240 lines
33 KiB
XML
240 lines
33 KiB
XML
<document id="5EBBD0EA32B50020C254948338D2E09C" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel & Polotow, Daniele" docDate="2022" docId="FB2A5333186FFFF647343033AAFAE83F" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba heliani Griswold, Ubick, Ledford & Polotow, 2022, new species" docType="treatment" docUuid="8B8279AA-15DB-4E40-AB1A-CAF914A3E658" docUuidSource="ZooBank" docVersion="2" lastPageNumber="45" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="44" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3E7FD36F95740A8BDD874B989F18C719">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:name id="E690CFBB997B8A6A7F8DF28D96BE44FC" type="personal">
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<mods:namePart id="40FCAFA91AEDC99BC05305522C4E5C9B">Griswold, Charles</mods:namePart>
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<mods:affiliation id="F24E1124C4CC05E5B39EF9DB3322D494">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="FC884562ED3B94FFB6A47B374201FB8E">Ubick, Darrell</mods:namePart>
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<mods:affiliation id="F2D40849F64784D2EA7099D64488FCE4">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="9F897110571CC8509CEEE623F856C076">Author</mods:roleTerm>
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<mods:namePart id="B259CCE9EB8751306E310B9D81E24E1B">Ledford, Joel</mods:namePart>
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<mods:affiliation id="51E601C65CB73219FC81F4FC399E7694">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="ACCBC0A7DCD6083000BA7CBB64808C3A">Polotow, Daniele</mods:namePart>
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<mods:affiliation id="AB98F7200D0C7676142132FFE207C3E3">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:title id="EC105E225C46EE2388181A0D0897F416">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="F99D3055C89D5D2F656AA17968E58B4B">
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<mods:date id="D7464EC89B03FB33FBA50A838E921D7F">2022</mods:date>
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<mods:number id="A651D3376E1715F18A112436901EE180">2022-12-15</mods:number>
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<mods:number id="6C91A5843BCC65DEC69588F0489309DA">67</mods:number>
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<mods:identifier id="D06B0C63512650620B4905282A9A5BCE" type="DOI">10.5281/zenodo.13158554</mods:identifier>
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<treatment id="FB2A5333186FFFF647343033AAFAE83F" LSID="urn:lsid:zoobank.org:act:8B8279AA-15DB-4E40-AB1A-CAF914A3E658" httpUri="http://treatment.plazi.org/id/FB2A5333186FFFF647343033AAFAE83F" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
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<subSubSection id="3B99B1AE186FFFF147343033AA9CE81F" box="[123,427,316,339]" pageId="43" pageNumber="44" type="nomenclature">
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<paragraph id="733CE225186FFFF147343033AA9CE81F" blockId="43.[123,491,316,369]" box="[123,427,316,339]" pageId="43" pageNumber="44">
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<heading id="28745549186FFFF147343033AA9CE81F" box="[123,427,316,339]" fontSize="10" level="2" pageId="43" pageNumber="44" reason="4">
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<emphasis id="41F73E37186FFFF147343033AA9CE81F" bold="true" box="[123,427,316,339]" pageId="43" pageNumber="44">
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<taxonomicName id="B48399A6186FFFF147343033AA2EE81F" ID-CoL="C8KJD" authorityName="Griswold & Ubick & Ledford & Polotow" authorityYear="2022" box="[123,281,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="heliani" status="sp. nov.">
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<emphasis id="41F73E37186FFFF147343033AA2EE81F" bold="true" box="[123,281,316,339]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="5AC4834C186FFFF146683033AA9CE81F" box="[295,427,316,339]" pageId="43" pageNumber="44" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3B99B1AE186FFFF147343053AADCE83D" box="[123,491,348,369]" pageId="43" pageNumber="44" type="description">
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<paragraph id="733CE225186FFFF147343053AADCE83D" blockId="43.[123,491,316,369]" box="[123,491,348,369]" pageId="43" pageNumber="44">
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<figureCitation id="EBB8FEA0186FFFF147343053AA3CE83D" box="[123,267,348,369]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="43" pageNumber="44">Figures 5 D, E</figureCitation>
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,
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<figureCitation id="EBB8FEA0186FFFF1465A3053AA55E83D" box="[277,354,348,369]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">72 A, B</figureCitation>
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,
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<figureCitation id="EBB8FEA0186FFFF146223053AAF5E83D" box="[365,450,348,369]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="43" pageNumber="44">Maps 11</figureCitation>
