-elliptic, tapering to rounded ends, one side flat in side view, widest point towards one end, 0-septate, hyaline.
</paragraph>
<caption doi="10.3897/mycokeys.80.64435.figure7" httpUri="https://binary.pensoft.net/fig/540238" pageId="0" pageNumber="19" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">Figure 7.</emphasis>
<taxonomicName class="Leotiomycetes" family="Chlorociboriaceae" genus="Chlorociboria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Chlorociboria metrosideri" order="Helotiales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="metrosideri">
<emphasis italics="true" pageId="0" pageNumber="19">Chlorociboria metrosideri</emphasis>
</taxonomicName>
(PDD 116740)
<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
fresh apothecia
<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
margin of receptacle in vertical section
<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
surface of receptacle in squash mount showing rough-walled hairs
<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
apex of asci and paraphyses
<emphasis bold="true" pageId="0" pageNumber="19">E</emphasis>
immature ascus with 8 spores, and mature asci with 4 spores. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
); 20
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="19">B, C</emphasis>
); 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="19">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="additional specimen examined">
<paragraph pageId="0" pageNumber="19">Additional specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="19">
New Zealand -
<emphasis bold="true" pageId="0" pageNumber="19">Auckland</emphasis>
• Rangitoto Island, Kidney Fern Glen;
<geoCoordinate degrees="36.805544" direction="south" orientation="latitude" precision="1" value="-36.805546">-36.805544</geoCoordinate>
,
<geoCoordinate degrees="174.860064" direction="east" orientation="longitude" precision="1" value="174.86006">174.860064</geoCoordinate>
; on fallen, partly rotten
<taxonomicName class="Leotiomycetes" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Metrosideros excelsa" order="Myrtales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="excelsa">
<emphasis italics="true" pageId="0" pageNumber="19">Metrosideros excelsa</emphasis>
</taxonomicName>
leaves; P.R. Johnston (D2329) leg.; 23 Apr 2012; PDD 102723.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="notes">
<paragraph pageId="0" pageNumber="19">Notes.</paragraph>
<paragraph pageId="0" pageNumber="19">
The substrate in both specimens was partly rotted leaves. It is possible that this fungus has a broader host range as most host-specialised, leaf-inhabiting
<taxonomicName class="Leotiomycetes" higherTaxonomySource="GBIF,CoL" kingdom="Fungi" lsidName="" pageId="0" pageNumber="19" phylum="Ascomycota" rank="class">Leotiomycetes</taxonomicName>
are found on recently fallen leaves of their preferred host. Cultures are slow growing (on PDA, 9 mm after 8 weeks) with sparse mycelium and pale brownish pigmentation, remaining sterile.
</paragraph>
</subSubSection>
</treatment>
</document>