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, 16.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="3B99B1AE186FFFF147E4308EA995E895" pageId="43" pageNumber="44" type="materials_examined">
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<paragraph id="733CE225186FFFF147E4308EA995E895" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
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<emphasis id="41F73E37186FFFF147E4308EAA66E8D4" bold="true" box="[171,337,385,408]" pageId="43" pageNumber="44">Type material.</emphasis>
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<materialsCitation id="C3EBE878186FFFF14616308EA995E895" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CAS" country="Madagascar" elevation="530" location="Foret Classee Andriantantely" municipality="Helian Ratsirarson" pageId="43" pageNumber="44" specimenCode="CASENT9006106" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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<typeStatus id="AC385C87186FFFF14616308EAA8BE8D4" box="[345,444,385,408]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
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female (
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||
<specimenCode id="23254A5E186FFFF14550308EA9D0E8D4" box="[543,743,385,408]" collectionCode="CASENT" pageId="43" pageNumber="44">CASENT9006106</specimenCode>
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) from
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<quantity id="B47B4FC0186FFFF1447B308EA846E8D4" box="[820,881,385,408]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">
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<elevation id="F8AE0516186FFFF1447B308EA846E8D4" box="[820,881,385,408]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">530m</elevation>
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</quantity>
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elevation in
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<location id="765CB4FE186FFFF1434C308EAA2CE8F5" LSID="urn:lsid:plazi:treatment:FB2A5333186FFFF647343033AAFAE83F:765CB4FE186FFFF1434C308EAA2CE8F5" country="Madagascar" municipality="Helian Ratsirarson" name="Foret Classee Andriantantely" pageId="43" pageNumber="44" stateProvince="Toamasina">Forêt Classée Andriantantely</location>
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(18°41.7ʹS, 048°48.8ʹE),
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<collectingRegion id="B1472CC7186FFFF1457D30ADA839E8F5" box="[562,782,418,441]" country="Madagascar" name="Toamasina" pageId="43" pageNumber="44">Toamasina Province</collectingRegion>
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,
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<collectingCountry id="0B94A2B5186FFFF1445530ADA8AEE8F5" box="[794,921,418,441]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
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, collected
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<date id="073DC4E5186FFFF1435F30ADABEDE895" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">
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||
<collectingDate id="17793D0D186FFFF1435F30ADABEDE895" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4–7 December 1998</collectingDate>
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</date>
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by
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<collectingMunicipality id="9358785F186FFFF1464A30CDAAE7E895" box="[261,464,450,473]" pageId="43" pageNumber="44">Helian Ratsirarson</collectingMunicipality>
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, deposited in
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<collectionCode id="15927AE0186FFFF1452830CDA9A9E895" box="[615,670,450,473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="44" type="Museum">CAS</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3B99B1AE186FFFF147E430EDA9E2EB75" pageId="43" pageNumber="44" type="etymology">
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<paragraph id="733CE225186FFFF147E430EDA9E2EB75" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
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<emphasis id="41F73E37186FFFF147E430EDAA1DE8B5" bold="true" box="[171,298,482,505]" pageId="43" pageNumber="44">Etymology.</emphasis>
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The species epithet is a patronym in honor of Mr. Helian Ratsirarson, Malagasy entomologist, Lakeside International Scholar at CAS and member of the Fisher-Griswold Arthropod Team that collected widely throughout
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<collectingCountry id="0B94A2B5186FFFF1451E332DA9F8EB75" box="[593,719,546,569]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3B99B1AE186FFFF147E4334DAB88EA56" pageId="43" pageNumber="44" type="diagnosis">
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<paragraph id="733CE225186FFFF147E4334DAB88EA56" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
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||
<emphasis id="41F73E37186FFFF147E4334DAAFAEB15" bold="true" box="[171,461,578,601]" pageId="43" pageNumber="44">
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Diagnosis.
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<taxonomicName id="B48399A6186FFFF14664334DAAFAEB15" authorityName="Griswold & Ubick & Ledford & Polotow" authorityYear="2022" box="[299,461,578,601]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="heliani">
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<emphasis id="41F73E37186FFFF14664334DAAFAEB15" bold="true" box="[299,461,578,601]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
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</taxonomicName>
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</emphasis>
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belong to Group II, the Epigynum lateral projection group, or
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<taxonomicName id="B48399A6186FFFF14734336DAA76EB35" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,321,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
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group (Map 16). They have a divided cribellum. Females of
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<taxonomicName id="B48399A6186FFFF14483336DAF5DEB35" authorityName="Griswold & Ubick & Ledford & Polotow" authorityYear="2022" box="[972,1130,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="heliani">
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||
<emphasis id="41F73E37186FFFF14483336DAF5DEB35" bold="true" box="[972,1130,610,633]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
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</taxonomicName>
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||
can be distinguished from those of other
|
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<taxonomicName id="B48399A6186FFFF1455D338DA985EBD5" box="[530,690,642,665]" pageId="43" pageNumber="44">
|
||
<emphasis id="41F73E37186FFFF1455D338DA96BEBD5" bold="true" box="[530,604,642,665]" italics="true" pageId="43" pageNumber="44">Uduba</emphasis>
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||
species
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</taxonomicName>
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by the female genitalia (
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<figureCitation id="EBB8FEA0186FFFF14488338DAF68EBD5" box="[967,1119,642,665]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Figs. 72 A, B</figureCitation>
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): the epigynal plate is broadly oval (width 1.83 times length) with the copulatory openings beneath transverse grooves at mid-level of plate (
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<figureCitation id="EBB8FEA0186FFFF1457933CCA9A9EB96" box="[566,670,707,730]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 A</figureCitation>
|
||
), CO origin at 0.29 times of epigynum length; vulva with spermathecal ducts forming three longitudinal loops extending anteriad of the CO (
|
||
<figureCitation id="EBB8FEA0186FFFF1432533ECAB86EA56" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3B99B1AE186FFFF147E4322CA85CEF31" pageId="43" pageNumber="44" type="description">
|
||
<paragraph id="733CE225186FFFF147E4322CA87CEF11" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
|
||
<emphasis id="41F73E37186FFFF147E4322CAA4BEA76" bold="true" box="[171,380,803,826]" pageId="43" pageNumber="44">Description. Male</emphasis>
|
||
: Unknown.
|
||
<emphasis id="41F73E37186FFFF14541322CA956EA76" bold="true" box="[526,609,803,826]" pageId="43" pageNumber="44">Female</emphasis>
|
||
(
|
||
<typeStatus id="AC385C87186FFFF1453A322CA9EBEA76" box="[629,732,803,826]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
, CASENT9006106): Total length 6.10. Markings as in
|
||
<figureCitation id="EBB8FEA0186FFFF14667324CAA84EA16" box="[296,435,835,858]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="43" pageNumber="44">Figs. 5 D, E</figureCitation>
|
||
. Carapace 2.90 long, 2.00 wide, 1.20 high; clypeus 0.15 high. Eye diameters: AME 0.10, ALE and PME 0.12, PLE 0.16. Chelicerae 1.30 long; sternum 1.30 long, 1.10 wide; labium 0.60 long; palpal coxae 0.95 long.
|
||
<emphasis id="41F73E37186FFFF145FB328CA9CAEAD6" bold="true" box="[692,765,899,922]" pageId="43" pageNumber="44">Ratios</emphasis>
|
||
– carapace length / width = 1.40, carapace height / width = 1.40, PER / carapace width = 0.34, PER/OAL = 3.07, PER/AER = 1.34, OAL/OQL = 1.27, OQP/ OQA = 1.48, clypeus height / AME = 1.50, cheliceral length / clypeus height = 8.67, sternum length / width = 1.18, palpal coxa length / width = 2.11, femur I length / carapace width = 0.90, metatarsus I length / carapace width = 0.75, femur IV length / carapace width = 1.00, palpal tarsus length / carapace width = 0.50, palpal tibia length / palpal patella length = 2.00, palpal tarsus length / palpal tibia length = 2.00, palpal tarsus length / palpal femur length = 1.11, palpal tibia length / palpal patella length = 1.00. Cribellum divided. Spination: palpus– femur d0-1-0-2, patella p1-0, tibia p2-1, tarsus p2-1, r0-1-0; leg I–femur d1-0-0, tibia v1-1-1-2, metatarsus v2-2-2; leg II–femur d1-0-0, tibia v1-1-1-2, metatarsus p0-0-1, v2-2-2; leg III–femur d0-1-1-0, p0-1-1-0, r0-0-0-1, tibia d0-1-0, p1-0-1, v2-2-2, r0-1-1-0, metatarsus d0-1-0-0, p1-2-2, v2-2-2, r1-1-1; leg IV–femur d0-1-0, tibia v1-2-2, r0-1-1-0, metatarsus p0-0-1-2, v2-2-2, r0-1-1-1. Scopulae: weak ventral beneath tarsi I and II. Leg measurements: I: 1.80 + 0.90 + 1.50 + 1.40 + 1.10 = 6.70; II: 1.70 + 0.80 + 1.30 + 1.25 + 1.00 = 6.05; III: 1.25 + 0.70 + 0.80 + 1.10 + 0.60 = 4.45; IV: 2.00 + 0.80 + 1.50 + 1.60 + 1.00 = 6.90; palpus: 0.90 + 0.50 + 0.50 + NA + 1.00 = 2.90. Leg formula 4123.
|
||
<emphasis id="41F73E37186FFFF1461E346AA927EC30" bold="true" box="[337,528,1381,1404]" pageId="43" pageNumber="44">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="EBB8FEA0186FFFF1456D346AA989EC30" box="[546,702,1381,1404]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Figs. 72 A, B</figureCitation>
|
||
): epigynal plate without depressed atrium, posterior margin at epigastric furrow convex but concave in center, plate width 1.83 times length, ML and LL not clearly defined, copulatory openings beneath transverse grooves at mid-level of plate, distance between copulatory openings 0.295 times epigynum width, CO origin at 0.29 times of epigynum length (
|
||
<figureCitation id="EBB8FEA0186FFFF1461134EAAAFFECB0" box="[350,456,1509,1532]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 A</figureCitation>
|
||
), extent of plate laterad of copulatory openings equals 0.16 times plate width, plate with weak lateral projections, lateral margins well defined. Vulva with spermathecal ducts forming three anterior, longitudinal loops (
|
||
<figureCitation id="EBB8FEA0186FFFF1444A3729A842EF71" box="[773,885,1574,1597]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 B</figureCitation>
|
||
), vulva length 0.80 times width, vulva width 6.25 times distance between fertilization ducts.
|
||
</paragraph>
|
||
<paragraph id="733CE225186FFFF147E43769A85CEF31" blockId="43.[123,1174,385,1725]" box="[171,875,1638,1661]" pageId="43" pageNumber="44">
|
||
<emphasis id="41F73E37186FFFF147E43769AA2DEF31" bold="true" box="[171,282,1638,1661]" pageId="43" pageNumber="44">Variation.</emphasis>
|
||
Unknown: only the unique
|
||
<typeStatus id="AC385C87186FFFF145043769A99FEF31" box="[587,680,1638,1661]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
female is known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3B99B1AE186FFFF647E43789A985E87F" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="materials_examined">
|
||
<paragraph id="733CE225186FFFF147E43789AFA2EFF1" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
|
||
<emphasis id="41F73E37186FFFF147E43789AAB9EFD1" bold="true" box="[171,398,1670,1693]" pageId="43" pageNumber="44">Material examined.</emphasis>
|
||
<materialsCitation id="C3EBE878186FFFF146D63789AFA2EFF1" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectorName="H. Ratsirarson" country="Madagascar" elevation="530" location="Foret Classee Andriantantely" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
|
||
<collectingCountry id="0B94A2B5186FFFF146D63789A96CEFD1" box="[409,603,1670,1693]" name="Madagascar" pageId="43" pageNumber="44">MADAGASCAR</collectingCountry>
|
||
:
|
||
<emphasis id="41F73E37186FFFF145263789A873EFD1" box="[617,836,1670,1693]" italics="true" pageId="43" pageNumber="44">
|
||
<collectingRegion id="B1472CC7186FFFF145263789A873EFD1" box="[617,836,1670,1693]" country="Madagascar" name="Toamasina" pageId="43" pageNumber="44">Toamasina Province</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<location id="765CB4FE186FFFF1441A3789AFB9EFD1" LSID="urn:lsid:plazi:treatment:FB2A5333186FFFF647343033AAFAE83F:765CB4FE186FFFF1441A3789AFB9EFD1" box="[853,1166,1670,1693]" country="Madagascar" name="Foret Classee Andriantantely" pageId="43" pageNumber="44" stateProvince="Toamasina">Forêt Classée Andriantantely</location>
|
||
, 18°41.7ʹS, 048°48.8ʹE, elev.
|
||
<quantity id="B47B4FC0186FFFF146F237A9AACBEFF1" box="[445,508,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">
|
||
<elevation id="F8AE0516186FFFF146F237A9AACBEFF1" box="[445,508,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">530m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="073DC4E5186FFFF1454337A9A9C5EFF1" box="[524,754,1702,1725]" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">
|
||
<collectingDate id="17793D0D186FFFF1454337A9A9C5EFF1" box="[524,754,1702,1725]" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4–7 December 1998</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DE7687F3186FFFF1444C37A9A895EFF1" box="[771,930,1702,1725]" pageId="43" pageNumber="44">H. Ratsirarson</collectorName>
|
||
[#HJ-121] (
|
||
<typeStatus id="AC385C87186FFFF1437F37A9AFA5EFF1" box="[1072,1170,1702,1725]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
,
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="733CE2251868FFF6473431D3AAF5E9BF" blockId="44.[123,1174,220,371]" box="[123,450,220,243]" pageId="44" pageNumber="45">
|
||
<materialsCitation id="C3EBE8781868FFF6473431D3AA47E9BF" box="[123,368,220,243]" collectionCode="CASENT" pageId="44" pageNumber="45" specimenCode="CASENT9006106" specimenCount="1" specimenCount-female="1">
|
||
<specimenCode id="23254A5E1868FFF6473431D3AA74E9BF" box="[123,323,220,243]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9006106</specimenCode>
|
||
,
|
||
<specimenCount id="658529AC1868FFF6460131D3AA47E9BF" box="[334,368,220,243]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="15927AE01868FFF6463131D3AA81E9BF" box="[382,438,220,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="44" pageNumber="45" type="Museum">CAS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="733CE2251868FFF647E431F3A985E87F" blockId="44.[123,1174,220,371]" pageId="44" pageNumber="45">
|
||
<emphasis id="41F73E371868FFF647E431F3AA68E85F" bold="true" box="[171,351,252,275]" pageId="44" pageNumber="45">Natural history.</emphasis>
|
||
An individual of the cribellate
|
||
<taxonomicName id="B48399A61868FFF645E131F3A87AE85F" authorityName="Griswold & Ubick & Ledford & Polotow" authorityYear="2022" box="[686,845,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="heliani">
|
||
<emphasis id="41F73E371868FFF645E131F3A87AE85F" bold="true" box="[686,845,252,275]" italics="true" pageId="44" pageNumber="45">Uduba heliani</emphasis>
|
||
</taxonomicName>
|
||
has been collected in mid-elevation rainforest; no other habitat data are available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3B99B1AE1868FFF647E43033AAFAE83F" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="733CE2251868FFF647E43033AAFAE83F" blockId="44.[123,1174,220,371]" pageId="44" pageNumber="45">
|
||
<emphasis id="41F73E371868FFF647E43033AAD6E81F" bold="true" box="[171,481,316,339]" pageId="44" pageNumber="45">
|
||
Distribution.
|
||
<taxonomicName id="B48399A61868FFF6460C3033AAD6E81F" authorityName="Griswold & Ubick & Ledford & Polotow" authorityYear="2022" box="[323,481,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="heliani">
|
||
<emphasis id="41F73E371868FFF6460C3033AAD6E81F" bold="true" box="[323,481,316,339]" italics="true" pageId="44" pageNumber="45">Uduba heliani</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
are known only from the
|
||
<typeStatus id="AC385C871868FFF645B93033A813E81F" box="[758,804,316,339]" pageId="44" pageNumber="45">type</typeStatus>
|
||
locality near the east coast of central
|
||
<collectingCountry id="0B94A2B51868FFF647E93053AA10E83F" box="[166,295,348,371]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
|
||
<emphasis id="41F73E371868FFF646603053AA0FE83F" bold="true" box="[303,312,348,371]" pageId="44" pageNumber="45">(</emphasis>
|
||
<figureCitation id="EBB8FEA01868FFF646773053AAA1E83F" box="[312,406,348,371]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="44" pageNumber="45">Maps 11</figureCitation>
|
||
, 16
|
||
<emphasis id="41F73E371868FFF646F23053AAFAE83F" bold="true" box="[445,461,348,371]" pageId="44" pageNumber="45">).</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |