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<document id="F6DCEB48A486A9E63960C2EB072E5B00" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331801FF9D47343430A9ECEFF1" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba pseudoevanescens Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docVersion="2" lastPageNumber="72" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="70" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="78ED658297B7A950D5EB4D473344860E">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="3E226509C339B8917981265FFF13D6B6" type="personal">
<mods:role id="1A2E68CB18449074C06C1CA94BD639E7">
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<mods:namePart id="E61C690FE8492762F929AB85A47AA365">Griswold, Charles</mods:namePart>
<mods:affiliation id="4A889DB0277884FBD4C99DB7FD5C8E2E">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="485FBCC96BD61F89506E4B8F6B9DA94C" type="personal">
<mods:role id="1EFA0F60C96AC248F4BF3FF898B46BD9">
<mods:roleTerm id="310D394FB6E4E474CF60BB025F15B993">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="635649AB26E74B56B439EDEEFAC6F938">Ubick, Darrell</mods:namePart>
<mods:affiliation id="2809F44C4594DD4E96DE3EA5B3048B76">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="C2D828E06779494B332163B4FA3BC389" type="personal">
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<mods:namePart id="95A2E1A8DBC5A3623E3DEBBAE04BF116">Ledford, Joel</mods:namePart>
<mods:affiliation id="21692686BFB73B13811F02536B5649C5">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
</mods:name>
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<mods:role id="C023F685C495F127E8D20B7ECD22A79C">
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</mods:role>
<mods:namePart id="FC79A6625ABFB38E81429403B3B0B844">Polotow, Daniele</mods:namePart>
<mods:affiliation id="A35EF03160F1407EEC32DA15F94FA944">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
</mods:name>
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<mods:title id="D905DE61B6D58B49D46B6B02CB932D26">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="01243D0FE147BD97B7317361CFF638AA">
<mods:date id="E7F0042EE32EF2834DFE385EB43974D2">2022</mods:date>
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<mods:number id="6B1F573D8A2786BDA3823620A777B6E4">2022-12-15</mods:number>
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<mods:number id="4FAF9B38F109176E7826F49F73718C85">67</mods:number>
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<treatment id="FB2A53331801FF9D47343430A9ECEFF1" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1" httpUri="http://treatment.plazi.org/id/FB2A53331801FF9D47343430A9ECEFF1" lastPageId="71" lastPageNumber="72" pageId="69" pageNumber="70">
<subSubSection id="3B99B1AE1801FF9F47343430A911EC1A" box="[123,550,1343,1366]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="733CE2251801FF9F47343430A911EC1A" blockId="69.[123,756,1343,1396]" box="[123,550,1343,1366]" pageId="69" pageNumber="70">
<heading id="287455491801FF9F47343430A911EC1A" box="[123,550,1343,1366]" fontSize="10" level="2" pageId="69" pageNumber="70" reason="4">
<emphasis id="41F73E371801FF9F47343430A911EC1A" bold="true" box="[123,550,1343,1366]" pageId="69" pageNumber="70">
<taxonomicName id="B48399A61801FF9F47343430AAA3EC1A" ID-CoL="C8KJV" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,404,1343,1366]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudoevanescens" status="sp. nov.">
<emphasis id="41F73E371801FF9F47343430AAA3EC1A" bold="true" box="[123,404,1343,1366]" italics="true" pageId="69" pageNumber="70">Uduba pseudoevanescens</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1801FF9F46ED3430A911EC1A" box="[418,550,1343,1366]" pageId="69" pageNumber="70" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1801FF9F47343450A9C3EC38" box="[123,756,1375,1396]" pageId="69" pageNumber="70" type="description">
<paragraph id="733CE2251801FF9F47343450A9C3EC38" blockId="69.[123,756,1343,1396]" box="[123,756,1375,1396]" pageId="69" pageNumber="70">
<figureCitation id="EBB8FEA01801FF9F47343450ABD9EC38" box="[123,238,1375,1396]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="69" pageNumber="70">Figures 7 A</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F47B73450AA2AEC38" box="[248,285,1375,1396]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="69" pageNumber="70">8 B</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F46673450AA60EC38" box="[296,343,1375,1396]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="69" pageNumber="70">11 C</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F462D3450AAA4EC38" box="[354,403,1375,1396]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="69" pageNumber="70">20 C</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F46D23450AADDEC38" box="[413,490,1375,1396]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="69" pageNumber="70">54 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F46BB3450A975EC38" box="[500,578,1375,1396]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="69" pageNumber="70">71 A, B</figureCitation>
,
<figureCitation id="EBB8FEA01801FF9F45033450A941EC38" box="[588,630,1375,1396]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="69" pageNumber="70">80 J</figureCitation>
, Maps 4, 16.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1801FF9F47E4348AAF3FEF11" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="733CE2251801FF9F47E4348AAF3FEF11" blockId="69.[123,1174,1413,1725]" pageId="69" pageNumber="70">
<emphasis id="41F73E371801FF9F47E4348AAA64ECD0" bold="true" box="[171,339,1413,1436]" pageId="69" pageNumber="70">Type material.</emphasis>
<materialsCitation id="C3EBE8781801FF9F4611348AAA89EC90" collectingDate="1992-08-01" collectingDateMax="1992-08-03" collectingDateMin="1992-08-01" collectionCode="CASENT" collectorName="V. and B. Roth" country="Madagascar" location="Perinet" municipality="California Academy of Sciences" pageId="69" pageNumber="70" specimenCode="CASENT9006008" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C871801FF9F4611348AAAF6ECD0" box="[350,449,1413,1436]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E1801FF9F4568348AA9D8ECD0" box="[551,751,1413,1436]" collectionCode="CASENT" pageId="69" pageNumber="70">CASENT9006008</specimenCode>
) from
<location id="765CB4FE1801FF9F440F348AA8B9ECD0" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1:765CB4FE1801FF9F440F348AA8B9ECD0" box="[832,910,1413,1436]" country="Madagascar" municipality="California Academy of Sciences" name="Perinet" pageId="69" pageNumber="70" stateProvince="Toamasina">Périnet</location>
, 18º55ʹS; 48º25ʹE,
<collectingRegion id="B1472CC71801FF9F4328348AAA06ECF0" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B51801FF9F467134AAAA8AECF0" box="[318,445,1445,1468]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
, collected
<date id="073DC4E51801FF9F457934AAA9C5ECF0" box="[566,754,1445,1468]" pageId="69" pageNumber="70" value="1992-08-01" valueMax="1992-08-03" valueMin="1992-08-01">
<collectingDate id="17793D0D1801FF9F457934AAA9C5ECF0" box="[566,754,1445,1468]" pageId="69" pageNumber="70" value="1992-08-01" valueMax="1992-08-03" valueMin="1992-08-01">1–3 August 1992</collectingDate>
</date>
, by
<collectorName id="DE7687F31801FF9F446D34AAA8F1ECF0" box="[802,966,1445,1468]" pageId="69" pageNumber="70">V. and B. Roth</collectorName>
, deposited in
<collectingMunicipality id="9358785F1801FF9F432F34AAAA89EC90" pageId="69" pageNumber="70">California Academy of Sciences</collectingMunicipality>
</materialsCitation>
.
<materialsCitation id="C3EBE8781801FF9F468134CAAFA5EC90" box="[462,1170,1477,1500]" collectingDate="1992-08-01" collectingDateMax="1992-08-03" collectingDateMin="1992-08-01" collectionCode="CAS" collectorName="V. and B. Roth" country="Madagascar" location="Perinet" municipality="California Academy of Sciences" pageId="69" pageNumber="70" specimenCode="CASENT9006005" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="AC385C871801FF9F468134CAA91CEC90" box="[462,555,1477,1500]" pageId="69" pageNumber="70" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1801FF9F45D834CAA855EC90" box="[663,866,1477,1500]" collectionCode="CASENT" pageId="69" pageNumber="70">CASENT9006005</specimenCode>
), same data, also in
<collectionCode id="15927AE01801FF9F431434CAAFA5EC90" box="[1115,1170,1477,1500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="69" pageNumber="70" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781801FF9F473434EAAAD9EF71" collectingDate="1992-03-05" collectingDateMax="1992-07-20" collectingDateMin="1992-03-05" collectionCode="MCZ, CAS" collectorName="V. and B. Roth" country="Madagascar" location="Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT9006004, CASENT9006005" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871801FF9F473434EAABEFECB0" box="[123,216,1509,1532]" pageId="69" pageNumber="70" type="paratype">Paratype</typeStatus>
females from
<location id="765CB4FE1801FF9F463634EAA937ECB0" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1:765CB4FE1801FF9F463634EAA937ECB0" box="[377,512,1509,1532]" country="Madagascar" name="Ranomafana" pageId="69" pageNumber="70" stateProvince="Fianarantsoa">Ranomafana</location>
in
<collectingRegion id="B1472CC71801FF9F456834EAA82DECB0" box="[551,794,1509,1532]" country="Madagascar" name="Fianarantsoa" pageId="69" pageNumber="70">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51801FF9F446934EAA892ECB0" box="[806,933,1509,1532]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
, collected
<date id="073DC4E51801FF9F435134EAAB87EF50" pageId="69" pageNumber="70" value="1992-07-19" valueMax="1992-07-20" valueMin="1992-07-19">
<collectingDate id="17793D0D1801FF9F435134EAAB87EF50" pageId="69" pageNumber="70" value="1992-07-19" valueMax="1992-07-20" valueMin="1992-07-19">19–20 July 1992</collectingDate>
</date>
by
<collectorName id="DE7687F31801FF9F4790370AAAB2EF50" box="[223,389,1541,1564]" pageId="69" pageNumber="70">V. and B. Roth</collectorName>
(
<specimenCode id="23254A5E1801FF9F46D8370AA955EF50" box="[407,610,1541,1564]" collectionCode="CASENT" pageId="69" pageNumber="70">CASENT9006004</specimenCode>
), deposited in
<collectionCode id="15927AE01801FF9F4446370AA871EF50" box="[777,838,1541,1564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="69" pageNumber="70" type="Museum">MCZ</collectionCode>
, (
<specimenCode id="23254A5E1801FF9F4414370AAF13EF50" box="[859,1060,1541,1564]" collectionCode="CASENT" pageId="69" pageNumber="70">CASENT9006005</specimenCode>
) collected
<date id="073DC4E51801FF9F47343729AA2DEF71" box="[123,282,1574,1597]" pageId="69" pageNumber="70" value="1992-03-05">
<collectingDate id="17793D0D1801FF9F47343729AA2DEF71" box="[123,282,1574,1597]" pageId="69" pageNumber="70" value="1992-03-05">5 March 1992</collectingDate>
</date>
, deposited in
<collectionCode id="15927AE01801FF9F46F83729AAD9EF71" box="[439,494,1574,1597]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="69" pageNumber="70" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781801FF9F46B43729AF33EF11" collectingDate="1953-09" collectionCode="MNHN" collectorName="J. Millot &amp; Paris" country="Madagascar" location="Route d'Anosibe de Moramanga" pageId="69" pageNumber="70" specimenCode="CASENT9006006" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871801FF9F46B43729A96FEF71" box="[507,600,1574,1597]" pageId="69" pageNumber="70" type="paratype">Paratype</typeStatus>
male (
<specimenCode id="23254A5E1801FF9F45E83729A847EF71" box="[679,880,1574,1597]" collectionCode="CASENT" pageId="69" pageNumber="70">CASENT9006006</specimenCode>
) collected at
<quantity id="B47B4FC01801FF9F43473729AF7BEF71" box="[1032,1100,1574,1597]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="69" pageNumber="70" unit="km" value="57.0">57 km</quantity>
<location id="765CB4FE1801FF9F43193729AAA0EF11" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1:765CB4FE1801FF9F43193729AAA0EF11" country="Madagascar" name="Route d'Anosibe de Moramanga" pageId="69" pageNumber="70" stateProvince="Fianarantsoa">Route d’Anosibe de Moramanga</location>
,
<date id="073DC4E51801FF9F46EC3749A965EF11" box="[419,594,1606,1629]" pageId="69" pageNumber="70" value="1953-09">
<collectingDate id="17793D0D1801FF9F46EC3749A965EF11" box="[419,594,1606,1629]" pageId="69" pageNumber="70" value="1953-09">September 1953</collectingDate>
</date>
by
<collectorName id="DE7687F31801FF9F45323749A9ECEF11" box="[637,731,1606,1629]" pageId="69" pageNumber="70">J. Millot</collectorName>
, deposited in
<collectionCode id="15927AE01801FF9F443D3749A8F4EF11" box="[882,963,1606,1629]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="69" pageNumber="70">MNHN</collectionCode>
<collectorName id="DE7687F31801FF9F44843749AF33EF11" box="[971,1028,1606,1629]" pageId="69" pageNumber="70">Paris</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1801FF9F47E43769AA76EFF1" pageId="69" pageNumber="70" type="etymology">
<paragraph id="733CE2251801FF9F47E43769AA76EFF1" blockId="69.[123,1174,1413,1725]" pageId="69" pageNumber="70">
<emphasis id="41F73E371801FF9F47E43769AA1DEF31" bold="true" box="[171,298,1638,1661]" pageId="69" pageNumber="70">Etymology.</emphasis>
The species epithet refers to the similarity of this species to the ecribellate
<taxonomicName id="B48399A61801FF9F47343789A93AEFD1" authority="Dahl 1901" authorityName="Dahl" authorityYear="1901" box="[123,525,1670,1693]" class="Arachnida" family="Tengellidae" genus="Calamistrula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371801FF9F47343789AAB8EFD1" bold="true" box="[123,399,1670,1693]" italics="true" pageId="69" pageNumber="70">Calamistrula evanescens</emphasis>
Dahl 1901
</taxonomicName>
, whereas this new species
<taxonomicName id="B48399A61801FF9F44773789AF64EFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[824,1107,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371801FF9F44773789AF64EFD1" bold="true" box="[824,1107,1670,1693]" italics="true" pageId="69" pageNumber="70">Uduba pseudoevanescens</emphasis>
</taxonomicName>
has a divided cribellum.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1802FF9C47E431D3AFA1EB39" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="733CE2251802FF9C47E431D3AFA1EB39" blockId="70.[122,1174,220,1725]" pageId="70" pageNumber="71">
<emphasis id="41F73E371802FF9C47E431D3A970E9BF" bold="true" box="[171,583,220,243]" pageId="70" pageNumber="71">
Diagnosis.
<taxonomicName id="B48399A61802FF9C466531D3A970E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[298,583,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371802FF9C466531D3A970E9BF" bold="true" box="[298,583,220,243]" italics="true" pageId="70" pageNumber="71">Uduba pseudoevanescens</emphasis>
</taxonomicName>
</emphasis>
belong to Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A61802FF9C47A531F3AA84E85F" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[234,435,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). Males (
<figureCitation id="EBB8FEA01802FF9C459F31F3A852E85F" box="[720,869,252,275]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Figs. 54 A–C</figureCitation>
) can be distinguished from those of other
<taxonomicName id="B48399A61802FF9C466E3013AAF3E87F" box="[289,452,284,307]" pageId="70" pageNumber="71">
<emphasis id="41F73E371802FF9C466E3013AA5CE87F" bold="true" box="[289,363,284,307]" italics="true" pageId="70" pageNumber="71">Uduba</emphasis>
species
</taxonomicName>
except
<taxonomicName id="B48399A61802FF9C45503013A988E87F" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[543,703,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371802FF9C45503013A988E87F" bold="true" box="[543,703,284,307]" italics="true" pageId="70" pageNumber="71">U. evanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61802FF9C45B33013A8AFE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[764,920,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E371802FF9C45B33013A8AFE87F" bold="true" box="[764,920,284,307]" italics="true" pageId="70" pageNumber="71">U. rakotofrah</emphasis>
</taxonomicName>
in that the palp has a notched MA and the pointed TA2 that extends far apicad but differs from
<taxonomicName id="B48399A61802FF9C44CC3033AF2DE81F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[899,1050,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E371802FF9C44CC3033AF2DE81F" bold="true" box="[899,1050,316,339]" italics="true" pageId="70" pageNumber="71">U. rakotofrah</emphasis>
</taxonomicName>
in that TA3 and conductor extend apicad of all processes (in
<taxonomicName id="B48399A61802FF9C45C03053A81FE83F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[655,808,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E371802FF9C45C03053A81FE83F" bold="true" box="[655,808,348,371]" italics="true" pageId="70" pageNumber="71">U. rakotofrah</emphasis>
</taxonomicName>
the TA2 extends far apicad,
<figureCitation id="EBB8FEA01802FF9C432E3053ABE7E8D8" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="70" pageNumber="71">Figs. 57 A–C</figureCitation>
), TA3 is large and swollen (small and hidden in
<taxonomicName id="B48399A61802FF9C45AF3072A84EE8D8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[736,889,381,404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E371802FF9C45AF3072A84EE8D8" bold="true" box="[736,889,381,404]" italics="true" pageId="70" pageNumber="71">U. rakotofrah</emphasis>
</taxonomicName>
) and the basal lobe of the MA is entire with a concave margin (deeply forked in
<taxonomicName id="B48399A61802FF9C458C3092A86BE8F8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[707,860,413,436]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E371802FF9C458C3092A86BE8F8" bold="true" box="[707,860,413,436]" italics="true" pageId="70" pageNumber="71">U. rakotofrah</emphasis>
</taxonomicName>
). The females have the epigynum of the typical “
<taxonomicName id="B48399A61802FF9C462B30B2AAC6E898" authorityName="F. Dahl" authorityYear="1901" box="[356,497,445,468]" class="Arachnida" family="Tengellidae" genus="Calamistrula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371802FF9C462B30B2AAC6E898" box="[356,497,445,468]" italics="true" pageId="70" pageNumber="71">Calamistrula</emphasis>
</taxonomicName>
”
<typeStatus id="AC385C871802FF9C454A30B2A901E898" box="[517,566,445,468]" pageId="70" pageNumber="71">type</typeStatus>
, or
<taxonomicName id="B48399A61802FF9C452F30B2A819E898" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[608,814,445,468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371802FF9C452F30B2A819E898" bold="true" box="[608,814,445,468]" italics="true" pageId="70" pageNumber="71">Uduba evanescens</emphasis>
</taxonomicName>
group (Map 16), with large, earlike lobes extending laterally of the median sector (
<figureCitation id="EBB8FEA01802FF9C45EA30D2A82FE8B8" box="[677,792,477,500]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="70" pageNumber="71">Figs. 71 A</figureCitation>
,
<figureCitation id="EBB8FEA01802FF9C446C30D2A867E8B8" box="[803,848,477,500]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="70" pageNumber="71">80 J</figureCitation>
<emphasis id="41F73E371802FF9C441F30D2A86EE8B8" bold="true" box="[848,857,477,500]" pageId="70" pageNumber="71">)</emphasis>
. The lateral lobes are laterally rounded and larger than those on the epigyna of species
<taxonomicName id="B48399A61802FF9C445330F1A898EB59" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[796,943,510,533]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371802FF9C445330F1A898EB59" bold="true" box="[796,943,510,533]" italics="true" pageId="70" pageNumber="71">U. goodmani</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01802FF9C448E30F1AF6BEB59" box="[961,1116,510,533]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="70" pageNumber="71">Figs. 77 A, C</figureCitation>
) and
<taxonomicName id="B48399A61802FF9C47343311ABDDEB79" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,234,542,565]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="volana">
<emphasis id="41F73E371802FF9C47343311ABDDEB79" bold="true" box="[123,234,542,565]" italics="true" pageId="70" pageNumber="71">U. volana</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01802FF9C47B23311AAA9EB79" box="[253,414,542,565]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="70" pageNumber="71">Figs. 77 D, E</figureCitation>
). The genitalia of
<taxonomicName id="B48399A61802FF9C453C3311A873EB79" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[627,836,542,565]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371802FF9C453C3311A873EB79" bold="true" box="[627,836,542,565]" italics="true" pageId="70" pageNumber="71">Uduba evanescens</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01802FF9C44183311A8FFEB79" box="[855,968,542,565]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="70" pageNumber="71">Fig. 80 G</figureCitation>
) and
<taxonomicName id="B48399A61802FF9C43413311ABDDEB19" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371802FF9C43413311ABDDEB19" bold="true" italics="true" pageId="70" pageNumber="71">U. pseudoevanescens</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01802FF9C47B73331AA61EB19" box="[248,342,574,597]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="70" pageNumber="71">Fig. 80 J</figureCitation>
) are nearly identical and not diagnostic, but
<taxonomicName id="B48399A61802FF9C447D3331A8FAEB19" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[818,973,574,597]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371802FF9C447D3331A8FAEB19" bold="true" box="[818,973,574,597]" italics="true" pageId="70" pageNumber="71">U. evanescens</emphasis>
</taxonomicName>
is ecribellate (
<figureCitation id="EBB8FEA01802FF9C43253331AB87EB39" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="70" pageNumber="71">Fig. 20 D</figureCitation>
) and
<taxonomicName id="B48399A61802FF9C47A43351AAE3EB39" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[235,468,606,629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371802FF9C47A43351AAE3EB39" bold="true" box="[235,468,606,629]" italics="true" pageId="70" pageNumber="71">U. pseudoevanescens</emphasis>
</taxonomicName>
has a divided cribellum (
<figureCitation id="EBB8FEA01802FF9C45AB3351A87DEB39" box="[740,842,606,629]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="70" pageNumber="71">Fig. 20 C</figureCitation>
) or a vestige in the adult male.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1802FF9D47E43371ABF8ED15" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="description">
<paragraph id="733CE2251802FF9D47E43371AA33EA5A" blockId="70.[122,1174,220,1725]" lastBlockId="71.[122,1174,220,1725]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<emphasis id="41F73E371802FF9C47E43371AA40EBDA" bold="true" box="[171,375,638,662]" pageId="70" pageNumber="71">Description. Male</emphasis>
(
<typeStatus id="AC385C871802FF9C46C93370AADFEBDA" box="[390,488,639,662]" pageId="70" pageNumber="71" type="paratype">Paratype</typeStatus>
, CASENT9006006): Total length 13.29. Markings as in
<figureCitation id="EBB8FEA01802FF9C43043370ABB8EBFA" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="70" pageNumber="71">Figs. 8 B</figureCitation>
,
<figureCitation id="EBB8FEA01802FF9C47D43390ABF9EBFA" box="[155,206,671,694]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="70" pageNumber="71">11 C</figureCitation>
. Carapace 6.79 long, 4.93 wide, 2.00 high; clypeus 0.29 high. Eye diameters: AME 0.18, ALE and PME 0.27, PLE 0.34. Chelicerae 3.07 long; sternum 3.39 long, 2.43 wide; labium 1.43 long; palpal coxae 2.11 long.
<emphasis id="41F73E371802FF9C468C33D0A93BEBBA" bold="true" box="[451,524,735,758]" pageId="70" pageNumber="71">Ratios</emphasis>
: carapace length / width = 1.36, carapace height / width = 0.41, PER / carapace width = 0.97, PER / OAL = 2.53, PER / AER = 1.24, OAL /OQL = 1.23, OQP / OQA = 1.03, clypeus height / AME diameter width = 1.07, cheliceral length / clypeus height = 10.75, sternum length / width = 1.40, palpal coxa length / width = 2.19, femur I length / carapace width = 1.30, metatarsus I length / carapace width = 1.29, femur IV length / carapace width = 1.30, cymbium length / carapace width = 0.65, cymbium length / palpal patella length = 2.81, cymbium length / palpal tibia length = 3.00, cymbium length / palpal femur length = 1.22, palpal tibia length / palpal patella length = 0.94. Divided cribellum. Spination: palpus–femur d1-1-1, p0-0-2, patella p0-1-0, tibia p2-0-0; leg I–femur d1-1-1, p0-1-2, r0-1-0-1, tibia p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-1-1, v2-2-1, r1-0-1-0; leg II–femur d1-1-1, p1-1-2, r0-1-0-1, patella r1, tibia p0-1- 1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0-1, v2-2-2, r1-1-0; leg III–femur d1-1-0-1, p0-1-1-1, r0- 1-1-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1-2, v2-2-2, r1-2-2; leg IV–femur d1-1-0-1, p1-0-0-1, r0-0-0-1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-1, r1-2-2. Scopulae: cymbium, apicodorsal; tarsi and apical metatarsi I–II, ventral. Leg measurements: I: 6.43 + 2.57 + 6.21 + 6.36 + 3.57 = 25.14; II: 5.50 + 2.36 + 5.21 + 5.50 + 3.00 = 21.57; III: 4.64 + 1.86 + 3.36 + 4.43 + 2.36 = 16.64; IV: 6.43 + 2.29 + 5.64 + 6.57 + 2.86 = 23.79; palpus: 2.64 + 1.14 + 1.07 + NA + 3.21 = 8.07. Leg formula 1423.
<emphasis id="41F73E371802FF9C446335ECA8A8EDB6" bold="true" box="[812,927,1251,1274]" pageId="70" pageNumber="71">Male palp</emphasis>
(
<figureCitation id="EBB8FEA01802FF9C44E035ECAF75EDB6" box="[943,1090,1251,1274]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Figs. 54 A–C</figureCitation>
): palpal tibia 0.31 times cymbial length, RTA triangular (
<figureCitation id="EBB8FEA01802FF9C45CA340CA9D8EC56" box="[645,751,1283,1306]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Fig. 54 C</figureCitation>
), length 0.34 tibia length, length 1.375 times width, apex narrowed, sharply-pointed, VTA long, stout, cylindrical, upturned (
<figureCitation id="EBB8FEA01802FF9C4353342CAFBFEC76" box="[1052,1160,1315,1338]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Fig. 54 B</figureCitation>
), length 2.375 times width, length 0.459 times tibia width; tibia with slender spines but lacking stout spines; tegulum convex, with retroapical indentation, without ridge, tegulum length 1.13 times width; TA1 ends just before tegulum apex, TA2 and TA3 extend beyond TA1 and tegulum apex, TA3 a convex, blunt lobe, extending farthest apicad of all tegular processes (
<figureCitation id="EBB8FEA01802FF9C449E34ABAF0AECF7" box="[977,1085,1444,1467]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Fig. 54 A</figureCitation>
), TA2 a large, pointed, curved triangle; MA complex (
<figureCitation id="EBB8FEA01802FF9C453A34CBA83AEC97" box="[629,781,1476,1499]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Figs. 54 A, C</figureCitation>
), with proximal blunt lobe, median transverse notch, and apical curved lobe, origin at 0.17 times tegulum length, tegulum apex at 0.38 tegulum length from MA apex, length (including apical processes) 0.85 times width, MA large, length 0.415 times tegulum length, width 0.55 tegulum width (
<figureCitation id="EBB8FEA01802FF9C4465372AA8F7EF70" box="[810,960,1573,1596]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Figs. 54 A–C</figureCitation>
); tegulum with low apical mound that partially covers base of TA3 (
<figureCitation id="EBB8FEA01802FF9C45CF374AA9DEEF10" box="[640,745,1605,1628]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="70" pageNumber="71">Fig. 54 A</figureCitation>
); conductor fan convex.
<emphasis id="41F73E371802FF9C44BB374AAF70EF10" bold="true" box="[1012,1095,1605,1628]" pageId="70" pageNumber="71">Female</emphasis>
(
<typeStatus id="AC385C871802FF9C4319374AAB99EF30" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
): Total length 12.80. Markings as in
<figureCitation id="EBB8FEA01802FF9C45003769A998EF31" box="[591,687,1638,1661]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="70" pageNumber="71">Fig. 7 A</figureCitation>
. Carapace 6.20 long, 4.10 wide, 2.60 high; clypeus 0.40 high. Eye diameters: AME 0.20, ALE 0.26, PME 0.24, PLE 0.30. Chelicerae 3.00 long; sternum 2.70 long, 2.00 wide; labium 1.40 long; palpal coxae 2.00 long. Divided cribellum (
<figureCitation id="EBB8FEA01803FF9D47CC31D3ABDEE9BF" box="[131,233,220,243]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="71" pageNumber="72">Fig. 20 C</figureCitation>
). Spination: (
<typeStatus id="AC385C871803FF9D463931D3AAD7E9BF" box="[374,480,220,243]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
): palpus–femur d0-1-2, patella p0-1-0, tibia p2-0, r1-0, tarsus p2- 1; leg I–femur d1-0-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II–femur d0-1-0-0, p0-0-0-1, tibia v1-1-1-2, metatarsus v2-2-3; leg III–femur d1-0-0-1, p0-1-0-1, r0-1-1-0, tibia d0-0-1-0, p0-1- 1-0, v2-2-2, r0-1-1-0, metatarsus p0-0-2, v2-2-2, r0-0-2; leg IV–femur d0-1-1-0, tibia v1-1-2, r0- 1-1-0, metatarsus p0-0-2, v1-1-2-2, r0-0-2. Scopulae: ventral on tibia–tarsus I and II, metatarsus– tarsus III and IV and apical metatarsus III and IV. Leg measurements: I: 4.40 + 2.30 + 3.40 + 3.00 + 1.80 = 14.90; II: 3.80 + 2.00 + 3.00 + 2.80 + 1.70 = 13.30; III: 3.00 + 1.50 + 1.90 + 2.10 + 1.50 = 10.00; IV: 4.30 + 1.90 + 3.40 + 3.60 + 1.80 = 15.00; palpus: 2.10 + 1.10 + 1.10 + NA + 2.50 = 6.80. Leg formula 4123.
<emphasis id="41F73E371803FF9D46D630D2A962E8B8" bold="true" box="[409,597,477,500]" pageId="71" pageNumber="72">Female genitalia</emphasis>
(Forêt Marovato, MNHN, CASENT9006007): epigynal plate without depressed atrium, lateral lobes extending to sides forming earlike lobes (
<figureCitation id="EBB8FEA01803FF9D437130F1ABA6EB79" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="71" pageNumber="72">Figs. 71 A</figureCitation>
,
<figureCitation id="EBB8FEA01803FF9D47D23311ABFAEB79" box="[157,205,542,565]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="71" pageNumber="72">80 J</figureCitation>
), plate width 2.95 times length; width between bases of LL 0.55 times epigynum width; lateral ears wide, (LL side) width 0.19 times width epigynum, arising far from epigastric furrow, lateral ears (LL side) width 0.92 times LL side length; median lobe a flat plate with longitudinal, weakly convex median hoods, with copulatory openings beneath lateral hoods, close together, distance between copulatory openings 0.16 times epigynum width, CO origin at 0.475 of epigynum length. Vulva (
<figureCitation id="EBB8FEA01803FF9D465433B0AAA7EB9A" box="[283,400,703,726]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="71" pageNumber="72">Figs. 71 B</figureCitation>
,
<figureCitation id="EBB8FEA01803FF9D46D333B0AAFDEB9A" box="[412,458,703,726]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="71" pageNumber="72">80 J</figureCitation>
) with spermathecal ducts forming three loose, longitudinal loops, vulva length 0.54 times width, fertilization ducts well separated, vulva width 5.33 times distance between FD.
</paragraph>
<paragraph id="733CE2251803FF9D47E4322FABF8ED15" blockId="71.[122,1174,220,1725]" pageId="71" pageNumber="72">
<emphasis id="41F73E371803FF9D47E4322FAA6AEA7B" bold="true" box="[171,349,800,823]" pageId="71" pageNumber="72">Variation. Male</emphasis>
variation is unknown: only a single male is known.
<emphasis id="41F73E371803FF9D44ED322FA8C2EA7B" bold="true" box="[930,1013,800,823]" pageId="71" pageNumber="72">Female</emphasis>
(N = 5): Total length 9.30–14.50; carapace length / width = 1.46–1.58, carapace height / width = 0.54-0.66, PER / carapace width = 0.46-0.48, PER / OAL = 2.91–3.26, PER / AER = 2.91–3.26, OAL / OQL = 1.07–1.21, OQP / OQA = 1.06–1.13, clypeus height / AME = 1.00–1.92, cheliceral length / clypeus height = 6.60–10.50, sternum length / width = 1.25–1.38, palpal coxa length / width = 2.00–2.56, femur I length / carapace width = 1.00–1.14, metatarsus I length / carapace width = 0.70–0.86, femur IV length / carapace width = 0.98–1.09, palpal tarsus length / carapace width = 0.44–0.61, palpal tibia length / palpal patella length = 1.75–2.27, palpal tarsus length / palpal tibia length = 1.56–2.27, palpal tibia length / palpal patella length = 1.00-1.20. Female genitalia as in
<figureCitation id="EBB8FEA01803FF9D436D352EABB8ED15" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="71" pageNumber="72">Figs. 71 A, B</figureCitation>
,
<figureCitation id="EBB8FEA01803FF9D47D4354DABFCED15" box="[155,203,1090,1113]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="71" pageNumber="72">80 J</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1803FF9D47E4356DA921ECB0" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="733CE2251803FF9D47E4356DA873EC37" blockId="71.[122,1174,220,1725]" pageId="71" pageNumber="72">
<emphasis id="41F73E371803FF9D47E4356DAAA7ED35" bold="true" box="[171,400,1122,1145]" pageId="71" pageNumber="72">Material examined.</emphasis>
<materialsCitation id="C3EBE8781803FF9D46D2356DA8BCEDD5" collectingDate="1946-11" collectionCode="MNHN" collectorName="Madagascar E. &amp; J. Millot" country="Madagascar" location="Ankaraoka" municipality="Province" pageId="71" pageNumber="72" specimenCode="CASENT9006002" specimenCount="1" specimenCount-female="1">
<collectingCountry id="0B94A2B51803FF9D46D2356DA968ED35" box="[413,607,1122,1145]" name="Madagascar" pageId="71" pageNumber="72">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371803FF9D4524356DA851ED35" box="[619,870,1122,1145]" italics="true" pageId="71" pageNumber="72">
<collectingMunicipality id="9358785F1803FF9D4524356DA9FEED35" box="[619,713,1122,1145]" pageId="71" pageNumber="72">Province</collectingMunicipality>
not specified
</emphasis>
[“
<emphasis id="41F73E371803FF9D44C7356DAF03ED35" box="[904,1076,1122,1145]" italics="true" pageId="71" pageNumber="72">
<collectorName id="DE7687F31803FF9D44C7356DAF03ED35" box="[904,1076,1122,1145]" pageId="71" pageNumber="72">Madagascar E.</collectorName>
</emphasis>
”]: F. de
<location id="765CB4FE1803FF9D4735358DABC5EDD5" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1:765CB4FE1803FF9D4735358DABC5EDD5" box="[122,242,1154,1177]" country="Madagascar" municipality="Province" name="Ankaraoka" pageId="71" pageNumber="72">Ankaraoka</location>
,
<date id="073DC4E51803FF9D47B2358DAA9AEDD5" box="[253,429,1154,1177]" pageId="71" pageNumber="72" value="1946-11">
<collectingDate id="17793D0D1803FF9D47B2358DAA9AEDD5" box="[253,429,1154,1177]" pageId="71" pageNumber="72" value="1946-11">November 1946</collectingDate>
</date>
,
<collectorName id="DE7687F31803FF9D46F6358DA921EDD5" box="[441,534,1154,1177]" pageId="71" pageNumber="72">J. Millot</collectorName>
, (
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,
<specimenCount id="658529AC1803FF9D45B1358DA828EDD5" box="[766,799,1154,1177]" count="1" pageId="71" pageNumber="72" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01803FF9D4462358DA8B4EDD5" box="[813,899,1154,1177]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="71" pageNumber="72">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781803FF9D44D9358DA9A5EDF5" collectingDate="1946-11" collectionCode="MNHN" collectorName="F. Marovato &amp; Millot" country="Madagascar" pageId="71" pageNumber="72" specimenCode="CASENT9006007" specimenCount="1" specimenCount-female="1">
<collectorName id="DE7687F31803FF9D44D9358DAF2EEDD5" box="[918,1049,1154,1177]" pageId="71" pageNumber="72">F. Marovato</collectorName>
,
<date id="073DC4E51803FF9D436A358DAB84EDF5" pageId="71" pageNumber="72" value="1946-11">
<collectingDate id="17793D0D1803FF9D436A358DAB84EDF5" pageId="71" pageNumber="72" value="1946-11">November 1946</collectingDate>
</date>
, J.
<collectorName id="DE7687F31803FF9D479535ADAA2AEDF5" box="[218,285,1186,1209]" pageId="71" pageNumber="72">Millot</collectorName>
(
<specimenCode id="23254A5E1803FF9D466235ADAAC0EDF5" box="[301,503,1186,1209]" collectionCode="CASENT" pageId="71" pageNumber="72">CASENT9006007</specimenCode>
,
<specimenCount id="658529AC1803FF9D454B35ADA912EDF5" box="[516,549,1186,1209]" count="1" pageId="71" pageNumber="72" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01803FF9D457A35ADA9BCEDF5" box="[565,651,1186,1209]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="71" pageNumber="72">MNHN</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781803FF9D45EF35ADAAEBEDB6" collectingDate="1953-09" collectionCode="MNHN" collectorName="J. Millot &amp; Paris" country="Madagascar" location="Battage" municipality="Sandragate" pageId="71" pageNumber="72" specimenCode="CASENT9006006" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
“
<collectingMunicipality id="9358785F1803FF9D45E635ADA815EDF5" box="[681,802,1186,1209]" pageId="71" pageNumber="72">Sandragate</collectingMunicipality>
(
<location id="765CB4FE1803FF9D447C35ADA8BFEDF5" LSID="urn:lsid:plazi:treatment:FB2A53331801FF9D47343430A9ECEFF1:765CB4FE1803FF9D447C35ADA8BFEDF5" box="[819,904,1186,1209]" country="Madagascar" municipality="Sandragate" name="Battage" pageId="71" pageNumber="72">Battage</location>
) bord du torrent”
<locationDeviation id="628EFB071803FF9D431D35ADA933ED96" location="Route d'Anosibe de Moramanga" pageId="71" pageNumber="72">
<quantity id="B47B4FC01803FF9D431D35ADAFA2EDF5" box="[1106,1173,1186,1209]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="71" pageNumber="72" unit="km" value="57.0">57 km</quantity>
Route d’Anosibe de Moramanga
</locationDeviation>
,
<date id="073DC4E51803FF9D455235CCA9EBED96" box="[541,732,1219,1242]" pageId="71" pageNumber="72" value="1953-09">
<collectingDate id="17793D0D1803FF9D455235CCA9EBED96" box="[541,732,1219,1242]" pageId="71" pageNumber="72" value="1953-09">September 1953</collectingDate>
</date>
,
<collectorName id="DE7687F31803FF9D45BB35CCA86AED96" box="[756,861,1219,1242]" pageId="71" pageNumber="72">J. Millot</collectorName>
,
<collectionCode id="15927AE01803FF9D443A35CCA8FEED96" box="[885,969,1219,1242]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="71" pageNumber="72">MNHN</collectionCode>
<collectorName id="DE7687F31803FF9D449535CCAF25ED96" box="[986,1042,1219,1242]" pageId="71" pageNumber="72">Paris</collectorName>
(
<typeStatus id="AC385C871803FF9D436535CCAFA6ED96" box="[1066,1169,1219,1242]" pageId="71" pageNumber="72" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1803FF9D473435ECAA74EDB6" box="[123,323,1251,1274]" collectionCode="CASENT" pageId="71" pageNumber="72">CASENT9006006</specimenCode>
,
<specimenCount id="658529AC1803FF9D460035ECAA47EDB6" box="[335,368,1251,1274]" count="1" pageId="71" pageNumber="72" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01803FF9D463035ECAAE2EDB6" box="[383,469,1251,1274]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="71" pageNumber="72">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781803FF9D46A835ECA885EC56" collectingDate="1992-07-19" collectingDateMax="1992-07-20" collectingDateMin="1992-07-19" collectionCode="MCZ" collectorName="V. and B. Roth" country="Madagascar" location="Ranomafana" pageId="71" pageNumber="72" specimenCode="CASENT9006004" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis id="41F73E371803FF9D46A835ECA9E8EDB6" box="[487,735,1251,1274]" italics="true" pageId="71" pageNumber="72">
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:
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, ca. 21º12ʹS; 47º27ʹE,
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,
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(
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,
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,
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,
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;
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<collectorName id="DE7687F31803FF9D448C340CAFB9EC56" box="[963,1166,1283,1306]" pageId="71" pageNumber="72">Ranomafana N. P.</collectorName>
, township of
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,
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,
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(
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,
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,
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,
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</materialsCitation>
.
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<collectingRegion id="B1472CC71803FF9D4327342CAA01EC16" country="Madagascar" name="Toamasina" pageId="71" pageNumber="72">Toamasina Province</collectingRegion>
</emphasis>
:
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, 18º55ʹS; 48º25ʹE,
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,
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(
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,
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,
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(
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,
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,
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.
</paragraph>
<paragraph id="733CE2251803FF9D47E4348BA921ECB0" blockId="71.[122,1174,220,1725]" pageId="71" pageNumber="72">
<emphasis id="41F73E371803FF9D47E4348BAA52ECD7" bold="true" box="[171,357,1412,1435]" pageId="71" pageNumber="72">Natural history.</emphasis>
Natural history data are few, but most collections of
<taxonomicName id="B48399A61803FF9D4493348BABDDECF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371803FF9D4493348BABDDECF7" bold="true" italics="true" pageId="71" pageNumber="72">Uduba pseudoevanescens</emphasis>
</taxonomicName>
are from forested areas. At least
<specimenCount id="658529AC1803FF9D450534ABA9F7ECF7" box="[586,704,1444,1467]" count="1" pageId="71" pageNumber="72" type="female">one female</specimenCount>
was collected from an “… open tube, excavated in soil and rotting wood” (V. and B. Roth, label data) suggesting that these cribellate spiders dig burrows and use silk to line them.
</paragraph>
</subSubSection>
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<emphasis id="41F73E371803FF9D47E4370AAA0AEF50" bold="true" box="[171,317,1541,1564]" pageId="71" pageNumber="72">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61803FF9D4681370AA9DEEF50" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[462,745,1541,1564]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371803FF9D4681370AA9DEEF50" bold="true" box="[462,745,1541,1564]" italics="true" pageId="71" pageNumber="72">Uduba pseudoevanescens</emphasis>
</taxonomicName>
is distributed along the eastern side of the escarpment from Ranomafana in
<collectingRegion id="B1472CC71803FF9D4546372AA9C0EF70" box="[521,759,1573,1596]" country="Madagascar" name="Fianarantsoa" pageId="71" pageNumber="72">Fianarantsoa Province</collectingRegion>
in the south to Ankaraoka in the north (Maps 4, 16
<emphasis id="41F73E371803FF9D4642374AAA21EF10" bold="true" box="[269,278,1605,1628]" pageId="71" pageNumber="72">)</emphasis>
. There is one case of sympatry with the closely related ecribellate species
<taxonomicName id="B48399A61803FF9D4734376AAA2CEF30" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,283,1637,1660]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371803FF9D4734376AAA2CEF30" bold="true" box="[123,283,1637,1660]" italics="true" pageId="71" pageNumber="72">U. evanescens</emphasis>
</taxonomicName>
at Ranomafana: females of
<taxonomicName id="B48399A61803FF9D4512376AA87DEF30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[605,842,1637,1660]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371803FF9D4512376AA87DEF30" bold="true" box="[605,842,1637,1660]" italics="true" pageId="71" pageNumber="72">U. pseudoevanescens</emphasis>
</taxonomicName>
have been collected there on
<date id="073DC4E51803FF9D47343789AAF6EFD1" box="[123,449,1670,1693]" pageId="71" pageNumber="72" value="1992-03-05" valueMax="1992-07-19" valueMin="1992-03-05">5 March and 19–20 July 1992</date>
by V. and B. Roth, whereas we have collections of
<taxonomicName id="B48399A61803FF9D44B73789AFA2EFD1" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[1016,1173,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371803FF9D44B73789AFA2EFD1" bold="true" box="[1016,1173,1670,1693]" italics="true" pageId="71" pageNumber="72">U. evanescens</emphasis>
</taxonomicName>
from the Ranomafana area virtually throughout the year.
</paragraph>
</subSubSection>
</treatment>
</document>

285
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<mods:title id="9EF1B83955B4252C2E394B807FF22740">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="0137481D0DF75FA6217BA73F44B52F6E">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="2AA05B7EC9FBF18FCBC6210D41B1BF38">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<emphasis id="41F73E371804FF9A47343271AAFFEAD9" bold="true" box="[123,456,894,917]" pageId="64" pageNumber="65">
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<emphasis id="41F73E371804FF9A47343271AA02EAD9" bold="true" box="[123,309,894,917]" italics="true" pageId="64" pageNumber="65">Uduba milamina</emphasis>
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,
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<subSubSection id="3B99B1AE1804FF9A47343291AA93EAFF" box="[123,420,926,947]" pageId="64" pageNumber="65" type="description">
<paragraph id="733CE2251804FF9A47343291AA93EAFF" blockId="64.[123,456,894,947]" box="[123,420,926,947]" pageId="64" pageNumber="65">
<figureCitation id="EBB8FEA01804FF9A47343291AA11EAFF" box="[123,294,926,947]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="64" pageNumber="65">Figures 76 C, F, I</figureCitation>
, Maps 8, 15.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1804FF9A47E432CCA920ED17" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="733CE2251804FF9A47E432CCA920ED17" blockId="64.[123,1174,963,1725]" pageId="64" pageNumber="65">
<emphasis id="41F73E371804FF9A47E432CCAA62EA96" bold="true" box="[171,341,963,986]" pageId="64" pageNumber="65">Type material.</emphasis>
<materialsCitation id="C3EBE8781804FF9A462E32CCAA09ED57" collectingDate="2001-12-21" collectingDateMax="2001-12-24" collectingDateMin="2001-12-21" collectionCode="CAS" collectorName="R. Harin'Hala" country="Madagascar" location="Fianarantsoa Province" pageId="64" pageNumber="65" specimenCode="CASENT9010797" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871804FF9A462E32CCAAF3EA96" box="[353,452,963,986]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E1804FF9A456132CCA9C0EA96" box="[558,759,963,986]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT9010797</specimenCode>
) collected at the Ranomafana-Radio Tower in
<collectingRegion id="B1472CC71804FF9A47A532EBAAD7EAB7" box="[234,480,996,1019]" country="Madagascar" name="Fianarantsoa" pageId="64" pageNumber="65">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51804FF9A46BF32EBA958EAB7" box="[496,623,996,1019]" name="Madagascar" pageId="64" pageNumber="65">Madagascar</collectingCountry>
, by
<collectorName id="DE7687F31804FF9A45E732EBA872EAB7" box="[680,837,996,1019]" pageId="64" pageNumber="65">R. Harin’Hala</collectorName>
from
<date id="073DC4E51804FF9A44DE32EBAFA6EAB7" box="[913,1169,996,1019]" pageId="64" pageNumber="65" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">
<collectingDate id="17793D0D1804FF9A44DE32EBAFA6EAB7" box="[913,1169,996,1019]" pageId="64" pageNumber="65" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">21–24 December 2001</collectingDate>
</date>
, deposited in
<collectionCode id="15927AE01804FF9A4648350BAA09ED57" box="[263,318,1028,1051]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="64" pageNumber="65" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781804FF9A4605350BA924ED17" collectingDate="1993-11-19" collectingDateMax="1993-11-20" collectingDateMin="1993-11-19" collectionCode="FMNH" country="Madagascar" elevation="720" location="Andringitra Reserve" municipality="Steve Goodman" pageId="64" pageNumber="65" specimenCode="CASENT9006100" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871804FF9A4605350BAA90ED57" box="[330,423,1028,1051]" pageId="64" pageNumber="65" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1804FF9A4547350BA9E6ED57" box="[520,721,1028,1051]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT9006100</specimenCode>
) collected from soil at
<quantity id="B47B4FC01804FF9A449E350BAF39ED57" box="[977,1038,1028,1051]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="64" pageNumber="65" unit="m" value="720.0">
<elevation id="F8AE05161804FF9A449E350BAF39ED57" box="[977,1038,1028,1051]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="64" pageNumber="65" unit="m" value="720.0">720m</elevation>
</quantity>
elevation in the
<location id="765CB4FE1804FF9A47E1352BAAA5ED77" LSID="urn:lsid:plazi:treatment:FB2A53331804FF9B47343271AAF7EC1A:765CB4FE1804FF9A47E1352BAAA5ED77" box="[174,402,1060,1083]" country="Madagascar" municipality="Steve Goodman" name="Andringitra Reserve" pageId="64" pageNumber="65" stateProvince="Fianarantsoa">Andringitra Reserve</location>
,
<collectingRegion id="B1472CC71804FF9A46EB352BA9ABED77" box="[420,668,1060,1083]" country="Madagascar" name="Fianarantsoa" pageId="64" pageNumber="65">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51804FF9A45E2352BA81BED77" box="[685,812,1060,1083]" name="Madagascar" pageId="64" pageNumber="65">Madagascar</collectingCountry>
, by
<collectingMunicipality id="9358785F1804FF9A4426352BAF2BED77" box="[873,1052,1060,1083]" pageId="64" pageNumber="65">Steve Goodman</collectingMunicipality>
on
<date id="073DC4E51804FF9A431C352BAA1CED17" pageId="64" pageNumber="65" value="1993-11-19" valueMax="1993-11-20" valueMin="1993-11-19">
<collectingDate id="17793D0D1804FF9A431C352BAA1CED17" pageId="64" pageNumber="65" value="1993-11-19" valueMax="1993-11-20" valueMin="1993-11-19">19–20 November 1993</collectingDate>
</date>
, deposited in
<collectionCode id="15927AE01804FF9A468D354BA924ED17" box="[450,531,1092,1115]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="64" pageNumber="65" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1804FF9A47E4356BAA09EDD7" pageId="64" pageNumber="65" type="etymology">
<paragraph id="733CE2251804FF9A47E4356BAA09EDD7" blockId="64.[123,1174,963,1725]" pageId="64" pageNumber="65">
<emphasis id="41F73E371804FF9A47E4356BAA1DED37" bold="true" box="[171,298,1124,1147]" pageId="64" pageNumber="65">Etymology.</emphasis>
The species epithet is from a Malagasy word
<emphasis id="41F73E371804FF9A4464356BA8A7ED37" box="[811,912,1124,1147]" italics="true" pageId="64" pageNumber="65">milamina</emphasis>
meaning neat; an adjec tive in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1804FF9A47E435ABA988EC70" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="733CE2251804FF9A47E435ABA988EC70" blockId="64.[123,1174,963,1725]" pageId="64" pageNumber="65">
<emphasis id="41F73E371804FF9A47E435ABAAE9EDF7" bold="true" box="[171,478,1188,1211]" pageId="64" pageNumber="65">
Diagnosis.
<taxonomicName id="B48399A61804FF9A466A35ABAAE9EDF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,478,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="milamina">
<emphasis id="41F73E371804FF9A466A35ABAAE9EDF7" bold="true" box="[293,478,1188,1211]" italics="true" pageId="64" pageNumber="65">Uduba milamina</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.c, the
<taxonomicName id="B48399A61804FF9A45AA35ABA8B5EDF7" authorityName="Simon" authorityYear="1906" box="[741,898,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01804FF9A449C35ABAF10EDF7" box="[979,1063,1188,1211]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="64" pageNumber="65">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate, with a colulus; female with epigynal plate with median lobe surrounded by depressed atrium; epigynal plate short or nearly absent posteriad of atrium, height less than 0.12 times epigynal length, median lobe narrow, atrium width greater than three times ML width (
<figureCitation id="EBB8FEA01804FF9A463B342AA937EC70" box="[372,512,1317,1340]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="64" pageNumber="65">Figs. 76 C, I</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1804FF9B47E4344AAACDEAB9" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
<paragraph id="733CE2251804FF9A47E4344AAFA2EC10" blockId="64.[123,1174,963,1725]" box="[171,1173,1349,1372]" pageId="64" pageNumber="65">
<emphasis id="41F73E371804FF9A47E4344AAA4DEC10" bold="true" box="[171,378,1349,1372]" pageId="64" pageNumber="65">Description. Male</emphasis>
unknown.
<emphasis id="41F73E371804FF9A46B5344AA97AEC10" bold="true" box="[506,589,1349,1372]" pageId="64" pageNumber="65">Female</emphasis>
(
<typeStatus id="AC385C871804FF9A452F344AA9FDEC10" box="[608,714,1349,1372]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
): Total length 15.00. Markings typical of
</paragraph>
<paragraph id="733CE2251804FF9B4734346AAB8DEB58" blockId="64.[123,1174,963,1725]" lastBlockId="65.[122,1175,220,1366]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<taxonomicName id="B48399A61804FF9A4734346AABF2EC30" authorityName="Simon" authorityYear="1880" box="[123,197,1381,1404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371804FF9A4734346AABF2EC30" bold="true" box="[123,197,1381,1404]" italics="true" pageId="64" pageNumber="65">Uduba</emphasis>
</taxonomicName>
. Carapace 7.70 long, 4.65 wide, 3.20 high; clypeus 1.10 high. Eye diameters: AME 0.22, ALE and PME 0.26, PLE 0.34. Chelicerae 3.40 long; sternum 3.20 long, 2.30 wide; labium 1.60 long; palpal coxae 2.40 long. With a colulus. Spination: palpus–femur d0-0-1, patella p1-0, tibia p2-0, tarsus p2-1; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2-1, metatarsus v2-2-3; leg II–femur d1- 0-0, p0-0-1, tibia v1-1-1-0, metatarsus v2-2-3; leg III–femur d0-1-0-0, p0-0-1, tibia p0-1-1-0, v1- 1-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1-0-0, tibia v1-1-2, r0-1-1-0, metatarsus p0-1-2, v2-1-2-2, r0-2-2. Scopulae: strong ventral beneath tarsi I–IV and metatarsus I and II, on apices of metatarsi III and IV and on apices of tibiae I and II. Leg measurements: I: 5.60 + 2.50 + 4.80 + 4.00 + 2.30 = 19.20; II: 4.60 + 2.30 + 3.90 + 3.70 + 2.10 = 16.60; III: 3.80 + 1.80 + 2.10 + 3.00 + 1.80 = 12.50; IV: 5.10 + 2.10 + 4.30 + 5.00 + 2.30 = 18.80; palpus: 2.70 + 1.30 + 1.40 + NA + 2.50 = 7.90. Leg formula 1423.
<emphasis id="41F73E371804FF9A453B37A9A807EFF1" bold="true" box="[628,816,1702,1725]" pageId="64" pageNumber="65">Female genitalia</emphasis>
: epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01805FF9B46A731D3A963E9BF" box="[488,596,220,243]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="65" pageNumber="66">Fig. 76 C</figureCitation>
), plate width 1.21 times length; atrium sides weakly concave, atrium width 1.48 times atrium length; epigynum length 1.68 times atrium length, atrium width 0.73 times epigynum width, atrium origin at 0.12 times epigynum length from epigastric groove; median lobe narrow (
<figureCitation id="EBB8FEA01805FF9B46FB3033A92DE81F" box="[436,538,316,339]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="65" pageNumber="66">Fig. 76 C</figureCitation>
), wider towards base, atrium width at side of ML 1.27 times ML width; epigynal ML length 2.36 times width, length equal to atrium length, ML extending to apex of atrium, ML 0.62 times epigynal length; lateral lobes narrow, atrium width 9.25 times LL width; copulatory openings beneath median lateral margins of atrium. Vulva (
<figureCitation id="EBB8FEA01805FF9B44F83093AF2CE8FF" box="[951,1051,412,435]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="65" pageNumber="66">Fig. 76 F</figureCitation>
) with spermathecal ducts making broad, curved copulatory duct and single anterior spiral, vulva length 0.61 times width, fertilization ducts close together, vulva width 4.67 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251805FF9B47E43312AACDEAB9" blockId="65.[122,1175,220,1366]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E43312AA2DEB78" bold="true" box="[171,282,541,564]" pageId="65" pageNumber="66">Variation.</emphasis>
(N=2) Total length 15.00–16.00.
<emphasis id="41F73E371805FF9B45313312A9F0EB78" bold="true" box="[638,711,541,564]" pageId="65" pageNumber="66">Ratios</emphasis>
–carapace length / width = 1.54–1.66, carapace height / width = 0.57–0.69, PER / carapace width = 0.42–0.46, PER/OAL = 3.23–3.53, PER/AER = 1.39–1.41, OAL/OQL = 1.07–1.20, OQP/ OQA = 1.19–1.26, clypeus height / AME = 2.25–5.00, cheliceral length / clypeus height = 3.09–7.33, sternum length / width = 1.35–1.39, palpal coxa length / width = 2.09–2.18, metatarsus I length / carapace width = 0.83–0.86, femur IV length / carapace width = 1.10–1.11, palpal tarsus length / carapace width = 0.48–0.54, palpal tibia length / palpal patella length = 1.69–1.92, palpal tarsus length / palpal tibia length = 1.57–1.79, palpal tarsus length / palpal femur length = 0.85–0.93, palpal tibia length / palpal patella length = 1.08. Spines beneath the leg tarsi may be absent (
<typeStatus id="AC385C871805FF9B45E73211A83DEA79" box="[680,778,798,821]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
, Ranomafana) or present (
<typeStatus id="AC385C871805FF9B437E3211AFA6EA79" box="[1073,1169,798,821]" pageId="65" pageNumber="66" type="paratype">paratype</typeStatus>
, Andringitra). Epigynal plate width 1.16–1.21 times length (
<figureCitation id="EBB8FEA01805FF9B444F3231A8BCEA19" box="[768,907,830,853]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="65" pageNumber="66">Figs. 76 C, I</figureCitation>
) epigynum length 1.68– 1.94 times atrium length, atrium origin at 0.10–0.12 times epigynum length from epigastric groove; epigynal median lobe length 2.36–4.75 times width, ML length 1.00–1.19 times atrium length; broad, curved copulatory duct making one or one and a half anterior spirals, vulva length 0.61– 1.20 times width, fertilization ducts close together, vulva width 4.67–5.00 times distance between fertilization ducts (
<figureCitation id="EBB8FEA01805FF9B460832D1AADDEAB9" box="[327,490,990,1013]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="65" pageNumber="66">Figs. 76 C, F, I</figureCitation>
<emphasis id="41F73E371805FF9B46A532D1AAC4EAB9" bold="true" box="[490,499,990,1013]" pageId="65" pageNumber="66">)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9B47E432F1A8D6EC5A" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="733CE2251805FF9B47E432F1A978EDFA" blockId="65.[122,1175,220,1366]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E432F1AAB9ED59" bold="true" box="[171,398,1022,1045]" pageId="65" pageNumber="66">Material examined.</emphasis>
<materialsCitation id="C3EBE8781805FF9B46D732F1A826ED1A" collectingDate="2001-12-21" collectingDateMax="2001-12-24" collectingDateMin="2001-12-21" collectionCode="CAS" collectorName="R. Harin'Hala" country="Madagascar" location="Ranomafana Radio Tower" pageId="65" pageNumber="66" specimenCode="CASENT9010797" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectingCountry id="0B94A2B51805FF9B46D732F1A96DED59" box="[408,602,1022,1045]" name="Madagascar" pageId="65" pageNumber="66">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371805FF9B452932F1A857ED59" box="[614,864,1022,1045]" italics="true" pageId="65" pageNumber="66">
<collectingRegion id="B1472CC71805FF9B452932F1A857ED59" box="[614,864,1022,1045]" country="Madagascar" name="Fianarantsoa" pageId="65" pageNumber="66">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1805FF9B443C32F1AFB8ED59" LSID="urn:lsid:plazi:treatment:FB2A53331804FF9B47343271AAF7EC1A:765CB4FE1805FF9B443C32F1AFB8ED59" box="[883,1167,1022,1045]" country="Madagascar" name="Ranomafana Radio Tower" pageId="65" pageNumber="66" stateProvince="Fianarantsoa">Ranomafana Radio Tower</location>
, 21°15.05ʹS, 47°24.43ʹE,
<collectingMethod id="AAC29A321805FF9B46C93511A96AED79" box="[390,605,1054,1077]" pageId="65" pageNumber="66">malaise canopy trap</collectingMethod>
at an open area at the forest edge,
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<collectingDate id="17793D0D1805FF9B44923511AB84ED1A" pageId="65" pageNumber="66" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">21–24 December 2001</collectingDate>
</date>
,
<collectorName id="DE7687F31805FF9B47F23530AA62ED1A" box="[189,341,1087,1110]" pageId="65" pageNumber="66">R. Harin’Hala</collectorName>
(
<typeStatus id="AC385C871805FF9B462C3530AAF2ED1A" box="[355,453,1087,1110]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E1805FF9B46803530A9AFED1A" box="[463,664,1087,1110]" collectionCode="CASENT" pageId="65" pageNumber="66">CASENT9010797</specimenCode>
,
<specimenCount id="658529AC1805FF9B45ED3530A9F4ED1A" box="[674,707,1087,1110]" count="1" pageId="65" pageNumber="66" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01805FF9B459E3530A83EED1A" box="[721,777,1087,1110]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781805FF9B44533530A97CEDFA" collectingDate="1993-11-19" collectingDateMax="1993-11-20" collectingDateMin="1993-11-19" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" county="Andringitra Reserve" elevation="720" latitude="-22.222223" location="Ambalamanenjana-Ambatomboay" longLatPrecision="21" longitude="47.02222" municipality="Iantara River" pageId="65" pageNumber="66" specimenCode="CASENT9006100" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa Province" typeStatus="paratype">
edge of
<collectingCounty id="9A5D9AA91805FF9B44203530AF7CED1A" box="[879,1099,1087,1110]" pageId="65" pageNumber="66">Andringitra Reserve</collectingCounty>
, ca.
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<quantity id="B47B4FC01805FF9B43343530ABAAED3A" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="65" pageNumber="66" unit="km" value="45.0">45 km</quantity>
S Ambalavao
</locationDeviation>
, on east bank of
<collectingMunicipality id="9358785F1805FF9B45493550A9A3ED3A" box="[518,660,1119,1142]" pageId="65" pageNumber="66">Iantara River</collectingMunicipality>
, along
<location id="765CB4FE1805FF9B45A63550AF6FED3A" LSID="urn:lsid:plazi:treatment:FB2A53331804FF9B47343271AAF7EC1A:765CB4FE1805FF9B45A63550AF6FED3A" box="[745,1112,1119,1142]" country="Madagascar" county="Andringitra Reserve" latitude="-22.222223" longLatPrecision="21" longitude="47.02222" municipality="Iantara River" name="Ambalamanenjana-Ambatomboay" pageId="65" pageNumber="66" stateProvince="Fianarantsoa Province">Ambalamanenjana-Ambatomboay</location>
trail, camp #1,
<geoCoordinate id="16B784E21805FF9B47A43570AA52EDDA" box="[235,357,1151,1174]" degrees="22" direction="south" minutes="13" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="20" value="-22.222223">22°13ʹ20ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21805FF9B463A3570AAC5EDDA" box="[373,498,1151,1174]" degrees="47" direction="east" minutes="01" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="20" value="47.02222">47°01ʹ20ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01805FF9B45753570A94DEDDA" box="[570,634,1151,1174]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="65" pageNumber="66" unit="m" value="720.0">
<elevation id="F8AE05161805FF9B45753570A94DEDDA" box="[570,634,1151,1174]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="65" pageNumber="66" unit="m" value="720.0">720m</elevation>
</quantity>
, from soil,
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<collectingDate id="17793D0D1805FF9B45B13570A8C9EDDA" box="[766,1022,1151,1174]" pageId="65" pageNumber="66" value="1993-11-19" valueMax="1993-11-20" valueMin="1993-11-19">19–20 November 1993</collectingDate>
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,
<collectorName id="DE7687F31805FF9B43433570AFA2EDDA" box="[1036,1173,1151,1174]" pageId="65" pageNumber="66">S. Goodman</collectorName>
(
<typeStatus id="AC385C871805FF9B47CD3590ABD5EDFA" box="[130,226,1183,1206]" pageId="65" pageNumber="66" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1805FF9B47A13590AA81EDFA" box="[238,438,1183,1206]" collectionCode="CASENT" pageId="65" pageNumber="66">CASENT9006100</specimenCode>
,
<specimenCount id="658529AC1805FF9B468D3590AAD4EDFA" box="[450,483,1183,1206]" count="1" pageId="65" pageNumber="66" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01805FF9B46BD3590A973EDFA" box="[498,580,1183,1206]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="65" pageNumber="66" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251805FF9B47E435B0A8D6EC5A" blockId="65.[122,1175,220,1366]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E435B0AA69ED9A" bold="true" box="[171,350,1215,1238]" pageId="65" pageNumber="66">Natural history.</emphasis>
The
<typeStatus id="AC385C871805FF9B46DD35B0AAD9ED9A" box="[402,494,1215,1238]" pageId="65" pageNumber="66" type="paratype">paratype</typeStatus>
female of
<taxonomicName id="B48399A61805FF9B451135B0A820ED9A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[606,791,1215,1238]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="milamina">
<emphasis id="41F73E371805FF9B451135B0A820ED9A" bold="true" box="[606,791,1215,1238]" italics="true" pageId="65" pageNumber="66">Uduba milamina</emphasis>
</taxonomicName>
was collected in soil and the female
<typeStatus id="AC385C871805FF9B473435D0ABEFEDBA" box="[123,216,1247,1270]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
of this ecribellate spider was collected in a malaise canopy trap at an open area at the forest edge, suggesting that this species is terrestrial but may wander in vegetation.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9B47E43410AAF7EC1A" pageId="65" pageNumber="66" type="distribution">
<paragraph id="733CE2251805FF9B47E43410AAF7EC1A" blockId="65.[122,1175,220,1366]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E43410AACAEC7A" bold="true" box="[171,509,1311,1334]" pageId="65" pageNumber="66">
Distribution.
<taxonomicName id="B48399A61805FF9B460C3410AACAEC7A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[323,509,1311,1334]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="milamina">
<emphasis id="41F73E371805FF9B460C3410AACAEC7A" bold="true" box="[323,509,1311,1334]" italics="true" pageId="65" pageNumber="66">Uduba milamina</emphasis>
</taxonomicName>
</emphasis>
are known from upland eastern forest from central and southern
<collectingCountry id="0B94A2B51805FF9B47EA3430AA11EC1A" box="[165,294,1343,1366]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
(Maps 8, 15).
</paragraph>
</subSubSection>
</treatment>
</document>

249
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<mods:title id="2623F707F8C7A2B193B786D1FB0E6BA6">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<taxonomicName id="B48399A61805FF9B4734348FAA3AECDB" ID-CoL="C8KJS" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,269,1408,1431]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orona" status="sp. nov.">
<emphasis id="41F73E371805FF9B4734348FAA3AECDB" bold="true" box="[123,269,1408,1431]" italics="true" pageId="65" pageNumber="66">Uduba orona</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1805FF9B4654348FAA97ECDB" box="[283,416,1408,1431]" pageId="65" pageNumber="66" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9B473434AFAAA3ECF9" box="[123,404,1440,1461]" pageId="65" pageNumber="66" type="description">
<paragraph id="733CE2251805FF9B473434AFAAA3ECF9" blockId="65.[123,416,1408,1461]" box="[123,404,1440,1461]" pageId="65" pageNumber="66">
<figureCitation id="EBB8FEA01805FF9B473434AFAA20ECF9" box="[123,279,1440,1461]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="65" pageNumber="66">Figures 75 D, E</figureCitation>
, Maps 4, 15.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9B47E434CAAF79EF50" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="733CE2251805FF9B47E434CAAF79EF50" blockId="65.[123,1174,1477,1725]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E434CAAA79EC90" bold="true" box="[171,334,1477,1500]" pageId="65" pageNumber="66">Type material.</emphasis>
<materialsCitation id="C3EBE8781805FF9B461A34CAAF79EF50" collectingDate="1993-12-12" collectingDateMax="1993-12-13" collectingDateMin="1993-12-12" collectionCode="FMNH" collectorName="Steve Goodman" country="Madagascar" elevation="1625" latitude="-22.194166" location="Andringitra Reserve" longLatPrecision="21" longitude="46.97111" pageId="65" pageNumber="66" specimenCode="CASENT9006101" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871805FF9B461A34CAAA8FEC90" box="[341,440,1477,1500]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E1805FF9B455A34CAA9E9EC90" box="[533,734,1477,1500]" collectionCode="CASENT" pageId="65" pageNumber="66">CASENT9006101</specimenCode>
) from soil at
<quantity id="B47B4FC01805FF9B442234CAA880EC90" box="[877,951,1477,1500]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="65" pageNumber="66" unit="m" value="1625.0">
<elevation id="F8AE05161805FF9B442234CAA880EC90" box="[877,951,1477,1500]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="65" pageNumber="66" unit="m" value="1625.0">1625m</elevation>
</quantity>
elevation on ridge
<locationDeviation id="628EFB071805FF9B43CA34CAAA45ECB0" location="Volotsangana River" pageId="65" pageNumber="66">E of Volotsangana River</locationDeviation>
,
<geoCoordinate id="16B784E21805FF9B46C734EAA937ECB0" box="[392,512,1509,1532]" degrees="22" direction="south" minutes="11" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="39" value="-22.194166">22°11ʹ39ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21805FF9B455A34EAA9A5ECB0" box="[533,658,1509,1532]" degrees="46" direction="east" minutes="58" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="16" value="46.97111">46°58ʹ16ʺE</geoCoordinate>
,
<location id="765CB4FE1805FF9B45ED34EAA8B1ECB0" LSID="urn:lsid:plazi:treatment:FB2A53331805FF984734348FA82EECFF:765CB4FE1805FF9B45ED34EAA8B1ECB0" box="[674,902,1509,1532]" country="Madagascar" latitude="-22.194166" longLatPrecision="21" longitude="46.97111" name="Andringitra Reserve" pageId="65" pageNumber="66" stateProvince="Fianarantsoa">Andringitra Reserve</location>
,
<collectingRegion id="B1472CC71805FF9B44D634EAAFA5ECB0" box="[921,1170,1509,1532]" country="Madagascar" name="Fianarantsoa" pageId="65" pageNumber="66">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51805FF9B4734370AABCDEF50" box="[123,250,1541,1564]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
, collected by
<collectorName id="DE7687F31805FF9B46DC370AA908EF50" box="[403,575,1541,1564]" pageId="65" pageNumber="66">Steve Goodman</collectorName>
on
<date id="073DC4E51805FF9B4525370AA855EF50" box="[618,866,1541,1564]" pageId="65" pageNumber="66" value="1993-12-12" valueMax="1993-12-13" valueMin="1993-12-12">
<collectingDate id="17793D0D1805FF9B4525370AA855EF50" box="[618,866,1541,1564]" pageId="65" pageNumber="66" value="1993-12-12" valueMax="1993-12-13" valueMin="1993-12-12">12–13 December 1993</collectingDate>
</date>
, deposited in
<collectionCode id="15927AE01805FF9B44B6370AAF7DEF50" box="[1017,1098,1541,1564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="65" pageNumber="66" type="Museum">FMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9B47E43729A9F7EF11" pageId="65" pageNumber="66" type="etymology">
<paragraph id="733CE2251805FF9B47E43729A9F7EF11" blockId="65.[123,1174,1477,1725]" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E43729AA1DEF71" bold="true" box="[171,298,1574,1597]" pageId="65" pageNumber="66">Etymology.</emphasis>
The species epithet is from a Malagasy word for nose,
<emphasis id="41F73E371805FF9B44EF3729A8E9EF71" box="[928,990,1574,1597]" italics="true" pageId="65" pageNumber="66">orona</emphasis>
, referring to the shape of the median lobe (ML); a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1805FF9847E43769AAACE85F" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="733CE2251805FF9847E43769AAACE85F" blockId="65.[123,1174,1477,1725]" lastBlockId="66.[123,1175,220,1459]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="41F73E371805FF9B47E43769AA8FEF31" bold="true" box="[171,440,1638,1661]" pageId="65" pageNumber="66">
Diagnosis.
<taxonomicName id="B48399A61805FF9B46693769AA8FEF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,440,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orona">
<emphasis id="41F73E371805FF9B46693769AA8FEF31" bold="true" box="[294,440,1638,1661]" italics="true" pageId="65" pageNumber="66">Uduba orona</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.c, the
<taxonomicName id="B48399A61805FF9B45883769A852EF31" authorityName="Simon" authorityYear="1906" box="[711,869,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01805FF9B44F63769AF39EF31" box="[953,1038,1638,1661]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="65" pageNumber="66">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate with a colulus. The female has the epigynal plate with median lobe surrounded by depressed atrium; the median lobe is broad and extends to or bulges across epigastric furrow, ML length greater than 1.75 times width (
<figureCitation id="EBB8FEA01806FF9844DA31D3AF37E9BF" box="[917,1024,220,243]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="66" pageNumber="67">Fig. 75 D</figureCitation>
). The male of
<taxonomicName id="B48399A61806FF98473431F3AA39E85F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,270,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="orona">
<emphasis id="41F73E371806FF98473431F3AA39E85F" bold="true" box="[123,270,252,275]" italics="true" pageId="66" pageNumber="67">Uduba orona</emphasis>
</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1806FF9847E43013AA4AEC3F" pageId="66" pageNumber="67" type="description">
<paragraph id="733CE2251806FF9847E43013AFA2E87F" blockId="66.[123,1175,220,1459]" box="[171,1173,284,307]" pageId="66" pageNumber="67">
<emphasis id="41F73E371806FF9847E43013AA4CE87F" bold="true" box="[171,379,284,307]" pageId="66" pageNumber="67">Description. Male</emphasis>
unknown.
<emphasis id="41F73E371806FF9846B33013A978E87F" bold="true" box="[508,591,284,307]" pageId="66" pageNumber="67">Female</emphasis>
(
<typeStatus id="AC385C871806FF98452D3013A9F1E87F" box="[610,710,284,307]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
): Total length 14.50. Markings typical of
</paragraph>
<paragraph id="733CE2251806FF9847343033AA3CEDDF" blockId="66.[123,1175,220,1459]" pageId="66" pageNumber="67">
<taxonomicName id="B48399A61806FF9847343033ABF2E81F" authorityName="Simon" authorityYear="1880" box="[123,197,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371806FF9847343033ABF2E81F" bold="true" box="[123,197,316,339]" italics="true" pageId="66" pageNumber="67">Uduba</emphasis>
</taxonomicName>
. Carapace 7.60 long, 5.20 wide, 3.00 high; clypeus 0.55 high. Eye diameters: AME, ALE and PLE 0.26, PME 0.22. Chelicerae 3.50 long; sternum 3.40 long, 2.60 wide; labium 1.70 long; palpal coxae 2.55 long.
<emphasis id="41F73E371806FF9846393073AAFBE8DF" bold="true" box="[374,460,380,403]" pageId="66" pageNumber="67">Ratios–</emphasis>
carapace length / width = 1.46, carapace height / width = 0.58, PER / carapace width = 0.41, PER/OAL = 3.24, PER/AER = 1.35, OAL/OQL = 1.22, OQP/ OQA = 1.09, clypeus height / AME = 2.12, cheliceral length / clypeus height = 6.36, sternum length / width = 1.31, palpal coxa length / width = 2.32, femur I length / carapace width =1.13, metatarsus I length / carapace width = 0.79, femur IV length / carapace width = 1.10, palpal tarsus length / carapace width = 0.48, palpal tibia length / palpal patella length = 1.92, palpal tarsus length / palpal tibia length = 1.67, palpal tarsus length / palpal femur length = 0.86, palpal tibia length / palpal patella length = 1.15. With a colulus. Spination: palpus–femur d0-0-1, patella p1-0, tibia d0-1-0, p2-0, tarsus p2-1; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2-0, metatarsus v2-2-3; leg II–femur d1-0-0-1, p0- 1-1-0, tibia p0-0-1-0, v1-1-1-0, metatarsus v2-2-3; leg III–femur p1-1-1-1, r0-1-0-0, tibia p0-1-1- 0, v1-2-1, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1-0-0, tibia v2-2-2, metatarsus p0-1-2, v1-2-2-2, r1-1-2. Scopulae: strong ventral beneath tarsi I–IV and metatarsus I and II and apices of metatarsi III and IV. Leg measurements: I: 5.90 + 2.70 + 5.10 + 4.10 + 2.30 = 20.10; II: 5.10 + 2.50 + 4.10 + 3.60 + 2.10 = 17.40; III: 4.00 + 2.10 + 2.50 + 3.40 + 1.90 = 13.90; IV: 5.70 + 2.30 + 4.80 + 5.50 + 2.40 = 20.70; palpus: 2.90 + 1.30 + 1.50 + NA + 2.50 = 8.20. Leg formula 4123.
<emphasis id="41F73E371806FF9847973253AAA3EA3F" bold="true" box="[216,404,860,883]" pageId="66" pageNumber="67">Female genitalia</emphasis>
: epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01806FF9843253253AB85EADF" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="66" pageNumber="67">Fig. 75 D</figureCitation>
), plate width 1.28 times length; atrium sides concave, atrium width 1.35 times atrium length; epigynum length 1.35 times atrium length, atrium width 0.78 times epigynum width; atrium width at side of ML equals ML width; atrium and ML extend as lobe posteriad of epigastric groove (
<figureCitation id="EBB8FEA01806FF98432532B3AB81EABF" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="66" pageNumber="67">Fig. 75 D</figureCitation>
); median lobe triangular, wider at base, median lobe length 1.85 times width, ML length equals atrium length, 0.69 times epigynal length; lateral lobes narrow, atrium width 8.75 times LL width; copulatory openings beneath anterolateral margins of atrium. Vulva (
<figureCitation id="EBB8FEA01806FF9844FE3513AF2FED7F" box="[945,1048,1052,1075]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="66" pageNumber="67">Fig. 75 E</figureCitation>
) with spermathecal ducts making a large lateral copulatory duct, anteriorly two narrow spirals, and a posteriorly a lobate base, vulva length 0.62 times width, vulva width 5.25 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251806FF9847E43593A8E3EDFF" blockId="66.[123,1175,220,1459]" box="[171,980,1180,1203]" pageId="66" pageNumber="67">
<emphasis id="41F73E371806FF9847E43593AA2DEDFF" bold="true" box="[171,282,1180,1203]" pageId="66" pageNumber="67">Variation.</emphasis>
Unknown: only the
<typeStatus id="AC385C871806FF9846B43593A96FEDFF" box="[507,600,1180,1203]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="B48399A61806FF9845803593A855EDFF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[719,866,1180,1203]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="orona">
<emphasis id="41F73E371806FF9845803593A855EDFF" bold="true" box="[719,866,1180,1203]" italics="true" pageId="66" pageNumber="67">Uduba orona</emphasis>
</taxonomicName>
is known.
</paragraph>
<paragraph id="733CE2251806FF9847E435B3ABEFEC7F" blockId="66.[123,1175,220,1459]" pageId="66" pageNumber="67">
<materialsCitation id="C3EBE8781806FF9847E435B3ABE3EC7F" collectingDate="1993-12-12" collectingDateMax="1993-12-13" collectingDateMin="1993-12-12" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1625" latitude="-22.194166" location="Andringitra Reserve" longLatPrecision="21" longitude="46.97111" pageId="66" pageNumber="67" specimenCode="CASENT9006101" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="holotype">
M
<emphasis id="41F73E371806FF98478D35B3AAB9ED9F" bold="true" box="[194,398,1212,1235]" pageId="66" pageNumber="67">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51806FF9846D435B3A96AED9F" box="[411,605,1212,1235]" name="Madagascar" pageId="66" pageNumber="67">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371806FF98452235B3A85EED9F" box="[621,873,1212,1235]" italics="true" pageId="66" pageNumber="67">
<collectingRegion id="B1472CC71806FF98452235B3A85EED9F" box="[621,873,1212,1235]" country="Madagascar" name="Fianarantsoa" pageId="66" pageNumber="67">Fianarantsoa Province</collectingRegion>
</emphasis>
: ca.
<locationDeviation id="628EFB071806FF9844E535B3AFA5ED9F" box="[938,1170,1212,1235]" location="Ambalavao" pageId="66" pageNumber="67">
<quantity id="B47B4FC01806FF9844E535B3A8DAED9F" box="[938,1005,1212,1235]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="66" pageNumber="67" unit="km" value="38.0">38 km</quantity>
S Ambalavao
</locationDeviation>
,
<location id="765CB4FE1806FF98473435D3AA6DEDBF" LSID="urn:lsid:plazi:treatment:FB2A53331805FF984734348FA82EECFF:765CB4FE1806FF98473435D3AA6DEDBF" box="[123,346,1244,1267]" country="Madagascar" latitude="-22.194166" longLatPrecision="21" longitude="46.97111" name="Andringitra Reserve" pageId="66" pageNumber="67" stateProvince="Fianarantsoa">Andringitra Reserve</location>
, on ridge
<locationDeviation id="628EFB071806FF98468335D3A9E3EDBF" box="[460,724,1244,1267]" location="Volotsangana River" pageId="66" pageNumber="67">E of Volotsangana River</locationDeviation>
, camp #4,
<geoCoordinate id="16B784E21806FF98441C35D3A8FCEDBF" box="[851,971,1244,1267]" degrees="22" direction="south" minutes="11" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="39" value="-22.194166">22°11ʹ39ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21806FF98449435D3AF6FEDBF" box="[987,1112,1244,1267]" degrees="46" direction="east" minutes="58" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="16" value="46.97111">46°58ʹ16ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01806FF98473435F3ABFFEC5F" box="[123,200,1276,1299]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="66" pageNumber="67" unit="m" value="1625.0">
<elevation id="F8AE05161806FF98473435F3ABFFEC5F" box="[123,200,1276,1299]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="66" pageNumber="67" unit="m" value="1625.0">1625m</elevation>
</quantity>
, ex soil sample,
<date id="073DC4E51806FF9846CD35F3A94AEC5F" box="[386,637,1276,1299]" pageId="66" pageNumber="67" value="1993-12-12" valueMax="1993-12-13" valueMin="1993-12-12">
<collectingDate id="17793D0D1806FF9846CD35F3A94AEC5F" box="[386,637,1276,1299]" pageId="66" pageNumber="67" value="1993-12-12" valueMax="1993-12-13" valueMin="1993-12-12">12–13 December 1993</collectingDate>
</date>
,
<collectorName id="DE7687F31806FF9845C535F3A822EC5F" box="[650,789,1276,1299]" pageId="66" pageNumber="67">S. Goodman</collectorName>
, (
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,
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,
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,
<collectionCode id="15927AE01806FF9847343413ABFAEC7F" box="[123,205,1308,1331]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="66" pageNumber="67" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251806FF9847E43433AA4AEC3F" blockId="66.[123,1175,220,1459]" pageId="66" pageNumber="67">
<emphasis id="41F73E371806FF9847E43433AA68EC1F" bold="true" box="[171,351,1340,1363]" pageId="66" pageNumber="67">Natural history.</emphasis>
The
<typeStatus id="AC385C871806FF9846DA3433AAC5EC1F" box="[405,498,1340,1363]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
female of the ecribellate
<taxonomicName id="B48399A61806FF98444C3433A8A2EC1F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[771,917,1340,1363]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="orona">
<emphasis id="41F73E371806FF98444C3433A8A2EC1F" bold="true" box="[771,917,1340,1363]" italics="true" pageId="66" pageNumber="67">Uduba orona</emphasis>
</taxonomicName>
was collected from soil in a montane rainforest.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1806FF9847E43473A82EECFF" pageId="66" pageNumber="67" type="distribution">
<paragraph id="733CE2251806FF9847E43473A82EECFF" blockId="66.[123,1175,220,1459]" pageId="66" pageNumber="67">
<emphasis id="41F73E371806FF9847E43473AAE3ECDF" bold="true" box="[171,468,1404,1427]" pageId="66" pageNumber="67">
Distribution.
<taxonomicName id="B48399A61806FF98460C3473AAE3ECDF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[323,468,1404,1427]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="orona">
<emphasis id="41F73E371806FF98460C3473AAE3ECDF" bold="true" box="[323,468,1404,1427]" italics="true" pageId="66" pageNumber="67">Uduba orona</emphasis>
</taxonomicName>
</emphasis>
are known only from the
<typeStatus id="AC385C871806FF9845A83473A822ECDF" box="[743,789,1404,1427]" pageId="66" pageNumber="67">type</typeStatus>
locality in montane rainforest at the Andringitra Reserve in far southern
<collectingCountry id="0B94A2B51806FF98454F3493A9B6ECFF" box="[512,641,1436,1459]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
(Maps 4, 15).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="195A414359A448C8CCFD2F5AD2A9A3E6">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="7857AC1E855404662C8B8D26EDE8A67F">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="C4CB8E4A98DE9D0D39ACC5AEC0A64564">Ubick, Darrell</mods:namePart>
<mods:affiliation id="278E267A82C3372B87301FDDDD996BE6">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="CB20D117A56702EAD2651889940C348D">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<paragraph id="733CE2251808FF96473431D3AAACE9BF" blockId="76.[123,814,220,273]" box="[123,411,220,243]" pageId="76" pageNumber="77">
<heading id="287455491808FF96473431D3AAACE9BF" box="[123,411,220,243]" fontSize="10" level="2" pageId="76" pageNumber="77" reason="4">
<emphasis id="41F73E371808FF96473431D3AAACE9BF" bold="true" box="[123,411,220,243]" pageId="76" pageNumber="77">
<taxonomicName id="B48399A61808FF96473431D3AA3EE9BF" ID-CoL="C8KJZ" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,265,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha" status="sp. nov.">
<emphasis id="41F73E371808FF96473431D3AA3EE9BF" bold="true" box="[123,265,220,243]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1808FF96465831D3AAACE9BF" box="[279,411,220,243]" pageId="76" pageNumber="77" rank="species">new species</taxonomicNameLabel>
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<paragraph id="733CE2251808FF96473431F3A819E85D" blockId="76.[123,814,220,273]" box="[123,814,252,273]" pageId="76" pageNumber="77">
<figureCitation id="EBB8FEA01808FF96473431F3AA3CE85D" box="[123,267,252,273]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: A–E = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="76" pageNumber="77">Figures 6 D, E</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96465A31F3AA0EE85D" box="[277,313,252,273]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="76" pageNumber="77">8 A</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96460B31F3AA42E85D" box="[324,373,252,273]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="76" pageNumber="77">11 D</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646CF31F3AA9EE85D" box="[384,425,252,273]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">29 I</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646FC31F3A937E85D" box="[435,512,252,273]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">59 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96454531F3A96DE85D" box="[522,602,252,273]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">71 G, H</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96452A31F3A986E85D" box="[613,689,252,273]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B, E</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9645F431F3A833E85D" box="[699,772,252,273]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="76" pageNumber="77">Maps 3</figureCitation>
, 18.
</paragraph>
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<paragraph id="733CE2251808FF9647E4302EAA76E8D4" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4302EAA67E874" bold="true" box="[171,336,289,312]" pageId="76" pageNumber="77">Type material.</emphasis>
<materialsCitation id="C3EBE8781808FF964618302EAA76E8D4" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" country="Madagascar" elevation="140" location="Parc National de Namoroka" municipality="Fisher-Griswold Arthropod Team" pageId="76" pageNumber="77" specimenCode="CASENT9018660, CASENT9082218" specimenCount="2" stateProvince="Mahajanga" typeStatus="holotype">
<typeStatus id="AC385C871808FF964618302EAA8DE874" box="[343,442,289,312]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
male (
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) and
<typeStatus id="AC385C871808FF964444302EA850E874" box="[779,871,289,312]" pageId="76" pageNumber="77" type="paratype">paratype</typeStatus>
female (
<specimenCode id="23254A5E1808FF964489302EAFB9E874" box="[966,1166,289,312]" collectionCode="CASENT" pageId="76" pageNumber="77">CASENT9082218</specimenCode>
)
<collectingMethod id="AAC29A321808FF964734304EAAAFE814" box="[123,408,321,344]" pageId="76" pageNumber="77">collected from pitfall traps</collectingMethod>
at
<quantity id="B47B4FC01808FF9646F5304EAAC0E814" box="[442,503,321,344]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="76" pageNumber="77" unit="m" value="140.0">
<elevation id="F8AE05161808FF9646F5304EAAC0E814" box="[442,503,321,344]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="76" pageNumber="77" unit="m" value="140.0">140m</elevation>
</quantity>
elevation in dry tropical forest at
<location id="765CB4FE1808FF96442B304EAFA6E814" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1808FF96442B304EAFA6E814" box="[868,1169,321,344]" country="Madagascar" municipality="Fisher-Griswold Arthropod Team" name="Parc National de Namoroka" pageId="76" pageNumber="77" stateProvince="Mahajanga">Parc National de Namoroka</location>
,
<collectingRegion id="B1472CC71808FF964734306EAA52E834" box="[123,357,353,376]" country="Madagascar" name="Mahajanga" pageId="76" pageNumber="77">Mahajanga Province</collectingRegion>
,
<collectingCountry id="0B94A2B51808FF964635306EAACEE834" box="[378,505,353,376]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
,
<date id="073DC4E51808FF96455F306EA833E834" box="[528,772,353,376]" pageId="76" pageNumber="77" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D1808FF96455F306EA833E834" box="[528,772,353,376]" pageId="76" pageNumber="77" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">4–8 November 2002</collectingDate>
</date>
,
<collectingMunicipality id="9358785F1808FF964455306EAFA5E834" box="[794,1170,353,376]" pageId="76" pageNumber="77">Fisher-Griswold Arthropod Team</collectingMunicipality>
, deposited in
<collectionCode id="15927AE01808FF964649308EAA0AE8D4" box="[262,317,385,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="76" pageNumber="77" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E430AEA816EBF5" pageId="76" pageNumber="77" type="discussion">
<paragraph id="733CE2251808FF9647E430AEA816EBF5" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E430AEAA24E8F4" bold="true" box="[171,275,417,440]" pageId="76" pageNumber="77">Remarks</emphasis>
. Males of the similar species
<taxonomicName id="B48399A61808FF96451530AEA9DEE8F4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[602,745,417,440]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96451530AEA9DEE8F4" bold="true" box="[602,745,417,440]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61808FF96446F30AEA8B3E8F4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[800,900,417,440]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96446F30AEA8B3E8F4" bold="true" box="[800,900,417,440]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
may be distinguished by the TA2 form. Association of the males and females of
<taxonomicName id="B48399A61808FF96459830CDA801E895" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[727,822,450,473]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96459830CDA801E895" bold="true" box="[727,822,450,473]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
is unproblematic in that several were collected together at Parc National de Namoroka. Two similar females, one from Manongarivo (CASENT9006108) and a second from Tsingy de Bemeraha (CASENT9009383), are more difficult to associate with species. Females of
<taxonomicName id="B48399A61808FF96452C332DA9C6EB75" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[611,753,546,569]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96452C332DA9C6EB75" bold="true" box="[611,753,546,569]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
have the epigynal plate posterior of the ML smooth or grooved and the CO hidden beneath the ML hood. The female from Manongarivo (CASENT9006108) that has the CO hidden beneath the ML hood is assigned to species
<taxonomicName id="B48399A61808FF96437E336DAFB8EB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1073,1167,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96437E336DAFB8EB35" bold="true" box="[1073,1167,610,633]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
. The female from Tsingy de Bemeraha (CASENT9009383) that has exposed copulatory openings differs from
<taxonomicName id="B48399A61808FF96464C33ADAA55EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[259,354,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96464C33ADAA55EBF5" bold="true" box="[259,354,674,697]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
and is assigned to
<taxonomicName id="B48399A61808FF96457C33ADA9A1EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[563,662,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96457C33ADA9A1EBF5" bold="true" box="[563,662,674,697]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E433CCAA20EA76" pageId="76" pageNumber="77" type="etymology">
<paragraph id="733CE2251808FF9647E433CCAA20EA76" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E433CCAA1DEB96" bold="true" box="[171,298,707,730]" pageId="76" pageNumber="77">Etymology.</emphasis>
The species epithet is a patronym in honor of Mr. Harin’Hala Rin’Ha, entomologist and explorer-naturalist who conducted most of the fieldwork for the Schlinger
<emphasis id="41F73E371808FF9644B333ECAF1BEBB6" box="[1020,1068,739,762]" italics="true" pageId="76" pageNumber="77">et al</emphasis>
., Malaise Trap Survey of
<collectingCountry id="0B94A2B51808FF96466E320CAA95EA56" box="[289,418,771,794]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
arthropods. His efforts have revealed numerous new species of insects and arachnids.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E4324CAF4FED77" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="733CE2251808FF9647E4324CAF4FED77" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4324CAA8FEA16" bold="true" box="[171,440,835,858]" pageId="76" pageNumber="77">
Diagnosis.
<taxonomicName id="B48399A61808FF964667324CAA8FEA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,440,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF964667324CAA8FEA16" bold="true" box="[296,440,835,858]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
</emphasis>
are members of Group IV, the
<taxonomicName id="B48399A61808FF96445D324CA8AEEA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[786,921,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">Uduba rinha</taxonomicName>
group (Map 18). They have a divided cribellum. Males can be distinguished from those of other
<taxonomicName id="B48399A61808FF9644DA326CA8E8EA36" box="[917,991,867,890]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371808FF9644DA326CA8E8EA36" bold="true" box="[917,991,867,890]" italics="true" pageId="76" pageNumber="77">Uduba</emphasis>
</taxonomicName>
except
<taxonomicName id="B48399A61808FF96437C326CAFA2EA36" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1075,1173,867,890]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96437C326CAFA2EA36" bold="true" box="[1075,1173,867,890]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
by the simple, trapezoidal MA (
<figureCitation id="EBB8FEA01808FF96469B328CA953EAD6" box="[468,612,899,922]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Figs. 59 A–C</figureCitation>
<emphasis id="41F73E371808FF96452B328CA95AEAD6" bold="true" box="[612,621,899,922]" pageId="76" pageNumber="77">)</emphasis>
;
<taxonomicName id="B48399A61808FF964532328CA9EBEAD6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[637,732,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF964532328CA9EBEAD6" bold="true" box="[637,732,899,922]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A61808FF964340328CAF45EAD6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1039,1138,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF964340328CAF45EAD6" bold="true" box="[1039,1138,899,922]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
by having the TA2 a flexible hook or curlicue (
<figureCitation id="EBB8FEA01808FF96451C32ACA985EAF6" box="[595,690,931,954]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">Fig. 29 I</figureCitation>
) (
<taxonomicName id="B48399A61808FF96458532ACA81BEAF6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[714,812,931,954]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96458532ACA81BEAF6" bold="true" box="[714,812,931,954]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
has the TA2 an erect, rectangular spike,
<figureCitation id="EBB8FEA01808FF96478E32CCAA1DEA96" box="[193,298,963,986]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="76" pageNumber="77">Fig. 46 B</figureCitation>
). The females can be distinguished from those of other cribellate
<taxonomicName id="B48399A61808FF9644BD32CCAF0BEA96" box="[1010,1084,963,986]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371808FF9644BD32CCAF0BEA96" bold="true" box="[1010,1084,963,986]" italics="true" pageId="76" pageNumber="77">Uduba</emphasis>
</taxonomicName>
by having the epigynum lateral lobes fused medially (
<figureCitation id="EBB8FEA01808FF96452732EBA9EDEAB7" box="[616,730,996,1019]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9645AC32EBA820EAB7" box="[739,791,996,1019]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B</figureCitation>
). The females of
<taxonomicName id="B48399A61808FF96448332EBAF1EEAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[972,1065,996,1019]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96448332EBAF1EEAB7" bold="true" box="[972,1065,996,1019]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
have copulatory openings that are hidden beneath the sides of the ML (
<figureCitation id="EBB8FEA01808FF964416350BA8E0ED57" box="[857,983,1028,1051]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9644A8350BAF16ED57" box="[999,1057,1028,1051]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B</figureCitation>
); the lone female of
<taxonomicName id="B48399A61808FF9647A4352BAA79ED77" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[235,334,1060,1083]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF9647A4352BAA79ED77" bold="true" box="[235,334,1060,1083]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
has the copulatory openings exposed just posteriad of the ML (
<figureCitation id="EBB8FEA01808FF96434E352BAF5BED77" box="[1025,1132,1060,1083]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Fig. 71 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9447E4354BAFA1EB1F" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77" type="description">
<paragraph id="733CE2251808FF9747E4354BA96BEA94" blockId="76.[122,1174,289,1725]" lastBlockId="77.[123,1174,220,1725]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4354BAA40ED17" bold="true" box="[171,375,1092,1115]" pageId="76" pageNumber="77">Description. Male</emphasis>
(
<typeStatus id="AC385C871808FF9646C8354BAAC6ED17" box="[391,497,1092,1115]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
): Total length 10.00. Markings as in
<figureCitation id="EBB8FEA01808FF964431354BA8D4ED17" box="[894,995,1092,1115]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="76" pageNumber="77">Figs. 8 A</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9644A0354BAF11ED17" box="[1007,1062,1092,1115]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="76" pageNumber="77">11 D</figureCitation>
. Carapace 5.50 long, 3.60 wide, 1.70 high; clypeus 0.20 high. Eye diameters: AME 0.16, ALE 0.20, PME 0.22, PLE 0.24. Chelicerae 2.10 long; sternum 2.30 long, 1.80 wide; labium 1.00 long; palpal coxae 1.50 long. Divided cribellum. Spination. (MRAC 174.445): palpus–femur d1-1-1; leg I–femur d0- 0-2, p0-1-2, r1-1-1-1, patella p1, r1, tibia d1-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1- 2, v2-2-2, r1-1-1; leg II–femur d1-1-1, p0-1-0-1, r0-1-0-1, patella p1, tibia d0-1-0, p0-1-1-0, v1-2- 2, r0-1-0-0, metatarsus p1-1-2, v2-2-2, r1-1-1; leg III–femur d1-1-1, p1-1-1-1, r1-1-1-1, patella r1, tibia d0-1-0, p0-1-1-0, v1-0-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-1, p0-1-0-1, r0-0-0-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v1- 1-1-2-1, r1-2-1. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements (
<typeStatus id="AC385C871808FF96431A346AAB99ECD0" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
): I: 4.50 + 2.00 + 4.30 + 4.30 + 2.60 = 17.70; II: 3.80 + 1.80 + 3.30 + 3.30 + 2.20 = 14.40; III: 3.20 + 1.40 + 2.00 + 2.50 + 1.60 = 10.70; IV: 3.20 + 1.80 + 3.30 + 2.60 + 1.60 = 12.50; palpus: 2.00 + 0.80 + 0.80 + NA + 2.30 = 5.90. Leg formula 1243.
<emphasis id="41F73E371808FF96446934CAA8AEEC90" bold="true" box="[806,921,1477,1500]" pageId="76" pageNumber="77">Male palp</emphasis>
(MRAC 174.445): palpal tibia 0.36 cymbial length, RTA long, curved to ventral, RTA length 0.59 times tibia length, length 2.00 times width, with sharply-pointed apex (
<figureCitation id="EBB8FEA01808FF9645E5370AA826EF50" box="[682,785,1541,1564]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Fig. 59 C</figureCitation>
), VTA short, conical, blunt (
<figureCitation id="EBB8FEA01808FF964371370AAB87EF71" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Figs. 59 A–C</figureCitation>
), length 1.44 times width, length 0.48 times tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 1.08 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.21 times tegulum length; TA3 a transverse hook, TA3 extends apicad of TA2 by 1.175 tegulum length, TA2 small, curlicue-shaped, soft and flexible, extends partially in front of TA3 (
<figureCitation id="EBB8FEA01808FF96465C37A9AAB6EFF1" box="[275,385,1702,1725]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">Figs. 29 I</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646C137A9AAF1EFF1" box="[398,454,1702,1725]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">59 A</figureCitation>
); MA a broad, flat trapezoid, origin at 0.40 times tegulum length, extends nearly to tegulum apex, tegulum apex at 0.09 tegulum length from MA apex, length (including apical processes) 1.05 times width, MA large, length 0.38 tegulum length, width 0.39 tegulum width (
<figureCitation id="EBB8FEA01809FF9747B73013AABDE87F" box="[248,394,284,307]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="77" pageNumber="78">Figs. 59 A, B</figureCitation>
); conductor fan entire (
<figureCitation id="EBB8FEA01809FF9745C73013A9C6E87F" box="[648,753,284,307]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="77" pageNumber="78">Fig. 59 C</figureCitation>
).
<emphasis id="41F73E371809FF97444A3013A86FE87F" bold="true" box="[773,856,284,307]" pageId="77" pageNumber="78">Female</emphasis>
(
<typeStatus id="AC385C871809FF9744283013A8FDE87F" box="[871,970,284,307]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
): Total length 8.60. Markings as in
<figureCitation id="EBB8FEA01809FF9746533033AAACE81F" box="[284,411,316,339]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: A–E = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="77" pageNumber="78">Figs. 6 D, E</figureCitation>
. Carapace 4.80 long, 3.40 wide, 1.90 high; clypeus 0.30 high. Eye diameter: AME 0.14, ALE 0.18, PME 0.20. Chelicera 2.20 long; sternum 2.10 long, 0.80 wide; labium 1.20 long; palpal coxae 1.60 long. Divided cribellum. Spination: palpus–femur d0-1-2, patella p1- 0, tibia p2-0, tarsus p2-1; leg I–femur d1-0-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II– femur d1-0-0, p0-0-1, tibia p0-1-0, v1-1-2, metatarsus p0-1-0, v2-2-3; leg III–femur d1-0-0-1, p1- 0-1, r0-1-0, tibia d1-1-0, p0-1-1-0, v1-1-2, r0-1-1-0, metatarsus d1-0-0-0, p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-0-0-0, tibia v1-2-2, r1-0-1, metatarsus p0-0-0-2, v2-1-2-2, r0-0-1- 2. Scopulae: ventral on tarsi I–IV and apices of metatarsi I and II. Leg measurements: I: 3.50 + 1.80 + 2.80 + 2.40 + 1.90 = 12.40; II: 3.00 + 1.60 + 2.30 + 2.10 + 1.70 = 10.70; III: 2.50 + 1.20 + 1.30 + 1.70 + 1.30 = 8.00; IV: 3.50 + 1.60 + 2.70 + 2.80 + 1.50 = 12.10; palpus: 1.80 + 0.80 + 0.90 + NA + 1.50 = 5.00. Leg formula 1423.
<emphasis id="41F73E371809FF9745313371A80CEBD9" bold="true" box="[638,827,638,661]" pageId="77" pageNumber="78">Female genitalia</emphasis>
(
<typeStatus id="AC385C871809FF9744053371A89AEBD9" box="[842,941,638,661]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
): epigynal plate without depressed atrium, plate width 1.32 times length (
<figureCitation id="EBB8FEA01809FF9745E03390A82EEBFA" box="[687,793,671,694]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 G</figureCitation>
); median lobe and lateral lobes differentiated only in anterior half of plate, distance from posterior margin of ML to epigastric furrow 0.83 times epigynum length; median lobe (
<figureCitation id="EBB8FEA01809FF97450033D0A98DEBBA" box="[591,698,735,758]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">Fig. 81 B</figureCitation>
), a flat plate with lateral margins extending around to posterior, nearly meeting in middle, broad, ML width between bases of LL 0.59 times epigynum width, ML width 4.20 times LL width at sides of ML; median lobe width 0.38 times epigynum width, ML width 0.45 times ML length, with copulatory openings hidden beneath lateral margins of ML (
<figureCitation id="EBB8FEA01809FF974609326FAA85EA3B" box="[326,434,864,887]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 G</figureCitation>
), distance between copulatory openings 0.56 times epigynum width, CO origin at 0.70 of epigynum length. Vulva (
<figureCitation id="EBB8FEA01809FF974538328FA9D5EADB" box="[631,738,896,919]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 H</figureCitation>
) with spermathecal ducts forming three longitudinal loops, vulva length 0.61 times width, fertilization ducts widely separated, vulva width 2.68 times distance between FD (
<figureCitation id="EBB8FEA01809FF9746A932CEA978EA94" box="[486,591,961,984]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">Fig. 81 E</figureCitation>
).
</paragraph>
<paragraph id="733CE2251809FF9747E432EEA993EF50" blockId="77.[123,1174,220,1725]" pageId="77" pageNumber="78">
<emphasis id="41F73E371809FF9747E432EEAA6BEAB4" bold="true" box="[171,348,993,1016]" pageId="77" pageNumber="78">Variation. Male</emphasis>
(N= 5): Total length 8.14–10.00; carapace length / width = 1.40–1.54, carapace height / width = 0.42–0.56, PER / carapace width = 0.36–0.81, PER / OAL = 2.48–2.92, PER / AER = 1.31–1.42, OAL / OQL = 1.07–1.15, OQP / OQA = 1.10–1.43, clypeus height / AME = 1.18–1.56, cheliceral length / clypeus height = 8.00–11.60, sternum length / width = 1.28–1.41, palpal coxa length / width = 2.41–3.35, femur I length / carapace width = 1.22–1.26, metatarsus I length / carapace width = 1.17–1.35, femur IV length / carapace width = 0.89–1.45, cymbium length / carapace width = 0.56–0.70, cymbium length / palpal patella length = 2.36–2.91, cymbium length / palpal tibia length = 2.50–2.91, cymbium length / palpal femur length = 0.71–1.15, palpal tibia length / palpal patella length = 0.93–1.09.
<emphasis id="41F73E371809FF9745ED35ECA9C2EDB6" bold="true" box="[674,757,1251,1274]" pageId="77" pageNumber="78">Female</emphasis>
(N= 3): Total length 8.60–10.60; carapace length / width = 1.41–1.56, carapace height / width = 0.54–0.62, PER / carapace width = 0.39– 0.44, PER / OAL = 3.13–3.19, PER / AER = 1.32–1.39, OAL / OQL = 1.05–1.29, OQP / OQA = 1.12–1.23, clypeus height / AME = 1.75–2.50, cheliceral length / clypeus height = 6.50–7.71, sternum length / width = 1.17–1.35, palpal coxa length / width = 2.29–2.38, femur I length / carapace width = 0.97–1.05, metatarsus I length / carapace width = 0.71–0.85, femur IV length / carapace width = 0.95–1.05, cymbium length / carapace width = 0.41–0.44, palpal tarsus length / palpal patella length = 1.67–1.88, palpal tarsus length / palpal tibia length = 1.50–1.67, palpal tarsus length / palpal femur length = 0.81–1.00, palpal tibia length / palpal patella length = 1.10–1.13. Female genitalia vary as in
<figureCitation id="EBB8FEA01809FF9746E8370AA909EF50" box="[423,574,1541,1564]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Figs. 71 G, H</figureCitation>
,
<figureCitation id="EBB8FEA01809FF974505370AA993EF50" box="[586,676,1541,1564]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">81 B, E.</figureCitation>
</paragraph>
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<materialsCitation id="C3EBE8781809FF9747E4372AA96AEF10" collectingDate="1988-10" collectionCode="USNM" collectorName="W. Steiner" country="Madagascar" location="Antananarivo Province" pageId="77" pageNumber="78" specimenCode="CASENT9006040" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
M
<emphasis id="41F73E371809FF97478D372AAABFEF70" bold="true" box="[194,392,1573,1596]" pageId="77" pageNumber="78">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51809FF9746C1372AA967EF70" box="[398,592,1573,1596]" name="Madagascar" pageId="77" pageNumber="78">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371809FF974515372AA865EF70" box="[602,850,1573,1596]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF974515372AA865EF70" box="[602,850,1573,1596]" country="Madagascar" name="Antananarivo" pageId="77" pageNumber="78">Antananarivo Province</collectingRegion>
</emphasis>
:
<collectingRegion id="B1472CC71809FF974411372AA8C5EF70" box="[862,1010,1573,1596]" country="Madagascar" name="Antananarivo" pageId="77" pageNumber="78">Antananarivo</collectingRegion>
,
<date id="073DC4E51809FF9744B3372AAFA6EF70" box="[1020,1169,1573,1596]" pageId="77" pageNumber="78" value="1988-10">
<collectingDate id="17793D0D1809FF9744B3372AAFA6EF70" box="[1020,1169,1573,1596]" pageId="77" pageNumber="78" value="1988-10">October 1988</collectingDate>
</date>
,
<collectorName id="DE7687F31809FF974734374AABDCEF10" box="[123,235,1605,1628]" pageId="77" pageNumber="78">W. Steiner</collectorName>
(
<specimenCode id="23254A5E1809FF9747B4374AAAF2EF10" box="[251,453,1605,1628]" collectionCode="CASENT" pageId="77" pageNumber="78">CASENT9006040</specimenCode>
,
<specimenCount id="658529AC1809FF97469D374AAAC4EF10" box="[466,499,1605,1628]" count="1" pageId="77" pageNumber="78" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01809FF97454B374AA961EF10" box="[516,598,1605,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="77" pageNumber="78" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781809FF974525374AA83BEFD1" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.961666" location="Manongarivo" longLatPrecision="21" longitude="48.433334" pageId="77" pageNumber="78" specimenCode="CASENT9006108" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<emphasis id="41F73E371809FF974525374AA862EF10" box="[618,853,1605,1628]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF974525374AA862EF10" box="[618,853,1605,1628]" country="Madagascar" name="Antsiranana" pageId="77" pageNumber="78">Antsiranana Province</collectingRegion>
</emphasis>
: R.S.
<location id="765CB4FE1809FF9744D1374AAF07EF10" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1809FF9744D1374AAF07EF10" box="[926,1072,1605,1628]" country="Madagascar" latitude="-13.961666" longLatPrecision="21" longitude="48.433334" name="Manongarivo" pageId="77" pageNumber="78" stateProvince="Antsiranana">Manongarivo</location>
,
<locationDeviation id="628EFB071809FF974371374AAA52EF30" location="Antanambao" pageId="77" pageNumber="78">
<quantity id="B47B4FC01809FF974371374AAFA2EF10" box="[1086,1173,1605,1628]" metricMagnitude="4" metricUnit="m" metricValue="1.08" pageId="77" pageNumber="78" unit="km" value="10.8">10.8 km</quantity>
229°SW Antanambao
</locationDeviation>
,
<geoCoordinate id="16B784E21809FF97463D376AAADBEF31" box="[370,492,1637,1661]" degrees="13" direction="south" minutes="57" orientation="latitude" pageId="77" pageNumber="78" precision="15" seconds="42" value="-13.961666">13°57ʹ42ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21809FF9746B53769A9B2EF31" box="[506,645,1638,1661]" degrees="048" direction="east" minutes="26" orientation="longitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="48.433334">048°26ʹ00ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01809FF9745853769A83EEF31" box="[714,777,1638,1661]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="77" pageNumber="78" unit="m" value="400.0">
<elevation id="F8AE05161809FF9745853769A83EEF31" box="[714,777,1638,1661]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="77" pageNumber="78" unit="m" value="400.0">400m</elevation>
</quantity>
, sifted litter, rainforest,
<date id="073DC4E51809FF9743403769AB84EFD1" pageId="77" pageNumber="78" value="1998-11-08">
<collectingDate id="17793D0D1809FF9743403769AB84EFD1" pageId="77" pageNumber="78" value="1998-11-08">8 November 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31809FF9747F23789AA0AEFD1" box="[189,317,1670,1693]" pageId="77" pageNumber="78">B. L. Fisher</collectorName>
[BLF1996] (
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,
<specimenCount id="658529AC1809FF9745D23789A989EFD1" box="[669,702,1670,1693]" count="1" pageId="77" pageNumber="78" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01809FF9745833789A833EFD1" box="[716,772,1670,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="77" pageNumber="78" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781809FF9444593789A82DE81F" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" collectorName="Fisher &amp; Griswold &amp; Male &amp; Female" country="Madagascar" elevation="140" lastPageId="78" lastPageNumber="79" latitude="-16.466667" location="Parc National de Namoroka" longLatPrecision="21" longitude="45.35" pageId="77" pageNumber="78" specimenCode="CASENT9018660, CASENT9082218, CASENT9065644, CASENT9065645, CASENT9065646, CASENT9065647, CASENT9065648" specimenCount="13" specimenCount-female="2" specimenCount-male="11" stateProvince="Mahajanga" typeStatus="holotype">
<emphasis id="41F73E371809FF9744593789A8C2EFD1" box="[790,1013,1670,1693]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF9744593789A8C2EFD1" box="[790,1013,1670,1693]" country="Madagascar" name="Mahajanga" pageId="77" pageNumber="78">Mahajanga Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1809FF97434B3789AA39EFF1" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1809FF97434B3789AA39EFF1" country="Madagascar" latitude="-16.466667" longLatPrecision="21" longitude="45.35" name="Parc National de Namoroka" pageId="77" pageNumber="78" stateProvince="Mahajanga">Parc National de Namoroka</location>
,
<locationDeviation id="628EFB071809FF97465637A9A956EFF1" box="[281,609,1702,1725]" location="Vilanandro" pageId="77" pageNumber="78">
<quantity id="B47B4FC01809FF97465637A9AA53EFF1" box="[281,356,1702,1725]" metricMagnitude="3" metricUnit="m" metricValue="9.8" pageId="77" pageNumber="78" unit="km" value="9.8">9.8 km</quantity>
300°WNW Vilanandro
</locationDeviation>
,
<geoCoordinate id="16B784E21809FF97452437A9A9D2EFF1" box="[619,741,1702,1725]" degrees="16" direction="south" minutes="28" orientation="latitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="-16.466667">16°28ʹ00ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21809FF9745BD37A9A84AEFF1" box="[754,893,1702,1725]" degrees="045" direction="east" minutes="21" orientation="longitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="45.35">045°21ʹ00ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01809FF9744F037A9A8C9EFF1" box="[959,1022,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="77" pageNumber="78" unit="m" value="140.0">
<elevation id="F8AE05161809FF9744F037A9A8C9EFF1" box="[959,1022,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="77" pageNumber="78" unit="m" value="140.0">140m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321809FF97434637A9AF4EEFF1" box="[1033,1145,1702,1725]" pageId="77" pageNumber="78">pitfall trap</collectingMethod>
in tropical dry forest,
<date id="073DC4E5180AFF94461931D3A909E9BF" box="[342,574,220,243]" pageId="78" pageNumber="79" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D180AFF94461931D3A909E9BF" box="[342,574,220,243]" pageId="78" pageNumber="79" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">4–8 November 2002</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF94450131D3A9B9E9BF" box="[590,654,220,243]" pageId="78" pageNumber="79">Fisher</collectorName>
,
<collectorName id="DE7687F3180AFF9445EE31D3A833E9BF" box="[673,772,220,243]" pageId="78" pageNumber="79">Griswold</collectorName>
<emphasis id="41F73E37180AFF94445F31D3A87DE9BF" box="[784,842,220,243]" italics="true" pageId="78" pageNumber="79">et al.</emphasis>
[BLF6444], (
<typeStatus id="AC385C87180AFF9444A831D3AF7BE9BF" box="[999,1100,220,243]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
<collectorName id="DE7687F3180AFF94431731D3AFA6E9BF" box="[1112,1169,220,243]" pageId="78" pageNumber="79">Male</collectorName>
,
<specimenCode id="23254A5E180AFF94473431F3AA7AE85F" box="[123,333,252,275]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9018660</specimenCode>
,
<specimenCount id="658529AC180AFF94462031F3AAA6E85F" box="[367,401,252,275]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9446FC31F3AADCE85F" box="[435,491,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C87180AFF94455231F3A9B3E85F" box="[541,644,252,275]" pageId="78" pageNumber="79" type="paratype">Paratype</typeStatus>
<collectorName id="DE7687F3180AFF9445D031F3A9C4E85F" box="[671,755,252,275]" pageId="78" pageNumber="79">Female</collectorName>
,
<specimenCode id="23254A5E180AFF94445A31F3A8D0E85F" box="[789,999,252,275]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9082218</specimenCode>
,
<specimenCount id="658529AC180AFF94434631F3AF1CE85F" box="[1033,1067,252,275]" count="1" pageId="78" pageNumber="79" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0180AFF94430131F3AFBEE85F" box="[1102,1161,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9447CD3013AA7BE87F" box="[130,332,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065644</specimenCode>
,
<specimenCount id="658529AC180AFF9446193013AA40E87F" box="[342,375,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<specimenCount id="658529AC180AFF9446323013AAA8E87F" box="[381,415,284,307]" count="1" pageId="78" pageNumber="79" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0180AFF9446E33013AAD4E87F" box="[428,483,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9446B23013A9F1E87F" box="[509,710,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065645</specimenCode>
,
<specimenCount id="658529AC180AFF94459E3013A9C5E87F" box="[721,754,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445B03013A801E87F" box="[767,822,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF94441F3013AF2DE87F" box="[848,1050,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065646</specimenCode>
,
<specimenCount id="658529AC180AFF94436C3013AF72E87F" box="[1059,1093,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF94431D3013AFBEE87F" box="[1106,1161,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9447CD3033AA7CE81F" box="[130,331,316,339]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065647</specimenCode>
,
<specimenCount id="658529AC180AFF9446193033AA40E81F" box="[342,375,316,339]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9446CA3033AA8BE81F" box="[389,444,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9446993033A997E81F" box="[470,672,316,339]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065648</specimenCode>
,
<specimenCount id="658529AC180AFF9445E43033A9FBE81F" box="[683,716,316,339]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445953033A825E81F" box="[730,786,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE878180AFF94446D3033A82EE8DF" collectingDate="2007-10-09" collectingDateMax="2007-10-16" collectingDateMin="2007-10-09" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="20" location="Analamanitra Forest" pageId="78" pageNumber="79" specimenCode="CASENT9042527" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="holotype">
<location id="765CB4FE180AFF94446D3033A8C8E81F" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE180AFF94446D3033A8C8E81F" box="[802,1023,316,339]" country="Madagascar" name="Analamanitra Forest" pageId="78" pageNumber="79" stateProvince="Mahajanga">Analamanitra Forest</location>
,
<locationDeviation id="628EFB07180AFF9443453033ABE6E83F" location="Misinjo" pageId="78" pageNumber="79">
<quantity id="B47B4FC0180AFF9443453033AF79E81F" box="[1034,1102,316,339]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="78" pageNumber="79" unit="km" value="14.0">14 km</quantity>
NE of Misinjo
</locationDeviation>
, 16°8ʹS, 45°42ʹE, elev.
<quantity id="B47B4FC0180AFF9446813053A937E83F" box="[462,512,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="78" pageNumber="79" unit="m" value="20.0">
<elevation id="F8AE0516180AFF9446813053A937E83F" box="[462,512,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="78" pageNumber="79" unit="m" value="20.0">20m</elevation>
</quantity>
, dense dry forest, malaise,
<date id="073DC4E5180AFF9444673053A8CCE83F" box="[808,1019,348,371]" pageId="78" pageNumber="79" value="2007-10-09" valueMax="2007-10-16" valueMin="2007-10-09">
<collectingDate id="17793D0D180AFF9444673053A8CCE83F" box="[808,1019,348,371]" pageId="78" pageNumber="79" value="2007-10-09" valueMax="2007-10-16" valueMin="2007-10-09">9–16 October 2007</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF9443483053AF50E83F" box="[1031,1127,348,371]" pageId="78" pageNumber="79">M. Irwin</collectorName>
and
<collectorName id="DE7687F3180AFF9447343073AA2CE8DF" box="[123,283,380,403]" pageId="78" pageNumber="79">R. Harin’Hala</collectorName>
[MG-38-04] (
<specimenCode id="23254A5E180AFF94468A3073A9B9E8DF" box="[453,654,380,403]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9042527</specimenCode>
,
<specimenCount id="658529AC180AFF9445ED3073A9F4E8DF" box="[674,707,380,403]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445963073A826E8DF" box="[729,785,380,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE878180AFF9444633073A9F0E89F" collectingDate="1991-10-01" collectingDateMax="1991-10-25" collectingDateMin="1991-10-01" collectionCode="MRAC" collectorName="A. Pauly" country="Madagascar" location="25 km W Morarano Chrome" pageId="78" pageNumber="79" specimenCode="MRAC174.445" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="41F73E37180AFF9444633073AF3BE8DF" box="[812,1036,380,403]" italics="true" pageId="78" pageNumber="79">
<collectingRegion id="B1472CC7180AFF9444633073AF3BE8DF" box="[812,1036,380,403]" country="Madagascar" name="Toamasina" pageId="78" pageNumber="79">Toamasina Province</collectingRegion>
</emphasis>
:
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<quantity id="B47B4FC0180AFF94436C3073AF59E8DF" box="[1059,1134,380,403]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="78" pageNumber="79" unit="km" value="25.0">25 km</quantity>
W Morarano Chrome
</location>
, (coordinates approximate) 17º45ʹS, 48º10ʹE, forest, yellow plate,
<date id="073DC4E5180AFF94432F3093AA27E89F" pageId="78" pageNumber="79" value="1991-10-01" valueMax="1991-10-25" valueMin="1991-10-01">
<collectingDate id="17793D0D180AFF94432F3093AA27E89F" pageId="78" pageNumber="79" value="1991-10-01" valueMax="1991-10-25" valueMin="1991-10-01">1–25 October 1991</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF94465630B3AA4FE89F" box="[281,376,444,467]" pageId="78" pageNumber="79">A. Pauly</collectorName>
(
<specimenCode id="23254A5E180AFF9446C830B3A907E89F" box="[391,560,444,467]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="78" pageNumber="79" type="Museum">MRAC174.445</specimenCode>
,
<specimenCount id="658529AC180AFF94457330B3A969E89F" box="[572,606,444,467]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF94452330B3A988E89F" box="[620,703,444,467]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="78" pageNumber="79" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225180AFF9447E430D3AFA1EB1F" blockId="78.[123,1174,220,659]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E430D3AA56E8BF" bold="true" box="[171,353,476,499]" pageId="78" pageNumber="79">Natural history.</emphasis>
Collections are from dry forest and there is even
<specimenCount id="658529AC180AFF9444C430D3AF17E8BF" box="[907,1056,476,499]" count="1" pageId="78" pageNumber="79" type="generic">one specimen</specimenCount>
labeled as coming from
<collectingRegion id="B1472CC7180AFF94464330F3AA97EB5F" box="[268,416,508,531]" country="Madagascar" name="Antananarivo" pageId="78" pageNumber="79">Antananarivo</collectingRegion>
, although the habitat for this specimen, urban or not, is not specified. Specimens were taken on the forest floor in sifted litter and yellow pan traps and above the floor in a malaise trap.
<taxonomicName id="B48399A6180AFF94467A3333AAF5EB1F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[309,450,572,595]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E37180AFF94467A3333AAF5EB1F" bold="true" box="[309,450,572,595]" italics="true" pageId="78" pageNumber="79">Uduba rinha</emphasis>
</taxonomicName>
are cribellate but we have no records of how they use cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF9447E43353AFA1EBDF" pageId="78" pageNumber="79" type="distribution">
<paragraph id="733CE225180AFF9447E43353AFA1EBDF" blockId="78.[123,1174,220,659]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E43353AA0AEB3F" bold="true" box="[171,317,604,627]" pageId="78" pageNumber="79">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6180AFF9446843353A96DEB3F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[459,602,604,627]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E37180AFF9446843353A96DEB3F" bold="true" box="[459,602,604,627]" italics="true" pageId="78" pageNumber="79">Uduba rinha</emphasis>
</taxonomicName>
is recorded on the west side of the great escarpment across the northern half of
<collectingCountry id="0B94A2B5180AFF9446D83373A92FEBDF" box="[407,536,636,659]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
on the high plateau and in western dry forests (
<figureCitation id="EBB8FEA0180AFF9443413373AF57EBDF" box="[1038,1120,636,659]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="78" pageNumber="79">Maps 3</figureCitation>
, 18).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="77DFF6242E2B8DE8ADCB68C2503C2F72">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="23AC2C05DDE023B6D6E37B34CBE6D37C">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="9B29828A458A91F7EAE18A395A6F5B7B">Ubick, Darrell</mods:namePart>
<mods:affiliation id="A9EFB9D0A3C6E9ABC4B5ABF472C8815D">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="17CCE382BB2283DC238DE51736DF144A">Ledford, Joel</mods:namePart>
<mods:affiliation id="0DE18E226B596082A6BE377769D5DCD4">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="485C3A62FDBD2374EA46061B1CFF099A">Polotow, Daniele</mods:namePart>
<mods:affiliation id="8F3113AF992DAE88678F5895CC19A923">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:date id="35DA670168948CB728830D07BF26FECD">2022</mods:date>
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<paragraph id="733CE225180AFF94473433B3AA95EB9F" blockId="78.[123,644,700,753]" box="[123,418,700,723]" pageId="78" pageNumber="79">
<heading id="28745549180AFF94473433B3AA95EB9F" box="[123,418,700,723]" fontSize="10" level="2" pageId="78" pageNumber="79" reason="4">
<emphasis id="41F73E37180AFF94473433B3AA95EB9F" bold="true" box="[123,418,700,723]" pageId="78" pageNumber="79">
<taxonomicName id="B48399A6180AFF94473433B3AA27EB9F" ID-CoL="C8KK2" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,272,700,723]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="salegy" status="sp. nov.">
<emphasis id="41F73E37180AFF94473433B3AA27EB9F" bold="true" box="[123,272,700,723]" italics="true" pageId="78" pageNumber="79">Uduba salegy</emphasis>
</taxonomicName>
,
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</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF94473433D3A9B3EBBD" box="[123,644,732,753]" pageId="78" pageNumber="79" type="description">
<paragraph id="733CE225180AFF94473433D3A9B3EBBD" blockId="78.[123,644,700,753]" box="[123,644,732,753]" pageId="78" pageNumber="79">
<figureCitation id="EBB8FEA0180AFF94473433D3ABDAEBBD" box="[123,237,732,753]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="78" pageNumber="79">Figures 7 E</figureCitation>
,
<figureCitation id="EBB8FEA0180AFF9447B833D3AA2AEBBD" box="[247,285,732,753]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="78" pageNumber="79">9 D</figureCitation>
,
<figureCitation id="EBB8FEA0180AFF94466733D3AA60EBBD" box="[296,343,732,753]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="78" pageNumber="79">11 A</figureCitation>
,
<figureCitation id="EBB8FEA0180AFF94462D33D3AA98EBBD" box="[354,431,732,753]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="78" pageNumber="79">60 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180AFF9446F633D3A931EBBD" box="[441,518,732,753]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="78" pageNumber="79">74 A, B</figureCitation>
, Maps 2, 13.
</paragraph>
</subSubSection>
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<emphasis id="41F73E37180AFF9447E4320AAA78EA50" bold="true" box="[171,335,773,796]" pageId="78" pageNumber="79">Type material.</emphasis>
<materialsCitation id="C3EBE878180AFF944618320AA849EA10" collectingDate="1993-11-21" collectingDateMax="1993-11-30" collectingDateMin="1993-11-21" collectionCode="USNM" country="Madagascar" elevation="1000" location="Parc National Montagne d'Ambre" municipality="Jonathan Coddington" pageId="78" pageNumber="79" specimenCode="CASENT9006044" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="AC385C87180AFF944618320AAA8DEA50" box="[343,442,773,796]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
male (
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) from montane rainforest at
<quantity id="B47B4FC0180AFF94434C320AAF7BEA50" box="[1027,1100,773,796]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="79" unit="m" value="1000.0">
<elevation id="F8AE0516180AFF94434C320AAF7BEA50" box="[1027,1100,773,796]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="79" unit="m" value="1000.0">1000m</elevation>
</quantity>
elevation in
<location id="765CB4FE180AFF94479F322AA97BEA70" LSID="urn:lsid:plazi:treatment:FB2A5333180AFF8A473433B3AACFEA79:765CB4FE180AFF94479F322AA97BEA70" box="[208,588,805,828]" country="Madagascar" municipality="Jonathan Coddington" name="Parc National Montagne d'Ambre" pageId="78" pageNumber="79" stateProvince="Antsiranana">Parc National Montagne d’Ambre</location>
,
<collectingRegion id="B1472CC7180AFF944516322AA870EA70" box="[601,839,805,828]" country="Madagascar" name="Antsiranana" pageId="78" pageNumber="79">Antsiranana Province</collectingRegion>
,
<collectingCountry id="0B94A2B5180AFF944419322AA8E2EA70" box="[854,981,805,828]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
, collected
<date id="073DC4E5180AFF94431C322AAA18EA10" pageId="78" pageNumber="79" value="1993-11-21" valueMax="1993-11-30" valueMin="1993-11-21">
<collectingDate id="17793D0D180AFF94431C322AAA18EA10" pageId="78" pageNumber="79" value="1993-11-21" valueMax="1993-11-30" valueMin="1993-11-21">21–30 November 1993</collectingDate>
</date>
by
<collectingMunicipality id="9358785F180AFF944629324AA962EA10" box="[358,597,837,860]" pageId="78" pageNumber="79">Jonathan Coddington</collectingMunicipality>
, deposited in the
<collectionCode id="15927AE0180AFF944462324AA849EA10" box="[813,894,837,860]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="78" pageNumber="79" type="Museum">USNM</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE878180AFF9444C0324AA8ECEAD0" collectingDate="1992-08-12" collectionCode="CAS" country="Madagascar" county="Barbara Roth" location="Parc National Montagne d'Ambre" municipality="Vince" pageId="78" pageNumber="79" specimenCode="CASENT9006045" specimenCount="2" specimenCount-female="2" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="AC385C87180AFF9444C0324AA8CEEA10" box="[911,1017,837,860]" pageId="78" pageNumber="79" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="658529AC180AFF944343324AAFA2EA10" box="[1036,1173,837,860]" count="2" pageId="78" pageNumber="79" type="female">two females</specimenCount>
(
<specimenCode id="23254A5E180AFF9447CD326AAA7AEA30" box="[130,333,869,892]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9006045</specimenCode>
), also from
<location id="765CB4FE180AFF94469B326AA87CEA30" LSID="urn:lsid:plazi:treatment:FB2A5333180AFF8A473433B3AACFEA79:765CB4FE180AFF94469B326AA87CEA30" box="[468,843,869,892]" country="Madagascar" county="Barbara Roth" municipality="Vince" name="Parc National Montagne d'Ambre" pageId="78" pageNumber="79" stateProvince="Antsiranana">Parc National Montagne d’Ambre</location>
,
<collectingRegion id="B1472CC7180AFF944419326AAF74EA30" box="[854,1091,869,892]" country="Madagascar" name="Antsiranana" pageId="78" pageNumber="79">Antsiranana Province</collectingRegion>
,
<collectingCountry id="0B94A2B5180AFF94431F326AAB8AEAD0" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
, collected
<date id="073DC4E5180AFF94467B328AAAEAEAD0" box="[308,477,901,924]" pageId="78" pageNumber="79" value="1992-08-12">
<collectingDate id="17793D0D180AFF94467B328AAAEAEAD0" box="[308,477,901,924]" pageId="78" pageNumber="79" value="1992-08-12">12 August 1992</collectingDate>
</date>
by
<collectingMunicipality id="9358785F180AFF944548328AA974EAD0" box="[519,579,901,924]" pageId="78" pageNumber="79">Vince</collectingMunicipality>
and
<collectingCounty id="9A5D9AA9180AFF944535328AA839EAD0" box="[634,782,901,924]" pageId="78" pageNumber="79">Barbara Roth</collectingCounty>
, deposited in
<collectionCode id="15927AE0180AFF9444EB328AA8ECEAD0" box="[932,987,901,924]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF9447E432AAA9CFEA90" pageId="78" pageNumber="79" type="etymology">
<paragraph id="733CE225180AFF9447E432AAA9CFEA90" blockId="78.[123,1174,773,1725]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E432AAAA1DEAF0" bold="true" box="[171,298,933,956]" pageId="78" pageNumber="79">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E37180AFF94455E32AAA96FEAF0" box="[529,600,933,956]" italics="true" pageId="78" pageNumber="79">Salegy</emphasis>
, is a popular music genre from
<collectingCountry id="0B94A2B5180AFF9444F732AAAF0EEAF0" box="[952,1081,933,956]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
that has its home in the north of
<collectingCountry id="0B94A2B5180AFF9446CC32CAA930EA90" box="[387,519,965,988]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF9447E432EAAA99ED90" pageId="78" pageNumber="79" type="diagnosis">
<paragraph id="733CE225180AFF9447E432EAAA99ED90" blockId="78.[123,1174,773,1725]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E432EAAA8EEAB0" bold="true" box="[171,441,997,1020]" pageId="78" pageNumber="79">
Diagnosis.
<taxonomicName id="B48399A6180AFF94466A32EAAA8EEAB0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,441,997,1020]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="salegy">
<emphasis id="41F73E37180AFF94466A32EAAA8EEAB0" bold="true" box="[293,441,997,1020]" italics="true" pageId="78" pageNumber="79">Uduba salegy</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.a, the
<taxonomicName id="B48399A6180AFF94444F32EAA8B5EAB0" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[768,898,997,1020]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA0180AFF94449B32EAAF10EAB0" box="[980,1063,997,1020]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="78" pageNumber="79">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are cribellate, with large, divided cribellum or vestige. Males can be distinguished from those of other
<taxonomicName id="B48399A6180AFF944503352AA9A1ED70" box="[588,662,1061,1084]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180AFF944503352AA9A1ED70" bold="true" box="[588,662,1061,1084]" italics="true" pageId="78" pageNumber="79">Uduba</emphasis>
</taxonomicName>
with a screw-shaped TA3 by having the TA3 J-shaped and extending apicad of TA2, MA small, simple (
<figureCitation id="EBB8FEA0180AFF944442354AA891ED10" box="[781,934,1093,1116]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="78" pageNumber="79">Figs. 60 A–C</figureCitation>
). The females can be distinguished from those of other
<taxonomicName id="B48399A6180AFF9446AC356AA91AED30" box="[483,557,1125,1148]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180AFF9446AC356AA91AED30" bold="true" box="[483,557,1125,1148]" italics="true" pageId="78" pageNumber="79">Uduba</emphasis>
</taxonomicName>
of the of Group I, the Epigynal atrium group, which have an epigynal plate with the median lobe surrounded by a depressed atrium, by having the median lobe end well anteriad of epigastric furrow, with the posterior margin of atrium entire, forming a posterior pocket (
<figureCitation id="EBB8FEA0180AFF94467735CAAA95ED90" box="[312,418,1221,1244]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="78" pageNumber="79">Fig. 74 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF8A47E435EAABE0EBB8" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="description">
<paragraph id="733CE225180AFF9547E435EAAA97EDD5" blockId="78.[123,1174,773,1725]" lastBlockId="79.[123,1174,220,1725]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E435EAAA40EDB0" bold="true" box="[171,375,1253,1276]" pageId="78" pageNumber="79">Description. Male</emphasis>
(
<typeStatus id="AC385C87180AFF9446C835EAAAC6EDB0" box="[391,497,1253,1276]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
): Total length 10.00. Markings as in
<figureCitation id="EBB8FEA0180AFF94443135EAA8D2EDB0" box="[894,997,1253,1276]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="78" pageNumber="79">Figs. 9 D</figureCitation>
,
<figureCitation id="EBB8FEA0180AFF9444BD35EAAF11EDB0" box="[1010,1062,1253,1276]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="78" pageNumber="79">11 A</figureCitation>
. Carapace 5.79 long, 4.07 wide, 1.79 high; clypeus 0.25 high. Eye diameters: AME 0.20, ALE and PME 0.21, PLE 0.23. Chelicerae 2.29 long; sternum 2.71 long, 2.04 wide; labium 1.07 long; palpal coxae 1.57 long. Divided cribellum. Spination: palpus–femur d0-0-1, p0-0-1, r0-0-1; leg I–femur d1-0-0-1, p0-0-2, r0-1-0-1, patella p1, tibia d0-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2-2- 3, r1-1-0; leg II–femur d0-1-0-1, p0-1-2, r0-1-0-1, patella p1, tibia d0-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-1-1, r1-1-1; leg III–femur d0-0-1-1, p0-1-1-1, r0-1-1-1, tibia d1-0- 1-0, p0-1-1-0, v1-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0- 1, p0-0-0-1, r0-0-0-1, patella r1, tibia d0-1-1-0, p0-1-0-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v2- 2-2, r0-1-2. Scopulae: cymbium, apicodorsal; weak beneath tarsi and apices of all metatarsi. Leg measurements: I: 5.00 + 2.14 + 5.07 + 5.14 + 2.93 = 20.09; II: 4.50 + 1.93 + 3.79 + 4.29 + 2.50 = 17.00; III: 3.71 + 1.79 + 2.00 + 3.07 + 1.93 = 12.50; IV: 5.00 + 1.93 + 4.00 + 5.07 + 2.36 = 18.36; palpus: 2.29 + 0.93 + 0.86 + NA + 2.50 = 6.57. Leg formula 1423.
<emphasis id="41F73E37180AFF9444093769A88EEF31" bold="true" box="[838,953,1638,1661]" pageId="78" pageNumber="79">Male palp</emphasis>
(
<figureCitation id="EBB8FEA0180AFF9444873769AF6DEF31" box="[968,1114,1638,1661]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="78" pageNumber="79">Figs. 60 A–C</figureCitation>
): palpal tibia 0.38 times cymbial length, RTA straight, with pointed apex (
<figureCitation id="EBB8FEA0180AFF9444273789A8E4EFD1" box="[872,979,1670,1693]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="78" pageNumber="79">Fig. 60 C</figureCitation>
), RTA length 0.47 tibia, length 1.78 times width, VTA slender, conical, elongate (
<figureCitation id="EBB8FEA0180AFF94446637A9A8F6EFF1" box="[809,961,1702,1725]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="78" pageNumber="79">Figs. 60 B, C</figureCitation>
), length 2.00 times width, length 0.44 tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.84 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.16 times tegulum length; TA3 J-shaped (
<figureCitation id="EBB8FEA0180BFF95455D3013A94BE87F" box="[530,636,284,307]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="79" pageNumber="80">Fig. 60 A</figureCitation>
), apicad of TA3 to TA2 by 1.125 tegulum length, TA2 a large triangular blade, behind TA3 apex; MA very small, simple (
<figureCitation id="EBB8FEA0180BFF9544C13033AF13E81F" box="[910,1060,316,339]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="79" pageNumber="80">Figs. 60 A, C</figureCitation>
), origin at 0.44 tegulum length, length (including apical processes) 0.73 times width, length 0.19 tegulum length, width 0.22 tegulum width; conductor fan entire (
<figureCitation id="EBB8FEA0180BFF9544673072A894E8D8" box="[808,931,381,404]" captionStart="FIGURE" captionStartId="151.[155,167,1613,1632]" captionTargetBox="[129,1163,996,1598]" captionTargetId="figure-73@151.[120,1176,972,1604]" captionTargetPageId="151" captionText="FIGURE 60. Uduba salegy, new species, Holotype (CASENT9006044) from Parc National Montagne d’Ambre, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13158709" httpUri="https://zenodo.org/record/13158709/files/figure.png" pageId="79" pageNumber="80">Fig. 60 C</figureCitation>
).
<emphasis id="41F73E37180BFF95448C3072AF20E8D8" bold="true" box="[963,1047,381,404]" pageId="79" pageNumber="80">Female</emphasis>
(
<typeStatus id="AC385C87180BFF9543603072AFA5E8D8" box="[1071,1170,381,404]" pageId="79" pageNumber="80" type="paratype">Paratype</typeStatus>
, CASENT9006045): Total length 15.00. Markings as in male and as in
<figureCitation id="EBB8FEA0180BFF9544C33092A8DCE8F8" box="[908,1003,413,436]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="79" pageNumber="80">Fig. 7 E</figureCitation>
. Carapace 7.57 long, 5.00 wide, 2.64 high; clypeus 0.43 high. Eye diameters: AME 0.21, ALE 0.25, PME 0.27, PLE 0.30. Chelicerae 3.50 long; sternum 3.39 long, 2.50 wide; labium 1.71 long; palpal coxae 2.57 long. Divided cribellum. Spination: palpus–femur d0-0-1, tibia p1-0, tarsus p2-0, v0-0-1; leg I– femur d1-0-1, p0-0-1, tibia v2-2-2-2, metatarsus p0-0-1, v1-2-2; leg II–femur d1-0-0, p0-0-1, tibia v1-1-2, metatarsus p0-0-1, v2-2-2; leg III–femur d0-1-0-1, p0-0-1-1, tibia d1-0-1-0, p0-1-1-0, v0- 1-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1-0-0, tibia v1-1-2, r0-1-1-0, metatarsus p0-0-1-1, v1-2-2, r0-2-1. Scopulae: strong ventral beneath all tarsi and beneath metatarsi I and II and apices of metatarsi III and IV. Leg measurements: I: 5.00 + 2.50 + 4.43 + 3.79 + 2.36 = 18.07; II: 4.50 + 2.29 + 3.21 + 3.00 + 2.14 = 15.14; III: 3.79 + 2.14 + 1.86 + 2.64 + 1.64 = 12.07; IV: 5.14 + 2.29 + 4.07 + 4.64 + 2.36 = 18.50; palpus: 2.29 + 1.29 + 1.36 + NA + 2.64 = 7.57. Leg formula 4123.
<emphasis id="41F73E37180BFF95465733F0AAEBEA5A" bold="true" box="[280,476,767,790]" pageId="79" pageNumber="80">Female genitalia:</emphasis>
epigynal plate with median lobe surrounded by depressed atrium, plate width 1.18 times length (
<figureCitation id="EBB8FEA0180BFF954684322FA904EA7B" box="[459,563,800,823]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="79" pageNumber="80">Fig. 74 A</figureCitation>
<emphasis id="41F73E37180BFF95457C322FA90BEA7B" bold="true" box="[563,572,800,823]" pageId="79" pageNumber="80">)</emphasis>
; atrium sides deeply concave, atrium wide, width 2.27 times atrium length, atrium with deep transverse concavity along posterior margin (
<figureCitation id="EBB8FEA0180BFF954354324FAFBEEA1B" box="[1051,1161,832,855]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="79" pageNumber="80">Fig. 74 A</figureCitation>
); epigynum length 2.50 times atrium length, atrium width 0.77 times epigynum width; atrium width at side of ML 1.42 times ML width; atrium and ML end well anteriad of epigastric groove, atrium origin at 0.24 of epigynal plate length, median lobe narrow, slightly wider at base, median lobe length 2.08 times width, ML extends well anteriad of atrium apex, ML length extends 0.18 times epigynum length anteriad of atrium, ML length 0.45 times epigynal length; LL narrow, atrium width 7.15 times LL width; copulatory openings hidden beneath lateral margins of atrium, distance between CO 0.61 width of epigynal plate, CO distance from epigastric furrow 0.36 times epigynum length. Vulva (
<figureCitation id="EBB8FEA0180BFF954604354DAA98ED15" box="[331,431,1090,1113]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="79" pageNumber="80">Fig. 74 B</figureCitation>
) with spermathecal ducts making four broad, transverse curves, vulva length 0.82 times width, fertilization ducts close-together, vulva width 11.25 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE225180BFF8A47E435ADAA87E85F" blockId="79.[123,1174,220,1725]" lastBlockId="80.[123,1174,220,821]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="41F73E37180BFF9547E435ADAA6BEDF5" bold="true" box="[171,348,1186,1209]" pageId="79" pageNumber="80">
Variation.
<emphasis id="41F73E37180BFF95466B35ADAA6BEDF5" bold="true" box="[292,348,1186,1209]" italics="true" pageId="79" pageNumber="80">Male</emphasis>
</emphasis>
(N = 4): Total length = 10.00–12.57, carapace length / width = 1.37–1.48, carapace height / width = 0.44–0.55, PER / carapace width = 0.62–0.82, PER/OAL = 2.42–2.62, PER/AER = 1.32–1.38, OAL/OQL = 1.07–1.15, OQP/ OQA = 1.18–1.29, clypeus height / AME = 0.91–1.27, cheliceral length / clypeus height = 9.14–14.80, sternum length / width = 1.30–1.40, palpal coxa length / width = 1.93–2.45, femur I length / carapace width = 1.20–1.23, metatarsus I length / carapace width = 1.23–1.30, femur IV length / carapace width = 1.19–1.25, cymbium length / carapace width = 0.55–0.63, cymbium length / palpal patella length = 2.67–2.75, cymbium length / palpal tibia length = 2.92–3.07, cymbium length / palpal femur length = 1.06–1.21, palpal tibia length / palpal patella length 0.88–0.92.
<emphasis id="41F73E37180BFF9545CF34ABA9E4ECF7" bold="true" box="[640,723,1444,1467]" italics="true" pageId="79" pageNumber="80">Female</emphasis>
(N = 5): Total length = 11.00–17.14, carapace length / width = 1.43–1.53, carapace height / width = 0.47–0.62, PER / carapace width = 0.67– 0.89, PER/OAL = 3.06–3.39, PER/AER = 1.33–1.45, OAL/OQL = 1.10–1.19, OQP/ OQA = 1.03– 1.25, clypeus height / AME = 1.23–2.00, cheliceral length / clypeus height = 8.17–10.25, sternum length / width = 1.29–1.36, palpal coxa length / width = 1.79–2.40, femur I length / carapace width = 0.92–1.05, metatarsus I length / carapace width = 0.68–0.79, femur IV length / carapace width = 0.96–1.11, palpal tarsus length / carapace width = 0.41–0.53, palpal tarsus length / palpal tibia length = 1.50–2.06, palpal tarsus length / palpal tibia length = 1.50–1.95, palpal tarsus length / palpal femur length = 0.82–1.16, palpal tibia length / palpal patella length = 0.94–1.06. Epigynum with median lobe length 1.80–3.33 times width; distance from atrium base to epigastric fold 0.19– 0.26 times epigynum length.
</paragraph>
<paragraph id="733CE2251814FF8A47E43013AF1AEBD8" blockId="80.[123,1174,220,821]" pageId="80" pageNumber="81">
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M
<emphasis id="41F73E371814FF8A478D3013AAA6E87F" bold="true" box="[194,401,284,307]" pageId="80" pageNumber="81">aterial examined.</emphasis>
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:
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</emphasis>
:
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, 2.79 air km NE of park entrance, 12°32ʹS, 49°10ʹE, elev. approx.
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,
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,
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, S.
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and J.
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(
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,
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), (
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,
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), (
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<collectingMethod id="AAC29A321814FF8A457D3093A999E8FF" box="[562,686,412,435]" pageId="80" pageNumber="81">pitfall traps</collectingMethod>
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;
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[VR-MA-1992- 01 (
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,
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,
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,
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), (
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,
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, 1jj,
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), [tracheal preparation of juvenile] (
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,
<specimenCode id="23254A5E1814FF8A457330F2A97EEB58" box="[572,585,509,532]" collectionCode="CASENT" pageId="80" pageNumber="81">2</specimenCode>
jj,
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)
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;
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235°SW Joffreville
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, montane rainforest, general collecting day,
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<emphasis id="41F73E371814FF8A45C23332A9FDEB18" box="[653,714,573,596]" italics="true" pageId="80" pageNumber="81">et al.,</emphasis>
[JJR0112] (
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montane rainforest, ex ground level,
<date id="073DC4E51814FF8A47F23372AAACEBD8" box="[189,411,637,660]" pageId="80" pageNumber="81" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">
<collectingDate id="17793D0D1814FF8A47F23372AAACEBD8" box="[189,411,637,660]" pageId="80" pageNumber="81" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">20–26 January 2001</collectingDate>
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,
<collectorName id="DE7687F31814FF8A46E83372AAE2EBD8" box="[423,469,637,660]" pageId="80" pageNumber="81">Lisa</collectorName>
J.
<collectorName id="DE7687F31814FF8A46B93372A976EBD8" box="[502,577,637,660]" pageId="80" pageNumber="81">Boutin</collectorName>
, [LJB0001.2] (
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.
</paragraph>
<paragraph id="733CE2251814FF8A47E43392ABE0EBB8" blockId="80.[123,1174,220,821]" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E43392AA57EBF8" bold="true" box="[171,352,669,692]" pageId="80" pageNumber="81">Natural history.</emphasis>
This species,
<emphasis id="41F73E371814FF8A46B33392A9AEEBF8" bold="true" box="[508,665,669,692]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="B48399A61814FF8A46B33392A9A5EBF8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[508,658,669,692]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="salegy">Uduba salegy</taxonomicName>
,
</emphasis>
is only known from montane rainforest where individuals were collected in pitfall traps and under objects. We know nothing of its use of cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8A47E433F2AACFEA79" pageId="80" pageNumber="81" type="distribution">
<paragraph id="733CE2251814FF8A47E433F2AACFEA79" blockId="80.[123,1174,220,821]" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E433F2AAE8EA58" bold="true" box="[171,479,765,788]" pageId="80" pageNumber="81">
Distribution.
<taxonomicName id="B48399A61814FF8A460833F2AAE8EA58" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[327,479,765,788]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="salegy">
<emphasis id="41F73E371814FF8A460833F2AAE8EA58" bold="true" box="[327,479,765,788]" italics="true" pageId="80" pageNumber="81">Uduba salegy</emphasis>
</taxonomicName>
</emphasis>
are known only from rainforests at Montagne d’Ambre in far northern
<collectingCountry id="0B94A2B51814FF8A47923211AA69EA79" box="[221,350,798,821]" name="Madagascar" pageId="80" pageNumber="81">Madagascar</collectingCountry>
(Maps 2, 13).
</paragraph>
</subSubSection>
</treatment>
</document>

303
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<mods:title id="094167D243A205B28AFE53C47C61C291">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="42B7622651E5938439A4C9A86C44DEAC">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="44632A50AE6B324352E0D789A0E4BB30">Ubick, Darrell</mods:namePart>
<mods:affiliation id="5D1C625D86B12C4621B387E398DDD292">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="21A52747C529BD1ADB6E6DE101C67B4E">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="E923D6601ACF109F03EF7FD193585D98">Polotow, Daniele</mods:namePart>
<mods:affiliation id="B68BC413958D8E38325B23EDC3FDCEF3">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<emphasis id="41F73E37180CFF92473431D3AA3AE9BF" bold="true" box="[123,269,220,243]" italics="true" pageId="72" pageNumber="73">Uduba rajery</emphasis>
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,
<taxonomicNameLabel id="5AC4834C180CFF92465431D3AA97E9BF" box="[283,416,220,243]" pageId="72" pageNumber="73" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="EBB8FEA0180CFF92473431F3ABDAE85D" box="[123,237,252,273]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="72" pageNumber="73">Figures 8 E</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF9247B831F3AA73E85D" box="[247,324,252,273]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">55 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF92460131F3AAA0E85D" box="[334,407,252,273]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">56 A–F</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF9246EE31F3AADCE85D" box="[417,491,252,273]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="72" pageNumber="73">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF9246B931F3A926E85D" box="[502,529,252,273]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="72" pageNumber="73">19</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180CFF9247E4302EA889E834" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="733CE225180CFF9247E4302EA889E834" blockId="72.[122,1174,289,1725]" pageId="72" pageNumber="73">
<emphasis id="41F73E37180CFF9247E4302EAA78E874" bold="true" box="[171,335,289,312]" pageId="72" pageNumber="73">Type material.</emphasis>
<materialsCitation id="C3EBE878180CFF924619302EA889E834" collectingDate="2005-11-21" collectingDateMax="2005-11-24" collectingDateMin="2005-11-21" collectionCode="CAS" collectorName="Brian L. Fisher" country="Madagascar" elevation="20" location="Foret Ambohidena" municipality="Ile Sainte Marie" pageId="72" pageNumber="73" specimenCode="CASENT9026393" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C87180CFF924619302EAA8EE874" box="[342,441,289,312]" pageId="72" pageNumber="73" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E180CFF92454B302EA9FAE874" box="[516,717,289,312]" collectionCode="CASENT" pageId="72" pageNumber="73">CASENT9026393</specimenCode>
) from a
<collectingMethod id="AAC29A32180CFF924465302EA8E6E874" box="[810,977,289,312]" pageId="72" pageNumber="73">yellow pan trap</collectingMethod>
in littoral rainforest at
<quantity id="B47B4FC0180CFF92478E304EABC7E814" box="[193,240,321,344]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="72" pageNumber="73" unit="m" value="20.0">
<elevation id="F8AE0516180CFF92478E304EABC7E814" box="[193,240,321,344]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="72" pageNumber="73" unit="m" value="20.0">20m</elevation>
</quantity>
elevation in
<location id="765CB4FE180CFF9246C9304EA964E814" LSID="urn:lsid:plazi:treatment:FB2A5333180CFF93473431D3A96AEB38:765CB4FE180CFF9246C9304EA964E814" box="[390,595,321,344]" country="Madagascar" municipality="Ile Sainte Marie" name="Foret Ambohidena" pageId="72" pageNumber="73" stateProvince="Toamasina">Forêt Ambohidena</location>
,
<collectingMunicipality id="9358785F180CFF92452E304EA821E814" box="[609,790,321,344]" pageId="72" pageNumber="73">Ile Sainte Marie</collectingMunicipality>
,
<collectingRegion id="B1472CC7180CFF92446B304EAF35E814" box="[804,1026,321,344]" country="Madagascar" name="Toamasina" pageId="72" pageNumber="73">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B5180CFF92435F304EAFB8E814" box="[1040,1167,321,344]" name="Madagascar" pageId="72" pageNumber="73">Madagascar</collectingCountry>
, collected
<date id="073DC4E5180CFF9247AC306EAAECE834" box="[227,475,353,376]" pageId="72" pageNumber="73" value="2005-11-21" valueMax="2005-11-24" valueMin="2005-11-21">
<collectingDate id="17793D0D180CFF9247AC306EAAECE834" box="[227,475,353,376]" pageId="72" pageNumber="73" value="2005-11-21" valueMax="2005-11-24" valueMin="2005-11-21">21–24 November 2005</collectingDate>
</date>
by
<collectorName id="DE7687F3180CFF924549306EA99CE834" box="[518,683,353,376]" pageId="72" pageNumber="73">Brian L. Fisher</collectorName>
<emphasis id="41F73E37180CFF9245FC306EA9DDE834" box="[691,746,353,376]" italics="true" pageId="72" pageNumber="73">et al.</emphasis>
, deposited in
<collectionCode id="15927AE0180CFF9244CC306EA88DE834" box="[899,954,353,376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="72" pageNumber="73" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180CFF9247E4308EA9D0EB55" pageId="72" pageNumber="73" type="etymology">
<paragraph id="733CE225180CFF9247E4308EA9D0EB55" blockId="72.[122,1174,289,1725]" pageId="72" pageNumber="73">
<emphasis id="41F73E37180CFF9247E4308EAA1DE8D4" bold="true" box="[171,298,385,408]" pageId="72" pageNumber="73">Etymology.</emphasis>
The species epithet is patronym is honor of the Malagasy musician Rajery, born Germain Randrianrisoa, who is a player of the bamboo tube zither (valiha) and who founded the modern valiha orchestra. In spite of a childhood accident that cost him the fingers of one hand he developed a unique and memorable style of playing the valiha, invented notation for this instrument, and is one of Madagascar’s finest recording artists.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180CFF9247E4332DA9DBEB96" pageId="72" pageNumber="73" type="diagnosis">
<paragraph id="733CE225180CFF9247E4332DA9DBEB96" blockId="72.[122,1174,289,1725]" pageId="72" pageNumber="73">
<emphasis id="41F73E37180CFF9247E4332DAA8EEB75" bold="true" box="[171,441,546,569]" pageId="72" pageNumber="73">
Diagnosis.
<taxonomicName id="B48399A6180CFF924668332DAA8EEB75" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,441,546,569]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180CFF924668332DAA8EEB75" bold="true" box="[295,441,546,569]" italics="true" pageId="72" pageNumber="73">Uduba rajery</emphasis>
</taxonomicName>
</emphasis>
belong to Group V, the
<taxonomicName id="B48399A6180CFF9245F5332DA870EB75" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[698,839,546,569]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rajery">Uduba rajery</taxonomicName>
group (
<figureCitation id="EBB8FEA0180CFF9244D4332DA8C5EB75" box="[923,1010,546,569]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="72" pageNumber="73">Map 19</figureCitation>
). Males can be distinguished from those of other
<taxonomicName id="B48399A6180CFF9246AF334DA91DEB15" box="[480,554,578,601]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180CFF9246AF334DA91DEB15" bold="true" box="[480,554,578,601]" italics="true" pageId="72" pageNumber="73">Uduba</emphasis>
</taxonomicName>
by the form of the male palp (
<figureCitation id="EBB8FEA0180CFF924425334DA8CEEB15" box="[874,1017,578,601]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">Figs. 55 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF92434C334DAF66EB15" box="[1027,1105,578,601]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">56 A–F</figureCitation>
<emphasis id="41F73E37180CFF92431F334DAF6EEB15" bold="true" box="[1104,1113,578,601]" pageId="72" pageNumber="73">)</emphasis>
, with a thick, swollen VTA on the tibia, the spine-like TA3 extending through a notch at the apex of the tegulum, and the large, triangular MA:
<taxonomicName id="B48399A6180CFF92456F338DA985EBD5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[544,690,642,665]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180CFF92456F338DA985EBD5" bold="true" box="[544,690,642,665]" italics="true" pageId="72" pageNumber="73">Uduba rajery</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A6180CFF924490338DAF43EBD5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[991,1140,642,665]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180CFF924490338DAF43EBD5" bold="true" box="[991,1140,642,665]" italics="true" pageId="72" pageNumber="73">U. rakotozafy</emphasis>
</taxonomicName>
by having the MA apex with two spikes (
<figureCitation id="EBB8FEA0180CFF92454233ADA949EBF5" box="[525,638,674,697]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">Figs. 55 C</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF9245C733ADA9CDEBF5" box="[648,762,674,697]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">56 B, D, E</figureCitation>
) (
<taxonomicName id="B48399A6180CFF92445E33ADA892EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[785,933,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180CFF92445E33ADA892EBF5" bold="true" box="[785,933,674,697]" italics="true" pageId="72" pageNumber="73">U. rakotozafy</emphasis>
</taxonomicName>
has an apical lobe and spike,
<figureCitation id="EBB8FEA0180CFF92478D33CCAA1CEB96" box="[194,299,707,730]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="72" pageNumber="73">Fig. 58 A</figureCitation>
). Females of
<taxonomicName id="B48399A6180CFF9246F233CCA967EB96" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[445,592,707,730]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180CFF9246F233CCA967EB96" bold="true" box="[445,592,707,730]" italics="true" pageId="72" pageNumber="73">Uduba rajery</emphasis>
</taxonomicName>
are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180CFF9347E433ECAFA7E87F" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="description">
<paragraph id="733CE225180CFF9347E433ECA86AE85F" blockId="72.[122,1174,289,1725]" lastBlockId="73.[123,1174,220,628]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<emphasis id="41F73E37180CFF9247E433ECAA01EBB6" bold="true" box="[171,310,739,762]" pageId="72" pageNumber="73">Description.</emphasis>
Male (
<typeStatus id="AC385C87180CFF9246CC33ECAAD0EBB6" box="[387,487,739,762]" pageId="72" pageNumber="73" type="holotype">holotype</typeStatus>
): Total length 6.00. Markings as in
<figureCitation id="EBB8FEA0180CFF92442C33ECA88AEBB6" box="[867,957,739,762]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="72" pageNumber="73">Fig. 8 E</figureCitation>
, with a striking pattern of black legs, tan coxae and trochanters, and black lateral stripes on a tan carapace. Carapace 3.80 long, 2.60 wide, 1.60 high; clypeus 0.25 high. Eye diameters:AME, PME and PLE 0.16, ALE 0.15. Chelicerae 1.70 long; sternum 2.00 long, 1.40 wide; labium 0.90 long; palpal coxae 1.30 long.
<emphasis id="41F73E37180CFF924734326CABE7EA36" bold="true" box="[123,208,867,890]" pageId="72" pageNumber="73">Ratios–</emphasis>
carapace length / width = 1.46, carapace height / width = 0.62, PER / carapace width = 0.39, PER / OAL = 2.83, PER / AER = 1.28, OAL / OQL = 1.06, OQP / OQA = 1.11, clypeus height / AME = 1.56, cheliceral length / clypeus height = 6.80, sternum length / width = 1.43, palpal coxa length / width = 2.60, femur I length / carapace width = 1.23, metatarsus I length / carapace width = 1.27, femur IV length / carapace width = 1.27, cymbium length / carapace width = 0.62, cymbium length / palpal patella length = 3.20, cymbium length / palpal tibia length = 2.29, cymbium length / palpal femur length = 1.23, palpal tibia length / palpal patella length = 1.40. Cribellum divided. Spination: palpus–femur d1-0-1, patella d0-1; leg I–femur d1-1-1, p0-0-2, tibia p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1, v2-2-2, r1-1-1; leg II–femur d1-1-1, p0-0-1, r0-0-1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-1, v2-2-3, r1-1-1; leg III–femur d1-1- 1, p0-1-1-0, r0-1-1-0, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2- 2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-1, p0-0-1, r0-0-1, patella r1, tibia d0-1-0, p0-1-1-0, v2- 2-3, r0-1-1-0, metatarsus d0-1-0, p1-1-1, v2-2-2, r1-1-1. Scopulae: cymbium, apicodorsal (
<figureCitation id="EBB8FEA0180CFF92437035EBAB87EC50" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">Figs. 56 A, C</figureCitation>
); very weak beneath all tarsi. Leg measurements: I: 3.20 + 1.40 + 3.10 + 3.30 + 2.20 = 13.20; II: 2.80 + 1.20 + 2.50 + 2.70 + 1.60 = 10.80; III: 2.40 + 1.10 + 1.60 + 2.20 + 1.20 = 8.50; IV: 3.30 + 1.20 + 2.80 + 3.60 + 1.70 = 12.60; palpus: 1.30 + 0.50 + 0.70 + NA + 1.60 = 4.10. Leg formula 1423.
<emphasis id="41F73E37180CFF924797346AAA7CEC30" bold="true" box="[216,331,1381,1404]" pageId="72" pageNumber="73">Male palp</emphasis>
: palpal tibia 0.52 times cymbial length, RTA broad, very short, sharply-curved to ventrad (
<figureCitation id="EBB8FEA0180CFF9247B0348AAA44ECD0" box="[255,371,1413,1436]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">Fig. 55 C</figureCitation>
), RTA length 0.09 tibia length, length 0.44 times width, VTA unique, large, swollen, transversely wider than long and truncate apically (
<figureCitation id="EBB8FEA0180CFF92441434AAA8C1ECF0" box="[859,1014,1445,1468]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">Figs. 55 B, C</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF92434B34AAAF0BECF0" box="[1028,1084,1445,1468]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">56 A</figureCitation>
), length 0.705 times width, length 0.44 tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.98 times width; TA1 serrate medially and with pointed apex, extends distad of tegulum apex, TA1 extends slightly past TA3 apex; TA3 elongate with recurved apex extending through notch at tegular anterior margin (
<figureCitation id="EBB8FEA0180CFF9245253729A9C9EF71" box="[618,766,1574,1597]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">Figs. 56 B, F</figureCitation>
), TA3 extends apicad of TA2 by 1.25 tegulum length, TA2 a mound partly forming the notch from which the TA3 apex emerges and partially covering mesal margin of MA (
<figureCitation id="EBB8FEA0180CFF92456E3769A9ABEF31" box="[545,668,1638,1661]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="72" pageNumber="73">Figs. 55 B</figureCitation>
,
<figureCitation id="EBB8FEA0180CFF9245E33769A83EEF31" box="[684,777,1638,1661]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="72" pageNumber="73">56 B, E</figureCitation>
); MA origin at 0.26 times tegulum length, MA apex extends distad of tegulum apex, length (including apical processes) 2.00 times width, MA very large, length 0.81 times tegulum length, width 0.395 times tegulum width, oval with a swollen, rounded base and with retrolateral and prolateral acutely-pointed apical projections (
<figureCitation id="EBB8FEA0180DFF9347F031F3AA1FE85F" box="[191,296,252,275]" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="73" pageNumber="74">Fig. 55 B</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF93467B31F3AABBE85F" box="[308,396,252,275]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="73" pageNumber="74">56 D, E</figureCitation>
); conductor fan entire.
<emphasis id="41F73E37180DFF9345C531F3A9EAE85F" bold="true" box="[650,733,252,275]" pageId="73" pageNumber="74">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="733CE225180DFF9347E43013AFA7E87F" blockId="73.[123,1174,220,628]" box="[171,1168,284,307]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E43013AA2DE87F" bold="true" box="[171,282,284,307]" pageId="73" pageNumber="74">Variation.</emphasis>
Unknown: the species
<taxonomicName id="B48399A6180DFF9345573013A99CE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[536,683,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180DFF9345573013A99CE87F" bold="true" box="[536,683,284,307]" italics="true" pageId="73" pageNumber="74">Uduba rajery</emphasis>
</taxonomicName>
is only known from
<specimenCount id="658529AC180DFF9344DE3013A8C1E87F" box="[913,1014,284,307]" count="1" pageId="73" pageNumber="74" type="male">one male</specimenCount>
, the
<typeStatus id="AC385C87180DFF9343643013AFBBE87F" box="[1067,1164,284,307]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180DFF9347E43033ABEBEB78" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="733CE225180DFF9347E43033AAF5E8FF" blockId="73.[123,1174,220,628]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E43033AABBE81F" bold="true" box="[171,396,316,339]" pageId="73" pageNumber="74">Material examined.</emphasis>
<materialsCitation id="C3EBE878180DFF9346DB3033AA88E8FF" collectingDate="2005-11-21" collectingDateMax="2005-11-24" collectingDateMin="2005-11-21" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="20" latitude="-16.824446" location="Foret Ambohidena" longLatPrecision="21" longitude="49.96417" municipality="Ile Sainte Marie" pageId="73" pageNumber="74" specimenCode="CASENT9026393" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<collectingCountry id="0B94A2B5180DFF9346DB3033A961E81F" box="[404,598,316,339]" name="Madagascar" pageId="73" pageNumber="74">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37180DFF93452D3033A80CE81F" box="[610,827,316,339]" italics="true" pageId="73" pageNumber="74">
<collectingRegion id="B1472CC7180DFF93452D3033A80CE81F" box="[610,827,316,339]" country="Madagascar" name="Toamasina" pageId="73" pageNumber="74">Toamasina Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F180DFF9344043033A8C9E81F" box="[843,1022,316,339]" pageId="73" pageNumber="74">Ile Sainte Marie</collectingMunicipality>
,
<location id="765CB4FE180DFF9343443033ABF2E83F" LSID="urn:lsid:plazi:treatment:FB2A5333180CFF93473431D3A96AEB38:765CB4FE180DFF9343443033ABF2E83F" country="Madagascar" latitude="-16.824446" longLatPrecision="21" longitude="49.96417" municipality="Ile Sainte Marie" name="Foret Ambohidena" pageId="73" pageNumber="74" stateProvince="Toamasina">Forêt Ambohidena</location>
,
<locationDeviation id="628EFB07180DFF93479A3053A903E83F" box="[213,564,348,371]" location="Ambodifotatra" pageId="73" pageNumber="74">
<quantity id="B47B4FC0180DFF93479A3053AA1AE83F" box="[213,301,348,371]" metricMagnitude="4" metricUnit="m" metricValue="2.28" pageId="73" pageNumber="74" unit="km" value="22.8">22.8 km</quantity>
44°NW Ambodifotatra
</locationDeviation>
,
<geoCoordinate id="16B784E2180DFF93450B3053A989E83F" box="[580,702,348,371]" degrees="16" direction="south" minutes="49" orientation="latitude" pageId="73" pageNumber="74" precision="15" seconds="28" value="-16.824446">16°49ʹ28ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E2180DFF93459F3053A879E83F" box="[720,846,348,371]" degrees="49" direction="east" minutes="57" orientation="longitude" pageId="73" pageNumber="74" precision="15" seconds="51" value="49.96417">49°57ʹ51ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC0180DFF9344D53053A8FBE83F" box="[922,972,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="73" pageNumber="74" unit="m" value="20.0">
<elevation id="F8AE0516180DFF9344D53053A8FBE83F" box="[922,972,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="73" pageNumber="74" unit="m" value="20.0">20m</elevation>
</quantity>
,
<collectingMethod id="AAC29A32180DFF9344923053AFA6E83F" box="[989,1169,348,371]" pageId="73" pageNumber="74">yellow pan trap</collectingMethod>
, littoral rainforest,
<date id="073DC4E5180DFF9346143073A959E8DF" box="[347,622,380,403]" pageId="73" pageNumber="74" value="2005-11-21" valueMax="2005-11-24" valueMin="2005-11-21">
<collectingDate id="17793D0D180DFF9346143073A959E8DF" box="[347,622,380,403]" pageId="73" pageNumber="74" value="2005-11-21" valueMax="2005-11-24" valueMin="2005-11-21">21–24 November 2005</collectingDate>
</date>
,
<collectorName id="DE7687F3180DFF9345C93073A829E8DF" box="[646,798,380,403]" pageId="73" pageNumber="74">B. L. Fisher</collectorName>
<emphasis id="41F73E37180DFF9344603073A85EE8DF" box="[815,873,380,403]" italics="true" pageId="73" pageNumber="74">et al</emphasis>
. [BLF12839] (
<typeStatus id="AC385C87180DFF93436A3073AFA5E8DF" box="[1061,1170,380,403]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E180DFF9347343093AA75E8FF" box="[123,322,412,435]" collectionCode="CASENT" pageId="73" pageNumber="74">CASENT9026393</specimenCode>
,
<specimenCount id="658529AC180DFF9346013093AA47E8FF" box="[334,368,412,435]" count="1" pageId="73" pageNumber="74" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180DFF9346313093AA81E8FF" box="[382,438,412,435]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="73" pageNumber="74" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225180DFF9347E430B3ABEBEB78" blockId="73.[123,1174,220,628]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E430B3AA68E89F" bold="true" box="[171,351,444,467]" pageId="73" pageNumber="74">Natural history.</emphasis>
The lone specimen of
<taxonomicName id="B48399A6180DFF93451B30B3A9D0E89F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[596,743,444,467]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180DFF93451B30B3A9D0E89F" bold="true" box="[596,743,444,467]" italics="true" pageId="73" pageNumber="74">Uduba rajery</emphasis>
</taxonomicName>
was collected in littoral rainforest. Like all adult male
<taxonomicName id="B48399A6180DFF93465630D3AA54E8BF" box="[281,355,476,499]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180DFF93465630D3AA54E8BF" bold="true" box="[281,355,476,499]" italics="true" pageId="73" pageNumber="74">Uduba</emphasis>
</taxonomicName>
it was wandering when trapped. The vestige of a divided cribellum on the male suggests that
<taxonomicName id="B48399A6180DFF93460330F2AA87EB58" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[332,432,509,532]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180DFF93460330F2AA87EB58" bold="true" box="[332,432,509,532]" italics="true" pageId="73" pageNumber="74">U. rajery</emphasis>
</taxonomicName>
females deploy cribellate sticky silk, but we know nothing of their behavior.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180DFF9347E43332A96AEB38" pageId="73" pageNumber="74" type="distribution">
<paragraph id="733CE225180DFF9347E43332A96AEB38" blockId="73.[123,1174,220,628]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E43332AAE0EB18" bold="true" box="[171,471,573,596]" pageId="73" pageNumber="74">
Distribution.
<taxonomicName id="B48399A6180DFF93460B3332AAE0EB18" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[324,471,573,596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180DFF93460B3332AAE0EB18" bold="true" box="[324,471,573,596]" italics="true" pageId="73" pageNumber="74">Uduba rajery</emphasis>
</taxonomicName>
</emphasis>
is known only from the
<typeStatus id="AC385C87180DFF9345AC3332A826EB18" box="[739,785,573,596]" pageId="73" pageNumber="74">type</typeStatus>
locality on Ile Sainte Marie, off the northeast coast of
<collectingCountry id="0B94A2B5180DFF93460E3352AAF5EB38" box="[321,450,605,628]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA0180DFF93469D3352A912EB38" box="[466,549,605,628]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="73" pageNumber="74">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF93457D3352A978EB38" box="[562,591,605,628]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="73" pageNumber="74">19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

334
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<document id="7AF7CBFA6860624217AD72CD61634B4F" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A5333180DFF9047343392AAA1EC5A" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba rakotofrah Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="EC2AD545-D95E-4F7E-B2F2-1996D02DB584" docUuidSource="ZooBank" docVersion="2" lastPageNumber="75" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="74" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="37C859EA1B95D751B0D689C18B807A6F">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="F513AC5719F12C8409D55BDA2FEFFDA6">Griswold, Charles</mods:namePart>
<mods:affiliation id="EDCDD9C2DAC0B7C7CEE75ACDF4B46965">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="FC0BB2F89AF89E0B1C7424786CB5A7A8">Ubick, Darrell</mods:namePart>
<mods:affiliation id="F68B4BE00DEDD7701BA55EA674C9AC2D">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="844C1B26577B0B2CE77DD585CDE5AA0E">Ledford, Joel</mods:namePart>
<mods:affiliation id="E8465A7C66F12DF398CE3930A0431832">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="9DDFD23E6D86228D8692C02012EE591A">Polotow, Daniele</mods:namePart>
<mods:affiliation id="5E902336332F3DF54CB895CE4A94FF86">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:date id="1FFC628C6503F9E1C6E6F19078C7060D">2022</mods:date>
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<taxonomicName id="B48399A6180DFF9347343392AA74EBF8" ID-CoL="C8KJX" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,323,669,692]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rakotofrah" status="sp. nov.">
<emphasis id="41F73E37180DFF9347343392AA74EBF8" bold="true" box="[123,323,669,692]" italics="true" pageId="73" pageNumber="74">Uduba rakotofrah</emphasis>
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,
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<figureCitation id="EBB8FEA0180DFF93473433B2ABCDEB9E" box="[123,250,701,722]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="73" pageNumber="74">Figures 29 B</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF93464B33B2AA66EB9E" box="[260,337,701,722]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="73" pageNumber="74">57 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF93461333B2AA91EB9E" box="[348,422,701,722]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="73" pageNumber="74">Maps 9</figureCitation>
, 16.
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<subSubSection id="3B99B1AE180DFF9347E433ECAA76EA16" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="733CE225180DFF9347E433ECAA76EA16" blockId="73.[122,1175,739,1725]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E433ECAA5BEBB6" bold="true" box="[171,364,739,762]" pageId="73" pageNumber="74">Type material.</emphasis>
<materialsCitation id="C3EBE878180DFF9346CB33ECAA76EA16" collectingDate="2015-02-21" collectingDateMax="2015-02-24" collectingDateMin="2015-02-21" collectionCode="CAS" country="Madagascar" county="Flavia Estevez" elevation="540" location="Anosyenne Mts." municipality="Brian Fisher" pageId="73" pageNumber="74" specimenCode="CASENT9062795, CASENT9062794" specimenCount="2" specimenCount-male="2" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C87180DFF9346CB33ECAAD8EBB6" box="[388,495,739,762]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
male (
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) and
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(
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), from a
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in rainforest at
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<elevation id="F8AE0516180DFF9345A6320CA811EA56" box="[745,806,771,794]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="73" pageNumber="74" unit="m" value="540.0">540m</elevation>
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elevation in the
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,
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,
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, collected
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<collectingDate id="17793D0D180DFF934564322CA827EA76" box="[555,784,803,826]" pageId="73" pageNumber="74" value="2015-02-21" valueMax="2015-02-24" valueMin="2015-02-21">21–24 February 2015</collectingDate>
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by
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and
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, deposited in
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.
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180DFF9347E4326CAA32ED17" pageId="73" pageNumber="74" type="etymology">
<paragraph id="733CE225180DFF9347E4326CAA32ED17" blockId="73.[122,1175,739,1725]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E4326CAA1DEA36" bold="true" box="[171,298,867,890]" pageId="73" pageNumber="74">Etymology.</emphasis>
The species epithet is patronym in honor of the Malagasy musician Rakotofrah, born Philibert Rabezoza. He was a renowned flautist and composer of music of the central highlands of
<collectingCountry id="0B94A2B5180DFF93479A32ACAA64EAF6" box="[213,339,931,954]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
. He became the most acclaimed 20
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century performer of the sodina flute, one of the oldest traditional instruments on the island and his picture with flute is even featured on Madagascar’s 1000 ariary banknote and 1500 FMG postage stamp. Through frequent international concerts and music festival performances, he promoted the music of the highlands of
<collectingCountry id="0B94A2B5180DFF934363350BABABED77" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
and became one of the most famous Malagasy artists, both within
<collectingCountry id="0B94A2B5180DFF934412352BA8E9ED77" box="[861,990,1060,1083]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
and on the world music scene.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180DFF9347E4356BABD2ECD0" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="733CE225180DFF9347E4356BABD2ECD0" blockId="73.[122,1175,739,1725]" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E4356BAAC6ED37" bold="true" box="[171,497,1124,1147]" pageId="73" pageNumber="74">
Diagnosis.
<taxonomicName id="B48399A6180DFF934668356BAAC6ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,497,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180DFF934668356BAAC6ED37" bold="true" box="[295,497,1124,1147]" italics="true" pageId="73" pageNumber="74">Uduba rakotofrah</emphasis>
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</emphasis>
belong to Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A6180DFF934734358BAA74EDD7" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,323,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). Males can be distinguished from those of other
<taxonomicName id="B48399A6180DFF934358358BAFA1EDD7" box="[1047,1174,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="41F73E37180DFF934358358BAF56EDD7" bold="true" box="[1047,1121,1156,1179]" italics="true" pageId="73" pageNumber="74">Uduba</emphasis>
spp.
</taxonomicName>
except
<taxonomicName id="B48399A6180DFF93478835ABAA87EDF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[199,432,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E37180DFF93478835ABAA87EDF7" bold="true" box="[199,432,1188,1211]" italics="true" pageId="73" pageNumber="74">U. pseudoevanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A6180DFF9346AB35ABA9B7EDF7" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[484,640,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E37180DFF9346AB35ABA9B7EDF7" bold="true" box="[484,640,1188,1211]" italics="true" pageId="73" pageNumber="74">U. evanescens</emphasis>
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in that the palp (
<figureCitation id="EBB8FEA0180DFF93447935ABA8F4EDF7" box="[822,963,1188,1211]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="73" pageNumber="74">Figs. 57 A–C</figureCitation>
<emphasis id="41F73E37180DFF93448C35ABA8FBEDF7" bold="true" box="[963,972,1188,1211]" pageId="73" pageNumber="74">)</emphasis>
has a notched MA and the pointed TA2 that extends far apicad, but differs from those in that TA2 extends apicad of all processes (
<figureCitation id="EBB8FEA0180DFF93465635EBAAB0EDB7" box="[281,391,1252,1275]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="73" pageNumber="74">Fig. 29 B</figureCitation>
) (in
<taxonomicName id="B48399A6180DFF93468C35EBA987EDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[451,688,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E37180DFF93468C35EBA987EDB7" bold="true" box="[451,688,1252,1275]" italics="true" pageId="73" pageNumber="74">U. pseudoevanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A6180DFF9345A235EBA8BBEDB7" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[749,908,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E37180DFF9345A235EBA8BBEDB7" bold="true" box="[749,908,1252,1275]" italics="true" pageId="73" pageNumber="74">U. evanescens</emphasis>
</taxonomicName>
the TA3 and conductor extend as far apical as TA2,
<figureCitation id="EBB8FEA0180DFF9346F8340AA91BEC50" box="[439,556,1285,1308]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="73" pageNumber="74">Figs. 29 A</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF934575340AA9A7EC50" box="[570,656,1285,1308]" captionStart="FIGURE" captionStartId="133.[155,167,892,911]" captionTargetBox="[138,1159,230,876]" captionTargetId="figure-16@133.[120,1176,216,877]" captionTargetPageId="133" captionText="FIGURE 35. Uduba evanescens (Dahl, 1901), (CASENT9006087) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158647" httpUri="https://zenodo.org/record/13158647/files/figure.png" pageId="73" pageNumber="74">35 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180DFF9345D1340AA9C2EC50" box="[670,757,1285,1308]" captionStart="FIGURE" captionStartId="147.[155,167,1615,1634]" captionTargetBox="[120,1176,960,1598]" captionTargetId="figure-52@147.[120,1176,960,1598]" captionTargetPageId="147" captionText="FIGURE 54. Uduba pseudoevanescens, new species, Paratype (CASENT9006006) from 57 km Route d’Anosibe de Moramanga, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158693" httpUri="https://zenodo.org/record/13158693/files/figure.png" pageId="73" pageNumber="74">54 A–C</figureCitation>
), TA3 is hidden (large and swollen in
<taxonomicName id="B48399A6180DFF934734342AAA53EC70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,356,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E37180DFF934734342AAA53EC70" bold="true" box="[123,356,1317,1340]" italics="true" pageId="73" pageNumber="74">U. pseudoevanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A6180DFF9346D7342AA903EC70" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[408,564,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E37180DFF9346D7342AA903EC70" bold="true" box="[408,564,1317,1340]" italics="true" pageId="73" pageNumber="74">U. evanescens</emphasis>
</taxonomicName>
) and the basal lobe of the MA is deeply forked (entire in
<taxonomicName id="B48399A6180DFF934734344AAA54EC10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,355,1349,1372]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E37180DFF934734344AAA54EC10" bold="true" box="[123,355,1349,1372]" italics="true" pageId="73" pageNumber="74">U. pseudoevanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A6180DFF9346D9344AA905EC10" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[406,562,1349,1372]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E37180DFF9346D9344AA905EC10" bold="true" box="[406,562,1349,1372]" italics="true" pageId="73" pageNumber="74">U. evanescens</emphasis>
</taxonomicName>
).
<taxonomicName id="B48399A6180DFF934507344AA838EC10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[584,783,1349,1372]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180DFF934507344AA838EC10" bold="true" box="[584,783,1349,1372]" italics="true" pageId="73" pageNumber="74">Uduba rakotofrah</emphasis>
</taxonomicName>
further differ from
<taxonomicName id="B48399A6180DFF934490344AAF4DEC10" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[991,1146,1349,1372]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E37180DFF934490344AAF4DEC10" bold="true" box="[991,1146,1349,1372]" italics="true" pageId="73" pageNumber="74">U. evanescens</emphasis>
</taxonomicName>
in having a divided cribellum; the latter is ecribellate (
<figureCitation id="EBB8FEA0180DFF9345E6346AA827EC30" box="[681,784,1381,1404]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="73" pageNumber="74">Fig. 20 D</figureCitation>
). Females of
<taxonomicName id="B48399A6180DFF9344EA346AAF59EC30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[933,1134,1381,1404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180DFF9344EA346AAF59EC30" bold="true" box="[933,1134,1381,1404]" italics="true" pageId="73" pageNumber="74">Uduba rakotofrah</emphasis>
</taxonomicName>
are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180DFF9047E434AAABF7EAB9" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="description">
<paragraph id="733CE225180DFF9047E434AAA815EBF8" blockId="73.[122,1175,739,1725]" lastBlockId="74.[123,1174,220,1302]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<emphasis id="41F73E37180DFF9347E434AAAA4FECF0" bold="true" box="[171,376,1445,1468]" pageId="73" pageNumber="74">Description. Male</emphasis>
(
<typeStatus id="AC385C87180DFF9346C834AAAADCECF0" box="[391,491,1445,1468]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
): Total length 9.00. Markings typical of
<taxonomicName id="B48399A6180DFF9344ED34AAA8DBECF0" box="[930,1004,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180DFF9344ED34AAA8DBECF0" bold="true" box="[930,1004,1445,1468]" italics="true" pageId="73" pageNumber="74">Uduba</emphasis>
</taxonomicName>
. Carapace 5.10 long, 3.30 wide, 1.90 high; clypeus 0.25 high. Eye diameters:AME and ALE 0.22, PME 0.20, PLE 0.24. Chelicerae 2.00 long; sternum 2.20 long, 1.80 wide; labium 0.90 long; palpal coxae 1.50 long. Divided cribellum. Spination (
<typeStatus id="AC385C87180DFF93468B370AA91FEF50" box="[452,552,1541,1564]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
): palpus–femur d1-1-1, p0-0-1, r0-0-1; leg I–femur d1-1- 1, p0-0-2, r0-1-0-1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus d1-1-0-0, p0-0-1-1, v2- 2-2, r1-1-0; leg II–femur d1-1-1, p0-1-1, r0-0-1-1, patella p1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus d1-1-0-0, p0-0-1-1, v2-2-2, r1-1-0-0; leg III–femur d1-1-0-1, p0-1-1-0, r0-1- 1-0, patella p1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-3, tibia d0-0-1-0, v2-1-2, r0-1-0-0, metatarsus d0-2-2, p1-1-2, v2-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; legs I–IV, leg scopulae very weak to absent. Leg measurements (
<typeStatus id="AC385C87180EFF9047CD31F3ABD1E85F" box="[130,230,252,275]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
): I: 4.00 + 1.70 + 3.70 + 3.70 + 2.20 = 15.30; II: 3.50 + 1.60 + 3.20 + 3.40 + 2.00 = 13.70; III: 3.10 + 1.30 + 2.10 + 2.90 + 1.50 = 10.90; IV: 4.00 + 1.50 + 3.50 + 4.30 + 2.00 = 15.30; palpus: 1.50 + 0.70 + 0.80 + NA + 2.20 = 5.20. Leg formula 4=1, 23.
<emphasis id="41F73E37180EFF9044EB3033AF2AE81F" bold="true" box="[932,1053,316,339]" pageId="74" pageNumber="75">Male palp</emphasis>
(
<typeStatus id="AC385C87180EFF90437E3033AFA6E81F" box="[1073,1169,316,339]" pageId="74" pageNumber="75" type="paratype">paratype</typeStatus>
, CASENT9064775) (
<figureCitation id="EBB8FEA0180EFF9046163053AAD8E83F" box="[345,495,348,371]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">Figs. 57 A–C</figureCitation>
): palpal tibia 0.36 times cymbial length, RTA small, triangular, with pointed apex (
<figureCitation id="EBB8FEA0180EFF9046053073AA85E8DF" box="[330,434,380,403]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">Fig. 57 A</figureCitation>
), length 0.19 tibia length, length equals width; VTA very long, cylindrical, upturned (
<figureCitation id="EBB8FEA0180EFF90467B3093AAF2E8FF" box="[308,453,412,435]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">Figs. 57 A, C</figureCitation>
), length 1.91 times width, length 0.81 times tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 1.02 times width; TA1 and TA3 extend to tegulum apex, TA2 extends beyond TA1 and tegulum apex (
<figureCitation id="EBB8FEA0180EFF90443830D3A8D5E8BF" box="[887,994,476,499]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">Fig. 57 B</figureCitation>
), TA3 a convex, blunt lobe, TA2 a large, sharply-pointed triangle, extending farthest apicad of all tegular processes (
<figureCitation id="EBB8FEA0180EFF9047CC3312ABDAEB78" box="[131,237,541,564]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="74" pageNumber="75">Fig. 29 B</figureCitation>
); MA complex, with proximal blunt lobe, median notch, and small apical curved lobe (
<figureCitation id="EBB8FEA0180EFF9047CC3332ABC2EB18" box="[131,245,573,596]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="74" pageNumber="75">Figs. 29 B</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF90464E3332AA01EB18" box="[257,310,573,596]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">57 B</figureCitation>
), origin at 0.15 times tegulum length, tegulum apex at 0.52 tegulum length from MA apex, length (including apical processes) 1.02 times width, MA length 0.30 times tegulum length, width 0.44 tegulum width; tegulum with apical mound that partially covers base of TA3; conductor fan margin irregular (
<figureCitation id="EBB8FEA0180EFF9046993392A977EBF8" box="[470,576,669,692]" captionStart="FIGURE" captionStartId="150.[155,167,854,873]" captionTargetBox="[125,1168,238,833]" captionTargetId="figure-11@150.[120,1176,216,851]" captionTargetPageId="150" captionText="FIGURE 57. Uduba rakotofrah, new species, paratype (CASENT9062794), from Anosyenne Mts., left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158701" httpUri="https://zenodo.org/record/13158701/files/figure.png" pageId="74" pageNumber="75">Fig. 57 C</figureCitation>
).
<emphasis id="41F73E37180EFF90451A3392A99FEBF8" bold="true" box="[597,680,669,692]" pageId="74" pageNumber="75">Female</emphasis>
: unknown.
</paragraph>
<paragraph id="733CE225180EFF9047E433B2ABF7EAB9" blockId="74.[123,1174,220,1302]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E433B2AA2DEB98" bold="true" box="[171,282,701,724]" pageId="74" pageNumber="75">Variation.</emphasis>
Male (N= 3): Total length 8.20–9.00; carapace length / width = 1.51–1.56, carapace height / width = 0.43–0.58, PER / carapace width = 0.41–0.43, PER / OAL = 2.57–2.65, PER / AER = 1.22–1.23, OAL / OQL = 1.08–1.18, OQP / OQA = 1.00–1.13, clypeus height / AME = 1.14– 1.36, cheliceral length / clypeus height = 6.67–8.00, sternum length / width = 1.22–1.41, palpal coxa length / width = 2.50–3.00, femur I length / carapace width = 1.14–1.25, metatarsus I length / carapace width 1.06–1.16, femur IV length / carapace width = 1.21–1.29, cymbium length / carapace width = 0.53–0.72, cymbium length / palpal patella length = 2.75–3.29, cymbium length / palpal tibia length = 2.75–3.29, cymbium length / palpal femur length = 1.16–1.47, palpal tibia length / palpal patella length = 1.00–1.14. Female variation is unknown: the species is only known from males.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9047E432F1AFA1ED9A" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="733CE225180EFF9047E432F1AAF5ED3A" blockId="74.[123,1174,220,1302]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E432F1AAA2ED59" bold="true" box="[171,405,1022,1045]" pageId="74" pageNumber="75">Material examined.</emphasis>
<materialsCitation id="C3EBE878180EFF9046EA32F1AA88ED3A" collectingDate="2015-02-21" collectingDateMax="2015-02-24" collectingDateMin="2015-02-21" collectionCode="CAS" collectorName="B. Fisher &amp; F. Estevez" country="Madagascar" elevation="540" latitude="-24.14" location="Anosyenne Mts." longLatPrecision="21" longitude="47.074165" pageId="74" pageNumber="75" specimenCode="CASENT9062795, CASENT9064775" specimenCount="3" specimenCount-male="3" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="0B94A2B5180EFF9046EA32F1A953ED59" box="[421,612,1022,1045]" name="Madagascar" pageId="74" pageNumber="75">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37180EFF90453032F1A80DED59" box="[639,826,1022,1045]" italics="true" pageId="74" pageNumber="75">
<collectingRegion id="B1472CC7180EFF90453032F1A80DED59" box="[639,826,1022,1045]" country="Madagascar" name="Toliara" pageId="74" pageNumber="75">Toliara Province</collectingRegion>
</emphasis>
:
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,
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,
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, elev.
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<elevation id="F8AE0516180EFF90467D3511AA46ED79" box="[306,369,1054,1077]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="74" pageNumber="75" unit="m" value="540.0">540m</elevation>
</quantity>
, rainforest,
<collectingMethod id="AAC29A32180EFF9046B93511A94AED79" box="[502,637,1054,1077]" pageId="74" pageNumber="75">malaise trap</collectingMethod>
,
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<collectingDate id="17793D0D180EFF9045C43511A84DED79" box="[651,890,1054,1077]" pageId="74" pageNumber="75" value="2015-02-21" valueMax="2015-02-24" valueMin="2015-02-21">21–24 February 2015</collectingDate>
</date>
,
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and
<collectorName id="DE7687F3180EFF9043683511AFA2ED79" box="[1063,1173,1054,1077]" pageId="74" pageNumber="75">F. Estevez</collectorName>
[BLF36109] (
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,
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,
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), (
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,
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,
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and
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,
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,
<collectionCode id="15927AE0180EFF9046313550AA81ED3A" box="[382,438,1119,1142]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="74" pageNumber="75" type="Museum">CAS</collectionCode>
)
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.
</paragraph>
<paragraph id="733CE225180EFF9047E43570AFA1ED9A" blockId="74.[123,1174,220,1302]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E43570AA57EDDA" bold="true" box="[171,352,1151,1174]" pageId="74" pageNumber="75">Natural history.</emphasis>
Collection records indicate that
<taxonomicName id="B48399A6180EFF90458A3570A8B8EDDA" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[709,911,1151,1174]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180EFF90458A3570A8B8EDDA" bold="true" box="[709,911,1151,1174]" italics="true" pageId="74" pageNumber="75">Uduba rakotofrah</emphasis>
</taxonomicName>
occurs in low elevation rain forest. Wandering males were collected above the forest floor in a malaise trap. The unknown females of
<taxonomicName id="B48399A6180EFF9047A035B0AAB1ED9A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[239,390,1215,1238]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180EFF9047A035B0AAB1ED9A" bold="true" box="[239,390,1215,1238]" italics="true" pageId="74" pageNumber="75">U. rakotofrah</emphasis>
</taxonomicName>
should have a divided cribellum, but we know nothing of their use of silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9047E435D0AAA1EC5A" pageId="74" pageNumber="75" type="distribution">
<paragraph id="733CE225180EFF9047E435D0AAA1EC5A" blockId="74.[123,1174,220,1302]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E435D0AA0AEDBA" bold="true" box="[171,317,1247,1270]" pageId="74" pageNumber="75">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6180EFF90468835D0A9B9EDBA" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[455,654,1247,1270]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotofrah">
<emphasis id="41F73E37180EFF90468835D0A9B9EDBA" bold="true" box="[455,654,1247,1270]" italics="true" pageId="74" pageNumber="75">Uduba rakotofrah</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C87180EFF9044DC35D0A8F6EDBA" box="[915,961,1247,1270]" pageId="74" pageNumber="75">type</typeStatus>
locality in southern
<collectingCountry id="0B94A2B5180EFF90473435F0ABCBEC5A" box="[123,252,1279,1302]" name="Madagascar" pageId="74" pageNumber="75">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA0180EFF90464435F0AA68EC5A" box="[267,351,1279,1302]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="74" pageNumber="75">Maps 9</figureCitation>
, 16).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="551FD0457EEEA8DE125D2D8109471009">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<emphasis id="41F73E37180EFF9047343430AAE5EC1A" bold="true" box="[123,466,1343,1366]" pageId="74" pageNumber="75">
<taxonomicName id="B48399A6180EFF9047343430AA77EC1A" ID-CoL="C8KJY" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,320,1343,1366]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotozafy" status="sp. nov.">
<emphasis id="41F73E37180EFF9047343430AA77EC1A" bold="true" box="[123,320,1343,1366]" italics="true" pageId="74" pageNumber="75">Uduba rakotozafy</emphasis>
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,
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9047343450A90DEC38" box="[123,570,1375,1396]" pageId="74" pageNumber="75" type="description">
<paragraph id="733CE225180EFF9047343450A90DEC38" blockId="74.[123,570,1343,1396]" box="[123,570,1375,1396]" pageId="74" pageNumber="75">
<figureCitation id="EBB8FEA0180EFF9047343450ABD8EC38" box="[123,239,1375,1396]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="74" pageNumber="75">Figures 8 G</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF9047B53450AA1EEC38" box="[250,297,1375,1396]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="74" pageNumber="75">11 E</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF90467C3450AA53EC38" box="[307,356,1375,1396]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="74" pageNumber="75">29 G</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF9046203450AA8BEC38" box="[367,444,1375,1396]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="74" pageNumber="75">58 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF9046883450A927EC38" box="[455,528,1375,1396]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="74" pageNumber="75">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA0180EFF9045543450A901EC38" box="[539,566,1375,1396]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="74" pageNumber="75">19</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9047E4348AAAC1EC90" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="733CE225180EFF9047E4348AAAC1EC90" blockId="74.[123,1174,1413,1725]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E4348AAA65ECD0" bold="true" box="[171,338,1413,1436]" pageId="74" pageNumber="75">Type material.</emphasis>
<materialsCitation id="C3EBE878180EFF904614348AAAC1EC90" collectingDate="1993-11-04" collectingDateMax="1993-11-05" collectingDateMin="1993-11-04" collectionCode="CAS" collectorName="C. Griswold" country="Madagascar" elevation="1000" location="Parc National Perinet" municipality="Andasibe" pageId="74" pageNumber="75" specimenCode="CASENT9006024" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C87180EFF904614348AAA89ECD0" box="[347,446,1413,1436]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E180EFF904542348AA9E1ECD0" box="[525,726,1413,1436]" collectionCode="CASENT" pageId="74" pageNumber="75">CASENT9006024</specimenCode>
) from forest at
<quantity id="B47B4FC0180EFF9044C6348AA8E4ECD0" box="[905,979,1413,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="m" value="1000.0">
<elevation id="F8AE0516180EFF9044C6348AA8E4ECD0" box="[905,979,1413,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="m" value="1000.0">1000m</elevation>
</quantity>
elevation at
<location id="765CB4FE180EFF904329348AAA1AECF0" LSID="urn:lsid:plazi:treatment:FB2A5333180EFF9147343430A8E1EF10:765CB4FE180EFF904329348AAA1AECF0" country="Madagascar" municipality="Andasibe" name="Parc National Perinet" pageId="74" pageNumber="75" stateProvince="Toamasina">Parc National Périnet</location>
, near
<collectingMunicipality id="9358785F180EFF90462334AAAAE4ECF0" box="[364,467,1445,1468]" pageId="74" pageNumber="75">Andasibe</collectingMunicipality>
,
<collectingRegion id="B1472CC7180EFF90469034AAA98DECF0" box="[479,698,1445,1468]" country="Madagascar" name="Toamasina" pageId="74" pageNumber="75">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B5180EFF90458934AAA872ECF0" box="[710,837,1445,1468]" name="Madagascar" pageId="74" pageNumber="75">Madagascar</collectingCountry>
, collected
<date id="073DC4E5180EFF9044F334AAAFA2ECF0" box="[956,1173,1445,1468]" pageId="74" pageNumber="75" value="1993-11-04" valueMax="1993-11-05" valueMin="1993-11-04">
<collectingDate id="17793D0D180EFF9044F334AAAFA2ECF0" box="[956,1173,1445,1468]" pageId="74" pageNumber="75" value="1993-11-04" valueMax="1993-11-05" valueMin="1993-11-04">4-5 November 1993</collectingDate>
</date>
by
<collectorName id="DE7687F3180EFF9047D234CAAA13EC90" box="[157,292,1477,1500]" pageId="74" pageNumber="75">C. Griswold</collectorName>
, deposited in
<collectionCode id="15927AE0180EFF9046F434CAAAC5EC90" box="[443,498,1477,1500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="74" pageNumber="75" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9047E434EAAAE2EF71" pageId="74" pageNumber="75" type="etymology">
<paragraph id="733CE225180EFF9047E434EAAAE2EF71" blockId="74.[123,1174,1413,1725]" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E434EAAA1DECB0" bold="true" box="[171,298,1509,1532]" pageId="74" pageNumber="75">Etymology.</emphasis>
The species epithet is a patronym in honor of the Malagasy musician Rakotozafy, a legendary player of his own customized
<emphasis id="41F73E37180EFF904507370AA986EF50" box="[584,689,1541,1564]" italics="true" pageId="74" pageNumber="75">marovany</emphasis>
zither and one of the most celebrated
<emphasis id="41F73E37180EFF90431C370AAFA1EF50" box="[1107,1174,1541,1564]" italics="true" pageId="74" pageNumber="75">valiha</emphasis>
players of the twentieth century.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180EFF9147E43749AAEDE85F" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="733CE225180EFF9147E43749AAEDE85F" blockId="74.[123,1174,1413,1725]" lastBlockId="75.[122,1174,220,1628]" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
<emphasis id="41F73E37180EFF9047E43749AADDEF11" bold="true" box="[171,490,1606,1629]" pageId="74" pageNumber="75">
Diagnosis.
<taxonomicName id="B48399A6180EFF90466A3749AADDEF11" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,490,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180EFF90466A3749AADDEF11" bold="true" box="[293,490,1606,1629]" italics="true" pageId="74" pageNumber="75">Uduba rakotozafy</emphasis>
</taxonomicName>
</emphasis>
belong to Group V, the
<taxonomicName id="B48399A6180EFF9045AC3749A859EF11" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[739,878,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rajery">Uduba rajery</taxonomicName>
group (
<figureCitation id="EBB8FEA0180EFF90448F3749AF21EF11" box="[960,1046,1606,1629]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="74" pageNumber="75">Map 19</figureCitation>
). Males can be distinguished from those of other
<taxonomicName id="B48399A6180EFF90455E3769A96CEF31" box="[529,603,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37180EFF90455E3769A96CEF31" bold="true" box="[529,603,1638,1661]" italics="true" pageId="74" pageNumber="75">Uduba</emphasis>
</taxonomicName>
by the form of the male palp, with a thick, swollen VTA on the tibia, the spine-like TA3 extending through a notch at the apex of the tegulum (
<figureCitation id="EBB8FEA0180EFF9043063789ABA6EFF1" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="74" pageNumber="75">Fig. 29 G</figureCitation>
), and the large, triangular MA (
<figureCitation id="EBB8FEA0180EFF9046BD37A9A9BEEFF1" box="[498,649,1702,1725]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="74" pageNumber="75">Figs. 58 A–C</figureCitation>
).
<taxonomicName id="B48399A6180EFF9045D137A9A851EFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[670,870,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180EFF9045D137A9A851EFF1" bold="true" box="[670,870,1702,1725]" italics="true" pageId="74" pageNumber="75">Uduba rakotozafy</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A6180FFF91473431D3ABEAE9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,221,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180FFF91473431D3ABEAE9BF" bold="true" box="[123,221,220,243]" italics="true" pageId="75" pageNumber="76">U. rajery</emphasis>
</taxonomicName>
by having the MA apex with an apical lobe and spike (
<taxonomicName id="B48399A6180FFF91446331D3A8B9E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[812,910,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rajery">
<emphasis id="41F73E37180FFF91446331D3A8B9E9BF" bold="true" box="[812,910,220,243]" italics="true" pageId="75" pageNumber="76">U. rajery</emphasis>
</taxonomicName>
has two spikes,
<figureCitation id="EBB8FEA0180FFF91437031D3ABB8E85F" captionStart="FIGURE" captionStartId="148.[155,167,825,844]" captionTargetBox="[120,1176,216,809]" captionTargetId="figure-11@148.[120,1176,216,809]" captionTargetPageId="148" captionText="FIGURE 55. Uduba rajery, new species, Holotype (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158695" httpUri="https://zenodo.org/record/13158695/files/figure.png" pageId="75" pageNumber="76">Figs. 55 B</figureCitation>
,
<figureCitation id="EBB8FEA0180FFF9147D431F3ABE5E85F" box="[155,210,252,275]" captionStart="FIGURE" captionStartId="149.[155,167,1218,1237]" captionTargetBox="[121,1176,216,1203]" captionTargetId="figure-16@149.[120,1176,216,1203]" captionTargetPageId="149" captionText="FIGURE 56. Uduba rajery, new species, Holotype male (CASENT9026393) from Forêt Ambohidena, Ile Sainte Marie, right palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, proventral. F. Bulb, prolateral. C, conductor; CmS, cymbial scopula (arrows); E, embolus; Eb, embolic base; MA, median apophysis; ST, subtegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C, F = 100 µm, D, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158697" httpUri="https://zenodo.org/record/13158697/files/figure.png" pageId="75" pageNumber="76">56 E</figureCitation>
). Females are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180FFF9147E43013A83EEF50" pageId="75" pageNumber="76" type="description">
<paragraph id="733CE225180FFF9147E43013AF10EC56" blockId="75.[122,1174,220,1628]" pageId="75" pageNumber="76">
<emphasis id="41F73E37180FFF9147E43013AA01E87F" bold="true" box="[171,310,284,307]" pageId="75" pageNumber="76">Description.</emphasis>
Make (
<typeStatus id="AC385C87180FFF9146C43013AAC2E87F" box="[395,501,284,307]" pageId="75" pageNumber="76" type="holotype">Holotype</typeStatus>
): Total length 9.29. Markings as in
<figureCitation id="EBB8FEA0180FFF9144343013A8D3E87F" box="[891,996,284,307]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="75" pageNumber="76">Figs. 8 G</figureCitation>
.
<figureCitation id="EBB8FEA0180FFF9144BF3013AF11E87F" box="[1008,1062,284,307]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="75" pageNumber="76">11 E</figureCitation>
, Carapace 4.86 long, 3.71 wide, 1.57 high; clypeus 0.21 high. Eye diameters: AME 0.14, ALE 0.18, PME 0.20, PLE 0.21. Chelicerae 1.93 long; sternum 2.25 long, 1.61 wide; labium 1.04 long; palpal coxae 1.57 long.
<emphasis id="41F73E37180FFF9147A03072AA73E8D8" bold="true" box="[239,324,381,404]" pageId="75" pageNumber="76">Ratios–</emphasis>
carapace length / width = 1.31, carapace height / width = 0.42, PER / carapace width = 0.84, PER / OAL = 2.64, PER / AER = 1.42, OAL / OQL = 1.27, OQP / OQA = 1.33, clypeus height / AME = 1.50, cheliceral length / clypeus height = 9.00, sternum length / width = 1.40, palpal coxa length / width = 2.59, femur I length / carapace width = 1.10, metatarsus I length / carapace width = 1.10, femur IV length / carapace width = 1.15, cymbium length / carapace width = 0.58, cymbium length / palpal patella length = 3.00, cymbium length / palpal tibia length = 2.73, cymbium length / palpal femur length = 1.15, palpal tibia length / palpal patella length = 1.10. Cribellum divided. Spination: palpus–femur d0-1-1, p0-0-1, r0-0-1, patella p1-0, tibia p1-0; leg I– femur d1-0-1-1, p0-0-2, r0-0-1, tibia p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1- 2; leg II–femur d1-1-1, p0-1-1-0, r0-1-1-0, tibia d0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-1; leg III–femur d1-1-1, p1-1-2, r1-1-2, tibia d0-1-0-0, p0-1-1-0, v2-2-2, r0-1- 1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-1, p1-1-1, r0-0-1, tibia d1-0-1-0, p0-0-1-0, v2-2-2, r0-1-0-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements: I: 4.07 + 1.64 + 3.64 + 4.07 + 2.64 = 16.07; II: 3.57 + 1.57 + 2.93 + 3.21 + 2.07 = 13.36; III: 3.07 + 1.43 + 1.79 + 2.71 + 1.50 = 10.50; IV: 4.29 + 1.50 + 3.43 + 4.29 + 2.14 = 15.64; palpus: 1.86 + 0.71 + 0.79 + NA + 2.14 = 5.50. Leg formula 1423.
<emphasis id="41F73E37180FFF914655328FAABAEADB" bold="true" box="[282,397,896,919]" pageId="75" pageNumber="76">Male palp</emphasis>
(
<figureCitation id="EBB8FEA0180FFF9146D3328FA918EADB" box="[412,559,896,919]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">Figs. 58 A–C</figureCitation>
): palpal tibia 0.41 cymbial length, RTA broad, RTA length 0.24 tibia length, length equals width, with bluntly-pointed apex (
<figureCitation id="EBB8FEA0180FFF91440632AEA885EAF4" box="[841,946,929,952]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">Fig. 58 C</figureCitation>
), VTA unique, large, hammer-shaped, transversely wider than long and with apical recurved hook (
<figureCitation id="EBB8FEA0180FFF9144FD32CEAF76EA94" box="[946,1089,961,984]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">Figs. 58 A–C</figureCitation>
), length 0.80 times width, length 0.32 tibia width; tibia lacking a stout retrolateral spine but with slender prolateral spine (
<figureCitation id="EBB8FEA0180FFF914662350EAAA4ED54" box="[301,403,1025,1048]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">Fig. 58 B</figureCitation>
); tegulum convex, without ridge, tegulum length 0.875 times width; TA1 with subbasal notch and broad apex, extends distad of tegulum apex, TA1 extends past TA3 apex by 1.15 times tegulum length; TA3 elongate with recurved apex extending through notch at tegular anterior margin (
<figureCitation id="EBB8FEA0180FFF91461B356DAAFDED35" box="[340,458,1122,1145]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="75" pageNumber="76">Figs. 29 G</figureCitation>
,
<figureCitation id="EBB8FEA0180FFF914699356DA93CED35" box="[470,523,1122,1145]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">58 A</figureCitation>
), TA3 extends apicad of TA2 by 1.09 tegulum length, TA2 a small low mound partly forming the notch from which the TA3 apex emerges; MA origin at 0.17 times tegulum length, MA apex extends distad of tegulum apex, MA length (including apical processes) 1.03 times width, MA very large, covering much of tegulum, length 0.83 times tegulum length, width 0.71 times tegulum width, triangular with apical projections, retrolateral blunt and prolateral acutely-pointed (
<figureCitation id="EBB8FEA0180FFF9146D0340CA922EC56" box="[415,533,1283,1306]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="75" pageNumber="76">Figs. 29 G</figureCitation>
,
<figureCitation id="EBB8FEA0180FFF91456E340CA96FEC56" box="[545,600,1283,1306]" captionStart="FIGURE" captionStartId="150.[155,167,1593,1612]" captionTargetBox="[126,1164,1021,1567]" captionTargetId="figure-93@150.[120,1176,1008,1581]" captionTargetPageId="150" captionText="FIGURE 58. Uduba rakotozafy, new species, Holotype (CASENT9006024) from Parc National Périnet, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. RTA, retrolateral tibial apophysis on male palp; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158703" httpUri="https://zenodo.org/record/13158703/files/figure.png" pageId="75" pageNumber="76">58 A</figureCitation>
); conductor fan entire.
<emphasis id="41F73E37180FFF91441B340CA890EC56" bold="true" box="[852,935,1283,1306]" pageId="75" pageNumber="76">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="733CE225180FFF9147E4342CABE8EC16" blockId="75.[122,1174,220,1628]" pageId="75" pageNumber="76">
<emphasis id="41F73E37180FFF9147E4342CAA2DEC76" bold="true" box="[171,282,1315,1338]" pageId="75" pageNumber="76">Variation.</emphasis>
Male. Variation unknown:
<taxonomicName id="B48399A6180FFF914507342CA827EC76" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[584,784,1315,1338]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180FFF914507342CA827EC76" bold="true" box="[584,784,1315,1338]" italics="true" pageId="75" pageNumber="76">Uduba rakotozafy</emphasis>
</taxonomicName>
is only known from
<specimenCount id="658529AC180FFF91434F342CAF51EC76" box="[1024,1126,1315,1338]" count="1" pageId="75" pageNumber="76" type="male">one male</specimenCount>
, the
<typeStatus id="AC385C87180FFF914734344CABEBEC16" box="[123,220,1347,1370]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="733CE225180FFF9147E4346BAF13ECF7" blockId="75.[122,1174,220,1628]" pageId="75" pageNumber="76">
<materialsCitation id="C3EBE878180FFF9147E4346BAF17ECF7" collectingDate="1993-11-04" collectingDateMax="1993-11-05" collectingDateMin="1993-11-04" collectionCode="CAS" collectorName="J. Coddington &amp; N. Scharff &amp; S. Larcher &amp; C. Griswold &amp; R. Andriamasimanana" country="Madagascar" elevation="1000" location="Andasibe" municipality="Parc National Perinet" pageId="75" pageNumber="76" specimenCode="CASENT9006024" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
M
<emphasis id="41F73E37180FFF91478D346BAAB8EC37" bold="true" box="[194,399,1380,1403]" pageId="75" pageNumber="76">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5180FFF9146D3346BA969EC37" box="[412,606,1380,1403]" name="Madagascar" pageId="75" pageNumber="76">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37180FFF914521346BA87BEC37" box="[622,844,1380,1403]" italics="true" pageId="75" pageNumber="76">
<collectingRegion id="B1472CC7180FFF914521346BA87BEC37" box="[622,844,1380,1403]" country="Madagascar" name="Toamasina" pageId="75" pageNumber="76">Toamasina Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F180FFF91442E346BAF61EC37" box="[865,1110,1380,1403]" pageId="75" pageNumber="76">Parc National Périnet</collectingMunicipality>
, near
<location id="765CB4FE180FFF914734348BABD5ECD7" LSID="urn:lsid:plazi:treatment:FB2A5333180EFF9147343430A8E1EF10:765CB4FE180FFF914734348BABD5ECD7" box="[123,226,1412,1435]" country="Madagascar" municipality="Parc National Perinet" name="Andasibe" pageId="75" pageNumber="76" stateProvince="Toamasina">Andasibe</location>
, 18°56ʹS, 48°24ʹE, elev.
<quantity id="B47B4FC0180FFF9146B4348BA97FECD7" box="[507,584,1412,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="75" pageNumber="76" unit="m" value="1000.0">
<elevation id="F8AE0516180FFF9146B4348BA97FECD7" box="[507,584,1412,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="75" pageNumber="76" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<date id="073DC4E5180FFF914518348BA808ECD7" box="[599,831,1412,1435]" pageId="75" pageNumber="76" value="1993-11-04" valueMax="1993-11-05" valueMin="1993-11-04">
<collectingDate id="17793D0D180FFF914518348BA808ECD7" box="[599,831,1412,1435]" pageId="75" pageNumber="76" value="1993-11-04" valueMax="1993-11-05" valueMin="1993-11-04">4–5 November 1993</collectingDate>
</date>
,
<collectorName id="DE7687F3180FFF914401348BA8DBECD7" box="[846,1004,1412,1435]" pageId="75" pageNumber="76">J. Coddington</collectorName>
,
<collectorName id="DE7687F3180FFF9144B4348BAF47ECD7" box="[1019,1136,1412,1435]" pageId="75" pageNumber="76">N. Scharff</collectorName>
,
<collectorName id="DE7687F3180FFF9143CF348BABFBECF7" pageId="75" pageNumber="76">S. Larcher</collectorName>
,
<collectorName id="DE7687F3180FFF91479534ABAA57ECF7" box="[218,352,1444,1467]" pageId="75" pageNumber="76">C. Griswold</collectorName>
,
<collectorName id="DE7687F3180FFF91462334ABA96EECF7" box="[364,601,1444,1467]" pageId="75" pageNumber="76">R. Andriamasimanana</collectorName>
(
<typeStatus id="AC385C87180FFF91452634ABA9E7ECF7" box="[617,720,1444,1467]" pageId="75" pageNumber="76" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E180FFF91459334ABA893ECF7" box="[732,932,1444,1467]" collectionCode="CASENT" pageId="75" pageNumber="76">CASENT9006024</specimenCode>
,
<specimenCount id="658529AC180FFF9144FF34ABA8E6ECF7" box="[944,977,1444,1467]" count="1" pageId="75" pageNumber="76" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180FFF9144AF34ABAF2FECF7" box="[992,1048,1444,1467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="75" pageNumber="76" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225180FFF9147E434CBA83EEF50" blockId="75.[122,1174,220,1628]" pageId="75" pageNumber="76">
<emphasis id="41F73E37180FFF9147E434CBAA69EC97" bold="true" box="[171,350,1476,1499]" pageId="75" pageNumber="76">Natural history.</emphasis>
The lone specimen of
<taxonomicName id="B48399A6180FFF91450134CBA824EC97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[590,787,1476,1499]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180FFF91450134CBA824EC97" bold="true" box="[590,787,1476,1499]" italics="true" pageId="75" pageNumber="76">Uduba rakotozafy</emphasis>
</taxonomicName>
was collected on the ground in rainforest. The vestige of a divided cribellum on the male suggests that
<taxonomicName id="B48399A6180FFF91441934EAA8DCECB0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[854,1003,1509,1532]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180FFF91441934EAA8DCECB0" bold="true" box="[854,1003,1509,1532]" italics="true" pageId="75" pageNumber="76">U. rakotozafy</emphasis>
</taxonomicName>
females deploy cribellate sticky silk, but we know nothing of their behavior.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180FFF9147E4372AA8E1EF10" pageId="75" pageNumber="76" type="distribution">
<paragraph id="733CE225180FFF9147E4372AA8E1EF10" blockId="75.[122,1174,220,1628]" pageId="75" pageNumber="76">
<emphasis id="41F73E37180FFF9147E4372AAA0AEF70" bold="true" box="[171,317,1573,1596]" pageId="75" pageNumber="76">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6180FFF91469D372AA9ADEF70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[466,666,1573,1596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rakotozafy">
<emphasis id="41F73E37180FFF91469D372AA9ADEF70" bold="true" box="[466,666,1573,1596]" italics="true" pageId="75" pageNumber="76">Uduba rakotozafy</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C87180FFF9144F9372AA8D3EF70" box="[950,996,1573,1596]" pageId="75" pageNumber="76">type</typeStatus>
locality at Parc National Périnet in the central eastern escarpment of
<collectingCountry id="0B94A2B5180FFF9145F4374AA80BEF10" box="[699,828,1605,1628]" name="Madagascar" pageId="75" pageNumber="76">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA0180FFF914404374AA8A8EF10" box="[843,927,1605,1628]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="75" pageNumber="76">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA0180FFF9144E3374AA8FEEF10" box="[940,969,1605,1628]" captionStart="MAP" captionStartId="185.[155,174,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-17@185.[648,1176,216,1157]" captionTargetPageId="185" captionText="MAP 19. Distribution of Group V, the Uduba rajery MAP 20. Distribution of Group VI, the Uduba danielae group: Uduba rakotozafy and U. rajery. group: Uduba jayjay and U. danielae." figureDoi="http://doi.org/10.5281/zenodo.13158797" httpUri="https://zenodo.org/record/13158797/files/figure.png" pageId="75" pageNumber="76">19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="CC73A0E5F4BC8DD131898F42755A2D3D" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331810FF8C47343570AF0FE8DF" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba taralily Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="CC92A913-00FD-40EF-AA3B-17271ED9F3CE" docUuidSource="ZooBank" docVersion="2" lastPageNumber="87" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="85" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B3EF6B1F3D458C0B6862DA987F2DF898">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="AA7E750BEB728D08F7EE0E6B691B1C68">Griswold, Charles</mods:namePart>
<mods:affiliation id="DBEF0146679988D248415F455B27EA10">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="AFC5CEAAB40AD56FC38FACAD1DD3ACBD">Ubick, Darrell</mods:namePart>
<mods:affiliation id="A6DA7E8F6F6BD8AF3021E9B86FB16825">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
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<mods:namePart id="8419E1E68A60FA9AF717F009FC766D65">Ledford, Joel</mods:namePart>
<mods:affiliation id="53355896EC919EC6D6511BC520598C33">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="2EA8AA394334CED5642C0B2F43D61FBA">Polotow, Daniele</mods:namePart>
<mods:affiliation id="ACD5A427D35D7A3A3D5D1012B6D3FB94">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:date id="54E021201B043EF0FA9732DA506F39D8">2022</mods:date>
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<emphasis id="41F73E371810FF8E47343570AA9AEDDA" bold="true" box="[123,429,1151,1174]" pageId="84" pageNumber="85">
<taxonomicName id="B48399A61810FF8E47343570AA2CEDDA" ID-CoL="C8KK5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,283,1151,1174]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="taralily" status="sp. nov.">
<emphasis id="41F73E371810FF8E47343570AA2CEDDA" bold="true" box="[123,283,1151,1174]" italics="true" pageId="84" pageNumber="85">Uduba taralily</emphasis>
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,
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<paragraph id="733CE2251810FF8E47343590AA97EDF8" blockId="84.[123,429,1151,1204]" box="[123,416,1183,1204]" pageId="84" pageNumber="85">
<figureCitation id="EBB8FEA01810FF8E47343590AA21EDF8" box="[123,278,1183,1204]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="84" pageNumber="85">Figures 63 A–C</figureCitation>
, Maps 10, 14.
</paragraph>
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<subSubSection id="3B99B1AE1810FF8E47E435CBABF8EC30" pageId="84" pageNumber="85" type="materials_examined">
<paragraph id="733CE2251810FF8E47E435CBABF8EC30" blockId="84.[123,1174,1220,1725]" pageId="84" pageNumber="85">
<emphasis id="41F73E371810FF8E47E435CBAA65ED97" bold="true" box="[171,338,1220,1243]" pageId="84" pageNumber="85">Type material.</emphasis>
<materialsCitation id="C3EBE8781810FF8E461335CBA8AAEC50" collectingDate="2006-10-25" collectingDateMax="2006-11-04" collectingDateMin="2006-10-25" collectionCode="CAS" collectorName="R. Harin'Hala &amp; M. Irwin" country="Madagascar" elevation="822" location="Miandritsara Forest" municipality="Km" pageId="84" pageNumber="85" specimenCode="CASENT9042524" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871810FF8E461335CBAA88ED97" box="[348,447,1220,1243]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
male (
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) from malaise at
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</quantity>
elevation in the
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, 40
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<locationDeviation id="628EFB071810FF8E46EE35EBA975EDB7" box="[417,578,1252,1275]" location="Ambositra" pageId="84" pageNumber="85">S of Ambositra</locationDeviation>
,
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,
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, collected
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<collectingDate id="17793D0D1810FF8E437735EBAA59EC50" pageId="84" pageNumber="85" value="2006-10-25" valueMax="2006-11-04" valueMin="2006-10-25">25 October–4 November 2006</collectingDate>
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by
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and
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, deposited in
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</materialsCitation>
.
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,
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, one from
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,
<collectingRegion id="B1472CC71810FF8E4548342AA9D9EC70" box="[519,750,1317,1340]" country="Madagascar" name="Antsiranana" pageId="84" pageNumber="85">Antsiranana Province</collectingRegion>
(
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) deposited in
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</materialsCitation>
,
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and one from
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,
<collectingRegion id="B1472CC71810FF8E46B4344AA986EC10" box="[507,689,1349,1372]" country="Madagascar" name="Toliara" pageId="84" pageNumber="85">Toliara Province</collectingRegion>
(
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#-0000-044-065) deposited in
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1810FF8E47E4348AA8B5ECF0" pageId="84" pageNumber="85" type="etymology">
<paragraph id="733CE2251810FF8E47E4348AA8B5ECF0" blockId="84.[123,1174,1220,1725]" pageId="84" pageNumber="85">
<emphasis id="41F73E371810FF8E47E4348AAA1DECD0" bold="true" box="[171,298,1413,1436]" pageId="84" pageNumber="85">Etymology.</emphasis>
The species epithet is from the Malagasy word
<emphasis id="41F73E371810FF8E4475348AA8BEECD0" box="[826,905,1413,1436]" italics="true" pageId="84" pageNumber="85">taralily</emphasis>
, which is an annual rice harvest festival held on the island of
<collectingCountry id="0B94A2B51810FF8E454234AAA9A6ECF0" box="[525,657,1445,1468]" name="Madagascar" pageId="84" pageNumber="85">Madagascar</collectingCountry>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1810FF8F47E434CAA97FE81F" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="diagnosis">
<paragraph id="733CE2251810FF8F47E434CAA97FE81F" blockId="84.[123,1174,1220,1725]" lastBlockId="85.[122,1175,220,1725]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="41F73E371810FF8E47E434CAAAFEEC90" bold="true" box="[171,457,1477,1500]" pageId="84" pageNumber="85">
Diagnosis.
<taxonomicName id="B48399A61810FF8E466834CAAAFEEC90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,457,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371810FF8E466834CAAAFEEC90" bold="true" box="[295,457,1477,1500]" italics="true" pageId="84" pageNumber="85">Uduba taralily</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.c, the
<taxonomicName id="B48399A61810FF8E446C34CAA8F5EC90" authorityName="Simon" authorityYear="1906" box="[803,962,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01810FF8E435734CAAF59EC90" box="[1048,1134,1477,1500]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="84" pageNumber="85">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. Males can be distinguished from those of other cribellate (with at least vestige of a divided cribellum)
<taxonomicName id="B48399A61810FF8E4569370AA947EF50" box="[550,624,1541,1564]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371810FF8E4569370AA947EF50" bold="true" box="[550,624,1541,1564]" italics="true" pageId="84" pageNumber="85">Uduba</emphasis>
</taxonomicName>
by having the male palp (
<figureCitation id="EBB8FEA01810FF8E44DF370AAF16EF50" box="[912,1057,1541,1564]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="84" pageNumber="85">Figs. 63 A–C</figureCitation>
) with TA2 highly sclerotized, black, concave medially against TA3, MA forked, concave in middle with proximal and distal small pointed projections (
<figureCitation id="EBB8FEA01810FF8E45733749A993EF11" box="[572,676,1606,1629]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="84" pageNumber="85">Fig. 63 B</figureCitation>
). The forked MA distinguishes
<taxonomicName id="B48399A61810FF8E44B93749AFA1EF11" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1014,1174,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371810FF8E44B93749AFA1EF11" bold="true" box="[1014,1174,1606,1629]" italics="true" pageId="84" pageNumber="85">Uduba taralily</emphasis>
</taxonomicName>
from other divided cribellum males:
<taxonomicName id="B48399A61810FF8E455E3769A9B3EF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[529,644,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371810FF8E455E3769A9B3EF31" bold="true" box="[529,644,1638,1661]" italics="true" pageId="84" pageNumber="85">U. taralily</emphasis>
</taxonomicName>
are distinguished from those of
<taxonomicName id="B48399A61810FF8E44A03769AF52EF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1007,1125,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371810FF8E44A03769AF52EF31" bold="true" box="[1007,1125,1638,1661]" italics="true" pageId="84" pageNumber="85">U. woodae</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61810FF8E47343789AA16EFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,289,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371810FF8E47343789AA16EFD1" bold="true" box="[123,289,1670,1693]" italics="true" pageId="84" pageNumber="85">U. lehibekokoa</emphasis>
</taxonomicName>
by having the MA with proximal and distal small pointed projections (
<figureCitation id="EBB8FEA01810FF8E43503789AFBFEFD1" box="[1055,1160,1670,1693]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="84" pageNumber="85">Fig. 63 B</figureCitation>
), tegulum apex at 0.40 tegulum length from MA apex, length (including apical processes) 0.69 times width:
<taxonomicName id="B48399A61811FF8F478731D3AA47E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[200,368,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371811FF8F478731D3AA47E9BF" bold="true" box="[200,368,220,243]" italics="true" pageId="85" pageNumber="86">U. lehibekokoa</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01811FF8F46CD31D3A922E9BF" box="[386,533,220,243]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="85" pageNumber="86">Figs. 50 A–C</figureCitation>
) and
<taxonomicName id="B48399A61811FF8F451931D3A9FBE9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[598,716,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371811FF8F451931D3A9FBE9BF" bold="true" box="[598,716,220,243]" italics="true" pageId="85" pageNumber="86">U. woodae</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01811FF8F459231D3A847E9BF" box="[733,880,220,243]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="85" pageNumber="86">Figs. 66 A–C</figureCitation>
) have the MA with only a proximal sharp point, and the MA smaller and farther distad, with the tegulum apex less than 0.35 tegulum length from MA apex, MA length (including apical processes) less than 0.50 times width. The female of
<taxonomicName id="B48399A61811FF8F46543033AA8BE81F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[283,444,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371811FF8F46543033AA8BE81F" bold="true" box="[283,444,316,339]" italics="true" pageId="85" pageNumber="86">Uduba taralily</emphasis>
</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1811FF8C47E43053AA8DE81F" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86" type="description">
<paragraph id="733CE2251811FF8F47E43053A894ED15" blockId="85.[122,1175,220,1725]" pageId="85" pageNumber="86">
<emphasis id="41F73E371811FF8F47E43053AA41E83F" bold="true" box="[171,374,348,371]" pageId="85" pageNumber="86">Description. Male</emphasis>
(
<typeStatus id="AC385C871811FF8F46CB3053AADAE83F" box="[388,493,348,371]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
). Total length 10.80. Markings typical of
<taxonomicName id="B48399A61811FF8F44EA3053A8D8E83F" box="[933,1007,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371811FF8F44EA3053A8D8E83F" bold="true" box="[933,1007,348,371]" italics="true" pageId="85" pageNumber="86">Uduba</emphasis>
</taxonomicName>
. Carapace 6.70 long, 4.60 wide, 2.00 high; clypeus 0.30 high. Eye diameters: AME 0.20, ALE 0.22, PME 0.24, PLE 0.30. Chelicerae 2.90 long; sternum 3.00 long, 2.20 wide; labium 1.30 long; palpal coxae 2.10 long. Divided cribellum. Spination. palpus–femur d1-1-3; leg I–femur d1-1-0-1, p0-0-0-2, r0-1-0- 1, patella p1, r1, tibia p1-0-0-0, v2-2-0-1, r1-0-0-0, metatarsus p0-0-1-2, v2-2-2, r0-1-0-1; leg II– femur d1-1-1, p0-1-0-2, r0-1-1-1, patella p1, r1, tibia d0-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p0-1-0, v2-2-2, r0-1-0; leg III–femur d1-1-1, p0-1-0-2, r0-1-0-2, patella p1, r1, tibia d0- 1-0, p0-1-1-0, v2-1-2, r0-1-1-0, metatarsus p1-2-0-2, v2-2-0-2, r1-1-0-2; leg IV–femur d1-1-1, p0- 1-1-1, r0-0-0-1, patella p1, r1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1-2, v1- 1-1-1-1-2, r1-1-0-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements: I: 5.80 + 2.50 + 4.50 + 4.90 + 3.10 = 20.80; II: 5.50 + 2.30 + 4.50 + 5.00 + 3.20 = 20.50; III: 4.70 + 2.10 + 3.30 + 4.10 + 2.60 = 16.80; IV: 6.30 + 2.10 + 5.40 + 7.00 + 3.30 = 24.10; palpus: 2.30 + 0.70 + 1.10 + NA + 3.10 = 7.20. Leg formula 4123.
<emphasis id="41F73E371811FF8F459D33D0A871EBBA" bold="true" box="[722,838,735,758]" pageId="85" pageNumber="86">Male palp</emphasis>
: palpal tibia 0.395 times cymbial length, RTA broadly triangular, with pointed apex sharply-curved ventrally (
<figureCitation id="EBB8FEA01811FF8F44A833F0AF64EA5A" box="[999,1107,767,790]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Fig. 63 C</figureCitation>
), RTA length 0.315 times tibia length, length equals width, VTA length 1.33 times width, length 0.43 times tibia width, VTA conical, straight, with rounded apex (
<figureCitation id="EBB8FEA01811FF8F459F324FA850EA1B" box="[720,871,832,855]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Figs. 63 A, B</figureCitation>
); tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.94 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.30 times tegulum length; TA3 transverse, tapering to a blunt apex, concave dorsally, TA3 apex reflexed (
<figureCitation id="EBB8FEA01811FF8F450732AEA9EEEAF4" box="[584,729,929,952]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Figs. 63 B, C</figureCitation>
), visible in front of TA2; TA2 highly sclerotized, black, concave medially against TA3 apex and even with TA3 apex (
<figureCitation id="EBB8FEA01811FF8F448F32CEAF62EA94" box="[960,1109,961,984]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Figs. 63 A, B</figureCitation>
); MA transverse, concave in middle with proximal and distal small pointed projections (
<figureCitation id="EBB8FEA01811FF8F44BE32EEAF6DEAB4" box="[1009,1114,993,1016]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Fig. 63 B</figureCitation>
), origin at 0.44 times tegulum length, tegulum apex at 0.40 tegulum length from MA apex, length (including apical processes) 0.69 times width, MA small, length 0.20 tegulum length, width 0.27 times tegulum width; conductor fan entire (
<figureCitation id="EBB8FEA01811FF8F451E354DA98BED15" box="[593,700,1090,1113]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="85" pageNumber="86">Fig. 63 C</figureCitation>
).
<emphasis id="41F73E371811FF8F459F354DA814ED15" bold="true" box="[720,803,1090,1113]" pageId="85" pageNumber="86">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="733CE2251811FF8F47E4356DA9F6EC37" blockId="85.[122,1175,220,1725]" pageId="85" pageNumber="86">
<emphasis id="41F73E371811FF8F47E4356DAA1BED35" bold="true" box="[171,300,1122,1145]" pageId="85" pageNumber="86">Variation. (</emphasis>
N = 3): Total length 10.89–14.30, carapace length / width = 1.41–1.46, carapace height / width = 0.43–0.50, PER / carapace width = 0.36–0.37, PER / OAL = 2.73–2.87, PER / AER = 1.25–1.38, OAL / OQL = 1.07–1.12, OQP / OQA =1.03–1.17, clypeus height / AME diameter width = 1.50–1.67, cheliceral length / clypeus height = 8.25–9.67, sternum length / width = 1.36– 1.38, palpal coxa length / width = 2.63–2.67, femur I length / carapace width = 1.26–1.33, metatarsus I length / carapace width = 1.07–1.39, femur IV length / carapace width = 1.35–1.37, cymbium length / carapace width = 0.55–0.67, cymbium length / palpal patella length = 3.60–4.43, cymbium length / palpal tibia length = 2.82–3.27, cymbium length / palpal femur length = 1.29–1.35, palpal tibia length / palpal patella length = 1.10–1.57.
</paragraph>
<paragraph id="733CE2251811FF8F47E4348BAA08EFD1" blockId="85.[122,1175,220,1725]" pageId="85" pageNumber="86">
M
<emphasis id="41F73E371811FF8F478D348BAAB0ECD7" bold="true" box="[194,391,1412,1435]" pageId="85" pageNumber="86">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51811FF8F46C2348BA978ECD7" box="[397,591,1412,1435]" name="Madagascar" pageId="85" pageNumber="86">MADAGASCAR</collectingCountry>
: A
<emphasis id="41F73E371811FF8F4526348BA876ECD7" box="[617,833,1412,1435]" italics="true" pageId="85" pageNumber="86">ntsiranana Province</emphasis>
: Sahamalaza National Park,
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W Maropapango, MG69-Region Sofia, District Analalava, Ankarafa subhumid evergreen forest,
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,
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, elev.
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,
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, R. Harin’Hala and M. Irwin (
<typeStatus id="AC385C871811FF8F47CD34EAABD5ECB0" box="[130,226,1509,1532]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
, CASENT9064667,
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, CAS).
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</emphasis>
:
<collectingMunicipality id="9358785F1811FF8F440D34EAAF21ECB0" box="[834,1046,1509,1532]" pageId="85" pageNumber="86">Miandritsara Forest</collectingMunicipality>
, 40
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<locationDeviation id="628EFB071811FF8F432534EAABD9EF50" location="Ambositra" pageId="85" pageNumber="86">S of Ambositra</locationDeviation>
, 20°47.56ʹS, 47°10.54ʹE, elev.
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</quantity>
, malaise in low altitude rainforest,
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<collectingDate id="17793D0D1811FF8F434E370AAA18EF70" pageId="85" pageNumber="86" value="2006-10-25" valueMax="2006-11-04" valueMin="2006-10-25">25 October–4 November 2006</collectingDate>
</date>
,
<collectorName id="DE7687F31811FF8F4670372AAAECEF70" box="[319,475,1573,1596]" pageId="85" pageNumber="86">R. Harin’Hala</collectorName>
and
<collectorName id="DE7687F31811FF8F4557372AA94AEF70" box="[536,637,1573,1596]" pageId="85" pageNumber="86">M. Irwin</collectorName>
[MG-29-66] (
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,
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,
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,
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)
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.
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:
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SSE of Befandefa
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, 22°13.00ʹS, 43°19.80ʹE, elev.
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#10–12,
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(
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,
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#-0000-044- 065,
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,
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.
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<emphasis id="41F73E371811FF8F47E437A9AA6CEFF1" bold="true" box="[171,347,1702,1725]" pageId="85" pageNumber="86">Natural history</emphasis>
. Males of
<taxonomicName id="B48399A61811FF8F468037A9A958EFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[463,623,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371811FF8F468037A9A958EFF1" bold="true" box="[463,623,1702,1725]" italics="true" pageId="85" pageNumber="86">Uduba taralily</emphasis>
</taxonomicName>
have been collected in various forest habitats from only
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elevation by pitfalls up to a collection by malaise trap at over
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elevation. Collection of individuals in pitfalls and malaise traps suggest that males wander on the ground and up into the vegetation. Males have a vestige of a divided cribellum, but we know nothing of the use of cribellate silk by this species.
</paragraph>
</subSubSection>
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<emphasis id="41F73E371812FF8C47E43053AA0AE83F" bold="true" box="[171,317,348,371]" pageId="86" pageNumber="87">Distribution.</emphasis>
The three widely separated records of this species (Maps 10, 14) suggest that
<taxonomicName id="B48399A61812FF8C47343073AA2BE8DF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,284,380,403]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371812FF8C47343073AA2BE8DF" bold="true" box="[123,284,380,403]" italics="true" pageId="86" pageNumber="87">Uduba taralily</emphasis>
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occurs throughout dry forests west of the mountain spine of
<collectingCountry id="0B94A2B51812FF8C44FB3073AF05E8DF" box="[948,1074,380,403]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="B5E7E3B6734C5CF3E5A8D71F41D7FCF0">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="A24A5E4BE97C4A3E95438427639CB780">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="69D0827A8A5386E6330F0CF9D9A69BDB">Ubick, Darrell</mods:namePart>
<mods:affiliation id="10EB7B3667EA54946C60392DFD49C86F">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="EE6E95C505311314C7A598BADC0DE3BE">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<emphasis id="41F73E371812FF8C473430B3AA27E89F" bold="true" box="[123,272,444,467]" italics="true" pageId="86" pageNumber="87">Uduba valiha</emphasis>
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,
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<figureCitation id="EBB8FEA01812FF8C473430D3ABD9E8BD" box="[123,238,476,497]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: A–E = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="86" pageNumber="87">Figures 6 C</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C47B730D3AA2EE8BD" box="[248,281,476,497]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="86" pageNumber="87">8 F</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C466C30D3AA63E8BD" box="[291,340,476,497]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="86" pageNumber="87">28 B</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C461130D3AA9CE8BD" box="[350,427,476,497]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="86" pageNumber="87">64 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C46F930D3AAC9E8BD" box="[438,510,476,497]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="86" pageNumber="87">65 A–F</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C454730D3A96FE8BD" box="[520,600,476,497]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">72 G, H</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C452C30D3A99EE8BD" box="[611,681,476,497]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="86" pageNumber="87">80 H, I</figureCitation>
, Maps 10, 17.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1812FF8C47E4330DA847EBD5" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="733CE2251812FF8C47E4330DA847EBD5" blockId="86.[123,1174,514,1725]" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C47E4330DAA5AEB55" bold="true" box="[171,365,514,537]" pageId="86" pageNumber="87">Type material.</emphasis>
<materialsCitation id="C3EBE8781812FF8C46C8330DA8EAEB15" collectingDate="2006-04-23" collectionCode="CAS" collectorName="de Vevembe &amp; B. L. Fisher" country="Madagascar" elevation="600" location="Foret de Vevembe" pageId="86" pageNumber="87" specimenCode="CASENT9026394, CASENT9082181" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871812FF8C46C8330DAAC5EB55" box="[391,498,514,537]" pageId="86" pageNumber="87" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1812FF8C452B330DA80FEB55" box="[612,824,514,537]" collectionCode="CASENT" pageId="86" pageNumber="87">CASENT9026394</specimenCode>
) and
<specimenCount id="658529AC1812FF8C44D0330DAFA2EB55" box="[927,1173,514,537]" count="1" pageId="86" pageNumber="87" type="male" typeStatus="paratype">one paratype male</specimenCount>
(
<specimenCode id="23254A5E1812FF8C47CD332DAA7DEB75" box="[130,330,546,569]" collectionCode="CASENT" pageId="86" pageNumber="87">CASENT9082181</specimenCode>
) collected in rainforest at
<quantity id="B47B4FC01812FF8C4522332DA99EEB75" box="[621,681,546,569]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="86" pageNumber="87" unit="m" value="600.0">
<elevation id="F8AE05161812FF8C4522332DA99EEB75" box="[621,681,546,569]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="86" pageNumber="87" unit="m" value="600.0">600m</elevation>
</quantity>
elevation in
<location id="765CB4FE1812FF8C4474332DAF37EB75" LSID="urn:lsid:plazi:treatment:FB2A53331812FF82473430B3AAC0EA39:765CB4FE1812FF8C4474332DAF37EB75" box="[827,1024,546,569]" country="Madagascar" name="Foret de Vevembe" pageId="86" pageNumber="87" stateProvince="Fianarantsoa">
Forêt
<collectorName id="DE7687F31812FF8C4433332DAF37EB75" box="[892,1024,546,569]" pageId="86" pageNumber="87">de Vevembe</collectorName>
</location>
,
<collectingRegion id="B1472CC71812FF8C4342332DABEAEB15" country="Madagascar" name="Fianarantsoa" pageId="86" pageNumber="87">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51812FF8C47A7334DAA50EB15" box="[232,359,578,601]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
, on
<date id="073DC4E51812FF8C46D8334DA91CEB15" box="[407,555,578,601]" pageId="86" pageNumber="87" value="2006-04-23">
<collectingDate id="17793D0D1812FF8C46D8334DA91CEB15" box="[407,555,578,601]" pageId="86" pageNumber="87" value="2006-04-23">23 April 2006</collectingDate>
</date>
by
<collectorName id="DE7687F31812FF8C4519334DA9E7EB15" box="[598,720,578,601]" pageId="86" pageNumber="87">B.L. Fisher</collectorName>
<emphasis id="41F73E371812FF8C4598334DA830EB15" box="[727,775,578,601]" italics="true" pageId="86" pageNumber="87">et al</emphasis>
., deposited in
<collectionCode id="15927AE01812FF8C44E9334DA8EAEB15" box="[934,989,578,601]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="86" pageNumber="87" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781812FF8C44A7334DA85CEBD5" collectingDate="1993-09-24" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="785" location="Massif Andringitra" pageId="86" pageNumber="87" specimenCode="CASENT9006071" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871812FF8C44A7334DAF72EB15" box="[1000,1093,578,601]" pageId="86" pageNumber="87" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1812FF8C47CD336DAA7DEB35" box="[130,330,610,633]" collectionCode="CASENT" pageId="86" pageNumber="87">CASENT9006071</specimenCode>
) collected at
<quantity id="B47B4FC01812FF8C4692336DA92EEB35" box="[477,537,610,633]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="86" pageNumber="87" unit="m" value="785.0">
<elevation id="F8AE05161812FF8C4692336DA92EEB35" box="[477,537,610,633]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="86" pageNumber="87" unit="m" value="785.0">785m</elevation>
</quantity>
elevation from
<location id="765CB4FE1812FF8C4588336DA8A3EB35" LSID="urn:lsid:plazi:treatment:FB2A53331812FF82473430B3AAC0EA39:765CB4FE1812FF8C4588336DA8A3EB35" box="[711,916,610,633]" country="Madagascar" name="Massif Andringitra" pageId="86" pageNumber="87" stateProvince="Fianarantsoa">Massif Andringitra</location>
,
<collectingRegion id="B1472CC71812FF8C44EF336DAFA5EB35" box="[928,1170,610,633]" country="Madagascar" name="Fianarantsoa" pageId="86" pageNumber="87">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51812FF8C4734338DABCBEBD5" box="[123,252,642,665]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
on
<date id="073DC4E51812FF8C4669338DAACFEBD5" box="[294,504,642,665]" pageId="86" pageNumber="87" value="1993-09-24">
<collectingDate id="17793D0D1812FF8C4669338DAACFEBD5" box="[294,504,642,665]" pageId="86" pageNumber="87" value="1993-09-24">24 September 1993</collectingDate>
</date>
by
<collectorName id="DE7687F31812FF8C456C338DA9ACEBD5" box="[547,667,642,665]" pageId="86" pageNumber="87">B.L. Fisher</collectorName>
, deposited in
<collectionCode id="15927AE01812FF8C447B338DA85CEBD5" box="[820,875,642,665]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="86" pageNumber="87" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1812FF8C47E433ADAFA1EAB7" pageId="86" pageNumber="87" type="discussion">
<paragraph id="733CE2251812FF8C47E433ADAFA1EAB7" blockId="86.[123,1174,514,1725]" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C47E433ADAA24EBF5" bold="true" box="[171,275,674,697]" pageId="86" pageNumber="87">Remarks</emphasis>
. The form of the male palp is similar among
<taxonomicName id="B48399A61812FF8C45B333ADA8A6EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[764,913,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C45B333ADA8A6EBF5" bold="true" box="[764,913,674,697]" italics="true" pageId="86" pageNumber="87">Uduba valiha</emphasis>
</taxonomicName>
,
<taxonomicName id="B48399A61812FF8C44D033ADAF3FEBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[927,1032,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371812FF8C44D033ADAF3FEBF5" bold="true" box="[927,1032,674,697]" italics="true" pageId="86" pageNumber="87">U. fisheri</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61812FF8C437233ADAB9BEB96" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C437233ADAB9BEB96" bold="true" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
: females of
<taxonomicName id="B48399A61812FF8C466033CCAAA0EB96" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[303,407,707,730]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371812FF8C466033CCAAA0EB96" bold="true" box="[303,407,707,730]" italics="true" pageId="86" pageNumber="87">U. fisheri</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61812FF8C468533CCA97DEB96" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[458,586,707,730]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C468533CCA97DEB96" bold="true" box="[458,586,707,730]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
have been collected with and therefore associated with males but no females have been collected with
<taxonomicName id="B48399A61812FF8C453A33ECA9EDEBB6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[629,730,739,762]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C453A33ECA9EDEBB6" bold="true" box="[629,730,739,762]" italics="true" pageId="86" pageNumber="87">U. valiha</emphasis>
</taxonomicName>
. The form of the female genitalia of both the species
<taxonomicName id="B48399A61812FF8C47B9320CAA69EA56" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[246,350,771,794]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371812FF8C47B9320CAA69EA56" bold="true" box="[246,350,771,794]" italics="true" pageId="86" pageNumber="87">U. fisheri</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61812FF8C46DC320CA924EA56" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[403,531,771,794]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C46DC320CA924EA56" bold="true" box="[403,531,771,794]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
is similar to that of specimen CASENT9006071 from Massif Andringitra, suggesting this female belongs to the same group of species. Because this female is distinct from those of
<taxonomicName id="B48399A61812FF8C46CD324CAADDEA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[386,490,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371812FF8C46CD324CAADDEA16" bold="true" box="[386,490,835,858]" italics="true" pageId="86" pageNumber="87">U. fisheri</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61812FF8C4550324CA9A8EA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[543,671,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C4550324CA9A8EA16" bold="true" box="[543,671,835,858]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
, and geographically near the male localities of
<taxonomicName id="B48399A61812FF8C4734326CABD7EA36" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,224,867,890]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C4734326CABD7EA36" bold="true" box="[123,224,867,890]" italics="true" pageId="86" pageNumber="87">U. valiha</emphasis>
</taxonomicName>
, we tentatively associate the males collected in rainforest at
<quantity id="B47B4FC01812FF8C443D326CA898EA36" box="[882,943,867,890]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="86" pageNumber="87" unit="m" value="600.0">600m</quantity>
elevation in Forêt de Vevembe with the female collected at
<quantity id="B47B4FC01812FF8C455E328CA979EAD6" box="[529,590,899,922]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="86" pageNumber="87" unit="m" value="785.0">785m</quantity>
elevation from Massif Andringitra as
<taxonomicName id="B48399A61812FF8C44A9328CAF7DEAD6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[998,1098,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C44A9328CAF7DEAD6" bold="true" box="[998,1098,899,922]" italics="true" pageId="86" pageNumber="87">U. valiha</emphasis>
</taxonomicName>
. There appears to be an error on the label for specimens collected by S. Goodman at Forêt de Vinetalo [FMHD-00-223, i.e., CASENT9064656,
<specimenCount id="658529AC1812FF8C451932CCA940EA96" box="[598,631,963,986]" count="3" pageId="86" pageNumber="87" type="male">3♂</specimenCount>
, FMNH): the label states “[212°[
<emphasis id="41F73E371812FF8C435D32CCAF07EA96" box="[1042,1072,963,986]" italics="true" pageId="86" pageNumber="87">sic</emphasis>
] 46.6ʹS, 47°20.8ʹE” but according to maps and Google Earth the locality should be 21°46.6ʹS, 47°20.8ʹE.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1812FF8C47E4350BA963ED77" pageId="86" pageNumber="87" type="etymology">
<paragraph id="733CE2251812FF8C47E4350BA963ED77" blockId="86.[123,1174,514,1725]" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C47E4350BAA1DED57" bold="true" box="[171,298,1028,1051]" pageId="86" pageNumber="87">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E371812FF8C4544350BA963ED57" box="[523,596,1028,1051]" italics="true" pageId="86" pageNumber="87">valiha,</emphasis>
refers to a tube zither considered the “national instrument” of
<collectingCountry id="0B94A2B51812FF8C47AD352BAA56ED77" box="[226,353,1060,1083]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
, a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1812FF8C47E4354BA985EF50" pageId="86" pageNumber="87" type="diagnosis">
<paragraph id="733CE2251812FF8C47E4354BA985EF50" blockId="86.[123,1174,514,1725]" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C47E4354BAAF7ED17" bold="true" box="[171,448,1092,1115]" pageId="86" pageNumber="87">
Diagnosis.
<taxonomicName id="B48399A61812FF8C4667354BAAF7ED17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,448,1092,1115]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C4667354BAAF7ED17" bold="true" box="[296,448,1092,1115]" italics="true" pageId="86" pageNumber="87">Uduba valiha</emphasis>
</taxonomicName>
</emphasis>
belong to Group III, the
<taxonomicName id="B48399A61812FF8C4593354BA85AED17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[732,877,1092,1115]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">Uduba valiha</taxonomicName>
group (
<figureCitation id="EBB8FEA01812FF8C4489354BAF28ED17" box="[966,1055,1092,1115]" captionStart="MAP" captionStartId="184.[155,174,1176,1195]" captionTargetBox="[120,1166,216,1153]" captionTargetId="figure-11@184.[120,648,216,1161]" captionTargetPageId="184" captionText="MAP 17. Distribution of Group III, the Uduba valiha MAP 18. Distribution of Group IV, the Uduba rinha group: Uduba hiragasy, U. fisheri and U. valiha. group: Uduba rinha and U. irwini." figureDoi="http://doi.org/10.5281/zenodo.13158795" httpUri="https://zenodo.org/record/13158795/files/figure.png" pageId="86" pageNumber="87">Map 17</figureCitation>
). They are cribellate with divided cribellum or vestige. Males differ from other species by having a retromedian ridge on the tegulum (
<figureCitation id="EBB8FEA01812FF8C46E9358BA92CEDD7" box="[422,539,1156,1179]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="86" pageNumber="87">Figs. 64 C</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C4566358BA968EDD7" box="[553,607,1156,1179]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="86" pageNumber="87">65 E</figureCitation>
), and a deeply bifid MA (
<figureCitation id="EBB8FEA01812FF8C4433358BA8D1EDD7" box="[892,998,1156,1179]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="86" pageNumber="87">Fig. 28 B</figureCitation>
) with prolateral lobe slender, retrolateral lobe stout;
<taxonomicName id="B48399A61812FF8C46B035ABA952EDF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[511,613,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C46B035ABA952EDF7" bold="true" box="[511,613,1188,1211]" italics="true" pageId="86" pageNumber="87">U. valiha</emphasis>
</taxonomicName>
males differ from those of
<taxonomicName id="B48399A61812FF8C44C135ABAF38EDF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[910,1039,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C44C135ABAF38EDF7" bold="true" box="[910,1039,1188,1211]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
by having a short, stout retrolateral spine on the palpal tibia (
<figureCitation id="EBB8FEA01812FF8C45C235CBA9C1ED97" box="[653,758,1220,1243]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="86" pageNumber="87">Fig. 64 C</figureCitation>
) (spine absent in
<emphasis id="41F73E371812FF8C44F735CBAF77ED97" bold="true" box="[952,1088,1220,1243]" pageId="86" pageNumber="87">
<taxonomicName id="B48399A61812FF8C44F735CBAF0DED97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[952,1082,1220,1243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C44F735CBAF0DED97" bold="true" box="[952,1082,1220,1243]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
,
</emphasis>
<figureCitation id="EBB8FEA01812FF8C430735CBABB8EDB7" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="86" pageNumber="87">Fig. 44 C</figureCitation>
), by having the TA2 apically rolled (
<figureCitation id="EBB8FEA01812FF8C455335EBA9B4EDB7" box="[540,643,1252,1275]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="86" pageNumber="87">Fig. 28 B</figureCitation>
) (blunt, entire in
<taxonomicName id="B48399A61812FF8C440C35EBA8F4EDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[835,963,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C440C35EBA8F4EDB7" bold="true" box="[835,963,1252,1275]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01812FF8C449E35EBAF0FEDB7" box="[977,1080,1252,1275]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="86" pageNumber="87">Fig. 28 A</figureCitation>
) and the apex of the MA prolateral lobe rounded slender (
<figureCitation id="EBB8FEA01812FF8C45CE340AA9C6EC50" box="[641,753,1285,1308]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="86" pageNumber="87">Figs. 64 A</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C45B3340AA819EC50" box="[764,814,1285,1308]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="86" pageNumber="87">65 E</figureCitation>
) (truncate in
<taxonomicName id="B48399A61812FF8C44F2340AAF0AEC50" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[957,1085,1285,1308]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C44F2340AAF0AEC50" bold="true" box="[957,1085,1285,1308]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01812FF8C4305340AABB8EC70" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="86" pageNumber="87">Fig. 44 B</figureCitation>
). Females lack a median lobe or defined atrium on the epigynum (
<figureCitation id="EBB8FEA01812FF8C4400342AA8F5EC70" box="[847,962,1317,1340]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Figs. 72 G</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C4483342AAF36EC70" box="[972,1025,1317,1340]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="86" pageNumber="87">80 H</figureCitation>
) and have the vulva with spermathecal loops few, anteroposteriorly oriented (
<figureCitation id="EBB8FEA01812FF8C446F344AA8A2EC10" box="[800,917,1349,1372]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Figs. 72 H</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C44EE344AA8F9EC10" box="[929,974,1349,1372]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="86" pageNumber="87">80 I</figureCitation>
);
<taxonomicName id="B48399A61812FF8C44AC344AAF7FEC10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[995,1096,1349,1372]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371812FF8C44AC344AAF7FEC10" bold="true" box="[995,1096,1349,1372]" italics="true" pageId="86" pageNumber="87">U. valiha</emphasis>
</taxonomicName>
can be distinguished from those of similar
<taxonomicName id="B48399A61812FF8C46B2346AA9ABEC30" box="[509,668,1381,1404]" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C46B2346AA970EC30" bold="true" box="[509,583,1381,1404]" italics="true" pageId="86" pageNumber="87">Uduba</emphasis>
species
</taxonomicName>
by having the lateral epigynal margins strongly convex (
<figureCitation id="EBB8FEA01812FF8C47BB348AAA54ECD0" box="[244,355,1413,1436]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Fig. 72 G</figureCitation>
,
<figureCitation id="EBB8FEA01812FF8C463F348AAA9FECD0" box="[368,424,1413,1436]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="86" pageNumber="87">80 H</figureCitation>
) (but the lateral margins not developed into the lateral “ears” found in
<taxonomicName id="B48399A61812FF8C47D734AAAA02ECF0" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[152,309,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="evanescens">
<emphasis id="41F73E371812FF8C47D734AAAA02ECF0" bold="true" box="[152,309,1445,1468]" italics="true" pageId="86" pageNumber="87">U. evanescens</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61812FF8C462434AAA962ECF0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[363,597,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="pseudoevanescens">
<emphasis id="41F73E371812FF8C462434AAA962ECF0" bold="true" box="[363,597,1445,1468]" italics="true" pageId="86" pageNumber="87">U. pseudoevanescens</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01812FF8C452B34AAA9C2ECF0" box="[612,757,1445,1468]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="86" pageNumber="87">Figs. 71 A, E</figureCitation>
); distinguished from female
<taxonomicName id="B48399A61812FF8C436334AAAFA2ECF0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1068,1173,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371812FF8C436334AAAFA2ECF0" bold="true" box="[1068,1173,1445,1468]" italics="true" pageId="86" pageNumber="87">U. fisheri</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01812FF8C47CD34CAABDEEC90" box="[130,233,1477,1500]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Fig. 72 E</figureCitation>
). and
<taxonomicName id="B48399A61812FF8C466634CAAA9FEC90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[297,424,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371812FF8C466634CAAA9FEC90" bold="true" box="[297,424,1477,1500]" italics="true" pageId="86" pageNumber="87">U. hiragasy</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01812FF8C46F834CAA92BEC90" box="[439,540,1477,1500]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Fig. 72 C</figureCitation>
) by lacking a strong ridge along epigastric furrow and having lateral lobes with convex edges forming ridges or lobes on each side, these ridges extending around anteriorly and posteriorly of CO (
<figureCitation id="EBB8FEA01812FF8C4576370AA991EF50" box="[569,678,1541,1564]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="86" pageNumber="87">Fig. 72 G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1812FF8247E43729A883E8FF" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87" type="description">
<paragraph id="733CE2251812FF8D47E43729A9DAEF70" blockId="86.[123,1174,514,1725]" lastBlockId="87.[123,1174,220,1725]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
<emphasis id="41F73E371812FF8C47E43729AA41EF71" bold="true" box="[171,374,1574,1597]" pageId="86" pageNumber="87">Description. Male</emphasis>
(
<typeStatus id="AC385C871812FF8C46CA3729AADEEF71" box="[389,489,1574,1597]" pageId="86" pageNumber="87" type="holotype">holotype</typeStatus>
): Total length 9.00. Markings as in
<figureCitation id="EBB8FEA01812FF8C442E3729A885EF71" box="[865,946,1574,1597]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route d’Anosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="86" pageNumber="87">
Fig.
<specimenCount id="658529AC1812FF8C44DC3729A885EF71" box="[915,946,1574,1597]" count="8" pageId="86" pageNumber="87" type="generic">8 F</specimenCount>
</figureCitation>
. Carapace 4.60 long, 3.10 wide, 1.60 high; clypeus 0.25 high. Eye diameters:AME 0.18, ALE, PME and PLE 0.20. Chelicerae 1.80 long; sternum 2.10 long, 1.60 wide; labium 1.00 long; palpal coxae 1.50 long. Divid- ed cribellum. Spination (
<typeStatus id="AC385C871812FF8C46C73789AAD1EFD1" box="[392,486,1670,1693]" pageId="86" pageNumber="87" type="paratype">paratype</typeStatus>
CASENT9082181): palpus–femur d1-1-1, p0-0-1, tibia r0-1-0, p0-1-0; leg I–femur d1-1-1, p0-1-1-0, r0-1-1-1, patella p1, tibia d0-1-1-1, p0-1-1-0, v2-2-2-2, r0- 1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-1; leg II–femur d1-1-1, p0-1-1-0, r0-1-1-1, patella p1, tibia d0-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-1; leg III–femur d1-1-1, p1-0-1-1, r1-0-1-1, patella p1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1- 2, v2-2-1, r1-1-2; leg IV–femur d1-1-1, p0-1-1-1, r0-0-1-0, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1- 1-0, metatarsus p1-2-2, v2-2-2, r1-1-2. Scopulae: cymbium, apicodorsal (
<figureCitation id="EBB8FEA01813FF8D44E93053AF23E83F" box="[934,1044,348,371]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="87" pageNumber="88">Fig. 65 A</figureCitation>
); tarsi I–IV, ventral. Leg measurements (
<typeStatus id="AC385C871813FF8D46E53072A939E8D8" box="[426,526,381,404]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
): I: 4.20 + 1.60 + 4.00 + 4.30 + 2.50 = 16.60; II: 3.90 + 1.50 + 3.40 + 3.60 + 2.20 = 14.60; III: 3.30 + 1.40 + 2.40 + 3.30 + 1.90 = 12.30; IV: 4.60 + 1.40 + 4.00 + 5.20 + 2.30 = 17.50; palpus: 1.80 + 0.70 + 0.80 + NA + 2.00 = 5.30. Leg formula 4123.
<emphasis id="41F73E371813FF8D431330B2ABF7E8B8" bold="true" pageId="87" pageNumber="88">Male palp (</emphasis>
<typeStatus id="AC385C871813FF8D478F30D2AA14E8B8" box="[192,291,477,500]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
): palpal tibia 0.41 cymbial length, RTA triangular, procurved, with pointed apex (
<figureCitation id="EBB8FEA01813FF8D47CC30F1ABCFEB59" box="[131,248,510,533]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Figs. 64 C</figureCitation>
,
<figureCitation id="EBB8FEA01813FF8D464A30F1AA6BEB59" box="[261,348,510,533]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="87" pageNumber="88">65 A, C</figureCitation>
), RTA length 0.33 tibia, VTA triangular, short, length 0.30 tibia width (
<figureCitation id="EBB8FEA01813FF8D432F30F1ABE5EB79" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Figs. 64 B, C</figureCitation>
),
<collectingRegion id="B1472CC71813FF8D47A83311AA2DEB79" box="[231,282,542,565]" country="United States of America" name="Utah" pageId="87" pageNumber="88">UTA</collectingRegion>
small; tibia with slender prolateral and a stout retrolateral spine (
<figureCitation id="EBB8FEA01813FF8D44913311AF7EEB79" box="[990,1097,542,565]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Fig. 64 C</figureCitation>
); tegulum (
<figureCitation id="EBB8FEA01813FF8D47FD3331AA15EB19" box="[178,290,574,597]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Figs. 64 C</figureCitation>
,
<figureCitation id="EBB8FEA01813FF8D46633331AA56EB19" box="[300,353,574,597]" captionStart="FIGURE" captionStartId="155.[155,167,1242,1261]" captionTargetBox="[121,1173,216,1227]" captionTargetId="figure-16@155.[120,1173,216,1227]" captionTargetPageId="155" captionText="FIGURE 65. Uduba valiha, new species, Holotype male (CASENT9026394) from Forêt de Vevembe, left palpus, SEM. A. Tibia and tarsus, prolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, retroventral. F. Bulb, prolateral. Eb, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158719" httpUri="https://zenodo.org/record/13158719/files/figure.png" pageId="87" pageNumber="88">65 D</figureCitation>
) with longitudinal, retromedian ridge, ridge beginning near tegulum base and extending for 0.80 tegulum length; TA3 a pointed spine, mostly hidden behind TA2, TA2 small, rectangular, apex curled (
<figureCitation id="EBB8FEA01813FF8D46C33371AAC1EBD9" box="[396,502,638,661]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="87" pageNumber="88">Fig. 28 B</figureCitation>
); MA origin at 0.56 tegulum length, extends to tegulum apex, length (including apical processes) slightly greater than width, length 0.45 tegulum width, deeply bifid MA with prolateral lobe slender, with rounded apex, retrolateral lobe stout with apical hook (
<figureCitation id="EBB8FEA01813FF8D47CC33D0AA23EBBA" box="[131,276,735,758]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Figs. 64 A–C</figureCitation>
); tegulum with apical mound that partially covers prolateral lobe of MA (
<figureCitation id="EBB8FEA01813FF8D436D33D0AFBEEBBA" box="[1058,1161,735,758]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="87" pageNumber="88">Fig. 64 A</figureCitation>
); conductor fan entire.
<emphasis id="41F73E371813FF8D462833F0AA8DEA5A" bold="true" box="[359,442,767,790]" pageId="87" pageNumber="88">Female</emphasis>
(
<typeStatus id="AC385C871813FF8D468433F0A919EA5A" box="[459,558,767,790]" pageId="87" pageNumber="88" type="paratype">paratype</typeStatus>
): Total length 10.00. Markings as in
<figureCitation id="EBB8FEA01813FF8D448933F0AF12EA5A" box="[966,1061,767,790]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: A–E = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="87" pageNumber="88">Fig. 6 C</figureCitation>
. Carapace 4.79 long, 3.29 wide, 2.14 high; clypeus 0.21 high. Eye diameters: AME 0.18, ALE and PME 0.25, PLE 0.21. Chelicerae 2.43 long; sternum 2.18 long, 1.61 wide; labium 1.07 long; palpal coxae 1.61 long.
<emphasis id="41F73E371813FF8D47F8326FAA37EA3B" bold="true" box="[183,256,864,887]" pageId="87" pageNumber="88">Ratios</emphasis>
–carapace length / width = 1.46, carapace height / width = 0.65, PER / carapace width = 0.67, PER/OAL = 2.67, PER/AER = 1.31, OAL/OQL = 1.00, OQP/ OQA = 1.24, clypeus height / AME = 1.20, cheliceral length / clypeus height = 11.33, sternum length / width = 1.36, palpal coxa length / width = 2.25, femur I length / carapace width =1.00, metatarsus I length / carapace width = 0.76, femur IV length / carapace width = 1.00, palpal tarsus length / carapace width = 0.37, palpal tibia length / palpal patella length = 1.42, palpal tarsus length / palpal tibia length = 1.42, palpal tarsus length / palpal femur length = 0.81, palpal tibia length / palpal patella length = 1.00. Divided cribellum. Spination: palpus–femur d0-0-1-1, r0-0-1, tibia p2-1, r1-0, tarsus p2-1; leg I– femur d0-1-1-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II–femur d0-1-1-0, p0-0-0-1, tibia v1-1-1-2, metatarsus v2-2-3; leg III–femur d0-1-0-0, p0-1-0-1, r0-1-0-1, tibia d1-1-0-0, p0-1-1-0, v2-1-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d0-1-1-0, tibia v1- 2-2, r0-1-1-0, metatarsus p0-1-2, v0-2-2, r1-1-2, tarsus v0-1-0. Scopulae: ventral on metatarsus– tarsus I and II, ventral on apical metatarsus–tarsus III and IV. Leg measurements: I: 3.29 + 1.57 + 2.43 + 2.50 + 1.50 = 11.29; II: 2.79 + 1.43 + 2.07 + 1.86 + 1.36 = 9.50; III: 2.43 + 1.07 + 1.43 + 1.79 + 1.07 = 7.79; IV: 3.29 + 1.50 + 2.57 + 3.21 + 1.43 = 12.00; palpus: 1.50 + 0.86 + 0.86 + NA + 1.21 = 4.43. Leg formula 4123.
<emphasis id="41F73E371813FF8D4554344CA9E0EC16" bold="true" box="[539,727,1347,1370]" pageId="87" pageNumber="88">Female genitalia</emphasis>
: epigynal plate without depressed atrium (
<figureCitation id="EBB8FEA01813FF8D47CC346BABCEEC37" box="[131,249,1380,1403]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="87" pageNumber="88">Figs. 72 G</figureCitation>
,
<figureCitation id="EBB8FEA01813FF8D464B346BAA0AEC37" box="[260,317,1380,1403]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="87" pageNumber="88">80 H</figureCitation>
), plate width 1.43 times length, lateral lobes with convex edges forming ridges on each side, these convex, these ridges extending around anteriorly and posteriorly of CO, width between bases of LL 0.88 times epigynum width, LL side width 0.14 times width epigynum; medi- an lobe a flat plate, copulatory openings at mid-level of plate at 0.41 of epigynum length, distance between copulatory openings 0.58 times epigynum width. Vulva (
<figureCitation id="EBB8FEA01813FF8D447534EAA898ECB0" box="[826,943,1509,1532]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="87" pageNumber="88">Figs. 72 H</figureCitation>
,
<figureCitation id="EBB8FEA01813FF8D44F534EAA8D2ECB0" box="[954,997,1509,1532]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="87" pageNumber="88">80 I</figureCitation>
) with spermathecal ducts forming 2–3 loose, longitudinal loops, vulva length 0.88 times width, fertilization ducts very close, vulva width 16.50 times distance between FD.
</paragraph>
<paragraph id="733CE2251813FF8247E4374AA883E8FF" blockId="87.[123,1174,220,1725]" lastBlockId="88.[123,1174,220,885]" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
<emphasis id="41F73E371813FF8D47E4374AAA2DEF10" bold="true" box="[171,282,1605,1628]" pageId="87" pageNumber="88">Variation.</emphasis>
Male (N= 5): Total length 8.50–11.00; carapace length / width = 1.38–1.68, carapace height / width = 0.49–0.55, PER / carapace width = 0.34–0.38, PER/OAL = 2.19–2.60, PER/AER = 1.24–1.51, OAL/OQL = 1.13–1.37, OQP/ OQA = 1.19–1.39, clypeus height / AME = 1.11–1.67, cheliceral length / clypeus height = 7.00–10.50, sternum length / width = 1.29–1.41, palpal coxa length / width = 2.33–2.80, femur I length / carapace width = 1.16–1.35, metatarsus I length / carapace width = 1.11–1.39, femur IV length / carapace width = 1.18–1.48, cymbium length / carapace width = 0.60–0.70, cymbium length / palpal patella length = 2.86–3.83, cymbium length / palpal tibia length = 2.33–2.80, cymbium length / palpal femur length = 1.05–1.28, palpal tibia length / palpal patella length = 1.14–1.50. Compared to specimens from Vinetelo, specimens from Vevembe have the legs slightly relatively longer and the cymbium slightly shorter. Female variation unknown: only
<specimenCount id="658529AC181CFF8246AD3093A96BE8FF" box="[482,604,412,435]" count="1" pageId="88" pageNumber="89" type="female">one female</specimenCount>
, a
<typeStatus id="AC385C87181CFF8245333093A9ECE8FF" box="[636,731,412,435]" pageId="88" pageNumber="89" type="paratype">paratype</typeStatus>
, has been collected.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8247E430B3A9FBEA79" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="733CE225181CFF8247E430B3A858EBF8" blockId="88.[123,1174,220,885]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E430B3AABBE89F" bold="true" box="[171,396,444,467]" pageId="88" pageNumber="89">Material examined.</emphasis>
<materialsCitation id="C3EBE878181CFF8246DB30B3A84AEB78" collectingDate="2006-04-23" collectionCode="CAS" collectorName="de Vevembe &amp; B. L. Fisher" country="Madagascar" elevation="600" latitude="-22.791111" location="Foret de Vevembe" longLatPrecision="21" longitude="47.181946" pageId="88" pageNumber="89" specimenCode="CASENT9026394, CASENT9082181" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectingCountry id="0B94A2B5181CFF8246DB30B3A961E89F" box="[404,598,444,467]" name="Madagascar" pageId="88" pageNumber="89">MADAGASCAR</collectingCountry>
:
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<collectingRegion id="B1472CC7181CFF82452D30B3A86DE89F" box="[610,858,444,467]" country="Madagascar" name="Fianarantsoa" pageId="88" pageNumber="89">Fianarantsoa Province</collectingRegion>
</emphasis>
:
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Forêt
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</location>
,
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293°WNW Farafangana
</locationDeviation>
,
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,
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, elev.
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</quantity>
,
<collectingMethod id="AAC29A32181CFF82446930D3A8F8E8BF" box="[806,975,476,499]" pageId="88" pageNumber="89">yellow pan trap</collectingMethod>
, rainforest in transition to montane forest,
<date id="073DC4E5181CFF82468E30F2A945EB58" box="[449,626,509,532]" pageId="88" pageNumber="89" value="2006-04-23">
<collectingDate id="17793D0D181CFF82468E30F2A945EB58" box="[449,626,509,532]" pageId="88" pageNumber="89" value="2006-04-23">23 April 2006</collectingDate>
</date>
,
<collectorName id="DE7687F3181CFF8245C330F2A820EB58" box="[652,791,509,532]" pageId="88" pageNumber="89">B.L. Fisher</collectorName>
<emphasis id="41F73E37181CFF82446630F2A852EB58" box="[809,869,509,532]" italics="true" pageId="88" pageNumber="89">et al</emphasis>
. [BLF14119] (
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,
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,
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,
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), (
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,
<specimenCode id="23254A5E181CFF8245743312A834EB78" box="[571,771,541,564]" collectionCode="CASENT" pageId="88" pageNumber="89">CASENT9082181</specimenCode>
,
<specimenCount id="658529AC181CFF8244413312A807EB78" box="[782,816,541,564]" count="1" pageId="88" pageNumber="89" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0181CFF8244713312A842EB78" box="[830,885,541,564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="88" pageNumber="89" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE878181CFF8244C63312AF63EB38" collectionCode="FMNH" collectorName="de Vinetalo &amp; S. Goodman" country="Madagascar" elevation="100" location="Ambodivohita" municipality="Foret de Vinetalo" pageId="88" pageNumber="89" specimenCode="CASENT9064656" specimenCount="3" specimenCount-male="3" stateProvince="Fianarantsoa Province" typeStatus="holotype">
<collectingMunicipality id="9358785F181CFF8244C63312AF72EB78" box="[905,1093,541,564]" pageId="88" pageNumber="89">
Forêt
<collectorName id="DE7687F3181CFF8244863312AF72EB78" box="[969,1093,541,564]" pageId="88" pageNumber="89">de Vinetalo</collectorName>
</collectingMunicipality>
, at foot of
<location id="765CB4FE181CFF8247D83332AA03EB18" LSID="urn:lsid:plazi:treatment:FB2A53331812FF82473430B3AAC0EA39:765CB4FE181CFF8247D83332AA03EB18" box="[151,308,573,596]" country="Madagascar" municipality="Foret de Vinetalo" name="Ambodivohita" pageId="88" pageNumber="89" stateProvince="Fianarantsoa Province">Ambodivohita</location>
, 21°46.6ʹS, 47°20.8ʹE, elev.
<quantity id="B47B4FC0181CFF8245223332A99BEB18" box="[621,684,573,596]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">
<elevation id="F8AE0516181CFF8245223332A99BEB18" box="[621,684,573,596]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">100m</elevation>
</quantity>
, disturbed lowland rainforest in transition to montane forest,
<collectingMethod id="AAC29A32181CFF8246683352AA94EB38" box="[295,419,605,628]" pageId="88" pageNumber="89">pitfall traps</collectingMethod>
,
<collectorName id="DE7687F3181CFF8246E23352A905EB38" box="[429,562,605,628]" pageId="88" pageNumber="89">S. Goodman</collectorName>
[FMHD-00-223] (
<specimenCode id="23254A5E181CFF8245B63352A8F4EB38" box="[761,963,605,628]" collectionCode="CASENT" pageId="88" pageNumber="89">CASENT9064656</specimenCode>
,
<specimenCount id="658529AC181CFF8244823352A8D8EB38" box="[973,1007,605,628]" count="3" pageId="88" pageNumber="89" type="male">3♂</specimenCount>
,
<collectionCode id="15927AE0181CFF8244B33352AF7AEB38" box="[1020,1101,605,628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="88" pageNumber="89" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE878181CFF8243103352A85CEBF8" collectingDate="1993-09-24" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="785" location="Massif Andringitra" pageId="88" pageNumber="89" specimenCode="CASENT9006071" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa Province" typeStatus="paratype">
<location id="765CB4FE181CFF8243103352AA17EBD8" LSID="urn:lsid:plazi:treatment:FB2A53331812FF82473430B3AAC0EA39:765CB4FE181CFF8243103352AA17EBD8" country="Madagascar" name="Massif Andringitra" pageId="88" pageNumber="89" stateProvince="Fianarantsoa Province">Massif Andringitra</location>
,
<locationDeviation id="628EFB07181CFF82467D3372A924EBD8" box="[306,531,637,660]" location="Ambalavo" pageId="88" pageNumber="89">
<quantity id="B47B4FC0181CFF82467D3372AA4FEBD8" box="[306,376,637,660]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="88" pageNumber="89" unit="km" value="45.0">45 km</quantity>
S Ambalavo
</locationDeviation>
, 22°13ʹS, 47°1ʹE, elev.
<quantity id="B47B4FC0181CFF8244693372A852EBD8" box="[806,869,637,660]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="88" pageNumber="89" unit="m" value="785.0">
<elevation id="F8AE0516181CFF8244693372A852EBD8" box="[806,869,637,660]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="88" pageNumber="89" unit="m" value="785.0">785m</elevation>
</quantity>
, rainforest,
<collectingMethod id="AAC29A32181CFF8244BC3372AF5AEBD8" box="[1011,1133,637,660]" pageId="88" pageNumber="89">pitfall trap</collectingMethod>
,
<date id="073DC4E5181CFF8243343372AA1AEBF8" pageId="88" pageNumber="89" value="1993-09-24">
<collectingDate id="17793D0D181CFF8243343372AA1AEBF8" pageId="88" pageNumber="89" value="1993-09-24">24 September 1993</collectingDate>
</date>
,
<collectorName id="DE7687F3181CFF8246763392AA84EBF8" box="[313,435,669,692]" pageId="88" pageNumber="89">B.L. Fisher</collectorName>
(
<typeStatus id="AC385C87181CFF82468D3392A917EBF8" box="[450,544,669,692]" pageId="88" pageNumber="89" type="paratype">paratype</typeStatus>
<specimenCode id="23254A5E181CFF8245683392A9C7EBF8" box="[551,752,669,692]" collectionCode="CASENT" pageId="88" pageNumber="89">CASENT9006071</specimenCode>
,
<specimenCount id="658529AC181CFF8245B43392A82AEBF8" box="[763,797,669,692]" count="1" pageId="88" pageNumber="89" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0181CFF8244643392A854EBF8" box="[811,867,669,692]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="88" pageNumber="89" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225181CFF8247E433B2A9FBEA79" blockId="88.[123,1174,220,885]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E433B2AA57EB98" bold="true" box="[171,352,701,724]" pageId="88" pageNumber="89">Natural history.</emphasis>
All specimens of
<taxonomicName id="B48399A6181CFF82456633B2A9F7EB98" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[553,704,701,724]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E37181CFF82456633B2A9F7EB98" bold="true" box="[553,704,701,724]" italics="true" pageId="88" pageNumber="89">Uduba valiha</emphasis>
</taxonomicName>
have been collected in rainforest at low to mid elevation, from
<quantity id="B47B4FC0181CFF82461733D2AAA2EBB8" box="[344,405,733,756]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">100m</quantity>
to
<quantity id="B47B4FC0181CFF8246F633D2AACEEBB8" box="[441,505,733,756]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="88" pageNumber="89" unit="m" value="785.0">785m</quantity>
, and all were collected in pitfall or yellow pan traps, suggesting that males and females of this cribellate species both wander at some point in their lives. We know nothing of the use of cribellate silk in
<emphasis id="41F73E37181CFF8245113211A9FBEA79" bold="true" box="[606,716,798,821]" italics="true" pageId="88" pageNumber="89">
<taxonomicName id="B48399A6181CFF8245113211A9F3EA79" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[606,708,798,821]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="valiha">U. valiha</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8247E43231AAC0EA39" pageId="88" pageNumber="89" type="distribution">
<paragraph id="733CE225181CFF8247E43231AAC0EA39" blockId="88.[123,1174,220,885]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E43231AA0AEA19" bold="true" box="[171,317,830,853]" pageId="88" pageNumber="89">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6181CFF8246893231A96CEA19" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[454,603,830,853]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E37181CFF8246893231A96CEA19" bold="true" box="[454,603,830,853]" italics="true" pageId="88" pageNumber="89">Uduba valiha</emphasis>
</taxonomicName>
is known only from three forested localities in southeastern
<collectingCountry id="0B94A2B5181CFF8247813251AA78EA39" box="[206,335,862,885]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
(Maps 10, 17).
</paragraph>
</subSubSection>
</treatment>
</document>

262
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@ -0,0 +1,262 @@
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<mods:title id="F0F40DA99B111986E4AEDB8B786D0869">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="F24CFD98E9654797C7638B3B0A801F9A">Griswold, Charles</mods:namePart>
<mods:affiliation id="46C02EB72D5646E74441A0E26FB22BC1">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="0E7FE48B6609C8966A6B9DE07036F5A6">Ubick, Darrell</mods:namePart>
<mods:affiliation id="694FFDF5CED4D78640A2ACE8B342D738">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="786BBCFF1E052A07F0D692ABFE7CC86A">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="7BBB9BAA10FA63F4C6AAC43246B1D6CA">Polotow, Daniele</mods:namePart>
<mods:affiliation id="ECC74306DF0ADC5A664FFF58DD76EF20">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<paragraph id="733CE2251814FF8A47343251AA9AEA39" blockId="80.[123,429,862,915]" box="[123,429,862,885]" pageId="80" pageNumber="81">
<heading id="287455491814FF8A47343251AA9AEA39" box="[123,429,862,885]" fontSize="10" level="2" pageId="80" pageNumber="81" reason="4">
<emphasis id="41F73E371814FF8A47343251AA9AEA39" bold="true" box="[123,429,862,885]" pageId="80" pageNumber="81">
<taxonomicName id="B48399A61814FF8A47343251AA2CEA39" ID-CoL="C8KK3" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,283,862,885]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sarotra" status="sp. nov.">
<emphasis id="41F73E371814FF8A47343251AA2CEA39" bold="true" box="[123,283,862,885]" italics="true" pageId="80" pageNumber="81">Uduba sarotra</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1814FF8A46663251AA9AEA39" box="[297,429,862,885]" pageId="80" pageNumber="81" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8A47343271AA97EADF" box="[123,416,894,915]" pageId="80" pageNumber="81" type="description">
<paragraph id="733CE2251814FF8A47343271AA97EADF" blockId="80.[123,429,862,915]" box="[123,416,894,915]" pageId="80" pageNumber="81">
<figureCitation id="EBB8FEA01814FF8A47343271AA20EADF" box="[123,279,894,915]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="80" pageNumber="81">Figures 70 D, E</figureCitation>
, Maps 10, 21.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8A47E432ACAAA7ED57" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="733CE2251814FF8A47E432ACAAA7ED57" blockId="80.[122,1174,931,1725]" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E432ACAA51EAF6" bold="true" box="[171,358,931,954]" pageId="80" pageNumber="81">Type material.</emphasis>
<materialsCitation id="C3EBE8781814FF8A463732ACAAA7ED57" collectingDate="2003-11-18" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1325" location="Parc National de Marojejy" pageId="80" pageNumber="81" specimenCode="CASENT9030916, CASENT9064719, CASENT9064720" specimenCount="2" specimenCount-female="2" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="AC385C871814FF8A463732ACAAD4EAF6" box="[376,483,931,954]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
female (
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) and
<specimenCount id="658529AC1814FF8A44C632ACAFA2EAF6" box="[905,1173,931,954]" count="2" pageId="80" pageNumber="81" type="female" typeStatus="paratype">two paratype females</specimenCount>
(
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and
<specimenCode id="23254A5E1814FF8A46CD32CCA97EEA96" box="[386,585,963,986]" collectionCode="CASENT" pageId="80" pageNumber="81">CASENT9064720</specimenCode>
) from montane rainforest at
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<elevation id="F8AE05161814FF8A44C632CCA8E4EA96" box="[905,979,963,986]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="80" pageNumber="81" unit="m" value="1325.0">1325m</elevation>
</quantity>
elevation in
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,
<collectingRegion id="B1472CC71814FF8A462232EBA96EEAB7" box="[365,601,996,1019]" country="Madagascar" name="Antsiranana" pageId="80" pageNumber="81">Antsiranana Province</collectingRegion>
,
<collectingCountry id="0B94A2B51814FF8A452A32EBA9D3EAB7" box="[613,740,996,1019]" name="Madagascar" pageId="80" pageNumber="81">Madagascar</collectingCountry>
, collected
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<collectingDate id="17793D0D1814FF8A441232EBAF07EAB7" box="[861,1072,996,1019]" pageId="80" pageNumber="81" value="2003-11-18">18 November 2003</collectingDate>
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by
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, deposited in
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</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8A47E4352BABC6ED17" pageId="80" pageNumber="81" type="etymology">
<paragraph id="733CE2251814FF8A47E4352BABC6ED17" blockId="80.[122,1174,931,1725]" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E4352BAA1DED77" bold="true" box="[171,298,1060,1083]" pageId="80" pageNumber="81">Etymology.</emphasis>
The species epithet is from a Malagasy word for difficult,
<emphasis id="41F73E371814FF8A44D0352BA8DBED77" box="[927,1004,1060,1083]" italics="true" pageId="80" pageNumber="81">sarotra</emphasis>
; an adjective in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8A47E4356BA8C6EDB7" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="733CE2251814FF8A47E4356BA8C6EDB7" blockId="80.[122,1174,931,1725]" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E4356BAAFCED37" bold="true" box="[171,459,1124,1147]" pageId="80" pageNumber="81">
Diagnosis.
<taxonomicName id="B48399A61814FF8A4667356BAAFCED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,459,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sarotra">
<emphasis id="41F73E371814FF8A4667356BAAFCED37" bold="true" box="[296,459,1124,1147]" italics="true" pageId="80" pageNumber="81">Uduba sarotra</emphasis>
</taxonomicName>
</emphasis>
are an Unclassified species (
<figureCitation id="EBB8FEA01814FF8A4443356BA854ED37" box="[780,867,1124,1147]" captionStart="MAP" captionStartId="186.[421,440,1168,1187]" captionTargetBox="[388,916,216,1150]" captionTargetId="figure-11@186.[388,916,216,1150]" captionTargetPageId="186" captionText="MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863)." figureDoi="http://doi.org/10.5281/zenodo.13158799" httpUri="https://zenodo.org/record/13158799/files/figure.png" pageId="80" pageNumber="81">Map 21</figureCitation>
) and females can be distinguished from those of other
<taxonomicName id="B48399A61814FF8A46EB358BAAD9EDD7" box="[420,494,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371814FF8A46EB358BAAD9EDD7" bold="true" box="[420,494,1156,1179]" italics="true" pageId="80" pageNumber="81">Uduba</emphasis>
</taxonomicName>
that have a divided cribellum by the form of the epigynum (
<figureCitation id="EBB8FEA01814FF8A4325358BAB83EDF7" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="80" pageNumber="81">Fig. 70 D</figureCitation>
): the epigynal plate lacks sclerotized lateral margin, not projecting laterally, ML short, limit- ed to anterior half of epigynal plate, CO hidden beneath ML, spermathecae path, if visible through epigynal cuticle, seen only posteriad of copulatory openings. Males are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1814FF8B47E4340BAA7BEA1A" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="description">
<paragraph id="733CE2251814FF8B47E4340BAF72E87F" blockId="80.[122,1174,931,1725]" lastBlockId="81.[123,1174,220,918]" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
<emphasis id="41F73E371814FF8A47E4340BAA4BEC57" bold="true" box="[171,380,1284,1307]" pageId="80" pageNumber="81">Description. Male</emphasis>
Unknown.
<emphasis id="41F73E371814FF8A454B340BA960EC57" bold="true" box="[516,599,1284,1307]" pageId="80" pageNumber="81">Female</emphasis>
(
<typeStatus id="AC385C871814FF8A4525340BA9E6EC57" box="[618,721,1284,1307]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
, CASENT9030916): Total length 10.50. Markings typical of
<emphasis id="41F73E371814FF8A4611342AAA98EC70" bold="true" box="[350,431,1317,1340]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="B48399A61814FF8A4611342AAA9DEC70" box="[350,426,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">Uduba</taxonomicName>
.
</emphasis>
Carapace 3.80 long, 2.80 wide, 1.40 high; clypeus 0.20 high. Eye diameters: AME 0.13, ALE 0.20, PME 0.18, PLE 0.22. Chelicerae 2.00 long; sternum 1.90 long, 1.55 wide; labium 0.80 long; palpal coxae 1.45 long. Divided cribellum. Spination: palpus–femur d0-1-1, patella p1-0, tibia p2-0, tarsus p2-1-0, v0-0-1; leg I–femur d0-1-0, p0-0-1, tibia v2-2-2-2, metatarsus v2-2-3; leg II–femur d0-1-0, tibia r0-1-0, metatarsus v2-2-3; leg III–femur d1-0-0, tibia d0-1-0, p0-1-1-0, v0-1-1-0, r0-1-1-0, metatarsus p1-2-2, v2-1-2, r1-1-2; leg IV–femur d1-0-0, tibia v1-1-2, r0-0-1, metatarsus p0-0-2, v0-2-2-2, r0-0-2. Scopulae: strong ventral on tarsi I-IV, metatarsus I, apex of metatarsi II and IV. Leg measurements: I: 2.80 + 1.40 + 2.40 + 2.20 + 1.60 = 10.40; II: 2.50 + 1.20 + 1.90 + 1.80 + 1.30 = 8.70; III: 2.00 + 1.10 + 1.10 + 1.50 + 1.10 = 6.80; IV: 2.80 + 1.30 + 2.40 + 2.60 + 1.40 = 10.50; palpus: 1.40 + 0.70 + 0.70 + NA + 1.30 = 4.10. Leg formula 4123.
<emphasis id="41F73E371814FF8A47973769AAA3EF31" bold="true" box="[216,404,1638,1661]" pageId="80" pageNumber="81">Female genitalia</emphasis>
: epigynal plate without depressed atrium, posterior margin at epigastric furrow convex (
<figureCitation id="EBB8FEA01814FF8A46633789AAADEFD1" box="[300,410,1670,1693]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="80" pageNumber="81">Fig. 70 D</figureCitation>
), plate width 1.37 times length, ML and LL not clearly defined, CO beneath longitudinal grooves in anterior half of plate, distance between CO 0.41 times epigynum width, CO origin at 0.44 of epigynum length, extant of plate laterad to CO equals 0.19 times plate width. Vulva (
<figureCitation id="EBB8FEA01815FF8B465F31F3AA43E85F" box="[272,372,252,275]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="81" pageNumber="82">Fig. 70 E</figureCitation>
) with spermathecal ducts forming two-and-a-half longitudinal loops, vulva length 0.67 times width, FD widely spaced, vulva width 3.86 times distance between FD.
</paragraph>
<paragraph id="733CE2251815FF8B47E43033AAF5EB39" blockId="81.[123,1174,220,918]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E43033AA2DE81F" bold="true" box="[171,282,316,339]" pageId="81" pageNumber="82">Variation.</emphasis>
Female (N= 3): Total length = 7.30–11.10. Carapace length / width = 1.36–1.50, carapace height / width = 0.50–0.73, PER / carapace width = 0.46–0.48, PER/OAL = 3.00–3.10, PER/AER = 1.17–1.40, OAL/OQL = 1.00–1.18, OQP/ OQA = 1.23–1.39, clypeus height / AME = 1.54–1.67, cheliceral length / clypeus height = 9.00–10.00, sternum length / width = 1.23–1.35, palpal coxa length / width = 2.00–2.17, femur I length / carapace width = 0.92–1.00, metatarsus I length / carapace width = 0.79–0.88, femur IV length / carapace width = 1.00–1.09, palpal tarsus length / carapace width = 0.46–0.53, palpal tibia length / palpal patella length = 1.70–2.17, palpal tarsus length / palpal tibia length = 1.86–2.17, palpal tarsus length / palpal femur length = 0.93– 1.00, palpal tibia length / palpal patella length = 0.90–1.00. Male variation unknown;
<taxonomicName id="B48399A61815FF8B436B3331AFA1EB19" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1060,1174,574,597]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sarotra">
<emphasis id="41F73E371815FF8B436B3331AFA1EB19" bold="true" box="[1060,1174,574,597]" italics="true" pageId="81" pageNumber="82">U. sarotra</emphasis>
</taxonomicName>
are known only from females.
</paragraph>
<paragraph id="733CE2251815FF8B47E43371A849EA5A" blockId="81.[123,1174,220,918]" pageId="81" pageNumber="82">
<materialsCitation id="C3EBE8781815FF8B47E43371A84DEA5A" collectingDate="2003-11-18" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1325" latitude="-14.443334" location="Antranohofa" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy" pageId="81" pageNumber="82" specimenCode="CASENT9030916, CASENT9064719, CASENT9064720" specimenCount="3" specimenCount-female="3" stateProvince="Antsiranana" typeStatus="holotype">
M
<emphasis id="41F73E371815FF8B478D3371AABAEBD9" bold="true" box="[194,397,638,661]" pageId="81" pageNumber="82">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51815FF8B46D73371A96DEBD9" box="[408,602,638,661]" name="Madagascar" pageId="81" pageNumber="82">MADAGASCAR</collectingCountry>
:
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<collectingRegion id="B1472CC71815FF8B45263371A861EBD9" box="[617,854,638,661]" country="Madagascar" name="Antsiranana" pageId="81" pageNumber="82">Antsiranana Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1815FF8B44263371AFB9EBD9" box="[873,1166,638,661]" pageId="81" pageNumber="82">Parc National de Marojejy</collectingMunicipality>
,
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,
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<quantity id="B47B4FC01815FF8B46503391AAB3EBF9" box="[287,388,670,693]" metricMagnitude="4" metricUnit="m" metricValue="2.66" pageId="81" pageNumber="82" unit="km" value="26.6">26.6 km</quantity>
31°NNE Andapa
</locationDeviation>
,
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318° NW Manantenina
</locationDeviation>
,
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,
<geoCoordinate id="16B784E21815FF8B473433B1AA32EB99" box="[123,261,702,725]" degrees="049" direction="east" minutes="44" orientation="longitude" pageId="81" pageNumber="82" precision="15" seconds="36" value="49.743332">049°44ʹ36ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01815FF8B460033B1AAABEB99" box="[335,412,702,725]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="81" pageNumber="82" unit="m" value="1325.0">
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</quantity>
, general collecting day spiders in montane rainforest,
<date id="073DC4E51815FF8B44B033B1AB84EBB9" pageId="81" pageNumber="82" value="2003-11-18">
<collectingDate id="17793D0D1815FF8B44B033B1AB84EBB9" pageId="81" pageNumber="82" value="2003-11-18">18 November 2003</collectingDate>
</date>
,
<collectorName id="DE7687F31815FF8B47F233D1AA0EEBB9" box="[189,313,734,757]" pageId="81" pageNumber="82">B. L. Fisher</collectorName>
, [BLF9082] (
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<emphasis id="41F73E371815FF8B468333D1A95CEBB9" bold="true" box="[460,619,734,757]" italics="true" pageId="81" pageNumber="82">Uduba sarotra</emphasis>
</taxonomicName>
<typeStatus id="AC385C871815FF8B453E33D1A9E0EBB9" box="[625,727,734,757]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1815FF8B45AE33D1A89EEBB9" box="[737,937,734,757]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9030916</specimenCode>
,
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,
<collectionCode id="15927AE01815FF8B44AE33D1AF2FEBB9" box="[993,1048,734,757]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
), (
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,
<specimenCode id="23254A5E1815FF8B473433F0AA74EA5A" box="[123,323,767,790]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9064719</specimenCode>
,
<specimenCount id="658529AC1815FF8B460133F0AA47EA5A" box="[334,368,767,790]" count="1" pageId="81" pageNumber="82" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01815FF8B463133F0AA82EA5A" box="[382,437,767,790]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C871815FF8B469E33F0A918EA5A" box="[465,559,767,790]" pageId="81" pageNumber="82" type="paratype">paratype</typeStatus>
<specimenCode id="23254A5E1815FF8B457933F0A9C9EA5A" box="[566,766,767,790]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9064720</specimenCode>
,
<specimenCount id="658529AC1815FF8B444533F0A81CEA5A" box="[778,811,767,790]" count="1" pageId="81" pageNumber="82" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01815FF8B447533F0A845EA5A" box="[826,882,767,790]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251815FF8B47E43210AA7BEA1A" blockId="81.[123,1174,220,918]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E43210AA56EA7A" bold="true" box="[171,353,799,822]" pageId="81" pageNumber="82">Natural history.</emphasis>
Collection records suggest that the cribellate
<taxonomicName id="B48399A61815FF8B44163210A8CCEA7A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[857,1019,799,822]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sarotra">
<emphasis id="41F73E371815FF8B44163210A8CCEA7A" bold="true" box="[857,1019,799,822]" italics="true" pageId="81" pageNumber="82">Uduba sarotra</emphasis>
</taxonomicName>
occur only in montane rainforest.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1815FF8B47E43250A8CBEADA" pageId="81" pageNumber="82" type="distribution">
<paragraph id="733CE2251815FF8B47E43250A8CBEADA" blockId="81.[123,1174,220,918]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E43250AA0AEA3A" bold="true" box="[171,317,863,886]" pageId="81" pageNumber="82">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61815FF8B46893250A951EA3A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[454,614,863,886]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sarotra">
<emphasis id="41F73E371815FF8B46893250A951EA3A" bold="true" box="[454,614,863,886]" italics="true" pageId="81" pageNumber="82">Uduba sarotra</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C871815FF8B44253250A8AFEA3A" box="[874,920,863,886]" pageId="81" pageNumber="82">type</typeStatus>
locality at Parc National de Marojejy in
<collectingRegion id="B1472CC71815FF8B46743270A915EADA" box="[315,546,895,918]" country="Madagascar" name="Antsiranana" pageId="81" pageNumber="82">Antsiranana Province</collectingRegion>
in northeastern
<collectingCountry id="0B94A2B51815FF8B459C3270A863EADA" box="[723,852,895,918]" name="Madagascar" pageId="81" pageNumber="82">Madagascar</collectingCountry>
(Maps 10, 21).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4C6D9AC26A170F81DF3E3019C8BC6CB3">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:identifier id="FA181E364DEF0CCA6727FE182B46528D" type="DOI">10.5281/zenodo.13158554</mods:identifier>
<mods:identifier id="D6AD6A7B47E1DEE85A346DFADE92B251" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="80B64B08FBDDDDA28E574590342278F9" type="Zenodo-Dep">13158554</mods:identifier>
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<subSubSection id="3B99B1AE1815FF8B473432CFAAFBEA9B" box="[123,460,960,983]" pageId="81" pageNumber="82" type="nomenclature">
<paragraph id="733CE2251815FF8B473432CFAAFBEA9B" blockId="81.[123,1174,960,1042]" box="[123,460,960,983]" pageId="81" pageNumber="82">
<heading id="287455491815FF8B473432CFAAFBEA9B" box="[123,460,960,983]" fontSize="10" level="2" pageId="81" pageNumber="82" reason="4">
<emphasis id="41F73E371815FF8B473432CFAAFBEA9B" bold="true" box="[123,460,960,983]" pageId="81" pageNumber="82">
<taxonomicName id="B48399A61815FF8B473432CFAA0DEA9B" ID-CoL="C8KK4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,314,960,983]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="schlingeri" status="sp. nov.">
<emphasis id="41F73E371815FF8B473432CFAA0DEA9B" bold="true" box="[123,314,960,983]" italics="true" pageId="81" pageNumber="82">Uduba schlingeri</emphasis>
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<taxonomicNameLabel id="5AC4834C1815FF8B460732CFAAFBEA9B" box="[328,460,960,983]" pageId="81" pageNumber="82" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="3B99B1AE1815FF8B473432EFAF29ED5E" pageId="81" pageNumber="82" type="description">
<paragraph id="733CE2251815FF8B473432EFAF29ED5E" blockId="81.[123,1174,960,1042]" pageId="81" pageNumber="82">
<figureCitation id="EBB8FEA01815FF8B473432EFAA27EAB9" box="[123,272,992,1013]" captionStart="FIGURE" captionStartId="106.[155,167,815,834]" captionTargetBox="[120,1176,216,796]" captionTargetId="figure-11@106.[120,1176,216,796]" captionTargetPageId="106" captionText="FIGURE 2. Uduba spp., nests and live individuals. A, C, U. schlingeri, ♀, CASENT9006058. B, D, U. evanescens (Dahl, 1901), ♀, CASENT9006088. A. Tubular nest surmounted by funnel of cribellate silk, Ranomafana. B. Nest opening on soil surface, Ranomafana. C. Tubular nest, Ranomafana. D. Nest opening on soil surface, Ranomafana. E. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. A–D. Photos by Dong Lin, Ranomafana, April 1998. E. Photo by Nikolaj Scharff, November 1993. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13158560" httpUri="https://zenodo.org/record/13158560/files/figure.png" pageId="81" pageNumber="82">Figures. 2 A, C</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B465432EFAA6AEAB9" box="[283,349,992,1013]" captionStart="FIGURE" captionStartId="107.[155,167,1531,1550]" captionTargetBox="[120,1176,216,1515]" captionTargetId="figure-16@107.[120,1176,216,1515]" captionTargetPageId="107" captionText="FIGURE 3. Habitus photos of Uduba schlingeri from Parc National Ranomafana. A, B, D, G. Female ♀ (CASENT9006075). C, E, F, H. ♂ Holotype male (CASENT9006052). A. Dorsal. B. Ventral. C. Dorsal. D. Carapace, dorsal. E. Ventral. F. Carapace, dorsal. G. Carapace, lateral. H. Carapace, lateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158562" httpUri="https://zenodo.org/record/13158562/files/figure.png" pageId="81" pageNumber="82">3 A–H</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B462832EFAABBEAB9" box="[359,396,992,1013]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="81" pageNumber="82">7 G</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B46D832EFAA8CEAB9" box="[407,443,992,1013]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="81" pageNumber="82">9 A</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B468932EFAAC0EAB9" box="[454,503,992,1013]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="81" pageNumber="82">11 G</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B454E32EFA967EAB9" box="[513,592,992,1013]" captionStart="FIGURE" captionStartId="116.[155,167,925,944]" captionTargetBox="[120,1176,216,925]" captionTargetId="figure-11@116.[120,1176,216,910]" captionTargetPageId="116" captionText="FIGURE 12. Uduba spp., morphology photos. A–C. U. schlingeri, ♂ CASENT9006052 from Talatakely, Ranomafana National Park. D, F. U. schlingeri, ♀ CASENT9006075 from Vatoharanana, Ranomafana National Park. E. U. fisheri, ♀ CASENT9026389 from Anjozorobe. A. Cephalothorax, dorsal. B. Cephalothorax, ventral. C. Habitus, lateral. D. Face, clypeus and divided chilum, anterior. E. Pedicel, dorsal (anterior to top). F. Habitus, lateral. Scale bars: A, B, F = 2 mm, C = 2 mm, D =1 mm, E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158586" httpUri="https://zenodo.org/record/13158586/files/figure.png" pageId="81" pageNumber="82">12 A–D</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B451532EFA995EAB9" box="[602,674,992,1013]" captionStart="FIGURE" captionStartId="116.[155,167,1638,1657]" captionTargetBox="[121,1176,1059,1621]" captionTargetId="figure-105@116.[120,1176,1059,1622]" captionTargetPageId="116" captionText="FIGURE 13. Uduba schlingeri, from Ranomafana National Park, leg photos. A–C, E, F. ♀ (CASENT9006075). D. ♂ (CASENT9006052). A. Leg I, prolateral. B. Metatarsus and tarsus IV, retrolateral, showing calamistrum-arrow to calamistrum. C. Leg IV, retrolateral. D. ♂ tibia II retrolateral, base showing tibial crack (arrow). E. Trochanter IV, left. F. Trochanter I, left. Scale bars: A, C = 2 mm, B = 1 mm, D, E, F = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13158588" httpUri="https://zenodo.org/record/13158588/files/figure.png" pageId="81" pageNumber="82">13 A–F</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B45E332EFA9CDEAB9" box="[684,762,992,1013]" captionStart="FIGURE" captionStartId="117.[155,167,913,932]" captionTargetBox="[120,1176,216,897]" captionTargetId="figure-16@117.[120,1176,216,897]" captionTargetPageId="117" captionText="FIGURE 14. Uduba spp., females, legs, SEM. A–E, G. U. schlingeri ♀ (CASENT9025468) from Vatoharanana, Ranomafana National Park. F, H. U. fisheri, ♀ (CASENT9026389) from Anjozorobe. A. Tarsus I, left, retrolateral. B. Tarsus IV, left, retrolateral. C. Tarsus I, left, dorsal. D. Tarsus IV, left, dorsal. E. Palpal tarsus, right, retrolateral. F. Palpus, trochanter–tarsus, prolateral. G. Metatarsus IV base, retrodorsal, showing calamistrum [arrow]. H. Metatarsus IV, retrodorsal, showing calamistrum [arrow]. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158590" httpUri="https://zenodo.org/record/13158590/files/figure.png" pageId="81" pageNumber="82">14 A–H</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B444B32EFA801EAB9" box="[772,822,992,1013]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="81" pageNumber="82">20 G</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B440E32EFA8B8EAB9" box="[833,911,992,1013]" captionStart="FIGURE" captionStartId="122.[155,167,1045,1064]" captionTargetBox="[120,1176,216,1027]" captionTargetId="figure-11@122.[120,1176,216,1027]" captionTargetPageId="122" captionText="FIGURE 21. Spinning organs of Uduba schlingeri, male CASENT9006052 from Talatakely, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Whole spinnerets. B. Left ALS, showing Nu of MAP. C. Apex of left PMS. D. Left PMS, showing Nu of MS and Fls triplet. AC, aciniform gland spigot(s); Fls, flankers of pseudoflagelliform gland spigot; MAP, major ampullate gland spigot(s); mAP, minor ampullate gland spigot(s); MS, modified spigot, or pseudoflagelliform gland spigot; Nu, nubbin; PI, piriform gland spigot(s); Tp, tartipore. Scale bars: A = 100 µm, B = 20 µm, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158612" httpUri="https://zenodo.org/record/13158612/files/figure.png" pageId="81" pageNumber="82">21 A–D</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B44D632EFA8D0EAB9" box="[921,999,992,1013]" captionStart="FIGURE" captionStartId="123.[155,167,1028,1047]" captionTargetBox="[120,1176,216,1011]" captionTargetId="figure-16@123.[120,1176,216,1011]" captionTargetPageId="123" captionText="FIGURE 22. Spinning organs of Uduba schlingeri, female CASENT9025468 from Vatoharanana, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Whole spinnerets. B. Right ALS. C. PMS. D. Left PLS. AC, aciniform gland spigot(s); CY, cylindrical gland spigot(s); Fls, flankers of pseudoflagelliform gland spigot; MAP, major ampullate gland spigot(s); mAP, minor ampullate gland spigot(s); MS, modified spigot, or pseudoflagelliform gland spigot; PI, piriform gland spigot(s); Tp, tartipore. Scale bars: A, C = 100 µm, B = 20 µm, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158615" httpUri="https://zenodo.org/record/13158615/files/figure.png" pageId="81" pageNumber="82">22 A–D</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B44BE32EFAF77EAB9" box="[1009,1088,992,1013]" captionStart="FIGURE" captionStartId="124.[155,167,1031,1050]" captionTargetBox="[120,1176,217,1016]" captionTargetId="figure-11@124.[120,1176,216,1016]" captionTargetPageId="124" captionText="FIGURE 23. Details of spinning organs of Uduba schlingeri, female CASENT9025468 from Vatoharanana, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Cribellum and anterior of PMS. B. Right ALS, MAP and TP. C. Cribellum spinning field, close-up of cribellar spigot bundles. D. Right ALS, PI and TP. MAP, major ampullate gland spigot(s); PI, piriform gland spigot(s); TP, tartipore. Scale bars: A = 100 µm, C = 3 µm, B, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158617" httpUri="https://zenodo.org/record/13158617/files/figure.png" pageId="81" pageNumber="82">23 A–D</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B430632EFAFA6EAB9" box="[1097,1169,992,1013]" captionStart="FIGURE" captionStartId="125.[155,167,1419,1438]" captionTargetBox="[121,1176,216,1403]" captionTargetId="figure-16@125.[120,1176,216,1404]" captionTargetPageId="125" captionText="FIGURE 24. Details of spinning organs of Uduba schlingeri, female CASENT9025468 from Vatoharanana, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Right PMS apex, showing AC, mAP and TP. B. Left PLS (image flipped to appear as right), showing several AC, a CY and the triplet of MS and two MSFl. C. Right PMS, showing AC (not labelled). D. Left PLS (image flipped to appear as right), showing the triplet of MS and two MSFls, a CY and several AC (not labelled). E. Right PMS, posterolateral surface showing CY. F. Right ALS, showing PIs and TPs. AC, aciniform gland spigot(s); CY, cylindrical gland spigot(s); Fls, flanking spigot(s) of modified spigot; mAP, minor ampullate gland spigot(s); MS, modified spigot, or pseudoflagelliform gland spigot; TP, tartipore. Scale bars: A–F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158621" httpUri="https://zenodo.org/record/13158621/files/figure.png" pageId="81" pageNumber="82">24 A–F</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B47D432F2AA32ED5E" box="[155,261,1021,1042]" captionStart="FIGURE" captionStartId="126.[155,167,1186,1205]" captionTargetBox="[120,1176,216,1169]" captionTargetId="figure-11@126.[120,1176,216,1169]" captionTargetPageId="126" captionText="FIGURE 25. Details of spinning organs of male Uduba spp., scanning electron micrographs (SEM). A, C. U. fisheri (CASENT9026389) from Anjozorobe. B, D, E. U. schlingeri, (CASENT9006052) from Talatakely, Parc Nacional Ranomafana. A. Cribellum, lacking spigots. B. Left PMS showing mAP and TP. C. Left PLS showing nubbins (Nu) that remain from MS-MSFls triplet. D. Left ALS, showing MAP, TP, and MAP Nu. E. Left PLS showing nubbins (Fl) that remain from MS-MSFl triplet. Fls, nubbins of flankers of pseudoflagelliform gland spigot; MAP, major ampullate gland spigot(s); MAP-Nu, major ampullate gland spigot nubbin; mAP, minor ampullate gland spigot(s); MS, nubbin of modified spigot, or pseudoflagelliform gland spigot; TP, tartipore. Scale bars: A–D = 10 µm, E = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.13158623" httpUri="https://zenodo.org/record/13158623/files/figure.png" pageId="81" pageNumber="82">25 B, D, E</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B465F32F2AA6AED5E" box="[272,349,1021,1042]" captionStart="FIGURE" captionStartId="127.[155,167,738,757]" captionTargetBox="[120,1176,216,721]" captionTargetId="figure-16@127.[120,1176,216,721]" captionTargetPageId="127" captionText="FIGURE 26. Spinning organs of Uduba spp. A. B. U. schlingeri ♀, (CASENT9006075), Vatoharanana, Parc Nacional Ranomafana. C. U. lamba, ♂, holotype, (CASENT9042542), R.N.I. Betampona. Cl, colulus, Cr, cribellum. Scale bars: A, B = 0.5 mm, C = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158625" httpUri="https://zenodo.org/record/13158625/files/figure.png" pageId="81" pageNumber="82">26 A, B</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B462732F2AAA0ED5E" box="[360,407,1021,1042]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="81" pageNumber="82">29 E</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B46EE32F2AAD9ED5E" box="[417,494,1021,1042]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="81" pageNumber="82">61 A–C</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B46B632F2A973ED5E" box="[505,580,1021,1042]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="81" pageNumber="82">62 A–E</figureCitation>
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<figureCitation id="EBB8FEA01815FF8B450132F2A9ABED5E" box="[590,668,1021,1042]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="81" pageNumber="82">73 D, E</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B45E932F2A9C4ED5E" box="[678,755,1021,1042]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="81" pageNumber="82">80 A, B</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B45B132F2A87EED5E" box="[766,841,1021,1042]" captionStart="FIGURE" captionStartId="171.[155,167,990,1009]" captionTargetBox="[120,1176,216,971]" captionTargetId="figure-16@171.[120,1176,216,971]" captionTargetPageId="171" captionText="FIGURE 83. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Vulva and book lungs, dorsal. B. Vulva, dorsal. C. Right side of vulva, dorsal, inset shows HS. D. Right side of vulva, dorsal, inset shows texture of spermatheca. E. Right side of vulva, dorsal, close-up of pores of HS. Scale bars: A = 500 µm, B = 100 µm, C = 430 µm, D, = 300 µm, E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13158761" httpUri="https://zenodo.org/record/13158761/files/figure.png" pageId="81" pageNumber="82">83 A–E</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B441B32F2A896ED5E" box="[852,929,1021,1042]" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="81" pageNumber="82">84 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B44E432F2A8C3ED5E" box="[939,1012,1021,1042]" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="81" pageNumber="82">Maps 5</figureCitation>
, 14.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1815FF8B47E4352DAF77ED96" pageId="81" pageNumber="82" type="materials_examined">
<paragraph id="733CE2251815FF8B47E4352DAF77ED96" blockId="81.[123,1174,1058,1725]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E4352DAA79ED75" bold="true" box="[171,334,1058,1081]" pageId="81" pageNumber="82">Type material.</emphasis>
<materialsCitation id="C3EBE8781815FF8B461A352DAA0BED36" collectingDate="1998-04-05" collectingDateMax="1998-04-18" collectingDateMin="1998-04-05" collectionCode="CAS" collectorName="C. E. Griswold" country="Madagascar" county="Ranomafana" location="Talatakely" municipality="Parc National Ranomafana" pageId="81" pageNumber="82" specimenCode="CASENT9006052" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871815FF8B461A352DAA8FED75" box="[341,440,1058,1081]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1815FF8B454F352DA9FEED75" box="[512,713,1058,1081]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9006052</specimenCode>
) from rainforest at
<location id="765CB4FE1815FF8B44D8352DAF37ED75" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1815FF8B44D8352DAF37ED75" box="[919,1024,1058,1081]" country="Madagascar" county="Ranomafana" municipality="Parc National Ranomafana" name="Talatakely" pageId="81" pageNumber="82" stateProvince="Fianarantsoa">Talatakely</location>
,
<collectingMunicipality id="9358785F1815FF8B4341352DAA28ED16" pageId="81" pageNumber="82">
Parc National
<collectingCounty id="9A5D9AA91815FF8B47DA354CAA28ED16" box="[149,287,1091,1114]" pageId="81" pageNumber="82">Ranomafana</collectingCounty>
</collectingMunicipality>
,
<collectingRegion id="B1472CC71815FF8B4666354CA92DED16" box="[297,538,1091,1114]" country="Madagascar" name="Fianarantsoa" pageId="81" pageNumber="82">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51815FF8B456A354CA993ED16" box="[549,676,1091,1114]" name="Madagascar" pageId="81" pageNumber="82">Madagascar</collectingCountry>
, collected
<date id="073DC4E51815FF8B4457354CA8F2ED16" box="[792,965,1091,1114]" pageId="81" pageNumber="82" value="1998-04-05" valueMax="1998-04-18" valueMin="1998-04-05">
<collectingDate id="17793D0D1815FF8B4457354CA8F2ED16" box="[792,965,1091,1114]" pageId="81" pageNumber="82" value="1998-04-05" valueMax="1998-04-18" valueMin="1998-04-05">5–18 April 1998</collectingDate>
</date>
by
<collectorName id="DE7687F31815FF8B44A1354CAFA6ED16" box="[1006,1169,1091,1114]" pageId="81" pageNumber="82">C. E. Griswold</collectorName>
, deposited in
<collectionCode id="15927AE01815FF8B464A356CAA0BED36" box="[261,316,1123,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781815FF8B4608356CA856ED36" box="[327,865,1123,1146]" collectingDate="1998-04-05" collectingDateMax="1998-04-18" collectingDateMin="1998-04-05" collectionCode="CASENT" collectorName="C. E. Griswold" country="Madagascar" county="Ranomafana" location="Talatakely" municipality="Parc National Ranomafana" pageId="81" pageNumber="82" specimenCode="CASENT9006052" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871815FF8B4608356CAA93ED36" box="[327,420,1123,1146]" pageId="81" pageNumber="82" type="paratype">Paratype</typeStatus>
male, same data, also
<specimenCode id="23254A5E1815FF8B45D6356CA856ED36" box="[665,865,1123,1146]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9006052</specimenCode>
</materialsCitation>
, deposited in
<collectionCode id="15927AE01815FF8B44B8356CAF19ED36" box="[1015,1070,1123,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
;
<materialsCitation id="C3EBE8781815FF8B4375356CA98CEDF6" collectingDate="1998-04-28" collectionCode="CAS" collectorName="Talatakely, P. N &amp; C. E. Griswold" country="Madagascar" location="Ranomafana" pageId="81" pageNumber="82" specimenCode="CASENT9006053" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871815FF8B4375356CAFA1ED36" box="[1082,1174,1123,1146]" pageId="81" pageNumber="82" type="paratype">paratype</typeStatus>
male (
<specimenCode id="23254A5E1815FF8B47F1358CAAB0EDD6" box="[190,391,1155,1178]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9006053</specimenCode>
) from forest at research cabin at
<collectorName id="DE7687F31815FF8B45A1358CA8A3EDD6" box="[750,916,1155,1178]" pageId="81" pageNumber="82">Talatakely, P. N</collectorName>
.
<location id="765CB4FE1815FF8B44EF358CAF1DEDD6" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1815FF8B44EF358CAF1DEDD6" box="[928,1066,1155,1178]" country="Madagascar" name="Ranomafana" pageId="81" pageNumber="82" stateProvince="Fianarantsoa">Ranomafana</location>
, collected
<date id="073DC4E51815FF8B473435ACAA24EDF6" box="[123,275,1187,1210]" pageId="81" pageNumber="82" value="1998-04-28">
<collectingDate id="17793D0D1815FF8B473435ACAA24EDF6" box="[123,275,1187,1210]" pageId="81" pageNumber="82" value="1998-04-28">28 April 1998</collectingDate>
</date>
by
<collectorName id="DE7687F31815FF8B460E35ACAADEEDF6" box="[321,489,1187,1210]" pageId="81" pageNumber="82">C. E. Griswold</collectorName>
, deposited in
<collectionCode id="15927AE01815FF8B45CB35ACA98CEDF6" box="[644,699,1187,1210]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="82" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781815FF8B458735ACAF0CED96" collectingDate="1992-03-23" collectionCode="MCZ" collectorName="V. and B. Roth" country="Madagascar" location="Parc National Ranomafana" municipality="Ranomafana" pageId="81" pageNumber="82" specimenCode="CASENT9006079" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C871815FF8B458735ACA812EDF6" box="[712,805,1187,1210]" pageId="81" pageNumber="82" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1815FF8B44C735ACAF66EDF6" box="[904,1105,1187,1210]" collectionCode="CASENT" pageId="81" pageNumber="82">CASENT9006079</specimenCode>
) from
<location id="765CB4FE1815FF8B473435CCAA97ED96" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1815FF8B473435CCAA97ED96" box="[123,416,1219,1242]" country="Madagascar" municipality="Ranomafana" name="Parc National Ranomafana" pageId="81" pageNumber="82" stateProvince="Fianarantsoa">
Parc National
<collectingMunicipality id="9358785F1815FF8B465935CCAA97ED96" box="[278,416,1219,1242]" pageId="81" pageNumber="82">Ranomafana</collectingMunicipality>
</location>
, collected
<date id="073DC4E51815FF8B455B35CCA98BED96" box="[532,700,1219,1242]" pageId="81" pageNumber="82" value="1992-03-23">
<collectingDate id="17793D0D1815FF8B455B35CCA98BED96" box="[532,700,1219,1242]" pageId="81" pageNumber="82" value="1992-03-23">23 March 1992</collectingDate>
</date>
,
<collectorName id="DE7687F31815FF8B458835CCA85FED96" box="[711,872,1219,1242]" pageId="81" pageNumber="82">V. and B. Roth</collectorName>
, deposited in
<collectionCode id="15927AE01815FF8B44B135CCAF0CED96" box="[1022,1083,1219,1242]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="81" pageNumber="82" type="Museum">MCZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1815FF8B47E435ECA8A5EC37" pageId="81" pageNumber="82" type="etymology">
<paragraph id="733CE2251815FF8B47E435ECA8A5EC37" blockId="81.[123,1174,1058,1725]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E435ECAA1DEDB6" bold="true" box="[171,298,1251,1274]" pageId="81" pageNumber="82">Etymology.</emphasis>
The species epithet is a patronym in honor of Evert I. Schlinger, entomological explorer, charismatic teacher, and generous mentor. The eponymous Schlinger Foundation provid- ed extensive support for Arachnology at the California Academy of Sciences and also for the Malaise Trap survey of
<collectingCountry id="0B94A2B51815FF8B46C9344BA930EC17" box="[390,519,1348,1371]" name="Madagascar" pageId="81" pageNumber="82">Madagascar</collectingCountry>
arthropods, i.e., the Schlinger
<emphasis id="41F73E371815FF8B442C344BA894EC17" box="[867,931,1348,1371]" italics="true" pageId="81" pageNumber="82">et al.,</emphasis>
Malaise Trap Survey, which endeavor produced many of the spiders examined by our revision.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1815FF8B47E4348BAAE0EF31" pageId="81" pageNumber="82" type="diagnosis">
<paragraph id="733CE2251815FF8B47E4348BAAE0EF31" blockId="81.[123,1174,1058,1725]" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E4348BAADFECD7" bold="true" box="[171,488,1412,1435]" pageId="81" pageNumber="82">
Diagnosis.
<taxonomicName id="B48399A61815FF8B4668348BAADFECD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,488,1412,1435]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371815FF8B4668348BAADFECD7" bold="true" box="[295,488,1412,1435]" italics="true" pageId="81" pageNumber="82">Uduba schlingeri</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.c, the
<taxonomicName id="B48399A61815FF8B440E348BA8D6ECD7" authorityName="Simon" authorityYear="1906" box="[833,993,1412,1435]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01815FF8B4378348BAFBAECD7" box="[1079,1165,1412,1435]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="81" pageNumber="82">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are cribellate with cribellum divided (
<figureCitation id="EBB8FEA01815FF8B434B34AAAF58ECF0" box="[1028,1135,1445,1468]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="81" pageNumber="82">Fig. 20 G</figureCitation>
) or with vestige of a divided cribellum (
<figureCitation id="EBB8FEA01815FF8B46B334CAA954EC90" box="[508,611,1477,1500]" captionStart="FIGURE" captionStartId="122.[155,167,1045,1064]" captionTargetBox="[120,1176,216,1027]" captionTargetId="figure-11@122.[120,1176,216,1027]" captionTargetPageId="122" captionText="FIGURE 21. Spinning organs of Uduba schlingeri, male CASENT9006052 from Talatakely, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Whole spinnerets. B. Left ALS, showing Nu of MAP. C. Apex of left PMS. D. Left PMS, showing Nu of MS and Fls triplet. AC, aciniform gland spigot(s); Fls, flankers of pseudoflagelliform gland spigot; MAP, major ampullate gland spigot(s); mAP, minor ampullate gland spigot(s); MS, modified spigot, or pseudoflagelliform gland spigot; Nu, nubbin; PI, piriform gland spigot(s); Tp, tartipore. Scale bars: A = 100 µm, B = 20 µm, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158612" httpUri="https://zenodo.org/record/13158612/files/figure.png" pageId="81" pageNumber="82">Fig. 21 A</figureCitation>
). Males have the palp (
<figureCitation id="EBB8FEA01815FF8B441634CAA8D1EC90" box="[857,998,1477,1500]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="81" pageNumber="82">Figs. 61 A–C</figureCitation>
<emphasis id="41F73E371815FF8B44AA34CAA8D9EC90" bold="true" box="[997,1006,1477,1500]" pageId="81" pageNumber="82">)</emphasis>
with TA2 highly sclerotized, black, TA2 forming a chela around TA3 apex (
<figureCitation id="EBB8FEA01815FF8B444634EAA84CECB0" box="[777,891,1509,1532]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="81" pageNumber="82">Figs. 29 E</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B44C934EAA8ECECB0" box="[902,987,1509,1532]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="81" pageNumber="82">62 C, E</figureCitation>
), TA3 transverse, tapering to a sharp point (
<figureCitation id="EBB8FEA01815FF8B46DD370AA910EF50" box="[402,551,1541,1564]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="81" pageNumber="82">Figs. 62 D, E</figureCitation>
). Females have an epigynal plate with a median lobe surrounded by a depressed atrium (
<figureCitation id="EBB8FEA01815FF8B4699372AA97EEF70" box="[470,585,1573,1596]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="81" pageNumber="82">Figs. 80 A</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B451A372AA9BBEF70" box="[597,652,1573,1596]" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="81" pageNumber="82">84 A</figureCitation>
); median lobe extends to epigastric furrow (
<figureCitation id="EBB8FEA01815FF8B4325372AAB98EF10" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="81" pageNumber="82">Fig. 73 D</figureCitation>
).
<taxonomicName id="B48399A61815FF8B478B374AAAB5EF10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[196,386,1605,1628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371815FF8B478B374AAAB5EF10" bold="true" box="[196,386,1605,1628]" italics="true" pageId="81" pageNumber="82">Uduba schlingeri</emphasis>
</taxonomicName>
are distinguished from the ecribellate
<taxonomicName id="B48399A61815FF8B4457374AA8E2EF10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[792,981,1605,1628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="andriamihajai">
<emphasis id="41F73E371815FF8B4457374AA8E2EF10" bold="true" box="[792,981,1605,1628]" italics="true" pageId="81" pageNumber="82">U. andriamihajai</emphasis>
</taxonomicName>
in having a divid- ed cribellum (
<figureCitation id="EBB8FEA01815FF8B465D3769AABFEF31" box="[274,392,1638,1661]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="81" pageNumber="82">Figs. 20 G</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B46DB3769AAFCEF31" box="[404,459,1638,1661]" captionStart="FIGURE" captionStartId="124.[155,167,1031,1050]" captionTargetBox="[120,1176,217,1016]" captionTargetId="figure-11@124.[120,1176,216,1016]" captionTargetPageId="124" captionText="FIGURE 23. Details of spinning organs of Uduba schlingeri, female CASENT9025468 from Vatoharanana, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Cribellum and anterior of PMS. B. Right ALS, MAP and TP. C. Cribellum spinning field, close-up of cribellar spigot bundles. D. Right ALS, PI and TP. MAP, major ampullate gland spigot(s); PI, piriform gland spigot(s); TP, tartipore. Scale bars: A = 100 µm, C = 3 µm, B, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158617" httpUri="https://zenodo.org/record/13158617/files/figure.png" pageId="81" pageNumber="82">23 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1815FF8E47E43789A819EAF9" lastPageId="84" lastPageNumber="85" pageId="81" pageNumber="82" type="description">
<paragraph id="733CE2251815FF8947E43789AB93E85F" blockId="81.[123,1174,1058,1725]" lastBlockId="83.[123,1175,220,1725]" lastPageId="83" lastPageNumber="84" pageId="81" pageNumber="82">
<emphasis id="41F73E371815FF8B47E43789AA40EFD1" bold="true" box="[171,375,1670,1693]" pageId="81" pageNumber="82">Description. Male</emphasis>
(
<typeStatus id="AC385C871815FF8B46C93789AAC7EFD1" box="[390,496,1670,1693]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
): Total length 14.29. Markings as in
<figureCitation id="EBB8FEA01815FF8B44323789AF0AEFD1" box="[893,1085,1670,1693]" captionStart="FIGURE" captionStartId="107.[155,167,1531,1550]" captionTargetBox="[120,1176,216,1515]" captionTargetId="figure-16@107.[120,1176,216,1515]" captionTargetPageId="107" captionText="FIGURE 3. Habitus photos of Uduba schlingeri from Parc National Ranomafana. A, B, D, G. Female ♀ (CASENT9006075). C, E, F, H. ♂ Holotype male (CASENT9006052). A. Dorsal. B. Ventral. C. Dorsal. D. Carapace, dorsal. E. Ventral. F. Carapace, dorsal. G. Carapace, lateral. H. Carapace, lateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158562" httpUri="https://zenodo.org/record/13158562/files/figure.png" pageId="81" pageNumber="82">Figs. 3 C, E, F, H</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B43073789AF46EFD1" box="[1096,1137,1670,1693]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="81" pageNumber="82">9 A</figureCitation>
,
<figureCitation id="EBB8FEA01815FF8B43323789ABA6EFF1" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="81" pageNumber="82">11 G</figureCitation>
. Carapace 9.07 long, 6.43 wide, 2.86 high; clypeus 0.43 high. Eye diameters: AME 0.39, ALE and PME 0.41, PLE 0.45. Chelicerae 4.07 long; sternum 4.11 long, 3.11 wide; labium 1.96 long; palpal coxae 2.86 long. Divided cribellum (
<figureCitation id="EBB8FEA01816FF88450531F3A987E85F" box="[586,688,252,275]" captionStart="FIGURE" captionStartId="122.[155,167,1045,1064]" captionTargetBox="[120,1176,216,1027]" captionTargetId="figure-11@122.[120,1176,216,1027]" captionTargetPageId="122" captionText="FIGURE 21. Spinning organs of Uduba schlingeri, male CASENT9006052 from Talatakely, Parc Nacional Ranomafana, scanning electron micrographs (SEM). A. Whole spinnerets. B. Left ALS, showing Nu of MAP. C. Apex of left PMS. D. Left PMS, showing Nu of MS and Fls triplet. AC, aciniform gland spigot(s); Fls, flankers of pseudoflagelliform gland spigot; MAP, major ampullate gland spigot(s); mAP, minor ampullate gland spigot(s); MS, modified spigot, or pseudoflagelliform gland spigot; Nu, nubbin; PI, piriform gland spigot(s); Tp, tartipore. Scale bars: A = 100 µm, B = 20 µm, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13158612" httpUri="https://zenodo.org/record/13158612/files/figure.png" pageId="82" pageNumber="83">Fig. 21 A</figureCitation>
). Spination (Mt Papango CASENT9062276): palpus–femur d0-0-1-1, p0-0-0-1, r0-0-0-1; leg I–femur d1-1-0-1, p0-1-1-1, r1-1-1-1, patella p1, r1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p0-1-1-0, v2-2-3, r0-1-1-0; leg II–femur d1-1-0-1, p0-1-1-1, r1-1-1-1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p0-1-1-0, v2- 2-3, r0-1-1-0; leg III–femur d1-0-1-1, p0-1-1-1, r0-1-1-1, patella r1, tibia d0-1-1-0, p0-0-0-1, v1- 2-2, r0-0-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-1, p0-1-1-1, r0-0-0-1, patella r1, tibia-tarsus of CASENT9062276 lost; (leg IV of
<typeStatus id="AC385C871816FF88458330B2A818E898" box="[716,815,445,468]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
): femur d1-1-0-1, p0-1-1-1, r0-1- 1-1, patella r1, tibia d0-0-1-0, p0-1-0-1, v2-2-2, r0-1-0-1, metatarsus p0-1-1-2, v1-2-2-2, r1-2-2). Scopulae: cymbium, apicodorsal (
<figureCitation id="EBB8FEA01816FF8846BF30F1A9BCEB59" box="[496,651,510,533]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">Figs. 62 A, C</figureCitation>
); beneath tarsi and apices of all metatarsi. Leg measurements (
<typeStatus id="AC385C871816FF88466C3311AAB0EB79" box="[291,391,542,565]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
): I: 8.93 + 3.64 + 9.21 + 10.00 + 5.14 = 36.93; II: 8.43 + 3.36 + 7.64 + 8.64 + 4.50 = 32.57; III: 6.79 + 3.21 + 4.86 + 7.36 + 3.50 = 25.71; IV: 9.14 + 3.14 + 8.36 + 11.07 + 4.64 = 36.36; palpus: 4.14 + 1.43 + 1.64 + NA + 4.07 = 11.29. Leg formula 1423.
<emphasis id="41F73E371816FF88436E3351AFA2EB39" bold="true" box="[1057,1173,606,629]" pageId="82" pageNumber="83">Male palp</emphasis>
(
<figureCitation id="EBB8FEA01816FF8847CC3371ABCEEBD9" box="[131,249,638,661]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="82" pageNumber="83">Figs. 29 E</figureCitation>
,
<figureCitation id="EBB8FEA01816FF8846483371AA6AEBD9" box="[263,349,638,661]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">61 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01816FF8846243371AA8AEBD9" box="[363,445,638,661]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">62 A–E</figureCitation>
<emphasis id="41F73E371816FF8846F23371AAF1EBD9" bold="true" box="[445,454,638,661]" pageId="82" pageNumber="83">)</emphasis>
(CASENT9006062): palpal tibia 0.42 times cymbial length, RTA broadly triangular, with pointed apex (
<figureCitation id="EBB8FEA01816FF8845573390A9B6EBFA" box="[536,641,671,694]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">Fig. 61 C</figureCitation>
), RTA length 0.22 tibia length, length 0.89 width, VTA cylindrical, straight, with rounded apex (
<figureCitation id="EBB8FEA01816FF88453433B0A82FEB9A" box="[635,792,703,726]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">Figs. 61 B, C</figureCitation>
,
<figureCitation id="EBB8FEA01816FF88446833B0A8B3EB9A" box="[807,900,703,726]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">62 B, C</figureCitation>
), VTA length 3.00 times width, length 0.39 times tibia width, tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.975 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.57 times tegulum length; TA3 transverse, tapering to a sharp point (
<figureCitation id="EBB8FEA01816FF8844DC322FAF1FEA7B" box="[915,1064,800,823]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">Figs. 61 A, B</figureCitation>
), concave dorsally (
<figureCitation id="EBB8FEA01816FF8847AF324FAA43EA1B" box="[224,372,832,855]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">Figs. 62 D, E</figureCitation>
), TA3 apex visible in front of TA2; TA2 highly sclerotized, black, concave medially against TA3, in this species forming a chela around TA3 apex (
<figureCitation id="EBB8FEA01816FF8844D5326FAF27EA3B" box="[922,1040,864,887]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="82" pageNumber="83">Figs. 29 E</figureCitation>
,
<figureCitation id="EBB8FEA01816FF884351326FAF62EA3B" box="[1054,1109,864,887]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">62 E</figureCitation>
); MA transverse, concave in middle with proximal and distal enlargements (
<figureCitation id="EBB8FEA01816FF884420328FA8D5EADB" box="[879,994,896,919]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">Figs. 61 A</figureCitation>
,
<figureCitation id="EBB8FEA01816FF8844A1328FAF11EADB" box="[1006,1062,896,919]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">62 D</figureCitation>
), origin at 0.35 times tegulum length, tegulum apex at 0.425 tegulum length from MA apex, length (including apical processes) 0.61 times width, MA length 0.275 tegulum length, MA width 0.44 times tegulum width; conductor fan entire (
<figureCitation id="EBB8FEA01816FF88452A32EEA823EAB4" box="[613,788,993,1016]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="82" pageNumber="83">Figs. 61 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01816FF88446032EEA8A6EAB4" box="[815,913,993,1016]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="82" pageNumber="83">62 A–E</figureCitation>
).
<emphasis id="41F73E371816FF8844FB32EEAF3BEAB4" bold="true" box="[948,1036,993,1016]" pageId="82" pageNumber="83">Female</emphasis>
(
<typeStatus id="AC385C871816FF88436832EEAFA6EAB4" box="[1063,1169,993,1016]" pageId="82" pageNumber="83" type="paratype">Paratype</typeStatus>
, CASENT9006079): Total length 17.14. Markings as in
<figureCitation id="EBB8FEA01816FF88459D350EA84EED54" box="[722,889,1025,1048]" captionStart="FIGURE" captionStartId="107.[155,167,1531,1550]" captionTargetBox="[120,1176,216,1515]" captionTargetId="figure-16@107.[120,1176,216,1515]" captionTargetPageId="107" captionText="FIGURE 3. Habitus photos of Uduba schlingeri from Parc National Ranomafana. A, B, D, G. Female ♀ (CASENT9006075). C, E, F, H. ♂ Holotype male (CASENT9006052). A. Dorsal. B. Ventral. C. Dorsal. D. Carapace, dorsal. E. Ventral. F. Carapace, dorsal. G. Carapace, lateral. H. Carapace, lateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158562" httpUri="https://zenodo.org/record/13158562/files/figure.png" pageId="82" pageNumber="83">Figs. 3 A, B, D</figureCitation>
,
<figureCitation id="EBB8FEA01816FF8844C9350EA886ED54" box="[902,945,1025,1048]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="82" pageNumber="83">7 G</figureCitation>
. Carapace 8.71 long, 6.00 wide, 3.71 high; clypeus 0.64 high. Eye diameters:AME, PME and PLE 0.36, ALE 0.38. Chelicerae 4.14 long; sternum 3.93 long, 3.04 wide; labium 1.96 long; palpal coxae 3.04 long. Cribellum divided (
<figureCitation id="EBB8FEA01816FF884644356DAAB6ED35" box="[267,385,1122,1145]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="82" pageNumber="83">Figs. 20 G</figureCitation>
,
<figureCitation id="EBB8FEA01816FF8846C3356DAAD6ED35" box="[396,481,1122,1145]" captionStart="FIGURE" captionStartId="127.[155,167,738,757]" captionTargetBox="[120,1176,216,721]" captionTargetId="figure-16@127.[120,1176,216,721]" captionTargetPageId="127" captionText="FIGURE 26. Spinning organs of Uduba spp. A. B. U. schlingeri ♀, (CASENT9006075), Vatoharanana, Parc Nacional Ranomafana. C. U. lamba, ♂, holotype, (CASENT9042542), R.N.I. Betampona. Cl, colulus, Cr, cribellum. Scale bars: A, B = 0.5 mm, C = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158625" httpUri="https://zenodo.org/record/13158625/files/figure.png" pageId="82" pageNumber="83">26 A, B</figureCitation>
). Spination (Talatakely, CASENT9006056): palpus–femur d0-0- 2; leg I–femur d1-0-1-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II–femur d1-0-10, p0-0-1- 1, tibia v1-1-2, metatarsus v2-2-3; leg III–femur d1-0-0-0, p0-0-0-2, r0-1-0-1, tibia d0-1-0, p0-1- 1-0, v1-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d0-1-1-0, r0-0-0-1, tibia v1- 1-2, r0-1-1-0, metatarsus p0-0-1-1, v2-2-2, r0-0-1-1. Scopulae: strong ventral beneath all tarsi and metatarsi and apices of tibiae I and II (
<figureCitation id="EBB8FEA01816FF884569340CA98BEC56" box="[550,700,1283,1306]" captionStart="FIGURE" captionStartId="116.[155,167,1638,1657]" captionTargetBox="[121,1176,1059,1621]" captionTargetId="figure-105@116.[120,1176,1059,1622]" captionTargetPageId="116" captionText="FIGURE 13. Uduba schlingeri, from Ranomafana National Park, leg photos. A–C, E, F. ♀ (CASENT9006075). D. ♂ (CASENT9006052). A. Leg I, prolateral. B. Metatarsus and tarsus IV, retrolateral, showing calamistrum-arrow to calamistrum. C. Leg IV, retrolateral. D. ♂ tibia II retrolateral, base showing tibial crack (arrow). E. Trochanter IV, left. F. Trochanter I, left. Scale bars: A, C = 2 mm, B = 1 mm, D, E, F = 0.25 mm" figureDoi="http://doi.org/10.5281/zenodo.13158588" httpUri="https://zenodo.org/record/13158588/files/figure.png" pageId="82" pageNumber="83">Figs. 13 A, C</figureCitation>
,
<figureCitation id="EBB8FEA01816FF884586340CA877EC56" box="[713,832,1283,1306]" captionStart="FIGURE" captionStartId="117.[155,167,913,932]" captionTargetBox="[120,1176,216,897]" captionTargetId="figure-16@117.[120,1176,216,897]" captionTargetPageId="117" captionText="FIGURE 14. Uduba spp., females, legs, SEM. A–E, G. U. schlingeri ♀ (CASENT9025468) from Vatoharanana, Ranomafana National Park. F, H. U. fisheri, ♀ (CASENT9026389) from Anjozorobe. A. Tarsus I, left, retrolateral. B. Tarsus IV, left, retrolateral. C. Tarsus I, left, dorsal. D. Tarsus IV, left, dorsal. E. Palpal tarsus, right, retrolateral. F. Palpus, trochanter–tarsus, prolateral. G. Metatarsus IV base, retrodorsal, showing calamistrum [arrow]. H. Metatarsus IV, retrodorsal, showing calamistrum [arrow]. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158590" httpUri="https://zenodo.org/record/13158590/files/figure.png" pageId="82" pageNumber="83">14 A–E, G</figureCitation>
). Leg measurements (
<typeStatus id="AC385C871816FF88437F340CAFA5EC56" box="[1072,1170,1283,1306]" pageId="82" pageNumber="83" type="paratype">Paratype</typeStatus>
, CASENT9006079): I: 6.93 + 3.21 + 6.07 + 5.00 + 3.14 = 24.36; II: 6.43 + 3.14 + 4.79 + 4.29 + 2.86 = 21.50; III: 5.00 + 2.50 + 2.64 + 3.79 + 1.86 = 15.79; IV: 6.64 + 2.86 + 5.43 + 6.14 + 2.71 = 23.79; palpus: 3.43 + 1.71 + 1.86 + NA + 2.79 = 9.79. Leg formula 1423.
<emphasis id="41F73E371816FF88434E346BAB9AECD7" bold="true" pageId="82" pageNumber="83">Female genitalia</emphasis>
(Talatakely, CASENT9006057): epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01816FF88478034ABAA74ECF7" box="[207,323,1444,1467]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="82" pageNumber="83">Figs. 73 D</figureCitation>
,
<figureCitation id="EBB8FEA01816FF88461F34ABAAB1ECF7" box="[336,390,1444,1467]" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="82" pageNumber="83">84 A</figureCitation>
), plate width 1.31 times length; atrium sides concave, atrium width 1.81 times atrium length; epigynum length 1.95 times atrium length, epigynum width 1.42 times atrium width; atrium width at side of ML 0.86 times ML width, 1.71 times LL width; atrium origin near epigastric groove, atrium origin at 0.12 of epigynal plate length, median lobe extends posteriad to epigastric groove, median lobe length 2.07 times width, ML extends anteriad of atrium apex by 0.15 times epigynum length, ML length 0.71 times epigynal length; lateral lobes narrow, atrium width at side of ML 1.71 times LL width; CO hidden beneath posterolateral margins of atrium, distance between CO 0.70 times width of epigynal plate, CO distance from epigastric furrow 0.29 times epigynum length. Vulva (
<figureCitation id="EBB8FEA01816FF88468137A9A975EFF1" box="[462,578,1702,1725]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="82" pageNumber="83">Figs. 73 E</figureCitation>
,
<figureCitation id="EBB8FEA01816FF88450137A9A9B4EFF1" box="[590,643,1702,1725]" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="82" pageNumber="83">84 B</figureCitation>
) with spermathecal ducts making two transverse curves, vulva length 1.04 times width, FD close-together, vulva width 8.17 times distance between FD.
</paragraph>
<paragraph id="733CE2251817FF8947E43013A9F7EADB" blockId="83.[123,1175,220,1725]" pageId="83" pageNumber="84">
<emphasis id="41F73E371817FF8947E43013AA6AE87F" bold="true" box="[171,349,284,307]" pageId="83" pageNumber="84">Variation. Male</emphasis>
(N = 9): Total length = 14.70–24.29, carapace length / width = 1.41–1.78, carapace height / width = 0.42–0.71, PER / carapace width = 0.44–0.92, PER / OAL = 2.43–3.21, PER / AER = 1.21–1.42, OAL / OQL = 1.00–1.09, OQP / OQA = 1.02–1.16, clypeus height / AME = 1.33–2.00, cheliceral length / clypeus height = 6.44–10.83, sternum length / width = 1.24–1.43, palpal coxa length / width = 1.53–3.10, femur I length / carapace width = 1.27–1.59, metatarsus I length / carapace width = 1.29–1.60, femur IV length / carapace width = 1.30–1.56, cymbium length / carapace width = 0.62–0.72, cymbium length / palpal patella length = 2.85–3.08, cymbium length / palpal tibia length = 2.48–2.80, cymbium length / palpal femur length = 0.98–1.24, palpal tibia length / palpal patella length = 1.04–1.15. Markings as in
<figureCitation id="EBB8FEA01817FF8944193311AF2EEB79" box="[854,1049,542,565]" captionStart="FIGURE" captionStartId="107.[155,167,1531,1550]" captionTargetBox="[120,1176,216,1515]" captionTargetId="figure-16@107.[120,1176,216,1515]" captionTargetPageId="107" captionText="FIGURE 3. Habitus photos of Uduba schlingeri from Parc National Ranomafana. A, B, D, G. Female ♀ (CASENT9006075). C, E, F, H. ♂ Holotype male (CASENT9006052). A. Dorsal. B. Ventral. C. Dorsal. D. Carapace, dorsal. E. Ventral. F. Carapace, dorsal. G. Carapace, lateral. H. Carapace, lateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158562" httpUri="https://zenodo.org/record/13158562/files/figure.png" pageId="83" pageNumber="84">Figs. 3 C, E, F, H</figureCitation>
,
<figureCitation id="EBB8FEA01817FF89436A3311AF79EB79" box="[1061,1102,542,565]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="83" pageNumber="84">9 A</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8943153311AFA6EB79" box="[1114,1169,542,565]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="83" pageNumber="84">11 G</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8947343331ABE6EB19" box="[123,209,574,597]" captionStart="FIGURE" captionStartId="116.[155,167,925,944]" captionTargetBox="[120,1176,216,925]" captionTargetId="figure-11@116.[120,1176,216,910]" captionTargetPageId="116" captionText="FIGURE 12. Uduba spp., morphology photos. A–C. U. schlingeri, ♂ CASENT9006052 from Talatakely, Ranomafana National Park. D, F. U. schlingeri, ♀ CASENT9006075 from Vatoharanana, Ranomafana National Park. E. U. fisheri, ♀ CASENT9026389 from Anjozorobe. A. Cephalothorax, dorsal. B. Cephalothorax, ventral. C. Habitus, lateral. D. Face, clypeus and divided chilum, anterior. E. Pedicel, dorsal (anterior to top). F. Habitus, lateral. Scale bars: A, B, F = 2 mm, C = 2 mm, D =1 mm, E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158586" httpUri="https://zenodo.org/record/13158586/files/figure.png" pageId="83" pageNumber="84">12 A–C</figureCitation>
. Palpus as in
<figureCitation id="EBB8FEA01817FF8946243331AAE0EB19" box="[363,471,574,597]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="83" pageNumber="84">Figs. 29E</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8946AB3331A90DEB19" box="[484,570,574,597]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="83" pageNumber="84">61 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8945083331A9ABEB19" box="[583,668,574,597]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="83" pageNumber="84">62 A–E</figureCitation>
.
<emphasis id="41F73E371817FF8945E63331A9CBEB19" bold="true" box="[681,764,574,597]" pageId="83" pageNumber="84">Female</emphasis>
(N = 7): Total length = 14.00–27.14, carapace length / width = 1.41–1.69, carapace height / width = 0.42–0.71, PER / carapace width = 0.44–0.92, PER / OAL = 2.43–3.21, PER / AER = 1.21–1.42, OAL / OQL = 1.00–1.09, OQP / OQA = 1.02–1.16, clypeus height / AME = 1.33–2.00, cheliceral length / clypeus height = 6.44–10.83, sternum length / width = 1.24–1.48, palpal coxa length / width = 1.82–2.18, femur I length / carapace width = 1.00–1.25, metatarsus I length / carapace width = 0.73–0.85, femur IV length / carapace width = 0.97–1.16, palpal tarsus length / carapace width = 0.37–0.48, palpal tarsus length / palpal patella length = 1.58–1.82, palpal tarsus length / palpal tibia length = 1.48–1.54, palpal tarsus length / palpal femur length = 0.76–0.89, palpal tibia length / palpal patella length = 1.04–1.15. Markings vary as in
<figureCitation id="EBB8FEA01817FF894615326FA915EA3B" box="[346,546,864,887]" captionStart="FIGURE" captionStartId="107.[155,167,1531,1550]" captionTargetBox="[120,1176,216,1515]" captionTargetId="figure-16@107.[120,1176,216,1515]" captionTargetPageId="107" captionText="FIGURE 3. Habitus photos of Uduba schlingeri from Parc National Ranomafana. A, B, D, G. Female ♀ (CASENT9006075). C, E, F, H. ♂ Holotype male (CASENT9006052). A. Dorsal. B. Ventral. C. Dorsal. D. Carapace, dorsal. E. Ventral. F. Carapace, dorsal. G. Carapace, lateral. H. Carapace, lateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158562" httpUri="https://zenodo.org/record/13158562/files/figure.png" pageId="83" pageNumber="84">Figs. 3 A, B, D, G</figureCitation>
,
<figureCitation id="EBB8FEA01817FF894562326FA96FEA3B" box="[557,600,864,887]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="83" pageNumber="84">7 G</figureCitation>
,
<figureCitation id="EBB8FEA01817FF89452B326FA984EA3B" box="[612,691,864,887]" captionStart="FIGURE" captionStartId="116.[155,167,925,944]" captionTargetBox="[120,1176,216,925]" captionTargetId="figure-11@116.[120,1176,216,910]" captionTargetPageId="116" captionText="FIGURE 12. Uduba spp., morphology photos. A–C. U. schlingeri, ♂ CASENT9006052 from Talatakely, Ranomafana National Park. D, F. U. schlingeri, ♀ CASENT9006075 from Vatoharanana, Ranomafana National Park. E. U. fisheri, ♀ CASENT9026389 from Anjozorobe. A. Cephalothorax, dorsal. B. Cephalothorax, ventral. C. Habitus, lateral. D. Face, clypeus and divided chilum, anterior. E. Pedicel, dorsal (anterior to top). F. Habitus, lateral. Scale bars: A, B, F = 2 mm, C = 2 mm, D =1 mm, E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158586" httpUri="https://zenodo.org/record/13158586/files/figure.png" pageId="83" pageNumber="84">12 D, F</figureCitation>
. Epigynum varies as in
<figureCitation id="EBB8FEA01817FF8944F6326FAF1AEA3B" box="[953,1069,864,887]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="83" pageNumber="84">Figs. 73 D</figureCitation>
,
<figureCitation id="EBB8FEA01817FF894375326FAF58EA3B" box="[1082,1135,864,887]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="83" pageNumber="84">80 A</figureCitation>
,
<figureCitation id="EBB8FEA01817FF894334326FABA6EADB" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="83" pageNumber="84">84 A</figureCitation>
; vulva varies as in
<figureCitation id="EBB8FEA01817FF89462C328FAAE2EADB" box="[355,469,896,919]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="83" pageNumber="84">Figs. 73 E</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8946AE328FA92FEADB" box="[481,536,896,919]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="83" pageNumber="84">80 B</figureCitation>
,
<figureCitation id="EBB8FEA01817FF89456A328FA94DEADB" box="[549,634,896,919]" captionStart="FIGURE" captionStartId="171.[155,167,990,1009]" captionTargetBox="[120,1176,216,971]" captionTargetId="figure-16@171.[120,1176,216,971]" captionTargetPageId="171" captionText="FIGURE 83. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Vulva and book lungs, dorsal. B. Vulva, dorsal. C. Right side of vulva, dorsal, inset shows HS. D. Right side of vulva, dorsal, inset shows texture of spermatheca. E. Right side of vulva, dorsal, close-up of pores of HS. Scale bars: A = 500 µm, B = 100 µm, C = 430 µm, D, = 300 µm, E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13158761" httpUri="https://zenodo.org/record/13158761/files/figure.png" pageId="83" pageNumber="84">83 B–E</figureCitation>
,
<figureCitation id="EBB8FEA01817FF8945CA328FA9F7EADB" box="[645,704,896,919]" captionStart="FIGURE" captionStartId="172.[155,167,1152,1171]" captionTargetBox="[120,1176,216,1132]" captionTargetId="figure-11@172.[120,1176,216,1132]" captionTargetPageId="172" captionText="FIGURE 84. Uduba schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park, ♀ genitalia, SEM images. A. Epigynum, ventral. B. Vulva, dorsal. C. Vulva and book lungs, dorsal. BL, book lungs; BS, spermathecal base; CD, copulatory duct; CO, copulatory opening; HS, spermathecal head, LL, lateral lobes of epigynum; ML, median lobe of epigynum. Scale bars: A, B = 300 µm, C = 750 µm." figureDoi="http://doi.org/10.5281/zenodo.13158763" httpUri="https://zenodo.org/record/13158763/files/figure.png" pageId="83" pageNumber="84">84 B.</figureCitation>
</paragraph>
<paragraph id="733CE2251817FF8E47E432AEABEFEA58" blockId="83.[123,1175,220,1725]" lastBlockId="84.[123,1174,220,1110]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
<materialsCitation id="C3EBE8781817FF8947E432AEA9D0EAB4" collectingDate="2001-03-04" collectingDateMax="2001-03-19" collectingDateMin="2001-03-04" collectionCode="CAS" collectorName="Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="1125" latitude="-12.520278" location="Parc National Montagne d'Ambre" longLatPrecision="21" longitude="49.17917" pageId="83" pageNumber="84" specimenCode="CASENT9001832" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
M
<emphasis id="41F73E371817FF89478D32AEAAA6EAF4" bold="true" box="[194,401,929,952]" pageId="83" pageNumber="84">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51817FF8946EF32AEA955EAF4" box="[416,610,929,952]" name="Madagascar" pageId="83" pageNumber="84">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371817FF89453A32AEA850EAF4" box="[629,871,929,952]" italics="true" pageId="83" pageNumber="84">
<collectingRegion id="B1472CC71817FF89453A32AEA850EAF4" box="[629,871,929,952]" country="Madagascar" name="Antsiranana" pageId="83" pageNumber="84">Antsiranana Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1817FF89443132AEABE9EA94" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF89443132AEABE9EA94" country="Madagascar" latitude="-12.520278" longLatPrecision="21" longitude="49.17917" name="Parc National Montagne d'Ambre" pageId="83" pageNumber="84" stateProvince="Antsiranana">Parc National Montagne d’Ambre</location>
,
<geoCoordinate id="16B784E21817FF8947A732CEAA55EA94" box="[232,354,961,984]" degrees="12" direction="south" minutes="31" orientation="latitude" pageId="83" pageNumber="84" precision="15" seconds="13" value="-12.520278">12°31ʹ13ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21817FF89462032CEAADBEA94" box="[367,492,961,984]" degrees="49" direction="east" minutes="10" orientation="longitude" pageId="83" pageNumber="84" precision="15" seconds="45" value="49.17917">49°10ʹ45ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01817FF89456232CEA941EA94" box="[557,630,961,984]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="83" pageNumber="84" unit="m" value="1125.0">
<elevation id="F8AE05161817FF89456232CEA941EA94" box="[557,630,961,984]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="83" pageNumber="84" unit="m" value="1125.0">1125m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321817FF8945CF32CEA833EA94" box="[640,772,961,984]" pageId="83" pageNumber="84">malaise trap</collectingMethod>
,
<date id="073DC4E51817FF89444032CEA8F8EA94" box="[783,975,961,984]" pageId="83" pageNumber="84" value="2001-03-04" valueMax="2001-03-19" valueMin="2001-03-04">
<collectingDate id="17793D0D1817FF89444032CEA8F8EA94" box="[783,975,961,984]" pageId="83" pageNumber="84" value="2001-03-04" valueMax="2001-03-19" valueMin="2001-03-04">4–19 March 2001</collectingDate>
</date>
, M.
<collectorName id="DE7687F31817FF8944B132CEAF0DEA94" box="[1022,1082,961,984]" pageId="83" pageNumber="84">Irwin</collectorName>
,
<collectorName id="DE7687F31817FF89430A32CEABFCEAB4" pageId="83" pageNumber="84">R. Harin’Hala</collectorName>
[MA-01-01D-05] (
<specimenCode id="23254A5E1817FF8946D032EEA95EEAB4" box="[415,617,993,1016]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9001832</specimenCode>
,
<specimenCount id="658529AC1817FF89453932EEA9A0EAB4" box="[630,663,993,1016]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF8945E832EEA9E8EAB4" box="[679,735,993,1016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="83" pageNumber="84" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781817FF8945BB32EEAADBED15" collectingDate="1997-09-12" collectingDateMax="1997-09-17" collectingDateMin="1997-09-12" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="900" location="Camp" municipality="Angodongodona" pageId="83" pageNumber="84" specimenCode="CASENT9006080, CASENT9006070" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa">
<emphasis id="41F73E371817FF8945BB32EEA8DBEAB4" box="[756,1004,993,1016]" italics="true" pageId="83" pageNumber="84">
<collectingRegion id="B1472CC71817FF8945BB32EEA8DBEAB4" box="[756,1004,993,1016]" country="Madagascar" name="Fianarantsoa" pageId="83" pageNumber="84">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<locationDeviation id="628EFB071817FF8944B332EEAB9BED54" location="Ivohibe" pageId="83" pageNumber="84">
<quantity id="B47B4FC01817FF8944B332EEAF04EAB4" box="[1020,1075,993,1016]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="83" pageNumber="84" unit="km" value="9.0">9 km</quantity>
NE Ivohibe
</locationDeviation>
,
<quantity id="B47B4FC01817FF8947F9350EAA37ED54" box="[182,256,1025,1048]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="83" pageNumber="84" unit="km" value="6.5">6.5 km</quantity>
ESE
<collectingMunicipality id="9358785F1817FF894676350EAAD9ED54" box="[313,494,1025,1048]" pageId="83" pageNumber="84">Angodongodona</collectingMunicipality>
, 22°25.6ʹS, 046°56.3ʹE,
<location id="765CB4FE1817FF8945BD350EA805ED54" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF8945BD350EA805ED54" box="[754,818,1025,1048]" country="Madagascar" municipality="Angodongodona" name="Camp" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Camp</location>
#5, elev.
<quantity id="B47B4FC01817FF8944DA350EA8E2ED54" box="[917,981,1025,1048]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="83" pageNumber="84" unit="m" value="900.0">
<elevation id="F8AE05161817FF8944DA350EA8E2ED54" box="[917,981,1025,1048]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="83" pageNumber="84" unit="m" value="900.0">900m</elevation>
</quantity>
, undisturbed rainforest,
<collectingMethod id="AAC29A321817FF89478A352EAA77ED74" box="[197,320,1057,1080]" pageId="83" pageNumber="84">pitfall traps</collectingMethod>
#13–15,
<date id="073DC4E51817FF8946EB352EA9A9ED74" box="[420,670,1057,1080]" pageId="83" pageNumber="84" value="1997-09-12" valueMax="1997-09-17" valueMin="1997-09-12">
<collectingDate id="17793D0D1817FF8946EB352EA9A9ED74" box="[420,670,1057,1080]" pageId="83" pageNumber="84" value="1997-09-12" valueMax="1997-09-17" valueMin="1997-09-12">12–17 September 1997</collectingDate>
</date>
,
<collectorName id="DE7687F31817FF8945E6352EA819ED74" box="[681,814,1057,1080]" pageId="83" pageNumber="84">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1817FF894473352EAF31ED74" box="[828,1030,1057,1080]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9006080</specimenCode>
,
<specimenCount id="658529AC1817FF89435E352EAF05ED74" box="[1041,1074,1057,1080]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF894370352EAFB9ED74" box="[1087,1166,1057,1080]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
) (
<specimenCode id="23254A5E1817FF8947CD354DAA7BED15" box="[130,332,1090,1113]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9006070</specimenCode>
,
<specimenCount id="658529AC1817FF894612354DAA48ED15" box="[349,383,1090,1113]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF8946DB354DAAD1ED15" box="[404,486,1090,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF8946B0354DA861EDF5" collectingDate="2003-10-26" collectingDateMax="2003-10-31" collectingDateMin="2003-10-26" collectionCode="FMNH, FHNH" collectorName="Olsen" country="Madagascar" county="Parc National de Midousy" elevation="875" location="Andrauorena River" municipality="Mt. Papango" pageId="83" pageNumber="84" specimenCode="CASENT9064783, CASENT9062276, CASENT9062277, CASENT9062778" specimenCount="4" specimenCount-male="4" stateProvince="Fianarantsoa">
<collectingCounty id="9A5D9AA91817FF8946B0354DA81EED15" box="[511,809,1090,1113]" pageId="83" pageNumber="84">Parc National de Midousy</collectingCounty>
, NE slope
<collectingMunicipality id="9358785F1817FF8944F9354DAF7EED15" box="[950,1097,1090,1113]" pageId="83" pageNumber="84">Mt. Papango</collectingMunicipality>
, along
<location id="765CB4FE1817FF894734356DAA70ED35" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF894734356DAA70ED35" box="[123,327,1122,1145]" country="Madagascar" county="Parc National de Midousy" municipality="Mt. Papango" name="Andrauorena River" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Andrauorena River</location>
,
<locationDeviation id="628EFB071817FF894619356DA910ED35" box="[342,551,1122,1145]" location="Befotaka" pageId="83" pageNumber="84">
<quantity id="B47B4FC01817FF894619356DAABBED35" box="[342,396,1122,1145]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="83" pageNumber="84" unit="km" value="2.0">2 km</quantity>
SW Befotaka
</locationDeviation>
, 23°50.1ʹS, 46°57.8ʹE, elev.
<quantity id="B47B4FC01817FF894414356DA8ADED35" box="[859,922,1122,1145]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="83" pageNumber="84" unit="m" value="875.0">
<elevation id="F8AE05161817FF894414356DA8ADED35" box="[859,922,1122,1145]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="83" pageNumber="84" unit="m" value="875.0">875m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321817FF8944E9356DA8DFED35" box="[934,1000,1122,1145]" pageId="83" pageNumber="84">pitfall</collectingMethod>
,
<date id="073DC4E51817FF8944BA356DAB85EDD5" pageId="83" pageNumber="84" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">
<collectingDate id="17793D0D1817FF8944BA356DAB85EDD5" pageId="83" pageNumber="84" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">26–31 October 2003</collectingDate>
</date>
, L. E.
<collectorName id="DE7687F31817FF894654358DAA6BEDD5" box="[283,348,1154,1177]" pageId="83" pageNumber="84">Olsen</collectorName>
, (
<specimenCode id="23254A5E1817FF894634358DA97CEDD5" box="[379,587,1154,1177]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9064783</specimenCode>
,
<specimenCount id="658529AC1817FF89452D358DA9B3EDD5" box="[610,644,1154,1177]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF8945D3358DA9DAEDD5" box="[668,749,1154,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1817FF89445B358DA8D4EDD5" box="[788,995,1154,1177]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9062276</specimenCode>
,
<specimenCount id="658529AC1817FF8944B4358DAF2AEDD5" box="[1019,1053,1154,1177]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF89437A358DAFBEEDD5" box="[1077,1161,1154,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1817FF8947CD35ADAA7CEDF5" box="[130,331,1186,1209]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9062277</specimenCode>
,
<specimenCount id="658529AC1817FF89461635ADAA4DEDF5" box="[345,378,1186,1209]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF8946C435ADAAEBEDF5" box="[395,476,1186,1209]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1817FF8946B635ADA9F5EDF5" box="[505,706,1186,1209]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9062778</specimenCode>
,
<specimenCount id="658529AC1817FF89459F35ADA9C6EDF5" box="[720,753,1186,1209]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF89444D35ADA878EDF5" box="[770,847,1186,1209]" pageId="83" pageNumber="84">FHNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF89442B35ADA860EDB6" collectingDate="1992-05-09" collectingDateMax="1992-05-10" collectingDateMin="1992-05-09" collectionCode="MCZ, CAS" collectorName="V. and B. Roth &amp; S. Kariko" country="Madagascar" location="Vatoharanana" municipality="Ranomafana" pageId="83" pageNumber="84" specimenCode="CASENT9025468, CASENT9006075" specimenCount="2" specimenCount-female="2" stateProvince="Fianarantsoa">
Parc Nacional
<collectingMunicipality id="9358785F1817FF89434735ADAFA5EDF5" box="[1032,1170,1186,1209]" pageId="83" pageNumber="84">Ranomafana</collectingMunicipality>
,
<location id="765CB4FE1817FF89473435CCAA27ED96" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF89473435CCAA27ED96" box="[123,272,1219,1242]" country="Madagascar" municipality="Ranomafana" name="Vatoharanana" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Vatoharanana</location>
, ca. 21º12ʹS, 47º27ʹE,
<date id="073DC4E51817FF89457835CCA9CDED96" box="[567,762,1219,1242]" pageId="83" pageNumber="84" value="1992-05-09" valueMax="1992-05-10" valueMin="1992-05-09">
<collectingDate id="17793D0D1817FF89457835CCA9CDED96" box="[567,762,1219,1242]" pageId="83" pageNumber="84" value="1992-05-09" valueMax="1992-05-10" valueMin="1992-05-09">9–10 May 1992</collectingDate>
</date>
,
<collectorName id="DE7687F31817FF89445E35CCA8E7ED96" box="[785,976,1219,1242]" pageId="83" pageNumber="84">V. and B. Roth</collectorName>
, and
<collectorName id="DE7687F31817FF89436E35CCAFA2ED96" box="[1057,1173,1219,1242]" pageId="83" pageNumber="84">S. Kariko</collectorName>
(
<specimenCode id="23254A5E1817FF8947CD35ECAA7CEDB6" box="[130,331,1251,1274]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9025468</specimenCode>
,
<specimenCount id="658529AC1817FF89462C35ECAAB3EDB6" box="[355,388,1251,1274]" count="1" pageId="83" pageNumber="84" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01817FF8946D335ECAAEFEDB6" box="[412,472,1251,1274]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="83" pageNumber="84" type="Museum">MCZ</collectionCode>
), (
<specimenCode id="23254A5E1817FF8946B035ECA9FFEDB6" box="[511,712,1251,1274]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9006075</specimenCode>
,
<specimenCount id="658529AC1817FF8945AF35ECA836EDB6" box="[736,769,1251,1274]" count="1" pageId="83" pageNumber="84" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01817FF89445635ECA878EDB6" box="[793,847,1251,1274]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="83" pageNumber="84" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF89442135ECAF0DEC76" collectingDate="1992-05-05" collectionCode="MCZ" collectorName="V. and B. Roth &amp; S. Kariko" country="Madagascar" location="Vatoharanana" pageId="83" pageNumber="84" specimenCode="CASENT9006078" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<location id="765CB4FE1817FF89442135ECA8C9EDB6" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF89442135ECA8C9EDB6" box="[878,1022,1251,1274]" country="Madagascar" name="Vatoharanana" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Vatoharanana</location>
(coordinates approximate) 21º16ʹS, 47º28ʹE, general collecting, [“Ten-1, KARIKO-ROTH VOUCHER COLLECTION”],
<date id="073DC4E51817FF89465F342CAAA0EC76" box="[272,407,1315,1338]" pageId="83" pageNumber="84" value="1992-05-05">
<collectingDate id="17793D0D1817FF89465F342CAAA0EC76" box="[272,407,1315,1338]" pageId="83" pageNumber="84" value="1992-05-05">5 May 1992</collectingDate>
</date>
,
<collectorName id="DE7687F31817FF8946ED342CA977EC76" box="[418,576,1315,1338]" pageId="83" pageNumber="84">V. and B. Roth</collectorName>
and
<collectorName id="DE7687F31817FF894538342CA9E8EC76" box="[631,735,1315,1338]" pageId="83" pageNumber="84">S. Kariko</collectorName>
(
<specimenCode id="23254A5E1817FF8945A1342CA880EC76" box="[750,951,1315,1338]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9006078</specimenCode>
,
<specimenCount id="658529AC1817FF89448B342CA8D2EC76" box="[964,997,1315,1338]" count="1" pageId="83" pageNumber="84" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01817FF8944BB342CAF04EC76" box="[1012,1075,1315,1338]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="83" pageNumber="84" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF89430A342CA906ECD7" collectingDate="2000-10-03" collectingDateMax="2000-10-09" collectingDateMin="2000-10-03" collectionCode="FMNH" collectorName="SW Ranomafana &amp; S. Goodman" country="Madagascar" elevation="1025" location="Vatoharanana" pageId="83" pageNumber="84" specimenCode="CASENT9064665" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<location id="765CB4FE1817FF89430A342CABF2EC16" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF89430A342CABF2EC16" country="Madagascar" name="Vatoharanana" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Vatoharanana</location>
, 4.0 km
<collectorName id="DE7687F31817FF89466F344CAAE2EC16" box="[288,469,1347,1370]" pageId="83" pageNumber="84">SW Ranomafana</collectorName>
(ville), 21°14.7ʹS, 047°26.0ʹE, elev.
<quantity id="B47B4FC01817FF894412344CA89DEC16" box="[861,938,1347,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="83" pageNumber="84" unit="m" value="1025.0">
<elevation id="F8AE05161817FF894412344CA89DEC16" box="[861,938,1347,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="83" pageNumber="84" unit="m" value="1025.0">1025m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321817FF8944FB344CAF36EC16" box="[948,1025,1347,1370]" pageId="83" pageNumber="84">pitfalls</collectingMethod>
, slightly transitional lowland to lower montane rainforest,
<date id="073DC4E51817FF89453F346BA80EEC37" box="[624,825,1380,1403]" pageId="83" pageNumber="84" value="2000-10-03" valueMax="2000-10-09" valueMin="2000-10-03">
<collectingDate id="17793D0D1817FF89453F346BA80EEC37" box="[624,825,1380,1403]" pageId="83" pageNumber="84" value="2000-10-03" valueMax="2000-10-09" valueMin="2000-10-03">3–9 October 2000</collectingDate>
</date>
, coll.
<collectorName id="DE7687F31817FF894431346BAF32EC37" box="[894,1029,1380,1403]" pageId="83" pageNumber="84">S. Goodman</collectorName>
[
<collectionCode id="15927AE01817FF89435A346BAF51EC37" box="[1045,1126,1380,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="83" pageNumber="84" type="Museum">FMNH</collectionCode>
#00- 225] (
<specimenCode id="23254A5E1817FF89478C348BAABBECD7" box="[195,396,1412,1435]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9064665</specimenCode>
,
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)
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,
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,
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231°SW Ranomafana
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,
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, elev.
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, montane rainforest, general collecting at night,
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,
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[BLF8395] (
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), [
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] (
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,
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)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF89451334EAA9C0EF10" collectingDate="1998-04-26" collectingDateMax="1998-04-27" collectingDateMin="1998-04-26" collectionCode="CAS" collectorName="C. Griswold &amp; D. Kavanaugh &amp; N. Penny &amp; D. Ubick &amp; Raherilalao, J. &amp; Schweikert &amp; Ranorainarisoa &amp; Dong Lin" country="Madagascar" elevation="1000" location="Vohiparara" municipality="Ranomafana" pageId="83" pageNumber="84" specimenCode="CASENT9006058" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
Parc National
<collectingMunicipality id="9358785F1817FF8945B434EAA8B2ECB0" box="[763,901,1509,1532]" pageId="83" pageNumber="84">Ranomafana</collectingMunicipality>
,
<location id="765CB4FE1817FF8944DD34EAAF31ECB0" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF8944DD34EAAF31ECB0" box="[914,1030,1509,1532]" country="Madagascar" municipality="Ranomafana" name="Vohiparara" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Vohiparara</location>
, piste touristique, 21º13.6ʹS, 47º24.0ʹE, elev. ca.
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<elevation id="F8AE05161817FF894540370AA96BEF50" box="[527,604,1541,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="83" pageNumber="84" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
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<collectingDate id="17793D0D1817FF894526370AA81BEF50" box="[617,812,1541,1564]" pageId="83" pageNumber="84" value="1998-04-26" valueMax="1998-04-27" valueMin="1998-04-26">26–27 April 1998</collectingDate>
</date>
,
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,
<collectorName id="DE7687F31817FF894480370AAF59EF50" box="[975,1134,1541,1564]" pageId="83" pageNumber="84">D. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31817FF894334370AAB8DEF70" pageId="83" pageNumber="84">N. Penny</collectorName>
,
<collectorName id="DE7687F31817FF894788372AAA1BEF70" box="[199,300,1573,1596]" pageId="83" pageNumber="84">D. Ubick</collectorName>
, M.
<collectorName id="DE7687F31817FF894613372AAAC1EF70" box="[348,502,1573,1596]" pageId="83" pageNumber="84">Raherilalao, J.</collectorName>
,
<collectorName id="DE7687F31817FF89454C372AA94CEF70" box="[515,635,1573,1596]" pageId="83" pageNumber="84">Schweikert</collectorName>
and S.
<collectorName id="DE7687F31817FF894584372AA85BEF70" box="[715,876,1573,1596]" pageId="83" pageNumber="84">Ranorainarisoa</collectorName>
[
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<collectorName id="DE7687F31817FF894433372AA8D5EF70" box="[892,994,1573,1596]" pageId="83" pageNumber="84">Dong Lin</collectorName>
nest photos, this publication,
<figureCitation id="EBB8FEA01817FF89464D374AAAAAEF10" box="[258,413,1605,1628]" captionStart="FIGURE" captionStartId="106.[155,167,815,834]" captionTargetBox="[120,1176,216,796]" captionTargetId="figure-11@106.[120,1176,216,796]" captionTargetPageId="106" captionText="FIGURE 2. Uduba spp., nests and live individuals. A, C, U. schlingeri, ♀, CASENT9006058. B, D, U. evanescens (Dahl, 1901), ♀, CASENT9006088. A. Tubular nest surmounted by funnel of cribellate silk, Ranomafana. B. Nest opening on soil surface, Ranomafana. C. Tubular nest, Ranomafana. D. Nest opening on soil surface, Ranomafana. E. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. A–D. Photos by Dong Lin, Ranomafana, April 1998. E. Photo by Nikolaj Scharff, November 1993. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13158560" httpUri="https://zenodo.org/record/13158560/files/figure.png" pageId="83" pageNumber="84">Figures 2 A, C</figureCitation>
</emphasis>
] (
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,
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)
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;
<materialsCitation id="C3EBE8781817FF89444E374AA891EFD1" collectingDate="1998-04-09" collectingDateMax="1998-04-30" collectingDateMin="1998-04-09" collectionCode="CAS" collectorName="C. Griswold &amp; D. Kavanaugh &amp; N. Penny &amp; D. Ubick &amp; M. Raherilalao &amp; Schweikert &amp; Ranorainarisoa" country="Madagascar" location="Research" municipality="Talatakely" pageId="83" pageNumber="84" specimenCode="CASENT9006053" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingMunicipality id="9358785F1817FF89444E374AA85DEF10" box="[769,874,1605,1628]" pageId="83" pageNumber="84">Talatakely</collectingMunicipality>
,
<location id="765CB4FE1817FF894437374AA8EDEF10" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF894437374AA8EDEF10" box="[888,986,1605,1628]" country="Madagascar" municipality="Talatakely" name="Research" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Research</location>
cabin, 21º14.9ʹS, 47º25.6ʹE,
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,
<collectorName id="DE7687F31817FF8946FF3769A901EF31" box="[432,566,1638,1661]" pageId="83" pageNumber="84">C. Griswold</collectorName>
,
<collectorName id="DE7687F31817FF89450D3769A9D7EF31" box="[578,736,1638,1661]" pageId="83" pageNumber="84">D. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31817FF8945A33769A87AEF31" box="[748,845,1638,1661]" pageId="83" pageNumber="84">N. Penny</collectorName>
,
<collectorName id="DE7687F31817FF8944133769A8F5EF31" box="[860,962,1638,1661]" pageId="83" pageNumber="84">D. Ubick</collectorName>
,
<collectorName id="DE7687F31817FF8944813769AF45EF31" box="[974,1138,1638,1661]" pageId="83" pageNumber="84">M. Raherilalao</collectorName>
, J.,
<collectorName id="DE7687F31817FF8947343789ABC4EFD1" box="[123,243,1670,1693]" pageId="83" pageNumber="84">Schweikert</collectorName>
and S.
<collectorName id="DE7687F31817FF8946093789AAD0EFD1" box="[326,487,1670,1693]" pageId="83" pageNumber="84">Ranorainarisoa</collectorName>
(
<typeStatus id="AC385C871817FF8946B93789A96FEFD1" box="[502,600,1670,1693]" pageId="83" pageNumber="84" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1817FF89452C3789A81CEFD1" box="[611,811,1670,1693]" collectionCode="CASENT" pageId="83" pageNumber="84">CASENT9006053</specimenCode>
,
<specimenCount id="658529AC1817FF8944793789A86FEFD1" box="[822,856,1670,1693]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01817FF8944293789A8A9EFD1" box="[870,926,1670,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="83" pageNumber="84" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781817FF8E44FE3789AFBEE85F" collectingDate="1998-04-09" collectingDateMax="1998-04-30" collectingDateMin="1998-04-09" collectionCode="CAS" collectorName="C. E. Griswold &amp; D. H. Kavanaugh &amp; N. D. Penny &amp; M. J. Raherilalao &amp; Ranorianarisoa &amp; Schweikert &amp; Ubick" country="Madagascar" lastPageId="84" lastPageNumber="85" location="Talatakely" pageId="83" pageNumber="84" specimenCode="CASENT9006054, CASENT9006055, CASENT9006057, CASENT9006051" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="765CB4FE1817FF8944FE3789AF2DEFD1" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1817FF8944FE3789AF2DEFD1" box="[945,1050,1670,1693]" country="Madagascar" name="Talatakely" pageId="83" pageNumber="84" stateProvince="Fianarantsoa">Talatakely</location>
, 21º14.9ʹS, 47º25.6ʹE,
<date id="073DC4E51817FF8947BE37A9AA94EFF1" box="[241,419,1702,1725]" pageId="83" pageNumber="84" value="1998-04-09" valueMax="1998-04-30" valueMin="1998-04-09">
<collectingDate id="17793D0D1817FF8947BE37A9AA94EFF1" box="[241,419,1702,1725]" pageId="83" pageNumber="84" value="1998-04-09" valueMax="1998-04-30" valueMin="1998-04-09">9–30 April 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31817FF8946E037A9A963EFF1" box="[431,596,1702,1725]" pageId="83" pageNumber="84">C. E. Griswold</collectorName>
,
<collectorName id="DE7687F31817FF89452F37A9A828EFF1" box="[608,799,1702,1725]" pageId="83" pageNumber="84">D. H. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31817FF89446437A9A89AEFF1" box="[811,941,1702,1725]" pageId="83" pageNumber="84">N. D. Penny</collectorName>
,
<collectorName id="DE7687F31817FF8944F337A9AF4EEFF1" box="[956,1145,1702,1725]" pageId="83" pageNumber="84">M. J. Raherilalao</collectorName>
, J. S.
<collectorName id="DE7687F31810FF8E47E531D3AA6EE9BF" box="[170,345,220,243]" pageId="84" pageNumber="85">Ranorianarisoa</collectorName>
, J.
<collectorName id="DE7687F31810FF8E46EC31D3A912E9BF" box="[419,549,220,243]" pageId="84" pageNumber="85">Schweikert</collectorName>
and D.
<collectorName id="DE7687F31810FF8E45FD31D3A9C0E9BF" box="[690,759,220,243]" pageId="84" pageNumber="85">Ubick</collectorName>
(
<specimenCode id="23254A5E1810FF8E445831D3A8DCE9BF" box="[791,1003,220,243]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006054</specimenCode>
,
<specimenCount id="658529AC1810FF8E434431D3AF19E9BF" box="[1035,1070,220,243]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
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), (
<specimenCode id="23254A5E1810FF8E47CD31F3AA7CE85F" box="[130,331,252,275]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006055</specimenCode>
,
<specimenCount id="658529AC1810FF8E461331F3AA4AE85F" box="[348,381,252,275]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E46DF31F3AAF0E85F" box="[400,455,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E1810FF8E46A831F3A987E85F" box="[487,688,252,275]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006057</specimenCode>
,
<specimenCount id="658529AC1810FF8E458E31F3A9D5E85F" box="[705,738,252,275]" count="1" pageId="84" pageNumber="85" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01810FF8E45BA31F3A81BE85F" box="[757,812,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
),
<specimenCode id="23254A5E1810FF8E440A31F3AF3AE85F" box="[837,1037,252,275]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006051</specimenCode>
,
<specimenCount id="658529AC1810FF8E435231F3AF09E85F" box="[1053,1086,252,275]" count="3" pageId="84" pageNumber="85" type="male">3♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E431E31F3AFBEE85F" box="[1105,1161,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
</materialsCitation>
);
<materialsCitation id="C3EBE8781810FF8E47343013AAF2E83F" collectingDate="1998-04-05" collectingDateMax="1998-04-18" collectingDateMin="1998-04-05" collectionCode="CAS" collectorName="C. E. Griswold &amp; D. H. Kavanaugh &amp; N. D. Penny &amp; M. J. Raherilalao &amp; J. S. Ranorianarisoa &amp; Schweikert &amp; Ubick" country="Madagascar" location="Talatakely" pageId="84" pageNumber="85" specimenCode="CASENT9006056" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="765CB4FE1810FF8E47343013ABD3E87F" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E47343013ABD3E87F" box="[123,228,284,307]" country="Madagascar" name="Talatakely" pageId="84" pageNumber="85" stateProvince="Fianarantsoa">Talatakely</location>
, 21º14.9ʹS, 47º25.6ʹE, “trail F, burrow at tree base”
<date id="073DC4E51810FF8E445E3013A8F5E87F" box="[785,962,284,307]" pageId="84" pageNumber="85" value="1998-04-05" valueMax="1998-04-18" valueMin="1998-04-05">
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,
<collectorName id="DE7687F31810FF8E44823013AF46E87F" box="[973,1137,284,307]" pageId="84" pageNumber="85">C. E. Griswold</collectorName>
,
<collectorName id="DE7687F31810FF8E43333013AA2EE81F" pageId="84" pageNumber="85">D. H. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31810FF8E466A3033AA9EE81F" box="[293,425,316,339]" pageId="84" pageNumber="85">N. D. Penny</collectorName>
,
<collectorName id="DE7687F31810FF8E46F73033A941E81F" box="[440,630,316,339]" pageId="84" pageNumber="85">M. J. Raherilalao</collectorName>
,
<collectorName id="DE7687F31810FF8E45CD3033A86AE81F" box="[642,861,316,339]" pageId="84" pageNumber="85">J. S. Ranorianarisoa</collectorName>
, J.
<collectorName id="DE7687F31810FF8E44CC3033A8CCE81F" box="[899,1019,316,339]" pageId="84" pageNumber="85">Schweikert</collectorName>
and D.
<collectorName id="DE7687F31810FF8E431B3033AFA2E81F" box="[1108,1173,316,339]" pageId="84" pageNumber="85">Ubick</collectorName>
(
<specimenCode id="23254A5E1810FF8E47CD3053AA7BE83F" box="[130,332,348,371]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006056</specimenCode>
,
<specimenCount id="658529AC1810FF8E46183053AA4FE83F" box="[343,376,348,371]" count="1" pageId="84" pageNumber="85" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01810FF8E46C93053AA8AE83F" box="[390,445,348,371]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781810FF8E469F3053AF35E8DF" collectingDate="1992-05-31" collectingDateMax="1998-04-18" collectingDateMin="1992-05-31" collectionCode="CAS" collectorName="Ranomafana N. P. &amp; V. Roth" country="Madagascar" location="Ranomafana" municipality="Talatakely" pageId="84" pageNumber="85" specimenCode="CASENT9006062" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingMunicipality id="9358785F1810FF8E469F3053A90EE83F" box="[464,569,348,371]" pageId="84" pageNumber="85">Talatakely</collectingMunicipality>
, 21º14.9ʹS, 47º25.6ʹE,
<date id="073DC4E51810FF8E447E3053A8D5E83F" box="[817,994,348,371]" pageId="84" pageNumber="85" value="1998-04-05" valueMax="1998-04-18" valueMin="1998-04-05">
<collectingDate id="17793D0D1810FF8E447E3053A8D5E83F" box="[817,994,348,371]" pageId="84" pageNumber="85" value="1998-04-05" valueMax="1998-04-18" valueMin="1998-04-05">5–18 April 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31810FF8E44A13053ABBDE8DF" pageId="84" pageNumber="85">Ranomafana N. P.</collectorName>
, township of
<location id="765CB4FE1810FF8E466D3073AA9BE8DF" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E466D3073AA9BE8DF" box="[290,428,380,403]" country="Madagascar" municipality="Talatakely" name="Ranomafana" pageId="84" pageNumber="85" stateProvince="Fianarantsoa">Ranomafana</location>
,
<date id="073DC4E51810FF8E46F73073A97BE8DF" box="[440,588,380,403]" pageId="84" pageNumber="85" value="1992-05-31">
<collectingDate id="17793D0D1810FF8E46F73073A97BE8DF" box="[440,588,380,403]" pageId="84" pageNumber="85" value="1992-05-31">31 May 1992</collectingDate>
</date>
,
<collectorName id="DE7687F31810FF8E45183073A99EE8DF" box="[599,681,380,403]" pageId="84" pageNumber="85">V. Roth</collectorName>
, (
<specimenCode id="23254A5E1810FF8E45F33073A8B1E8DF" box="[700,902,380,403]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006062</specimenCode>
,
<specimenCount id="658529AC1810FF8E44DD3073A884E8DF" box="[914,947,380,403]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E448D3073A8CDE8DF" box="[962,1018,380,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781810FF8E43403073A83CE89F" collectingDate="1992-03-23" collectionCode="CAS" collectorName="Ranomafana N. P. &amp; Roth" country="Madagascar" pageId="84" pageNumber="85" specimenCode="CASENT9006079" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectorName id="DE7687F31810FF8E43403073AB93E8FF" pageId="84" pageNumber="85">Ranomafana N.P.</collectorName>
, at camp (coordinates approximate) 21º16ʹS, 47º28ʹE, general collecting,
<date id="073DC4E51810FF8E44AD3093AFA6E8FF" box="[994,1169,412,435]" pageId="84" pageNumber="85" value="1992-03-23">
<collectingDate id="17793D0D1810FF8E44AD3093AFA6E8FF" box="[994,1169,412,435]" pageId="84" pageNumber="85" value="1992-03-23">23 March 1992</collectingDate>
</date>
, V. and B.
<collectorName id="DE7687F31810FF8E47A830B3AA2CE89F" box="[231,283,444,467]" pageId="84" pageNumber="85">Roth</collectorName>
(
<typeStatus id="AC385C871810FF8E466430B3AABAE89F" box="[299,397,444,467]" pageId="84" pageNumber="85" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1810FF8E46D530B3A955E89F" box="[410,610,444,467]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006079</specimenCode>
,
<specimenCount id="658529AC1810FF8E452130B3A9A7E89F" box="[622,656,444,467]" count="1" pageId="84" pageNumber="85" type="female">1♀</specimenCount>
, 3jj,
<collectionCode id="15927AE01810FF8E458430B3A834E89F" box="[715,771,444,467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="84" pageNumber="85" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781810FF8E445730B3A84EE8BF" collectingDate="1988-10-22" collectingDateMax="1988-10-31" collectingDateMin="1988-10-22" collectionCode="USNM" collectorName="Steiner" country="Madagascar" elevation="1100" location="7 km W Ranomafana" pageId="84" pageNumber="85" specimenCode="CASENT9006061" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="765CB4FE1810FF8E445730B3AF35E89F" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E445730B3AF35E89F" box="[792,1026,444,467]" country="Madagascar" name="7 km W Ranomafana" pageId="84" pageNumber="85" stateProvince="Fianarantsoa">
<quantity id="B47B4FC01810FF8E445730B3A878E89F" box="[792,847,444,467]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="84" pageNumber="85" unit="km" value="7.0">7 km</quantity>
W Ranomafana
</location>
, elev.
<quantity id="B47B4FC01810FF8E430830B3AFA5E89F" box="[1095,1170,444,467]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="84" pageNumber="85" unit="m" value="1100.0">
<elevation id="F8AE05161810FF8E430830B3AFA5E89F" box="[1095,1170,444,467]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="84" pageNumber="85" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<date id="073DC4E51810FF8E473430D3AA5CE8BF" box="[123,363,476,499]" pageId="84" pageNumber="85" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">
<collectingDate id="17793D0D1810FF8E473430D3AA5CE8BF" box="[123,363,476,499]" pageId="84" pageNumber="85" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">22–31 October 1988</collectingDate>
</date>
, W.
<collectorName id="DE7687F31810FF8E46E730D3AAC3E8BF" box="[424,500,476,499]" pageId="84" pageNumber="85">Steiner</collectorName>
(
<specimenCode id="23254A5E1810FF8E454430D3A9E3E8BF" box="[523,724,476,499]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006061</specimenCode>
,
<specimenCount id="658529AC1810FF8E45A730D3A83DE8BF" box="[744,778,476,499]" count="2" pageId="84" pageNumber="85" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E446F30D3A845E8BF" box="[800,882,476,499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781810FF8E44C330D3A98FEB18" collectingDate="1998-03-09" collectingDateMax="1998-03-15" collectingDateMin="1998-03-09" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1050" location="Antanimena" municipality="Foret Analavelona" pageId="84" pageNumber="85" specimenCode="CASENT9065671" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="41F73E371810FF8E44C330D3AF72E8BF" box="[908,1093,476,499]" italics="true" pageId="84" pageNumber="85">
<collectingRegion id="B1472CC71810FF8E44C330D3AF72E8BF" box="[908,1093,476,499]" country="Madagascar" name="Toliara" pageId="84" pageNumber="85">Toliara Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1810FF8E431230D3AA32EB58" pageId="84" pageNumber="85">Forêt Analavelona</collectingMunicipality>
,
<location id="765CB4FE1810FF8E464130F2AAA4EB58" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E464130F2AAA4EB58" box="[270,403,509,532]" country="Madagascar" municipality="Foret Analavelona" name="Antanimena" pageId="84" pageNumber="85" stateProvince="Toliara">Antanimena</location>
,
<locationDeviation id="628EFB071810FF8E46EF30F2A9F4EB58" box="[416,707,509,532]" location="Andranoheza" pageId="84" pageNumber="85">
<quantity id="B47B4FC01810FF8E46EF30F2AACFEB58" box="[416,504,509,532]" metricMagnitude="4" metricUnit="m" metricValue="1.25" pageId="84" pageNumber="85" unit="km" value="12.5">12.5 km</quantity>
NW Andranoheza
</locationDeviation>
, 22°40.7ʹS, 44°11.5ʹE, elev.
<quantity id="B47B4FC01810FF8E44B730F2AF73EB58" box="[1016,1092,509,532]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="84" pageNumber="85" unit="m" value="1050.0">
<elevation id="F8AE05161810FF8E44B730F2AF73EB58" box="[1016,1092,509,532]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="84" pageNumber="85" unit="m" value="1050.0">1050m</elevation>
</quantity>
, transitional mid-altitude forest with elements of eastern and western forests,
<collectingMethod id="AAC29A321810FF8E44C93312AF33EB78" box="[902,1028,541,564]" pageId="84" pageNumber="85">pitfall traps</collectingMethod>
,
<date id="073DC4E51810FF8E435D3312AB84EB18" pageId="84" pageNumber="85" value="1998-03-09" valueMax="1998-03-15" valueMin="1998-03-09">
<collectingDate id="17793D0D1810FF8E435D3312AB84EB18" pageId="84" pageNumber="85" value="1998-03-09" valueMax="1998-03-15" valueMin="1998-03-09">9–15 March 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31810FF8E478F3332AA70EB18" box="[192,327,573,596]" pageId="84" pageNumber="85">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1810FF8E46183332A917EB18" box="[343,544,573,596]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9065671</specimenCode>
,
<specimenCount id="658529AC1810FF8E45613332A978EB18" box="[558,591,573,596]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E45103332A986EB18" box="[607,689,573,596]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781810FF8E458A3332AADDEBF8" collectingDate="1995-11-07" collectingDateMax="1995-11-17" collectingDateMin="1995-11-07" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1200" location="Reserve Naturelle Integrale d'Andohahela" pageId="84" pageNumber="85" specimenCode="FMNH#12, CASENT9006082, FMNH#71-7, 2, CASENT9006081" specimenCount="2" specimenCount-female="2" stateProvince="Toliara" typeStatus="paratype">
<location id="765CB4FE1810FF8E458A3332AFA5EB18" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E458A3332AFA5EB18" box="[709,1170,573,596]" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="84" pageNumber="85" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
,
<locationDeviation id="628EFB071810FF8E47343352AAB5EB38" box="[123,386,605,628]" location="Eminiminy" pageId="84" pageNumber="85">
<quantity id="B47B4FC01810FF8E47343352ABE5EB38" box="[123,210,605,628]" metricMagnitude="4" metricUnit="m" metricValue="1.35" pageId="84" pageNumber="85" unit="km" value="13.5">13.5 km</quantity>
NW Eminiminy
</locationDeviation>
, parcel #1, camp #3, 24°35.0ʹS, 46°44.1ʹE, elev.
<quantity id="B47B4FC01810FF8E44DF3352A8EBEB38" box="[912,988,605,628]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="84" pageNumber="85" unit="m" value="1200.0">
<elevation id="F8AE05161810FF8E44DF3352A8EBEB38" box="[912,988,605,628]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="84" pageNumber="85" unit="m" value="1200.0">1200m</elevation>
</quantity>
,
<date id="073DC4E51810FF8E44A73352AB83EBD8" pageId="84" pageNumber="85" value="1995-11-07" valueMax="1995-11-17" valueMin="1995-11-07">
<collectingDate id="17793D0D1810FF8E44A73352AB83EBD8" pageId="84" pageNumber="85" value="1995-11-07" valueMax="1995-11-17" valueMin="1995-11-07">7–17 November 1995</collectingDate>
</date>
,
<collectorName id="DE7687F31810FF8E47823372AA52EBD8" box="[205,357,637,660]" pageId="84" pageNumber="85">S. Goodman</collectorName>
[
<specimenCode id="23254A5E1810FF8E46333372AACCEBD8" box="[380,507,637,660]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH#12</specimenCode>
] (
<specimenCode id="23254A5E1810FF8E45513372A9C4EBD8" box="[542,755,637,660]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006082</specimenCode>
,
<specimenCount id="658529AC1810FF8E44433372A819EBD8" box="[780,814,637,660]" count="1" pageId="84" pageNumber="85" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01810FF8E44073372A8ACEBD8" box="[840,923,637,660]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
), [
<specimenCode id="23254A5E1810FF8E448D3372AF56EBD8" box="[962,1121,637,660]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH#71-7</specimenCode>
-
<specimenCode id="23254A5E1810FF8E43263372AF40EBD8" box="[1129,1143,637,660]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">2</specimenCode>
-S] (
<specimenCode id="23254A5E1810FF8E47CD3392AA7CEBF8" box="[130,331,669,692]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006081</specimenCode>
,
<specimenCount id="658529AC1810FF8E46133392AA4AEBF8" box="[348,381,669,692]" count="1" pageId="84" pageNumber="85" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01810FF8E46DE3392AAD4EBF8" box="[401,483,669,692]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781810FF8E46B43392ABE3EA58" collectingDate="1995-10-19" collectingDateMax="1995-10-28" collectingDateMin="1995-10-19" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="440" location="Reserve Naturelle Integrale d'Andohahela" pageId="84" pageNumber="85" specimenCode="CASENT9006063, CASENT9006060, CASENT9006059" specimenCount="4" specimenCount-male="4" stateProvince="Toliara" typeStatus="paratype">
<location id="765CB4FE1810FF8E46B43392A8E3EBF8" LSID="urn:lsid:plazi:treatment:FB2A53331815FF8E473432CFAF01ED1A:765CB4FE1810FF8E46B43392A8E3EBF8" box="[507,980,669,692]" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="84" pageNumber="85" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
,
<locationDeviation id="628EFB071810FF8E44AA3392ABF4EB98" location="Eminiminy" pageId="84" pageNumber="85">
<quantity id="B47B4FC01810FF8E44AA3392AF2CEBF8" box="[997,1051,669,692]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="84" pageNumber="85" unit="km" value="8.0">8 km</quantity>
NW Eminiminy
</locationDeviation>
, parcel #1, camp #1, 24°37.6ʹS, 46°45.9ʹE, elev.
<quantity id="B47B4FC01810FF8E459C33B2A825EB98" box="[723,786,701,724]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="84" pageNumber="85" unit="m" value="440.0">
<elevation id="F8AE05161810FF8E459C33B2A825EB98" box="[723,786,701,724]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="84" pageNumber="85" unit="m" value="440.0">440m</elevation>
</quantity>
,
<date id="073DC4E51810FF8E445233B2A8CAEB98" box="[797,1021,701,724]" pageId="84" pageNumber="85" value="1995-10-19" valueMax="1995-10-28" valueMin="1995-10-19">
<collectingDate id="17793D0D1810FF8E445233B2A8CAEB98" box="[797,1021,701,724]" pageId="84" pageNumber="85" value="1995-10-19" valueMax="1995-10-28" valueMin="1995-10-19">19–28 October 1995</collectingDate>
</date>
,
<collectorName id="DE7687F31810FF8E434733B2AFA6EB98" box="[1032,1169,701,724]" pageId="84" pageNumber="85">S. Goodman</collectorName>
, (
<specimenCode id="23254A5E1810FF8E47CD33D2AA7CEBB8" box="[130,331,733,756]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006063</specimenCode>
,
<specimenCount id="658529AC1810FF8E461033D2AAB7EBB8" box="[351,384,733,756]" count="2" pageId="84" pageNumber="85" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E46D933D2AAD0EBB8" box="[406,487,733,756]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1810FF8E454633D2A9E4EBB8" box="[521,723,733,756]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006060</specimenCode>
,
<specimenCount id="658529AC1810FF8E45A933D2A830EBB8" box="[742,775,733,756]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E445233D2A859EBB8" box="[797,878,733,756]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1810FF8E44DF33D2AF6EEBB8" box="[912,1113,733,756]" collectionCode="CASENT" pageId="84" pageNumber="85">CASENT9006059</specimenCode>
,
<specimenCount id="658529AC1810FF8E432233D2AFB8EBB8" box="[1133,1167,733,756]" count="1" pageId="84" pageNumber="85" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01810FF8E473433F2ABFAEA58" box="[123,205,765,788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="84" pageNumber="85" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251810FF8E47E43211A819EAF9" blockId="84.[123,1174,220,1110]" pageId="84" pageNumber="85">
<emphasis id="41F73E371810FF8E47E43211AA68EA79" bold="true" box="[171,351,798,821]" pageId="84" pageNumber="85">Natural history.</emphasis>
Many records from pitfall traps reveal that males of
<taxonomicName id="B48399A61810FF8E44D53211AF6EEA79" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[922,1113,798,821]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371810FF8E44D53211AF6EEA79" bold="true" box="[922,1113,798,821]" italics="true" pageId="84" pageNumber="85">Uduba schlingeri</emphasis>
</taxonomicName>
wan- der, even up into vegetation where they have been collected in malaise taps. Females of this cribellate species are recorded as burrowing, e.g., at Talatakely where a record states “trail F, burrow at tree base.” They make a vertical tube of silk that may extend above the soil surface, and which may be surmounted by a funnel of cribellate silk (
<figureCitation id="EBB8FEA01810FF8E45D53291A816EAF9" box="[666,801,926,949]" captionStart="FIGURE" captionStartId="106.[155,167,815,834]" captionTargetBox="[120,1176,216,796]" captionTargetId="figure-11@106.[120,1176,216,796]" captionTargetPageId="106" captionText="FIGURE 2. Uduba spp., nests and live individuals. A, C, U. schlingeri, ♀, CASENT9006058. B, D, U. evanescens (Dahl, 1901), ♀, CASENT9006088. A. Tubular nest surmounted by funnel of cribellate silk, Ranomafana. B. Nest opening on soil surface, Ranomafana. C. Tubular nest, Ranomafana. D. Nest opening on soil surface, Ranomafana. E. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. A–D. Photos by Dong Lin, Ranomafana, April 1998. E. Photo by Nikolaj Scharff, November 1993. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13158560" httpUri="https://zenodo.org/record/13158560/files/figure.png" pageId="84" pageNumber="85">Figs. 2 A, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1810FF8E47E432B1AF01ED1A" pageId="84" pageNumber="85" type="distribution">
<paragraph id="733CE2251810FF8E47E432B1AF01ED1A" blockId="84.[123,1174,220,1110]" pageId="84" pageNumber="85">
<emphasis id="41F73E371810FF8E47E432B1AA0AEA99" bold="true" box="[171,317,958,981]" pageId="84" pageNumber="85">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61810FF8E468632B1A9BFEA99" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[457,648,958,981]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371810FF8E468632B1A9BFEA99" bold="true" box="[457,648,958,981]" italics="true" pageId="84" pageNumber="85">Uduba schlingeri</emphasis>
</taxonomicName>
has been collected in rainforests from Montagne d’Ambre from far northern
<collectingCountry id="0B94A2B51810FF8E46EE32D1A915EAB9" box="[417,546,990,1013]" name="Madagascar" pageId="84" pageNumber="85">Madagascar</collectingCountry>
south through the eastern mountains to Andohahela (
<figureCitation id="EBB8FEA01810FF8E431632D1ABBDED59" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="84" pageNumber="85">Maps 5</figureCitation>
, 14). A southwestern collection record of
<taxonomicName id="B48399A61810FF8E451832F1A9DFED59" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[599,744,1022,1045]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371810FF8E451832F1A9DFED59" bold="true" box="[599,744,1022,1045]" italics="true" pageId="84" pageNumber="85">U. schlingeri</emphasis>
</taxonomicName>
from Forêt Analavelona pitfall traps at elevation
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, in “transitional mid-altitude forest with elements of eastern and western forests”, represents a spider collected in a disjunct outlier of typical eastern rainforest vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>

229
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<mods:title id="1C965579F69971E9E491BD3879D03241">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="D8AAE446D824D93E66FE78AC292E53A6">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="9538EDB6A1EAD09FC6A4E49997B3235D">Ubick, Darrell</mods:namePart>
<mods:affiliation id="C1563E703B43430971FA64AD93DFB0BE">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<emphasis id="41F73E37181CFF8247343291AA21EAF9" bold="true" box="[123,278,926,949]" italics="true" pageId="88" pageNumber="89">Uduba volana</emphasis>
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,
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<subSubSection id="3B99B1AE181CFF82473432B1AAF7EA9F" box="[123,448,958,979]" pageId="88" pageNumber="89" type="description">
<paragraph id="733CE225181CFF82473432B1AAF7EA9F" blockId="88.[123,448,926,979]" box="[123,448,958,979]" pageId="88" pageNumber="89">
<figureCitation id="EBB8FEA0181CFF82473432B1ABD9EA9F" box="[123,238,958,979]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="88" pageNumber="89">Figures 7 C</figureCitation>
,
<figureCitation id="EBB8FEA0181CFF8247B732B1AA74EA9F" box="[248,323,958,979]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="88" pageNumber="89">77 D–F</figureCitation>
,
<figureCitation id="EBB8FEA0181CFF82460332B1AAA2EA9F" box="[332,405,958,979]" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="88" pageNumber="89">Maps 5</figureCitation>
, 16.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8247E432EBAAB2ED77" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="733CE225181CFF8247E432EBAAB2ED77" blockId="88.[122,1174,996,1725]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E432EBAA67EAB7" bold="true" box="[171,336,996,1019]" pageId="88" pageNumber="89">Type material.</emphasis>
<materialsCitation id="C3EBE878181CFF82461832EBAAB2ED77" collectingDate="1993-11-02" collectionCode="CAS" collectorName="C. Griswold" country="Madagascar" elevation="1400" location="Ambohimanga" pageId="88" pageNumber="89" specimenCode="CASENT9006072" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="AC385C87181CFF82461832EBAA8DEAB7" box="[343,442,996,1019]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
female (
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) dug from a burrow at
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<elevation id="F8AE0516181CFF8244AD32EBAF1CEAB7" box="[994,1067,996,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="88" pageNumber="89" unit="m" value="1400.0">1400m</elevation>
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elevation from
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,
<collectingRegion id="B1472CC7181CFF824613350BA961ED57" box="[348,598,1028,1051]" country="Madagascar" name="Antananarivo" pageId="88" pageNumber="89">Antananarivo Province</collectingRegion>
,
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, collected
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<collectingDate id="17793D0D181CFF82441A350BAF21ED57" box="[853,1046,1028,1051]" pageId="88" pageNumber="89" value="1993-11-02">2 November 1993</collectingDate>
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by
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, deposited in
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.
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8247E4354BAA9CED37" pageId="88" pageNumber="89" type="etymology">
<paragraph id="733CE225181CFF8247E4354BAA9CED37" blockId="88.[122,1174,996,1725]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E4354BAA1DED17" bold="true" box="[171,298,1092,1115]" pageId="88" pageNumber="89">Etymology.</emphasis>
The species epithet is from the Malagasy
<emphasis id="41F73E37181CFF82445E354BA899ED17" box="[785,942,1092,1115]" italics="true" pageId="88" pageNumber="89">volana volana</emphasis>
, meaning a crescent moon; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8247E4358BABD1EC50" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="733CE225181CFF8247E4358BABD1EC50" blockId="88.[122,1174,996,1725]" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E4358BAAF7EDD7" bold="true" box="[171,448,1156,1179]" pageId="88" pageNumber="89">
Diagnosis.
<taxonomicName id="B48399A6181CFF82466A358BAAF7EDD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,448,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="volana">
<emphasis id="41F73E37181CFF82466A358BAAF7EDD7" bold="true" box="[293,448,1156,1179]" italics="true" pageId="88" pageNumber="89">Uduba volana</emphasis>
</taxonomicName>
</emphasis>
are members of Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A6181CFF82473435ABAA76EDF7" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,321,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). They have a divided cribellum. Females can be distinguished from those of other
<taxonomicName id="B48399A6181CFF82461B35CBAAA9ED97" box="[340,414,1220,1243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37181CFF82461B35CBAAA9ED97" bold="true" box="[340,414,1220,1243]" italics="true" pageId="88" pageNumber="89">Uduba</emphasis>
</taxonomicName>
by having the epigynal plate wide (width 1.52 times length) and with the lateral ears very slender, LL side width 1.25 times LL side length (
<figureCitation id="EBB8FEA0181CFF82443135EBAF21EDB7" box="[894,1046,1252,1275]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="88" pageNumber="89">Figs. 77 D, E</figureCitation>
). Males are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181CFF8347E4342AA805EAED" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="description">
<paragraph id="733CE225181CFF8347E4342AAB93EB91" blockId="88.[122,1174,996,1725]" lastBlockId="89.[122,1174,220,995]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<emphasis id="41F73E37181CFF8247E4342AAA4FEC70" bold="true" box="[171,376,1317,1340]" pageId="88" pageNumber="89">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E37181CFF8246B0342AA965EC70" bold="true" box="[511,594,1317,1340]" pageId="88" pageNumber="89">Female</emphasis>
(
<typeStatus id="AC385C87181CFF82452D342AA9F1EC70" box="[610,710,1317,1340]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
): Total length 13.71. Markings as in
<figureCitation id="EBB8FEA0181CFF824319342AABA7EC10" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="88" pageNumber="89">Fig. 7 C</figureCitation>
. Carapace 5.79 long, 3.71 wide, 2.57 high; clypeus 0.29 high. Eye diameters: AME 0.21, ALE, PME and PLE 0.27. Chelicerae 2.79 long; sternum 2.71 long, 2.00 wide; labium 1.21 long; palpal coxae 1.96 long.
<emphasis id="41F73E37181CFF824660348AAAB2ECD0" bold="true" box="[303,389,1413,1436]" pageId="88" pageNumber="89">Ratios–</emphasis>
carapace length / width = 1.56, carapace height / width = 0.69, PER / carapace width = 0.67, PER/OAL = 2.77, PER/AER = 1.33, OAL/OQL = 1.00, OQP/ OQA = 1.17, clypeus height / AME = 1.33, cheliceral length / clypeus height = 9.75, sternum length / width = 1.36, palpal coxa length / width = 1.83, femur I length / carapace width = 1.08, metatarsus I length / carapace width = 0.73, femur IV length / carapace width = 1.06, palpal tarsus length / carapace width = 0.38, palpal tibia length / palpal patella length = 1.33, palpal tarsus length / palpal tibia length = 1.33, palpal tarsus length / palpal femur length = 0.69, palpal tibia length / palpal patella length = 1.00. Divided cribellum. Spination: palpus–femur d0-0-2, patella p1-0, tibia p2-0, tarsus p2-1; leg I–femur d1-0-0, p0-0-2, tibia v2-2-2, metatarsus v2-2-3; leg II–femur d1-0-0, p0-0-1, tibia p0-1-1-0, v1-1-2, metatarsus v2-2-3; leg III–femur d1-1-0-0, p1-0-1-1, tibia d1-0-0, p0-1-1-0, v1-0-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2, tarsus v1-0; leg IV–femur d1-0-0, tibia v0-1- 2, r1-0-1, metatarsus p1-2-2, v2-1-2-2, r1-2-2. Scopulae: strong ventral beneath tarsi I–IV, beneath metatarsi I and II and beneath apices of metatarsi III and IV. Leg measurements: I: 4.00 + 1.86 + 2.93 + 2.71 + 1.64 = 13.14; II: 3.36 + 1.79 + 2.36 + 2.29 + 1.57 = 11.36; III: 2.71 + 1.43 + 1.29 + 1.86 + 1.14 = 8.43; IV: 3.93+ 1.79 + 2.93 + 2.93 + 1.71 = 13.29; palpus: 2.07 + 1.07 + 1.07 + NA + 1.43 = 5.64. Leg formula 4123.
<emphasis id="41F73E37181DFF8345593070A9E4E8DA" bold="true" box="[534,723,383,406]" pageId="89" pageNumber="90">Female genitalia</emphasis>
: epigynal plate without depressed atrium, lateral lobes extending to sides forming slender, earlike lobes (
<figureCitation id="EBB8FEA0181DFF83447E30AFA8FBE8FB" box="[817,972,416,439]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="89" pageNumber="90">Figs. 77 D, E</figureCitation>
), plate width 1.52 times length; width between bases of LL 0.33 times epigynum width; lateral ears (LL side) width 0.17 times width epigynum, lateral ears (LL side) very slender, width 1.25 times LL side length, arising far from epigastric furrow; median lobe a flat plate, each side with a bulge beneath the transverse hood formed by the posterior margin of the LL, width 0.33 times epigynum width, 0.66 times ML length, with copulatory openings beneath near ML bulge beneath LL hood at 0.51 epigynum length from epigastric furrow, distance between copulatory openings 0.60 times epigynum width. Vulva (
<figureCitation id="EBB8FEA0181DFF83465F338AAA44EBD0" box="[272,371,645,668]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="89" pageNumber="90">Fig. 77 F</figureCitation>
) with spermathecal ducts forming 2–3 loose, transverse loops, vulva length 0.68 times width, fertilization ducts widely separated, vulva width 5.91 times distance between FD.
</paragraph>
<paragraph id="733CE225181DFF8347E433E8A82DEBB2" blockId="89.[122,1174,220,995]" box="[171,794,743,766]" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E433E8AA2DEBB2" bold="true" box="[171,282,743,766]" pageId="89" pageNumber="90">Variation.</emphasis>
Unknown: only the
<typeStatus id="AC385C87181DFF8346B433E8A96FEBB2" box="[507,600,743,766]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
female is known.
</paragraph>
<paragraph id="733CE225181DFF8347E43208AF02EA2C" blockId="89.[122,1174,220,995]" pageId="89" pageNumber="90">
<materialsCitation id="C3EBE878181DFF8347E43208AF06EA2C" collectingDate="1993-11-02" collectionCode="CAS" collectorName="C. Griswold &amp; S. Larcher &amp; N. Scharff" country="Madagascar" elevation="1400" location="Ambohimanga" pageId="89" pageNumber="90" specimenCode="CASENT9006072" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo" typeStatus="holotype">
M
<emphasis id="41F73E37181DFF83478D3208AAA7EA52" bold="true" box="[194,400,775,798]" pageId="89" pageNumber="90">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5181DFF8346D13208A957EA52" box="[414,608,775,798]" name="Madagascar" pageId="89" pageNumber="90">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37181DFF83453D3208A844EA52" box="[626,883,775,798]" italics="true" pageId="89" pageNumber="90">
<collectingRegion id="B1472CC7181DFF83453D3208A844EA52" box="[626,883,775,798]" country="Madagascar" name="Antananarivo" pageId="89" pageNumber="90">Antananarivo Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE181DFF8344C63208AF1DEA52" LSID="urn:lsid:plazi:treatment:FB2A5333181CFF8347343291A9ADEAAF:765CB4FE181DFF8344C63208AF1DEA52" box="[905,1066,775,798]" country="Madagascar" name="Ambohimanga" pageId="89" pageNumber="90" stateProvince="Antananarivo">Ambohimanga</location>
, 18º44ʹS, 47º34ʹE, elev.
<quantity id="B47B4FC0181DFF8346523227AA5EEA73" box="[285,361,808,831]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="89" pageNumber="90" unit="m" value="1400.0">
<elevation id="F8AE0516181DFF8346523227AA5EEA73" box="[285,361,808,831]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="89" pageNumber="90" unit="m" value="1400.0">1400m</elevation>
</quantity>
, “from short burrows in road cut, night collecting,”
<date id="073DC4E5181DFF83448C3227AFA6EA73" box="[963,1169,808,831]" pageId="89" pageNumber="90" value="1993-11-02">
<collectingDate id="17793D0D181DFF83448C3227AFA6EA73" box="[963,1169,808,831]" pageId="89" pageNumber="90" value="1993-11-02">2 November 1993</collectingDate>
</date>
,
<collectorName id="DE7687F3181DFF8347343246AA36EA2C" box="[123,257,841,864]" pageId="89" pageNumber="90">C. Griswold</collectorName>
,
<collectorName id="DE7687F3181DFF8346423246AA4BEA2C" box="[269,380,841,864]" pageId="89" pageNumber="90">S. Larcher</collectorName>
,
<collectorName id="DE7687F3181DFF8346C53246AACEEA2C" box="[394,505,841,864]" pageId="89" pageNumber="90">N. Scharff</collectorName>
[CEG009] (
<typeStatus id="AC385C87181DFF8345CF3246A9D5EA2C" box="[640,738,841,864]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E181DFF8345A23246A882EA2C" box="[749,949,841,864]" collectionCode="CASENT" pageId="89" pageNumber="90">CASENT9006072</specimenCode>
,
<specimenCount id="658529AC181DFF83448E3246A8D5EA2C" box="[961,994,841,864]" count="1" pageId="89" pageNumber="90" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0181DFF8344BE3246AF1EEA2C" box="[1009,1065,841,864]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="89" pageNumber="90" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225181DFF8347E43266A805EAED" blockId="89.[122,1174,220,995]" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E43266AA69EACC" bold="true" box="[171,350,873,896]" pageId="89" pageNumber="90">Natural history.</emphasis>
The cribellate female of
<taxonomicName id="B48399A6181DFF8345273266A835EACC" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[616,770,873,896]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="volana">
<emphasis id="41F73E37181DFF8345273266A835EACC" bold="true" box="[616,770,873,896]" italics="true" pageId="89" pageNumber="90">Uduba volana</emphasis>
</taxonomicName>
was collected from a short burrow on an incline in disturbed secondary forest in the central highlands.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181DFF8347E432A4A9ADEAAF" pageId="89" pageNumber="90" type="distribution">
<paragraph id="733CE225181DFF8347E432A4A9ADEAAF" blockId="89.[122,1174,220,995]" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E432A4AA0AEA8E" bold="true" box="[171,317,939,962]" pageId="89" pageNumber="90">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6181DFF83468532A4A952EA8E" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[458,613,939,962]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="volana">
<emphasis id="41F73E37181DFF83468532A4A952EA8E" bold="true" box="[458,613,939,962]" italics="true" pageId="89" pageNumber="90">Uduba volana</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C87181DFF83443F32A4A8A9EA8E" box="[880,926,939,962]" pageId="89" pageNumber="90">type</typeStatus>
locality in
<collectingRegion id="B1472CC7181DFF83435932A4ABC8EAAF" country="Madagascar" name="Antananarivo" pageId="89" pageNumber="90">Antananarivo Province</collectingRegion>
, in central
<collectingCountry id="0B94A2B5181DFF83463732C3AACEEAAF" box="[376,505,972,995]" name="Madagascar" pageId="89" pageNumber="90">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA0181DFF83454732C3A96BEAAF" box="[520,604,972,995]" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="89" pageNumber="90">Maps 5</figureCitation>
, 16.).
</paragraph>
</subSubSection>
</treatment>
</document>

716
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@ -0,0 +1,716 @@
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<mods:title id="B2E0776EBAD8ACEEA3D7AE1198006341">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="52BEA9F1176BA2B73A8D847D25A5570F">Griswold, Charles</mods:namePart>
<mods:affiliation id="56AC0AEBA1BF33CF79E4310151F9A529">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="3F36F773109666E471DBE64E50CF432E">Ubick, Darrell</mods:namePart>
<mods:affiliation id="9842C55D056E0CC9700749F315EB4A6C">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="EA8FD4EFE3D76B119C67BBAB3038D322">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:roleTerm id="64DA3D51C1AA324D45BB05011D2F4B52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FF6A4B1186452D10ACCB1A34C6E0D95">Polotow, Daniele</mods:namePart>
<mods:affiliation id="13273A8D3B9D14E1C8B9018A9DD58422">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:typeOfResource id="47B42395F9B31845B937F08776CF3B35">text</mods:typeOfResource>
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<mods:titleInfo id="92721681E0D60ED1FD005BEE1E578408">
<mods:title id="E06DFE9ED39A72C391C30F9A41EAE103">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="D492868A5B35E35FD27407644F6D8271">
<mods:date id="7A7633F45CAB069779C2E320F1B84EA2">2022</mods:date>
<mods:detail id="819FFC07B5A03CD4BA9E416E9EC8692D" type="pubDate">
<mods:number id="EA3E3D5EDAF06629836554DE2DF7CE91">2022-12-15</mods:number>
</mods:detail>
<mods:detail id="334A2B2E912FADB157C22AAA16ED5781" type="volume">
<mods:number id="89D7C2286032827B2C5B97FCAA760906">67</mods:number>
</mods:detail>
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<mods:start id="0FAE37206FC4FBC2BD697540E73BF5FA">1</mods:start>
<mods:end id="F5F54108ADCF84382E9E03F1BC9C7D32">193</mods:end>
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<mods:classification id="93924179797968114B8F7A930BC2858D">journal article</mods:classification>
<mods:identifier id="4FE3F661CC975CEEFE7C56CA8F8BF62B" type="DOI">10.5281/zenodo.13158554</mods:identifier>
<mods:identifier id="2B83D01C3BFAF7B2892F543B3BCF6B41" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="98289697D4E1DEFC142F0622AC07E7A6" type="Zenodo-Dep">13158554</mods:identifier>
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<subSubSection id="3B99B1AE181DFF8347343502AA86ED68" box="[123,433,1037,1060]" pageId="89" pageNumber="90" type="nomenclature">
<paragraph id="733CE225181DFF8347343502AA86ED68" blockId="89.[123,835,1037,1091]" box="[123,433,1037,1060]" pageId="89" pageNumber="90">
<heading id="28745549181DFF8347343502AA86ED68" box="[123,433,1037,1060]" fontSize="10" level="2" pageId="89" pageNumber="90" reason="4">
<emphasis id="41F73E37181DFF8347343502AA86ED68" bold="true" box="[123,433,1037,1060]" pageId="89" pageNumber="90">
<taxonomicName id="B48399A6181DFF8347343502AA28ED68" ID-CoL="C8KK8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,287,1037,1060]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="woodae" status="sp. nov.">
<emphasis id="41F73E37181DFF8347343502AA28ED68" bold="true" box="[123,287,1037,1060]" italics="true" pageId="89" pageNumber="90">Uduba woodae</emphasis>
</taxonomicName>
,
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181DFF8347343521A874ED0F" box="[123,835,1070,1091]" pageId="89" pageNumber="90" type="description">
<paragraph id="733CE225181DFF8347343521A874ED0F" blockId="89.[123,835,1037,1091]" box="[123,835,1070,1091]" pageId="89" pageNumber="90">
<figureCitation id="EBB8FEA0181DFF8347343521AA32ED0F" box="[123,261,1070,1091]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="89" pageNumber="90">Figures 7 J, K</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF83465F3521AA03ED0F" box="[272,308,1070,1091]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="89" pageNumber="90">9 B</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF8346703521AABBED0F" box="[319,396,1070,1091]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="89" pageNumber="90">66 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF8346D93521AAE9ED0F" box="[406,478,1070,1091]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="89" pageNumber="90">67 A–F</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF8346A63521A906ED0F" box="[489,561,1070,1091]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="89" pageNumber="90">78 A–F</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF8345743521A9BEED0F" box="[571,649,1070,1091]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="89" pageNumber="90">79 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF8345DC3521A9F2ED0F" box="[659,709,1070,1091]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="89" pageNumber="90">80 D</figureCitation>
, Maps 8, 14.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181DFF8347E43556A952ED9F" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="733CE225181DFF8347E43556A952ED9F" blockId="89.[123,1174,1113,1725]" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E43556AA79ED3C" bold="true" box="[171,334,1113,1136]" pageId="89" pageNumber="90">Type material.</emphasis>
<materialsCitation id="C3EBE878181DFF83461B3556A8F1EDFE" collectingDate="2003-05-13" collectingDateMax="2003-05-21" collectingDateMin="2003-05-13" collectionCode="CAS" country="Madagascar" elevation="20" location="Lake Ranobe" municipality="Frontier Wilderness Project" pageId="89" pageNumber="90" specimenCode="CASENT9017834" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C87181DFF83461B3556AA80ED3C" box="[340,439,1113,1136]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E181DFF8346B03556A9F0ED3C" box="[511,711,1113,1136]" collectionCode="CASENT" pageId="89" pageNumber="90">CASENT9017834</specimenCode>
)
<collectingMethod id="AAC29A32181DFF83459A3556AF4DED3C" box="[725,1146,1113,1136]" pageId="89" pageNumber="90">from general collecting and pitfall traps</collectingMethod>
in spiny forest at
<quantity id="B47B4FC0181DFF8346563575AA7FEDDD" box="[281,328,1146,1169]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="90" unit="m" value="20.0">
<elevation id="F8AE0516181DFF8346563575AA7FEDDD" box="[281,328,1146,1169]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="90" unit="m" value="20.0">20m</elevation>
</quantity>
elevation at
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,
<collectingRegion id="B1472CC7181DFF8345243575A829EDDD" box="[619,798,1146,1169]" country="Madagascar" name="Toliara" pageId="89" pageNumber="90">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B5181DFF8344673575A890EDDD" box="[808,935,1146,1169]" name="Madagascar" pageId="89" pageNumber="90">Madagascar</collectingCountry>
, collected
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<collectingDate id="17793D0D181DFF8343543575AB86EDFE" pageId="89" pageNumber="90" value="2003-05-13" valueMax="2003-05-21" valueMin="2003-05-13">13–21 May 2003</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F181DFF8346513594A96FEDFE" box="[286,600,1179,1202]" pageId="89" pageNumber="90">Frontier Wilderness Project</collectingMunicipality>
[MGF072], deposited in
<collectionCode id="15927AE0181DFF8344DD3594A8F1EDFE" box="[914,966,1179,1202]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="89" pageNumber="90" type="Museum">CAS</collectionCode>
</materialsCitation>
.
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<typeStatus id="AC385C87181DFF8344923594AF0CEDFE" box="[989,1083,1179,1202]" pageId="89" pageNumber="90" type="paratype">Paratype</typeStatus>
female (
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), same data, also in
<collectionCode id="15927AE0181DFF83456535B3A956ED9F" box="[554,609,1212,1235]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="89" pageNumber="90" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181DFF8347E435D3AF7EEC19" pageId="89" pageNumber="90" type="etymology">
<paragraph id="733CE225181DFF8347E435D3AF7EEC19" blockId="89.[123,1174,1113,1725]" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E435D3AA1DEDBF" bold="true" box="[171,298,1244,1267]" pageId="89" pageNumber="90">Etymology.</emphasis>
The species epithet is patronym in honor of Dr. Hannah Wood, collector of many
<taxonomicName id="B48399A6181DFF83473435F2ABF2EC58" box="[123,197,1277,1300]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37181DFF83473435F2ABF2EC58" bold="true" box="[123,197,1277,1300]" italics="true" pageId="89" pageNumber="90">Uduba</emphasis>
</taxonomicName>
and many other rare and cryptic arachnid specimens from around the world, student of the evolution, functional morphology and behavior of the assassin spiders (Palpimanoidea) and with a focus on the pelican spiders (
<taxonomicName id="B48399A6181DFF8346F83431A901EC19" authorityName="C.L.Koch &amp; Berendt" authorityYear="1854" box="[439,566,1342,1365]" class="Arachnida" family="Archaeidae" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="family">Archaeidae</taxonomicName>
), ancient survivors from tropical Gondwanaland.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181DFF8047E43450A8EDE85F" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="discussion">
<paragraph id="733CE225181DFF8047E43450A8EDE85F" blockId="89.[123,1174,1113,1725]" lastBlockId="90.[123,1175,220,1725]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis id="41F73E37181DFF8347E43450AA2EEC3A" bold="true" box="[171,281,1375,1398]" pageId="89" pageNumber="90">Remarks.</emphasis>
Our concept of
<taxonomicName id="B48399A6181DFF8346863450A95AEC3A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[457,621,1375,1398]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37181DFF8346863450A95AEC3A" bold="true" box="[457,621,1375,1398]" italics="true" pageId="89" pageNumber="90">Uduba woodae</emphasis>
</taxonomicName>
is a cribellate species found in the drier southwestern parts of
<collectingCountry id="0B94A2B5181DFF8347B5348FAA4FECDB" box="[250,376,1408,1431]" name="Madagascar" pageId="89" pageNumber="90">Madagascar</collectingCountry>
. An epigynum that has a clearly defined median lobe and surrounding atrium is associated with a male palpus that has a screw shaped TA3, e.g., as in
<taxonomicName id="B48399A6181DFF8344E434AFAF03ECFB" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[939,1076,1440,1463]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E37181DFF8344E434AFAF03ECFB" bold="true" box="[939,1076,1440,1463]" italics="true" pageId="89" pageNumber="90">Uduba dahli</emphasis>
</taxonomicName>
,
<taxonomicName id="B48399A6181DFF83430D34AFAB85EC94" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="41F73E37181DFF83430D34AFAB85EC94" bold="true" italics="true" pageId="89" pageNumber="90">U. platnicki</emphasis>
</taxonomicName>
, and
<taxonomicName id="B48399A6181DFF8347A034CEAA48EC94" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[239,383,1473,1496]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E37181DFF8347A034CEAA48EC94" bold="true" box="[239,383,1473,1496]" italics="true" pageId="89" pageNumber="90">U. schlingeri</emphasis>
</taxonomicName>
. Females of
<taxonomicName id="B48399A6181DFF83454334CEA985EC94" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[524,690,1473,1496]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37181DFF83454334CEA985EC94" bold="true" box="[524,690,1473,1496]" italics="true" pageId="89" pageNumber="90">Uduba woodae</emphasis>
</taxonomicName>
have a divided cribellum and a median lobe plus atrium
<typeStatus id="AC385C87181DFF8347B734EDAA11ECB5" box="[248,294,1506,1529]" pageId="89" pageNumber="90">type</typeStatus>
epigynum that has the part of the epigynal plate that is posteriad of the atrium high, nearly equal in length to the ML itself (
<figureCitation id="EBB8FEA0181DFF834550370CA9FAEF56" box="[543,717,1539,1562]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="89" pageNumber="90">Figs. 78 A, C, E</figureCitation>
,
<figureCitation id="EBB8FEA0181DFF834597370CA83AEF56" box="[728,781,1539,1562]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="89" pageNumber="90">79 B</figureCitation>
). Females from southwestern
<collectingCountry id="0B94A2B5181DFF83431F370CABF7EF76" name="Madagascar" pageId="89" pageNumber="90">Madagascar</collectingCountry>
(
<collectingRegion id="B1472CC7181DFF834795372CAAA1EF76" box="[218,406,1571,1594]" country="Madagascar" name="Toliara" pageId="89" pageNumber="90">Toliara Province</collectingRegion>
) that have divided cribella and this
<typeStatus id="AC385C87181DFF83442D372CA8A6EF76" box="[866,913,1571,1594]" pageId="89" pageNumber="90">type</typeStatus>
of epigynum include CASENT9000536 from Forêt de Beroboka (
<figureCitation id="EBB8FEA0181DFF834517374BA989EF17" box="[600,702,1604,1627]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="89" pageNumber="90">Fig. 78 A</figureCitation>
) and CASENT9030930 from Mikea Forest, NW of Manombo, which are here considered to be examples of
<taxonomicName id="B48399A6181DFF83446E376AA8A3EF30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[801,916,1637,1660]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37181DFF83446E376AA8A3EF30" bold="true" box="[801,916,1637,1660]" italics="true" pageId="89" pageNumber="90">U. woodae</emphasis>
</taxonomicName>
. An old series of female specimens collected by Carougeau during
<date id="073DC4E5181DFF834571378AA804EFD0" box="[574,819,1669,1692]" pageId="89" pageNumber="90" value="1901-11-19" valueMax="1901-11-30" valueMin="1901-11-19">19–30 November 1901</date>
and deposited in the Paris Museum (MNHN) also shows these epigynal characters. The collection locality for these specimens may be Ambovombe, which is in far southern
<collectingCountry id="0B94A2B5181EFF80450B31D3A9F4E9BF" box="[580,707,220,243]" name="Madagascar" pageId="90" pageNumber="91">Madagascar</collectingCountry>
, also in
<collectingRegion id="B1472CC7181EFF80446B31D3A8EFE9BF" box="[804,984,220,243]" country="Madagascar" name="Toliara" pageId="90" pageNumber="91">Toliara Province</collectingRegion>
. All these unique collections of cribellate females are considered to belong to species
<taxonomicName id="B48399A6181EFF80441131F3A8E4E85F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[862,979,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37181EFF80441131F3A8E4E85F" bold="true" box="[862,979,252,275]" italics="true" pageId="90" pageNumber="91">U. woodae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181EFF8047E43013AAB2EB9A" pageId="90" pageNumber="91" type="diagnosis">
<paragraph id="733CE225181EFF8047E43013AAB2EB9A" blockId="90.[123,1175,220,1725]" pageId="90" pageNumber="91">
<emphasis id="41F73E37181EFF8047E43013AAFBE87F" bold="true" box="[171,460,284,307]" pageId="90" pageNumber="91">
Diagnosis.
<taxonomicName id="B48399A6181EFF8046683013AAFBE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,460,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37181EFF8046683013AAFBE87F" bold="true" box="[295,460,284,307]" italics="true" pageId="90" pageNumber="91">Uduba woodae</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.b, the
<taxonomicName id="B48399A6181EFF80446C3013A8F2E87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[803,965,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="woodae">Uduba woodae</taxonomicName>
group (Map 14) of Group I, the Epigynal atrium group. They are cribellate, with divided cribellum or vestige. Males have the palp (
<figureCitation id="EBB8FEA0181EFF8046573053AA95E83F" box="[280,418,348,371]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs 66 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8046E23053AACBE83F" box="[429,508,348,371]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 A–F</figureCitation>
) with TA2 highly sclerotized, black, concave medially against TA3 (
<figureCitation id="EBB8FEA0181EFF8047F73072AA13E8D8" box="[184,292,381,404]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Fig. 66 B</figureCitation>
), MA concave in middle with proximal sharp point (
<figureCitation id="EBB8FEA0181EFF8044283072A8EAE8D8" box="[871,989,381,404]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 C</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8044A53072AF13E8D8" box="[1002,1060,381,404]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 D</figureCitation>
), origin at 0.45 times tegulum length. Males are distinguished from
<taxonomicName id="B48399A6181EFF8045A83092A8B9E8F8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[743,910,413,436]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E37181EFF8045A83092A8B9E8F8" bold="true" box="[743,910,413,436]" italics="true" pageId="90" pageNumber="91">U. lehibekokoa</emphasis>
</taxonomicName>
by having a sharp point on the MA, the TA3 apex blunt (
<figureCitation id="EBB8FEA0181EFF80469230B2A966E898" box="[477,593,445,468]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 C</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF80451230B2A9A3E898" box="[605,660,445,468]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 B</figureCitation>
), the VTA that is short and straight (
<figureCitation id="EBB8FEA0181EFF80435130B2ABB8E8B8" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 A, C</figureCitation>
), length less than 2 times width, and specimens that are smaller in size, total length = 9.00–
<quantity id="B47B4FC0181EFF80473430F1ABD6EB59" box="[123,225,510,533]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="90" pageNumber="91" unit="mm" value="12.4">12.40mm</quantity>
(
<taxonomicName id="B48399A6181EFF8047BA30F1AA97EB59" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[245,416,510,533]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E37181EFF8047BA30F1AA97EB59" bold="true" box="[245,416,510,533]" italics="true" pageId="90" pageNumber="91">U. lehibekokoa</emphasis>
</taxonomicName>
males have the MA smooth, the TA3 apex pointed, VTA long and curved apically, length greater than 2 times width,
<figureCitation id="EBB8FEA0181EFF8045EF3311A806EB79" box="[672,817,542,565]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="90" pageNumber="91">Figs. 50 A–C</figureCitation>
, and specimens that are larger in size, total length 14.00–
<quantity id="B47B4FC0181EFF8046303331AAD0EB19" box="[383,487,574,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="90" pageNumber="91" unit="mm" value="15.5">15.50mm</quantity>
). Females have the epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA0181EFF80462D3351A925EB39" box="[354,530,606,629]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="90" pageNumber="91">Figs. 78 A, C, E</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8045513351A963EB39" box="[542,596,606,629]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="90" pageNumber="91">79 B</figureCitation>
), ML elongate, length at least 1.5 times width, atrium narrow and ML broad. The median lobe ends well anteriad of epigastric furrow, the post-atrial plate of the epigynum is very high and the epigynum length is greater than 1.5 times atrial length (
<figureCitation id="EBB8FEA0181EFF80432F3390ABC7EB9A" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="90" pageNumber="91">Figs. 78 A, C, E</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8047B333B0AA05EB9A" box="[252,306,703,726]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="90" pageNumber="91">79 B</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF80467033B0AA4FEB9A" box="[319,376,703,726]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="90" pageNumber="91">80 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE181EFF8647E433D0A93DED28" lastPageId="92" lastPageNumber="93" pageId="90" pageNumber="91" type="description">
<paragraph id="733CE225181EFF8147E433D0AA72EBFA" blockId="90.[123,1175,220,1725]" lastBlockId="91.[123,1174,220,1725]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
<emphasis id="41F73E37181EFF8047E433D0AA42EBBA" bold="true" box="[171,373,735,758]" pageId="90" pageNumber="91">Description. Male</emphasis>
(
<typeStatus id="AC385C87181EFF8046CC33D0AADAEBBA" box="[387,493,735,758]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
): Total length 9.80. Markings as in
<figureCitation id="EBB8FEA0181EFF80441033D0A880EBBA" box="[863,951,735,758]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="90" pageNumber="91">Fig. 9 B</figureCitation>
. Carapace 6.10 long, 4.20 wide, 2.10 high; clypeus 0.30 high. Eye diameters:AME, ALE and PLE 0.22, PME 0.20. Chelicerae 2.40 long; sternum 2.70 long, 2.10 wide; labium 1.30 long; palpal coxae 1.90 long. Divid- ed cribellum. Spination: palpus–femur d1-1-3, tibia p0-1-0; leg I–femur d1-1-1-1, p0-1-0-2, r0-1- 1-0, patella p1, r1, tibia d0-1-1-0, p1-0-1-0, v2-2-2-2, r1-0-1-0, metatarsus p1-2-2, v2-2-1-2, r1-1- 1; leg II–femur d1-0-1-1, p0-1-1-1, r1-1-1-1, patella p1, r1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0- 1-1-0, metatarsus p1-1-1, v2-2-1-2, r1-1-1; leg III–femur d1-1-1, p0-1-1-1, r0-1-1-1, patella p1, r1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0-0, p1-1-2, v2-2-2, r1-1-2; leg IV– femur d2-01-1, p0-1-1-0, r0-1-1-0, patella p1, r1, tibia d0-0-1-0, p0-1-1-0, v2-1-2-2, r0-1-1-0, metatarsus d0-1-0-0, p1-1-1-2, v1-1-1-2-2, r1-1-2. Scopulae: cymbium, apicodorsal (
<figureCitation id="EBB8FEA0181EFF804353350EAFBEED54" box="[1052,1161,1025,1048]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">Fig. 67 A</figureCitation>
); strong beneath all tarsi. Leg measurements: I: 5.90 + 2.30 + 5.70 + 5.60 + 3.50 = 23.00; II: 5.00 + 2.10 + 4.40 + 4.90 + 3.00 = 19.40; III: 4.40 + 1.70 + 3.00 + 4.40 + 2.50 = 16.00; IV: 6.00 + 2.10 + 5.30 + 7.10 + 3.10 = 23.60; palpus: 2.10 + 0.80 + 0.90 + NA + 2.50 = 6.30. Leg formula 4123.
<emphasis id="41F73E37181EFF8047F1358DAA05EDD5" bold="true" box="[190,306,1154,1177]" pageId="90" pageNumber="91">Male palp</emphasis>
: palpal tibia 0.36 times cymbial length, RTA very broad, crescentic, with point- ed apex (
<figureCitation id="EBB8FEA0181EFF80479035ADAA62EDF5" box="[223,341,1186,1209]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 A</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF80462C35ADAAADEDF5" box="[355,410,1186,1209]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 A</figureCitation>
), RTA length 0.26 tibia length, length 0.75 times width, VTA conical, distally hooked, with rounded apex (
<figureCitation id="EBB8FEA0181EFF80454A35CCA9A0ED96" box="[517,663,1219,1242]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 A, C</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8045EC35CCA9CDED96" box="[675,762,1219,1242]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 B, C</figureCitation>
), length 1.67 times width, VTA length 0.29 times tibia width; tibia with slender prolateral spine (
<figureCitation id="EBB8FEA0181EFF80444835ECA848EDB6" box="[775,895,1251,1274]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 A</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8044C135ECA8FFEDB6" box="[910,968,1251,1274]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 C</figureCitation>
); tegulum convex, without ridge (
<figureCitation id="EBB8FEA0181EFF804656340CAA9DEC56" box="[281,426,1283,1306]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 B, C</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8046FA340CAADBEC56" box="[437,492,1283,1306]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 D</figureCitation>
), tegulum length 0.92 times width; TA1 rounded, blunt apically, extends distad of tegulum apex, TA1 extends past TA3 apex by 1.27 times tegulum length; TA3 transverse, tapering, concave dorsally, rounded TA3 apex reflexed, visible in front of TA2 (
<figureCitation id="EBB8FEA0181EFF804307344CABB8EC37" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Fig. 66 B</figureCitation>
); TA2 highly sclerotized, black, concave medially against TA3 (
<figureCitation id="EBB8FEA0181EFF80441C346BA8DBEC37" box="[851,1004,1380,1403]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Figs. 66 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0181EFF8044B5346BAF63EC37" box="[1018,1108,1380,1403]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">67 B, E</figureCitation>
); MA transverse, simple, concave in middle with proximal sharp point (
<figureCitation id="EBB8FEA0181EFF80441F348BA8DDECD7" box="[848,1002,1412,1435]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="90" pageNumber="91">Figs. 67 D, E</figureCitation>
), origin at 0.45 times tegulum length, tegulum apex at 0.32 tegulum length from MA apex, MA length (including apical processes) 0.41 times width, MA small, length 0.16 tegulum length, width 0.35 times tegulum width; conductor fan entire (
<figureCitation id="EBB8FEA0181EFF8046AB34EAA967ECB0" box="[484,592,1509,1532]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="90" pageNumber="91">Fig. 66 C</figureCitation>
).
<emphasis id="41F73E37181EFF80452B34EAA9FEECB0" bold="true" box="[612,713,1509,1532]" pageId="90" pageNumber="91">Female (</emphasis>
<typeStatus id="AC385C87181EFF80458634EAA812ECB0" box="[713,805,1509,1532]" pageId="90" pageNumber="91" type="paratype">paratype</typeStatus>
<emphasis id="41F73E37181EFF80446A34EAA801ECB0" bold="true" box="[805,822,1509,1532]" pageId="90" pageNumber="91">):</emphasis>
Total length 11.40. Markings as in
<figureCitation id="EBB8FEA0181EFF8047D8370AAA21EF50" box="[151,278,1541,1564]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="90" pageNumber="91">Figs. 7 J, K</figureCitation>
. Carapace 5.50 long, 3.80 wide, 2.70 high; clypeus 0.30 high. Eye diameters: AME and PME 0.18, ALE and PLE 0.22. Chelicerae 2.80 long; sternum 2.40 long, 1.90 wide; labium 2.20 long; palpal coxae 2.80 long. Divided cribellum. Spination: palpus–femur d0-0-2, p1, tibia p0- 1-0, tarsus p2-1-1; leg I–femur d0-1-0-1, p0-0-0-1, tibia v1-1-1-2, metatarsus p0-0-1, v2-2-3; leg II–femur d0-1-0-0, p0-0-0-1, tibia v1-1-1-2, metatarsus v2-2-3; leg III–femur d0-1-0-1, p0-1-0-1, r0-1-0-1, tibia p0-1-1-0, v1-1-2, r0-1-1-0, metatarsus p1-1-1, v2-2-2, r1-1-2; leg IV–femur d1-0-0- 1, tibia v1-1-2, r0-1-1-0, metatarsus p0-0-2-2, v2-2-2, r0-2-2. Scopulae: strong ventral beneath all tarsi and apices of metatarsi II–IV. Leg measurements: I: 3.10 + 1.90 + 2.90 + 2.40 + 1.70 = 12.00; II: 3.10 + 1.70 + 2.30 + 2.10 + 1.60 = 10.80; III: 2.70 + 1.40 + 1.50 + 2.20 + 1.40 = 9.20; IV: 3.70 + 1.60 + 2.80 + 3.60 + 1.70 = 13.40; palpus: 1.90 + 0.90 + 1.00 + NA + 1.70 = 5.50. Leg formula 4123.
<emphasis id="41F73E37181FFF81464D3053AAF7E83F" bold="true" box="[258,448,348,371]" pageId="91" pageNumber="92">Female genitalia</emphasis>
: epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA0181FFF8147CC3072ABD4E8D8" box="[131,227,381,404]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="91" pageNumber="92">Fig. 78 E</figureCitation>
), plate width 1.24 times length; atrium width 1.83 times atrium length, atrium sides evenly convex; epigynum length 2.50 times atrium length, atrium width 0.53 times epigynum width; atrium width at side of ML 0.47 times ML width; atrium and ML arise well anteriad of epigastric groove, atrium origin at 0.26 of epigynal plate length, ML relatively large, bell-shaped, ML length 0.62 times epigynal length, ML length 1.82 times width, ML extends far anteriad of atrium apex; LL broad, atrium width 2.06 times LL width; CO hidden beneath posterolateral margin of atrium, distance between CO 0.50 times width of epigynal plate, CO distance from epigastric furrow 0.34 times epigynum length. Vulva (
<figureCitation id="EBB8FEA0181FFF81452E3351A9F8EB39" box="[609,719,606,629]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="91" pageNumber="92">Fig. 78 F</figureCitation>
), with spermathecal ducts making three, transverse curves, vulva length 0.60 times width, FD close-together, vulva width 7.57 times distance between FD.
</paragraph>
<paragraph id="733CE225181FFF8147E433B0A92EEC97" blockId="91.[123,1174,220,1725]" pageId="91" pageNumber="92">
<emphasis id="41F73E37181FFF8147E433B0AA68EB9A" bold="true" box="[171,351,703,726]" pageId="91" pageNumber="92">Variation. Male</emphasis>
(N = 5): Total length = 9.00–12.40, carapace length / width = 1.34–1.57, carapace height / width = 0.47–0.52, PER / carapace width = 0.36–0.38, PER / OAL = 2.94–4.76, PER / AER = 1.28–1.38, OAL / OQL = 1.06–1.16, OQP / OQA 1.07–1.20, clypeus height / AME = 1.25–1.59, cheliceral length / clypeus height = 8.00–11.50, sternum length / width = 1.24–1.38, palpal coxa length / width = 2.22–3.20, femur I length / carapace width =1.16–1.40, metatarsus I length / carapace width = 1.16–1.47, femur IV length / carapace width = 1.22–1.45, cymbium length / carapace width = 0.61–0.70, cymbium length / palpal patella length = 3.10–3.75, cymbium length / palpal tibia length = 2.58–3.29, cymbium length / palpal femur length = 1.11–1.44, palpal tibia length / palpal patella length = 1.00–1.38. Palpus as in
<figureCitation id="EBB8FEA0181FFF81446432CEA88CEA94" box="[811,955,961,984]" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="91" pageNumber="92">Figs. 66 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0181FFF81448832CEAF26EA94" box="[967,1041,961,984]" captionStart="FIGURE" captionStartId="157.[155,167,1229,1248]" captionTargetBox="[120,1176,216,1211]" captionTargetId="figure-16@157.[120,1176,216,1211]" captionTargetPageId="157" captionText="FIGURE 67. Uduba woodae, new species, CASENT9017834 from Lake Ranobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. E, embolus; MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bars: A–C = 200µm, D–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.13158725" httpUri="https://zenodo.org/record/13158725/files/figure.png" pageId="91" pageNumber="92">67 A–F</figureCitation>
.
<emphasis id="41F73E37181FFF81435032CEAF45EA94" bold="true" box="[1055,1138,961,984]" pageId="91" pageNumber="92">Female</emphasis>
(N = 7): Total length = 11.00–22.43, carapace length / width = 1.23–1.49, carapace height/ width = 0.44–0.72, PER/ carapace width = 0.42–0.70, PER / OAL = 2.79–4.00, PER / AER = 1.25–1.59, OAL / OQL = 1.00–1.17, OQP / OQA = 1.04–1.23, clypeus height/ AME diameter width = 1.56– 2.08, cheliceral length / clypeus height = 7.00–10.00, sternum length / width = 1.18–1.27, palpal coxa length / width = 1.94–3.11, femur I length / carapace width = 0.82–1.06, metatarsus I length / carapace width = 0.63–0.72, femur IV length / carapace width = 0.95–1.04, palpal tarsus length / carapace width = 0.39–0.45, palpal tibia length / palpal patella length = 1.52–2.00, palpal tarsus length / palpal tibia length = 1.35–2.18, palpal tarsus length / palpal femur length = 0.71–0.96, palpal tibia length / palpal patella length = 0.92–1.13. Genitalia with epigynal plate (
<figureCitation id="EBB8FEA0181FFF8144AC35ECAFA5EDB6" box="[995,1170,1251,1274]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="91" pageNumber="92">Figs. 78 A, C, E</figureCitation>
,
<figureCitation id="EBB8FEA0181FFF814734340CAB9AEC56" box="[123,173,1283,1306]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="91" pageNumber="92">79 B</figureCitation>
) width 0.94–1.24 times length; atrium width 1.57–1.83 times atrium length, atrium sides nearly straight to evenly convex; epigynum length 1.69–2.56 times atrium length, atrium width at side of ML 0.21–0.85 times ML width; ML length 0.51–0.62 times epigynal length, median lobe length 1.57–3.00 times width, ML length extends 0.34–0.85 times epigynum length anteriad of atrium. Vulva (
<figureCitation id="EBB8FEA0181FFF814784348BAAB3ECD7" box="[203,388,1412,1435]" captionStart="FIGURE" captionStartId="166.[155,167,903,922]" captionTargetBox="[120,1176,216,887]" captionTargetId="figure-11@166.[120,1176,216,887]" captionTargetPageId="166" captionText="FIGURE 78. Uduba woodae, new species, female genitalia drawings. A, B. Forêt de Beroboka (CASENT9000536). C, D. Ambovombe, Carougeau, Muséum Paris (CASENT9006092). E, F. Paratype (CASENT9017834) from Lake Ranobe. A, C, E. Epigynum, ventral. B, D, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158747" httpUri="https://zenodo.org/record/13158747/files/figure.png" pageId="91" pageNumber="92">Figs. 78 B, D, F</figureCitation>
,
<figureCitation id="EBB8FEA0181FFF8146DA348BAAFAECD7" box="[405,461,1412,1435]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="91" pageNumber="92">79 A</figureCitation>
,
<figureCitation id="EBB8FEA0181FFF814693348BA922ECD7" box="[476,533,1412,1435]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="91" pageNumber="92">80 D</figureCitation>
) with spermathecal ducts making two to three transverse curves, vulva length 0.57–0.625 times width, FD widely-spaced to close-together, vulva width 3.30–7.57 times distance between FD.
</paragraph>
<paragraph id="733CE225181FFF8647E434EAAA3AEA33" blockId="91.[123,1174,220,1725]" lastBlockId="92.[123,1174,220,1189]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
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M
<emphasis id="41F73E37181FFF81478D34EAAABAECB0" bold="true" box="[194,397,1509,1532]" pageId="91" pageNumber="92">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5181FFF8146D634EAA96CECB0" box="[409,603,1509,1532]" name="Madagascar" pageId="91" pageNumber="92">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37181FFF81452534EAA851ECB0" box="[618,870,1509,1532]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="B1472CC7181FFF81452534EAA851ECB0" box="[618,870,1509,1532]" country="Madagascar" name="Fianarantsoa" pageId="91" pageNumber="92">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectingCounty id="9A5D9AA9181FFF81443534EAA8D1ECB0" box="[890,998,1509,1532]" pageId="91" pageNumber="92">Italaviana</collectingCounty>
,
<locationDeviation id="628EFB07181FFF8144BA34EAABD2EF50" location="Antsirabe" pageId="91" pageNumber="92">
<quantity id="B47B4FC0181FFF8144BA34EAAF0EECB0" box="[1013,1081,1509,1532]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="91" pageNumber="92" unit="km" value="35.0">35 km</quantity>
SSE of Antsirabe
</locationDeviation>
, 20°10.40ʹS, 47°05.16ʹE, elev.
<quantity id="B47B4FC0181FFF81457A370AA9B4EF50" box="[565,643,1541,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="91" pageNumber="92" unit="m" value="1360.0">
<elevation id="F8AE0516181FFF81457A370AA9B4EF50" box="[565,643,1541,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="91" pageNumber="92" unit="m" value="1360.0">1360m</elevation>
</quantity>
,
<collectingMethod id="AAC29A32181FFF8145DE370AA823EF50" box="[657,788,1541,1564]" pageId="91" pageNumber="92">malaise trap</collectingMethod>
in
<taxonomicName id="B48399A6181FFF814475370AA8B9EF50" box="[826,910,1541,1564]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="91" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E37181FFF814475370AA8B9EF50" box="[826,910,1541,1564]" italics="true" pageId="91" pageNumber="92">Uapaca</emphasis>
</taxonomicName>
, forest,
<date id="073DC4E5181FFF8144A6370AAB84EF70" pageId="91" pageNumber="92" value="2005-04-24" valueMax="2005-05-08" valueMin="2005-04-24">
<collectingDate id="17793D0D181FFF8144A6370AAB84EF70" pageId="91" pageNumber="92" value="2005-04-24" valueMax="2005-05-08" valueMin="2005-04-24">24 April–8 May 2005</collectingDate>
</date>
,
<collectorName id="DE7687F3181FFF81478E372AAA13EF70" box="[193,292,1573,1596]" pageId="91" pageNumber="92">M. Irwin</collectorName>
and
<collectorName id="DE7687F3181FFF81462F372AAACCEF70" box="[352,507,1573,1596]" pageId="91" pageNumber="92">R. Harin’Hala</collectorName>
[MA-24-71] (
<specimenCode id="23254A5E181FFF8145D4372AA852EF70" box="[667,869,1573,1596]" collectionCode="CASENT" pageId="91" pageNumber="92">CASENT9042522</specimenCode>
,
<specimenCount id="658529AC181FFF81443C372AA8A2EF70" box="[883,917,1573,1596]" count="1" pageId="91" pageNumber="92" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0181FFF8144E9372AA8E9EF70" box="[934,990,1573,1596]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="91" pageNumber="92" type="Museum">CAS</collectionCode>
).
<emphasis id="41F73E37181FFF8144BB372AA975EF10" italics="true" pageId="91" pageNumber="92">
<collectingMunicipality id="9358785F181FFF8144BB372AAF09EF70" box="[1012,1086,1573,1596]" pageId="91" pageNumber="92">Border</collectingMunicipality>
of
<collectingRegion id="B1472CC7181FFF814328372AAA66EF10" country="Madagascar" name="Fianarantsoa" pageId="91" pageNumber="92">Fianarantsoa Province</collectingRegion>
and
<location id="765CB4FE181FFF8146DC374AA975EF10" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE181FFF8146DC374AA975EF10" box="[403,578,1605,1628]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Tulear Province" pageId="91" pageNumber="92" stateProvince="Fianarantsoa">Tulear Province</location>
</emphasis>
:
<locationDeviation id="628EFB07181FFF814519374AA8AFEF10" box="[598,920,1605,1628]" location="Ilikaka Saphir" pageId="91" pageNumber="92">
<quantity id="B47B4FC0181FFF814519374AA9A5EF10" box="[598,658,1605,1628]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="91" pageNumber="92" unit="km" value="9.0">9 km</quantity>
SW of Ilikaka Saphir
</locationDeviation>
[sic., Sapphire] town, 22°37.73ʹS, 45°21.67ʹE, elev.
<quantity id="B47B4FC0181FFF81468F376AA93BEF30" box="[448,524,1637,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="91" pageNumber="92" unit="m" value="1036.0">
<elevation id="F8AE0516181FFF81468F376AA93BEF30" box="[448,524,1637,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="91" pageNumber="92" unit="m" value="1036.0">1036m</elevation>
</quantity>
, malaise in
<taxonomicName id="B48399A6181FFF8145DE376AA9D2EF30" box="[657,741,1637,1660]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="91" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E37181FFF8145DE376AA9D2EF30" box="[657,741,1637,1660]" italics="true" pageId="91" pageNumber="92">Uapaca</emphasis>
</taxonomicName>
forest,
<date id="073DC4E5181FFF814475376AA8E0EF30" box="[826,983,1637,1660]" pageId="91" pageNumber="92" value="2002-03-06">
<collectingDate id="17793D0D181FFF814475376AA8E0EF30" box="[826,983,1637,1660]" pageId="91" pageNumber="92" value="2002-03-06">6 March 2002</collectingDate>
</date>
,
<collectorName id="DE7687F3181FFF8144AB376AAF72EF30" box="[996,1093,1637,1660]" pageId="91" pageNumber="92">M. Irwin</collectorName>
and
<collectorName id="DE7687F3181FFF814332376AABC4EFD1" pageId="91" pageNumber="92">R. Harin’Hala</collectorName>
[MG-12-11] (
<specimenCode id="23254A5E181FFF8146C23789A961EFD1" box="[397,598,1670,1693]" collectionCode="CASENT" pageId="91" pageNumber="92">CASENT9032853</specimenCode>
,
<specimenCount id="658529AC181FFF81452C3789A9B3EFD1" box="[611,644,1670,1693]" count="1" pageId="91" pageNumber="92" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0181FFF8145DC3789A9FDEFD1" box="[659,714,1670,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="91" pageNumber="92" type="Museum">CAS</collectionCode>
),
<date id="073DC4E5181FFF8145903789AF3EEFD1" box="[735,1033,1670,1693]" pageId="91" pageNumber="92" value="2002-02-27" valueMax="2002-03-06" valueMin="2002-02-27">
<collectingDate id="17793D0D181FFF8145903789AF3EEFD1" box="[735,1033,1670,1693]" pageId="91" pageNumber="92" value="2002-02-27" valueMax="2002-03-06" valueMin="2002-02-27">27 February–6 March 2002</collectingDate>
</date>
[MG-12-10] (
<specimenCode id="23254A5E181FFF8147CD37A9AA7CEFF1" box="[130,331,1702,1725]" collectionCode="CASENT" pageId="91" pageNumber="92">CASENT9032855</specimenCode>
,
<specimenCount id="658529AC181FFF81461937A9AA40EFF1" box="[342,375,1702,1725]" count="1" pageId="91" pageNumber="92" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0181FFF8146CB37A9AA8BEFF1" box="[388,444,1702,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="91" pageNumber="92" type="Museum">CAS</collectionCode>
).
<emphasis id="41F73E37181FFF81468137A9A999EFF1" box="[462,686,1702,1725]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="B1472CC7181FFF81468137A9A999EFF1" box="[462,686,1702,1725]" country="Madagascar" name="Mahajanga" pageId="91" pageNumber="92">Mahajanga Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE181FFF8145F437A9A827EFF1" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE181FFF8145F437A9A827EFF1" box="[699,784,1702,1725]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Western" pageId="91" pageNumber="92" stateProvince="Fianarantsoa">Western</location>
slope of
<location id="765CB4FE181FFF81442137A9AF04EFF1" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE181FFF81442137A9AF04EFF1" box="[878,1075,1702,1725]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Anjanaharibe-Sud" pageId="91" pageNumber="92" stateProvince="Fianarantsoa">Anjanaharibe-Sud</location>
,
<locationDeviation id="628EFB07181FFF86437237A9AA16E9BF" lastPageId="92" lastPageNumber="93" location="Befingotra" pageId="91" pageNumber="92">
<quantity id="B47B4FC0181FFF81437237A9AFA2EFF1" box="[1085,1173,1702,1725]" metricMagnitude="4" metricUnit="m" metricValue="1.35" pageId="91" pageNumber="92" unit="km" value="13.5">13.5 km</quantity>
SW Befingotra
</locationDeviation>
, 14°47.0ʹS, 49°26.5ʹE, elev.
<quantity id="B47B4FC01818FF86451231D3A99DE9BF" box="[605,682,220,243]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="92" pageNumber="93" unit="m" value="1600.0">
<elevation id="F8AE05161818FF86451231D3A99DE9BF" box="[605,682,220,243]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="92" pageNumber="93" unit="m" value="1600.0">1600m</elevation>
</quantity>
, undisturbed transitional montane rainforest,
<location id="765CB4FE1818FF86473431F3AB8CE85F" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF86473431F3AB8CE85F" box="[123,187,252,275]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Camp" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Camp</location>
#1,
<collectingMethod id="AAC29A321818FF8647A531F3AA1DE85F" box="[234,298,252,275]" pageId="92" pageNumber="93">pitfall</collectingMethod>
7,
<date id="073DC4E51818FF86460331F3A9A1E85F" box="[332,662,252,275]" pageId="92" pageNumber="93" value="1999-10-26" valueMax="1999-11-01" valueMin="1999-10-26">
<collectingDate id="17793D0D1818FF86460331F3A9A1E85F" box="[332,662,252,275]" pageId="92" pageNumber="93" value="1999-10-26" valueMax="1999-11-01" valueMin="1999-10-26">26 October–1 November 1999</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8645EE31F3A811E85F" box="[673,806,252,275]" pageId="92" pageNumber="93">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1818FF86447A31F3A8C9E85F" box="[821,1022,252,275]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9064657</specimenCode>
,
<specimenCount id="658529AC1818FF86434531F3AF1CE85F" box="[1034,1067,252,275]" count="1" pageId="92" pageNumber="93" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01818FF86437631F3AFBCE85F" box="[1081,1163,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="92" pageNumber="93" type="Museum">FMNH</collectionCode>
).
<emphasis id="41F73E371818FF8647343012AA1DE878" box="[123,298,285,308]" italics="true" pageId="92" pageNumber="93">
<collectingRegion id="B1472CC71818FF8647343012AA1DE878" box="[123,298,285,308]" country="Madagascar" name="Toliara" pageId="92" pageNumber="93">Toliara Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1818FF8646773012AAF0E878" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF8646773012AAF0E878" box="[312,455,285,308]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Lake Ranobe" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Lake Ranobe</location>
, 23°02.524ʹS, 43°37.214ʹE, elev.
<quantity id="B47B4FC01818FF8644663012A86CE878" box="[809,859,285,308]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="92" pageNumber="93" unit="m" value="20.0">
<elevation id="F8AE05161818FF8644663012A86CE878" box="[809,859,285,308]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="92" pageNumber="93" unit="m" value="20.0">20m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321818FF86442A3012AB98E818" pageId="92" pageNumber="93">general collecting and pitfall traps</collectingMethod>
in spiny forest,
<date id="073DC4E51818FF86463B3032A90EE818" box="[372,569,317,340]" pageId="92" pageNumber="93" value="2003-05-13" valueMax="2003-05-21" valueMin="2003-05-13">
<collectingDate id="17793D0D1818FF86463B3032A90EE818" box="[372,569,317,340]" pageId="92" pageNumber="93" value="2003-05-13" valueMax="2003-05-21" valueMin="2003-05-13">13–21 May 2003</collectingDate>
</date>
,
<location id="765CB4FE1818FF8645053032A84CE818" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF8645053032A84CE818" box="[586,891,317,340]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Frontier Wilderness Project" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Frontier Wilderness Project</location>
[MGF072] (
<typeStatus id="AC385C871818FF8643403032AF59E818" box="[1039,1134,317,340]" pageId="92" pageNumber="93" type="holotype">holotype</typeStatus>
<specimenCount id="658529AC1818FF8643343032AFB8E818" box="[1147,1167,317,340]" count="1" pageId="92" pageNumber="93" type="male">♂</specimenCount>
,
<specimenCode id="23254A5E1818FF8647343051AA74E839" box="[123,323,350,373]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9017834</specimenCode>
,
<typeStatus id="AC385C871818FF86461C3051AA98E839" box="[339,431,350,373]" pageId="92" pageNumber="93" type="paratype">paratype</typeStatus>
<specimenCount id="658529AC1818FF8646F33051AAE7E839" box="[444,464,350,373]" count="1" pageId="92" pageNumber="93" type="female">♀</specimenCount>
,
<specimenCode id="23254A5E1818FF8646AC3051A99CE839" box="[483,683,350,373]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9064773</specimenCode>
,
<collectionCode id="15927AE01818FF8645F33051A9C4E839" box="[700,755,350,373]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
), [
<specimenCode id="23254A5E1818FF86445D3051A84EE839" box="[786,889,350,373]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">MGF067</specimenCode>
] (
<specimenCode id="23254A5E1818FF8644DC3051AF6BE839" box="[915,1116,350,373]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9017831</specimenCode>
,
<specimenCount id="658529AC1818FF8643223051AFB8E839" box="[1133,1167,350,373]" count="1" pageId="92" pageNumber="93" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01818FF8647343070AB84E8DA" box="[123,179,383,406]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1818FF8647893070AA60E8DA" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF8647893070AA60E8DA" box="[198,343,383,406]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Mikea Forest" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Mikea Forest</location>
,
<locationDeviation id="628EFB071818FF86462C3070A914E8DA" box="[355,547,383,406]" location="Manombo" pageId="92" pageNumber="93">NW of Manombo</locationDeviation>
, 22°54.22ʹS, 43°28.53ʹE, elev.
<quantity id="B47B4FC01818FF8644203070A896E8DA" box="[879,929,383,406]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="92" pageNumber="93" unit="m" value="30.0">
<elevation id="F8AE05161818FF8644203070A896E8DA" box="[879,929,383,406]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="92" pageNumber="93" unit="m" value="30.0">30m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321818FF8644E23070AF18E8DA" box="[941,1071,383,406]" pageId="92" pageNumber="93">malaise trap</collectingMethod>
in deciduous dry forest,
<date id="073DC4E51818FF8646133090A956E8FA" box="[348,609,415,438]" pageId="92" pageNumber="93" value="2002-10-13" valueMax="2002-10-21" valueMin="2002-10-13">
<collectingDate id="17793D0D1818FF8646133090A956E8FA" box="[348,609,415,438]" pageId="92" pageNumber="93" value="2002-10-13" valueMax="2002-10-21" valueMin="2002-10-13">13–21 October 2002</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8645333090A9D9E8FA" box="[636,750,415,438]" pageId="92" pageNumber="93">M. Irwin</collectorName>
,
<collectorName id="DE7687F31818FF8644453090A88EE8FA" box="[778,953,415,438]" pageId="92" pageNumber="93">R. Harin’Hala</collectorName>
[MA-02-18A-36] (
<specimenCode id="23254A5E1818FF8647CD30CFAA7BE89B" box="[130,332,448,471]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9030930</specimenCode>
,
<specimenCount id="658529AC1818FF86461330CFAA4AE89B" box="[348,381,448,471]" count="1" pageId="92" pageNumber="93" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01818FF8646DF30CFAAFFE89B" box="[400,456,448,471]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1818FF8646AE30CFA982E89B" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF8646AE30CFA982E89B" box="[481,693,448,471]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Foret de Beroboka" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">
Forêt
<collectorName id="DE7687F31818FF86456A30CFA982E89B" box="[549,693,448,471]" pageId="92" pageNumber="93">de Beroboka</collectorName>
</location>
,
<locationDeviation id="628EFB071818FF86458930CFAF32E89B" box="[710,1029,448,471]" location="Ankidranoka" pageId="92" pageNumber="93">
<quantity id="B47B4FC01818FF86458930CFA827E89B" box="[710,784,448,471]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="92" pageNumber="93" unit="km" value="5.9">5.9 km</quantity>
131°SE Ankidranoka
</locationDeviation>
,
<geoCoordinate id="16B784E21818FF86435930CFAFA7E89B" box="[1046,1168,448,471]" degrees="22" direction="south" minutes="13" orientation="latitude" pageId="92" pageNumber="93" precision="15" seconds="59" value="-22.233055">22°13ʹ59ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21818FF86473430EEABCFE8B4" box="[123,248,481,504]" degrees="43" direction="east" minutes="21" orientation="longitude" pageId="92" pageNumber="93" precision="15" seconds="59" value="43.366386">43°21ʹ59ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01818FF86467730EEAA5CE8B4" box="[312,363,481,504]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="92" pageNumber="93" unit="m" value="80.0">
<elevation id="F8AE05161818FF86467730EEAA5CE8B4" box="[312,363,481,504]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="92" pageNumber="93" unit="m" value="80.0">80m</elevation>
</quantity>
, tropical dry forest,
<collectingMethod id="AAC29A321818FF86450F30EEA985E8B4" box="[576,690,481,504]" pageId="92" pageNumber="93">pitfall trap</collectingMethod>
,
<date id="073DC4E51818FF8645F230EEA8BDE8B4" box="[701,906,481,504]" pageId="92" pageNumber="93" value="2002-03-12" valueMax="2002-03-16" valueMin="2002-03-12">
<collectingDate id="17793D0D1818FF8645F230EEA8BDE8B4" box="[701,906,481,504]" pageId="92" pageNumber="93" value="2002-03-12" valueMax="2002-03-16" valueMin="2002-03-12">12–16 March 2002</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8644DB30EEABD4EB54" pageId="92" pageNumber="93">Fisher-Griswold Arthropod Team</collectorName>
[BLF6068] (
<specimenCode id="23254A5E1818FF86463B330EA909EB54" box="[372,574,513,536]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9000536</specimenCode>
,
<specimenCount id="658529AC1818FF864505330EA95CEB54" box="[586,619,513,536]" count="1" pageId="92" pageNumber="93" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01818FF864534330EA984EB54" box="[635,691,513,536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1818FF86458B330EA8D1EB54" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF86458B330EA8D1EB54" box="[708,998,513,536]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Andohahela National Park" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Andohahela National Park</location>
,
<collectorName id="DE7687F31818FF8644BD330EAF6CEB54" box="[1010,1115,513,536]" pageId="92" pageNumber="93">Tsimelahy</collectorName>
,
<collectorName id="DE7687F31818FF864326330EAB9AEB75" pageId="92" pageNumber="93">Parcelle</collectorName>
#II, 24°56.21ʹS, 46°37.60ʹE, elev.
<quantity id="B47B4FC01818FF864570332DA94BEB75" box="[575,636,546,569]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="92" pageNumber="93" unit="m" value="180.0">
<elevation id="F8AE05161818FF864570332DA94BEB75" box="[575,636,546,569]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="92" pageNumber="93" unit="m" value="180.0">180m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321818FF8645DA332DA82AEB75" box="[661,797,546,569]" pageId="92" pageNumber="93">malaise trap</collectingMethod>
in transitional forest,
<date id="073DC4E51818FF86436D332DAB84EB15" pageId="92" pageNumber="93" value="2003-05-03" valueMax="2003-05-15" valueMin="2003-05-03">
<collectingDate id="17793D0D1818FF86436D332DAB84EB15" pageId="92" pageNumber="93" value="2003-05-03" valueMax="2003-05-15" valueMin="2003-05-03">3–15 May 2003</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8647F1334DAA17EB15" box="[190,288,578,601]" pageId="92" pageNumber="93">M. Irwin</collectorName>
,
<collectorName id="DE7687F31818FF864664334DAAB0EB15" box="[299,391,578,601]" pageId="92" pageNumber="93">F. Parker</collectorName>
, and
<collectorName id="DE7687F31818FF86468C334DA96CEB15" box="[451,603,578,601]" pageId="92" pageNumber="93">R. Harin’Hala</collectorName>
[MA-02-20-24] (
<specimenCode id="23254A5E1818FF864457334DA8D6EB15" box="[792,993,578,601]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9021818</specimenCode>
,
<specimenCount id="658529AC1818FF8644A2334DAF39EB15" box="[1005,1038,578,601]" count="1" pageId="92" pageNumber="93" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01818FF864353334DAF63EB15" box="[1052,1108,578,601]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1818FF864328334DAA23EB36" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF864328334DAA23EB36" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Forest des Milua" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Forest des Milua</location>
,
<locationDeviation id="628EFB071818FF86466C336CA93BEB36" box="[291,524,611,634]" location="Tanadava" pageId="92" pageNumber="93">
<quantity id="B47B4FC01818FF86466C336CAA5EEB36" box="[291,361,611,634]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="92" pageNumber="93" unit="km" value="19.0">19 km</quantity>
SW Tanadava
</locationDeviation>
, 21°52.00ʹS, 43°39.60ʹE, elev.
<quantity id="B47B4FC01818FF864429336CA8A0EB36" box="[870,919,611,634]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="92" pageNumber="93" unit="m" value="50.0">
<elevation id="F8AE05161818FF864429336CA8A0EB36" box="[870,919,611,634]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="92" pageNumber="93" unit="m" value="50.0">50m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321818FF8644E8336CA8C3EB36" box="[935,1012,611,634]" pageId="92" pageNumber="93">pitfalls</collectingMethod>
,
<date id="073DC4E51818FF86434D336CAB84EBD7" pageId="92" pageNumber="93" value="2003-03-16" valueMax="2003-03-25" valueMin="2003-03-16">
<collectingDate id="17793D0D1818FF86434D336CAB84EBD7" pageId="92" pageNumber="93" value="2003-03-16" valueMax="2003-03-25" valueMin="2003-03-16">16–25 March 2003</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8647F0338BAA5EEBD7" box="[191,361,644,667]" pageId="92" pageNumber="93">V. Sourivalalam</collectorName>
(
<specimenCode id="23254A5E1818FF864635338BA906EBD7" box="[378,561,644,667]" collectionCode="FMHD" pageId="92" pageNumber="93">FMHD#2003-10</specimenCode>
,
<specimenCount id="658529AC1818FF864571338BA968EBD7" box="[574,607,644,667]" count="1" pageId="92" pageNumber="93" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01818FF864520338BA9F6EBD7" box="[623,705,644,667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="92" pageNumber="93" type="Museum">FMNH</collectionCode>
);
<location id="765CB4FE1818FF864599338BA8EBEBD7" LSID="urn:lsid:plazi:treatment:FB2A5333181DFF8647343502AFA2EDE9:765CB4FE1818FF864599338BA8EBEBD7" box="[726,988,644,667]" country="Madagascar" county="Italaviana" latitude="-22.233055" longLatPrecision="21" longitude="43.366386" municipality="Border" name="Beza Mahafaly Reserve" pageId="92" pageNumber="93" stateProvince="Fianarantsoa">Beza Mahafaly Reserve</location>
,
<collectorName id="DE7687F31818FF8644A6338BAF08EBD7" box="[1001,1087,644,667]" pageId="92" pageNumber="93">Parcelle</collectorName>
#I near research station, 23°41.19ʹS, 44°35.46ʹE, elev.
<quantity id="B47B4FC01818FF8645CA33ABA9F3EBF7" box="[645,708,676,699]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="92" pageNumber="93" unit="m" value="165.0">
<elevation id="F8AE05161818FF8645CA33ABA9F3EBF7" box="[645,708,676,699]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="92" pageNumber="93" unit="m" value="165.0">165m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321818FF86459B33ABA86EEBF7" box="[724,857,676,699]" pageId="92" pageNumber="93">malaise trap</collectingMethod>
in dry deciduous forest,
<date id="073DC4E51818FF86433433ABAAF4EB90" pageId="92" pageNumber="93" value="2001-11-28" valueMax="2001-12-04" valueMin="2001-11-28">
<collectingDate id="17793D0D1818FF86433433ABAAF4EB90" pageId="92" pageNumber="93" value="2001-11-28" valueMax="2001-12-04" valueMin="2001-11-28">28 November–4 December 2001</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF86469C33CAA96BEB90" box="[467,604,709,732]" pageId="92" pageNumber="93">M. E. Irwin</collectorName>
,
<collectorName id="DE7687F31818FF86452333CAA9C2EB90" box="[620,757,709,732]" pageId="92" pageNumber="93">F. D. Parker</collectorName>
and
<collectorName id="DE7687F31818FF86447C33CAA8F8EB90" box="[819,975,709,732]" pageId="92" pageNumber="93">R. Harin’Hala</collectorName>
[MA-02-14A-04] (
<specimenCode id="23254A5E1818FF8647CD33E9AA7CEBB1" box="[130,331,742,765]" collectionCode="CASENT" pageId="92" pageNumber="93">CASENT9042545</specimenCode>
,
<specimenCount id="658529AC1818FF86462F33E9AAB6EBB1" box="[352,385,742,765]" count="1" pageId="92" pageNumber="93" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01818FF8646D733E9AAE7EBB1" box="[408,464,742,765]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="92" pageNumber="93" type="Museum">CAS</collectionCode>
).
<emphasis id="41F73E371818FF8646A433E9A9AFEBB1" box="[491,664,742,765]" italics="true" pageId="92" pageNumber="93">
<collectingCountry id="0B94A2B51818FF8646A433E9A9AFEBB1" box="[491,664,742,765]" name="Madagascar" pageId="92" pageNumber="93">MADAGASCAR</collectingCountry>
</emphasis>
:
<emphasis id="41F73E371818FF8645FF33E9A81CEBB1" box="[688,811,742,765]" italics="true" pageId="92" pageNumber="93">no locality</emphasis>
: probably
<collectorName id="DE7687F31818FF8644FE33E9AF08EBB1" box="[945,1087,742,765]" pageId="92" pageNumber="93">Ambovombe</collectorName>
,
<date id="073DC4E51818FF86431C33E9AA1BEA51" pageId="92" pageNumber="93" value="1901-11-19" valueMax="1901-11-30" valueMin="1901-11-19">
<collectingDate id="17793D0D1818FF86431C33E9AA1BEA51" pageId="92" pageNumber="93" value="1901-11-19" valueMax="1901-11-30" valueMin="1901-11-19">19–30 November 1901</collectingDate>
</date>
,
<collectorName id="DE7687F31818FF8646773209AA99EA51" box="[312,430,774,797]" pageId="92" pageNumber="93">Carougeau</collectorName>
, (
<specimenCount id="658529AC1818FF86468C3209AAD9EA51" box="[451,494,774,797]" count="7" pageId="92" pageNumber="93" type="female">7 ♀</specimenCount>
,
<collectionCode id="15927AE01818FF8646B23209A964EA51" box="[509,595,774,797]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="92" pageNumber="93">MNHN</collectionCode>
)
</materialsCitation>
,
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listed as follows: (
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), (
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.
</paragraph>
<paragraph id="733CE2251818FF8647E43286A93DED28" blockId="92.[123,1174,220,1189]" pageId="92" pageNumber="93">
<emphasis id="41F73E371818FF8647E43286AA57EAEC" bold="true" box="[171,352,905,928]" pageId="92" pageNumber="93">Natural history.</emphasis>
The species
<taxonomicName id="B48399A61818FF8646BF3286A9A1EAEC" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[496,662,905,928]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371818FF8646BF3286A9A1EAEC" bold="true" box="[496,662,905,928]" italics="true" pageId="92" pageNumber="93">Uduba woodae</emphasis>
</taxonomicName>
appears to occur mostly in dry parts of
<collectingCountry id="0B94A2B51818FF86431F3286AB88EA8C" name="Madagascar" pageId="92" pageNumber="93">Madagascar</collectingCountry>
(Maps 8, 14), and is recorded from dry deciduous forest, tropical dry forest, transitional dry forest in Andohahela,
<taxonomicName id="B48399A61818FF86462232C5AAF6EAAD" box="[365,449,970,993]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="92" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E371818FF86462232C5AAF6EAAD" box="[365,449,970,993]" italics="true" pageId="92" pageNumber="93">Uapaca</emphasis>
</taxonomicName>
forest and even spiny forest. Collection of individuals in malaise traps in addition to terrestrial pitfall traps suggests that wandering males may climb into vegetation in their search for females; females may also wander, as one was collected in a terrestrial pitfall trap.
<taxonomicName id="B48399A61818FF8647FD3523AA61ED0F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[178,342,1068,1091]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371818FF8647FD3523AA61ED0F" bold="true" box="[178,342,1068,1091]" italics="true" pageId="92" pageNumber="93">Uduba woodae</emphasis>
</taxonomicName>
spiders are cribellate, with a divided cribellum or vestige, but we know nothing of the use of silk by this species.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1818FF8647E43562AFA2EDE9" pageId="92" pageNumber="93" type="distribution">
<paragraph id="733CE2251818FF8647E43562AFA2EDE9" blockId="92.[123,1174,220,1189]" pageId="92" pageNumber="93">
<emphasis id="41F73E371818FF8647E43562AADCEDC8" bold="true" box="[171,491,1133,1156]" pageId="92" pageNumber="93">
Distribution.
<taxonomicName id="B48399A61818FF86460A3562AADCEDC8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[325,491,1133,1156]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371818FF86460A3562AADCEDC8" bold="true" box="[325,491,1133,1156]" italics="true" pageId="92" pageNumber="93">Uduba woodae</emphasis>
</taxonomicName>
</emphasis>
specimens have been collected in Southern
<collectingCountry id="0B94A2B51818FF8644803562AF67EDC8" box="[975,1104,1133,1156]" name="Madagascar" pageId="92" pageNumber="93">Madagascar</collectingCountry>
in the drier portions of
<collectingRegion id="B1472CC71818FF86467C3581AA8CEDE9" box="[307,443,1166,1189]" country="Madagascar" name="Fianarantsoa" pageId="92" pageNumber="93">Fianarantsoa</collectingRegion>
and Tulear Provinces with an outlier in the northeast (Maps 8, 14).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="178CF8F553E7FC56E7E27B93A61FA4F2">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="0C1FA1A046FFBABED6D35A2795976CCF">Griswold, Charles</mods:namePart>
<mods:affiliation id="AC031B138F5081A7AD812F3882FEEA57">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="A65B4432C46D848915CEC1C8AA41789E">Ubick, Darrell</mods:namePart>
<mods:affiliation id="2FE101ECA4159E6DC55DC773F6813D89">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="DF43C902E3ADFF0A732C77C4EFC10750">Ledford, Joel</mods:namePart>
<mods:affiliation id="E7A64A747E616F01516033C672D0699D">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="5DF6272E3BF8B743730B092518E523BA">Polotow, Daniele</mods:namePart>
<mods:affiliation id="12F2064E915E7D91FBF838E118BC84B1">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:date id="61032207949285BA76E46B5198DE9138">2022</mods:date>
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<paragraph id="733CE2251851FFCF473433F0AAF5EA5A" blockId="21.[123,540,767,820]" box="[123,450,767,790]" pageId="21" pageNumber="22">
<heading id="287455491851FFCF473433F0AAF5EA5A" box="[123,450,767,790]" fontSize="10" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="41F73E371851FFCF473433F0AAF5EA5A" bold="true" box="[123,450,767,790]" pageId="21" pageNumber="22">
<taxonomicName id="B48399A61851FFCF473433F0AA18EA5A" ID-CoL="C8KJ5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,303,767,790]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
<emphasis id="41F73E371851FFCF473433F0AA18EA5A" bold="true" box="[123,303,767,790]" italics="true" pageId="21" pageNumber="22">Uduba barbarae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1851FFCF467233F0AAF5EA5A" box="[317,450,767,790]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1851FFCF47343210A92BEA78" box="[123,540,799,820]" pageId="21" pageNumber="22" type="description">
<paragraph id="733CE2251851FFCF47343210A92BEA78" blockId="21.[123,540,767,820]" box="[123,540,799,820]" pageId="21" pageNumber="22">
<figureCitation id="EBB8FEA01851FFCF47343210ABD8EA78" box="[123,239,799,820]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="21" pageNumber="22">Figures 7 H</figureCitation>
,
<figureCitation id="EBB8FEA01851FFCF47B53210AA70EA78" box="[250,327,799,820]" captionStart="FIGURE" captionStartId="114.[155,167,1160,1179]" captionTargetBox="[297,999,216,1140]" captionTargetId="figure-11@114.[297,999,216,1140]" captionTargetPageId="114" captionText="FIGURE 10. Habitus drawings, Uduba barbarae ♀ (CASENT9066066), Nosy Be Island. A. Lateral. B. Dorsal. C. Ventral. Illustrations by Jenny Speckels. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158582" httpUri="https://zenodo.org/record/13158582/files/figure.png" pageId="21" pageNumber="22">10 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01851FFCF461E3210AAA9EA78" box="[337,414,799,820]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="21" pageNumber="22">76 A, B</figureCitation>
, Maps 4, 15.
</paragraph>
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<paragraph id="733CE2251851FFCF47E4324BAF53EAF7" blockId="21.[122,1174,836,1725]" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E4324BAA52EA17" bold="true" box="[171,357,836,859]" pageId="21" pageNumber="22">Type material.</emphasis>
<materialsCitation id="C3EBE8781851FFCF4638324BAF53EAF7" collectingDate="1992-08-11" collectingDateMax="1992-08-14" collectingDateMin="1992-08-11" collectionCode="CAS" county="Barbara Roth" latitude="-13.416333" location="Lokobe Forest" longLatPrecision="1" longitude="48.30736" municipality="Vince" pageId="21" pageNumber="22" specimenCode="CASENT9006067, CASENT9006064, CASENT9006066, CASENT9006068, CASENT9006069" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<typeStatus id="AC385C871851FFCF4638324BAAD5EA17" box="[375,482,836,859]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E1851FFCF4514324BA818EA17" box="[603,815,836,859]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9006067</specimenCode>
) and
<specimenCount id="658529AC1851FFCF44CA324BAFA2EA17" box="[901,1173,836,859]" count="4" pageId="21" pageNumber="22" type="female" typeStatus="paratype">four paratype females</specimenCount>
(
<specimenCode id="23254A5E1851FFCF47CD326BAA7DEA37" box="[130,330,868,891]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9006064</specimenCode>
,
<specimenCode id="23254A5E1851FFCF462E326BA91FEA37" box="[353,552,868,891]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9006066</specimenCode>
,
<specimenCode id="23254A5E1851FFCF4570326BA831EA37" box="[575,774,868,891]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9006068</specimenCode>
and
<specimenCode id="23254A5E1851FFCF4402326BAF21EA37" box="[845,1046,868,891]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9006069</specimenCode>
), collected along forest trails in
<location id="765CB4FE1851FFCF4621328BA926EAD7" LSID="urn:lsid:plazi:treatment:FB2A53331851FFCC473433F0A863ED1A:765CB4FE1851FFCF4621328BA926EAD7" box="[366,529,900,923]" county="Barbara Roth" latitude="-13.416333" longLatPrecision="1" longitude="48.30736" municipality="Vince" name="Lokobe Forest" pageId="21" pageNumber="22">Lokobe Forest</location>
, near sea level on the island of
<location id="765CB4FE1851FFCF44C2328BA8D8EAD7" LSID="urn:lsid:plazi:treatment:FB2A53331851FFCC473433F0A863ED1A:765CB4FE1851FFCF44C2328BA8D8EAD7" box="[909,1007,900,923]" county="Barbara Roth" latitude="-13.416333" longLatPrecision="1" longitude="48.30736" municipality="Vince" name="Nosy Be" pageId="21" pageNumber="22">Nosy Be</location>
(
<geoCoordinate id="16B784E21851FFCF434C328BAFA7EAD7" box="[1027,1168,900,923]" degrees="13" direction="south" minutes="24" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="58.8" value="-13.416333">13º24ʹ58.8ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21851FFCF473432ABAA3EEAF7" box="[123,265,932,955]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="26.5" value="48.30736">48º18ʹ26.5ʺE</geoCoordinate>
),
<date id="073DC4E51851FFCF465132ABAAD9EAF7" box="[286,494,932,955]" pageId="21" pageNumber="22" value="1992-08-11" valueMax="1992-08-14" valueMin="1992-08-11">
<collectingDate id="17793D0D1851FFCF465132ABAAD9EAF7" box="[286,494,932,955]" pageId="21" pageNumber="22" value="1992-08-11" valueMax="1992-08-14" valueMin="1992-08-11">11–14 August 1992</collectingDate>
</date>
by
<collectingMunicipality id="9358785F1851FFCF455732ABA963EAF7" box="[536,596,932,955]" pageId="21" pageNumber="22">Vince</collectingMunicipality>
and
<collectingCounty id="9A5D9AA91851FFCF45C432ABA82BEAF7" box="[651,796,932,955]" pageId="21" pageNumber="22">Barbara Roth</collectingCounty>
[VR0003], deposited in
<collectionCode id="15927AE01851FFCF436632ABAF57EAF7" box="[1065,1120,932,955]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="22" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1851FFCF47E432CBA934EAB0" pageId="21" pageNumber="22" type="etymology">
<paragraph id="733CE2251851FFCF47E432CBA934EAB0" blockId="21.[122,1174,836,1725]" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E432CBAA1DEA97" bold="true" box="[171,298,964,987]" pageId="21" pageNumber="22">Etymology.</emphasis>
The species epithet is to honor Mrs. Barbara Roth, collector of this and many other interesting arachnids in
<collectingCountry id="0B94A2B51851FFCF463132EAAACBEAB0" box="[382,508,997,1020]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1851FFCF47E4350AAAB8EDF0" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="733CE2251851FFCF47E4350AAAB8EDF0" blockId="21.[122,1174,836,1725]" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E4350AAAEDED50" bold="true" box="[171,474,1029,1052]" pageId="21" pageNumber="22">
Diagnosis.
<taxonomicName id="B48399A61851FFCF4669350AAAEDED50" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,474,1029,1052]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="41F73E371851FFCF4669350AAAEDED50" bold="true" box="[294,474,1029,1052]" italics="true" pageId="21" pageNumber="22">Uduba barbarae</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.c, the
<taxonomicName id="B48399A61851FFCF4465350AA8FFED50" authorityName="Simon" authorityYear="1906" box="[810,968,1029,1052]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01851FFCF4354350AAF47ED50" box="[1051,1136,1029,1052]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="21" pageNumber="22">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate, with a colulus (
<figureCitation id="EBB8FEA01851FFCF44ED352AAF00ED70" box="[930,1079,1061,1084]" captionStart="FIGURE" captionStartId="114.[155,167,1160,1179]" captionTargetBox="[297,999,216,1140]" captionTargetId="figure-11@114.[297,999,216,1140]" captionTargetPageId="114" captionText="FIGURE 10. Habitus drawings, Uduba barbarae ♀ (CASENT9066066), Nosy Be Island. A. Lateral. B. Dorsal. C. Ventral. Illustrations by Jenny Speckels. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158582" httpUri="https://zenodo.org/record/13158582/files/figure.png" pageId="21" pageNumber="22">Figs. 10 A–C</figureCitation>
); female with epigynal plate with median lobe surrounded by depressed atrium, ML fills most of atrium, atrium width at side of ML only 0.14 ML width, plate extensive posteriad of atrium, height greater than 0.15 times epigynal length; median lobe broad, width atrium less than 3 times ML width (
<figureCitation id="EBB8FEA01851FFCF432F358AABE7EDF0" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="21" pageNumber="22">Figs. 76 A, B</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1851FFCC47E435CAAF59EBB8" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="733CE2251851FFCC47E435CAA83BE89F" blockId="21.[122,1174,836,1725]" lastBlockId="22.[123,1174,220,1110]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E435CAAA4CED90" bold="true" box="[171,379,1221,1244]" pageId="21" pageNumber="22">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E371851FFCF454735CAA96CED90" bold="true" box="[520,603,1221,1244]" pageId="21" pageNumber="22">Female</emphasis>
(
<typeStatus id="AC385C871851FFCF452135CAA9E2ED90" box="[622,725,1221,1244]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
, CASENT9006067): Total length 28.43. Markings as in
<figureCitation id="EBB8FEA01851FFCF466E35EAAAB1EDB0" box="[289,390,1253,1276]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="21" pageNumber="22">Figs. 7 H</figureCitation>
,
<figureCitation id="EBB8FEA01851FFCF46DE35EAAAD2EDB0" box="[401,485,1253,1276]" captionStart="FIGURE" captionStartId="114.[155,167,1160,1179]" captionTargetBox="[297,999,216,1140]" captionTargetId="figure-11@114.[297,999,216,1140]" captionTargetPageId="114" captionText="FIGURE 10. Habitus drawings, Uduba barbarae ♀ (CASENT9066066), Nosy Be Island. A. Lateral. B. Dorsal. C. Ventral. Illustrations by Jenny Speckels. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158582" httpUri="https://zenodo.org/record/13158582/files/figure.png" pageId="21" pageNumber="22">10 A–C</figureCitation>
. Carapace 15.71 long, 10.86 wide, 5.71 high; clypeus 1.00 high. Eye diameters: AME 0.50, ALE 0.45, PME 0.43, PLE 0.54. Chelicerae 8.14 long; sternum 6.43 long, 4.50 wide; labium 3.39 long; palpal coxae 5.54 long. With a colulus. Spination: (
<typeStatus id="AC385C871851FFCF436F342AAFBDEC70" box="[1056,1162,1317,1340]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
): palpus – femur d0-1-2, patella p1-0, tibia p2-1, r1-0, tarsus p2-1, r1-0; leg I – femur d1-0-0, p0- 0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II – femur d0-1-0-1, p0-0-0-1, tibia p0-1-1-0, v2-1-2-2, metatarsus v2-2-3; leg III – femur d1-0-0-1, p1-1-0-1-1, r0-1-1-1, tibia d1-0-0-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-1-2-2, r1-1-2, tarsus v0-1-0; leg IV – femur d1-1-0-1, p0-1-0-1, r0- 0-0-1, tibia p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1-2, v2-1-2-2, r1-1-1-2. Scopulae: ventral on tarsi and metatarsi I-IV and tibia I, apical half of tibia II, and apices tibiae III and IV. Leg measurements [
<typeStatus id="AC385C871851FFCF47A43709AA60EF51" box="[235,343,1542,1565]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
]: I: 12.36 + 5.21 + 9.00 + 7.86 + 4.29 = 38.71; II: 9.71 + 5.07 + 7.86 + 8.14 + 4.00 = 34.79; III: 8.21 + 4.29 + 5.00 + 6.79 + 3.07 = 27.36; IV: 10.57 + 4.64 + 8.71 + 11.14 + 4.57 = 39.64; palpus: 5.79 + 2.93 + 3.07 + NA + 5.00 = 16.79. Leg formula 4123.
<emphasis id="41F73E371851FFCF43463749AB83EF31" bold="true" pageId="21" pageNumber="22">Female genitalia</emphasis>
(
<typeStatus id="AC385C871851FFCF47893769AA11EF31" box="[198,294,1638,1661]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
, CASENT9006066): epigynal plate with median lobe surrounded by depressed atrium; epigynal plate narrow (
<figureCitation id="EBB8FEA01851FFCF46893789A90FEFD1" box="[454,568,1670,1693]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="21" pageNumber="22">Figs. 76 A</figureCitation>
), width equals length; atrium sides narrow, atrium width 1.67 times atrium length; epigynum length 3.33 times atrium length, atrium width 0.50 times epigynum width; ML fills most of atrium, atrium width at side of ML only 0.14 ML width; atrium and median lobe anteromedian, arise at 0.30 epigynum length from epigastric groove; median lobe broad, triangular, median lobe length 1.49 times width, ML length 1.67 times atrium length, 0.50 times epigynal length, ML width 0.70 atrium width, ML apex extends far anteriad of atrium apex by nearly one-half ML length; lateral lobes broad, atrium width 2.00 times LL width; copulatory openings beneath anterolateral margins of atrium. Vulva (
<figureCitation id="EBB8FEA01852FFCC45E73073A821E8DF" box="[680,790,380,403]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="22" pageNumber="23">Figs. 76 B</figureCitation>
) with spermathecal ducts making at least four broad spiral curves, vulva length 0.75 times width, fertilization ducts close together, vulva width 12.00 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251852FFCC47E430D3AF59EBB8" blockId="22.[123,1174,220,1110]" pageId="22" pageNumber="23">
<emphasis id="41F73E371852FFCC47E430D3AA41E8BF" bold="true" box="[171,374,476,499]" pageId="22" pageNumber="23">Variation. Female</emphasis>
(N = 5): Total length 21.00–33.14. Carapace length / width = 1.38–1.47, carapace height/ width = 0.47–0.56, PER/ carapace width = 0.40–0.80, PER/OAL = 1.59–3.27, PER/AER = 2.73–3.27, OAL/OQL = 1.00–1.13, OQP/ OQA = 1.03–1.13, clypeus height/ AME diameter width = 1.57–2.56, cheliceral length / clypeus height = 6.67–10.50, sternum length / width = 1.30–1.43, palpal coxa length / width = 2.03–2.41, femur I length / carapace width = 0.94–1.14, metatarsus I length / carapace width = 0.71–0.76, femur IV/ carapace width = 0.94–1.05, palpal tarsus length / carapace width = 0.41–0.49, palpal tarsus length / palpal patella length = 1.50–1.73, palpal tarsus length / palpal tibia length = 1.33–1.67, palpal tarsus length / palpal femur length = 0.81–0.95, palpal tibia length / palpal patella length = 1.03–1.18.
<emphasis id="41F73E371852FFCC440E33D2A84CEBB8" bold="true" box="[833,891,733,756]" pageId="22" pageNumber="23">Male</emphasis>
variation is unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1852FFCC47E433F2AF2EED59" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="733CE2251852FFCC47E433F2A8B2EA99" blockId="22.[123,1174,220,1110]" pageId="22" pageNumber="23">
<emphasis id="41F73E371852FFCC47E433F2AAB9EA58" bold="true" box="[171,398,765,788]" pageId="22" pageNumber="23">Material examined.</emphasis>
<materialsCitation id="C3EBE8781852FFCC46D633F2AF66EA19" collectingDate="1992-08-20" collectingDateMax="1992-08-26" collectingDateMin="1992-08-20" collectionCode="CAS" collectorName="V. and B. Roth" country="Madagascar" latitude="-12.909444" location="English" longLatPrecision="21" longitude="49.109997" municipality="Ankaranana" pageId="22" pageNumber="23" specimenCode="CASENT9006065" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<collectingCountry id="0B94A2B51852FFCC46D633F2A96CEA58" box="[409,603,765,788]" name="Madagascar" pageId="22" pageNumber="23">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371852FFCC452833F2A863EA58" box="[615,852,765,788]" italics="true" pageId="22" pageNumber="23">
<collectingRegion id="B1472CC71852FFCC452833F2A863EA58" box="[615,852,765,788]" country="Madagascar" name="Antsiranana" pageId="22" pageNumber="23">Antsiranana Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1852FFCC442A33F2A8DEEA58" box="[869,1001,765,788]" pageId="22" pageNumber="23">Ankaranana</collectingMunicipality>
,
<location id="765CB4FE1852FFCC44B733F2AF7EEA58" LSID="urn:lsid:plazi:treatment:FB2A53331851FFCC473433F0A863ED1A:765CB4FE1852FFCC44B733F2AF7EEA58" box="[1016,1097,765,788]" country="Madagascar" latitude="-12.909444" longLatPrecision="21" longitude="49.109997" municipality="Ankaranana" name="English" pageId="22" pageNumber="23" stateProvince="Antsiranana">English</location>
camp,
<geoCoordinate id="16B784E21852FFCC47343211ABC5EA79" box="[123,242,798,821]" degrees="12" direction="south" minutes="54" orientation="latitude" pageId="22" pageNumber="23" precision="15" seconds="34" value="-12.909444">12º54ʹ34ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21852FFCC464F3211AA5AEA79" box="[256,365,798,821]" degrees="49" direction="east" minutes="6" orientation="longitude" pageId="22" pageNumber="23" precision="15" seconds="36" value="49.109997">49º6ʹ36ʺE</geoCoordinate>
, “
<quantity id="B47B4FC01852FFCC46CC3211AA84EA79" box="[387,435,798,821]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="cm" value="3.0">3cm</quantity>
diameter hole x
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deep in trail, few leaves around entrance tied by few silken threads”,
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<collectingMunicipality id="9358785F1852FFCC43123231ABACEA39" pageId="22" pageNumber="23">Nosy Be</collectingMunicipality>
,
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, “in 5/8” x 4-6” burrow on forest trail”,
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.
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<paragraph id="733CE2251852FFCC47E432D1AF2EED59" blockId="22.[123,1174,220,1110]" pageId="22" pageNumber="23">
<emphasis id="41F73E371852FFCC47E432D1AA56EAB9" bold="true" box="[171,353,990,1013]" pageId="22" pageNumber="23">Natural history.</emphasis>
Specimens of
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<emphasis id="41F73E371852FFCC454632D1A9F7EAB9" bold="true" box="[521,704,990,1013]" italics="true" pageId="22" pageNumber="23">Uduba barbarae</emphasis>
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were collected in lowland rainforest. This ecribellate species forms deep burrows with a small amount of silk near the entrance.
</paragraph>
</subSubSection>
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<emphasis id="41F73E371852FFCC47E43510A936ED7A" bold="true" box="[171,513,1055,1078]" pageId="22" pageNumber="23">
Distribution.
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<emphasis id="41F73E371852FFCC46073510A936ED7A" bold="true" box="[328,513,1055,1078]" italics="true" pageId="22" pageNumber="23">Uduba barbarae</emphasis>
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are known only from the island of Nosy Be and western coastal
<collectingRegion id="B1472CC71852FFCC47853530AA85ED1A" box="[202,434,1087,1110]" country="Madagascar" name="Antsiranana" pageId="22" pageNumber="23">Antsiranana Province</collectingRegion>
in northern
<collectingCountry id="0B94A2B51852FFCC45763530A98DED1A" box="[569,698,1087,1110]" name="Madagascar" pageId="22" pageNumber="23">Madagascar</collectingCountry>
(Maps 4, 15).
</paragraph>
</subSubSection>
</treatment>
</document>

417
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<mods:title id="1D35ECA60EE0AC339D97806F7F4D825F">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<emphasis id="41F73E371855FFCB473433F2AA5EEA58" bold="true" box="[123,361,765,788]" italics="true" pageId="17" pageNumber="18">Uduba andriamihajai</emphasis>
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,
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<paragraph id="733CE2251855FFCB47343212AAD6EA7E" blockId="17.[123,507,765,818]" box="[123,481,797,818]" pageId="17" pageNumber="18">
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; Maps 6, 14.
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</subSubSection>
<subSubSection id="3B99B1AE1855FFCB47E4324CAA76EA96" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="733CE2251855FFCB47E4324CAA76EA96" blockId="17.[122,1174,835,1725]" pageId="17" pageNumber="18">
<emphasis id="41F73E371855FFCB47E4324CAA78EA16" bold="true" box="[171,335,835,858]" pageId="17" pageNumber="18">Type material.</emphasis>
<materialsCitation id="C3EBE8781855FFCB4618324CAA76EA96" collectingDate="2000-03-13" collectionCode="CAS" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="1670" latitude="-20.384445" location="Korikory" longLatPrecision="21" longitude="47.667225" pageId="17" pageNumber="18" specimenCode="CASENT9010142, CASENT9064716, CASENT9006000, CASENT9064717" specimenCount="10" specimenCount-male="10" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871855FFCB4618324CAA8DEA16" box="[343,442,835,858]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1855FFCB454B324CA9FBEA16" box="[516,716,835,858]" collectionCode="CASENT" pageId="17" pageNumber="18">CASENT9010142</specimenCode>
)
<collectingMethod id="AAC29A321855FFCB4594324CA8AFEA16" box="[731,920,835,858]" pageId="17" pageNumber="18">from malaise trap</collectingMethod>
in forest at
<quantity id="B47B4FC01855FFCB4353324CAF52EA16" box="[1052,1125,835,858]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="17" pageNumber="18" unit="m" value="1670.0">
<elevation id="F8AE05161855FFCB4353324CAF52EA16" box="[1052,1125,835,858]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="17" pageNumber="18" unit="m" value="1670.0">1670m</elevation>
</quantity>
elevation at
<location id="765CB4FE1855FFCB47A9326CAA7FEA36" LSID="urn:lsid:plazi:treatment:FB2A53331855FFC8473433F2AA3AEF58:765CB4FE1855FFCB47A9326CAA7FEA36" box="[230,328,867,890]" country="Madagascar" latitude="-20.384445" longLatPrecision="21" longitude="47.667225" name="Korikory" pageId="17" pageNumber="18" stateProvince="Fianarantsoa">Korikory</location>
(
<geoCoordinate id="16B784E21855FFCB4614326CAAE1EA36" box="[347,470,867,890]" degrees="20" direction="south" minutes="23" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="04" value="-20.384445">20°23ʹ04ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21855FFCB46A8326CA953EA36" box="[487,612,867,890]" degrees="47" direction="east" minutes="40" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="02" value="47.667225">47°40ʹ02ʺE</geoCoordinate>
),
<collectingRegion id="B1472CC71855FFCB4533326CA846EA36" box="[636,881,867,890]" country="Madagascar" name="Fianarantsoa" pageId="17" pageNumber="18">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51855FFCB4430326CA8C9EA36" box="[895,1022,867,890]" name="Madagascar" pageId="17" pageNumber="18">Madagascar</collectingCountry>
, collected
<date id="073DC4E51855FFCB4334326CAA35EAD6" pageId="17" pageNumber="18" value="2000-03-13">
<collectingDate id="17793D0D1855FFCB4334326CAA35EAD6" pageId="17" pageNumber="18" value="2000-03-13">13 March 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31855FFCB465F328CAAA5EAD6" box="[272,402,899,922]" pageId="17" pageNumber="18">M. E. Irwin</collectorName>
and
<collectorName id="DE7687F31855FFCB4683328CA95CEAD6" box="[460,619,899,922]" pageId="17" pageNumber="18">E. I. Schlinger</collectorName>
(SI-001), deposited in
<collectionCode id="15927AE01855FFCB4421328CA892EAD6" box="[878,933,899,922]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="18" type="Museum">CAS</collectionCode>
.
<specimenCount id="658529AC1855FFCB44FD328CAFA5EAD6" box="[946,1170,899,922]" count="5" pageId="17" pageNumber="18" type="male" typeStatus="paratype">Five paratype males</specimenCount>
, same data, (
<specimenCode id="23254A5E1855FFCB464632ACAAE4EAF6" box="[265,467,931,954]" collectionCode="CASENT" pageId="17" pageNumber="18">CASENT9064716</specimenCode>
,
<specimenCount id="658529AC1855FFCB46AB32ACA92DEAF6" box="[484,538,931,954]" count="3" pageId="17" pageNumber="18" type="male">3♂♂</specimenCount>
) (
<specimenCode id="23254A5E1855FFCB457732ACA835EAF6" box="[568,770,931,954]" collectionCode="CASENT" pageId="17" pageNumber="18">CASENT9006000</specimenCode>
,
<specimenCount id="658529AC1855FFCB445B32ACA802EAF6" box="[788,821,931,954]" count="1" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
) and (
<specimenCode id="23254A5E1855FFCB44C632ACAF65EAF6" box="[905,1106,931,954]" collectionCode="CASENT" pageId="17" pageNumber="18">CASENT9064717</specimenCode>
,
<specimenCount id="658529AC1855FFCB432A32ACAFB1EAF6" box="[1125,1158,931,954]" count="1" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), deposited in
<collectionCode id="15927AE01855FFCB464932CCAA0AEA96" box="[262,317,963,986]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="18" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1855FFCB47E432EBA98EED17" pageId="17" pageNumber="18" type="etymology">
<paragraph id="733CE2251855FFCB47E432EBA98EED17" blockId="17.[122,1174,835,1725]" pageId="17" pageNumber="18">
<emphasis id="41F73E371855FFCB47E432EBAA1DEAB7" bold="true" box="[171,298,996,1019]" pageId="17" pageNumber="18">Etymology.</emphasis>
The species epithet is a patronym in honor of Dr. Benjamin Andriamihaja, biochemist and soil scientist, director of the Institute for the Conservation of Tropical Environments (ICTE), ardent conservationist and facilitator of science in
<collectingCountry id="0B94A2B51855FFCB45B7352BA841ED77" box="[760,886,1060,1083]" name="Madagascar" pageId="17" pageNumber="18">Madagascar</collectingCountry>
. His continuing assistance to our scientific research is gratefully acknowledged.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1855FFCB47E4356BA8F3EDB7" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="733CE2251855FFCB47E4356BA8F3EDB7" blockId="17.[122,1174,835,1725]" pageId="17" pageNumber="18">
<emphasis id="41F73E371855FFCB47E4356BA921ED37" bold="true" box="[171,534,1124,1147]" pageId="17" pageNumber="18">
Diagnosis.
<taxonomicName id="B48399A61855FFCB4668356BA921ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,534,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andriamihajai">
<emphasis id="41F73E371855FFCB4668356BA921ED37" bold="true" box="[295,534,1124,1147]" italics="true" pageId="17" pageNumber="18">Uduba andriamihajai</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.b, the
<taxonomicName id="B48399A61855FFCB4423356BAF39ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[876,1038,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="woodae">Uduba woodae</taxonomicName>
group (Map 14) of Group I, the Epigynal atrium group. Specimens are ecribellate, with a colulus. The male palp has the TA2 highly sclerotized, black, concave medially against TA3 and forming a chela around TA3 apex (
<figureCitation id="EBB8FEA01855FFCB47BD35CBAA9FED97" box="[242,424,1220,1243]" captionStart="FIGURE" captionStartId="130.[155,167,827,846]" captionTargetBox="[137,1167,231,802]" captionTargetId="figure-11@130.[120,1176,216,814]" captionTargetPageId="130" captionText="FIGURE 30. Uduba andriamihajai, new species, Holotype (CASENT9010142), Korikory, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158633" httpUri="https://zenodo.org/record/13158633/files/figure.png" pageId="17" pageNumber="18">Figs. 30 A, B, C</figureCitation>
). Males of
<taxonomicName id="B48399A61855FFCB456A35CBA823ED97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[549,788,1220,1243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andriamihajai">
<emphasis id="41F73E371855FFCB456A35CBA823ED97" bold="true" box="[549,788,1220,1243]" italics="true" pageId="17" pageNumber="18">Uduba andriamihajai</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="B48399A61855FFCB473435EBAA0CEDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,315,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371855FFCB473435EBAA0CEDB7" bold="true" box="[123,315,1252,1275]" italics="true" pageId="17" pageNumber="18">Uduba schlingeri</emphasis>
</taxonomicName>
in that the latter has a divided cribellum. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1855FFC847E4340AAA64ECD8" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="733CE2251855FFC847E4340AABDCEB7F" blockId="17.[122,1174,835,1725]" lastBlockId="18.[123,1175,220,1556]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="41F73E371855FFCB47E4340AAA01EC50" bold="true" box="[171,310,1285,1308]" pageId="17" pageNumber="18">Description.</emphasis>
Male (
<typeStatus id="AC385C871855FFCB46CE340AAADCEC50" box="[385,491,1285,1308]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
): Total length 10.00. Markings typical for
<taxonomicName id="B48399A61855FFCB44E1340AA8CFEC50" authorityName="Simon" authorityYear="1880" box="[942,1016,1285,1308]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371855FFCB44E1340AA8CFEC50" bold="true" box="[942,1016,1285,1308]" italics="true" pageId="17" pageNumber="18">Uduba</emphasis>
</taxonomicName>
, as in
<figureCitation id="EBB8FEA01855FFCB4371340AAFA6EC50" box="[1086,1169,1285,1308]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: A–E = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="17" pageNumber="18">Fig. 6B</figureCitation>
. Carapace 5.60 long, 4.30 wide, 2.10 high; clypeus 0.24 high. Eye diameters: AME and ALE 0.20, PME and PLE 0.24. Chelicerae 2.60 long; sternum 2.60 long, 1.80 wide; labium 1.20 long; palpal coxae 1.90 long. With colulus. Spination (
<typeStatus id="AC385C871855FFCB4579346AA9A1EC30" box="[566,662,1381,1404]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, CASENT9064717): palpus – femur d0-1-2; leg I – femur d1-1-1, p0-0-2, r0-0-1, patella p1, tibia d0-0-1-1, p0-1-1-0, v1-2-2-2, r0-1-1-0, metatarsus p1-1-1, v1-1-1-1, r0-1-1-0; leg II – femur d1-1-1, p0-1-1-0, r0-1-1-0, patella p1, tibia d0-1-1- 0, p0-1-1-0, v1-0-0-1, r0-1-1-0, metatarsus p0-1-1-0, v2-1-1-1, r0-1-1-0; leg III – femur d1-1-1, p0- 1-1-1, r0-1-1-0, patella r1, tibia d0-1-0, p0-1-1-0, v2-0-2, r0-1-1-0, metatarsus p1-2-2, v2-1-2, r1- 1-2, tarsus v0-1-0; leg IV – femur d1-1-1, p0-0-1, r0-0-1, tibia d0-1-0, p0-1-1-0, v2-1-3, r0-1-1-0, metatarsus p0-1-2, v2-2-2-2, r1-1-1-2. Scopulae: cymbium, apicodorsal; beneath all tarsi. Leg measurements (
<typeStatus id="AC385C871855FFCB466B3749AABFEF11" box="[292,392,1606,1629]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
): I: 5.50 + 2.10 + 5.80 + 6.00 + 3.10 = 22.50; II: 5.00 + 2.00 + 4.90 + 5.40 + 2.80 = 20.10; III: 4.40 + 1.70 + 2.90 + 4.20 + 2.20 = 15.40; IV: 5.70 + 1.70 + 5.60 + 6.80 + 2.90 = 22.70; palpus: 2.20 + 1.00 + 1.10 + NA + 2.60 = 6.90. Leg formula 4123.
<emphasis id="41F73E371855FFCB43503789AFA3EFD1" bold="true" box="[1055,1172,1670,1693]" pageId="17" pageNumber="18">Male palp</emphasis>
(
<typeStatus id="AC385C871855FFCB47CE37A9ABD5EFF1" box="[129,226,1702,1725]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
) (
<figureCitation id="EBB8FEA01855FFCB47B637A9AA9BEFF1" box="[249,428,1702,1725]" captionStart="FIGURE" captionStartId="130.[155,167,827,846]" captionTargetBox="[137,1167,231,802]" captionTargetId="figure-11@130.[120,1176,216,814]" captionTargetPageId="130" captionText="FIGURE 30. Uduba andriamihajai, new species, Holotype (CASENT9010142), Korikory, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158633" httpUri="https://zenodo.org/record/13158633/files/figure.png" pageId="17" pageNumber="18">Figs. 30 A, B, C</figureCitation>
): palpal tibia 0.42 times cymbial length, RTA broadly triangular (
<figureCitation id="EBB8FEA01855FFC8432537A9AB87E9BF" captionStart="FIGURE" captionStartId="130.[155,167,827,846]" captionTargetBox="[137,1167,231,802]" captionTargetId="figure-11@130.[120,1176,216,814]" captionTargetPageId="130" captionText="FIGURE 30. Uduba andriamihajai, new species, Holotype (CASENT9010142), Korikory, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158633" httpUri="https://zenodo.org/record/13158633/files/figure.png" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Fig. 30 C</figureCitation>
), RTA length 0.30 tibia length, length equals width, with pointed apex, VTA length 1.67 times width, length 0.405 times tibia width, VTA cylindrical, straight, with rounded apex; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.91 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.54 times tegulum length; TA3 transverse, tapering to a point, concave dorsally, TA3 apex straight, visible in front of TA2 (
<figureCitation id="EBB8FEA01856FFC844F53053AF64E83F" box="[954,1107,348,371]" captionStart="FIGURE" captionStartId="130.[155,167,827,846]" captionTargetBox="[137,1167,231,802]" captionTargetId="figure-11@130.[120,1176,216,814]" captionTargetPageId="130" captionText="FIGURE 30. Uduba andriamihajai, new species, Holotype (CASENT9010142), Korikory, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158633" httpUri="https://zenodo.org/record/13158633/files/figure.png" pageId="18" pageNumber="19">Figs. 30 B, C</figureCitation>
); TA2 highly sclerotized, black, concave medially against TA3, in this species forming a chela around TA3 apex (
<figureCitation id="EBB8FEA01856FFC847BE3093AA6EE8FF" box="[241,345,412,435]" captionStart="FIGURE" captionStartId="130.[155,167,827,846]" captionTargetBox="[137,1167,231,802]" captionTargetId="figure-11@130.[120,1176,216,814]" captionTargetPageId="130" captionText="FIGURE 30. Uduba andriamihajai, new species, Holotype (CASENT9010142), Korikory, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158633" httpUri="https://zenodo.org/record/13158633/files/figure.png" pageId="18" pageNumber="19">Fig. 30 A</figureCitation>
), base of TA2 with small bump just above MA base; MA transverse, concave in middle with proximal and distal enlargements, origin at 0.31 times tegulum length, tegulum apex at 0.29 tegulum length from MA apex, length (including apical processes) 0.72 times width, MA large, length 0.375 tegulum length, width 0.47 times tegulum width; conductor fan entire.
<emphasis id="41F73E371856FFC8437330F3AFB8EB5F" bold="true" box="[1084,1167,508,531]" pageId="18" pageNumber="19">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="733CE2251856FFC847E43333A908EA1F" blockId="18.[123,1175,220,1556]" pageId="18" pageNumber="19">
<emphasis id="41F73E371856FFC847E43333AA2DEB1F" bold="true" box="[171,282,572,595]" pageId="18" pageNumber="19">Variation.</emphasis>
(
<emphasis id="41F73E371856FFC846643333AA09EB1F" bold="true" box="[299,318,572,595]" pageId="18" pageNumber="19">N</emphasis>
= 6): Total length = 9.50–14.00, carapace length / width = 1.33–1.69, carapace height / width = 0.44–0.57, PER / carapace width = 0.36–0.38, PER/OAL = 2.70–3.27, PER/AER = 1.30–1.40, OAL / OQL = 1.00–1.15, OQP / OQA = 1.11–1.24, clypeus height / AME = 1.14– 1.54, cheliceral length / clypeus height = 8.00–10.80, sternum length / width = 1.24–1.50, palpal coxa length / width = 2.33–2.86, femur I length / carapace width =1.28–1.38, metatarsus I length / carapace width = 1.36–1.49, femur IV length / carapace width =1.31–1.38, cymbium length / carapace width = 0.59–0.71, cymbium length / palpal patella length = 2.60–3.56, cymbium length / palpal tibia length = 2.42–3.20, cymbium length / palpal femur length = 1.06–1.26, palpal tibia length / palpal patella length = 1.10–1.30.
</paragraph>
<paragraph id="733CE2251856FFC847E43253A835ED98" blockId="18.[123,1175,220,1556]" pageId="18" pageNumber="19">
<materialsCitation id="C3EBE8781856FFC847E43253AFA5EA3F" box="[171,1170,860,883]" collectionCode="FMNH" country="Madagascar" location="MADAGASCAR" pageId="18" pageNumber="19" specimenCode="CASENT9006076" specimenCount="2" specimenCount-male="2">
M
<emphasis id="41F73E371856FFC8478D3253AABCEA3F" bold="true" box="[194,395,860,883]" pageId="18" pageNumber="19">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51856FFC846DB3253A961EA3F" box="[404,598,860,883]" name="Madagascar" pageId="18" pageNumber="19">MADAGASCAR</collectingCountry>
: locality unknown (
<specimenCode id="23254A5E1856FFC8447F3253A8CDEA3F" box="[816,1018,860,883]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9006076</specimenCode>
,
<specimenCount id="658529AC1856FFC843483253AF1FEA3F" box="[1031,1064,860,883]" count="2" pageId="18" pageNumber="19" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE01856FFC843763253AFBCEA3F" box="[1081,1163,860,883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="19" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781856FFC847343273AAF0EABF" collectingDate="2000-03-13" collectionCode="CAS" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="1670" latitude="-20.384445" location="Korikory" longLatPrecision="21" longitude="47.667225" pageId="18" pageNumber="19" specimenCode="CASENT9010142, CASENT9064716, CASENT9064717, CASENT9006000" specimenCount="6" specimenCount-male="6" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="41F73E371856FFC847343273AA41EADF" box="[123,374,892,915]" italics="true" pageId="18" pageNumber="19">
<collectingRegion id="B1472CC71856FFC847343273AA41EADF" box="[123,374,892,915]" country="Madagascar" name="Fianarantsoa" pageId="18" pageNumber="19">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1856FFC846C43273AADEEADF" LSID="urn:lsid:plazi:treatment:FB2A53331855FFC8473433F2AA3AEF58:765CB4FE1856FFC846C43273AADEEADF" box="[395,489,892,915]" country="Madagascar" latitude="-20.384445" longLatPrecision="21" longitude="47.667225" name="Korikory" pageId="18" pageNumber="19" stateProvince="Fianarantsoa">Korikory</location>
,
<locationDeviation id="628EFB071856FFC846B33273A980EADF" box="[508,695,892,915]" location="Fandriana" pageId="18" pageNumber="19">SE of Fandriana</locationDeviation>
,
<geoCoordinate id="16B784E21856FFC845873273A875EADF" box="[712,834,892,915]" degrees="20" direction="south" minutes="23" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="04" value="-20.384445">20°23ʹ04ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21856FFC8441B3273A8E6EADF" box="[852,977,892,915]" degrees="47" direction="east" minutes="40" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="02" value="47.667225">47°40ʹ02ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01856FFC843513273AF5DEADF" box="[1054,1130,892,915]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="18" pageNumber="19" unit="m" value="1670.0">
<elevation id="F8AE05161856FFC843513273AF5DEADF" box="[1054,1130,892,915]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="18" pageNumber="19" unit="m" value="1670.0">1670m</elevation>
</quantity>
,
<date id="073DC4E51856FFC843343273AA36EAFF" pageId="18" pageNumber="19" value="2000-03-13">
<collectingDate id="17793D0D1856FFC843343273AA36EAFF" pageId="18" pageNumber="19" value="2000-03-13">13 March 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31856FFC846413293AAB8EAFF" box="[270,399,924,947]" pageId="18" pageNumber="19">M. E. Irwin</collectorName>
and
<collectorName id="DE7687F31856FFC846883293A952EAFF" box="[455,613,924,947]" pageId="18" pageNumber="19">E. I. Schlinger</collectorName>
[SI-001] (
<typeStatus id="AC385C871856FFC845963293A877EAFF" box="[729,832,924,947]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1856FFC844023293AF22EAFF" box="[845,1045,924,947]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9010142</specimenCode>
,
<specimenCount id="658529AC1856FFC8436E3293AF74EAFF" box="[1057,1091,924,947]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01856FFC8431D3293AFBEEAFF" box="[1106,1161,924,947]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="19" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C871856FFC847CD32B3ABD9EA9F" box="[130,238,956,979]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
,
<specimenCode id="23254A5E1856FFC847B132B3AAF1EA9F" box="[254,454,956,979]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9064716</specimenCode>
,
<specimenCount id="658529AC1856FFC8469A32B3AAC1EA9F" box="[469,502,956,979]" count="3" pageId="18" pageNumber="19" type="male">3♂</specimenCount>
,
<collectionCode id="15927AE01856FFC8454732B3A908EA9F" box="[520,575,956,979]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="19" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C871856FFC8451132B3A9F7EA9F" box="[606,704,956,979]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1856FFC8458032B3A8AFEA9F" box="[719,920,956,979]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9064717</specimenCode>
,
<specimenCount id="658529AC1856FFC844E932B3A8FFEA9F" box="[934,968,956,979]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01856FFC8449532B3AF26EA9F" box="[986,1041,956,979]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="19" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C871856FFC8437F32B3AFA5EA9F" box="[1072,1170,956,979]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1856FFC8473432D3AA74EABF" box="[123,323,988,1011]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9006000</specimenCode>
,
<specimenCount id="658529AC1856FFC8461C32D3AA43EABF" box="[339,372,988,1011]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01856FFC846C832D3AA88EABF" box="[391,447,988,1011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="19" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781856FFC8469832D3A996ED7F" collectingDate="1997-03-05" collectingDateMax="1997-03-11" collectingDateMin="1997-03-05" collectionCode="FMNH" collectorName="S. E. Goodman" country="Madagascar" county="Andringitra" elevation="1990" location="Philippia" municipality="Anjavidilay" pageId="18" pageNumber="19" specimenCode="CASENT9064648" specimenCount="12" specimenCount-male="12" stateProvince="Fianarantsoa" typeStatus="holotype">
RNI
<collectingCounty id="9A5D9AA91856FFC8454132D3A9BAEABF" box="[526,653,988,1011]" pageId="18" pageNumber="19">Andringitra</collectingCounty>
,
<collectingMunicipality id="9358785F1856FFC845D532D3A823EABF" box="[666,788,988,1011]" pageId="18" pageNumber="19">Anjavidilay</collectingMunicipality>
,
<locationDeviation id="628EFB071856FFC8446832D3AF24EABF" box="[807,1043,988,1011]" location="Atanifotsy" pageId="18" pageNumber="19">
<quantity id="B47B4FC01856FFC8446832D3A845EABF" box="[807,882,988,1011]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="19" unit="km" value="8.5">8.5 km</quantity>
SE Atanifotsy
</locationDeviation>
, 22°09.5ʹS, 46°57.6ʹE, elev.
<quantity id="B47B4FC01856FFC8466732F3AA42ED5F" box="[296,373,1020,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="18" pageNumber="19" unit="m" value="1990.0">
<elevation id="F8AE05161856FFC8466732F3AA42ED5F" box="[296,373,1020,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="18" pageNumber="19" unit="m" value="1990.0">1990m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321856FFC846CF32F3AAD8ED5F" box="[384,495,1020,1043]" pageId="18" pageNumber="19">pitfall trap</collectingMethod>
in
<emphasis id="41F73E371856FFC8455E32F3A943ED5F" box="[529,628,1020,1043]" italics="true" pageId="18" pageNumber="19">
<location id="765CB4FE1856FFC8455E32F3A943ED5F" LSID="urn:lsid:plazi:treatment:FB2A53331855FFC8473433F2AA3AEF58:765CB4FE1856FFC8455E32F3A943ED5F" box="[529,628,1020,1043]" country="Madagascar" county="Andringitra" municipality="Anjavidilay" name="Philippia" pageId="18" pageNumber="19" stateProvince="Fianarantsoa">Philippia</location>
</emphasis>
-dominated sclerophyllus forest,
<date id="073DC4E51856FFC8449C32F3AFA6ED5F" box="[979,1169,1020,1043]" pageId="18" pageNumber="19" value="1997-03-05" valueMax="1997-03-11" valueMin="1997-03-05">
<collectingDate id="17793D0D1856FFC8449C32F3AFA6ED5F" box="[979,1169,1020,1043]" pageId="18" pageNumber="19" value="1997-03-05" valueMax="1997-03-11" valueMin="1997-03-05">5–11 March 1997</collectingDate>
</date>
,
<collectorName id="DE7687F31856FFC847343513AA15ED7F" box="[123,290,1052,1075]" pageId="18" pageNumber="19">S. E. Goodman</collectorName>
(
<specimenCode id="23254A5E1856FFC8467C3513AACBED7F" box="[307,508,1052,1075]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9064648</specimenCode>
,
<specimenCount id="658529AC1856FFC845453513A90FED7F" box="[522,568,1052,1075]" count="12" pageId="18" pageNumber="19" type="male">12♂</specimenCount>
,
<collectionCode id="15927AE01856FFC845073513A9ADED7F" box="[584,666,1052,1075]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="19" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781856FFC845E13513ABE3EDD8" collectingDate="1997-10-24" collectingDateMax="1997-10-30" collectingDateMin="1997-10-24" collectionCode="FMNH" collectorName="Goodman" country="Madagascar" county="Ivohibe" elevation="1575" location="Camp" municipality="Andranomainty River" pageId="18" pageNumber="19" specimenCode="CASENT9006077" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
R. S. de
<collectingCounty id="9A5D9AA91856FFC8445E3513A852ED7F" box="[785,869,1052,1075]" pageId="18" pageNumber="19">Ivohibe</collectingCounty>
, at source of
<collectingMunicipality id="9358785F1856FFC844B53513ABE5ED1F" pageId="18" pageNumber="19">Andranomainty River</collectingMunicipality>
,
<locationDeviation id="628EFB071856FFC847AA3533AAFAED1F" box="[229,461,1084,1107]" location="Ivohibe" pageId="18" pageNumber="19">
<quantity id="B47B4FC01856FFC847AA3533AA05ED1F" box="[229,306,1084,1107]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="19" unit="km" value="6.5">6.5 km</quantity>
ESE Ivohibe
</locationDeviation>
, S22.496°, E46.955°,
<location id="765CB4FE1856FFC845873533A83FED1F" LSID="urn:lsid:plazi:treatment:FB2A53331855FFC8473433F2AA3AEF58:765CB4FE1856FFC845873533A83FED1F" box="[712,776,1084,1107]" country="Madagascar" county="Ivohibe" municipality="Andranomainty River" name="Camp" pageId="18" pageNumber="19" stateProvince="Fianarantsoa">Camp</location>
3, elev.
<quantity id="B47B4FC01856FFC844203533A88BED1F" box="[879,956,1084,1107]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="18" pageNumber="19" unit="m" value="1575.0">
<elevation id="F8AE05161856FFC844203533A88BED1F" box="[879,956,1084,1107]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="18" pageNumber="19" unit="m" value="1575.0">1575m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321856FFC844843533AF74ED1F" box="[971,1091,1084,1107]" pageId="18" pageNumber="19">pitfall trap</collectingMethod>
, undisturbed mossy forest,
<date id="073DC4E51856FFC8462A3552A97EED38" box="[357,585,1117,1140]" pageId="18" pageNumber="19" value="1997-10-24" valueMax="1997-10-30" valueMin="1997-10-24">
<collectingDate id="17793D0D1856FFC8462A3552A97EED38" box="[357,585,1117,1140]" pageId="18" pageNumber="19" value="1997-10-24" valueMax="1997-10-30" valueMin="1997-10-24">24–30 October 1997</collectingDate>
</date>
, S. E.
<collectorName id="DE7687F31856FFC845D73552A836ED38" box="[664,769,1117,1140]" pageId="18" pageNumber="19">Goodman</collectorName>
[SEG1997] (
<specimenCode id="23254A5E1856FFC844D93552AF68ED38" box="[918,1119,1117,1140]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9006077</specimenCode>
,
<specimenCount id="658529AC1856FFC843223552AFB8ED38" box="[1133,1167,1117,1140]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01856FFC847343572ABFAEDD8" box="[123,205,1149,1172]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="19" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781856FFC847903572A9C9ED98" collectingDate="2008-05-25" collectingDateMax="2008-06-01" collectingDateMin="2008-05-25" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="550" latitude="-17.916111" location="Betampona" longLatPrecision="21" longitude="49.201946" municipality="Reserve Nationale Integrale Betampona" pageId="18" pageNumber="19" specimenCode="CASENT9042359" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="41F73E371856FFC847903572AA81EDD8" box="[223,438,1149,1172]" italics="true" pageId="18" pageNumber="19">
<collectingRegion id="B1472CC71856FFC847903572AA81EDD8" box="[223,438,1149,1172]" country="Madagascar" name="Toamasina" pageId="18" pageNumber="19">Toamasina Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1856FFC8468A3572A842EDD8" box="[453,885,1149,1172]" pageId="18" pageNumber="19">Réserve Nationale Intégrale Betampona</collectingMunicipality>
,
<location id="765CB4FE1856FFC844CE3572A8CAEDD8" LSID="urn:lsid:plazi:treatment:FB2A53331855FFC8473433F2AA3AEF58:765CB4FE1856FFC844CE3572A8CAEDD8" box="[897,1021,1149,1172]" country="Madagascar" latitude="-17.916111" longLatPrecision="21" longitude="49.201946" municipality="Reserve Nationale Integrale Betampona" name="Betampona" pageId="18" pageNumber="19" stateProvince="Toamasina">Betampona</location>
,
<locationDeviation id="628EFB071856FFC843463572ABD8EDF8" location="Toamasina" pageId="18" pageNumber="19">
<quantity id="B47B4FC01856FFC843463572AF56EDD8" box="[1033,1121,1149,1172]" metricMagnitude="4" metricUnit="m" metricValue="3.51" pageId="18" pageNumber="19" unit="km" value="35.1">35.1 km</quantity>
NW
<collectingRegion id="B1472CC71856FFC847343592ABD8EDF8" box="[123,239,1181,1204]" country="Madagascar" name="Toamasina" pageId="18" pageNumber="19">Toamasina</collectingRegion>
</locationDeviation>
,
<geoCoordinate id="16B784E21856FFC847B43592AA42EDF8" box="[251,373,1181,1204]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="58" value="-17.916111">17°54ʹ58ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21856FFC846CB3592A939EDF8" box="[388,526,1181,1204]" degrees="049" direction="east" minutes="12" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="07" value="49.201946">049°12ʹ07ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01856FFC8451C3592A9A5EDF8" box="[595,658,1181,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="m" value="550.0">
<elevation id="F8AE05161856FFC8451C3592A9A5EDF8" box="[595,658,1181,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="m" value="550.0">550m</elevation>
</quantity>
, rainforest,
<collectingMethod id="AAC29A321856FFC8445A3592A8ACEDF8" box="[789,923,1181,1204]" pageId="18" pageNumber="19">malaise trap</collectingMethod>
,
<date id="073DC4E51856FFC844E73592AFA6EDF8" box="[936,1169,1181,1204]" pageId="18" pageNumber="19" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">
<collectingDate id="17793D0D1856FFC844E73592AFA6EDF8" box="[936,1169,1181,1204]" pageId="18" pageNumber="19" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">25 May–1 June 2008</collectingDate>
</date>
,
<collectorName id="DE7687F31856FFC8473435B2ABC3ED98" box="[123,244,1213,1236]" pageId="18" pageNumber="19">B.L. Fisher</collectorName>
[BLF19593_23] (
<specimenCode id="23254A5E1856FFC846F635B2A9B5ED98" box="[441,642,1213,1236]" collectionCode="CASENT" pageId="18" pageNumber="19">CASENT9042359</specimenCode>
,
<specimenCount id="658529AC1856FFC845C135B2A998ED98" box="[654,687,1213,1236]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01856FFC845F135B2A9C1ED98" box="[702,758,1213,1236]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="19" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251856FFC847E435D2AA64ECD8" blockId="18.[123,1175,220,1556]" pageId="18" pageNumber="19">
<emphasis id="41F73E371856FFC847E435D2AA69EDB8" bold="true" box="[171,350,1245,1268]" pageId="18" pageNumber="19">Natural history.</emphasis>
The ecribellate species
<taxonomicName id="B48399A61856FFC8451335D2A87DEDB8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[604,842,1245,1268]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="andriamihajai">
<emphasis id="41F73E371856FFC8451335D2A87DEDB8" bold="true" box="[604,842,1245,1268]" italics="true" pageId="18" pageNumber="19">Uduba andriamihajai</emphasis>
</taxonomicName>
is known only from males collected in pitfall and malaise traps, suggesting that these spiders wander on the ground and up into vegetation when mature. Nothing is known of their use of silk. Most records are from high elevations: from
<quantity id="B47B4FC01856FFC847B53432AA00EC18" box="[250,311,1341,1364]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="m" value="550.0">550m</quantity>
in rainforest to
<quantity id="B47B4FC01856FFC846A83432A906EC18" box="[487,561,1341,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="18" pageNumber="19" unit="m" value="1575.0">1575m</quantity>
and
<quantity id="B47B4FC01856FFC845273432A986EC18" box="[616,689,1341,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="18" pageNumber="19" unit="m" value="1670.0">1670m</quantity>
in rainforest and mossy forest, even extending to high-elevation
<emphasis id="41F73E371856FFC846293452AAFEEC38" box="[358,457,1373,1396]" italics="true" pageId="18" pageNumber="19">Philippia</emphasis>
-dominated sclerophyllus forest at
<quantity id="B47B4FC01856FFC8440B3452A8B9EC38" box="[836,910,1373,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="18" pageNumber="19" unit="m" value="1990.0">1990m</quantity>
at Andringitra in southeastern
<collectingCountry id="0B94A2B51856FFC847813472AA7BECD8" box="[206,332,1405,1428]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1856FFC847E43492AA3AEF58" pageId="18" pageNumber="19" type="distribution">
<paragraph id="733CE2251856FFC847E43492AA3AEF58" blockId="18.[123,1175,220,1556]" pageId="18" pageNumber="19">
<emphasis id="41F73E371856FFC847E43492AA0AECF8" bold="true" box="[171,317,1437,1460]" pageId="18" pageNumber="19">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61856FFC846863492A98FECF8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[457,696,1437,1460]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="andriamihajai">
<emphasis id="41F73E371856FFC846863492A98FECF8" bold="true" box="[457,696,1437,1460]" italics="true" pageId="18" pageNumber="19">Uduba andriamihajai</emphasis>
</taxonomicName>
is recorded from rainforest along the mountain spine of central-east
<collectingCountry id="0B94A2B51856FFC846DD34B2A924EC98" box="[402,531,1469,1492]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
and even from high elevation sclerophyll vegetation. One record, from Reserve Nationale Intégrale Betampona, is from an isolated remnant patch of forest (Maps 6, 14).
</paragraph>
</subSubSection>
</treatment>
</document>

465
data/FB/2A/53/FB2A53331856FFCF47343732A89FEB9B.xml Normal file
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<document id="3B71D61D00C524CF2E25EBA1D21F66B2" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331856FFCF47343732A89FEB9B" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba balsama Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="9B356CEA-6775-4D6A-B98D-135CC90954F3" docUuidSource="ZooBank" docVersion="2" lastPageNumber="22" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="19" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="69696F9CACABA4ADC48407F7DE154EFE">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="AFA13C800E39B0B6D081920956D0FA92">Griswold, Charles</mods:namePart>
<mods:affiliation id="BD680D5A4954F3BDC84C2953A67BB0A4">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="5E96821B805CCDE3D43A902CE75E1DD9">Ubick, Darrell</mods:namePart>
<mods:affiliation id="B2927FC0D9ADCA36D236CCC6623A83CC">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
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<mods:namePart id="F7250104169B481549CDEF991FB4BA57">Ledford, Joel</mods:namePart>
<mods:affiliation id="0CA17822D9FF126C508121C7243DD2BB">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="D0626E52393B021409F835A3A8CC0F63">Polotow, Daniele</mods:namePart>
<mods:affiliation id="76E87DB3A2A26E5C51D7997BEDD20306">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:title id="46317AD62794AF386803177E82430B09">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="2BFBAE6B81EC03F9A4D8E190AC3608EF">2022</mods:date>
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<heading id="287455491856FFC847343732AA8DEF18" box="[123,442,1597,1620]" fontSize="10" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="41F73E371856FFC847343732AA8DEF18" bold="true" box="[123,442,1597,1620]" pageId="18" pageNumber="19">
<taxonomicName id="B48399A61856FFC847343732AA1FEF18" ID-CoL="C8KJ4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,296,1597,1620]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="balsama" status="sp. nov.">
<emphasis id="41F73E371856FFC847343732AA1FEF18" bold="true" box="[123,296,1597,1620]" italics="true" pageId="18" pageNumber="19">Uduba balsama</emphasis>
</taxonomicName>
,
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</paragraph>
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<subSubSection id="3B99B1AE1856FFC847343752A97EEF3E" box="[123,585,1629,1650]" pageId="18" pageNumber="19" type="description">
<paragraph id="733CE2251856FFC847343752A97EEF3E" blockId="18.[123,585,1597,1650]" box="[123,585,1629,1650]" pageId="18" pageNumber="19">
<figureCitation id="EBB8FEA01856FFC847343752AA21EF3E" box="[123,278,1629,1650]" captionStart="FIGURE" captionStartId="130.[155,167,1462,1481]" captionTargetBox="[131,1165,933,1433]" captionTargetId="figure-46@130.[120,1176,924,1448]" captionTargetPageId="130" captionText="FIGURE 31. Uduba balsama, new species, Holotype (CASENT9030929), left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158635" httpUri="https://zenodo.org/record/13158635/files/figure.png" pageId="18" pageNumber="19">Figures 31 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01856FFC8466F3752AA5EEF3E" box="[288,361,1629,1650]" captionStart="FIGURE" captionStartId="131.[155,167,1216,1235]" captionTargetBox="[120,1176,216,1205]" captionTargetId="figure-16@131.[120,1176,216,1205]" captionTargetPageId="131" captionText="FIGURE 32. Uduba balsama, new species, (CASENT9000543), Kirindy, right male palpus, SEM. A. Tibia and tarsus, retrolateral; arrow to cymbial scopula. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral: note how embolus arises retrobasally, disappears behind cymbium, and apex appears distally from behind tegulum; arrow to cymbial scopula. D. Bulb, retrolateral. E. Bulb, retroventral: note how embolus apex extends from behind tegulum. F. Bulb, prolateral: note concave apical margin of TA3. C, conductor; Cm, cymbium; E, embolus; EB, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bars A = 200 µm, B, D, F = 100 µm, C = 300 µm, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158638" httpUri="https://zenodo.org/record/13158638/files/figure.png" pageId="18" pageNumber="19">32 A–F</figureCitation>
,
<figureCitation id="EBB8FEA01856FFC8463C3752AAF7EF3E" box="[371,448,1629,1650]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="18" pageNumber="19">74 D–E</figureCitation>
,
<figureCitation id="EBB8FEA01856FFC846853752A928EF3E" box="[458,543,1629,1650]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="18" pageNumber="19">Maps 11</figureCitation>
, 14.
</paragraph>
</subSubSection>
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<paragraph id="733CE2251856FFC947E43789A90EE81F" blockId="18.[123,1174,1670,1725]" lastBlockId="19.[123,1174,220,1725]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="41F73E371856FFC847E43789AA78EFD1" bold="true" box="[171,335,1670,1693]" pageId="18" pageNumber="19">Type material.</emphasis>
<materialsCitation id="C3EBE8781856FFC946193789AB85E85F" collectingDate="2002-04-11" collectingDateMax="2002-04-16" collectingDateMin="2002-04-11" collectionCode="CAS" collectorName="M. E. Irwin &amp; F. D. Parker &amp; R. Harin'Hala" country="Madagascar" elevation="825" lastPageId="19" lastPageNumber="20" location="Zombitse National Park" pageId="18" pageNumber="19" specimenCode="CASENT9030929" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C871856FFC846193789AA8EEFD1" box="[342,441,1670,1693]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
male (
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)
<collectingMethod id="AAC29A321856FFC845963789A8A1EFD1" box="[729,918,1670,1693]" pageId="18" pageNumber="19">from malaise trap</collectingMethod>
in deciduous spiny forest at
<quantity id="B47B4FC01856FFC847F737A9ABCCEFF1" box="[184,251,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="18" pageNumber="19" unit="m" value="825.0">
<elevation id="F8AE05161856FFC847F737A9ABCCEFF1" box="[184,251,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="18" pageNumber="19" unit="m" value="825.0">825 m</elevation>
</quantity>
elevation in
<location id="765CB4FE1856FFC846CB37A9A9B2EFF1" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1856FFC846CB37A9A9B2EFF1" box="[388,645,1702,1725]" country="Madagascar" name="Zombitse National Park" pageId="18" pageNumber="19" stateProvince="Toliara">Zombitse National Park</location>
, 22°50.43ʹS, 44°43.87ʹE,
<collectingRegion id="B1472CC71856FFC844DA37A9AF71EFF1" box="[917,1094,1702,1725]" country="Madagascar" name="Toliara" pageId="18" pageNumber="19">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51856FFC9431F37A9AB8AE9BF" lastPageId="19" lastPageNumber="20" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
, collected
<date id="073DC4E51857FFC9467A31D3AAC5E9BF" box="[309,498,220,243]" pageId="19" pageNumber="20" value="2002-04-11" valueMax="2002-04-16" valueMin="2002-04-11">
<collectingDate id="17793D0D1857FFC9467A31D3AAC5E9BF" box="[309,498,220,243]" pageId="19" pageNumber="20" value="2002-04-11" valueMax="2002-04-16" valueMin="2002-04-11">11–16 April 2002</collectingDate>
</date>
by
<collectorName id="DE7687F31857FFC9455131D3A996E9BF" box="[542,673,220,243]" pageId="19" pageNumber="20">M. E. Irwin</collectorName>
,
<collectorName id="DE7687F31857FFC945E031D3A806E9BF" box="[687,817,220,243]" pageId="19" pageNumber="20">F. D. Parker</collectorName>
and
<collectorName id="DE7687F31857FFC9442631D3AF32E9BF" box="[873,1029,220,243]" pageId="19" pageNumber="20">R. Harin’Hala</collectorName>
, deposited in
<collectionCode id="15927AE01857FFC9473431F3AB85E85F" box="[123,178,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="20" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781857FFC947F231F3A902E81F" collectingDate="2001-11-28" collectingDateMax="2001-12-03" collectingDateMin="2001-11-28" collectionCode="CAS" collectorName="de Kirindy" country="Madagascar" elevation="100" latitude="-20.045" location="Foret de Kirindy" longLatPrecision="21" longitude="44.662224" pageId="19" pageNumber="20" specimenCode="CASENT9013532" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="AC385C871857FFC947F231F3AA2DE85F" box="[189,282,252,275]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1857FFC9463531F3A975E85F" box="[378,578,252,275]" collectionCode="CASENT" pageId="19" pageNumber="20">CASENT9013532</specimenCode>
) from tropical dry forest at
<quantity id="B47B4FC01857FFC9443E31F3A899E85F" box="[881,942,252,275]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="100.0">
<elevation id="F8AE05161857FFC9443E31F3A899E85F" box="[881,942,252,275]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="100.0">100m</elevation>
</quantity>
elevation in
<location id="765CB4FE1857FFC9437331F3ABFDE87F" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1857FFC9437331F3ABFDE87F" country="Madagascar" latitude="-20.045" longLatPrecision="21" longitude="44.662224" name="Foret de Kirindy" pageId="19" pageNumber="20" stateProvince="Toliara">
Forêt
<collectorName id="DE7687F31857FFC9433331F3ABFDE87F" pageId="19" pageNumber="20">de Kirindy</collectorName>
</location>
,
<geoCoordinate id="16B784E21857FFC947963013AA71E87F" box="[217,326,284,307]" degrees="20" direction="south" minutes="2" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="42" value="-20.045">20°2ʹ42ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21857FFC9461B3013AAE6E87F" box="[340,465,284,307]" degrees="44" direction="east" minutes="39" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="44" value="44.662224">44°39ʹ44ʺE</geoCoordinate>
, collected
<date id="073DC4E51857FFC945093013A8F7E87F" box="[582,960,284,307]" pageId="19" pageNumber="20" value="2001-11-28" valueMax="2001-12-03" valueMin="2001-11-28">
<collectingDate id="17793D0D1857FFC945093013A8F7E87F" box="[582,960,284,307]" pageId="19" pageNumber="20" value="2001-11-28" valueMax="2001-12-03" valueMin="2001-11-28">28 November to 3 December 2001</collectingDate>
</date>
by the Fisher-Griswold Arthropod Team, deposited in
<collectionCode id="15927AE01857FFC946B13033A902E81F" box="[510,565,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="20" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1857FFC947E43053AA7BEB59" pageId="19" pageNumber="20" type="etymology">
<paragraph id="733CE2251857FFC947E43053AA7BEB59" blockId="19.[123,1174,220,1725]" pageId="19" pageNumber="20">
<emphasis id="41F73E371857FFC947E43053AA1DE83F" bold="true" box="[171,298,348,371]" pageId="19" pageNumber="20">Etymology.</emphasis>
The species epithet honors Mrs.
<taxonomicName id="B48399A61857FFC945DE3053A85DE83F" authority="Rajemison" authorityName="Rajemison" box="[657,874,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="balsama">Balsama Rajemison</taxonomicName>
, curator of the collection of Entomology at the Parc Botanique et Zoologique de Tsimbazaza (PBZT), Lakeside International Scholar and member of the Fisher-Griswold arthropod team at the California Academy of Sciences. She is gratefully acknowledged for her hospitality in her home country and for her essential efforts in securing permissions and permits for research and collecting in
<collectingCountry id="0B94A2B51857FFC9440730D2A8F1E8B8" box="[840,966,477,500]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
. Name treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1857FFC947E43311AAE0EBBA" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="733CE2251857FFC947E43311AAE0EBBA" blockId="19.[123,1174,220,1725]" pageId="19" pageNumber="20">
<emphasis id="41F73E371857FFC947E43311AAE4EB79" bold="true" box="[171,467,542,565]" pageId="19" pageNumber="20">
Diagnosis.
<taxonomicName id="B48399A61857FFC946693311AAE4EB79" box="[294,467,542,565]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="balsama">
<emphasis id="41F73E371857FFC946693311AAE4EB79" bold="true" box="[294,467,542,565]" italics="true" pageId="19" pageNumber="20">Uduba balsama</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.b, the
<taxonomicName id="B48399A61857FFC944693311A8F0EB79" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[806,967,542,565]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="woodae">Uduba woodae</taxonomicName>
group (Map 14) of Group I, the Epigynal atrium group. They are ecribellate
<taxonomicName id="B48399A61857FFC945A03331A80EEB19" authorityName="Simon" authorityYear="1880" box="[751,825,574,597]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371857FFC945A03331A80EEB19" bold="true" box="[751,825,574,597]" italics="true" pageId="19" pageNumber="20">Uduba</emphasis>
</taxonomicName>
, with a colulus; male palp with TA2 highly sclerotized, black, concave medially against TA3 but not forming a chela around TA3 apex, TA3 reflexed (
<figureCitation id="EBB8FEA01857FFC946283371A931EBD9" box="[359,518,638,661]" captionStart="FIGURE" captionStartId="130.[155,167,1462,1481]" captionTargetBox="[131,1165,933,1433]" captionTargetId="figure-46@130.[120,1176,924,1448]" captionTargetPageId="130" captionText="FIGURE 31. Uduba balsama, new species, Holotype (CASENT9030929), left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158635" httpUri="https://zenodo.org/record/13158635/files/figure.png" pageId="19" pageNumber="20">Figs. 31 A–C</figureCitation>
); female epigynal plate with median lobe surrounded by depressed atrium, plate extensive posteriad of atrium, plate height greater than 0.15 times epigynal length, median lobe broad, bell-shaped, atrium nearly twice ML width, ML and atrium apices nearly at same level (
<figureCitation id="EBB8FEA01857FFC9467A33D0AAFCEBBA" box="[309,459,735,758]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="19" pageNumber="20">Figs. 74 D–E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1857FFCF47E433F0AA72EBDF" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
<paragraph id="733CE2251857FFCE47E433F0AA97EAFB" blockId="19.[123,1174,220,1725]" lastBlockId="20.[122,1175,220,1725]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="41F73E371857FFC947E433F0AA4DEA5A" bold="true" box="[171,378,767,790]" pageId="19" pageNumber="20">Description. Male</emphasis>
(
<typeStatus id="AC385C871857FFC946C233F0AAC0EA5A" box="[397,503,767,790]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
): Total length 10.50. Markings typical for
<taxonomicName id="B48399A61857FFC9449933F0AF17EA5A" authorityName="Simon" authorityYear="1880" box="[982,1056,767,790]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371857FFC9449933F0AF17EA5A" bold="true" box="[982,1056,767,790]" italics="true" pageId="19" pageNumber="20">Uduba</emphasis>
</taxonomicName>
. Carapace 5.60 long, 4.25 wide, 2.40 high; clypeus 0.30 high. Eye diameters: AME 0.20, ALE and PLE 0.22, PME 0.20. Chelicerae 2.50 long; sternum 2.50 long, 2.00 wide; labium 1.20 long; palpal coxae 1.80 long. With a colulus. Spination: Palpus – femur d0-1-1, p0-0-1, r0-0-1; leg I – femur d1-1-1, p0-1-2, r0-1-0-1, patella p1, r1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2- 2-1, r1-1-2; leg II – femur d1-1-1, p0-1-1, r0-1-1, patella p1, r1, tibia d0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-1, r1-1-2; leg III – femur d1-1-1, p0-0-1-1, r0-1-1-1, patella p1, r1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v1-2-2-2, r1-1-2; leg IV – femur d1-1-0-1, p0-0-0-1, r0-0-0-1, patella p1, r1, tibia d0-1-1-0, p0-1-0-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v1-1-1-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; beneath tarsi and apices of all metatarsi. Leg measurements: I: 5.10 + 2.10 + 5.10 + 5.30 + 3.40 = 21.00; II: 4.60 + 2.00 + 4.10 + 4.80 + 2.90 = 18.40; III: 4.00 + 1.50 + 2.70 + 4.30 + 2.30 = 14.80; IV: 5.20 + 1.80 + 4.80 + 6.80 + 3.00 = 21.60; palpus: 2.10 + 0.80 + 1.00 + NA + 3.10 = 7.00. Leg formula 4123.
<emphasis id="41F73E371857FFC944AF358DAF65EDD5" bold="true" box="[992,1106,1154,1177]" pageId="19" pageNumber="20">Male palp</emphasis>
(
<figureCitation id="EBB8FEA01857FFC9432F358DABE3EDF5" captionStart="FIGURE" captionStartId="130.[155,167,1462,1481]" captionTargetBox="[131,1165,933,1433]" captionTargetId="figure-46@130.[120,1176,924,1448]" captionTargetPageId="130" captionText="FIGURE 31. Uduba balsama, new species, Holotype (CASENT9030929), left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158635" httpUri="https://zenodo.org/record/13158635/files/figure.png" pageId="19" pageNumber="20">Figs. 31 A–C</figureCitation>
): palpal tibia 0.33 times cymbial length, RTA broadly triangular, RTA length 0.375 tibia length, length 1.33 times width, curved ventrally to pointed apex; VTA curved, with rounded apex (
<figureCitation id="EBB8FEA01857FFC947CC35ECAA2CEDB6" box="[131,283,1251,1274]" captionStart="FIGURE" captionStartId="130.[155,167,1462,1481]" captionTargetBox="[131,1165,933,1433]" captionTargetId="figure-46@130.[120,1176,924,1448]" captionTargetPageId="130" captionText="FIGURE 31. Uduba balsama, new species, Holotype (CASENT9030929), left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158635" httpUri="https://zenodo.org/record/13158635/files/figure.png" pageId="19" pageNumber="20">Figs. 31 A, C</figureCitation>
), VTA length 2.28 times width, length 0.30 times tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 0.94 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.36 times tegulum length; TA3 transverse, concave dorsally, tapering to a blunt, upturned point, TA3 apex visible beside TA2; TA2 highly sclerotized, black, concave medially against TA3 (
<figureCitation id="EBB8FEA01857FFC94555346BA9B3EC37" box="[538,644,1380,1403]" captionStart="FIGURE" captionStartId="131.[155,167,1216,1235]" captionTargetBox="[120,1176,216,1205]" captionTargetId="figure-16@131.[120,1176,216,1205]" captionTargetPageId="131" captionText="FIGURE 32. Uduba balsama, new species, (CASENT9000543), Kirindy, right male palpus, SEM. A. Tibia and tarsus, retrolateral; arrow to cymbial scopula. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral: note how embolus arises retrobasally, disappears behind cymbium, and apex appears distally from behind tegulum; arrow to cymbial scopula. D. Bulb, retrolateral. E. Bulb, retroventral: note how embolus apex extends from behind tegulum. F. Bulb, prolateral: note concave apical margin of TA3. C, conductor; Cm, cymbium; E, embolus; EB, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bars A = 200 µm, B, D, F = 100 µm, C = 300 µm, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158638" httpUri="https://zenodo.org/record/13158638/files/figure.png" pageId="19" pageNumber="20">Fig. 32 E</figureCitation>
); MA transverse, concave, simple, origin at 0.51 times tegulum length, tegulum apex at 0.31 tegulum length from MA apex, length (including apical processes) 0.36 times width, MA small, length 0.18 tegulum length, width 0.46 times tegulum width; conductor fan entire (
<figureCitation id="EBB8FEA01857FFC946E034CBA977EC97" box="[431,576,1476,1499]" captionStart="FIGURE" captionStartId="131.[155,167,1216,1235]" captionTargetBox="[120,1176,216,1205]" captionTargetId="figure-16@131.[120,1176,216,1205]" captionTargetPageId="131" captionText="FIGURE 32. Uduba balsama, new species, (CASENT9000543), Kirindy, right male palpus, SEM. A. Tibia and tarsus, retrolateral; arrow to cymbial scopula. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral: note how embolus arises retrobasally, disappears behind cymbium, and apex appears distally from behind tegulum; arrow to cymbial scopula. D. Bulb, retrolateral. E. Bulb, retroventral: note how embolus apex extends from behind tegulum. F. Bulb, prolateral: note concave apical margin of TA3. C, conductor; Cm, cymbium; E, embolus; EB, embolic base; MA, median apophysis; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bars A = 200 µm, B, D, F = 100 µm, C = 300 µm, E = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158638" httpUri="https://zenodo.org/record/13158638/files/figure.png" pageId="19" pageNumber="20">Figs. 32 A–F</figureCitation>
).
<emphasis id="41F73E371857FFC9451C34CBA991EC97" bold="true" box="[595,678,1476,1499]" pageId="19" pageNumber="20">Female</emphasis>
(
<typeStatus id="AC385C871857FFC945FA34CBA82EEC97" box="[693,793,1476,1499]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
): Total length 14.20. Markings typical for
<taxonomicName id="B48399A61857FFC9478134EAAA2FECB0" authorityName="Simon" authorityYear="1880" box="[206,280,1509,1532]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371857FFC9478134EAAA2FECB0" bold="true" box="[206,280,1509,1532]" italics="true" pageId="19" pageNumber="20">Uduba</emphasis>
</taxonomicName>
. Carapace 7.40 long, 4.50 wide, 2.70 high; clypeus 0.40 high. Eye diameters: AME 0.20, ALE and PME 0.22, PLE 0.26. Chelicerae 3.30 long; sternum 2.90 long, 2.40 wide; labium 1.60 long; palpal coxae 2.20 long.
<emphasis id="41F73E371857FFC946BF372AA90EEF70" bold="true" box="[496,569,1573,1596]" pageId="19" pageNumber="20">Ratios</emphasis>
: carapace length / width = 1.64, carapace height / width = 0.60, PER / carapace width = 0.45, PER / OAL= 3.00, PER / AER = 1.36, OAL / OQL = 1.10, OQP / OQA = 1.07, clypeus height/ AME diameter width = 2.00, cheliceral length / clypeus height = 8.25, sternum length / width = 1.21, palpal coxa length / width = 2.00, femur I length / carapace width = 0.98, metatarsus I length / carapace width = 0.69, femur IV length / carapace width = 1.07, palpal tarsus length / carapace width = 0.47, palpal tibia length / palpal patella length = 1.91, palpal tarsus length / palpal tibia length =1.75, palpal tarsus length / palpal femur length = 0.95, palpal tibia length / palpal patella length = 1.09. With a colulus. Spination: palpus – femur d0-0-2, patella p1, tibia p0-1-0, tarsus p2-1, v0-0-1; leg I – femur d1-0-1, p0-0-1, tibia v2-2-2-2, metatarsus v2-2-3; leg II – femur d1-0-0, p0-0-1, tibia p0-0-1, v1-1-1-2, metatarsus v2-2-3; leg III – femur d1-0-0-1, p0-0-1-2, r0-0-0-1, tibia d1-1-0-0, p0-1-1-0, v1-1-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-2-2, tarsus v0-1-0; leg IV – femur d1-1-1, r0-0-1, tibia p1-0-0, v1-1-2, r0-1-1-0, metatarsus d0- 1-0-0, p1-1-1-2, v1-1-2-2, r1-1-1-2. Scopulae: strong ventral beneath all tarsi, beneath metatarsus I and apical half of metatarsus II and apices of metatarsi III and IV. Leg measurements: I: 4.40 + 2.30 + 3.60 + 3.10 + 2.10 = 15.50; II: 4.00 + 2.10 + 2.90 + 2.60 + 1.90 = 13.50; III: 3.10 + 1.90 + 2.00 + 2.70 + 1.60 = 11.30; IV: 4.80 + 1.90 + 3.70 + 4.70 + 2.20 = 17.30; palpus: 2.20 + 1.10 + 1.20 + NA + 2.10 = 6.60. Leg formula 4123.
<emphasis id="41F73E371850FFCE45353331A800EB19" bold="true" box="[634,823,574,597]" pageId="20" pageNumber="21">Female genitalia</emphasis>
: epigynal plate (
<figureCitation id="EBB8FEA01850FFCE44A23331AF61EB19" box="[1005,1110,574,597]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="20" pageNumber="21">Fig. 74 D</figureCitation>
) with median lobe surrounded by depressed atrium, plate width 1.125 times length; atrium sides nearly straight, atrium width 2.08 times atrium length; epigynum length 3.33 times atrium length, atrium width 0.56 times epigynum width; atrium width at side of ML 0.50 times ML width; atrium and median lobe arise well anteriad of epigastric groove, atrium origin at 0.425 of epigynal plate length, median lobe triangular, broadest at base, median lobe length 1.08 times width, ML apex near atrium apex, ML length 0.35 times epigynal length; lateral lobes narrow, atrium width 3.125 times LL width; copulatory openings hidden beneath posterolateral margin of atrium, distance between CO 0.55 times width of epigynal plate, CO distance from epigastric furrow 0.425 times epigynum length. Vulva (
<figureCitation id="EBB8FEA01850FFCE4650326FAABAEA3B" box="[287,397,864,887]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="20" pageNumber="21">Fig. 74 E</figureCitation>
) with spermathecal ducts making four tight, transverse curves, vulva length 0.73 times width, fertilization ducts widely separated, vulva width 3.67 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251850FFCE47E432CEA985EDB6" blockId="20.[122,1175,220,1725]" pageId="20" pageNumber="21">
<emphasis id="41F73E371850FFCE47E432CEAA69EA94" bold="true" box="[171,350,961,984]" pageId="20" pageNumber="21">Variation. Male</emphasis>
(N = 3): Total length = 10.50–12.00, carapace length / width = 1.31–1.45, carapace height / width = 0.46–0.56, PER / carapace width = 0.36–0.39, PER / OAL = 2.70–3.11, PER / AER = 1.33–1.36, OAL / OQL = 1.00–1.04, OQP / OQA = 1.08–1.19, clypeus height / AME diameter width = 0.83–1.36, cheliceral length / clypeus height = 8.33–15.00, sternum length / width = 1.18–1.26, palpal coxa length / width = 2.38–3.00, femur I length / carapace width = 1.20–1.26, metatarsus I length / carapace width = 1.25–1.31, femur IV length / carapace width = 1.22–1.36, cymbium length / carapace width = 0.63–0.73, cymbium length / palpal patella length = 3.00–3.88, cymbium length / palpal tibia length = 3.00–3.38, cymbium length / palpal femur length = 0.94– 1.48, palpal tibia length / palpal patella length = 1.00–1.25.
<emphasis id="41F73E371850FFCE445335CCA858ED96" bold="true" box="[796,879,1219,1242]" pageId="20" pageNumber="21">Female</emphasis>
variation is unknown: the female is known from the single
<typeStatus id="AC385C871850FFCE46AD35ECA909EDB6" box="[482,574,1251,1274]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
specimen.
</paragraph>
<paragraph id="733CE2251850FFCF47E4340CAAFCE81F" blockId="20.[122,1175,220,1725]" lastBlockId="21.[123,1174,220,727]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<materialsCitation id="C3EBE8781850FFCF47E4340CAAFFE81F" collectingDate="1996-10-04" collectingDateMax="2010-12-01" collectingDateMin="1996-10-04" collectionCode="FMNH, FMNH-INS, CAS" collectorName="R. N. de Marojejy &amp; S. Goodman &amp; Parc Nacional Bemaraha, S &amp; S. M. Goodman &amp; M. E. Irwin &amp; F. D. Parker &amp; R. Harin'Hala &amp; de Kirindy &amp; B. L. Fisher" country="Madagascar" county="Manantenina River" elevation="775" lastPageId="21" lastPageNumber="22" latitude="-21.218334" location="Manantenina River" longLatPrecision="21" longitude="45.310555" municipality="Camp" pageId="20" pageNumber="21" specimenCode="CASENT9064659, CASENT9064654, FMNH2001-91, CASENT9030929, CASENT9000543, CASENT9013532, CASENT9042528" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Antsiranana" typeStatus="holotype">
M
<emphasis id="41F73E371850FFCE478D340CAAB9EC56" bold="true" box="[194,398,1283,1306]" pageId="20" pageNumber="21">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51850FFCE46D5340CA96BEC56" box="[410,604,1283,1306]" name="Madagascar" pageId="20" pageNumber="21">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371850FFCE4525340CA86FEC56" box="[618,856,1283,1306]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="B1472CC71850FFCE4525340CA86FEC56" box="[618,856,1283,1306]" country="Madagascar" name="Antsiranana" pageId="20" pageNumber="21">Antsiranana Province</collectingRegion>
</emphasis>
:
<collectorName id="DE7687F31850FFCE4425340CAF0CEC56" box="[874,1083,1283,1306]" pageId="20" pageNumber="21">R. N. de Marojejy</collectorName>
,
<locationDeviation id="628EFB071850FFCE4301340CAA77EC76" location="Manantenina" pageId="20" pageNumber="21">
<quantity id="B47B4FC01850FFCE4301340CAFA2EC56" box="[1102,1173,1283,1306]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="10.0">10 km</quantity>
NW Manantenina
</locationDeviation>
, along tributary of
<collectingCounty id="9A5D9AA91850FFCE456F342CA9C7EC76" box="[544,752,1315,1338]" pageId="20" pageNumber="21">Manantenina River</collectingCounty>
,
<collectingMunicipality id="9358785F1850FFCE444C342CA874EC76" box="[771,835,1315,1338]" pageId="20" pageNumber="21">Camp</collectingMunicipality>
2, 14°26.0ʹS, 49°44.7ʹE, elev.
<quantity id="B47B4FC01850FFCE4734344CAB8DEC16" box="[123,186,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="20" pageNumber="21" unit="m" value="775.0">
<elevation id="F8AE05161850FFCE4734344CAB8DEC16" box="[123,186,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="20" pageNumber="21" unit="m" value="775.0">775m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321850FFCE478A344CAA32EC16" box="[197,261,1347,1370]" pageId="20" pageNumber="21">pitfall</collectingMethod>
in relatively undisturbed lowland-montane rainforest,
<date id="073DC4E51850FFCE4403344CAF1DEC16" box="[844,1066,1347,1370]" pageId="20" pageNumber="21" value="1996-10-14" valueMax="1996-10-23" valueMin="1996-10-14">
<collectingDate id="17793D0D1850FFCE4403344CAF1DEC16" box="[844,1066,1347,1370]" pageId="20" pageNumber="21" value="1996-10-14" valueMax="1996-10-23" valueMin="1996-10-14">14–23 October 1996</collectingDate>
</date>
,
<collectorName id="DE7687F31850FFCE437A344CAB9BEC37" pageId="20" pageNumber="21">S. Goodman</collectorName>
, (
<specimenCode id="23254A5E1850FFCE47F2346BAAB1EC37" box="[189,390,1380,1403]" collectionCode="CASENT" pageId="20" pageNumber="21">CASENT9064659</specimenCode>
,
<specimenCount id="658529AC1850FFCE46DE346BAA84EC37" box="[401,435,1380,1403]" count="2" pageId="20" pageNumber="21" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE01850FFCE468F346BA926EC37" box="[448,529,1380,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21" type="Museum">FMNH</collectionCode>
),
<collectorName id="DE7687F31850FFCE456B346BA9D6EC37" box="[548,737,1380,1403]" pageId="20" pageNumber="21">R. N. de Marojejy</collectorName>
,
<locationDeviation id="628EFB071850FFCE45A0346BA8DAEC37" box="[751,1005,1380,1403]" location="Manantenina" pageId="20" pageNumber="21">
<quantity id="B47B4FC01850FFCE45A0346BA811EC37" box="[751,806,1380,1403]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="21" unit="km" value="8.0">8 km</quantity>
NW Manantenina
</locationDeviation>
, along tributary of
<location id="765CB4FE1850FFCE47D7348BAA53ECD7" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE47D7348BAA53ECD7" box="[152,356,1412,1435]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Manantenina River" pageId="20" pageNumber="21" stateProvince="Antsiranana">Manantenina River</location>
,
<location id="765CB4FE1850FFCE463C348BAA84ECD7" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE463C348BAA84ECD7" box="[371,435,1412,1435]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Camp" pageId="20" pageNumber="21" stateProvince="Antsiranana">Camp</location>
1, 14°26.2ʹS, 49°46.5ʹE, elev.
<quantity id="B47B4FC01850FFCE45B3348BA80BECD7" box="[764,828,1412,1435]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="21" unit="m" value="450.0">
<elevation id="F8AE05161850FFCE45B3348BA80BECD7" box="[764,828,1412,1435]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="21" unit="m" value="450.0">450m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321850FFCE4408348BA8B0ECD7" box="[839,903,1412,1435]" pageId="20" pageNumber="21">pitfall</collectingMethod>
in relatively undisturbed lowland rainforest,
<date id="073DC4E51850FFCE460234ABA917ECF7" box="[333,544,1444,1467]" pageId="20" pageNumber="21" value="1996-10-04" valueMax="1996-10-13" valueMin="1996-10-04">
<collectingDate id="17793D0D1850FFCE460234ABA917ECF7" box="[333,544,1444,1467]" pageId="20" pageNumber="21" value="1996-10-04" valueMax="1996-10-13" valueMin="1996-10-04">4–13 October 1996</collectingDate>
</date>
,
<collectorName id="DE7687F31850FFCE456334ABA984ECF7" box="[556,691,1444,1467]" pageId="20" pageNumber="21">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1850FFCE458D34ABA8BBECF7" box="[706,908,1444,1467]" collectionCode="CASENT" pageId="20" pageNumber="21">CASENT9064654</specimenCode>
,
<specimenCount id="658529AC1850FFCE44D734ABA88EECF7" box="[920,953,1444,1467]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01850FFCE448734ABAF2DECF7" box="[968,1050,1444,1467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21" type="Museum">FMNH</collectionCode>
).
<emphasis id="41F73E371850FFCE436334ABABCAEC97" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="B1472CC71850FFCE436334ABABCAEC97" country="Madagascar" name="Mahajanga" pageId="20" pageNumber="21">Mahajanga Province</collectingRegion>
</emphasis>
:
<collectorName id="DE7687F31850FFCE464134CBA90AEC97" box="[270,573,1476,1499]" pageId="20" pageNumber="21">Parc Nacional Bemaraha, S</collectorName>
bank
<location id="765CB4FE1850FFCE45CA34CBA87EEC97" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE45CA34CBA87EEC97" box="[645,841,1476,1499]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Manambolo River" pageId="20" pageNumber="21" stateProvince="Antsiranana">Manambolo River</location>
,
<location id="765CB4FE1850FFCE441634CBAF16EC97" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE441634CBAF16EC97" box="[857,1057,1476,1499]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Tombeau Vazimba" pageId="20" pageNumber="21" stateProvince="Antsiranana">Tombeau Vazimba</location>
,
<locationDeviation id="628EFB071850FFCE436034CBABDFECB0" location="Bekopaka" pageId="20" pageNumber="21">
<quantity id="B47B4FC01850FFCE436034CBAF4CEC97" box="[1071,1147,1476,1499]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="21" unit="km" value="3.5">3.5 km</quantity>
E Bekopaka
</locationDeviation>
,
<geoCoordinate id="16B784E21850FFCE47BC34EAAA57ECB0" box="[243,352,1509,1532]" degrees="19" direction="south" minutes="8" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="24" value="-19.14">19°8ʹ24ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21850FFCE462234EAAAEAECB0" box="[365,477,1509,1532]" degrees="44" direction="east" minutes="49" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="2" value="44.817223">44°49ʹ2ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01850FFCE456F34EAA968ECB0" box="[544,607,1509,1532]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="100.0">
<elevation id="F8AE05161850FFCE456F34EAA968ECB0" box="[544,607,1509,1532]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="100.0">100m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321850FFCE452434EAA9DFECB0" box="[619,744,1509,1532]" pageId="20" pageNumber="21">pitfall traps</collectingMethod>
,
<date id="073DC4E51850FFCE45BB34EAAF61ECB0" box="[756,1110,1509,1532]" pageId="20" pageNumber="21" value="2001-11-30" valueMax="2001-12-06" valueMin="2001-11-30">
<collectingDate id="17793D0D1850FFCE45BB34EAAF61ECB0" box="[756,1110,1509,1532]" pageId="20" pageNumber="21" value="2001-11-30" valueMax="2001-12-06" valueMin="2001-11-30">30 November–6 December 2001</collectingDate>
</date>
,
<collectorName id="DE7687F31850FFCE432E34EAABD0EF50" pageId="20" pageNumber="21">S.M. Goodman</collectorName>
, (
<specimenCode id="23254A5E1850FFCE47B7370AAA96EF50" box="[248,417,1541,1564]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21" type="Museum">FMNH2001-91</specimenCode>
) (
<collectionCode id="15927AE01850FFCE46FA370AA90FEF50" box="[437,568,1541,1564]" pageId="20" pageNumber="21">FMNH-INS</collectionCode>
0000 044 665,
<specimenCount id="658529AC1850FFCE4592370AA9C9EF50" box="[733,766,1541,1564]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01850FFCE4444370AA86AEF50" box="[779,861,1541,1564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21" type="Museum">FMNH</collectionCode>
).
<emphasis id="41F73E371850FFCE4421370AAF2AEF50" box="[878,1053,1541,1564]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="B1472CC71850FFCE4421370AAF2AEF50" box="[878,1053,1541,1564]" country="Madagascar" name="Toliara" pageId="20" pageNumber="21">Toliara Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1850FFCE437E370AAA22EF70" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE437E370AAA22EF70" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Zombitse National Park" pageId="20" pageNumber="21" stateProvince="Antsiranana">Zombitse National Park</location>
, near national road, 22°50.43ʹS, 44°43.87ʹE, elev.
<quantity id="B47B4FC01850FFCE440A372AA8B3EF70" box="[837,900,1573,1596]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="20" pageNumber="21" unit="m" value="825.0">
<elevation id="F8AE05161850FFCE440A372AA8B3EF70" box="[837,900,1573,1596]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="20" pageNumber="21" unit="m" value="825.0">825m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321850FFCE44DC372AAF2CEF70" box="[915,1051,1573,1596]" pageId="20" pageNumber="21">malaise trap</collectingMethod>
, deciduous spiny forest,
<date id="073DC4E51850FFCE4642374AAAE7EF10" box="[269,464,1605,1628]" pageId="20" pageNumber="21" value="2002-04-11" valueMax="2002-04-16" valueMin="2002-04-11">
<collectingDate id="17793D0D1850FFCE4642374AAAE7EF10" box="[269,464,1605,1628]" pageId="20" pageNumber="21" value="2002-04-11" valueMax="2002-04-16" valueMin="2002-04-11">11–16 April 2002</collectingDate>
</date>
,
<collectorName id="DE7687F31850FFCE4690374AA951EF10" box="[479,614,1605,1628]" pageId="20" pageNumber="21">M. E. Irwin</collectorName>
,
<collectorName id="DE7687F31850FFCE453A374AA9CCEF10" box="[629,763,1605,1628]" pageId="20" pageNumber="21">F. D. Parker</collectorName>
and
<collectorName id="DE7687F31850FFCE4478374AA8E5EF10" box="[823,978,1605,1628]" pageId="20" pageNumber="21">R. Harin’Hala</collectorName>
[MA-02-13B-24] (
<typeStatus id="AC385C871850FFCE47CD3769ABDEEF31" box="[130,233,1638,1661]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1850FFCE47B33769AAF3EF31" box="[252,452,1638,1661]" collectionCode="CASENT" pageId="20" pageNumber="21">CASENT9030929</specimenCode>
,
<specimenCount id="658529AC1850FFCE469A3769AAC0EF31" box="[469,503,1638,1661]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01850FFCE45443769A974EF31" box="[523,579,1638,1661]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1850FFCE45113769A82DEF31" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1850FFCE45113769A82DEF31" box="[606,794,1638,1661]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Foret de Kirindy" pageId="20" pageNumber="21" stateProvince="Antsiranana">
Forêt
<collectorName id="DE7687F31850FFCE45EB3769A82DEF31" box="[676,794,1638,1661]" pageId="20" pageNumber="21">de Kirindy</collectorName>
</location>
,
<locationDeviation id="628EFB071850FFCE44603769AFA6EF31" box="[815,1169,1638,1661]" location="Marofandilia" pageId="20" pageNumber="21">
<quantity id="B47B4FC01850FFCE44603769A8B9EF31" box="[815,910,1638,1661]" metricMagnitude="4" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="21" unit="km" value="15.5">15.5 km</quantity>
64°ENE Marofandilia
</locationDeviation>
,
<geoCoordinate id="16B784E21850FFCE47343789ABDFEFD1" box="[123,232,1670,1693]" degrees="20" direction="south" minutes="2" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="42" value="-20.045">20°2ʹ42ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21850FFCE47B73789AA42EFD1" box="[248,373,1670,1693]" degrees="44" direction="east" minutes="39" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="44" value="44.662224">44°39ʹ44ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01850FFCE46F03789AAC9EFD1" box="[447,510,1670,1693]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="100.0">
<elevation id="F8AE05161850FFCE46F03789AAC9EFD1" box="[447,510,1670,1693]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="100.0">100m</elevation>
</quantity>
, tropical dry forest,
<collectingMethod id="AAC29A321850FFCE45A93789A86AEFD1" box="[742,861,1670,1693]" pageId="20" pageNumber="21">pitfall trap</collectingMethod>
,
<date id="073DC4E51850FFCE44233789AB84EFF1" pageId="20" pageNumber="21" value="2001-11-28" valueMax="2001-12-03" valueMin="2001-11-28">
<collectingDate id="17793D0D1850FFCE44233789AB84EFF1" pageId="20" pageNumber="21" value="2001-11-28" valueMax="2001-12-03" valueMin="2001-11-28">28 November–3 December 2001</collectingDate>
</date>
, Fisher-Griswold Arthropod Team [BLF4600] (
<specimenCode id="23254A5E1850FFCE45F437A9A8B2EFF1" box="[699,901,1702,1725]" collectionCode="CASENT" pageId="20" pageNumber="21">CASENT9000543</specimenCode>
,
<specimenCount id="658529AC1850FFCE44DC37A9A883EFF1" box="[915,948,1702,1725]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01850FFCE448A37A9A8CBEFF1" box="[965,1020,1702,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
), general collecting ground spiders, Fisher-Griswold Arthropod Team [BLF4682] (
<typeStatus id="AC385C871851FFCF441131D3A8F7E9BF" box="[862,960,220,243]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1851FFCF448531D3AFA5E9BF" box="[970,1170,220,243]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9013532</specimenCode>
,
<specimenCount id="658529AC1851FFCF473431F3ABABE85F" box="[123,156,252,275]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01851FFCF47E431F3ABD4E85F" box="[171,227,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="22" type="Museum">CAS</collectionCode>
);
<location id="765CB4FE1851FFCF47B831F3AA4BE85F" LSID="urn:lsid:plazi:treatment:FB2A53331856FFCF47343732A89FEB9B:765CB4FE1851FFCF47B831F3AA4BE85F" box="[247,380,252,275]" country="Madagascar" county="Manantenina River" latitude="-21.218334" longLatPrecision="21" longitude="45.310555" municipality="Camp" name="Makay Mts." pageId="21" pageNumber="22" stateProvince="Antsiranana">Makay Mts.</location>
,
<geoCoordinate id="16B784E21851FFCF46C631F3A934E85F" box="[393,515,252,275]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="21" pageNumber="22" precision="15" seconds="06" value="-21.218334">21°13ʹ06ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21851FFCF455E31F3A9ACE85F" box="[529,667,252,275]" degrees="045" direction="east" minutes="18" orientation="longitude" pageId="21" pageNumber="22" precision="15" seconds="38" value="45.310555">045°18ʹ38ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01851FFCF459031F3A817E85F" box="[735,800,252,275]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="21" pageNumber="22" unit="m" value="510.0">
<elevation id="F8AE05161851FFCF459031F3A817E85F" box="[735,800,252,275]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="21" pageNumber="22" unit="m" value="510.0">510m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321851FFCF446131F3A8D6E85F" box="[814,993,252,275]" pageId="21" pageNumber="22">yellow pan traps</collectingMethod>
in gallery forest on sandy soil,
<date id="073DC4E51851FFCF46083013A9EEE87F" box="[327,729,284,307]" pageId="21" pageNumber="22" value="2010-11-24" valueMax="2010-12-01" valueMin="2010-11-24">
<collectingDate id="17793D0D1851FFCF46083013A9EEE87F" box="[327,729,284,307]" pageId="21" pageNumber="22" value="2010-11-24" valueMax="2010-12-01" valueMin="2010-11-24">24 November–1 December 2010</collectingDate>
</date>
,
<collectorName id="DE7687F31851FFCF45BB3013A8A2E87F" box="[756,917,284,307]" pageId="21" pageNumber="22">B. L. Fisher</collectorName>
<emphasis id="41F73E371851FFCF44E33013A8CEE87F" box="[940,1017,284,307]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
[BLF25203] (
<specimenCode id="23254A5E1851FFCF47CD3033AA7CE81F" box="[130,331,316,339]" collectionCode="CASENT" pageId="21" pageNumber="22">CASENT9042528</specimenCode>
,
<specimenCount id="658529AC1851FFCF46183033AA4EE81F" box="[343,377,316,339]" count="1" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01851FFCF46C83033AA88E81F" box="[391,447,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="22" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251851FFCF47E43053AA72EBDF" blockId="21.[123,1174,220,727]" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E43053AA56E83F" bold="true" box="[171,353,348,371]" pageId="21" pageNumber="22">Natural history.</emphasis>
The species
<taxonomicName id="B48399A61851FFCF46BE3053A997E83F" box="[497,672,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="balsama">
<emphasis id="41F73E371851FFCF46BE3053A997E83F" bold="true" box="[497,672,348,371]" italics="true" pageId="21" pageNumber="22">Uduba balsama</emphasis>
</taxonomicName>
is unusual among
<taxonomicName id="B48399A61851FFCF443E3053A88DE83F" authorityName="Simon" authorityYear="1880" box="[881,954,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371851FFCF443E3053A88DE83F" box="[881,954,348,371]" italics="true" pageId="21" pageNumber="22">Uduba</emphasis>
</taxonomicName>
in its occurrence in some of the drier areas of
<collectingCountry id="0B94A2B51851FFCF46EF3073A929E8DF" box="[416,542,380,403]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
. Individuals have been collected in dry habitats in southwestern
<collectingCountry id="0B94A2B51851FFCF479B3093AA64E8FF" box="[212,339,412,435]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
, i.e., from a malaise trap in deciduous spiny forest at
<quantity id="B47B4FC01851FFCF44C23093A8E7E8FF" box="[909,976,412,435]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="21" pageNumber="22" unit="m" value="825.0">825 m</quantity>
elevation in Zombitse National Park and by general collecting (method unspecified) and from pitfall traps from tropical dry forest at
<quantity id="B47B4FC01851FFCF466030D3AA5BE8BF" box="[303,364,476,499]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="100.0">100m</quantity>
elevation in Forêt de Kirindy and from yellow pan traps in gallery forest on sandy soil in the Mackay Mountains. Other possibly conspecific individuals have been collected with pitfall traps in relatively undisturbed lowland rainforest from
<quantity id="B47B4FC01851FFCF442B3313A8E6EB7F" box="[868,977,540,563]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="21" pageNumber="22" unit="m" value="600.0" valueMax="800.0" valueMin="400.0">400-800m</quantity>
near Marojejy in northeastern
<collectingCountry id="0B94A2B51851FFCF464A3333AAB4EB1F" box="[261,387,572,595]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
. Data suggest that at least mature males wander on the ground and up into vegetation, where they are collected by malaise traps. We know nothing of the use of silk by these ecribellate spiders.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1851FFCF47E43391A89FEB9B" pageId="21" pageNumber="22" type="distribution">
<paragraph id="733CE2251851FFCF47E43391A89FEB9B" blockId="21.[123,1174,220,727]" pageId="21" pageNumber="22">
<emphasis id="41F73E371851FFCF47E43391AAC7EBF9" bold="true" box="[171,496,670,693]" pageId="21" pageNumber="22">
Distribution.
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<emphasis id="41F73E371851FFCF460C3391AAC7EBF9" bold="true" box="[323,496,670,693]" italics="true" pageId="21" pageNumber="22">Uduba balsama</emphasis>
</taxonomicName>
</emphasis>
are widely distributed in the drier parts of western
<collectingCountry id="0B94A2B51851FFCF435A3391AFA1EBF9" box="[1045,1174,670,693]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
with at least one outlier from wet rainforest in the northeast (
<figureCitation id="EBB8FEA01851FFCF445E33CFA846EB9B" box="[785,881,704,727]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="21" pageNumber="22">Maps 11</figureCitation>
, 14).
</paragraph>
</subSubSection>
</treatment>
</document>

367
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<mods:title id="013CAC6A625D1B7918D35BA6B18C485C">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="FCC9C23EC2D94B0934A7568FE27FD01B">Griswold, Charles</mods:namePart>
<mods:affiliation id="F982FB569C481FC90B0492B071599DC3">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="8965C63BC6060DA32BAB4118F8ACF4AA">Ubick, Darrell</mods:namePart>
<mods:affiliation id="66B34C68DB61970F3EFC66A68CE1C9FD">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
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<mods:namePart id="7379550E73300BE18055E5F8B754277E">Ledford, Joel</mods:namePart>
<mods:affiliation id="78A4E0C1BFE1ECC3AE5CEAC9B83782C2">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:affiliation id="1D675DC3CAF8D9C9134FE9D2979455F2">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<emphasis id="41F73E37185EFFC0473433F2AA1EEA58" bold="true" box="[123,297,765,788]" italics="true" pageId="26" pageNumber="27">Uduba danielae</emphasis>
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,
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</paragraph>
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<paragraph id="733CE225185EFFC047343212AADBEA7E" blockId="26.[123,492,765,818]" box="[123,492,797,818]" pageId="26" pageNumber="27">
<figureCitation id="EBB8FEA0185EFFC047343212AA3DEA7E" box="[123,266,797,818]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="26" pageNumber="27">Figures 5 A, B</figureCitation>
,
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, Maps 10, 20.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE185EFFC047E4324CAB81ED77" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="733CE225185EFFC047E4324CAB81ED77" blockId="26.[123,1174,835,1725]" pageId="26" pageNumber="27">
<emphasis id="41F73E37185EFFC047E4324CAA67EA16" bold="true" box="[171,336,835,858]" pageId="26" pageNumber="27">Type material.</emphasis>
<materialsCitation id="C3EBE878185EFFC04618324CA8E9EAF6" collectingDate="2001-04-17" collectingDateMax="2001-04-22" collectingDateMin="2001-04-17" collectionCode="CAS" country="Madagascar" county="Fisher-Griswold Arthropod Team" elevation="1620" latitude="-18.171389" location="Jardin Botanique" longLatPrecision="21" longitude="47.281944" municipality="Reserve Speciale d'Ambohitantely" pageId="26" pageNumber="27" specimenCode="CASENT9007875, CASENT9064777" specimenCount="2" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="AC385C87185EFFC04618324CAA8DEA16" box="[343,442,835,858]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
(
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) and
<typeStatus id="AC385C87185EFFC04580324CA81CEA16" box="[719,811,835,858]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
females (
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) collected from sifted litter in montane rainforest at
<quantity id="B47B4FC0185EFFC045C5326CA9E3EA36" box="[650,724,867,890]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="26" pageNumber="27" unit="m" value="1620.0">
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</quantity>
elevation in
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at
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,
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,
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,
<collectingRegion id="B1472CC7185EFFC045E0328CA89CEAD6" box="[687,939,899,922]" country="Madagascar" name="Antananarivo" pageId="26" pageNumber="27">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B5185EFFC044F8328CAF01EAD6" box="[951,1078,899,922]" name="Madagascar" pageId="26" pageNumber="27">Madagascar</collectingCountry>
, collect- ed
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<collectingDate id="17793D0D185EFFC047D332ACAA62EAF6" box="[156,341,931,954]" pageId="26" pageNumber="27" value="2001-04-17" valueMax="2001-04-22" valueMin="2001-04-17">17-22 April 2001</collectingDate>
</date>
by the
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, deposited in
<collectionCode id="15927AE0185EFFC044E832ACA8E9EAF6" box="[935,990,931,954]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="26" pageNumber="27" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE878185EFFC044A632ACAB85ED77" collectingDate="2003-03-20" collectionCode="CAS" country="Madagascar" elevation="1574" pageId="26" pageNumber="27" specimenCode="CASENT9014779, CASENT9015002, CASENT9014798" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="AC385C87185EFFC044A632ACAF64EAF6" box="[1001,1107,931,954]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
, three additional females (
<specimenCode id="23254A5E185EFFC0461932CCA928EA96" box="[342,543,963,986]" collectionCode="CASENT" pageId="26" pageNumber="27">CASENT9014779</specimenCode>
,
<specimenCode id="23254A5E185EFFC0456132CCA9C3EA96" box="[558,756,963,986]" collectionCode="CASENT" pageId="26" pageNumber="27">CASENT9015002</specimenCode>
and
<specimenCode id="23254A5E185EFFC0447F32CCA8CEEA96" box="[816,1017,963,986]" collectionCode="CASENT" pageId="26" pageNumber="27">CASENT9014798</specimenCode>
), collected by raking tree trunks in montane rainforest at
<quantity id="B47B4FC0185EFFC0451F32ECA9AEEAB6" box="[592,665,995,1018]" metricMagnitude="3" metricUnit="m" metricValue="1.574" pageId="26" pageNumber="27" unit="m" value="1574.0">
<elevation id="F8AE0516185EFFC0451F32ECA9AEEAB6" box="[592,665,995,1018]" metricMagnitude="3" metricUnit="m" metricValue="1.574" pageId="26" pageNumber="27" unit="m" value="1574.0">1574m</elevation>
</quantity>
elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely,
<date id="073DC4E5185EFFC04639350BA92CED57" box="[374,539,1028,1051]" pageId="26" pageNumber="27" value="2003-03-20">
<collectingDate id="17793D0D185EFFC04639350BA92CED57" box="[374,539,1028,1051]" pageId="26" pageNumber="27" value="2003-03-20">20 March 2003</collectingDate>
</date>
by Daniela Andriamalala and Diana Silva, all deposited in
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE185EFFC047E4354BA96BEDF7" pageId="26" pageNumber="27" type="etymology">
<paragraph id="733CE225185EFFC047E4354BA96BEDF7" blockId="26.[123,1174,835,1725]" pageId="26" pageNumber="27">
<emphasis id="41F73E37185EFFC047E4354BAA1DED17" bold="true" box="[171,298,1092,1115]" pageId="26" pageNumber="27">Etymology.</emphasis>
The species epithet is an honor to Dr. Daniela Andriamalala, Malagasy arachnologist, expert on jumping spiders (
<taxonomicName id="B48399A6185EFFC046A0356BA96CED37" authorityName="Blackwall" authorityYear="1841" box="[495,603,1124,1147]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">Salticidae</taxonomicName>
) and goblin spiders (
<taxonomicName id="B48399A6185EFFC04408356BA8F5ED37" box="[839,962,1124,1147]" class="Arachnida" family="Oonopidae" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">Oonopidae</taxonomicName>
), Lakeside International Scholar at CAS, member of the Fisher-Griswold Arthropod Team and collector of many new and interesting arthropods from
<collectingCountry id="0B94A2B5185EFFC0469735ABA961EDF7" box="[472,598,1188,1211]" name="Madagascar" pageId="26" pageNumber="27">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE185EFFC047E435CBABD2EF50" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="733CE225185EFFC047E435CBABD2EF50" blockId="26.[123,1174,835,1725]" pageId="26" pageNumber="27">
<emphasis id="41F73E37185EFFC047E435CBAAEFED97" bold="true" box="[171,472,1220,1243]" pageId="26" pageNumber="27">
Diagnosis.
<taxonomicName id="B48399A6185EFFC0466735CBAAEFED97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,472,1220,1243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E37185EFFC0466735CBAAEFED97" bold="true" box="[296,472,1220,1243]" italics="true" pageId="26" pageNumber="27">Uduba danielae</emphasis>
</taxonomicName>
</emphasis>
are members of group VI, the
<taxonomicName id="B48399A6185EFFC0447F35CBA8EEED97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[816,985,1220,1243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="danielae">Uduba danielae</taxonomicName>
group (Map 20). Females can be distinguished from those of other
<taxonomicName id="B48399A6185EFFC045D835EBA9D6EDB7" authorityName="Simon" authorityYear="1880" box="[663,737,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37185EFFC045D835EBA9D6EDB7" bold="true" box="[663,737,1252,1275]" italics="true" pageId="26" pageNumber="27">Uduba</emphasis>
</taxonomicName>
with a divided cribellum by the form of the female genitalia (
<figureCitation id="EBB8FEA0185EFFC04613340AAADCEC50" box="[348,491,1285,1308]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="26" pageNumber="27">Figs. 70 A, B</figureCitation>
).
<taxonomicName id="B48399A6185EFFC046B0340AA99AEC50" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[511,685,1285,1308]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E37185EFFC046B0340AA99AEC50" bold="true" box="[511,685,1285,1308]" italics="true" pageId="26" pageNumber="27">Uduba danielae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="B48399A6185EFFC044D3340AA8C9EC50" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[924,1022,1285,1308]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E37185EFFC044D3340AA8C9EC50" bold="true" box="[924,1022,1285,1308]" italics="true" pageId="26" pageNumber="27">U. jayjay</emphasis>
</taxonomicName>
by having the median lobe bell-shaped (
<figureCitation id="EBB8FEA0185EFFC046DF342AAACFEC70" box="[400,504,1317,1340]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="26" pageNumber="27">Fig. 70 A</figureCitation>
), sides gently curved (
<taxonomicName id="B48399A6185EFFC045A3342AA878EC70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[748,847,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E37185EFFC045A3342AA878EC70" bold="true" box="[748,847,1317,1340]" italics="true" pageId="26" pageNumber="27">U. jayjay</emphasis>
</taxonomicName>
with median lobe trapezoidal, lateral margins strongly convex), copulatory openings at anterior margin of obliquely transverse grooves, origin at 0.26 epigynum length (
<taxonomicName id="B48399A6185EFFC04572346AA997EC30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[573,672,1381,1404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E37185EFFC04572346AA997EC30" bold="true" box="[573,672,1381,1404]" italics="true" pageId="26" pageNumber="27">U. jayjay</emphasis>
</taxonomicName>
[
<figureCitation id="EBB8FEA0185EFFC045E0346AA82DEC30" box="[687,794,1381,1404]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="26" pageNumber="27">Fig. 70 C</figureCitation>
] with copulatory openings beneath anterior margin of obliquely transverse grooves, origin at 0.48 epigynum length) and the vulva with spermathecal ducts (
<figureCitation id="EBB8FEA0185EFFC0461B34AAAA8EECF0" box="[340,441,1445,1468]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="26" pageNumber="27">Fig. 70 B</figureCitation>
) beginning anteromedially and forming z-shaped path, lateral, medial, lateral, to posterior (
<taxonomicName id="B48399A6185EFFC0463A34CAAAE0EC90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[373,471,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E37185EFFC0463A34CAAAE0EC90" bold="true" box="[373,471,1477,1500]" italics="true" pageId="26" pageNumber="27">U. jayjay</emphasis>
</taxonomicName>
[
<figureCitation id="EBB8FEA0185EFFC046AA34CAA97DEC90" box="[485,586,1477,1500]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="26" pageNumber="27">Fig. 70 F</figureCitation>
] with vulva with spermathecal ducts beginning anteromedially, forming gentle outside curve to vulva base, then entering at apex of BS bulb). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE185EFFC147E43729ABDCEC55" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="733CE225185EFFC147E43729AA92EB39" blockId="26.[123,1174,835,1725]" lastBlockId="27.[122,1174,220,1369]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="41F73E37185EFFC047E43729AA4BEF71" bold="true" box="[171,380,1574,1597]" pageId="26" pageNumber="27">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E37185EFFC045423729A957EF71" bold="true" box="[525,608,1574,1597]" pageId="26" pageNumber="27">Female</emphasis>
(
<typeStatus id="AC385C87185EFFC0453A3729A9EBEF71" box="[629,732,1574,1597]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
, CASENT9007875): Total length 6.60. Markings as in
<figureCitation id="EBB8FEA0185EFFC046673749AA85EF11" box="[296,434,1606,1629]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="26" pageNumber="27">Figs. 5 A, B</figureCitation>
. Carapace 3.20 long, 2.30 wide, 1.10 high; clypeus 0.15 high. Eye diameters: AME 0.12, ALE, PME and PLE 0.16. Chelicerae 1.50 long; sternum 1.50 long, 1.25 wide; labium 0.75 long; palpal coxae 1.10 long. Cribellum divided. Spination: palpus–femur d0-1- 2, patella p1-0, tibia p2-1, tarsus p0-1-1-0; leg I–femur d1-0-0, p0-0-2, tibia v1-1-2, metatarsus v2- 2-3; leg II–femur d1-1-0, p0-0-1, tibia v1-1-2, metatarsus v2-2-3; leg III–femur d1-1-0-1, p0-1-1- 2; leg IV–femur p1-1-0, tibia v1-1-2, r0-1-1-0, metatarsus p0-2-2, v2-2-2, r1-2-2. Scopulae: ventral on tarsi I and II. Leg measurements: I: 2.10 + 1.10 + 1.80 + 1.50 + 1.20 = 7.70; II: 1.90 + 1.00 + 1.40 + 1.30 + 1.10 = 6.70; III: 1.60 + 0.80 + 1.00 + 1.30 + 0.90 = 5.60; IV: 2.40 + 1.00 + 1.80 + 2.10 + 1.15 = 8.45; palpus: 1.00 + 0.55 + 0.60 + NA + 1.00 = 3.15. Leg formula 4123.
<emphasis id="41F73E37185FFFC1430D3053ABEAE8DF" bold="true" pageId="27" pageNumber="28">Female genitalia</emphasis>
(
<typeStatus id="AC385C87185FFFC147A43073AA7CE8DF" box="[235,331,380,403]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
, CASENT9014779): epigynal plate without depressed atrium, median and lateral lobes indistinct (
<figureCitation id="EBB8FEA0185FFFC1462E3092AAFBE8F8" box="[353,460,413,436]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="27" pageNumber="28">Fig. 70 A</figureCitation>
), plate width 1.39 times length; width between bases of LL 0.47 times epigynum width; LL side width 0.25 epigynal plate width; median lobe bell-shaped, broad- er posteriorly, length 0.53 times epigynum length; copulatory openings at anterior margin of obliquely transverse grooves, origin at 0.26 epigynum length, distance between CO 0.47 times epigynum width. Vulva (
<figureCitation id="EBB8FEA0185FFFC146C93312AAD8EB78" box="[390,495,541,564]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="27" pageNumber="28">Fig. 70 B</figureCitation>
) with spermathecal ducts beginning anteromedially and forming z-shaped path, vulva length 0.69 times width, fertilization ducts separated, vulva width 3.53 times distance between FD.
</paragraph>
<paragraph id="733CE225185FFFC147E43371AF64EADA" blockId="27.[122,1174,220,1369]" pageId="27" pageNumber="28">
<emphasis id="41F73E37185FFFC147E43371AA42EBD9" bold="true" box="[171,373,638,661]" pageId="27" pageNumber="28">Variation. Female</emphasis>
(N= 5): Total length 6.30-7.10. Ratios of carapace length / width = 1.39– 1.767, carapace height / width = 0.48-0.68, PER / carapace width = 0.39–0.44, PER/OAL = 2.16– 3.13, PER/AER = 1.05–1.34, OAL/OQL = 1.06–1.14, OQP/ OQA = 1.20–1.29, clypeus height / AME = 1.25–1.67, chelicera length / clypeus height = 8.67–13.00, sternum length / width = 1.07– 1.36, palpal coxa length / width = 2.10–2.22, femur I length / carapace width = 0.87–1.05, metatarsus I length / carapace width = 0.64–0.76, femur IV length / carapace width = 0.96–1.10, palpal tarsus length / carapace width = 0.43–0.57, palpal tibia length / palpal patella length = 1.82–2.22, palpal tarsus length / palpal tibia length = 1.67–2.40, palpal tarsus length / palpal femur length = 1.00–1.11, palpal tibia length / palpal patella length = 0.83–1.22.
<emphasis id="41F73E37185FFFC1440F3270A84DEADA" bold="true" box="[832,890,895,918]" pageId="27" pageNumber="28">Male</emphasis>
variation unknown.
</paragraph>
<paragraph id="733CE225185FFFC147E432AFAAF5EDF4" blockId="27.[122,1174,220,1369]" pageId="27" pageNumber="28">
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M
<emphasis id="41F73E37185FFFC1478D32AFAABFEAFB" bold="true" box="[194,392,928,951]" pageId="27" pageNumber="28">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5185FFFC146C132AFA967EAFB" box="[398,592,928,951]" name="Madagascar" pageId="27" pageNumber="28">MADAGASCAR</collectingCountry>
:
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<collectingRegion id="B1472CC7185FFFC1451532AFA863EAFB" box="[602,852,928,951]" country="Madagascar" name="Antananarivo" pageId="27" pageNumber="28">Antananarivo Province</collectingRegion>
:
</emphasis>
<collectingCounty id="9A5D9AA9185FFFC1442E32AFABF7EA9B" pageId="27" pageNumber="28">Réserve Spéciale d’Ambohitantely</collectingCounty>
,
<collectingMunicipality id="9358785F185FFFC1479D32CFAA3DEA9B" box="[210,266,960,983]" pageId="27" pageNumber="28">Forêt</collectingMunicipality>
d
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,
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59°
<collectorName id="DE7687F3185FFFC1452E32CFA82CEA9B" box="[609,795,960,983]" pageId="27" pageNumber="28">
NE d’
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</collectorName>
, in
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,
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,
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, elev.
<quantity id="B47B4FC0185FFFC1467032EFAABAEABB" box="[319,397,992,1015]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="27" pageNumber="28" unit="m" value="1620.0">
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, sifted litter in montane rainforest,
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</date>
,
<collectorName id="DE7687F3185FFFC144A732EFAA77ED5B" pageId="27" pageNumber="28">Fisher-Griswold Arthropod Team</collectorName>
(
<taxonomicName id="B48399A6185FFFC14613350FA915ED5B" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[348,546,1024,1047]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E37185FFFC14613350FA915ED5B" bold="true" box="[348,546,1024,1047]" italics="true" pageId="27" pageNumber="28">Uduba danielae</emphasis>
</taxonomicName>
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at
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72°NE Ankazobe
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, raking tree trunks, montane rainforest,
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,
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.
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<paragraph id="733CE225185FFFC147E435CEABDCEC55" blockId="27.[122,1174,220,1369]" pageId="27" pageNumber="28">
<emphasis id="41F73E37185FFFC147E435CEAA69ED94" bold="true" box="[171,350,1217,1240]" pageId="27" pageNumber="28">Natural history.</emphasis>
Specimens of the cribellate spider
<taxonomicName id="B48399A6185FFFC1459A35CEA8B4ED94" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[725,899,1217,1240]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E37185FFFC1459A35CEA8B4ED94" bold="true" box="[725,899,1217,1240]" italics="true" pageId="27" pageNumber="28">Uduba danielae</emphasis>
</taxonomicName>
were collected in leaf litter and on mossy tree trunks in montane rainforest. At least some individuals occur on vertical substrates.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE185FFFC147E4342DA9CFEC15" pageId="27" pageNumber="28" type="distribution">
<paragraph id="733CE225185FFFC147E4342DA9CFEC15" blockId="27.[122,1174,220,1369]" pageId="27" pageNumber="28">
<emphasis id="41F73E37185FFFC147E4342DAAC5EC75" bold="true" box="[171,498,1314,1337]" pageId="27" pageNumber="28">
Distribution.
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<emphasis id="41F73E37185FFFC1460B342DAAC5EC75" bold="true" box="[324,498,1314,1337]" italics="true" pageId="27" pageNumber="28">Uduba danielae</emphasis>
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are known only from the
<typeStatus id="AC385C87185FFFC14446342DA800EC75" box="[777,823,1314,1337]" pageId="27" pageNumber="28">type</typeStatus>
locality in montane rainforest in central
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,
<collectingCountry id="0B94A2B5185FFFC14680344DA967EC15" box="[463,592,1346,1369]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
(Maps 10, 20).
</paragraph>
</subSubSection>
</treatment>
</document>

325
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<mods:title id="608A4445C4E2ACF59BAD3E96691F4236">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<taxonomicName id="B48399A61860FFFE473430D4AA82E8BE" ID-CoL="7DCV4" authority="Simon, 1906" authorityName="Simon" authorityYear="1906" box="[123,437,475,498]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
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<emphasis id="41F73E371860FFFE473430D4AA15E8BE" bold="true" box="[123,290,475,498]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
<bibRefCitation id="17129FD41860FFFE466630D4AA82E8BE" author="SIMON, E." box="[297,437,475,498]" pageId="36" pageNumber="37" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon, 1906</bibRefCitation>
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, Maps 2, 15.
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<emphasis id="41F73E371860FFFE4734332DAA20EB7B" bold="true" box="[123,279,546,567]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
<materialsCitation id="C3EBE8781860FFFE466C332DAA97EB7B" box="[291,416,546,567]" country="Madagascar" county="Species" location="Suberbieville" municipality="Paris" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="17129FD41860FFFE466C332DAA97EB7B" author="SIMON, E." box="[291,416,546,567]" pageId="36" pageNumber="37" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<typeStatus id="AC385C871860FFFE46FF332DA933EB7B" box="[432,516,546,567]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
female from
<location id="765CB4FE1860FFFE45D7332DA82DEB7B" LSID="urn:lsid:plazi:treatment:FB2A53331860FFFF473430D4AF49EB1F:765CB4FE1860FFFE45D7332DA82DEB7B" box="[664,794,546,567]" country="Madagascar" county="Species" municipality="Paris" name="Suberbieville" pageId="36" pageNumber="37">Suberbieville</location>
,
<collectingCountry id="0B94A2B51860FFFE4464332DA896EB7B" box="[811,929,546,567]" name="Madagascar" pageId="36" pageNumber="37">Madagascar</collectingCountry>
, specimen AR231 from
<location id="765CB4FE1860FFFE47D4334FA924EB19" LSID="urn:lsid:plazi:treatment:FB2A53331860FFFF473430D4AF49EB1F:765CB4FE1860FFFE47D4334FA924EB19" box="[155,531,576,597]" country="Madagascar" county="Species" municipality="Paris" name="Museum National d'Histoire Naturelle" pageId="36" pageNumber="37">Muséum National d’Histoire Naturelle</location>
,
<collectingMunicipality id="9358785F1860FFFE4552334FA978EB19" box="[541,591,576,597]" pageId="36" pageNumber="37">Paris</collectingMunicipality>
, examined.
<emphasis id="41F73E371860FFFE458C334FA89CEB19" bold="true" box="[707,939,576,597]" pageId="36" pageNumber="37">
<collectingCounty id="9A5D9AA91860FFFE458C334FA839EB19" box="[707,782,576,597]" pageId="36" pageNumber="37">Species</collectingCounty>
status restored
</emphasis>
.
</paragraph>
<paragraph id="733CE2251860FFFE47343350AF35EB38" blockId="36.[123,1174,546,628]" box="[123,1026,607,628]" pageId="36" pageNumber="37">
<taxonomicName id="B48399A61860FFFE47343350ABC0EB38" ID-CoL="7DD6V" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[123,247,607,628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE47343350ABC0EB38" bold="true" box="[123,247,607,628]" italics="true" pageId="36" pageNumber="37">Uduba dahli</emphasis>
</taxonomicName>
:
<bibRefCitation id="17129FD41860FFFE464B3350AAA1EB38" author="LEHTINEN, P. T." box="[260,406,607,628]" pageId="36" pageNumber="37" pagination="199 - 468" refId="ref108999" refString="LEHTINEN, P. T. (1967). Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
[misidentification, not
<taxonomicName id="B48399A61860FFFE45363350A81DEB38" ID-CoL="7DD6V" authority="Simon 1903" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[633,810,607,628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE45363350A99BEB38" bold="true" box="[633,684,607,628]" italics="true" pageId="36" pageNumber="37">dahli</emphasis>
<bibRefCitation id="17129FD41860FFFE45FD3350A81DEB38" author="SIMON, E." box="[690,810,607,628]" pageId="36" pageNumber="37" pagination="669 - 1080" refId="ref109813" refString="SIMON, E. (1903). Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 669 - 1080. [second pdf with detailed publication dates of the single parts] &lt;doi: 10.5962 / bhl. title. 51973&gt;" type="book chapter" year="1903">Simon 1903</bibRefCitation>
</taxonomicName>
;
<emphasis id="41F73E371860FFFE447A3350A8CEEB38" bold="true" box="[821,1017,607,628]" pageId="36" pageNumber="37">Synonymy rejected</emphasis>
]
</paragraph>
<paragraph id="733CE2251860FFFE47E4338BAA68EB97" blockId="36.[123,1174,644,1725]" pageId="36" pageNumber="37">
<emphasis id="41F73E371860FFFE47E4338BAA7BEBD7" bold="true" box="[171,332,644,667]" pageId="36" pageNumber="37">Identification.</emphasis>
<materialsCitation id="C3EBE8781860FFFE461B338BAA6BEB97" country="Madagascar" county="Suberbieville" location="Suberbieville" municipality="Distribution" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
We have examined the
<typeStatus id="AC385C871860FFFE451E338BA999EBD7" box="[593,686,644,667]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B48399A61860FFFE459A338BAF3EEBD7" ID-CoL="7DCV4" authority="Simon 1906" authorityName="Simon" authorityYear="1906" box="[725,1033,644,667]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE459A338BA849EBD7" bold="true" box="[725,894,644,667]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
<bibRefCitation id="17129FD41860FFFE44C9338BAF3EEBD7" author="SIMON, E." box="[902,1033,644,667]" pageId="36" pageNumber="37" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</taxonomicName>
from
<location id="765CB4FE1860FFFE4301338BABF9EBF7" LSID="urn:lsid:plazi:treatment:FB2A53331860FFFF473430D4AF49EB1F:765CB4FE1860FFFE4301338BABF9EBF7" country="Madagascar" county="Suberbieville" municipality="Distribution" name="Suberbieville" pageId="36" pageNumber="37">Suberbieville</location>
,
<collectingCountry id="0B94A2B51860FFFE479633ABAA60EBF7" box="[217,343,676,699]" name="Madagascar" pageId="36" pageNumber="37">Madagascar</collectingCountry>
.
<location id="765CB4FE1860FFFE462A33ABAABBEBF7" LSID="urn:lsid:plazi:treatment:FB2A53331860FFFF473430D4AF49EB1F:765CB4FE1860FFFE462A33ABAABBEBF7" box="[357,396,676,699]" country="Madagascar" county="Suberbieville" municipality="Distribution" name="See" pageId="36" pageNumber="37">See</location>
discussion below under
<emphasis id="41F73E371860FFFE45D833ABA815EBF7" bold="true" box="[663,802,676,699]" pageId="36" pageNumber="37">
<collectingMunicipality id="9358785F1860FFFE45D833ABA815EBF7" box="[663,802,676,699]" pageId="36" pageNumber="37">Distribution</collectingMunicipality>
</emphasis>
for more information on the location of
<collectingCounty id="9A5D9AA91860FFFE478533CBAA6BEB97" box="[202,348,708,731]" pageId="36" pageNumber="37">Suberbieville</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1860FFFE47E433EBA82DEAF0" pageId="36" pageNumber="37" type="discussion">
<paragraph id="733CE2251860FFFE47E433EBA82DEAF0" blockId="36.[123,1174,644,1725]" pageId="36" pageNumber="37">
<emphasis id="41F73E371860FFFE47E433EBAA24EBB7" bold="true" box="[171,275,740,763]" pageId="36" pageNumber="37">Remarks</emphasis>
. Lehtinen accepted
<bibRefCitation id="17129FD41860FFFE46A133EBA9A4EBB7" author="SIMON, E." box="[494,659,740,763]" pageId="36" pageNumber="37" pagination="669 - 1080" refId="ref109813" refString="SIMON, E. (1903). Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 669 - 1080. [second pdf with detailed publication dates of the single parts] &lt;doi: 10.5962 / bhl. title. 51973&gt;" type="book chapter" year="1903">Simon’s (1903)</bibRefCitation>
use of
<taxonomicName id="B48399A61860FFFE45AA33EBA858EBB7" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[741,879,740,763]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE45AA33EBA858EBB7" bold="true" box="[741,879,740,763]" italics="true" pageId="36" pageNumber="37">Uduba dahli</emphasis>
</taxonomicName>
as a replacement name for the preoccupied
<taxonomicName id="B48399A61860FFFE4662320BA83FEA57" authority="Dahl 1901 b" authorityName="F. Dahl" authorityYear="1901" box="[301,776,772,795]" class="Arachnida" family="Udubidae" genus="Marussenca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="41F73E371860FFFE4662320BA9B7EA57" bold="true" box="[301,640,772,795]" italics="true" pageId="36" pageNumber="37">Marussenca madagascariensis</emphasis>
<bibRefCitation id="17129FD41860FFFE45C8320BA83FEA57" author="DAHL, F." box="[647,776,772,795]" pageId="36" pageNumber="37" pagination="244 - 255" refId="ref108020" refString="DAHL, F. (1901 b). Nachtrag zur Uebersicht der Zoropsiden. Sitzungsberichte der Gesellschaft Naturforschen- der Freunde zu Berlin 1901: 244 - 255." type="journal article" year="1901">Dahl 1901b</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="17129FD41860FFFE445B320BA8CEEA57" author="LEHTINEN, P. T." box="[788,1017,772,795]" pageId="36" pageNumber="37" pagination="199 - 468" refId="ref108999" refString="LEHTINEN, P. T. (1967). Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967: 272)</bibRefCitation>
also proposed as a new synonym
<taxonomicName id="B48399A61860FFFE4608322BA970EA77" authority="Simon 1906" authorityName="Simon" authorityYear="1906" box="[327,583,804,827]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE4608322BAA89EA77" bold="true" box="[327,446,804,827]" italics="true" pageId="36" pageNumber="37">U. funerea</emphasis>
<bibRefCitation id="17129FD41860FFFE468A322BA970EA77" author="SIMON, E." box="[453,583,804,827]" pageId="36" pageNumber="37" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</taxonomicName>
equal to
<taxonomicName id="B48399A61860FFFE45E5322BA8A7EA77" authority="Simon 1903" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[682,912,804,827]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE45E5322BA834EA77" bold="true" box="[682,771,804,827]" italics="true" pageId="36" pageNumber="37">U. dahli</emphasis>
<bibRefCitation id="17129FD41860FFFE4444322BA8A7EA77" author="SIMON, E." box="[779,912,804,827]" pageId="36" pageNumber="37" pagination="669 - 1080" refId="ref109813" refString="SIMON, E. (1903). Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 669 - 1080. [second pdf with detailed publication dates of the single parts] &lt;doi: 10.5962 / bhl. title. 51973&gt;" type="book chapter" year="1903">Simon 1903</bibRefCitation>
</taxonomicName>
. Whereas the females of each species are similar to each other, that of
<taxonomicName id="B48399A61860FFFE4518324BA9FBEA17" authorityName="Simon" authorityYear="1906" box="[599,716,836,859]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE4518324BA9FBEA17" bold="true" box="[599,716,836,859]" italics="true" pageId="36" pageNumber="37">U. funerea</emphasis>
</taxonomicName>
has an entire cribellum and that of
<emphasis id="41F73E371860FFFE4371324BABB3EA37" bold="true" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="B48399A61860FFFE4371324BAFA2EA17" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[1086,1173,836,859]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">U. dahli</taxonomicName>
(
</emphasis>
<taxonomicName id="B48399A61860FFFE47CB326BA96EEA37" authority="Dahl 1901 b" authorityName="F. Dahl" authorityYear="1901" box="[132,601,868,891]" class="Arachnida" family="Udubidae" genus="Marussenca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="41F73E371860FFFE47CB326BAAE2EA37" bold="true" box="[132,469,868,891]" italics="true" pageId="36" pageNumber="37">Marussenca madagascariensis</emphasis>
<bibRefCitation id="17129FD41860FFFE4694326BA96EEA37" author="DAHL, F." box="[475,601,868,891]" pageId="36" pageNumber="37" pagination="244 - 255" refId="ref108020" refString="DAHL, F. (1901 b). Nachtrag zur Uebersicht der Zoropsiden. Sitzungsberichte der Gesellschaft Naturforschen- der Freunde zu Berlin 1901: 244 - 255." type="journal article" year="1901">Dahl 1901b</bibRefCitation>
</taxonomicName>
) has a divided cribellum (
<figureCitation id="EBB8FEA01860FFFE4421326BA8E2EA37" box="[878,981,868,891]" captionStart="FIGURE" captionStartId="132.[155,167,1595,1614]" captionTargetBox="[120,1176,1020,1579]" captionTargetId="figure-75@132.[120,1176,1020,1579]" captionTargetPageId="132" captionText="FIGURE 34. Syntype ZMB25276a of Marussenca-madagascariensis Dahl 1901 (= lectotype ♀ Uduba dahli (Simon 1903)). A. Carapace, dorsal. B. Prosoma ventral. C. Spinning organs including divided cribellum. D. Epigynum, ventral. E. Vulva, dorsal. F. Left metatarsus IV, retrodorsal, arrow to calamistrum. G. Calamistrum, left metatarsus IV, retrodorsal, arrows to beginning and end of calamistrum. CA, calamistrum. Scale bars: A, B = 5.0 mm, C, D, E = 1.0 mm, F = 2.0 mm. G = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158645" httpUri="https://zenodo.org/record/13158645/files/figure.png" pageId="36" pageNumber="37">Fig. 34 C</figureCitation>
). We consider this difference in cribellar form to be species diagnostic for our revision and therefore we reject the synonymy of
<taxonomicName id="B48399A61860FFFE47A432AAAADAEAF0" authority="Simon 1906" authorityName="Simon" authorityYear="1906" box="[235,493,933,956]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE47A432AAAA54EAF0" bold="true" box="[235,355,933,956]" italics="true" pageId="36" pageNumber="37">U. funerea</emphasis>
<bibRefCitation id="17129FD41860FFFE462432AAAADAEAF0" author="SIMON, E." box="[363,493,933,956]" pageId="36" pageNumber="37" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="B48399A61860FFFE456232AAA821EAF0" authority="Simon 1903" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[557,790,933,956]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE456232AAA9B0EAF0" bold="true" box="[557,647,933,956]" italics="true" pageId="36" pageNumber="37">U. dahli</emphasis>
<bibRefCitation id="17129FD41860FFFE45C032AAA821EAF0" author="SIMON, E." box="[655,790,933,956]" pageId="36" pageNumber="37" pagination="669 - 1080" refId="ref109813" refString="SIMON, E. (1903). Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 669 - 1080. [second pdf with detailed publication dates of the single parts] &lt;doi: 10.5962 / bhl. title. 51973&gt;" type="book chapter" year="1903">Simon 1903</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1860FFFE47E432CAAA15EDF0" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="733CE2251860FFFE47E432CAAA15EDF0" blockId="36.[123,1174,644,1725]" pageId="36" pageNumber="37">
<emphasis id="41F73E371860FFFE47E432CAAAF9EA90" bold="true" box="[171,462,965,988]" pageId="36" pageNumber="37">
Diagnosis.
<taxonomicName id="B48399A61860FFFE466932CAAAF9EA90" authorityName="Simon" authorityYear="1906" box="[294,462,965,988]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE466932CAAAF9EA90" bold="true" box="[294,462,965,988]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.c, the
<taxonomicName id="B48399A61860FFFE459232CAA84CEA90" authorityName="Simon" authorityYear="1906" box="[733,891,965,988]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA01860FFFE448032CAAF13EA90" box="[975,1060,965,988]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="36" pageNumber="37">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are cribellate, with cribellum entire. Males are unknown. The females of
<taxonomicName id="B48399A61860FFFE47BF350AAAA0ED50" authorityName="Simon" authorityYear="1906" box="[240,407,1029,1052]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE47BF350AAAA0ED50" bold="true" box="[240,407,1029,1052]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="B48399A61860FFFE4401350AA8AFED50" authorityName="Simon" authorityYear="1880" box="[846,920,1029,1052]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371860FFFE4401350AA8AFED50" bold="true" box="[846,920,1029,1052]" italics="true" pageId="36" pageNumber="37">Uduba</emphasis>
</taxonomicName>
having an entire cribellum and an epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01860FFFE44F9352AAF29ED70" box="[950,1054,1061,1084]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="36" pageNumber="37">Fig. 79 C</figureCitation>
); the entire cribellum is unlike
<taxonomicName id="B48399A61860FFFE461E354AA90DED10" authority="Simon 1903" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[337,570,1093,1116]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="41F73E371860FFFE461E354AAA9BED10" bold="true" box="[337,428,1093,1116]" italics="true" pageId="36" pageNumber="37">U. dahli</emphasis>
<bibRefCitation id="17129FD41860FFFE46F9354AA90DED10" author="SIMON, E." box="[438,570,1093,1116]" pageId="36" pageNumber="37" pagination="669 - 1080" refId="ref109813" refString="SIMON, E. (1903). Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 669 - 1080. [second pdf with detailed publication dates of the single parts] &lt;doi: 10.5962 / bhl. title. 51973&gt;" type="book chapter" year="1903">Simon 1903</bibRefCitation>
</taxonomicName>
but like
<taxonomicName id="B48399A61860FFFE45EF354AA809ED10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[672,830,1093,1116]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="halabe">
<emphasis id="41F73E371860FFFE45EF354AA809ED10" bold="true" box="[672,830,1093,1116]" italics="true" pageId="36" pageNumber="37">Uduba halabe</emphasis>
</taxonomicName>
.
<taxonomicName id="B48399A61860FFFE4401354AA8C0ED10" authorityName="Simon" authorityYear="1906" box="[846,1015,1093,1116]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371860FFFE4401354AA8C0ED10" bold="true" box="[846,1015,1093,1116]" italics="true" pageId="36" pageNumber="37">Uduba funerea</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="B48399A61860FFFE465E356AAA4AED30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[273,381,1125,1148]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="halabe">
<emphasis id="41F73E371860FFFE465E356AAA4AED30" bold="true" box="[273,381,1125,1148]" italics="true" pageId="36" pageNumber="37">U. halabe</emphasis>
</taxonomicName>
by having the copulatory openings hidden beneath the lateral margins of the atrium (
<figureCitation id="EBB8FEA01860FFFE47B9358AAA6BEDD0" box="[246,348,1157,1180]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="36" pageNumber="37">Fig. 79 E</figureCitation>
); in
<taxonomicName id="B48399A61860FFFE46C3358AAAC1EDD0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[396,502,1157,1180]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="halabe">
<emphasis id="41F73E371860FFFE46C3358AAAC1EDD0" bold="true" box="[396,502,1157,1180]" italics="true" pageId="36" pageNumber="37">U. halabe</emphasis>
</taxonomicName>
the CO are located centrally beneath the epigynal median lobe (
<figureCitation id="EBB8FEA01860FFFE47CC35AAAA22EDF0" box="[131,277,1189,1212]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="36" pageNumber="37">Figs. 74 C, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1860FFFF47E435CAAB9EE838" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="733CE2251860FFFE47E435CAAF39EFF1" blockId="36.[123,1174,644,1725]" pageId="36" pageNumber="37">
<emphasis id="41F73E371860FFFE47E435CAAAA7ED90" bold="true" box="[171,400,1221,1244]" pageId="36" pageNumber="37">Description. Female</emphasis>
(
<typeStatus id="AC385C871860FFFE46D035CAA934ED90" box="[415,515,1221,1244]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
). We made a detailed examination of the female
<typeStatus id="AC385C871860FFFE434135CAAF5CED90" box="[1038,1131,1221,1244]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
but upon returning the specimens to the Muséum National d’Histoire Naturelle in Paris in the 1990’s we kept only a few somatic illustrations and detailed illustrations and measurements of the female genitalia. We are therefore unable to provide details of measurements and spination for the
<typeStatus id="AC385C871860FFFE4312342AAB9EEC10" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
and can only describe and illustrate the female genitalia in detail. Cribellum entire.
<emphasis id="41F73E371860FFFE430D344AABEAEC30" bold="true" pageId="36" pageNumber="37">Female genitalia</emphasis>
(
<typeStatus id="AC385C871860FFFE47A5346AAA67EC30" box="[234,336,1381,1404]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
) Epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01860FFFE436F346AAFBEEC30" box="[1056,1161,1381,1404]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="36" pageNumber="37">Fig. 79 D</figureCitation>
), plate width 1.49 times length; atrium sides concave, atrium wide, width 1.80 times atrium length; epigynum length 1.90 times atrium length, atrium width 0.58 times epigynum width; atrium width at side of ML 1.88 times ML width; atrium and median lobe base arise anteriad of epigastric groove (
<figureCitation id="EBB8FEA01860FFFE47CC34EAABDFECB0" box="[131,232,1509,1532]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="36" pageNumber="37">Fig. 79 C</figureCitation>
), atrium base at 0.20 of epigynal plate length, median lobe narrow, slightly wider at base, median lobe length 2.70 times width, ML extends to apex of atrium, ML length 0.60 times epigynal length; lateral lobes narrow, atrium width 4.50 times LL width; copulatory openings hidden beneath lateral sides of atrium, distance between CO 0.62 width of epigynal plate, CO distance from epigastric furrow 0.50 times epigynum length. Vulva (
<figureCitation id="EBB8FEA01860FFFE45BA3769A8B2EF31" box="[757,901,1638,1661]" captionStart="FIGURE" captionStartId="167.[155,167,1184,1203]" captionTargetBox="[148,1149,222,1167]" captionTargetId="figure-16@167.[120,1176,216,1171]" captionTargetPageId="167" captionText="FIGURE 79. Uduba spp., female genitalia drawings. A, B. U. woodae, new species, (CASENT9000536) Forêt de Beroboka. C–E. Uduba funerea Simon 1906, holotype. B, D. Epigynum, ventral. A, C. Vulva, dorsal. E. Vulva, anterodorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158751" httpUri="https://zenodo.org/record/13158751/files/figure.png" pageId="36" pageNumber="37">Figs. 79 D, E</figureCitation>
) with spermathecal ducts asymmetrical, on left making three broad, transverse curves, on right three opposed loops, vulva length 0.65 times width, vulva width 4.50 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251861FFFF47E431D3AF43E9BF" blockId="37.[123,1174,220,595]" box="[171,1140,220,243]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E431D3AA2DE9BF" bold="true" box="[171,282,220,243]" pageId="37" pageNumber="38">Variation.</emphasis>
Unknown: only the female
<typeStatus id="AC385C871861FFFF450431D3A99FE9BF" box="[587,680,220,243]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B48399A61861FFFF458031D3AF36E9BF" authority="Simon 1906" authorityName="Simon" authorityYear="1906" box="[719,1025,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371861FFFF458031D3A840E9BF" bold="true" box="[719,887,220,243]" italics="true" pageId="37" pageNumber="38">Uduba funerea</emphasis>
<bibRefCitation id="17129FD41861FFFF443031D3AF36E9BF" author="SIMON, E." box="[895,1025,220,243]" pageId="37" pageNumber="38" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</taxonomicName>
is known.
</paragraph>
<paragraph id="733CE2251861FFFF47E431F3AA83E87F" blockId="37.[123,1174,220,595]" pageId="37" pageNumber="38">
<materialsCitation id="C3EBE8781861FFFF47E431F3AA9AE87F" collectionCode="MNHN" country="Madagascar" location="Paris" pageId="37" pageNumber="38" specimenCount="1" typeStatus="holotype">
M
<emphasis id="41F73E371861FFFF478D31F3AABAE85F" bold="true" box="[194,397,252,275]" pageId="37" pageNumber="38">aterial examined.</emphasis>
Only the
<typeStatus id="AC385C871861FFFF454731F3A952E85F" box="[520,613,252,275]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
female, specimen AR231 at
<collectionCode id="15927AE01861FFFF44FA31F3AF3EE85F" box="[949,1033,252,275]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="37" pageNumber="38">MNHN</collectionCode>
,
<location id="765CB4FE1861FFFF435631F3AF66E85F" LSID="urn:lsid:plazi:treatment:FB2A53331860FFFF473430D4AF49EB1F:765CB4FE1861FFFF435631F3AF66E85F" box="[1049,1105,252,275]" country="Madagascar" name="Paris" pageId="37" pageNumber="38">Paris</location>
, from “Suberbieville”,
<collectingCountry id="0B94A2B51861FFFF46603013AA9AE87F" box="[303,429,284,307]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251861FFFF47E43032AB9EE838" blockId="37.[123,1174,220,595]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E43032AA57E818" bold="true" box="[171,352,317,340]" pageId="37" pageNumber="38">Natural history.</emphasis>
Unknown. The female has an entire cribellum, suggesting use of cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E43072AF49EB1F" pageId="37" pageNumber="38" type="distribution">
<paragraph id="733CE2251861FFFF47E43072AF49EB1F" blockId="37.[123,1174,220,595]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E43072AA0AE8D8" bold="true" box="[171,317,381,404]" pageId="37" pageNumber="38">Distribution.</emphasis>
<taxonomicName id="B48399A61861FFFF460B3072A9B4E8D8" authority="Simon 1906" authorityName="Simon" authorityYear="1906" box="[324,643,381,404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="funerea">
<emphasis id="41F73E371861FFFF460B3072AAC7E8D8" bold="true" box="[324,496,381,404]" italics="true" pageId="37" pageNumber="38">Uduba funerea</emphasis>
<bibRefCitation id="17129FD41861FFFF46B33072A9B4E8D8" author="SIMON, E." box="[508,643,381,404]" pageId="37" pageNumber="38" pagination="284 - 308" refId="ref109865" refString="SIMON, E. (1906). Etude sur les araignees de la section des cribellates. Annales de la Societe Entomologique de Belgique 50: 284 - 308." type="journal article" year="1906">Simon 1906</bibRefCitation>
</taxonomicName>
is known only from the
<typeStatus id="AC385C871861FFFF44E63072A8E0E8D8" box="[937,983,381,404]" pageId="37" pageNumber="38">type</typeStatus>
locality, “Suberbieville”, in Majunga Province,
<collectingCountry id="0B94A2B51861FFFF469C3093A965E8FF" box="[467,594,412,435]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, approximately
<quantity id="B47B4FC01861FFFF444E3093A866E8FF" box="[769,849,412,435]" metricMagnitude="5" metricUnit="m" metricValue="2.4" pageId="37" pageNumber="38" unit="km" value="240.0">240 km</quantity>
NNW of
<collectingRegion id="B1472CC71861FFFF44F63093AF7DE8FF" box="[953,1098,412,435]" country="Madagascar" name="Antananarivo" pageId="37" pageNumber="38">Antananarivo</collectingRegion>
(Maps 2, 15). Gold mining has been conducted in this region since the late 1800’s. The town, formerly called Suberbieville after the industrialist who exploited the gold of the Ikopa River, is now known as Maevatanana and is located on the right bank of the Ikopa River, south of its confluence with the Betsiboka River. It is located at about
<quantity id="B47B4FC01861FFFF45DE3313A9F7EB7F" box="[657,704,540,563]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="38" unit="m" value="70.0">70m</quantity>
elevation at approximately S16°57′00ʺ, E46°50′00ʺ and is the main stop of the RN 4, the road connecting
<collectingRegion id="B1472CC71861FFFF44053333A8ECEB1F" box="[842,987,572,595]" country="Madagascar" name="Antananarivo" pageId="37" pageNumber="38">Antananarivo</collectingRegion>
to
<collectingRegion id="B1472CC71861FFFF434F3333AF4DEB1F" box="[1024,1146,572,595]" country="Madagascar" name="Mahajanga" pageId="37" pageNumber="38">Mahajanga</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

319
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<mods:title id="D97444211300D26C9487B8059CD4E865">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="7406D7C54B64A9BFB7C721D3342441C8">Ubick, Darrell</mods:namePart>
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<mods:classification id="E0115AC5A4FDC06278FF5DF3BCE1E91D">journal article</mods:classification>
<mods:identifier id="586357ED55B702741C9B5A7144FDBB92" type="DOI">10.5281/zenodo.13158554</mods:identifier>
<mods:identifier id="DC33C29EE5C10430C8F3B7B1014047AE" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="356EAF1F45ADB662FEC547F459CC087F" type="Zenodo-Dep">13158554</mods:identifier>
<mods:identifier id="61075B6BBE4B0D34EC85B073B68B66E5" type="ZooBank">74E0D22E-9890-4FF0-80AB-8FCBBA976B2F</mods:identifier>
</mods:mods>
<treatment id="FB2A53331861FFFC47343371A9D8EDFA" LSID="urn:lsid:zoobank.org:act:775E6518-0481-40E3-A112-A0519FA0B787" httpUri="http://treatment.plazi.org/id/FB2A53331861FFFC47343371A9D8EDFA" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="3B99B1AE1861FFFF47343371AAFBEBD9" box="[123,460,638,661]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="733CE2251861FFFF47343371AAFBEBD9" blockId="37.[123,589,638,691]" box="[123,460,638,661]" pageId="37" pageNumber="38">
<heading id="287455491861FFFF47343371AAFBEBD9" box="[123,460,638,661]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="41F73E371861FFFF47343371AAFBEBD9" bold="true" box="[123,460,638,661]" pageId="37" pageNumber="38">
<taxonomicName id="B48399A61861FFFF47343371AA0DEBD9" ID-CoL="C8KJ9" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,314,638,661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="goodmani" status="sp. nov.">
<emphasis id="41F73E371861FFFF47343371AA0DEBD9" bold="true" box="[123,314,638,661]" italics="true" pageId="37" pageNumber="38">Uduba goodmani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1861FFFF46073371AAFBEBD9" box="[328,460,638,661]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47343391A97AEBFF" box="[123,589,670,691]" pageId="37" pageNumber="38" type="description">
<paragraph id="733CE2251861FFFF47343391A97AEBFF" blockId="37.[123,589,638,691]" box="[123,589,670,691]" pageId="37" pageNumber="38">
<figureCitation id="EBB8FEA01861FFFF47343391ABD8EBFF" box="[123,239,670,691]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="37" pageNumber="38">Figures 7 D</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF47B53391AA11EBFF" box="[250,294,670,691]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="37" pageNumber="38">20 F</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF467F3391AA4AEBFF" box="[304,381,670,691]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">77 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF46C73391AAF8EBFF" box="[392,463,670,691]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">80 E, F</figureCitation>
, Maps 6, 16.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E433CCA8D7EA37" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="733CE2251861FFFF47E433CCA8D7EA37" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E433CCAA67EB96" bold="true" box="[171,336,707,730]" pageId="37" pageNumber="38">Type material.</emphasis>
<materialsCitation id="C3EBE8781861FFFF461833CCAFA4EA57" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1875" location="Reserve Naturelle Integrale d'Andohahela" pageId="37" pageNumber="38" specimenCode="CASENT9006074" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C871861FFFF461833CCAA8DEB96" box="[343,442,707,730]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
female from camp 5 at
<quantity id="B47B4FC01861FFFF458E33CCA83BEB96" box="[705,780,707,730]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="37" pageNumber="38" unit="m" value="1875.0">
<elevation id="F8AE05161861FFFF458E33CCA83BEB96" box="[705,780,707,730]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="37" pageNumber="38" unit="m" value="1875.0">1875m</elevation>
</quantity>
elevation in
<location id="765CB4FE1861FFFF44D533CCAA66EBB6" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1861FFFF44D533CCAA66EBB6" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="37" pageNumber="38" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
(24º33.7ʹS, 46º43.3ʹE),
<collectingRegion id="B1472CC71861FFFF451333ECA826EBB6" box="[604,785,739,762]" country="Madagascar" name="Toliara" pageId="37" pageNumber="38">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51861FFFF445033ECA8A9EBB6" box="[799,926,739,762]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, collected
<date id="073DC4E51861FFFF435633ECAA58EA57" pageId="37" pageNumber="38" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">
<collectingDate id="17793D0D1861FFFF435633ECAA58EA57" pageId="37" pageNumber="38" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">27 November–5 December 1995</collectingDate>
</date>
by
<collectorName id="DE7687F31861FFFF46D0320BA91FEA57" box="[415,552,772,795]" pageId="37" pageNumber="38">S. Goodman</collectorName>
(SG1995-01) (
<specimenCode id="23254A5E1861FFFF459E320BA8ABEA57" box="[721,924,772,795]" collectionCode="CASENT" pageId="37" pageNumber="38">CASENT9006074</specimenCode>
), deposited in
<collectionCode id="15927AE01861FFFF430D320BAFA4EA57" box="[1090,1171,772,795]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="37" pageNumber="38" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781861FFFF4734322BA8ECEA37" collectingDate="2002-01-21" collectingDateMax="2002-01-25" collectingDateMin="2002-01-21" collectionCode="CAS" country="Madagascar" elevation="900" latitude="-24.76389" location="Col du Sedro in Parc National d'Andohahela" longLatPrecision="21" longitude="46.751667" municipality="Fisher-Griswold Arthropod Team" pageId="37" pageNumber="38" specimenCode="CASENT9009538" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="AC385C871861FFFF4734322BABEFEA77" box="[123,216,804,827]" pageId="37" pageNumber="38" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1861FFFF4677322BA936EA77" box="[312,513,804,827]" collectionCode="CASENT" pageId="37" pageNumber="38">CASENT9009538</specimenCode>
) from montane rainforest at
<quantity id="B47B4FC01861FFFF4474322BA84FEA77" box="[827,888,804,827]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="m" value="900.0">
<elevation id="F8AE05161861FFFF4474322BA84FEA77" box="[827,888,804,827]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="m" value="900.0">900m</elevation>
</quantity>
elevation at
<location id="765CB4FE1861FFFF434A322BAAFCEA17" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1861FFFF434A322BAAFCEA17" country="Madagascar" latitude="-24.76389" longLatPrecision="21" longitude="46.751667" municipality="Fisher-Griswold Arthropod Team" name="Col du Sedro in Parc National d'Andohahela" pageId="37" pageNumber="38" stateProvince="Toliara">Col du Sedro in Parc National d’Andohahela</location>
(
<geoCoordinate id="16B784E21861FFFF4694324BA961EA17" box="[475,598,836,859]" degrees="24" direction="south" minutes="45" orientation="latitude" pageId="37" pageNumber="38" precision="15" seconds="50" value="-24.76389">24°45ʹ50ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21861FFFF452B324BA9E3EA17" box="[612,724,836,859]" degrees="46" direction="east" minutes="45" orientation="longitude" pageId="37" pageNumber="38" precision="15" seconds="6" value="46.751667">46°45ʹ6ʺE</geoCoordinate>
),
<collectingRegion id="B1472CC71861FFFF45A7324BA8ABEA17" box="[744,924,836,859]" country="Madagascar" name="Toliara" pageId="37" pageNumber="38">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51861FFFF44E8324BAF11EA17" box="[935,1062,836,859]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, collected
<date id="073DC4E51861FFFF4734326BAA62EA37" box="[123,341,868,891]" pageId="37" pageNumber="38" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">
<collectingDate id="17793D0D1861FFFF4734326BAA62EA37" box="[123,341,868,891]" pageId="37" pageNumber="38" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">21–25 January 2002</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F1861FFFF46E6326BA839EA37" box="[425,782,868,891]" pageId="37" pageNumber="38">Fisher-Griswold Arthropod Team</collectingMunicipality>
, deposited in
<collectionCode id="15927AE01861FFFF44EB326BA8ECEA37" box="[932,987,868,891]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E4328BAA7DEA97" pageId="37" pageNumber="38" type="etymology">
<paragraph id="733CE2251861FFFF47E4328BAA7DEA97" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E4328BAA1DEAD7" bold="true" box="[171,298,900,923]" pageId="37" pageNumber="38">Etymology.</emphasis>
The species epithet is a patronym in honor of Steve Goodman, naturalist, author, collector and mammologist who has made substantial contributions to our knowledge of
<collectingCountry id="0B94A2B51861FFFF431F32ABAB88EA97" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E432EBA8CFEDF0" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="733CE2251861FFFF47E432EBA8CFEDF0" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E432EBAADEEAB7" bold="true" box="[171,489,996,1019]" pageId="37" pageNumber="38">
Diagnosis.
<taxonomicName id="B48399A61861FFFF466732EBAADEEAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,489,996,1019]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371861FFFF466732EBAADEEAB7" bold="true" box="[296,489,996,1019]" italics="true" pageId="37" pageNumber="38">Uduba goodmani</emphasis>
</taxonomicName>
</emphasis>
belong to Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A61861FFFF4734350BAA76ED57" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,321,1028,1051]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). They are ecribellate. Females can be distinguished from those of other
<taxonomicName id="B48399A61861FFFF479A352BAA44ED77" box="[213,371,1060,1083]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF479A352BAA28ED77" bold="true" box="[213,287,1060,1083]" italics="true" pageId="37" pageNumber="38">Uduba</emphasis>
species
</taxonomicName>
by the form of the female genitalia (
<figureCitation id="EBB8FEA01861FFFF45B2352BA8BAED77" box="[765,909,1060,1083]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">Figs. 77 A–C</figureCitation>
.
<figureCitation id="EBB8FEA01861FFFF44D7352BA8D1ED77" box="[920,998,1060,1083]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">80 E, F</figureCitation>
<emphasis id="41F73E371861FFFF44A9352BA8CFED77" bold="true" box="[998,1016,1060,1083]" pageId="37" pageNumber="38">):</emphasis>
epigynal plate a broad oval, width 1.50 times length, lateral lobes with convex edges forming ridges on each side (
<figureCitation id="EBB8FEA01861FFFF47CC356BABDEED37" box="[131,233,1124,1147]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 E</figureCitation>
); vulva broad, width one and-a-half times length with spermathecal ducts characteristic, forming one and-a-half loose, transverse loops (
<figureCitation id="EBB8FEA01861FFFF45CB358AA9DBEDD0" box="[644,748,1157,1180]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 F</figureCitation>
), copulatory duct lateral and extending mesally to lead to posterior, widely spaced fertilization ducts. Males are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFC47E435CAAA53EA19" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="733CE2251861FFFC47E435CAA96FE8DF" blockId="37.[123,1174,707,1725]" lastBlockId="38.[123,1174,220,1206]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E435CAAA40ED90" bold="true" box="[171,375,1221,1244]" pageId="37" pageNumber="38">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E371861FFFF46B135CAA966ED90" bold="true" box="[510,593,1221,1244]" pageId="37" pageNumber="38">Female</emphasis>
(
<typeStatus id="AC385C871861FFFF452F35CAA9FDED90" box="[608,714,1221,1244]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
): Total length 16.43. Markings as in
<figureCitation id="EBB8FEA01861FFFF431935CAABA6EDB0" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="37" pageNumber="38">Fig. 7 D</figureCitation>
. Carapace 8.50 long, 5.71 wide, 3.50 high; clypeus 0.43 high. Eye diameters: AME 0.27, ALE and PME 0.36, PLE 0.38. Chelicerae 3.79 long; sternum 3.71 long, 2.79 wide; labium 1.82 long; palpal coxae 2.82 long. With a colulus (
<figureCitation id="EBB8FEA01861FFFF456A342AA9BEEC70" box="[549,649,1317,1340]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="37" pageNumber="38">Fig. 20 F</figureCitation>
<emphasis id="41F73E371861FFFF45C6342AA9A5EC70" bold="true" box="[649,658,1317,1340]" pageId="37" pageNumber="38">)</emphasis>
. Spination: palpus–femur d0-1-1, p0-0-1, patella p1-0, tibia p2-1, tarsus p2-1, r1-0-0; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2-0, metatarsus v2- 2-3; leg II–femur d1-0-0, p0-0-1, tibia p0-0-1-0, v1-1-1-0, metatarsus v2-2-3; leg III–femur d0-0- 0-1, p1-1-0-0-2, r1-1-0-1, tibia d0-1-0, p0-1-1-0, v1-2-0, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1- 1-2; leg IV–femur d1-0-1-0, tibia v1-1-2, metatarsus p0-0-2-2, v2-1-2-2, r1-2-2. Scopulae: strong ventral beneath all tarsi and metatarsi and beneath tibiae I and II. Leg measurements (
<typeStatus id="AC385C871861FFFF437D34CAAFA5EC90" box="[1074,1170,1477,1500]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
, CASENT9009538): I: 4.20 + 2.00 + 3.60 + 3.10 + 2.00 = 14.90; II: 3.90 + 2.00 + 3.10 + 2.90 + 1.90 = 13.80; III: 3.20 + 1.50 + 1.90 + 2.80 + 1.55 = 10.95; IV: 4.20 + 1.80 + 3.60 + 4.10 + 1.90 = 15.60; palpus: 2.10 + 1.00 + 1.10 + NA + 1.80 = 6.00. Leg formula 4123.
<emphasis id="41F73E371861FFFF434E3729AB9AEF11" bold="true" pageId="37" pageNumber="38">Female genitalia</emphasis>
(
<typeStatus id="AC385C871861FFFF47F33749AA17EF11" box="[188,288,1606,1629]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
): epigynal plate without depressed atrium, lateral lobes extending to sides forming blunt, earlike lobes (
<figureCitation id="EBB8FEA01861FFFF461A3769AAD1EF31" box="[341,486,1638,1661]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">Figs. 77 A, C</figureCitation>
), plate width 1.63 times length; width between bases of LL 0.72 times epigynum width; lateral ears short and blunt (
<figureCitation id="EBB8FEA01861FFFF45E43789A823EFD1" box="[683,788,1670,1693]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 E</figureCitation>
), arising near epigastric furrow, lateral ears (LL side) width 0.75 times LL side length, (LL side) width 0.16 times width epigynum; median lobe a flat plate, each side with an oblique bulge where copulatory openings are found, ML width 0.64 times epigynum width, 1.36 times ML length, CO origin at 0.485 epigynum length from epigastric furrow, distance between copulatory openings 0.41 times epigynum width. Vulva (
<figureCitation id="EBB8FEA01862FFFC47CC3033ABDEE81F" box="[131,233,316,339]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Fig. 77 B</figureCitation>
) with copulatory ducts lateral, median to these spermathecal ducts forming a single longitudinal loop (
<figureCitation id="EBB8FEA01862FFFC46523053AAB7E83F" box="[285,384,348,371]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">Fig. 80 F</figureCitation>
), vulva length 0.53 times width, fertilization ducts widely separated, vulva width only 3.21 times distance between FD.
</paragraph>
<paragraph id="733CE2251862FFFC47E43093AA53EA19" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43093AA2DE8FF" bold="true" box="[171,282,412,435]" pageId="38" pageNumber="39">Variation.</emphasis>
(N=2): Carapace length / width = 1.49–1.51, carapace height / width = 0.61–0.64, PER / carapace width = 0.45–0.69, PER/OAL = 2.70–2.90, PER/AER = 1.30–1.32, OAL/OQL = 1.00–1.25, OQP/ OQA = 1.07–1.18, clypeus height / AME = 1.60–1.75, cheliceral length / clypeus height = 7.71–8.83, sternum length / width = 1.22–1.33, palpal coxa length / width = 2.11–2.39, femur I length / carapace width = 1.08–1.14, metatarsus I length / carapace width = 0.58–0.79, femur IV length / carapace width = 1.08–1.09, palpal tarsus length / carapace width = 0.44–0.46, palpal tibia length / palpal patella length = 1.59–1.80, palpal tarsus length / palpal tibia length = 1.40–1.64, palpal tarsus length / palpal femur length = 0.80–0.86, palpal tibia length / palpal patella length = 1.10–1.14. Epigynal plate (
<figureCitation id="EBB8FEA01862FFFC456D3392A98FEBF8" box="[546,696,669,692]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Figs. 77 A, C</figureCitation>
,
<figureCitation id="EBB8FEA01862FFFC458A3392A9CDEBF8" box="[709,762,669,692]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">80 E</figureCitation>
) width 1.50–1.63 times length, width between bases of LL 0.72–0.75 times epigynum width, CO origin at 0.43–0.48 epigynum length from epigastric furrow, distance between copulatory openings 0.33–0.41 times epigynum width; vulva (
<figureCitation id="EBB8FEA01862FFFC478933F2AA0CEA58" box="[198,315,765,788]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Figs. 77 B</figureCitation>
,
<figureCitation id="EBB8FEA01862FFFC460833F2AA4DEA58" box="[327,378,765,788]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">80 F</figureCitation>
) with spermathecal ducts forming one and-a-half loose, transverse loops, vulva length 0.53–0.64 times width, fertilization ducts widely spaced, vulva width 2.58–3.21 times distance between FD.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E43251ABC7ED3A" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="733CE2251862FFFC47E43251AAF5ED79" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43251AABCEA39" bold="true" box="[171,395,862,885]" pageId="38" pageNumber="39">Material examined.</emphasis>
<materialsCitation id="C3EBE8781862FFFC46D63251AF6FEAF9" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMNH" collectorName="SE Andranondambo &amp; S. Goodman" country="Madagascar" elevation="1875" location="Reserve Naturelle Integrale d'Andohahela" pageId="38" pageNumber="39" specimenCode="CASENT9006074" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="0B94A2B51862FFFC46D63251A96CEA39" box="[409,603,862,885]" name="Madagascar" pageId="38" pageNumber="39">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371862FFFC45293251A820EA39" box="[614,791,862,885]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="B1472CC71862FFFC45293251A820EA39" box="[614,791,862,885]" country="Madagascar" name="Toliara" pageId="38" pageNumber="39">Toliara Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1862FFFC44693251ABE9EAD9" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1862FFFC44693251ABE9EAD9" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="38" pageNumber="39" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
, 20.0 km
<collectorName id="DE7687F31862FFFC46093271A929EAD9" box="[326,542,894,917]" pageId="38" pageNumber="39">SE Andranondambo</collectorName>
, camp 5, 24º33.7ʹS, 46º43.3ʹE, elev.
<quantity id="B47B4FC01862FFFC44EF3271A8DAEAD9" box="[928,1005,894,917]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="38" pageNumber="39" unit="m" value="1875.0">
<elevation id="F8AE05161862FFFC44EF3271A8DAEAD9" box="[928,1005,894,917]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="38" pageNumber="39" unit="m" value="1875.0">1875m</elevation>
</quantity>
,
<date id="073DC4E51862FFFC44B83271AA75EAF9" pageId="38" pageNumber="39" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">
<collectingDate id="17793D0D1862FFFC44B83271AA75EAF9" pageId="38" pageNumber="39" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">27 November– 5 December 1995</collectingDate>
</date>
,
<collectorName id="DE7687F31862FFFC461F3291AAEEEAF9" box="[336,473,926,949]" pageId="38" pageNumber="39">S. Goodman</collectorName>
[SG1995-01] (
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,
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,
<specimenCount id="658529AC1862FFFC44823291A8D9EAF9" box="[973,1006,926,949]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01862FFFC44B03291AF66EAF9" box="[1023,1105,926,949]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="38" pageNumber="39" type="Museum">FMNH</collectionCode>
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</materialsCitation>
;
<materialsCitation id="C3EBE8781862FFFC43293291AA88ED79" collectingDate="2002-01-21" collectingDateMax="2002-01-25" collectingDateMin="2002-01-21" collectionCode="CAS" collectorName="Col du Sedro &amp; Fisher-Griswold Arthropod Team" country="Madagascar" elevation="900" latitude="-24.76389" location="Col du Sedro" longLatPrecision="21" longitude="46.751667" municipality="Parc National d'Andohahela" pageId="38" pageNumber="39" specimenCode="CASENT9009538" specimenCount="1" specimenCount-female="1" stateProvince="Toliara Province" typeStatus="paratype">
<collectingMunicipality id="9358785F1862FFFC43293291AAB5EA99" pageId="38" pageNumber="39">Parc National d’Andohahela</collectingMunicipality>
,
<collectorName id="DE7687F31862FFFC46D732B1A900EA99" box="[408,567,958,981]" pageId="38" pageNumber="39">
<location id="765CB4FE1862FFFC46D732B1A900EA99" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1862FFFC46D732B1A900EA99" box="[408,567,958,981]" country="Madagascar" latitude="-24.76389" longLatPrecision="21" longitude="46.751667" municipality="Parc National d'Andohahela" name="Col du Sedro" pageId="38" pageNumber="39" stateProvince="Toliara Province">Col du Sedro</location>
</collectorName>
,
<locationDeviation id="628EFB071862FFFC450132B1A8ACEA99" box="[590,923,958,981]" location="Mahamavo" pageId="38" pageNumber="39">
<quantity id="B47B4FC01862FFFC450132B1A996EA99" box="[590,673,958,981]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="38" pageNumber="39" unit="km" value="3.8">3.8 km</quantity>
113°ESE Mahamavo
</locationDeviation>
,
<locationDeviation id="628EFB071862FFFC44FE32B1ABDFEAB9" location="Tolagnaro" pageId="38" pageNumber="39">
<quantity id="B47B4FC01862FFFC44FE32B1AF25EA99" box="[945,1042,958,981]" metricMagnitude="4" metricUnit="m" metricValue="3.76" pageId="38" pageNumber="39" unit="km" value="37.6">37.6 km</quantity>
341°NNW Tolagnaro
</locationDeviation>
,
<geoCoordinate id="16B784E21862FFFC47B532D1AA43EAB9" box="[250,372,990,1013]" degrees="24" direction="south" minutes="45" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="50" value="-24.76389">24°45ʹ50ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21862FFFC46C732D1AACFEAB9" box="[392,504,990,1013]" degrees="46" direction="east" minutes="45" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="6" value="46.751667">46°45ʹ6ʺE</geoCoordinate>
, montane rainforest at
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</quantity>
elevation, general collecting ground spiders,
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<collectingDate id="17793D0D1862FFFC461F32F1A962ED59" box="[336,597,1022,1045]" pageId="38" pageNumber="39" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">21–25 January 2002</collectingDate>
</date>
,
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(
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,
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,
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,
<collectionCode id="15927AE01862FFFC46313511AA81ED79" box="[382,438,1054,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="39" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251862FFFC47E43530ABC7ED3A" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43530AA69ED1A" bold="true" box="[171,350,1087,1110]" pageId="38" pageNumber="39">Natural history.</emphasis>
Collection data suggest that the species
<taxonomicName id="B48399A61862FFFC44423530A8FBED1A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[781,972,1087,1110]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371862FFFC44423530A8FBED1A" bold="true" box="[781,972,1087,1110]" italics="true" pageId="38" pageNumber="39">Uduba goodmani</emphasis>
</taxonomicName>
occurs in montane rain forest.
</paragraph>
</subSubSection>
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<paragraph id="733CE2251862FFFC47E43570A9D8EDFA" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43570A931EDDA" bold="true" box="[171,518,1151,1174]" pageId="38" pageNumber="39">
Distribution.
<taxonomicName id="B48399A61862FFFC460A3570A931EDDA" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[325,518,1151,1174]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371862FFFC460A3570A931EDDA" bold="true" box="[325,518,1151,1174]" italics="true" pageId="38" pageNumber="39">Uduba goodmani</emphasis>
</taxonomicName>
</emphasis>
are known only from two nearby localities on mountains at Andohahela in far southeastern
<collectingCountry id="0B94A2B51862FFFC469C3590A963EDFA" box="[467,596,1183,1206]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
(Maps 6, 16
<emphasis id="41F73E371862FFFC45903590A9DFEDFA" bold="true" box="[735,744,1183,1206]" pageId="38" pageNumber="39">)</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

278
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<mods:title id="078AF7B06D405EEA61B5B498B2FDC81A">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="56805406F2E3FB66690611D7311EF4DA">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<emphasis id="41F73E371862FFFC473435D0AA88EDBA" bold="true" box="[123,447,1247,1270]" pageId="38" pageNumber="39">
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<emphasis id="41F73E371862FFFC473435D0AA1BEDBA" bold="true" box="[123,300,1247,1270]" italics="true" pageId="38" pageNumber="39">Uduba hainteny</emphasis>
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,
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<paragraph id="733CE2251862FFFC473435F0AAA4EC58" blockId="38.[123,447,1247,1300]" box="[123,403,1279,1300]" pageId="38" pageNumber="39">
<figureCitation id="EBB8FEA01862FFFC473435F0AA21EC58" box="[123,278,1279,1300]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="38" pageNumber="39">Figures 42 A–C</figureCitation>
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<figureCitation id="EBB8FEA01862FFFC466F35F0AA5FEC58" box="[288,360,1279,1300]" captionStart="MAP" captionStartId="176.[155,174,1068,1087]" captionTargetBox="[138,1176,216,1047]" captionTargetId="figure-26@176.[648,1176,216,1056]" captionTargetPageId="176" captionText="MAP 1. Distributions of Uduba jayjay, Uduba ibonia MAP 2. Distributions of Uduba salegy, Uduba platnicki, and Uduba hainteny. Uduba halabe, Uduba funerea Simon 1906 and Uduba dahli (Simon 1903)." figureDoi="http://doi.org/10.5281/zenodo.13158773" httpUri="https://zenodo.org/record/13158773/files/figure.png" pageId="38" pageNumber="39">Maps 1</figureCitation>
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<figureCitation id="EBB8FEA01862FFFC463A35F0AAB8EC58" box="[373,399,1279,1300]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="38" pageNumber="39">13</figureCitation>
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</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E4342AA9AEEC30" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="733CE2251862FFFC47E4342AA9AEEC30" blockId="38.[123,1174,1317,1725]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E4342AAA64EC70" bold="true" box="[171,339,1317,1340]" pageId="38" pageNumber="39">Type material.</emphasis>
<materialsCitation id="C3EBE8781862FFFC4611342AA9AEEC30" collectingDate="2006-12-02" collectingDateMax="2006-12-04" collectingDateMin="2006-12-02" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1000" location="Foret Ivohibe" pageId="38" pageNumber="39" specimenCode="CASENT9032857" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C871862FFFC4611342AAAF6EC70" box="[350,449,1317,1340]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1862FFFC455C342AA9EBEC70" box="[531,732,1317,1340]" collectionCode="CASENT" pageId="38" pageNumber="39">CASENT9032857</specimenCode>
) from rainforest at
<quantity id="B47B4FC01862FFFC44F1342AAF3FEC70" box="[958,1032,1317,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">
<elevation id="F8AE05161862FFFC44F1342AAF3FEC70" box="[958,1032,1317,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">1000m</elevation>
</quantity>
elevation in
<location id="765CB4FE1862FFFC4734344AAA27EC10" LSID="urn:lsid:plazi:treatment:FB2A53331862FFFD473435D0AAA1EC38:765CB4FE1862FFFC4734344AAA27EC10" box="[123,272,1349,1372]" country="Madagascar" name="Foret Ivohibe" pageId="38" pageNumber="39" stateProvince="Toliara">Forêt Ivohibe</location>
, 24˚34’08ʺS, 047˚12’14ʺE,
<collectingRegion id="B1472CC71862FFFC4570344AA9C4EC10" box="[575,755,1349,1372]" country="Madagascar" name="Toliara" pageId="38" pageNumber="39">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51862FFFC45B0344AA849EC10" box="[767,894,1349,1372]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
, collected
<date id="073DC4E51862FFFC44B8344AAB87EC30" pageId="38" pageNumber="39" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">
<collectingDate id="17793D0D1862FFFC44B8344AAB87EC30" pageId="38" pageNumber="39" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">2–4 December 2006</collectingDate>
</date>
by
<collectorName id="DE7687F31862FFFC4794346AAA6AEC30" box="[219,349,1381,1404]" pageId="38" pageNumber="39">B. L. Fisher</collectorName>
<emphasis id="41F73E371862FFFC462A346AAA95EC30" box="[357,418,1381,1404]" italics="true" pageId="38" pageNumber="39">et al.,</emphasis>
deposited in the
<collectionCode id="15927AE01862FFFC4512346AA9A3EC30" box="[605,660,1381,1404]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="39" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E4348AA924ECF0" pageId="38" pageNumber="39" type="etymology">
<paragraph id="733CE2251862FFFC47E4348AA924ECF0" blockId="38.[123,1174,1317,1725]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E4348AAA1DECD0" bold="true" box="[171,298,1413,1436]" pageId="38" pageNumber="39">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E371862FFFC4542348AA95AECD0" box="[525,621,1413,1436]" italics="true" pageId="38" pageNumber="39">hainteny,</emphasis>
refers to a traditional form of Malagasy oral literature and poetry; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E434CAA9EEEFF1" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="733CE2251862FFFC47E434CAA9EEEFF1" blockId="38.[123,1174,1317,1725]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E434CAAAE0EC90" bold="true" box="[171,471,1477,1500]" pageId="38" pageNumber="39">
Diagnosis.
<taxonomicName id="B48399A61862FFFC466934CAAAE0EC90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,471,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371862FFFC466934CAAAE0EC90" bold="true" box="[294,471,1477,1500]" italics="true" pageId="38" pageNumber="39">Uduba hainteny</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.a, the
<taxonomicName id="B48399A61862FFFC45AB34CAA850EC90" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[740,871,1477,1500]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA01862FFFC44F534CAAF38EC90" box="[954,1039,1477,1500]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="38" pageNumber="39">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate with a colulus; male palp (
<figureCitation id="EBB8FEA01862FFFC44DD34EAAF29ECB0" box="[914,1054,1509,1532]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="38" pageNumber="39">Figs. 42 A–C</figureCitation>
<emphasis id="41F73E371862FFFC435134EAAF10ECB0" bold="true" box="[1054,1063,1509,1532]" pageId="38" pageNumber="39">)</emphasis>
with apex of TA3 a slender, pointed screw, TA2 a large, elongate oval blade, visible behind and beyond TA3, apex rounded; MA simple with small prolateral hook. The species
<taxonomicName id="B48399A61862FFFC440C3729A8C3EF71" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[835,1012,1574,1597]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371862FFFC440C3729A8C3EF71" bold="true" box="[835,1012,1574,1597]" italics="true" pageId="38" pageNumber="39">Uduba hainteny</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="B48399A61862FFFC47D83749AA18EF11" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[151,303,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="ibonia">
<emphasis id="41F73E371862FFFC47D83749AA18EF11" bold="true" box="[151,303,1606,1629]" italics="true" pageId="38" pageNumber="39">Uduba ibonia</emphasis>
</taxonomicName>
except that the latter has a divided cribellum (
<figureCitation id="EBB8FEA01862FFFC446B3749A8BAEF11" box="[804,909,1606,1629]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="38" pageNumber="39">Fig. 20 B</figureCitation>
). The large and conspicuous TA2 (
<figureCitation id="EBB8FEA01862FFFC47BB3769AA68EF31" box="[244,351,1638,1661]" captionStart="FIGURE" captionStartId="139.[155,167,1219,1238]" captionTargetBox="[120,1176,216,1202]" captionTargetId="figure-16@139.[120,1176,216,1202]" captionTargetPageId="139" captionText="FIGURE 41. Uduba fisheri (CASENT9026389), from 3 km 41°NE Andranomay near Anjozorobe, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, prolateral. D. Bulb, retrolateral. E. Bulb, ventral. F. Bulb, prolateral. C, conductor; E, embolus; Eb, embolic base; MA, median apophysis; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis. Arrows to bump on tegulum. Scale bars A–C = 100 µm, D = 40 µm, E = 50 µm, F = 60 µm." figureDoi="http://doi.org/10.5281/zenodo.13158661" httpUri="https://zenodo.org/record/13158661/files/figure.png" pageId="38" pageNumber="39">Fig. 41 B</figureCitation>
) distinguishes
<taxonomicName id="B48399A61862FFFC45483769A9BDEF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[519,650,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371862FFFC45483769A9BDEF31" bold="true" box="[519,650,1638,1661]" italics="true" pageId="38" pageNumber="39">U. hainteny</emphasis>
</taxonomicName>
from
<taxonomicName id="B48399A61862FFFC459D3769A85DEF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[722,874,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371862FFFC459D3769A85DEF31" bold="true" box="[722,874,1638,1661]" italics="true" pageId="38" pageNumber="39">U. fandroana</emphasis>
</taxonomicName>
, which has a small, inconspicuous TA2 (
<figureCitation id="EBB8FEA01862FFFC46553789AA9CEFD1" box="[282,427,1670,1693]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="38" pageNumber="39">Figs. 39 A, B</figureCitation>
). The small MA distinguishes
<taxonomicName id="B48399A61862FFFC45BD3789A844EFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[754,883,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371862FFFC45BD3789A844EFD1" bold="true" box="[754,883,1670,1693]" italics="true" pageId="38" pageNumber="39">U. hainteny</emphasis>
</taxonomicName>
from
<taxonomicName id="B48399A61862FFFC44F93789AF2CEFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[950,1051,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lamba">
<emphasis id="41F73E371862FFFC44F93789AF2CEFD1" bold="true" box="[950,1051,1670,1693]" italics="true" pageId="38" pageNumber="39">U. lamba</emphasis>
</taxonomicName>
, which has a large, complex MA (
<figureCitation id="EBB8FEA01862FFFC462237A9A934EFF1" box="[365,515,1702,1725]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="38" pageNumber="39">Figs. 49 B, C</figureCitation>
). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1863FFFD47E431D3A8E9EDDF" pageId="39" pageNumber="40" type="description">
<paragraph id="733CE2251863FFFD47E431D3AFA2E9BF" blockId="39.[123,1175,220,1396]" box="[171,1173,220,243]" pageId="39" pageNumber="40">
<emphasis id="41F73E371863FFFD47E431D3AA01E9BF" bold="true" box="[171,310,220,243]" pageId="39" pageNumber="40">Description.</emphasis>
Male (
<typeStatus id="AC385C871863FFFD46C531D3AAC6E9BF" box="[394,497,220,243]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
, CASENT9032857): Total length 9.50. Markings typical for
</paragraph>
<paragraph id="733CE2251863FFFD473431F3A995ED3F" blockId="39.[123,1175,220,1396]" pageId="39" pageNumber="40">
<taxonomicName id="B48399A61863FFFD473431F3ABF2E85F" authorityName="Simon" authorityYear="1880" box="[123,197,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371863FFFD473431F3ABF2E85F" bold="true" box="[123,197,252,275]" italics="true" pageId="39" pageNumber="40">Uduba</emphasis>
</taxonomicName>
. Carapace 5.90 long, 4.30 wide, 1.79 high; clypeus 0.25 high. Eye diameters: AME and ALE 0.22, PME 0.25, PLE 0.26. Chelicerae 2.40 long; sternum 2.60 long, 2.10 wide; labium 1.40 long; palpal coxae 2.00 long.
<emphasis id="41F73E371863FFFD468A3033A92DE81F" bold="true" box="[453,538,316,339]" pageId="39" pageNumber="40">Ratios–</emphasis>
carapace length / width = 1.37, carapace height / width = 0.53, PER / carapace width = 0.37, PER / OAL = 2.86, PER / AER = 1.38, OAL / OQL = 0.93, OQP / OQA = 1.15, clypeus height / AME = 1.36, cheliceral length / clypeus height = 8.00, sternum length / width = 1.24, palpal coxa length / width = 2.50, femur I length / carapace width = 1.40, metatarsus I length / carapace width = 1.56, femur IV length / carapace width = 1.44, cymbium length / carapace width = 0.65, cymbium length / palpal patella length = 2.80, cymbium length / palpal tibia length = 2.80, cymbium length / palpal femur length = 1.40, palpal tibia length / palpal patella length = 1.00. With a colulus. Spination: palpus–femur d0-1-0-1, p0-0-1, r0-0-1; leg I–femur d1-1-0-1, p0-1-1-0, r0-1-1-0, tibia d0-0-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1- 2-1, v2-1-1, r1-1-1; leg II–femur d1-1-0-1, p0-1-1-0, r0-1-1-0, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2-2, r1-0-1-0; leg III–femur d1-1-0-1, p0-1-0-1-1, r0-1-0-1-1, patella r1, tibia d0-0-1-0, p0-1-1-0, v2-0-2, r0-1-1-0, metatarsus d0-1-0-0, p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-1, p0-1-0-1, r0-0-0-1, tibia d0-0-1-0, p0-1-1-0, v2-1-2, r0-1-1-0, metatarsus, p1-1-1-2, v2-1-2-2, r1-1-1-2. Scopulae: cymbium, apicodorsal; dense beneath tarsi I–IV. Leg measurements: I: 6.00 + 2.30 + 6.20 + 6.70 + 3.50 = 24.70; II: 5.30 + 2.30 + 5.30 + 5.80 + 3.00 = 21.70; III: 4.60 + 1.70 + 3.30 + 5.00 + 2.50 = 17.10; IV: 6.20 + 2.10 + 5.90 + 8.00 + 3.40 = 25.60; palpus: 2.00 + 1.00 + 1.00 + NA + 2.80 = 6.80. Leg formula 4123.
<emphasis id="41F73E371863FFFD44633233A8A8EA1F" bold="true" box="[812,927,828,851]" pageId="39" pageNumber="40">Male palp</emphasis>
(
<figureCitation id="EBB8FEA01863FFFD44E03233AF75EA1F" box="[943,1090,828,851]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="39" pageNumber="40">Figs. 42 A–C</figureCitation>
): palpal tibia 0.39 times cymbial length, RTA a triangle, length 0.20 tibia length, length equal to width, with pointed apex (
<figureCitation id="EBB8FEA01863FFFD465A3273AA49EADF" box="[277,382,892,915]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="39" pageNumber="40">Fig. 42 C</figureCitation>
), VTA conical, with broad base and slender apex, length 0.36 tibia width, VTA length 1.71 times width; tibia lacking stout spines; tegulum length 0.87 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.425 times tegulum length; TA3 a slender, pointed screw, TA3 ends before TA2 apex, TA2 a large, elongate oval blade (
<figureCitation id="EBB8FEA01863FFFD435332D3AFBFEABF" box="[1052,1160,988,1011]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="39" pageNumber="40">Fig. 42 B</figureCitation>
), visible behind and beyond TA3, apex rounded; MA simple with small prolateral hook (
<figureCitation id="EBB8FEA01863FFFD435032F3AFBFED5F" box="[1055,1160,1020,1043]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="39" pageNumber="40">Fig. 42 C</figureCitation>
), origin at 0.42 times tegulum length, tegulum apex at 0.44 tegulum length from MA apex, length (including apical processes) 0.625 times width, MA small, length 0.21 tegulum length, width 0.29 tegulum width; conductor fan concave (
<figureCitation id="EBB8FEA01863FFFD45653553A9A3ED3F" box="[554,660,1116,1139]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="39" pageNumber="40">Fig. 42 B</figureCitation>
).
</paragraph>
<paragraph id="733CE2251863FFFD47E43573A8E9EDDF" blockId="39.[123,1175,220,1396]" box="[171,990,1148,1171]" pageId="39" pageNumber="40">
<emphasis id="41F73E371863FFFD47E43573AA2DEDDF" bold="true" box="[171,282,1148,1171]" pageId="39" pageNumber="40">Variation.</emphasis>
Unknown, only the male
<typeStatus id="AC385C871863FFFD45793573A9A4EDDF" box="[566,659,1148,1171]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B48399A61863FFFD45F63573A85BEDDF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[697,876,1148,1171]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371863FFFD45F63573A85BEDDF" bold="true" box="[697,876,1148,1171]" italics="true" pageId="39" pageNumber="40">Uduba hainteny</emphasis>
</taxonomicName>
is known.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1863FFFD47E43593A803EC78" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="733CE2251863FFFD47E43593A843EDB8" blockId="39.[123,1175,220,1396]" pageId="39" pageNumber="40">
<emphasis id="41F73E371863FFFD47E43593AAA0EDFF" bold="true" box="[171,407,1180,1203]" pageId="39" pageNumber="40">Material examined.</emphasis>
<materialsCitation id="C3EBE8781863FFFD46E73593A847EDB8" collectingDate="2006-12-02" collectingDateMax="2006-12-04" collectingDateMin="2006-12-02" collectionCode="CAS" collectorName="N Tolagnaro &amp; B. L. Fisher" country="Madagascar" elevation="200" location="Foret Ivohibe" pageId="39" pageNumber="40" specimenCode="CASENT9032857" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="0B94A2B51863FFFD46E73593A95FEDFF" box="[424,616,1180,1203]" name="Madagascar" pageId="39" pageNumber="40">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371863FFFD45CF3593A876EDFF" box="[640,833,1180,1203]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="B1472CC71863FFFD45CF3593A876EDFF" box="[640,833,1180,1203]" country="Madagascar" name="Toliara" pageId="39" pageNumber="40">Toliara Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1863FFFD44153593A8C0EDFF" LSID="urn:lsid:plazi:treatment:FB2A53331862FFFD473435D0AAA1EC38:765CB4FE1863FFFD44153593A8C0EDFF" box="[858,1015,1180,1203]" country="Madagascar" name="Foret Ivohibe" pageId="39" pageNumber="40" stateProvince="Toliara">Forêt Ivohibe</location>
, 55.0 km
<collectorName id="DE7687F31863FFFD43CD3593ABDFED9F" pageId="39" pageNumber="40">N Tolagnaro</collectorName>
, 24˚34ʹ08ʺS, 047˚12ʹ14ʺE, elev.
<quantity id="B47B4FC01863FFFD450B35B3A9B3ED9F" box="[580,644,1212,1235]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="200.0">
<elevation id="F8AE05161863FFFD450B35B3A9B3ED9F" box="[580,644,1212,1235]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="200.0">200m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321863FFFD45C035B3A823ED9F" box="[655,788,1212,1235]" pageId="39" pageNumber="40">malaise trap</collectingMethod>
, rainforest,
<date id="073DC4E51863FFFD44DA35B3AF45ED9F" box="[917,1138,1212,1235]" pageId="39" pageNumber="40" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">
<collectingDate id="17793D0D1863FFFD44DA35B3AF45ED9F" box="[917,1138,1212,1235]" pageId="39" pageNumber="40" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">2–4 December 2006</collectingDate>
</date>
,
<collectorName id="DE7687F31863FFFD433235B3ABEAEDB8" pageId="39" pageNumber="40">B. L. Fisher</collectorName>
<emphasis id="41F73E371863FFFD47AB35D2AA22EDB8" box="[228,277,1245,1268]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
., [BLF15448] (
<typeStatus id="AC385C871863FFFD46F035D2A916EDB8" box="[447,545,1245,1268]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E1863FFFD456335D2A9C3EDB8" box="[556,756,1245,1268]" collectionCode="CASENT" pageId="39" pageNumber="40">CASENT9032857</specimenCode>
,
<specimenCount id="658529AC1863FFFD444F35D2A816EDB8" box="[768,801,1245,1268]" count="1" pageId="39" pageNumber="40" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01863FFFD447F35D2A85FEDB8" box="[816,872,1245,1268]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="40" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251863FFFD47E435F2A803EC78" blockId="39.[123,1175,220,1396]" pageId="39" pageNumber="40">
<emphasis id="41F73E371863FFFD47E435F2AA57EC58" bold="true" box="[171,352,1277,1300]" pageId="39" pageNumber="40">Natural history.</emphasis>
The
<typeStatus id="AC385C871863FFFD46D635F2AAC1EC58" box="[409,502,1277,1300]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="B48399A61863FFFD451635F2A83CEC58" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[601,779,1277,1300]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371863FFFD451635F2A83CEC58" bold="true" box="[601,779,1277,1300]" italics="true" pageId="39" pageNumber="40">Uduba hainteny</emphasis>
</taxonomicName>
was collected in lowland rainforest. Retrieval from a malaise trap reveals that the spiders will climb.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1863FFFD47E43432AAA1EC38" pageId="39" pageNumber="40" type="distribution">
<paragraph id="733CE2251863FFFD47E43432AAA1EC38" blockId="39.[123,1175,220,1396]" pageId="39" pageNumber="40">
<emphasis id="41F73E371863FFFD47E43432AA0AEC18" bold="true" box="[171,317,1341,1364]" pageId="39" pageNumber="40">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61863FFFD46833432A949EC18" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[460,638,1341,1364]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371863FFFD46833432A949EC18" bold="true" box="[460,638,1341,1364]" italics="true" pageId="39" pageNumber="40">Uduba hainteny</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C871863FFFD44C13432A88BEC18" box="[910,956,1341,1364]" pageId="39" pageNumber="40">type</typeStatus>
locality in southern
<collectingCountry id="0B94A2B51863FFFD47343452ABCBEC38" box="[123,252,1373,1396]" name="Madagascar" pageId="39" pageNumber="40">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA01863FFFD46443452AA6AEC38" box="[267,349,1373,1396]" captionStart="MAP" captionStartId="176.[155,174,1068,1087]" captionTargetBox="[138,1176,216,1047]" captionTargetId="figure-26@176.[648,1176,216,1056]" captionTargetPageId="176" captionText="MAP 1. Distributions of Uduba jayjay, Uduba ibonia MAP 2. Distributions of Uduba salegy, Uduba platnicki, and Uduba hainteny. Uduba halabe, Uduba funerea Simon 1906 and Uduba dahli (Simon 1903)." figureDoi="http://doi.org/10.5281/zenodo.13158773" httpUri="https://zenodo.org/record/13158773/files/figure.png" pageId="39" pageNumber="40">Maps 1</figureCitation>
,
<figureCitation id="EBB8FEA01863FFFD46233452AAB0EC38" box="[364,391,1373,1396]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="39" pageNumber="40">13</figureCitation>
<emphasis id="41F73E371863FFFD46C93452AAB8EC38" bold="true" box="[390,399,1373,1396]" pageId="39" pageNumber="40">)</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E7F7404238AE196B4A26734BC7C57E0E">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="BA84E0823090986D7C64CB3867E602A6">Ubick, Darrell</mods:namePart>
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<heading id="287455491864FFFA473430D4AAE5E8BE" box="[123,466,475,498]" fontSize="10" level="2" pageId="32" pageNumber="33" reason="4">
<emphasis id="41F73E371864FFFA473430D4AAE5E8BE" bold="true" box="[123,466,475,498]" pageId="32" pageNumber="33">
<taxonomicName id="B48399A61864FFFA473430D4AA77E8BE" ID-CoL="C8KJ7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,320,475,498]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fandroana" status="sp. nov.">
<emphasis id="41F73E371864FFFA473430D4AA77E8BE" bold="true" box="[123,320,475,498]" italics="true" pageId="32" pageNumber="33">Uduba fandroana</emphasis>
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,
<taxonomicNameLabel id="5AC4834C1864FFFA460130D4AAE5E8BE" box="[334,466,475,498]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="3B99B1AE1864FFFA473430F4AAFDEB5C" box="[123,458,507,528]" pageId="32" pageNumber="33" type="description">
<paragraph id="733CE2251864FFFA473430F4AAFDEB5C" blockId="32.[123,466,475,528]" box="[123,458,507,528]" pageId="32" pageNumber="33">
<figureCitation id="EBB8FEA01864FFFA473430F4ABDDEB5C" box="[123,234,507,528]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="32" pageNumber="33">Figures 9 F</figureCitation>
,
<figureCitation id="EBB8FEA01864FFFA47BB30F4AA76EB5C" box="[244,321,507,528]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="32" pageNumber="33">39 A–C</figureCitation>
, Maps 12, 13.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1864FFFA47E4332BAF00EBD7" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="733CE2251864FFFA47E4332BAF00EBD7" blockId="32.[122,1174,548,1725]" pageId="32" pageNumber="33">
<emphasis id="41F73E371864FFFA47E4332BAA78EB77" bold="true" box="[171,335,548,571]" pageId="32" pageNumber="33">Type material.</emphasis>
<materialsCitation id="C3EBE8781864FFFA4618332BAA52EBD7" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" country="Madagascar" elevation="1300" latitude="-18.473333" location="Fisher-Griswold Arthropod Team" longLatPrecision="21" longitude="47.96" pageId="32" pageNumber="33" specimenCode="CASENT9002691, BLF2370" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="AC385C871864FFFA4618332BAA8DEB77" box="[343,442,548,571]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1864FFFA454B332BA9FBEB77" box="[516,716,548,571]" collectionCode="CASENT" pageId="32" pageNumber="33">CASENT9002691</specimenCode>
) from montane rainforest at
<quantity id="B47B4FC01864FFFA434C332BAF7BEB77" box="[1027,1100,548,571]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="32" pageNumber="33" unit="m" value="1300.0">
<elevation id="F8AE05161864FFFA434C332BAF7BEB77" box="[1027,1100,548,571]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="32" pageNumber="33" unit="m" value="1300.0">1300m</elevation>
</quantity>
elevation at
<locationDeviation id="628EFB071864FFFA4784334BA923EB17" box="[203,532,580,603]" location="Anjozorobe" pageId="32" pageNumber="33">
<quantity id="B47B4FC01864FFFA4784334BAA16EB17" box="[203,289,580,603]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="32" pageNumber="33" unit="km" value="11.5">11.5 km</quantity>
147°SSE Anjozorobe
</locationDeviation>
, (
<geoCoordinate id="16B784E21864FFFA4565334BA992EB17" box="[554,677,580,603]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="32" pageNumber="33" precision="15" seconds="24" value="-18.473333">18°28ʹ24ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21864FFFA45FB334BA806EB17" box="[692,817,580,603]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="32" pageNumber="33" precision="15" seconds="36" value="47.96">47°57ʹ36ʺE</geoCoordinate>
),
<collectingRegion id="B1472CC71864FFFA440A334BAF75EB17" box="[837,1090,580,603]" country="Madagascar" name="Antananarivo" pageId="32" pageNumber="33">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51864FFFA431F334BAB8AEB37" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
, collected
<date id="073DC4E51864FFFA460F336BA903EB37" box="[320,564,612,635]" pageId="32" pageNumber="33" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="17793D0D1864FFFA460F336BA903EB37" box="[320,564,612,635]" pageId="32" pageNumber="33" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5–13 December 2000</collectingDate>
</date>
by the
<location id="765CB4FE1864FFFA45D5336BAF3FEB37" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1864FFFA45D5336BAF3FEB37" box="[666,1032,612,635]" country="Madagascar" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Fisher-Griswold Arthropod Team" pageId="32" pageNumber="33" stateProvince="Antananarivo">Fisher-Griswold Arthropod Team</location>
(
<specimenCode id="23254A5E1864FFFA4351336BAFBEEB37" box="[1054,1161,612,635]" collectionCode="BLF" pageId="32" pageNumber="33">BLF2370</specimenCode>
), deposited in the
<collectionCode id="15927AE01864FFFA4661338BAA52EBD7" box="[302,357,644,667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="33" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781864FFFA463E338BAF04EBD7" box="[369,1075,644,667]" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" country="Madagascar" elevation="1300" latitude="-18.473333" location="Fisher-Griswold Arthropod Team" longLatPrecision="21" longitude="47.96" pageId="32" pageNumber="33" specimenCode="CASENT9064784" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="AC385C871864FFFA463E338BAAF9EBD7" box="[369,462,644,667]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
male (
<specimenCode id="23254A5E1864FFFA4556338BA9D3EBD7" box="[537,740,644,667]" collectionCode="CASENT" pageId="32" pageNumber="33">CASENT9064784</specimenCode>
), same data, deposited in
<collectionCode id="15927AE01864FFFA44B3338BAF04EBD7" box="[1020,1075,644,667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="33" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1864FFFA47E433ABAA87EB97" pageId="32" pageNumber="33" type="etymology">
<paragraph id="733CE2251864FFFA47E433ABAA87EB97" blockId="32.[122,1174,548,1725]" pageId="32" pageNumber="33">
<emphasis id="41F73E371864FFFA47E433ABAA1DEBF7" bold="true" box="[171,298,676,699]" pageId="32" pageNumber="33">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E371864FFFA454033ABA9B4EBF7" box="[527,643,676,699]" italics="true" pageId="32" pageNumber="33">fandroana,</emphasis>
refers to an annual New Year’s festival in
<collectingCountry id="0B94A2B51864FFFA431F33ABAB8AEB97" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
, a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1864FFFA47E433EBA91EED50" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="733CE2251864FFFA47E433EBA91EED50" blockId="32.[122,1174,548,1725]" pageId="32" pageNumber="33">
<emphasis id="41F73E371864FFFA47E433EBAADBEBB7" bold="true" box="[171,492,740,763]" pageId="32" pageNumber="33">
Diagnosis.
<taxonomicName id="B48399A61864FFFA466833EBAADBEBB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,492,740,763]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371864FFFA466833EBAADBEBB7" bold="true" box="[295,492,740,763]" italics="true" pageId="32" pageNumber="33">Uduba fandroana</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.a, the
<taxonomicName id="B48399A61864FFFA440E33EBA8F2EBB7" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[833,965,740,763]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA01864FFFA435533EBAF58EBB7" box="[1050,1135,740,763]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="32" pageNumber="33">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate, with a colulus. Males have the palpus (
<figureCitation id="EBB8FEA01864FFFA47CC322BAA25EA77" box="[131,274,804,827]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="32" pageNumber="33">Figs. 39 A–C</figureCitation>
) with the MA relatively small, width less than 0.35 tegulum width, height less than 0.25 tegulum height, without prominent projections, TA2 small, inconspicuous, TA2 a small, slen- der, sharp blade, pointing distally and hidden behind TA3, TA3 large, strongly curved (
<figureCitation id="EBB8FEA01864FFFA4356326BABA7EAD7" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="32" pageNumber="33">Figs. 39 A– C</figureCitation>
); males of
<taxonomicName id="B48399A61864FFFA465F328BAAEFEAD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[272,472,900,923]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371864FFFA465F328BAAEFEAD7" bold="true" box="[272,472,900,923]" italics="true" pageId="32" pageNumber="33">Uduba fandroana</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="B48399A61864FFFA45AC328BA87CEAD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[739,843,900,923]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lamba">
<emphasis id="41F73E371864FFFA45AC328BA87CEAD7" bold="true" box="[739,843,900,923]" italics="true" pageId="32" pageNumber="33">U. lamba</emphasis>
</taxonomicName>
by the small, simple MA (
<taxonomicName id="B48399A61864FFFA4334328BAB88EAF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lamba">
<emphasis id="41F73E371864FFFA4334328BAB88EAF7" bold="true" italics="true" pageId="32" pageNumber="33">U. lamba</emphasis>
</taxonomicName>
have the MA large, concave in center with strong marginal ridge [
<figureCitation id="EBB8FEA01864FFFA44DE32ABAF29EAF7" box="[913,1054,932,955]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="32" pageNumber="33">Fig. 49 A–C</figureCitation>
]); males of
<taxonomicName id="B48399A61864FFFA473432CAAA26EA90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,273,965,988]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371864FFFA473432CAAA26EA90" bold="true" box="[123,273,965,988]" italics="true" pageId="32" pageNumber="33">U. fandroana</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="B48399A61864FFFA455A32CAA9A0EA90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[533,663,965,988]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371864FFFA455A32CAA9A0EA90" bold="true" box="[533,663,965,988]" italics="true" pageId="32" pageNumber="33">U. hainteny</emphasis>
</taxonomicName>
by the small, inconspicuous TA2 (
<taxonomicName id="B48399A61864FFFA435C32CAAFA2EA90" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1043,1173,965,988]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E371864FFFA435C32CAAFA2EA90" bold="true" box="[1043,1173,965,988]" italics="true" pageId="32" pageNumber="33">U. hainteny</emphasis>
</taxonomicName>
have the TA2 large, conspicuous and with the apex a rounded blade, TA2 equal to or larger than TA3 [
<figureCitation id="EBB8FEA01864FFFA47F9350AAA7AED50" box="[182,333,1029,1052]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="32" pageNumber="33">Figs. 42 A–C</figureCitation>
]). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1864FFFB47E4352AA8DCEB7F" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="733CE2251864FFFB47E4352AAF75E85F" blockId="32.[122,1174,548,1725]" lastBlockId="33.[122,1174,220,1203]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="41F73E371864FFFA47E4352AAA01ED70" bold="true" box="[171,310,1061,1084]" pageId="32" pageNumber="33">Description.</emphasis>
Male (
<typeStatus id="AC385C871864FFFA46C7352AAAC5ED70" box="[392,498,1061,1084]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
): Total length 10.20. Markings as in
<figureCitation id="EBB8FEA01864FFFA44DE352AA8DFED70" box="[913,1000,1061,1084]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="32" pageNumber="33">
Fig.
<specimenCount id="658529AC1864FFFA448A352AA8DFED70" box="[965,1000,1061,1084]" count="9" pageId="32" pageNumber="33" type="generic">9 F</specimenCount>
</figureCitation>
. Carapace 5.80 long, 3.80 wide, 2.00 high; clypeus 0.20 high. Eye diameters:AME and ALE 0.22, PME 0.20, PLE 0.26. Chelicerae 2.50 long; sternum 2.50 long, 2.00 wide; labium 1.20 long; palpal coxae 1.90 long. With a colulus. Spination (
<typeStatus id="AC385C871864FFFA46E5358AA93DEDD0" box="[426,522,1157,1180]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
, CASENT9064784), palpus–femur d1-1-1, p0-0-1, r0-0-1, patella p0-1, tibia p1-0; leg I–femur d1-1-1, p0-1-2, r0-1-0-1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2-2-3, r1-1-0; leg II–femur d1-0-1-1, p1-0-2, r1-1-0-1, tibia d0-1-1- 0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2-2-3, r1-1-0; leg III–femur d1-1-1, p1-1-1-1, r0-1-1-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-1, p1-1-1-1, r0-0-1-1, patella r1, tibia d0-1-0, p0-1-1-0, v2-2-2, r10-0-1, metatarsus d1-1-1-0, p1-1-1-1, v2-2-2, r1-0-01. Scopulae: cymbium, apicodorsal; strong beneath all tarsi, half of metatarsus I and apices of metatarsi II-IV. Leg measurements (
<typeStatus id="AC385C871864FFFA448C346AAF10EC30" box="[963,1063,1381,1404]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
): I: 5.70 + 2.20 + 5.90 + 6.50 + 3.50 = 23.80; II: 5.30 + 2.00 + 5.00 + 5.50 + 3.10 = 20.90; III: 4.40 + 1.70 + 3.40 + 4.70 + 2.50 = 16.70; IV: 6.00 + 1.90 + 5.80 + 7.50 + 3.40 = 24.60; palpus: 1.90 + 0.80 + 0.90 + NA + 2.80 = 6.40. Leg formula 4123.
<emphasis id="41F73E371864FFFA45D534C9A827EC91" bold="true" box="[666,784,1478,1501]" pageId="32" pageNumber="33">Male palp</emphasis>
(
<typeStatus id="AC385C871864FFFA446C34C9A8B0EC91" box="[803,903,1478,1501]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
): palpal tibia 0.38 times cymbial length, RTA a broad blade, slightly curved ventrad, RTA length 0.32 times tibia, 1.20 times width, with pointed apex (
<figureCitation id="EBB8FEA01864FFFA46983709A95DEF51" box="[471,618,1542,1565]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="32" pageNumber="33">Figs. 39 A–C</figureCitation>
); VTA conical, VTA length 1.44 times width, length 0.35 times tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 1.02 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.18 times tegulum length; TA3 a broad, sharp-pointed screw with proximal ridges, extends apicad of TA2 by 1.17 tegulum length, TA2 a small, slender, sharp blade, pointing distally and hidden behind TA3 (
<figureCitation id="EBB8FEA01864FFFA47CC37A9ABDCEFF1" box="[131,235,1702,1725]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="32" pageNumber="33">Fig. 39 A</figureCitation>
); MA simple, origin at 0.35 times tegulum length, tegulum apex at 0.31 tegulum length from MA apex, length (including apical processes) 0.53 times width, MA small (
<figureCitation id="EBB8FEA01865FFFB449831D3AF09E9BF" box="[983,1086,220,243]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="33" pageNumber="34">Fig. 39 B</figureCitation>
), length 0.17 tegulum length, width 0.32 tegulum width; conductor fan entire.
<emphasis id="41F73E371865FFFB443F31F3A8F4E85F" bold="true" box="[880,963,252,275]" pageId="33" pageNumber="34">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="733CE2251865FFFB47E43013A8DCEB7F" blockId="33.[122,1174,220,1203]" pageId="33" pageNumber="34">
<emphasis id="41F73E371865FFFB47E43013AA2DE87F" bold="true" box="[171,282,284,307]" pageId="33" pageNumber="34">Variation.</emphasis>
(N = 4): Total length = 9.10–11.50, carapace length / width = 1.40–1.55, carapace height / width = 0.49–0.58, PER / carapace width = 0.38–0.41, PER / OAL = 2.61–3.00, PER / AER = 1.30–1.34, OAL / OQL = 1.00–1.20, OQP / OQA 1.09–1.20, clypeus height / AME = 0.91–1.67, cheliceral length / clypeus height = 7.75–12.50, sternum length / width = 1.21–1.33, palpal coxa length / width = 2.56–2.71, femur I length / carapace width =1.28–1.55, metatarsus I length / carapace width = 1.38–1.71, femur IV length / carapace width =1.34–1.58, cymbium length / carapace width = 0.65–0.74, cymbium length / palpal patella length = 3.13–3.50, cymbium length / palpal tibia length = 2.60–3.11, cymbium length / palpal femur length = 1.13–1.47, palpal tibia length / palpal patella length = 1.09–1.25. Only males known, female variation unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1865FFFB47E43333A812ED3F" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="733CE2251865FFFB47E43333AF20ED5F" blockId="33.[122,1174,220,1203]" pageId="33" pageNumber="34">
<emphasis id="41F73E371865FFFB47E43333AABAEB1F" bold="true" box="[171,397,572,595]" pageId="33" pageNumber="34">Material examined.</emphasis>
<materialsCitation id="C3EBE8781865FFFB46D83333A9BEEBFF" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="1300" latitude="-18.473333" location="Anjozorobe" longLatPrecision="21" longitude="47.96" pageId="33" pageNumber="34" specimenCode="CASENT9002691, CASENT9064784" specimenCount="2" specimenCount-male="2" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="0B94A2B51865FFFB46D83333A96EEB1F" box="[407,601,572,595]" name="Madagascar" pageId="33" pageNumber="34">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371865FFFB45283333A854EB1F" box="[615,867,572,595]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="B1472CC71865FFFB45283333A854EB1F" box="[615,867,572,595]" country="Madagascar" name="Antananarivo" pageId="33" pageNumber="34">Antananarivo Province</collectingRegion>
</emphasis>
:
<locationDeviation id="628EFB071865FFFB443A3333AFB8EB1F" box="[885,1167,572,595]" location="Andranomay" pageId="33" pageNumber="34">
<quantity id="B47B4FC01865FFFB443A3333A89BEB1F" box="[885,940,572,595]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="34" unit="km" value="3.0">3 km</quantity>
41°NE Andranomay
</locationDeviation>
,
<quantity id="B47B4FC01865FFFB47343353ABE6EB3F" box="[123,209,604,627]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="34" unit="km" value="11.5">11.5 km</quantity>
147°SSE
<location id="765CB4FE1865FFFB460F3353AAF6EB3F" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1865FFFB460F3353AAF6EB3F" box="[320,449,604,627]" country="Madagascar" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Anjozorobe" pageId="33" pageNumber="34" stateProvince="Antananarivo">Anjozorobe</location>
,
<geoCoordinate id="16B784E21865FFFB46823353A970EB3F" box="[461,583,604,627]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="33" pageNumber="34" precision="15" seconds="24" value="-18.473333">18°28ʹ24ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21865FFFB451A3353A9E5EB3F" box="[597,722,604,627]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="33" pageNumber="34" precision="15" seconds="36" value="47.96">47°57ʹ36ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01865FFFB44593353A854EB3F" box="[790,867,604,627]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="34" unit="m" value="1300.0">
<elevation id="F8AE05161865FFFB44593353A854EB3F" box="[790,867,604,627]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="34" unit="m" value="1300.0">1300m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321865FFFB443F3353A88DEB3F" box="[880,954,604,627]" pageId="33" pageNumber="34">pitfalls</collectingMethod>
in montane rainforest,
<date id="073DC4E51865FFFB47E83373AAA5EBDF" box="[167,402,636,659]" pageId="33" pageNumber="34" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="17793D0D1865FFFB47E83373AAA5EBDF" box="[167,402,636,659]" pageId="33" pageNumber="34" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5–13 December 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31865FFFB46D13373AAE8EBDF" box="[414,479,636,659]" pageId="33" pageNumber="34">Fisher</collectorName>
,
<collectorName id="DE7687F31865FFFB46A13373A966EBDF" box="[494,593,636,659]" pageId="33" pageNumber="34">Griswold</collectorName>
<emphasis id="41F73E371865FFFB45163373A9A0EBDF" box="[601,663,636,659]" italics="true" pageId="33" pageNumber="34">et al.,</emphasis>
[BLF2370] (
<typeStatus id="AC385C871865FFFB44693373A8BAEBDF" box="[806,909,636,659]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1865FFFB44D53373AF55EBDF" box="[922,1122,636,659]" collectionCode="CASENT" pageId="33" pageNumber="34">CASENT9002691</specimenCode>
,
<specimenCount id="658529AC1865FFFB43223373AFB8EBDF" box="[1133,1167,636,659]" count="1" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01865FFFB47343393AB85EBFF" box="[123,178,668,691]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="34" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C871865FFFB479F3393AA07EBFF" box="[208,304,668,691]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1865FFFB460F3393A93FEBFF" box="[320,520,668,691]" collectionCode="CASENT" pageId="33" pageNumber="34">CASENT9064784</specimenCode>
,
<specimenCount id="658529AC1865FFFB45593393A90FEBFF" box="[534,568,668,691]" count="1" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01865FFFB45053393A9B6EBFF" box="[586,641,668,691]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="34" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781865FFFB45D73393A8C2EBBF" collectingDate="1997-12-07" collectingDateMax="1997-12-12" collectingDateMin="1997-12-07" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1450" location="Reserve Speciale d'Ambohitantely" pageId="33" pageNumber="34" specimenCode="CASENT9006223" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo Province" typeStatus="holotype">
<location id="765CB4FE1865FFFB45D73393AF25EBFF" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1865FFFB45D73393AF25EBFF" box="[664,1042,668,691]" country="Madagascar" name="Reserve Speciale d'Ambohitantely" pageId="33" pageNumber="34" stateProvince="Antananarivo Province">Réserve Spéciale d’Ambohitantely</location>
,
<locationDeviation id="628EFB071865FFFB436C3393ABDDEB9F" location="Ankazobe" pageId="33" pageNumber="34">
<quantity id="B47B4FC01865FFFB436C3393AF50EBFF" box="[1059,1127,668,691]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="33" pageNumber="34" unit="km" value="24.0">24 km</quantity>
NE Ankazobe
</locationDeviation>
, 18º10.1ʹS, 47º16.6ʹE, elev.
<quantity id="B47B4FC01865FFFB455733B3A953EB9F" box="[536,612,700,723]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="33" pageNumber="34" unit="m" value="1450.0">
<elevation id="F8AE05161865FFFB455733B3A953EB9F" box="[536,612,700,723]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="33" pageNumber="34" unit="m" value="1450.0">1450m</elevation>
</quantity>
, disturbed transitional montane mossy forest,
<collectingMethod id="AAC29A321865FFFB431933B3AB98EBBF" pageId="33" pageNumber="34">pitfall traps</collectingMethod>
7–8,
<date id="073DC4E51865FFFB47BA33D3AADFEBBF" box="[245,488,732,755]" pageId="33" pageNumber="34" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">
<collectingDate id="17793D0D1865FFFB47BA33D3AADFEBBF" box="[245,488,732,755]" pageId="33" pageNumber="34" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">7–12 December 1997</collectingDate>
</date>
,
<collectorName id="DE7687F31865FFFB46B733D3A9B5EBBF" box="[504,642,732,755]" pageId="33" pageNumber="34">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1865FFFB45DA33D3A86BEBBF" box="[661,860,732,755]" collectionCode="CASENT" pageId="33" pageNumber="34">CASENT9006223</specimenCode>
<specimenCount id="658529AC1865FFFB442733D3A8BEEBBF" box="[872,905,732,755]" count="1" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01865FFFB44D333D3A8D9EBBF" box="[924,1006,732,755]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="33" pageNumber="34" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781865FFFB434B33D3AFA6EA1F" collectingDate="2009-11-10" collectingDateMax="2009-11-15" collectingDateMin="2009-11-10" collectionCode="CAS" collectorName="Harin'Hala Rinha &amp; M. Irwin" country="Madagascar" county="Fitovinany Region" elevation="1127" location="Radio Tower" municipality="District of Ifanadiana" pageId="33" pageNumber="34" specimenCode="CASENT9017919" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="41F73E371865FFFB434B33D3ABD5EA5F" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="B1472CC71865FFFB434B33D3ABEBEA5F" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="34">Fianarantsoa Province</collectingRegion>
:
</emphasis>
<collectingCounty id="9A5D9AA91865FFFB47A133F3AA88EA5F" box="[238,447,764,787]" pageId="33" pageNumber="34">Fitovinany Region</collectingCounty>
,
<collectingMunicipality id="9358785F1865FFFB468033F3A9F6EA5F" box="[463,705,764,787]" pageId="33" pageNumber="34">District of Ifanadiana</collectingMunicipality>
,
<locationDeviation id="628EFB071865FFFB459D33F3A8C3EA5F" box="[722,1012,764,787]" location="Ranomafana" pageId="33" pageNumber="34">
<quantity id="B47B4FC01865FFFB459D33F3A821EA5F" box="[722,790,764,787]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="km" value="12.0">12 km</quantity>
W of Ranomafana
</locationDeviation>
,
<location id="765CB4FE1865FFFB434B33F3AFB8EA5F" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1865FFFB434B33F3AFB8EA5F" box="[1028,1167,764,787]" country="Madagascar" county="Fitovinany Region" municipality="District of Ifanadiana" name="Radio Tower" pageId="33" pageNumber="34" stateProvince="Fianarantsoa">Radio Tower</location>
, 21º15.05ʹS, 047º24.43ʹE, elev.
<quantity id="B47B4FC01865FFFB46823213A92FEA7F" box="[461,536,796,819]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="33" pageNumber="34" unit="m" value="1127.0">
<elevation id="F8AE05161865FFFB46823213A92FEA7F" box="[461,536,796,819]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="33" pageNumber="34" unit="m" value="1127.0">1127m</elevation>
</quantity>
, malaise,
<collectingMethod id="AAC29A321865FFFB45C53213A83DEA7F" box="[650,778,796,819]" pageId="33" pageNumber="34">canopy trap</collectingMethod>
in open area at forest edge,
<date id="073DC4E51865FFFB431C3213AA1BEA1F" pageId="33" pageNumber="34" value="2009-11-10" valueMax="2009-11-15" valueMin="2009-11-10">
<collectingDate id="17793D0D1865FFFB431C3213AA1BEA1F" pageId="33" pageNumber="34" value="2009-11-10" valueMax="2009-11-15" valueMin="2009-11-10">10–15 November 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31865FFFB46773233AACFEA1F" box="[312,504,828,851]" pageId="33" pageNumber="34">Harin’Hala Rinha</collectorName>
and
<collectorName id="DE7687F31865FFFB45603233A9A7EA1F" box="[559,656,828,851]" pageId="33" pageNumber="34">M. Irwin</collectorName>
[MG-09B-267] (
<specimenCode id="23254A5E1865FFFB44043233AF23EA1F" box="[843,1044,828,851]" collectionCode="CASENT" pageId="33" pageNumber="34">CASENT9017919</specimenCode>
,
<specimenCount id="658529AC1865FFFB436E3233AF75EA1F" box="[1057,1090,828,851]" count="1" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01865FFFB431E3233AFBEEA1F" box="[1105,1161,828,851]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="34" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781865FFFB47343253A963EAFF" collectingDate="1998-01-07" collectingDateMax="1998-01-14" collectingDateMin="1998-01-07" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1670" location="Ambositra" municipality="Foret d'Ankazomivady" pageId="33" pageNumber="34" specimenCode="CASENT9006103" specimenCount="3" specimenCount-male="3" stateProvince="Fianarantsoa Province" typeStatus="holotype">
<collectingMunicipality id="9358785F1865FFFB47343253AA4CEA3F" box="[123,379,860,883]" pageId="33" pageNumber="34">Forêt d’Ankazomivady</collectingMunicipality>
,
<locationDeviation id="628EFB071865FFFB46DF3253A9DDEA3F" box="[400,746,860,883]" location="Ambalamanakana" pageId="33" pageNumber="34">
<quantity id="B47B4FC01865FFFB46DF3253AAD4EA3F" box="[400,483,860,883]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="33" pageNumber="34" unit="km" value="4.7">4.7 km</quantity>
SW Ambalamanakana
</locationDeviation>
,
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SSW
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, 20º46.5ʹS, 47º10.1ʹE, elev.
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</quantity>
,
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#1–3, disturbed upper montane forest,
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</date>
,
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(
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,
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,
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)
</materialsCitation>
;
<materialsCitation id="C3EBE8781865FFFB452D3293A9C1EABF" collectingDate="1998-03-05" collectingDateMax="1998-03-11" collectingDateMin="1998-03-05" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" county="Andringitra" elevation="1990" location="Philippia" municipality="Anjavidilay" pageId="33" pageNumber="34" specimenCode="CASENT9064649" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa Province" typeStatus="holotype">
RNI
<collectingCounty id="9A5D9AA91865FFFB45DA3293A823EAFF" box="[661,788,924,947]" pageId="33" pageNumber="34">Andringitra</collectingCounty>
,
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,
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<quantity id="B47B4FC01865FFFB44E63293A8C4EAFF" box="[937,1011,924,947]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="33" pageNumber="34" unit="km" value="8.5">8.5 km</quantity>
SE Atanifotsy
</locationDeviation>
, 22°09.5ʹS, 46°57.6ʹE, elev.
<quantity id="B47B4FC01865FFFB46E932B3AAC5EA9F" box="[422,498,956,979]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="33" pageNumber="34" unit="m" value="1990.0">
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</quantity>
,
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in
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<location id="765CB4FE1865FFFB45D732B3A9CCEA9F" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1865FFFB45D732B3A9CCEA9F" box="[664,763,956,979]" country="Madagascar" county="Andringitra" municipality="Anjavidilay" name="Philippia" pageId="33" pageNumber="34" stateProvince="Fianarantsoa Province">Philippia</location>
</emphasis>
-dominated sclerophyllus forest,
<date id="073DC4E51865FFFB432D32B3ABC8EABF" pageId="33" pageNumber="34" value="1998-03-05" valueMax="1998-03-11" valueMin="1998-03-05">
<collectingDate id="17793D0D1865FFFB432D32B3ABC8EABF" pageId="33" pageNumber="34" value="1998-03-05" valueMax="1998-03-11" valueMin="1998-03-05">5–11 March 1998</collectingDate>
</date>
,
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(
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,
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,
<collectionCode id="15927AE01865FFFB45D232D3A9D8EABF" box="[669,751,988,1011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="33" pageNumber="34" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781865FFFB444E32D3AF24ED5F" collectingDate="2000-03-13" collectionCode="CAS" collectorName="E. Schlinger" country="Madagascar" elevation="1670" latitude="-20.384445" location="Korikory" longLatPrecision="21" longitude="47.667225" pageId="33" pageNumber="34" specimenCode="CASENT9006000" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa Province" typeStatus="holotype">
<location id="765CB4FE1865FFFB444E32D3A868EABF" LSID="urn:lsid:plazi:treatment:FB2A53331864FFFB473430D4A9D7EDFF:765CB4FE1865FFFB444E32D3A868EABF" box="[769,863,988,1011]" country="Madagascar" latitude="-20.384445" longLatPrecision="21" longitude="47.667225" name="Korikory" pageId="33" pageNumber="34" stateProvince="Fianarantsoa Province">Korikory</location>
,
<locationDeviation id="628EFB071865FFFB442332D3AF2CEABF" box="[876,1051,988,1011]" location="Fandriana" pageId="33" pageNumber="34">SE of Fandriana</locationDeviation>
,
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,
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, elev.
<quantity id="B47B4FC01865FFFB466332F3AA4FED5F" box="[300,376,1020,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="33" pageNumber="34" unit="m" value="1670.0">
<elevation id="F8AE05161865FFFB466332F3AA4FED5F" box="[300,376,1020,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="33" pageNumber="34" unit="m" value="1670.0">1670m</elevation>
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,
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<collectingDate id="17793D0D1865FFFB46CA32F3A91AED5F" box="[389,557,1020,1043]" pageId="33" pageNumber="34" value="2000-03-13">13 March 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31865FFFB457632F3A989ED5F" box="[569,702,1020,1043]" pageId="33" pageNumber="34">E. Schlinger</collectorName>
(
<specimenCode id="23254A5E1865FFFB458232F3A8A0ED5F" box="[717,919,1020,1043]" collectionCode="CASENT" pageId="33" pageNumber="34">CASENT9006000</specimenCode>
,
<specimenCount id="658529AC1865FFFB44EC32F3A8F3ED5F" box="[931,964,1020,1043]" count="1" pageId="33" pageNumber="34" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01865FFFB449C32F3AF3CED5F" box="[979,1035,1020,1043]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="34" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251865FFFB47E43513A812ED3F" blockId="33.[122,1174,220,1203]" pageId="33" pageNumber="34">
<emphasis id="41F73E371865FFFB47E43513AA54ED7F" bold="true" box="[171,355,1052,1075]" pageId="33" pageNumber="34">Natural history.</emphasis>
Records of
<taxonomicName id="B48399A61865FFFB46BF3513A98EED7F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[496,697,1052,1075]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371865FFFB46BF3513A98EED7F" bold="true" box="[496,697,1052,1075]" italics="true" pageId="33" pageNumber="34">Uduba fandroana</emphasis>
</taxonomicName>
are all from over
<quantity id="B47B4FC01865FFFB44C23513A8E0ED7F" box="[909,983,1052,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="m" value="1000.0">1000m</quantity>
elevation, in the vicinity of montane rainforest, from forest margins and from
<emphasis id="41F73E371865FFFB446E3533A8B3ED1F" box="[801,900,1084,1107]" italics="true" pageId="33" pageNumber="34">Philippia</emphasis>
-dominated sclerophyllus forest. Nothing is known of silk use by this ecribellate species.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1865FFFB47E43573A9D7EDFF" pageId="33" pageNumber="34" type="distribution">
<paragraph id="733CE2251865FFFB47E43573A9D7EDFF" blockId="33.[122,1174,220,1203]" pageId="33" pageNumber="34">
<emphasis id="41F73E371865FFFB47E43573AA0AEDDF" bold="true" box="[171,317,1148,1171]" pageId="33" pageNumber="34">Distribution.</emphasis>
Specimens of
<taxonomicName id="B48399A61865FFFB46AC3573A99DEDDF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[483,682,1148,1171]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E371865FFFB46AC3573A99DEDDF" bold="true" box="[483,682,1148,1171]" italics="true" pageId="33" pageNumber="34">Uduba fandroana</emphasis>
</taxonomicName>
have been collected in upland eastern forest from central to central-south
<collectingCountry id="0B94A2B51865FFFB46F83593A90FEDFF" box="[439,568,1180,1203]" name="Madagascar" pageId="33" pageNumber="34">Madagascar</collectingCountry>
(Maps 12, 13).
</paragraph>
</subSubSection>
</treatment>
</document>

434
data/FB/2A/53/FB2A53331868FFF447343093A8EAE8BF.xml Normal file
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@ -0,0 +1,434 @@
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<mods:title id="C37870DB76DAA7CC897D47BC3AD22F35">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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,
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<subSubSection id="3B99B1AE1868FFF6473430B3A903E89D" box="[123,564,444,465]" pageId="44" pageNumber="45" type="description">
<paragraph id="733CE2251868FFF6473430B3A903E89D" blockId="44.[123,564,412,465]" box="[123,564,444,465]" pageId="44" pageNumber="45">
<figureCitation id="EBB8FEA01868FFF6473430B3ABCDE89D" box="[123,250,444,465]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Figures 28 A</figureCitation>
,
<figureCitation id="EBB8FEA01868FFF6464B30B3AA66E89D" box="[260,337,444,465]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="44" pageNumber="45">44 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01868FFF6461330B3AA9DE89D" box="[348,426,444,465]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">72 C, D</figureCitation>
, Maps 10, 17.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1868FFF647E430EDA92AEB35" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="733CE2251868FFF647E430EDA92AEB35" blockId="44.[122,1174,482,1725]" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E430EDAA65E8B5" bold="true" box="[171,338,482,505]" pageId="44" pageNumber="45">Type material.</emphasis>
<materialsCitation id="C3EBE8781868FFF6461430EDA8D5EB75" collectingDate="2014-08-26" collectingDateMax="2014-08-29" collectingDateMin="2014-08-26" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1385" location="Tsinjoarivo Forest" pageId="44" pageNumber="45" specimenCode="CASENT9065679, CASENT9064772" specimenCount="2" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="AC385C871868FFF6461430EDAA89E8B5" box="[347,446,482,505]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
male,
<specimenCode id="23254A5E1868FFF6454330EDA9E6E8B5" box="[524,721,482,505]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9065679</specimenCode>
and
<typeStatus id="AC385C871868FFF6444430EDA850E8B5" box="[779,871,482,505]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
female,
<specimenCode id="23254A5E1868FFF6448530EDAFA5E8B5" box="[970,1170,482,505]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9064772</specimenCode>
, from montane rainforest at
<quantity id="B47B4FC01868FFF646EB330DAAD9EB55" box="[420,494,514,537]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="44" pageNumber="45" unit="m" value="1385.0">
<elevation id="F8AE05161868FFF646EB330DAAD9EB55" box="[420,494,514,537]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="44" pageNumber="45" unit="m" value="1385.0">1385m</elevation>
</quantity>
elevation in
<location id="765CB4FE1868FFF64534330DA877EB55" LSID="urn:lsid:plazi:treatment:FB2A53331868FFF447343093A8EAE8BF:765CB4FE1868FFF64534330DA877EB55" box="[635,832,514,537]" country="Madagascar" name="Tsinjoarivo Forest" pageId="44" pageNumber="45" stateProvince="Antananarivo">Tsinjoarivo Forest</location>
,
<collectingRegion id="B1472CC71868FFF64406330DAF73EB55" box="[841,1092,514,537]" country="Madagascar" name="Antananarivo" pageId="44" pageNumber="45">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51868FFF6431F330DAB8AEB75" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
, collected
<date id="073DC4E51868FFF64676332DA938EB75" box="[313,527,546,569]" pageId="44" pageNumber="45" value="2014-08-26" valueMax="2014-08-29" valueMin="2014-08-26">
<collectingDate id="17793D0D1868FFF64676332DA938EB75" box="[313,527,546,569]" pageId="44" pageNumber="45" value="2014-08-26" valueMax="2014-08-29" valueMin="2014-08-26">26–29 August 2014</collectingDate>
</date>
by
<collectorName id="DE7687F31868FFF64570332DA9F0EB75" box="[575,711,546,569]" pageId="44" pageNumber="45">B. L. Fisher</collectorName>
<emphasis id="41F73E371868FFF6459E332DA833EB75" box="[721,772,546,569]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
., deposited in
<collectionCode id="15927AE01868FFF644E4332DA8D5EB75" box="[939,994,546,569]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="44" pageNumber="45" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781868FFF644BF332DA92EEB35" collectingDate="2014-08-26" collectingDateMax="2014-08-29" collectingDateMin="2014-08-26" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1385" location="Tsinjoarivo Forest" pageId="44" pageNumber="45" specimenCode="CASENT9065707, CASENT9065595, CASENT9065602, CASENT9065698" specimenCount="4" specimenCount-male="4" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="AC385C871868FFF644BF332DAF6DEB75" box="[1008,1114,546,569]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="658529AC1868FFF64326332DAB8DEB15" count="4" pageId="44" pageNumber="45" type="male">four males</specimenCount>
, same collection data (
<specimenCode id="23254A5E1868FFF64696334DA996EB15" box="[473,673,578,601]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9065707</specimenCode>
,
<specimenCode id="23254A5E1868FFF645F6334DA8B7EB15" box="[697,896,578,601]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9065595</specimenCode>
,
<specimenCode id="23254A5E1868FFF644D8334DAF6AEB15" box="[919,1117,578,601]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9065602</specimenCode>
and
<specimenCode id="23254A5E1868FFF64734336DAA73EB35" box="[123,324,610,633]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9065698</specimenCode>
), deposited in
<collectionCode id="15927AE01868FFF646AD336DA92EEB35" box="[482,537,610,633]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="44" pageNumber="45" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1868FFF647E4338DA935EA76" pageId="44" pageNumber="45" type="etymology">
<paragraph id="733CE2251868FFF647E4338DA935EA76" blockId="44.[122,1174,482,1725]" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E4338DAA1DEBD5" bold="true" box="[171,298,642,665]" pageId="44" pageNumber="45">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E371868FFF64541338DA947EBD5" box="[526,624,642,665]" italics="true" pageId="44" pageNumber="45">hiragasy,</emphasis>
refers to a musical tradition in
<collectingCountry id="0B94A2B51868FFF6448A338DAF73EBD5" box="[965,1092,642,665]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
, with a daylong celebration of music, dances and performances. According to Wikipedia:
<taxonomicName id="B48399A61868FFF6434933ADAF42EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1030,1141,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="hiragasy">
“
<emphasis id="41F73E371868FFF6435D33ADAF42EBF5" box="[1042,1141,674,697]" italics="true" pageId="44" pageNumber="45">Hiragasy</emphasis>
</taxonomicName>
or
<emphasis id="41F73E371868FFF6473433CCABD7EB96" box="[123,224,707,730]" italics="true" pageId="44" pageNumber="45">hira gasy</emphasis>
is a musical tradition in
<collectingCountry id="0B94A2B51868FFF646A333CCA95CEB96" box="[492,619,707,730]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
, particularly among the
<emphasis id="41F73E371868FFF6443C33CCA8F5EB96" box="[883,962,707,730]" italics="true" pageId="44" pageNumber="45">Merina</emphasis>
ethnic group of the Highland regions around the capital of
<collectingRegion id="B1472CC71868FFF6456833ECA98BEBB6" box="[551,700,739,762]" country="Madagascar" name="Antananarivo" pageId="44" pageNumber="45">Antananarivo</collectingRegion>
. It is a day-long spectacle of music, dance, and
<emphasis id="41F73E371868FFF647E5320CABC3EA56" box="[170,244,771,794]" italics="true" pageId="44" pageNumber="45">kabary</emphasis>
oratory performed by a troupe or as a competition between two troupes. (Wikipedia,
<date id="073DC4E51868FFF64734322CABC0EA76" box="[123,247,803,826]" pageId="44" pageNumber="45" value="2021-09-29">ix-29-2021</date>
).” A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1868FFF647E4324CAA86EDB7" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="733CE2251868FFF647E4324CAA86EDB7" blockId="44.[122,1174,482,1725]" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E4324CAAE0EA16" bold="true" box="[171,471,835,858]" pageId="44" pageNumber="45">
Diagnosis.
<taxonomicName id="B48399A61868FFF64668324CAAE0EA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[295,471,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371868FFF64668324CAAE0EA16" bold="true" box="[295,471,835,858]" italics="true" pageId="44" pageNumber="45">Uduba hiragasy</emphasis>
</taxonomicName>
</emphasis>
belong to Group III, the
<taxonomicName id="B48399A61868FFF645A8324CA840EA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[743,887,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">Uduba valiha</taxonomicName>
group (
<figureCitation id="EBB8FEA01868FFF64483324CAF14EA16" box="[972,1059,835,858]" captionStart="MAP" captionStartId="184.[155,174,1176,1195]" captionTargetBox="[120,1166,216,1153]" captionTargetId="figure-11@184.[120,648,216,1161]" captionTargetPageId="184" captionText="MAP 17. Distribution of Group III, the Uduba valiha MAP 18. Distribution of Group IV, the Uduba rinha group: Uduba hiragasy, U. fisheri and U. valiha. group: Uduba rinha and U. irwini." figureDoi="http://doi.org/10.5281/zenodo.13158795" httpUri="https://zenodo.org/record/13158795/files/figure.png" pageId="44" pageNumber="45">Map 17</figureCitation>
). They are cribellate with divided cribellum or vestige. Males differ from other species by having a retromedian ridge on the tegulum (
<figureCitation id="EBB8FEA01868FFF646E8328CA908EAD6" box="[423,575,899,922]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="44" pageNumber="45">Figs. 44 B, C</figureCitation>
), and a deeply bifid MA (
<figureCitation id="EBB8FEA01868FFF64411328CA8FFEAD6" box="[862,968,899,922]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 A</figureCitation>
) with the prolateral lobe slender, retrolateral lobe stout;
<taxonomicName id="B48399A61868FFF6455132ACA9A8EAF6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[542,671,931,954]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E371868FFF6455132ACA9A8EAF6" bold="true" box="[542,671,931,954]" italics="true" pageId="44" pageNumber="45">U. hiragasy</emphasis>
</taxonomicName>
males differ from those of
<taxonomicName id="B48399A61868FFF6448132ACAF03EAF6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[974,1076,931,954]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371868FFF6448132ACAF03EAF6" bold="true" box="[974,1076,931,954]" italics="true" pageId="44" pageNumber="45">U. valiha</emphasis>
</taxonomicName>
by lacking the short, stout spine retrolateral spine from the palpal tibia (
<figureCitation id="EBB8FEA01868FFF6440C32CCA89AEA96" box="[835,941,963,986]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 A</figureCitation>
) (spine present in
<emphasis id="41F73E371868FFF6433432CCABF1EAB7" bold="true" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="B48399A61868FFF6433432CCAB88EAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">U. valiha</taxonomicName>
,
</emphasis>
<figureCitation id="EBB8FEA01868FFF6478332EBAA05EAB7" box="[204,306,996,1019]" captionStart="FIGURE" captionStartId="154.[155,167,1647,1666]" captionTargetBox="[120,1176,861,1630]" captionTargetId="figure-46@154.[120,1176,861,1630]" captionTargetPageId="154" captionText="FIGURE 64. Uduba valiha, new species, Holotype (CASENT9026394) from Forêt de Vevembe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum (mostly hidden behind TA2 in this species). White arrows to bump on tegulum, black arrow to stout retrolateral spine on tibia. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158717" httpUri="https://zenodo.org/record/13158717/files/figure.png" pageId="44" pageNumber="45">Fig. 64 C</figureCitation>
), by having the TA2 blunt, entire (
<figureCitation id="EBB8FEA01868FFF645EF32EBA831EAB7" box="[672,774,996,1019]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 A</figureCitation>
) (apically rolled in
<emphasis id="41F73E371868FFF6449732EBAF73EAB7" bold="true" box="[984,1092,996,1019]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="B48399A61868FFF6449732EBAF0BEAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[984,1084,996,1019]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">U. valiha</taxonomicName>
,
</emphasis>
<figureCitation id="EBB8FEA01868FFF6430532EBABBAED57" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 B</figureCitation>
) and the apex of the MA prolateral lobe slender truncate (
<figureCitation id="EBB8FEA01868FFF645B1350BA854ED57" box="[766,867,1028,1051]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 A</figureCitation>
<emphasis id="41F73E371868FFF6442C350BA85BED57" bold="true" box="[867,876,1028,1051]" pageId="44" pageNumber="45">)</emphasis>
(rounded in
<taxonomicName id="B48399A61868FFF644B8350BAF6AED57" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1015,1117,1028,1051]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371868FFF644B8350BAF6AED57" bold="true" box="[1015,1117,1028,1051]" italics="true" pageId="44" pageNumber="45">U. valiha</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01868FFF64324350BAB86ED77" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="44" pageNumber="45">Fig. 28 B</figureCitation>
). The females (
<figureCitation id="EBB8FEA01868FFF64615352BAAD8ED77" box="[346,495,1060,1083]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">Figs. 72 C, D</figureCitation>
) lack a median lobe or defined atrium from the epigynum and the vulva with a few, anteroposteriorly oriented, spermathecal loops; can be distinguished from those of similar
<taxonomicName id="B48399A61868FFF64663356BAA41ED37" authorityName="Simon" authorityYear="1880" box="[300,374,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371868FFF64663356BAA41ED37" bold="true" box="[300,374,1124,1147]" italics="true" pageId="44" pageNumber="45">Uduba</emphasis>
</taxonomicName>
(
<taxonomicName id="B48399A61868FFF646C8356BAAD9ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[391,494,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371868FFF646C8356BAAD9ED37" bold="true" box="[391,494,1124,1147]" italics="true" pageId="44" pageNumber="45">U. valiha</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61868FFF6456A356BA9B9ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[549,654,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371868FFF6456A356BA9B9ED37" bold="true" box="[549,654,1124,1147]" italics="true" pageId="44" pageNumber="45">U. fisheri</emphasis>
</taxonomicName>
) species by having the lateral epigynal margins weakly convex (more convex in
<taxonomicName id="B48399A61868FFF64696358BA908EDD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[473,575,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371868FFF64696358BA908EDD7" bold="true" box="[473,575,1156,1179]" italics="true" pageId="44" pageNumber="45">U. valiha</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01868FFF64503358BA9F7EDD7" box="[588,704,1156,1179]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">Figs. 72 G</figureCitation>
,
<figureCitation id="EBB8FEA01868FFF64584358BA87DEDD7" box="[715,842,1156,1179]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="44" pageNumber="45">
80 H and
<taxonomicName id="B48399A61868FFF6447A358BA87DEDD7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[821,842,1156,1179]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371868FFF6447A358BA87DEDD7" bold="true" box="[821,842,1156,1179]" italics="true" pageId="44" pageNumber="45">U</emphasis>
</taxonomicName>
</figureCitation>
. fisheri,
<figureCitation id="EBB8FEA01868FFF644E4358BAF77EDD7" box="[939,1088,1156,1179]" captionStart="FIGURE" captionStartId="170.[155,167,1416,1435]" captionTargetBox="[121,1176,216,1416]" captionTargetId="figure-11@170.[120,1176,216,1401]" captionTargetPageId="170" captionText="FIGURE 82. Uduba spp., ♀ genitalia, SEM images. A, B. U. platnicki, new species, CASENT9030253, from Mikira Forest. A. Vulva, dorsal. B. Spermathecal texture, close-up. C–F. U. fisheri, new species, CASENT9026389 from Anjozorobe. C. Vulva, dorsal. D. Vulva, anterior. E. Spermathecal texture, close-up. F. Epigynum, ventral. Scale bars: A = 100 µm, B, D, E = 10 µm, C, F = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158759" httpUri="https://zenodo.org/record/13158759/files/figure.png" pageId="44" pageNumber="45">Figs. 82 C–F</figureCitation>
), with a strong margin along the epigastric furrow (weak in
<taxonomicName id="B48399A61868FFF645EA35ABA83DEDF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[677,778,1188,1211]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="valiha">
<emphasis id="41F73E371868FFF645EA35ABA83DEDF7" bold="true" box="[677,778,1188,1211]" italics="true" pageId="44" pageNumber="45">U. valiha</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01868FFF6445635ABA8BAEDF7" box="[793,909,1188,1211]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">Figs. 72 H</figureCitation>
,
<figureCitation id="EBB8FEA01868FFF644D735ABA8FDEDF7" box="[920,970,1188,1211]" captionStart="FIGURE" captionStartId="170.[155,167,1416,1435]" captionTargetBox="[121,1176,216,1416]" captionTargetId="figure-11@170.[120,1176,216,1401]" captionTargetPageId="170" captionText="FIGURE 82. Uduba spp., ♀ genitalia, SEM images. A, B. U. platnicki, new species, CASENT9030253, from Mikira Forest. A. Vulva, dorsal. B. Spermathecal texture, close-up. C–F. U. fisheri, new species, CASENT9026389 from Anjozorobe. C. Vulva, dorsal. D. Vulva, anterior. E. Spermathecal texture, close-up. F. Epigynum, ventral. Scale bars: A = 100 µm, B, D, E = 10 µm, C, F = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.13158759" httpUri="https://zenodo.org/record/13158759/files/figure.png" pageId="44" pageNumber="45">82 F</figureCitation>
) and with the copulatory openings in the posterior part of plate (
<figureCitation id="EBB8FEA01868FFF6453B35CBA9EBED97" box="[628,732,1220,1243]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">Fig. 72 C</figureCitation>
<emphasis id="41F73E371868FFF6459435CBA9D3ED97" bold="true" box="[731,740,1220,1243]" pageId="44" pageNumber="45">)</emphasis>
at 0.31 of epigynum length (more anterior in
<taxonomicName id="B48399A61868FFF6478A35EBAA1AEDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[197,301,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="41F73E371868FFF6478A35EBAA1AEDB7" bold="true" box="[197,301,1252,1275]" italics="true" pageId="44" pageNumber="45">U. fisheri</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01868FFF6467335EBAA92EDB7" box="[316,421,1252,1275]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="44" pageNumber="45">Fig. 72 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1868FFF447E4340AA8BDE8FF" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="733CE2251868FFF747E4340AAA32ED95" blockId="44.[122,1174,482,1725]" lastBlockId="45.[122,1175,220,1725]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E4340AAA41EC50" bold="true" box="[171,374,1285,1308]" pageId="44" pageNumber="45">Description. Male</emphasis>
(
<typeStatus id="AC385C871868FFF646CC340AAAD0EC50" box="[387,487,1285,1308]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
): Total length 10.80. Markings typical for
<taxonomicName id="B48399A61868FFF644E9340AA8C7EC50" authorityName="Simon" authorityYear="1880" box="[934,1008,1285,1308]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371868FFF644E9340AA8C7EC50" bold="true" box="[934,1008,1285,1308]" italics="true" pageId="44" pageNumber="45">Uduba</emphasis>
</taxonomicName>
. Carapace 5.50 long, 3.60 wide, 1.90 high; clypeus 0.20 high. Eye diameters:AME 0.18, ALE and PME 0.20, PLE 0.22. Chelicerae 2.30 long; sternum 3.10 long, 2.30 wide; labium 1.40 long; palpal coxae 2.00 long. Divided cribellum. Spination: palpus–femur d1-1-1, p0-0-1, r0-0-1, tibia p1-0-0; leg I–femur d1- 1-0-1, p0-0-2, r0-0-1-1, tibia d0-1-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-1, v2-2-2-1, r1-0-1; leg II–femur d1-1-0-1, p0-1-0-1, r1-1-1-1, tibia d1-1-0-0, p0-1-1-0, v2-2-2-2, metatarsus p1-1-2, v2-2-1, r1-1-2; leg III–femur d1-1-1, p1-1-1-0, r1-1-1-0, patella p1, r1, tibia d1-0-1-0, p0- 1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-1, p1-1-0-1, r0-0- 0-1, patella r1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v1-2-2-1, r1-2-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, weak, ventral. Leg measurements: I: 4.60 + 1.90 + 4.50 + 4.90 + 2.90 = 18.80; II: 4.30 + 1.80 + 3.80 + 4.00 + 2.30 = 16.20; III: 3.80 + 1.70 + 2.60 + 3.50 + 1.90 = 13.50; IV: 5.00 + 1.70 + 4.30 + 5.50 + 2.30 = 18.80; palpus: 2.10 + 0.70 + 0.80 + NA + 2.10 = 5.70. Leg formula 1=4, 23.
<emphasis id="41F73E371868FFF6451C3789A9F0EFD1" bold="true" box="[595,711,1670,1693]" pageId="44" pageNumber="45">Male palp</emphasis>
: palpal tibia 0.42 cymbial length, RTA triangular, slightly curved, RTA length 0.30 tibia, with pointed apex, VTA triangular, length 0.32 tibia width,
<collectingRegion id="B1472CC71869FFF7478931D3ABCEE9BF" box="[198,249,220,243]" country="United States of America" name="Utah" pageId="45" pageNumber="46">UTA</collectingRegion>
small; tibia with a long, slender prolateral spine (
<figureCitation id="EBB8FEA01869FFF7445931D3A84AE9BF" box="[790,893,220,243]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="45" pageNumber="46">Fig. 44 A</figureCitation>
<emphasis id="41F73E371869FFF7443231D3A8B1E9BF" bold="true" box="[893,902,220,243]" pageId="45" pageNumber="46">)</emphasis>
, without stout retrolateral spine (
<figureCitation id="EBB8FEA01869FFF7479031F3AA7FE85F" box="[223,328,252,275]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="45" pageNumber="46">Fig. 44 C</figureCitation>
<emphasis id="41F73E371869FFF7460831F3AA67E85F" bold="true" box="[327,336,252,275]" pageId="45" pageNumber="46">)</emphasis>
; tegulum with longitudinal, retromedian ridge, ridge beginning near tegulum base and extending for 0.70 tegulum length; TA3 a pointed spine, mostly hidden behind TA2, TA2 small, trapezoidal (
<figureCitation id="EBB8FEA01869FFF746063033AA85E81F" box="[329,434,316,339]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="45" pageNumber="46">Fig. 28 A</figureCitation>
); MA origin at 0.65 tegulum length, extends to tegulum apex, length (including apical processes) slightly less than width, length 0.30 tegulum width, deeply bifid MA (
<figureCitation id="EBB8FEA01869FFF747CD3072ABC3E8D8" box="[130,244,381,404]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="45" pageNumber="46">Figs. 28 A</figureCitation>
,
<figureCitation id="EBB8FEA01869FFF747B03072AA1BE8D8" box="[255,300,381,404]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="45" pageNumber="46">44B</figureCitation>
<emphasis id="41F73E371869FFF746633072AA02E8D8" bold="true" box="[300,309,381,404]" pageId="45" pageNumber="46">)</emphasis>
with prolateral lobe slender, retrolateral lobe stout; conductor fan entire (
<figureCitation id="EBB8FEA01869FFF743073072ABBAE8F8" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="45" pageNumber="46">Fig. 44 C</figureCitation>
<emphasis id="41F73E371869FFF747C23092ABA1E8F8" bold="true" box="[141,150,413,436]" pageId="45" pageNumber="46">)</emphasis>
.
<emphasis id="41F73E371869FFF747EB3092ABC0E8F8" bold="true" box="[164,247,413,436]" pageId="45" pageNumber="46">Female</emphasis>
(
<typeStatus id="AC385C871869FFF7464A3092AA52E8F8" box="[261,357,413,436]" pageId="45" pageNumber="46" type="paratype">paratype</typeStatus>
, CASENT9065679): Total length 12.00. Markings typical for
<taxonomicName id="B48399A61869FFF7434D3092AF7BE8F8" authorityName="Simon" authorityYear="1880" box="[1026,1100,413,436]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371869FFF7434D3092AF7BE8F8" bold="true" box="[1026,1100,413,436]" italics="true" pageId="45" pageNumber="46">Uduba</emphasis>
</taxonomicName>
. Carapace 5.10 long, 3.50 wide, 1.80 high; clypeus 0.30 high. Eye diameters: AME and ALE 0.16, PME 0.20, PLE 0.24. Chelicerae 2.50 long; sternum 2.30 long, 1.80 wide; labium 1.10 long; palpal coxae 1.70 long.
<emphasis id="41F73E371869FFF747A030F1AA73EB59" bold="true" box="[239,324,510,533]" pageId="45" pageNumber="46">Ratios–</emphasis>
carapace length / width = 1.46, carapace height / width = 0.51, PER / carapace width = 0.42, PER / OAL = 3.04, PER / AER = 1.40, OAL / OQL = 1.20, OQP / OQA= 1.19, clypeus height / AME = 1.88, cheliceral length / clypeus height = 8.33, sternum length / width = 1.28, palpal coxa length / width = 2.43, femur I length / carapace width = 1.03, metatarsus I length / carapace width = 0.80, femur IV length / carapace width = 1.06, palpal tarsus length / carapace width = 0.46, palpal tarsus length / palpal patella length = 1.78, palpal tarsus length / palpal tibia length = 1.60, palpal tarsus length / palpal femur length = 0.94, palpal tibia length / palpal patella length = 1.11. Cribellum divided. Spination: palpus–femur d0-1-2, patella p1-0, tibia p2-2, tarsus p2-0-1, v0-0-1, r0-0-1; leg I–femur d1-0-1, p0-0-1, tibia v2-2-2-2, metatarsus p0-0-1, v2-2-2; leg II–femur d1-0-0, p0-0-1, tibia v1-1-1-2, metatarsus v2-2-3; leg III–femur d1-0-0-0, p0-1-0-1, r0-0- 0-1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-0, p0-0-0-1, tibia p0-1-1-0, v2-2-2, metatarsus p0-1-1-2, v2-2-2, r1-2-1. Scopulae: ventral on tarsus and apical metatarsus I and II, tarsus III and IV. Leg measurements: I: 3.60 + 1.80 + 3.00 + 2.80 + 1.80 = 13.00; II: 2.90 + 1.50 + 2.50 + 2.40 + 1.70 = 11.00; III: 2.80 + 1.30 + 1.70 + 2.20 + 1.50 = 9.50; IV: 3.70 + 1.50 + 3.10 + 3.70 + 2.00 = 14.00; palpus: 1.70 + 0.90 + 1.00 + NA + 1.60 = 5.20. Leg formula 4123.
<emphasis id="41F73E371869FFF7454A32EEA9F2EAB4" bold="true" box="[517,709,993,1016]" pageId="45" pageNumber="46">Female genitalia</emphasis>
: epigynal plate without depressed atrium (
<figureCitation id="EBB8FEA01869FFF747CC350EABDDED54" box="[131,234,1025,1048]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="45" pageNumber="46">Fig. 72 C</figureCitation>
), plate width 1.43 times length, lateral lobes with posterolateral edges and posteromedi- an longitudinal ridges on each side, width between bases of LL 0.68 times epigynum width, LL side width 0.16 times width epigynum; median lobe a flat plate, copulatory openings at mid-level of plate at 0.31 of epigynum length, distance between copulatory openings 0.42 times epigynum width. Vulva (
<figureCitation id="EBB8FEA01869FFF74658358DAAB0EDD5" box="[279,391,1154,1177]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="45" pageNumber="46">Fig. 72 D</figureCitation>
), with spermathecal ducts forming 2-3 loose, longitudinal loops, vulva length 0.88 times width, fertilization ducts widely separated, vulva width 4.86 times distance between FD.
</paragraph>
<paragraph id="733CE2251869FFF747E435ECABD5EF30" blockId="45.[122,1175,220,1725]" pageId="45" pageNumber="46">
<emphasis id="41F73E371869FFF747E435ECAA2DEDB6" bold="true" box="[171,282,1251,1274]" pageId="45" pageNumber="46">Variation.</emphasis>
Male (N= 5): Total length 9.70–10.80; carapace length / width = 1.43–1.53, carapace height / width = 0.43–0.60, PER / carapace width = 0.34–0.35, PER / OAL = 2.42–2.54, PER / AER = 1.29–1.33, OAL / OQL = 1.09–1.25, OQP / OQA = 1.14–1.26, clypeus height / AME = 1.11–1.88, cheliceral length / clypeus height = 8.40–12.50, sternum length / width = 1.30–1.41, palpal coxa length / width = 2.50–3.00, femur I length / carapace width = 1.22–1.29, metatarsus I length / carapace width = 1.23–1.36, femur IV length / carapace width = 1.29–1.43, cymbium length / carapace width = 0.58–0.60, cymbium length / palpal patella length = 2.50–3.00, cymbium length / palpal tibia length = 2.33–2.63, cymbium length / palpal femur length = 1.00–1.05, palpal tibia length / palpal patella length = 1.00–1.14; palpal tibia length = 0.42–0.44 times cymbial length, RTA length 0.21–0.32 times tibia length, VTA length 0.19–0.32 times tibia width; tegulum longitudinal, retroapical ridge extending for 0.50–0.70 tegulum length, MA origin at 0.55–0.65 times tegulum length (
<figureCitation id="EBB8FEA01869FFF74622374AAAD7EF10" box="[365,480,1605,1628]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="45" pageNumber="46">Figs. 28 A</figureCitation>
,
<figureCitation id="EBB8FEA01869FFF746A4374AA90AEF10" box="[491,573,1605,1628]" captionStart="FIGURE" captionStartId="141.[155,167,888,907]" captionTargetBox="[120,1176,216,870]" captionTargetId="figure-19@141.[120,1176,215,874]" captionTargetPageId="141" captionText="FIGURE 44. Uduba hiragasy, new species, paratype (CASENT9065698) from Tsinjoarivo Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Arrow to tegular ridge. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158669" httpUri="https://zenodo.org/record/13158669/files/figure.png" pageId="45" pageNumber="46">44 A–C</figureCitation>
<emphasis id="41F73E371869FFF74572374AA971EF10" bold="true" box="[573,582,1605,1628]" pageId="45" pageNumber="46">)</emphasis>
. Female variation unknown; only
<specimenCount id="658529AC1869FFF744F6374AAF06EF10" box="[953,1073,1605,1628]" count="1" pageId="45" pageNumber="46" type="female">one female</specimenCount>
has been collected.
</paragraph>
<paragraph id="733CE2251869FFF447E53789A857E81F" blockId="45.[122,1175,220,1725]" lastBlockId="46.[123,1174,220,499]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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M
<emphasis id="41F73E371869FFF7478D3789AABBEFD1" bold="true" box="[194,396,1670,1693]" pageId="45" pageNumber="46">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51869FFF746DA3789A960EFD1" box="[405,599,1670,1693]" name="Madagascar" pageId="45" pageNumber="46">MADAGASCAR</collectingCountry>
: A
<emphasis id="41F73E371869FFF7453A3789A857EFD1" box="[629,864,1670,1693]" italics="true" pageId="45" pageNumber="46">
ntananarivo
<location id="765CB4FE1869FFF7444D3789A857EFD1" LSID="urn:lsid:plazi:treatment:FB2A53331868FFF447343093A8EAE8BF:765CB4FE1869FFF7444D3789A857EFD1" box="[770,864,1670,1693]" country="Madagascar" county="Ankadivory" latitude="-19.71572" longLatPrecision="1" longitude="47.82084" municipality="Tsinjoarivo forest" name="Province" pageId="45" pageNumber="46">Province</location>
</emphasis>
:
<collectingMunicipality id="9358785F1869FFF7443E3789AF04EFD1" box="[881,1075,1670,1693]" pageId="45" pageNumber="46">Tsinjoarivo forest</collectingMunicipality>
,
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,
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,
<geoCoordinate id="16B784E21869FFF7460E37A9AAF3EFF1" box="[321,452,1702,1725]" degrees="47.82084" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="47.82084">47.82084°E</geoCoordinate>
, elev.
<quantity id="B47B4FC01869FFF7454A37A9A965EFF1" box="[517,594,1702,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="45" pageNumber="46" unit="m" value="1385.0">
<elevation id="F8AE05161869FFF7454A37A9A965EFF1" box="[517,594,1702,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="45" pageNumber="46" unit="m" value="1385.0">1385m</elevation>
</quantity>
, montane rainforest,
<collectingMethod id="AAC29A321869FFF7447A37A9A885EFF1" box="[821,946,1702,1725]" pageId="45" pageNumber="46">pitfall traps</collectingMethod>
,
<date id="073DC4E51869FFF744F137A9AFA6EFF1" box="[958,1169,1702,1725]" pageId="45" pageNumber="46" value="2014-08-26" valueMax="2014-08-29" valueMin="2014-08-26">
<collectingDate id="17793D0D1869FFF744F137A9AFA6EFF1" box="[958,1169,1702,1725]" pageId="45" pageNumber="46" value="2014-08-26" valueMax="2014-08-29" valueMin="2014-08-26">26–29 August 2014</collectingDate>
</date>
,
<collectorName id="DE7687F3186AFFF4473431D3ABCDE9BF" box="[123,250,220,243]" pageId="46" pageNumber="47">B.L. Fisher</collectorName>
<emphasis id="41F73E37186AFFF4464831D3AA0BE9BF" box="[263,316,220,243]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., [BLF 35410_12] (
<typeStatus id="AC385C87186AFFF4456931D3A9BAE9BF" box="[550,653,220,243]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
,
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,
<specimenCount id="658529AC186AFFF4443831D3A8AEE9BF" box="[887,921,220,243]" count="1" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
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)
</materialsCitation>
<materialsCitation id="C3EBE878186AFFF444B931D3A86AE81F" collectionCode="CAS" country="Madagascar" pageId="46" pageNumber="47" specimenCode="CASENT9064772, BLF 35410, CASENT9065707, CASENT9065595, CASENT9065602, CASENT9065698" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
and (
<typeStatus id="AC385C87186AFFF4437E31D3AFA6E9BF" box="[1073,1169,220,243]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E186AFFF4473431F3AA74E85F" box="[123,323,252,275]" collectionCode="CASENT" pageId="46" pageNumber="47">CASENT9064772</specimenCode>
,
<specimenCount id="658529AC186AFFF4461D31F3AA43E85F" box="[338,372,252,275]" count="1" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0186AFFF446C931F3AA8AE85F" box="[390,445,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="46" pageNumber="47" type="Museum">CAS</collectionCode>
), [
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_5] (
<typeStatus id="AC385C87186AFFF445DE31F3A9C6E85F" box="[657,753,252,275]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E186AFFF4444E31F3A8FEE85F" box="[769,969,252,275]" collectionCode="CASENT" pageId="46" pageNumber="47">CASENT9065707</specimenCode>
,
<specimenCount id="658529AC186AFFF4449731F3A8CDE85F" box="[984,1018,252,275]" count="1" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0186AFFF4434331F3AF74E85F" box="[1036,1091,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="46" pageNumber="47" type="Museum">CAS</collectionCode>
), [
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_9] (
<typeStatus id="AC385C87186AFFF447A13013AA79E87F" box="[238,334,284,307]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
,
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,
<specimenCount id="658529AC186AFFF445663013A97DE87F" box="[553,586,284,307]" count="1" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
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<emphasis id="41F73E37186AFFF447E43053AA56E83F" bold="true" box="[171,353,348,371]" pageId="46" pageNumber="47">Natural history.</emphasis>
The species
<taxonomicName id="B48399A6186AFFF446BD3053A993E83F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[498,676,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E37186AFFF446BD3053A993E83F" bold="true" box="[498,676,348,371]" italics="true" pageId="46" pageNumber="47">Uduba hiragasy</emphasis>
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occurs in montane rainforest. Both male and female specimens were collected in pitfall traps. The female has a developed cribellum and calamistrum, but it is unknown if the species wanders or makes retreats.
</paragraph>
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<emphasis id="41F73E37186AFFF447E430B3AA0AE89F" bold="true" box="[171,317,444,467]" pageId="46" pageNumber="47">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6186AFFF4468530B3A94CE89F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[458,635,444,467]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="hiragasy">
<emphasis id="41F73E37186AFFF4468530B3A94CE89F" bold="true" box="[458,635,444,467]" italics="true" pageId="46" pageNumber="47">Uduba hiragasy</emphasis>
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is known only from the
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locality in rainforest on the eastern side of the escarpment in east-central
<collectingCountry id="0B94A2B5186AFFF445FB30D3A802E8BF" box="[692,821,476,499]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
(Maps 10, 17).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="FE1D30918EA68A6E38E3E2C8FE4E4771">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="5D287D32F52B216A3B6EDDE5ED964912">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<emphasis id="41F73E37186AFFF447343312AA25EB78" bold="true" box="[123,274,541,564]" italics="true" pageId="46" pageNumber="47">Uduba ibonia</emphasis>
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,
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<figureCitation id="EBB8FEA0186AFFF447343332ABDDEB1E" box="[123,234,573,594]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="46" pageNumber="47">Figures 7 F</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF447BB3332AA20EB1E" box="[244,279,573,594]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="46" pageNumber="47">9 E</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF4466D3332AA7AEB1E" box="[290,333,573,594]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="46" pageNumber="47">11 F</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF446183332AABFEB1E" box="[343,392,573,594]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="46" pageNumber="47">20 B</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF446DC3332AA88EB1E" box="[403,447,573,594]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="46" pageNumber="47">29 F</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF446863332A921EB1E" box="[457,534,573,594]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">45 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF4456E3332A959EB1E" box="[545,622,573,594]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="46" pageNumber="47">73 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF445373332A9F5EB1E" box="[632,706,573,594]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="46" pageNumber="47">81 C, F</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF445833332A823EB1E" box="[716,788,573,594]" captionStart="MAP" captionStartId="176.[155,174,1068,1087]" captionTargetBox="[138,1176,216,1047]" captionTargetId="figure-26@176.[648,1176,216,1056]" captionTargetPageId="176" captionText="MAP 1. Distributions of Uduba jayjay, Uduba ibonia MAP 2. Distributions of Uduba salegy, Uduba platnicki, and Uduba hainteny. Uduba halabe, Uduba funerea Simon 1906 and Uduba dahli (Simon 1903)." figureDoi="http://doi.org/10.5281/zenodo.13158773" httpUri="https://zenodo.org/record/13158773/files/figure.png" pageId="46" pageNumber="47">Maps 1</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF4446F3332A80DEB1E" box="[800,826,573,594]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="46" pageNumber="47">13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186AFFF447E4336DA855EBB6" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="733CE225186AFFF447E4336DA855EBB6" blockId="46.[122,1174,610,1725]" pageId="46" pageNumber="47">
<emphasis id="41F73E37186AFFF447E4336DAA67EB35" bold="true" box="[171,336,610,633]" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation id="C3EBE878186AFFF44618336DA9C4EBF5" collectingDate="1988-10-22" collectingDateMax="1988-10-31" collectingDateMin="1988-10-22" collectionCode="USNM" collectorName="W. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="46" pageNumber="47" specimenCode="CASENT9006038" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C87186AFFF44618336DAA8DEB35" box="[343,442,610,633]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E186AFFF4454A336DA9FAEB35" box="[517,717,610,633]" collectionCode="CASENT" pageId="46" pageNumber="47">CASENT9006038</specimenCode>
) from montane rainforest at
<quantity id="B47B4FC0186AFFF4434B336DAF7AEB35" box="[1028,1101,610,633]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="46" pageNumber="47" unit="m" value="1100.0">
<elevation id="F8AE0516186AFFF4434B336DAF7AEB35" box="[1028,1101,610,633]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="46" pageNumber="47" unit="m" value="1100.0">1100m</elevation>
</quantity>
elevation at
<quantity id="B47B4FC0186AFFF44787338DABC8EBD5" box="[200,255,642,665]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="46" pageNumber="47" unit="km" value="7.0">7 km</quantity>
W
<location id="765CB4FE186AFFF44668338DAA86EBD5" LSID="urn:lsid:plazi:treatment:FB2A5333186AFFEA47343312A8C5EBB8:765CB4FE186AFFF44668338DAA86EBD5" box="[295,433,642,665]" country="Madagascar" name="Ranomafana" pageId="46" pageNumber="47" stateProvince="Fianarantsoa">Ranomafana</location>
, ca. 21º12ʹS, 47º27ʹE,
<collectingRegion id="B1472CC7186AFFF445E7338DA8ACEBD5" box="[680,923,642,665]" country="Madagascar" name="Fianarantsoa" pageId="46" pageNumber="47">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B5186AFFF444E8338DAF11EBD5" box="[935,1062,642,665]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
, collected
<date id="073DC4E5186AFFF4473433ADAA69EBF5" box="[123,350,674,697]" pageId="46" pageNumber="47" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">
<collectingDate id="17793D0D186AFFF4473433ADAA69EBF5" box="[123,350,674,697]" pageId="46" pageNumber="47" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">22–31 October 1988</collectingDate>
</date>
,
<collectorName id="DE7687F3186AFFF4462433ADAAEEEBF5" box="[363,473,674,697]" pageId="46" pageNumber="47">W. Steiner</collectorName>
, deposited in the
<collectionCode id="15927AE0186AFFF445ED33ADA9C4EBF5" box="[674,755,674,697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">USNM</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE878186AFFF445B033ADA869EBB6" collectingDate="1998-04-11" collectionCode="CAS" collectorName="D. Kavanaugh" country="Madagascar" county="Talatakely" location="Parc National Ranomafana" municipality="Ranomafana" pageId="46" pageNumber="47" specimenCode="CASENT9006035" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="AC385C87186AFFF445B033ADA86BEBF5" box="[767,860,674,697]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E186AFFF444F033ADAFBDEBF5" box="[959,1162,674,697]" collectionCode="CASENT" pageId="46" pageNumber="47">CASENT9006035</specimenCode>
), from
<location id="765CB4FE186AFFF447F833CCAAE8EB96" LSID="urn:lsid:plazi:treatment:FB2A5333186AFFEA47343312A8C5EBB8:765CB4FE186AFFF447F833CCAAE8EB96" box="[183,479,707,730]" country="Madagascar" county="Talatakely" municipality="Ranomafana" name="Parc National Ranomafana" pageId="46" pageNumber="47" stateProvince="Fianarantsoa">
Parc National
<collectingMunicipality id="9358785F186AFFF4461A33CCAAE8EB96" box="[341,479,707,730]" pageId="46" pageNumber="47">Ranomafana</collectingMunicipality>
</location>
,
<collectingCounty id="9A5D9AA9186AFFF446A533CCA964EB96" box="[490,595,707,730]" pageId="46" pageNumber="47">Talatakely</collectingCounty>
, 21º14.9ʹS, 47º25.6ʹE,
<collectingRegion id="B1472CC7186AFFF4441F33CCAF74EB96" box="[848,1091,707,730]" country="Madagascar" name="Fianarantsoa" pageId="46" pageNumber="47">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B5186AFFF4431F33CCAB8AEBB6" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
, collected
<date id="073DC4E5186AFFF4467C33ECAAF0EBB6" box="[307,455,739,762]" pageId="46" pageNumber="47" value="1998-04-11">
<collectingDate id="17793D0D186AFFF4467C33ECAAF0EBB6" box="[307,455,739,762]" pageId="46" pageNumber="47" value="1998-04-11">11 April 1998</collectingDate>
</date>
by
<collectorName id="DE7687F3186AFFF446BD33ECA9A7EBB6" box="[498,656,739,762]" pageId="46" pageNumber="47">D. Kavanaugh</collectorName>
, deposited in
<collectionCode id="15927AE0186AFFF4446833ECA869EBB6" box="[807,862,739,762]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="46" pageNumber="47" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186AFFF447E4320CAF42EA76" pageId="46" pageNumber="47" type="etymology">
<paragraph id="733CE225186AFFF447E4320CAF42EA76" blockId="46.[122,1174,610,1725]" pageId="46" pageNumber="47">
<emphasis id="41F73E37186AFFF447E4320CAA1DEA56" bold="true" box="[171,298,771,794]" pageId="46" pageNumber="47">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E37186AFFF44540320CA96EEA56" box="[527,601,771,794]" italics="true" pageId="46" pageNumber="47">ibonia,</emphasis>
refers to an epic poem that has been told for at least several hundred years in various forms across the island of
<collectingCountry id="0B94A2B5186AFFF4444E322CA8B2EA76" box="[769,901,803,826]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186AFFF447E4324CA867ED17" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="733CE225186AFFF447E4324CA867ED17" blockId="46.[122,1174,610,1725]" pageId="46" pageNumber="47">
<emphasis id="41F73E37186AFFF447E4324CAAF7EA16" bold="true" box="[171,448,835,858]" pageId="46" pageNumber="47">
Diagnosis.
<taxonomicName id="B48399A6186AFFF44667324CAAF7EA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,448,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ibonia">
<emphasis id="41F73E37186AFFF44667324CAAF7EA16" bold="true" box="[296,448,835,858]" italics="true" pageId="46" pageNumber="47">Uduba ibonia</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.a, the
<taxonomicName id="B48399A6186AFFF44597324CA86BEA16" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[728,860,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA0186AFFF444FC324CAF3EEA16" box="[947,1033,835,858]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="46" pageNumber="47">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are cribellate, with a very small divided cribellum (
<figureCitation id="EBB8FEA0186AFFF44353326CAFBFEA36" box="[1052,1160,867,890]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="46" pageNumber="47">Fig. 20 B</figureCitation>
). Males can be distinguished from those of other cribellate
<taxonomicName id="B48399A6186AFFF445A8328CA806EAD6" authorityName="Simon" authorityYear="1880" box="[743,817,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37186AFFF445A8328CA806EAD6" bold="true" box="[743,817,899,922]" italics="true" pageId="46" pageNumber="47">Uduba</emphasis>
</taxonomicName>
by having the palpus with TA3 a slender, pointed screw (
<figureCitation id="EBB8FEA0186AFFF4463932ACAAEEEAF6" box="[374,473,931,954]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="46" pageNumber="47">Fig. 29 F</figureCitation>
) and the TA2 an elongate blade (
<figureCitation id="EBB8FEA0186AFFF4447832ACA8A8EAF6" box="[823,927,931,954]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="46" pageNumber="47">Figs. 29 F</figureCitation>
,
<figureCitation id="EBB8FEA0186AFFF444E332ACA8CBEAF6" box="[940,1020,931,954]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">45 A, B</figureCitation>
<emphasis id="41F73E37186AFFF444B332ACAF3BEAF6" bold="true" box="[1020,1036,931,954]" pageId="46" pageNumber="47">).</emphasis>
The females can be distinguished from those of other cribellate
<taxonomicName id="B48399A6186AFFF445D532CCA9D3EA96" authorityName="Simon" authorityYear="1880" box="[666,740,963,986]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37186AFFF445D532CCA9D3EA96" bold="true" box="[666,740,963,986]" italics="true" pageId="46" pageNumber="47">Uduba</emphasis>
</taxonomicName>
that have an epigynal plate with median lobe surrounded by depressed atrium by having the posterior margin of atrium divided at ML apex, atrium with atrial side plate (AtSp) next to ML (
<figureCitation id="EBB8FEA0186AFFF44532350BA9D2ED57" box="[637,741,1028,1051]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="46" pageNumber="47">Fig. 73 B</figureCitation>
), ML short, broad, length about equal to width;
<taxonomicName id="B48399A6186AFFF44787352BAA06ED77" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[200,305,1060,1083]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ibonia">
<emphasis id="41F73E37186AFFF44787352BAA06ED77" bold="true" box="[200,305,1060,1083]" italics="true" pageId="46" pageNumber="47">U. ibonia</emphasis>
</taxonomicName>
females are distinguished from those of
<taxonomicName id="B48399A6186AFFF445B9352BA84EED77" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[758,889,1060,1083]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="41F73E37186AFFF445B9352BA84EED77" bold="true" box="[758,889,1060,1083]" italics="true" pageId="46" pageNumber="47">U. platnicki</emphasis>
</taxonomicName>
in that the former have a divided cribellum (
<figureCitation id="EBB8FEA0186AFFF44606354BAA84ED17" box="[329,435,1092,1115]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="46" pageNumber="47">Fig. 20 B</figureCitation>
), whereas
<taxonomicName id="B48399A6186AFFF44568354BA99EED17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[551,681,1092,1115]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="41F73E37186AFFF44568354BA99EED17" bold="true" box="[551,681,1092,1115]" italics="true" pageId="46" pageNumber="47">U. platnicki</emphasis>
</taxonomicName>
are ecribellate.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186AFFEA47E4356BA9CCEBF8" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="733CE225186AFFF547E4356BAF7DED35" blockId="46.[122,1174,610,1725]" lastBlockId="47.[122,1174,220,1725]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="41F73E37186AFFF447E4356BAA4FED37" bold="true" box="[171,376,1124,1147]" pageId="46" pageNumber="47">Description. Male</emphasis>
(
<typeStatus id="AC385C87186AFFF446C7356BAAC6ED37" box="[392,497,1124,1147]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
). Total length 14.29. Markings as in
<figureCitation id="EBB8FEA0186AFFF444CB356BA8DDED37" box="[900,1002,1124,1147]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="46" pageNumber="47">Figs. 9 E</figureCitation>
,
<specimenCount id="658529AC186AFFF444B8356BAF1DED37" box="[1015,1066,1124,1147]" count="11" pageId="46" pageNumber="47" type="generic">
<figureCitation id="EBB8FEA0186AFFF444B8356BAF14ED37" box="[1015,1059,1124,1147]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne d’Ambre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route d’Anosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="46" pageNumber="47">11 F</figureCitation>
.
</specimenCount>
Carapace 8.36 long, 6.07 wide, 2.50 high; clypeus 0.43 high. Eye diameters: AME 0.34, ALE 0.32, PME 0.36, PLE 0.43. Chelicerae 3.79 long; sternum 3.86 long, 2.93 wide; labium 1.82 long; palpal coxae 2.75 long. Cribellum divided, very small. Spination: palpus–femur d0-0-1, p0-0-1, r0-0-1, tibia p1-0; leg I–femur d1-1-0-1, p0-0-0-2, r0-0-0-1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2-2-3, r1-1-0; leg II–femur d1-1-0-1, p0-0-1, r0-1-0-1, tibia d0-0-0-1, p0-1-1- 0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2-2-3, r1-1-0; leg III–femur d1-1-1, p0-1-1-1, r0-1-1-0, tibia d0-0-1-0, p0-1-1-0, v2-0-2, r0-1-1-0, metatarsus p1-1-1-1, v2-2-2, r1-1-0-1; leg IV–femur d1- 1-0-1, p0-1-0-1, r0-0-0-1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r2- 1-2-1-2. Scopulae: cymbium, apicodorsal; strong beneath all tarsi. Leg measurements: I: 7.64 + 3.00 + 7.71 + 8.43 + 4.29 = 31.01; II: 7.00 + 2.86 + 6.36 + 7.21 + 3.79 = 27.21; III: 5.86 + 2.71 + 4.00 + 6.07 + 3.01 = 21.71; IV: 8.14 + 2.86 + 7.14 + 9.71 + 4.14 = 32.00; palpus: 3.14 + 1.29 + 1.43 + NA + 3.86 = 9.71. Leg formula 4123.
<emphasis id="41F73E37186AFFF445EC34EAA82DECB0" bold="true" box="[675,794,1509,1532]" pageId="46" pageNumber="47">Male palp</emphasis>
(Ranomafana, CASENT9006036) (
<figureCitation id="EBB8FEA0186AFFF447CC370AAA23EF50" box="[131,276,1541,1564]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">Figs. 45 A–C</figureCitation>
): palpal tibia 0.38 times cymbial length, RTA a short triangle (
<figureCitation id="EBB8FEA0186AFFF444EB370AAF3DEF50" box="[932,1034,1541,1564]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">Fig. 45 A</figureCitation>
), length 0.21 tibia length, length 0.86 times width, with pointed apex, VTA conical, with broad base and slender apex (
<figureCitation id="EBB8FEA0186AFFF447F33749AA11EF11" box="[188,294,1606,1629]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">Fig. 45 B</figureCitation>
), length 0.31 tibia width, VTA length 1.60 times width; tibia with a long, slender prolateral spine but lacking stout spines; tegulum convex, without ridge, tegulum length 0.90 times width (
<figureCitation id="EBB8FEA0186AFFF447873789AA6BEFD1" box="[200,348,1670,1693]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="46" pageNumber="47">Figs. 45 A, B</figureCitation>
), TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.285 times tegulum length; TA3 a slender, pointed screw, TA3 ends before TA2 apex, TA2 an elongate triangular blade, visible behind and beyond TA3, apex rounded (
<figureCitation id="EBB8FEA0186BFFF5446231D3A8A4E9BF" box="[813,915,220,243]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="47" pageNumber="48">Fig. 29 F</figureCitation>
); MA simple with small prolateral hook (
<figureCitation id="EBB8FEA0186BFFF5467D31F3AAA9E85F" box="[306,414,252,275]" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="47" pageNumber="48">Fig. 45 A</figureCitation>
), origin at 0.46 times tegulum length, tegulum apex at 0.29 tegulum length from MA apex, length (including apical processes) 0.78 times width, MA small (
<figureCitation id="EBB8FEA0186BFFF5436F3013ABB8E81F" captionStart="FIGURE" captionStartId="141.[155,167,1589,1608]" captionTargetBox="[130,1162,965,1564]" captionTargetId="figure-60@141.[120,1176,950,1576]" captionTargetPageId="141" captionText="FIGURE 45. Uduba ibonia, new species, (CASENT9006036), from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. C, conductor; Cm, cymbium; E, embolus; Eb, embolic base; MA, median apophysis; RTA, retrolateral tibial apophysis on male palp, typically arising apically; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158673" httpUri="https://zenodo.org/record/13158673/files/figure.png" pageId="47" pageNumber="48">Figs. 45 A, C</figureCitation>
), length 0.20 tegulum length, width 0.23 tegulum width; conductor fan concave.
<emphasis id="41F73E37186BFFF5430D3033AFA2E81F" bold="true" box="[1090,1173,316,339]" pageId="47" pageNumber="48">Female</emphasis>
(
<typeStatus id="AC385C87186BFFF547CD3053ABD3E83F" box="[130,228,348,371]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
, CASENT9006035): Total length 12.50. Markings as in
<figureCitation id="EBB8FEA0186BFFF5441B3053A89DE83F" box="[852,938,348,371]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="47" pageNumber="48">
Fig.
<specimenCount id="658529AC186BFFF544C73053A89DE83F" box="[904,938,348,371]" count="7" pageId="47" pageNumber="48" type="generic">7 F</specimenCount>
</figureCitation>
. Carapace 6.70 long, 5.50 wide, 2.50 high; clypeus 0.45 high. Eye diameters:AME and ALE 0.24, PME 0.22, PLE 0.26. Chelicerae 3.20 long; sternum 2.80 long, 2.30 wide; labium 1.50 long; palpal coxae 2.30 long. Cribellum divided, very small (
<figureCitation id="EBB8FEA0186BFFF5468230B2A903E898" box="[461,564,445,468]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="47" pageNumber="48">Fig. 20 B</figureCitation>
<emphasis id="41F73E37186BFFF5457B30B2A90AE898" bold="true" box="[564,573,445,468]" pageId="47" pageNumber="48">)</emphasis>
. Spination: palpus–femur d0-0-2, patella p1, tibia p2-1, tarsus p2-1-1; leg I–femur d1-0-0-2, tibia v2-2-2-1, metatarsus v2-2-3; leg II–femur d1-0-0, p0-0- 1, tibia v1-1-1-2, metatarsus v2-2-3; leg III–femur d0-0-0-1, p0-0-1-1, r0-0-1-1, tibia d0-1-0, p0- 1-1-0, v2-0-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-0, r0- 0-1, tibia v1-2-2, r0-1-1-0, metatarsus p1-1-0-2, v2-1-2-2, r1-2-1. Scopulae: strong ventral beneath all tarsi, beneath metatarsi I and II and apices of metatarsi III and IV. Leg measurements: I: 4.70 + 2.30 + 3.90 + 3.70 + 2.30 = 16.90; II: 4.20 + 2.20 + 3.20 + 3.10 + 2.10 = 14.80; III: 3.40 + 1.90 + 2.00 + 2.80 + 1.90 = 12.00; IV: 4.80 + 2.00 + 3.80 + 4.90 + 2.40 = 17.90; palpus: 2.40 + 1.20 + 1.40 + NA + 2.30 = 7.30. Leg formula 4123.
<emphasis id="41F73E37186BFFF545CA33B0A875EB9A" bold="true" box="[645,834,703,726]" pageId="47" pageNumber="48">Female genitalia</emphasis>
(
<figureCitation id="EBB8FEA0186BFFF5441D33B0A8D0EB9A" box="[850,999,703,726]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="47" pageNumber="48">Figs. 73 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0186BFFF544BB33B0AF7DEB9A" box="[1012,1098,703,726]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="47" pageNumber="48">81 C, F</figureCitation>
): epigynal plate with median lobe surrounded by depressed atrium, shallow beside ML and forming deep sulci laterally (
<figureCitation id="EBB8FEA0186BFFF5465133F0AABDEA5A" box="[286,394,767,790]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="47" pageNumber="48">Fig. 73 B</figureCitation>
), epigynal plate width 1.43 times length; atrium sides weakly concave, atrium width 1.87 times atrium length; epigynum length 1.79 times atrium length, atrium width 0.73 times epigynum width; atrium plate width at side of ML 0.18 times ML width (
<figureCitation id="EBB8FEA0186BFFF54352324FAFBFEA1B" box="[1053,1160,832,855]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="47" pageNumber="48">Fig. 81 C</figureCitation>
), atrium lateral sulcus width at side of ML 0.21 times ML width; atrium and median lobe arise well anteriad of epigastric groove (
<figureCitation id="EBB8FEA0186BFFF5468A328FA918EADB" box="[453,559,896,919]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="47" pageNumber="48">Fig. 81 C</figureCitation>
<emphasis id="41F73E37186BFFF54560328FA90FEADB" bold="true" box="[559,568,896,919]" pageId="47" pageNumber="48">)</emphasis>
, atrium origin at 0.26 of epigynal plate length, median lobe broad, widest at base, median lobe length 1.08 times width, ML does not extend to atrium apex, atrium extends 0.20 times epigynum length anteriad of ML, ML length 0.38 times epigynal length; lateral lobes narrow, atrium width 7.30 times LL width; copulatory openings beneath the atrial side plate (
<emphasis id="41F73E37186BFFF54660350EAA56ED54" box="[303,353,1025,1048]" italics="true" pageId="47" pageNumber="48">AtSp</emphasis>
) next to ML (
<figureCitation id="EBB8FEA0186BFFF546B6350EA96AED54" box="[505,605,1025,1048]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="47" pageNumber="48">Fig. 73 A</figureCitation>
<emphasis id="41F73E37186BFFF54511350EA950ED54" bold="true" box="[606,615,1025,1048]" pageId="47" pageNumber="48">)</emphasis>
, distance between CO 0.35 times width of epigynal plate, CO distance from epigastric furrow 0.64 times epigynum length. Vulva (
<figureCitation id="EBB8FEA0186BFFF54497352EAF7AED74" box="[984,1101,1057,1080]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="47" pageNumber="48">Figs. 73 A</figureCitation>
,
<figureCitation id="EBB8FEA0186BFFF54315352EAFBAED74" box="[1114,1165,1057,1080]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="47" pageNumber="48">81 F</figureCitation>
<emphasis id="41F73E37186BFFF543C3352EAFA2ED74" bold="true" box="[1164,1173,1057,1080]" pageId="47" pageNumber="48">)</emphasis>
with spermathecal ducts making three to four, transverse curves, vulva length 0.82 times width, fertilization ducts close-together, vulva width 8.80 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE225186BFFF547E4358DA9DAEFF1" blockId="47.[122,1174,220,1725]" pageId="47" pageNumber="48">
<emphasis id="41F73E37186BFFF547E4358DAA6BEDD5" bold="true" box="[171,348,1154,1177]" pageId="47" pageNumber="48">
Variation.
<emphasis id="41F73E37186BFFF5466B358DAA6BEDD5" bold="true" box="[292,348,1154,1177]" italics="true" pageId="47" pageNumber="48">Male</emphasis>
</emphasis>
(N = 4): Total length = 11.43–14.29, carapace length / width = 1.38–1.53, carapace height / width = 0.45–0.59, PER / carapace width = 0.38–0.40, PER / OAL = 2.33–3.06, PER / AER = 1.32–1.42, OAL / OQL = 1.00–1.08, OQP / OQA = 1.13–1.28, clypeus height / AME diameter width = 1.26–1.56, cheliceral length / clypeus height = 6.58–8.91, sternum length / width = 1.24–1.31, palpal coxa length / width = 2.07–2.27, femur I length / carapace width = 1.26–1.39, metatarsus I length / carapace width = 1.39–1.55, femur IV length / carapace width = 1.34–1.59, cymbium length / carapace width = 0.63–0.85, cymbium length / palpal patella length = 2.99–3.87, cymbium length / palpal tibia length = 2.70–3.87, cymbium length / palpal femur length = 1.23– 1.61, palpal tibia length / palpal patella length = 1.00–1.11.
<emphasis id="41F73E37186BFFF545B5348BA87AECD7" bold="true" box="[762,845,1412,1435]" italics="true" pageId="47" pageNumber="48">Female</emphasis>
(N = 2): Total length = 11.80– 14.50, carapace length / width = 1.38–1.53, carapace height / width = 0.41–0.67, PER / carapace width = 0.44–0.68, PER / OAL = 2.82–3.56, PER / AER = 1.36–1.44, OAL / OQL = 1.00–1.10, OQP / OQA = 1.17–1.28, clypeus height / AME = 1.26–2.55, cheliceral length / clypeus height = 7.11–8.83, sternum length / width = 1.22–1.32, palpal coxa length / width = 1.89–2.26, femur I length / carapace width = 1.00–1.04, metatarsus I length / carapace width = 0.80–0.82, femur IV length / carapace width = 1.00–1.08, palpal tarsus length / carapace width = 0.49–0.51, palpal tarsus length / palpal patella length = 1.80–1.92, palpal tarsus length / palpal tibia length = 1.60– 1.71, palpal tarsus length / palpal femur length = 0.95–1.09, palpal tibia length / palpal patella length = 1.05–1.17. Genitalia as in
<figureCitation id="EBB8FEA0186BFFF546B537A9A9BBEFF1" box="[506,652,1702,1725]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="47" pageNumber="48">Figs. 73 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0186BFFF545D737A9A9DAEFF1" box="[664,749,1702,1725]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="47" pageNumber="48">81 C, F.</figureCitation>
</paragraph>
<paragraph id="733CE2251874FFEA47E431D3A94DEB18" blockId="48.[123,1174,220,756]" pageId="48" pageNumber="49">
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M
<emphasis id="41F73E371874FFEA478D31D3AABAE9BF" bold="true" box="[194,397,220,243]" pageId="48" pageNumber="49">aterial examined.</emphasis>
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:
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<collectingRegion id="B1472CC71874FFEA452831D3A855E9BF" box="[615,866,220,243]" country="Madagascar" name="Fianarantsoa" pageId="48" pageNumber="49">Fianarantsoa Province</collectingRegion>
</emphasis>
:
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W Ranomafana
</location>
, ca. 21º12ʹS, 47º27ʹE, elev.
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, W.
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(
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,
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,
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,
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,
<materialsCitation id="C3EBE8781874FFEA47913013A8BAE87F" box="[222,909,284,307]" collectingDate="1988-10-08" collectingDateMax="1988-10-21" collectingDateMin="1988-10-08" collectionCode="USNM" collectorName="Steiner" country="Madagascar" pageId="48" pageNumber="49" specimenCode="CASENT9006037" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<date id="073DC4E51874FFEA47913013AA9CE87F" box="[222,427,284,307]" pageId="48" pageNumber="49" value="1988-10-08" valueMax="1988-10-21" valueMin="1988-10-08">
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</date>
, W.
<collectorName id="DE7687F31874FFEA46973013A913E87F" box="[472,548,284,307]" pageId="48" pageNumber="49">Steiner</collectorName>
(
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,
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</materialsCitation>
,
<materialsCitation id="C3EBE8781874FFEA44D63013A8C1E81F" collectingDate="1988-11-01" collectingDateMax="1988-11-07" collectingDateMin="1988-11-01" collectionCode="USNM" collectorName="Steiner" country="Madagascar" pageId="48" pageNumber="49" specimenCode="CASENT9006036" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
“island in stream; montane rainforest,”
<date id="073DC4E51874FFEA46603033A927E81F" box="[303,528,316,339]" pageId="48" pageNumber="49" value="1988-11-01" valueMax="1988-11-07" valueMin="1988-11-01">
<collectingDate id="17793D0D1874FFEA46603033A927E81F" box="[303,528,316,339]" pageId="48" pageNumber="49" value="1988-11-01" valueMax="1988-11-07" valueMin="1988-11-01">1–7 November 1988</collectingDate>
</date>
, W.
<collectorName id="DE7687F31874FFEA45723033A9BEE81F" box="[573,649,316,339]" pageId="48" pageNumber="49">Steiner</collectorName>
(
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,
<specimenCount id="658529AC1874FFEA44223033A8B9E81F" box="[877,910,316,339]" count="1" pageId="48" pageNumber="49" type="male">1♂</specimenCount>
,
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)
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;
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<collectingCounty id="9A5D9AA91874FFEA434D3033AA1CE83F" pageId="48" pageNumber="49">Parc National de Ranomafana</collectingCounty>
,
<collectingMunicipality id="9358785F1874FFEA46733053AAFAE83F" box="[316,461,348,371]" pageId="48" pageNumber="49">Vatoharanana</collectingMunicipality>
, 4.0 km SW
<location id="765CB4FE1874FFEA453C3053A9CDE83F" LSID="urn:lsid:plazi:treatment:FB2A5333186AFFEA47343312A8C5EBB8:765CB4FE1874FFEA453C3053A9CDE83F" box="[627,762,348,371]" country="Madagascar" county="Parc National de Ranomafana" municipality="Vatoharanana" name="Ranomafana" pageId="48" pageNumber="49" stateProvince="Fianarantsoa">Ranomafana</location>
(ville), 21°14.7ʹS, 047°26.0ʹE, elev.
<quantity id="B47B4FC01874FFEA47343073ABFFE8DF" box="[123,200,380,403]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="48" pageNumber="49" unit="m" value="1025.0">
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, “
<collectingMethod id="AAC29A321874FFEA47903073AA19E8DF" box="[223,302,380,403]" pageId="48" pageNumber="49">pitfalls</collectingMethod>
, slightly transitional lowland to lower montane rainforest”,
<date id="073DC4E51874FFEA44883073AFA6E8DF" box="[967,1169,380,403]" pageId="48" pageNumber="49" value="2000-10-03" valueMax="2000-10-09" valueMin="2000-10-03">
<collectingDate id="17793D0D1874FFEA44883073AFA6E8DF" box="[967,1169,380,403]" pageId="48" pageNumber="49" value="2000-10-03" valueMax="2000-10-09" valueMin="2000-10-03">3–9 October 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31874FFEA47343093AA3BE8FF" box="[123,268,412,435]" pageId="48" pageNumber="49">S. Goodman</collectorName>
[
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#00-225], (
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), (
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,
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)
</materialsCitation>
;
<materialsCitation id="C3EBE8781874FFEA47AD30B3A8A2E8BF" collectingDate="1998-04-11" collectionCode="CAS" collectorName="D. Kavanaugh" country="Madagascar" location="Talatakely" municipality="Parc National Ranomafana" pageId="48" pageNumber="49" specimenCode="CASENT9006035" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingMunicipality id="9358785F1874FFEA47AD30B3A93BE89F" box="[226,524,444,467]" pageId="48" pageNumber="49">Parc National Ranomafana</collectingMunicipality>
,
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, 21º14.9ʹS, 47º25.6ʹE, “walking on ground”,
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<collectingDate id="17793D0D1874FFEA433230B3ABCCE8BF" pageId="48" pageNumber="49" value="1998-04-11">11 April 1998</collectingDate>
</date>
,
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(
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,
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,
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,
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)
</materialsCitation>
;
<materialsCitation id="C3EBE8781874FFEA44E730D3AB8CEB78" collectingDate="1998-11-02" collectionCode="CAS" collectorName="V. F. Lee" country="Madagascar" elevation="1200" location="Talatakely" pageId="48" pageNumber="49" specimenCode="CASENT9006034" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="765CB4FE1874FFEA44E730D3AF26E8BF" LSID="urn:lsid:plazi:treatment:FB2A5333186AFFEA47343312A8C5EBB8:765CB4FE1874FFEA44E730D3AF26E8BF" box="[936,1041,476,499]" country="Madagascar" name="Talatakely" pageId="48" pageNumber="49" stateProvince="Fianarantsoa">Talatakely</location>
, 21º14.9ʹS, 47º25.6ʹE, elev. ca.
<quantity id="B47B4FC01874FFEA461730F2AA92EB58" box="[344,421,509,532]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="48" pageNumber="49" unit="m" value="1200.0">
<elevation id="F8AE05161874FFEA461730F2AA92EB58" box="[344,421,509,532]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="48" pageNumber="49" unit="m" value="1200.0">1200m</elevation>
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, “under log”,
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<collectingDate id="17793D0D1874FFEA450830F2A824EB58" box="[583,787,509,532]" pageId="48" pageNumber="49" value="1998-11-02">2 November 1998</collectingDate>
</date>
,
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(
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,
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,
<collectionCode id="15927AE01874FFEA47343312AB84EB78" box="[123,179,541,564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="48" pageNumber="49" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781874FFEA478B3312A941EB18" collectingDate="1998-10-30" collectingDateMax="1998-11-20" collectingDateMin="1998-10-30" collectionCode="CAS" collectorName="V. F. Lee &amp; K. Ribardo" country="Madagascar" elevation="1008" location="Talatakely" pageId="48" pageNumber="49" specimenCode="CASENT9006033" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="765CB4FE1874FFEA478B3312AA1AEB78" LSID="urn:lsid:plazi:treatment:FB2A5333186AFFEA47343312A8C5EBB8:765CB4FE1874FFEA478B3312AA1AEB78" box="[196,301,541,564]" country="Madagascar" name="Talatakely" pageId="48" pageNumber="49" stateProvince="Fianarantsoa">Talatakely</location>
, 21º15ʹS, 47º26ʹE, elev. ca.
<quantity id="B47B4FC01874FFEA451A3312A9E1EB78" box="[597,726,541,564]" metricMagnitude="3" metricUnit="m" metricValue="1.0074999999999998" metricValueMax="1.1" metricValueMin="0.915" pageId="48" pageNumber="49" unit="m" value="1007.5" valueMax="1100.0" valueMin="915.0">
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,
<date id="073DC4E51874FFEA45AF3312AF03EB78" box="[736,1076,541,564]" pageId="48" pageNumber="49" value="1998-10-30" valueMax="1998-11-20" valueMin="1998-10-30">
<collectingDate id="17793D0D1874FFEA45AF3312AF03EB78" box="[736,1076,541,564]" pageId="48" pageNumber="49" value="1998-10-30" valueMax="1998-11-20" valueMin="1998-10-30">30 October–20 November 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31874FFEA43723312AFA2EB78" box="[1085,1173,541,564]" pageId="48" pageNumber="49">V. F. Lee</collectorName>
and
<collectorName id="DE7687F31874FFEA47E63332AA16EB18" box="[169,289,573,596]" pageId="48" pageNumber="49">K. Ribardo</collectorName>
(
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,
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,
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.
</paragraph>
<paragraph id="733CE2251874FFEA47E43352A9CCEBF8" blockId="48.[123,1174,220,756]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA47E43352AA57EB38" bold="true" box="[171,352,605,628]" pageId="48" pageNumber="49">Natural history.</emphasis>
The species
<taxonomicName id="B48399A61874FFEA46A13352A9B1EB38" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[494,646,605,628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ibonia">
<emphasis id="41F73E371874FFEA46A13352A9B1EB38" bold="true" box="[494,646,605,628]" italics="true" pageId="48" pageNumber="49">Uduba ibonia</emphasis>
</taxonomicName>
lives in lowland, premontane and montane rainforest. Collection records suggest that this species may wander; the presence of a calamistrum and a very small cribellum suggests some use of cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEA47E433B2A8C5EBB8" pageId="48" pageNumber="49" type="distribution">
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<emphasis id="41F73E371874FFEA47E433B2AA0AEB98" bold="true" box="[171,317,701,724]" pageId="48" pageNumber="49">Distribution.</emphasis>
Specimens of
<taxonomicName id="B48399A61874FFEA469033B2A94FEB98" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[479,632,701,724]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ibonia">
<emphasis id="41F73E371874FFEA469033B2A94FEB98" bold="true" box="[479,632,701,724]" italics="true" pageId="48" pageNumber="49">Uduba ibonia</emphasis>
</taxonomicName>
are known only from the vicinity of Ranomafana National Park in
<collectingRegion id="B1472CC71874FFEA467B33D2A914EBB8" box="[308,547,733,756]" country="Madagascar" name="Fianarantsoa" pageId="48" pageNumber="49">Fianarantsoa Province</collectingRegion>
of southeastern
<collectingCountry id="0B94A2B51874FFEA459833D2A86FEBB8" box="[727,856,733,756]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA01874FFEA442833D2A88EEBB8" box="[871,953,733,756]" captionStart="MAP" captionStartId="176.[155,174,1068,1087]" captionTargetBox="[138,1176,216,1047]" captionTargetId="figure-26@176.[648,1176,216,1056]" captionTargetPageId="176" captionText="MAP 1. Distributions of Uduba jayjay, Uduba ibonia MAP 2. Distributions of Uduba salegy, Uduba platnicki, and Uduba hainteny. Uduba halabe, Uduba funerea Simon 1906 and Uduba dahli (Simon 1903)." figureDoi="http://doi.org/10.5281/zenodo.13158773" httpUri="https://zenodo.org/record/13158773/files/figure.png" pageId="48" pageNumber="49">Maps 1</figureCitation>
,
<figureCitation id="EBB8FEA01874FFEA448733D2A8D2EBB8" box="[968,997,733,756]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="48" pageNumber="49">13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

240
data/FB/2A/53/FB2A5333186FFFF647343033AAFAE83F.xml Normal file
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@ -0,0 +1,240 @@
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<mods:title id="3E7FD36F95740A8BDD874B989F18C719">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="F24E1124C4CC05E5B39EF9DB3322D494">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<emphasis id="41F73E37186FFFF147343033AA2EE81F" bold="true" box="[123,281,316,339]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C186FFFF146683033AA9CE81F" box="[295,427,316,339]" pageId="43" pageNumber="44" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF147343053AADCE83D" box="[123,491,348,369]" pageId="43" pageNumber="44" type="description">
<paragraph id="733CE225186FFFF147343053AADCE83D" blockId="43.[123,491,316,369]" box="[123,491,348,369]" pageId="43" pageNumber="44">
<figureCitation id="EBB8FEA0186FFFF147343053AA3CE83D" box="[123,267,348,369]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="43" pageNumber="44">Figures 5 D, E</figureCitation>
,
<figureCitation id="EBB8FEA0186FFFF1465A3053AA55E83D" box="[277,354,348,369]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">72 A, B</figureCitation>
,
<figureCitation id="EBB8FEA0186FFFF146223053AAF5E83D" box="[365,450,348,369]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="43" pageNumber="44">Maps 11</figureCitation>
, 16.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF147E4308EA995E895" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="733CE225186FFFF147E4308EA995E895" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E4308EAA66E8D4" bold="true" box="[171,337,385,408]" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="C3EBE878186FFFF14616308EA995E895" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CAS" country="Madagascar" elevation="530" location="Foret Classee Andriantantely" municipality="Helian Ratsirarson" pageId="43" pageNumber="44" specimenCode="CASENT9006106" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C87186FFFF14616308EAA8BE8D4" box="[345,444,385,408]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E186FFFF14550308EA9D0E8D4" box="[543,743,385,408]" collectionCode="CASENT" pageId="43" pageNumber="44">CASENT9006106</specimenCode>
) from
<quantity id="B47B4FC0186FFFF1447B308EA846E8D4" box="[820,881,385,408]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">
<elevation id="F8AE0516186FFFF1447B308EA846E8D4" box="[820,881,385,408]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">530m</elevation>
</quantity>
elevation in
<location id="765CB4FE186FFFF1434C308EAA2CE8F5" LSID="urn:lsid:plazi:treatment:FB2A5333186FFFF647343033AAFAE83F:765CB4FE186FFFF1434C308EAA2CE8F5" country="Madagascar" municipality="Helian Ratsirarson" name="Foret Classee Andriantantely" pageId="43" pageNumber="44" stateProvince="Toamasina">Forêt Classée Andriantantely</location>
(18°41.7ʹS, 048°48.8ʹE),
<collectingRegion id="B1472CC7186FFFF1457D30ADA839E8F5" box="[562,782,418,441]" country="Madagascar" name="Toamasina" pageId="43" pageNumber="44">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B5186FFFF1445530ADA8AEE8F5" box="[794,921,418,441]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
, collected
<date id="073DC4E5186FFFF1435F30ADABEDE895" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">
<collectingDate id="17793D0D186FFFF1435F30ADABEDE895" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4–7 December 1998</collectingDate>
</date>
by
<collectingMunicipality id="9358785F186FFFF1464A30CDAAE7E895" box="[261,464,450,473]" pageId="43" pageNumber="44">Helian Ratsirarson</collectingMunicipality>
, deposited in
<collectionCode id="15927AE0186FFFF1452830CDA9A9E895" box="[615,670,450,473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="43" pageNumber="44" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF147E430EDA9E2EB75" pageId="43" pageNumber="44" type="etymology">
<paragraph id="733CE225186FFFF147E430EDA9E2EB75" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E430EDAA1DE8B5" bold="true" box="[171,298,482,505]" pageId="43" pageNumber="44">Etymology.</emphasis>
The species epithet is a patronym in honor of Mr. Helian Ratsirarson, Malagasy entomologist, Lakeside International Scholar at CAS and member of the Fisher-Griswold Arthropod Team that collected widely throughout
<collectingCountry id="0B94A2B5186FFFF1451E332DA9F8EB75" box="[593,719,546,569]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF147E4334DAB88EA56" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="733CE225186FFFF147E4334DAB88EA56" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E4334DAAFAEB15" bold="true" box="[171,461,578,601]" pageId="43" pageNumber="44">
Diagnosis.
<taxonomicName id="B48399A6186FFFF14664334DAAFAEB15" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[299,461,578,601]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="heliani">
<emphasis id="41F73E37186FFFF14664334DAAFAEB15" bold="true" box="[299,461,578,601]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
</taxonomicName>
</emphasis>
belong to Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A6186FFFF14734336DAA76EB35" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,321,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). They have a divided cribellum. Females of
<taxonomicName id="B48399A6186FFFF14483336DAF5DEB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[972,1130,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="heliani">
<emphasis id="41F73E37186FFFF14483336DAF5DEB35" bold="true" box="[972,1130,610,633]" italics="true" pageId="43" pageNumber="44">Uduba heliani</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="B48399A6186FFFF1455D338DA985EBD5" box="[530,690,642,665]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF1455D338DA96BEBD5" bold="true" box="[530,604,642,665]" italics="true" pageId="43" pageNumber="44">Uduba</emphasis>
species
</taxonomicName>
by the female genitalia (
<figureCitation id="EBB8FEA0186FFFF14488338DAF68EBD5" box="[967,1119,642,665]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Figs. 72 A, B</figureCitation>
): the epigynal plate is broadly oval (width 1.83 times length) with the copulatory openings beneath transverse grooves at mid-level of plate (
<figureCitation id="EBB8FEA0186FFFF1457933CCA9A9EB96" box="[566,670,707,730]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 A</figureCitation>
), CO origin at 0.29 times of epigynum length; vulva with spermathecal ducts forming three longitudinal loops extending anteriad of the CO (
<figureCitation id="EBB8FEA0186FFFF1432533ECAB86EA56" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF147E4322CA85CEF31" pageId="43" pageNumber="44" type="description">
<paragraph id="733CE225186FFFF147E4322CA87CEF11" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E4322CAA4BEA76" bold="true" box="[171,380,803,826]" pageId="43" pageNumber="44">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E37186FFFF14541322CA956EA76" bold="true" box="[526,609,803,826]" pageId="43" pageNumber="44">Female</emphasis>
(
<typeStatus id="AC385C87186FFFF1453A322CA9EBEA76" box="[629,732,803,826]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
, CASENT9006106): Total length 6.10. Markings as in
<figureCitation id="EBB8FEA0186FFFF14667324CAA84EA16" box="[296,435,835,858]" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="43" pageNumber="44">Figs. 5 D, E</figureCitation>
. Carapace 2.90 long, 2.00 wide, 1.20 high; clypeus 0.15 high. Eye diameters: AME 0.10, ALE and PME 0.12, PLE 0.16. Chelicerae 1.30 long; sternum 1.30 long, 1.10 wide; labium 0.60 long; palpal coxae 0.95 long.
<emphasis id="41F73E37186FFFF145FB328CA9CAEAD6" bold="true" box="[692,765,899,922]" pageId="43" pageNumber="44">Ratios</emphasis>
– carapace length / width = 1.40, carapace height / width = 1.40, PER / carapace width = 0.34, PER/OAL = 3.07, PER/AER = 1.34, OAL/OQL = 1.27, OQP/ OQA = 1.48, clypeus height / AME = 1.50, cheliceral length / clypeus height = 8.67, sternum length / width = 1.18, palpal coxa length / width = 2.11, femur I length / carapace width = 0.90, metatarsus I length / carapace width = 0.75, femur IV length / carapace width = 1.00, palpal tarsus length / carapace width = 0.50, palpal tibia length / palpal patella length = 2.00, palpal tarsus length / palpal tibia length = 2.00, palpal tarsus length / palpal femur length = 1.11, palpal tibia length / palpal patella length = 1.00. Cribellum divided. Spination: palpus– femur d0-1-0-2, patella p1-0, tibia p2-1, tarsus p2-1, r0-1-0; leg I–femur d1-0-0, tibia v1-1-1-2, metatarsus v2-2-2; leg II–femur d1-0-0, tibia v1-1-1-2, metatarsus p0-0-1, v2-2-2; leg III–femur d0-1-1-0, p0-1-1-0, r0-0-0-1, tibia d0-1-0, p1-0-1, v2-2-2, r0-1-1-0, metatarsus d0-1-0-0, p1-2-2, v2-2-2, r1-1-1; leg IV–femur d0-1-0, tibia v1-2-2, r0-1-1-0, metatarsus p0-0-1-2, v2-2-2, r0-1-1-1. Scopulae: weak ventral beneath tarsi I and II. Leg measurements: I: 1.80 + 0.90 + 1.50 + 1.40 + 1.10 = 6.70; II: 1.70 + 0.80 + 1.30 + 1.25 + 1.00 = 6.05; III: 1.25 + 0.70 + 0.80 + 1.10 + 0.60 = 4.45; IV: 2.00 + 0.80 + 1.50 + 1.60 + 1.00 = 6.90; palpus: 0.90 + 0.50 + 0.50 + NA + 1.00 = 2.90. Leg formula 4123.
<emphasis id="41F73E37186FFFF1461E346AA927EC30" bold="true" box="[337,528,1381,1404]" pageId="43" pageNumber="44">Female genitalia</emphasis>
(
<figureCitation id="EBB8FEA0186FFFF1456D346AA989EC30" box="[546,702,1381,1404]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Figs. 72 A, B</figureCitation>
): epigynal plate without depressed atrium, posterior margin at epigastric furrow convex but concave in center, plate width 1.83 times length, ML and LL not clearly defined, copulatory openings beneath transverse grooves at mid-level of plate, distance between copulatory openings 0.295 times epigynum width, CO origin at 0.29 times of epigynum length (
<figureCitation id="EBB8FEA0186FFFF1461134EAAAFFECB0" box="[350,456,1509,1532]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 A</figureCitation>
), extent of plate laterad of copulatory openings equals 0.16 times plate width, plate with weak lateral projections, lateral margins well defined. Vulva with spermathecal ducts forming three anterior, longitudinal loops (
<figureCitation id="EBB8FEA0186FFFF1444A3729A842EF71" box="[773,885,1574,1597]" captionStart="FIGURE" captionStartId="160.[155,167,1602,1621]" captionTargetBox="[138,1163,964,1577]" captionTargetId="figure-93@160.[120,1176,960,1591]" captionTargetPageId="160" captionText="FIGURE 72. Uduba spp., female genitalia drawings. A, B. U. heliani, new species, holotype (CASENT9006106) from Forêt Classée Andriantantely. C, D. U. hiragasy new species, paratype (CASENT9064772) from Tsinjoarivo Forest. E. F. U. fisheri, new species, (CASENT9065339) from 2 km W Andrambovato along River Tatamaly. G, H. U. valiha, new species, paratype (CASENT9006071) from Massif Andringitra. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158735" httpUri="https://zenodo.org/record/13158735/files/figure.png" pageId="43" pageNumber="44">Fig. 72 B</figureCitation>
), vulva length 0.80 times width, vulva width 6.25 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE225186FFFF147E43769A85CEF31" blockId="43.[123,1174,385,1725]" box="[171,875,1638,1661]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E43769AA2DEF31" bold="true" box="[171,282,1638,1661]" pageId="43" pageNumber="44">Variation.</emphasis>
Unknown: only the unique
<typeStatus id="AC385C87186FFFF145043769A99FEF31" box="[587,680,1638,1661]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
female is known.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE186FFFF647E43789A985E87F" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="733CE225186FFFF147E43789AFA2EFF1" blockId="43.[123,1174,385,1725]" pageId="43" pageNumber="44">
<emphasis id="41F73E37186FFFF147E43789AAB9EFD1" bold="true" box="[171,398,1670,1693]" pageId="43" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="C3EBE878186FFFF146D63789AFA2EFF1" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectorName="H. Ratsirarson" country="Madagascar" elevation="530" location="Foret Classee Andriantantely" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<collectingCountry id="0B94A2B5186FFFF146D63789A96CEFD1" box="[409,603,1670,1693]" name="Madagascar" pageId="43" pageNumber="44">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37186FFFF145263789A873EFD1" box="[617,836,1670,1693]" italics="true" pageId="43" pageNumber="44">
<collectingRegion id="B1472CC7186FFFF145263789A873EFD1" box="[617,836,1670,1693]" country="Madagascar" name="Toamasina" pageId="43" pageNumber="44">Toamasina Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE186FFFF1441A3789AFB9EFD1" LSID="urn:lsid:plazi:treatment:FB2A5333186FFFF647343033AAFAE83F:765CB4FE186FFFF1441A3789AFB9EFD1" box="[853,1166,1670,1693]" country="Madagascar" name="Foret Classee Andriantantely" pageId="43" pageNumber="44" stateProvince="Toamasina">Forêt Classée Andriantantely</location>
, 18°41.7ʹS, 048°48.8ʹE, elev.
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<elevation id="F8AE0516186FFFF146F237A9AACBEFF1" box="[445,508,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="m" value="530.0">530m</elevation>
</quantity>
,
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<collectingDate id="17793D0D186FFFF1454337A9A9C5EFF1" box="[524,754,1702,1725]" pageId="43" pageNumber="44" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4–7 December 1998</collectingDate>
</date>
,
<collectorName id="DE7687F3186FFFF1444C37A9A895EFF1" box="[771,930,1702,1725]" pageId="43" pageNumber="44">H. Ratsirarson</collectorName>
[#HJ-121] (
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,
</materialsCitation>
</paragraph>
<paragraph id="733CE2251868FFF6473431D3AAF5E9BF" blockId="44.[123,1174,220,371]" box="[123,450,220,243]" pageId="44" pageNumber="45">
<materialsCitation id="C3EBE8781868FFF6473431D3AA47E9BF" box="[123,368,220,243]" collectionCode="CASENT" pageId="44" pageNumber="45" specimenCode="CASENT9006106" specimenCount="1" specimenCount-female="1">
<specimenCode id="23254A5E1868FFF6473431D3AA74E9BF" box="[123,323,220,243]" collectionCode="CASENT" pageId="44" pageNumber="45">CASENT9006106</specimenCode>
,
<specimenCount id="658529AC1868FFF6460131D3AA47E9BF" box="[334,368,220,243]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
</materialsCitation>
,
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).
</paragraph>
<paragraph id="733CE2251868FFF647E431F3A985E87F" blockId="44.[123,1174,220,371]" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E431F3AA68E85F" bold="true" box="[171,351,252,275]" pageId="44" pageNumber="45">Natural history.</emphasis>
An individual of the cribellate
<taxonomicName id="B48399A61868FFF645E131F3A87AE85F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[686,845,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="heliani">
<emphasis id="41F73E371868FFF645E131F3A87AE85F" bold="true" box="[686,845,252,275]" italics="true" pageId="44" pageNumber="45">Uduba heliani</emphasis>
</taxonomicName>
has been collected in mid-elevation rainforest; no other habitat data are available.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1868FFF647E43033AAFAE83F" pageId="44" pageNumber="45" type="distribution">
<paragraph id="733CE2251868FFF647E43033AAFAE83F" blockId="44.[123,1174,220,371]" pageId="44" pageNumber="45">
<emphasis id="41F73E371868FFF647E43033AAD6E81F" bold="true" box="[171,481,316,339]" pageId="44" pageNumber="45">
Distribution.
<taxonomicName id="B48399A61868FFF6460C3033AAD6E81F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[323,481,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="heliani">
<emphasis id="41F73E371868FFF6460C3033AAD6E81F" bold="true" box="[323,481,316,339]" italics="true" pageId="44" pageNumber="45">Uduba heliani</emphasis>
</taxonomicName>
</emphasis>
are known only from the
<typeStatus id="AC385C871868FFF645B93033A813E81F" box="[758,804,316,339]" pageId="44" pageNumber="45">type</typeStatus>
locality near the east coast of central
<collectingCountry id="0B94A2B51868FFF647E93053AA10E83F" box="[166,295,348,371]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
<emphasis id="41F73E371868FFF646603053AA0FE83F" bold="true" box="[303,312,348,371]" pageId="44" pageNumber="45">(</emphasis>
<figureCitation id="EBB8FEA01868FFF646773053AAA1E83F" box="[312,406,348,371]" captionStart="MAP" captionStartId="181.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1050]" captionTargetId="figure-17@181.[648,1176,216,1036]" captionTargetPageId="181" captionText="MAP 11. Distributions of Uduba balsama and Uduba MAP 12. Distributions of Uduba lamba, Uduba lakroa heliani. and Uduba fandroana." figureDoi="http://doi.org/10.5281/zenodo.13158787" httpUri="https://zenodo.org/record/13158787/files/figure.png" pageId="44" pageNumber="45">Maps 11</figureCitation>
, 16
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</paragraph>
</subSubSection>
</treatment>
</document>

279
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<mods:title id="ECB8146EB8BDDBD178A4F2A0628F30F0">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="DE936DBC143E9CB34D604A804C5A0A6B">Griswold, Charles</mods:namePart>
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<mods:namePart id="00B1A9E6D12EB85C245924338B1C5024">Ubick, Darrell</mods:namePart>
<mods:affiliation id="84240E5F66752DD1059FA8AAC37A7032">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="543373255D9CDCB582650400D57411F9">Ledford, Joel</mods:namePart>
<mods:affiliation id="701BD1F60A137A67A88C1AF008C3171D">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="2CA44180390F12BE2502E27E7DA1CB82">Polotow, Daniele</mods:namePart>
<mods:affiliation id="F2C3C300ED9218A2541DE128E2A4BE3A">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<emphasis id="41F73E371870FFEE473431D3AA3AE9BF" bold="true" box="[123,269,220,243]" italics="true" pageId="52" pageNumber="53">Uduba jayjay</emphasis>
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,
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<paragraph id="733CE2251870FFEE473431F3AAE0E85D" blockId="52.[123,471,220,273]" box="[123,471,252,273]" pageId="52" pageNumber="53">
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,
<figureCitation id="EBB8FEA01870FFEE465F31F3AA6DE85D" box="[272,346,252,273]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">70 C, F</figureCitation>
,
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, 20.
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<emphasis id="41F73E371870FFEE47E4302EAA61E874" bold="true" box="[171,342,289,312]" pageId="52" pageNumber="53">Type material.</emphasis>
<materialsCitation id="C3EBE8781870FFEE462B302EA9D7E8D4" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" country="Madagascar" elevation="1300" latitude="-18.473333" location="Fisher-Griswold Arthropod Team" longLatPrecision="21" longitude="47.96" pageId="52" pageNumber="53" specimenCode="CASENT9004079" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
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female (
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) collected in montane rainforest at
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elevation at
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<quantity id="B47B4FC01870FFEE4628304EAA92E814" box="[359,421,321,344]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="53" unit="km" value="3.0">3 km</quantity>
41°NE Andranomay
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,
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<quantity id="B47B4FC01870FFEE45E8304EA832E814" box="[679,773,321,344]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="52" pageNumber="53" unit="km" value="11.5">11.5 km</quantity>
147°SSE Anjozorobe
</locationDeviation>
,
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,
<collectingRegion id="B1472CC71870FFEE464C306EA937E834" box="[259,512,353,376]" country="Madagascar" name="Antananarivo" pageId="52" pageNumber="53">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51870FFEE4542306EA9BBE834" box="[525,652,353,376]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
, collected
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<collectingDate id="17793D0D1870FFEE4449306EA8C6E834" box="[774,1009,353,376]" pageId="52" pageNumber="53" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5–13 December 2000</collectingDate>
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by the
<location id="765CB4FE1870FFEE4305306EAAA5E8D4" LSID="urn:lsid:plazi:treatment:FB2A53331870FFEF473431D3A82BE8B4:765CB4FE1870FFEE4305306EAAA5E8D4" country="Madagascar" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Fisher-Griswold Arthropod Team" pageId="52" pageNumber="53" stateProvince="Antananarivo">Fisher-Griswold Arthropod Team</location>
[BLF2543] deposited in
<collectionCode id="15927AE01870FFEE45EA308EA9EBE8D4" box="[677,732,385,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1870FFEE47E430AEAACAEB15" pageId="52" pageNumber="53" type="etymology">
<paragraph id="733CE2251870FFEE47E430AEAACAEB15" blockId="52.[123,1175,289,1725]" pageId="52" pageNumber="53">
<emphasis id="41F73E371870FFEE47E430AEAA1DE8F4" bold="true" box="[171,298,417,440]" pageId="52" pageNumber="53">Etymology.</emphasis>
The species epithet is an honor to Mr. Jean-Jacques Rafanomezantsoa, sometimes called “JJ”, a Malagasy entomologist, Lakeside International Scholar at the California Academy of Sciences, and principal arachnid collector with the Fisher-Griswold Arthropod Survey of
<collectingCountry id="0B94A2B51870FFEE431F30EDAB88EB55" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
(2000–2007). As a rugged and resourceful field biologist and essential member of the Fisher-Griswold Arthropod Team, Jean-Jacques Rafanomezantsoa is collector of many new and interesting arthropods from
<collectingCountry id="0B94A2B51870FFEE4637334DAAC1EB15" box="[376,502,578,601]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1870FFEE47E4336DABD1EA36" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="733CE2251870FFEE47E4336DABD1EA36" blockId="52.[123,1175,289,1725]" pageId="52" pageNumber="53">
<emphasis id="41F73E371870FFEE47E4336DAA88EB35" bold="true" box="[171,447,610,633]" pageId="52" pageNumber="53">
Diagnosis.
<taxonomicName id="B48399A61870FFEE4665336DAA88EB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[298,447,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E371870FFEE4665336DAA88EB35" bold="true" box="[298,447,610,633]" italics="true" pageId="52" pageNumber="53">Uduba jayjay</emphasis>
</taxonomicName>
</emphasis>
are members of Group VI, the
<taxonomicName id="B48399A61870FFEE4466336DA8E3EB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[809,980,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="danielae">Uduba danielae</taxonomicName>
group (Map 20). Females can be distinguished from those of other
<taxonomicName id="B48399A61870FFEE45D8338DA9D6EBD5" authorityName="Simon" authorityYear="1880" box="[663,737,642,665]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371870FFEE45D8338DA9D6EBD5" bold="true" box="[663,737,642,665]" italics="true" pageId="52" pageNumber="53">Uduba</emphasis>
</taxonomicName>
with a divided cribellum by the form of the female genitalia (
<figureCitation id="EBB8FEA01870FFEE462E33ADAAC5EBF5" box="[353,498,674,697]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Figs. 70 C, F</figureCitation>
<emphasis id="41F73E371870FFEE46BD33ADAACCEBF5" bold="true" box="[498,507,674,697]" pageId="52" pageNumber="53">)</emphasis>
.
<taxonomicName id="B48399A61870FFEE454533ADA9AAEBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[522,669,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E371870FFEE454533ADA9AAEBF5" bold="true" box="[522,669,674,697]" italics="true" pageId="52" pageNumber="53">Uduba jayjay</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="B48399A61870FFEE44EC33ADAF14EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[931,1059,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E371870FFEE44EC33ADAF14EBF5" bold="true" box="[931,1059,674,697]" italics="true" pageId="52" pageNumber="53">U. danielae</emphasis>
</taxonomicName>
by having the median lobe trapezoidal, broader posteriorly (
<figureCitation id="EBB8FEA01870FFEE45DB33CCA9C8EB96" box="[660,767,707,730]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 C</figureCitation>
), lateral margins strongly convex (
<taxonomicName id="B48399A61870FFEE433433CCABEFEBB6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E371870FFEE433433CCABEFEBB6" bold="true" italics="true" pageId="52" pageNumber="53">U. danielae</emphasis>
</taxonomicName>
with median lobe bell-shaped, broader posteriorly, sides gently curved,
<figureCitation id="EBB8FEA01870FFEE449933ECAF0DEBB6" box="[982,1082,739,762]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 A</figureCitation>
) and the vulva (
<figureCitation id="EBB8FEA01870FFEE478A320CAA1DEA56" box="[197,298,771,794]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 F</figureCitation>
) with spermathecal ducts beginning anteromedially, forming gentle outside curve to vulva base, then entering at apex of BS bulb (
<taxonomicName id="B48399A61870FFEE45DC322CA824EA76" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[659,787,803,826]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="danielae">
<emphasis id="41F73E371870FFEE45DC322CA824EA76" bold="true" box="[659,787,803,826]" italics="true" pageId="52" pageNumber="53">U. danielae</emphasis>
</taxonomicName>
with spermathecal ducts beginning anteromedially and forming Z-shaped path, lateral, medial, lateral, to posterior,
<figureCitation id="EBB8FEA01870FFEE4490324CAF7EEA16" box="[991,1097,835,858]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 B</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1870FFEF47E4328CABDEE8FB" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="733CE2251870FFEE47E4328CA915EFF1" blockId="52.[123,1175,289,1725]" pageId="52" pageNumber="53">
<emphasis id="41F73E371870FFEE47E4328CAA40EAD6" bold="true" box="[171,375,899,922]" pageId="52" pageNumber="53">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E371870FFEE46B0328CA965EAD6" bold="true" box="[511,594,899,922]" pageId="52" pageNumber="53">Female</emphasis>
(
<typeStatus id="AC385C871870FFEE452E328CA9FCEAD6" box="[609,715,899,922]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
): Total length 6.30. Markings as in
<figureCitation id="EBB8FEA01870FFEE4304328CAB92EAF6" captionStart="FIGURE" captionStartId="109.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-16@109.[120,1176,216,1254]" captionTargetPageId="109" captionText="FIGURE 5. Habitus photos of Uduba spp., females. A, B. U. danielae, Holotype (CASENT9007875) from montane rainforest at 1620m elevation in Jardin Botanique at Réserve Spéciale d’Ambohitantely. C, F. U. jayjay, Holotype (CASENT9004079) from montane rainforest at 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe. D, E. U. heliani, Holotype (CASENT9006106) from Forêt Classée Andriantantely. A, C, D. Dorsal. B, E, F. Ventral. Scale bars: A, B, D, E = 2 mm, C, F = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158568" httpUri="https://zenodo.org/record/13158568/files/figure.png" pageId="52" pageNumber="53">Figs. 5 C, F</figureCitation>
. Carapace 3.30 long, 2.20 wide, 1.00 high; clypeus 0.15 high. Eye diameters: AME 0.08, ALE 0.14, PME 0.13, PLE 0.16. Chelicerae 1.50 long; sternum 1.55 long, 1.20 wide; labium 0.70 long; palpal coxae 1.00 long.
<emphasis id="41F73E371870FFEE46CA32ECAAECEAB6" bold="true" box="[389,475,995,1018]" pageId="52" pageNumber="53">Ratios–</emphasis>
carapace length / width = 1.50, carapace height/ width = 0.45, PER/ carapace width = 0.41, PER/OAL = 3.00, PER/AER = 1.29, OAL/OQL = 1.15, OQP/ OQA = 1.23, clypeus height/ AME diameter width = 1.88, cheliceral length / clypeus height = 10.00, sternum length / width = 1.29, palpal coxa length / width = 2.50, femur I length / carapace width = 0.95, metatarsus I length / carapace width = 1.18, femur IV length / carapace width = 1.09, palpal tarsus length / carapace width = 0.41, palpal tibia length / palpal patella length = 1.50, palpal tarsus length / palpal tibia length = 1.80, palpal tarsus length / palpal femur length = 0.75, palpal tibia length / palpal patella length = 0.83. Cribellum divided. Spination: palpus–femur d0-1-2, patella p1-0, tibia p2-1, tarsus p2-1; leg I–femur d0-1-0, p0-0-1, tibia v1-1-2-0, metatarsus v2-2-2; leg II– femur p0-0-1, tibia v1-1-1 (retro), metatarsus v2-2-3; leg III–femur d1-0-0, p0-0-1, r0-0-1, tibia d0- 1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1-0-0, tibia v2- 2-2, metatarsus p1-2-2, v2-2-2, r0-2-2. Scopulae: ventral on tarsi I and II. Leg measurements: I: 2.10 + 1.20 + 1.80 + 2.60 + 1.30 = 9.00; II: 2.00 + 1.10 + 1.40 + 1.35 + 1.10 = 6.95; III: 1.60 + 0.80 + 0.90 + 1.30 + 0.90 = 5.50; IV: 2.40 + 1.00 + 1.90 + 2.20 + 1.30 = 8.80; palpus: 1.20 + 0.60 + 0.50 + NA + 0.90 = 3.20. Leg formula 1423.
<emphasis id="41F73E371870FFEE459F34AAA8B8ECF0" bold="true" box="[720,911,1445,1468]" pageId="52" pageNumber="53">Female genitalia</emphasis>
: epigynal plate without depressed atrium, median and lateral lobes indistinct (
<figureCitation id="EBB8FEA01870FFEE458434CAA800EC90" box="[715,823,1477,1500]" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 C</figureCitation>
), plate width 1.48 times length; width between bases of LL 0.52 times epigynum width; LL side width 0.19 epigynal plate width; median lobe trapezoidal, broader posteriorly, lateral margins strongly convex, length 0.62 times epigynum length; copulatory openings beneath anterior margin of obliquely transverse grooves, origin at 0.48 epigynum length, distance between CO 0.26 times epigynum width. Vulva (
<figureCitation id="EBB8FEA01870FFEE43073749ABBCEF31" captionStart="FIGURE" captionStartId="159.[155,167,1587,1606]" captionTargetBox="[144,1163,964,1556]" captionTargetId="figure-81@159.[120,1176,942,1573]" captionTargetPageId="159" captionText="FIGURE 70. Uduba spp., female genitalia drawings. A, B. U. danielae, new species, paratype (CASENT9014779) from Ambohitantely. D, E. Uduba sarotra, new species, holotype (CASENT9030916) from Parc National de Marojejy. C, F. U. jayjay, new species, holotype (CASENT9004079) from Anjozorobe. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158731" httpUri="https://zenodo.org/record/13158731/files/figure.png" pageId="52" pageNumber="53">Fig. 70 F</figureCitation>
) with spermathecal ducts beginning anteromedially, forming gentle outside curve to vulva base, then entering at apex of BS bulb, vulva length 0.60 times width, fertilization ducts separated, vulva width 4.17 times distance between FD.
</paragraph>
<paragraph id="733CE2251871FFEF47E431D3A82DE9BF" blockId="53.[123,1174,220,504]" box="[171,794,220,243]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E431D3AA2DE9BF" bold="true" box="[171,282,220,243]" pageId="53" pageNumber="54">Variation.</emphasis>
Unknown: only the
<typeStatus id="AC385C871871FFEF46B431D3A96FE9BF" box="[507,600,220,243]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
female is known.
</paragraph>
<paragraph id="733CE2251871FFEF47E431F3AAF4E839" blockId="53.[123,1174,220,504]" pageId="53" pageNumber="54">
<materialsCitation id="C3EBE8781871FFEF47E431F3AA88E839" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectionCode="CAS" country="Madagascar" elevation="1300" latitude="-18.473333" location="Fisher-Griswold Arthropod Team" longLatPrecision="21" longitude="47.96" pageId="53" pageNumber="54" specimenCode="CASENT9004079" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo" typeStatus="holotype">
M
<emphasis id="41F73E371871FFEF478D31F3AABBE85F" bold="true" box="[194,396,252,275]" pageId="53" pageNumber="54">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51871FFEF46DA31F3A960E85F" box="[405,599,252,275]" name="Madagascar" pageId="53" pageNumber="54">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371871FFEF452B31F3A868E85F" box="[612,863,252,275]" italics="true" pageId="53" pageNumber="54">
<collectingRegion id="B1472CC71871FFEF452B31F3A868E85F" box="[612,863,252,275]" country="Madagascar" name="Antananarivo" pageId="53" pageNumber="54">Antananarivo Province</collectingRegion>
</emphasis>
:
<locationDeviation id="628EFB071871FFEF443D31F3AFB8E85F" box="[882,1167,252,275]" location="Andranomay" pageId="53" pageNumber="54">
<quantity id="B47B4FC01871FFEF443D31F3A89CE85F" box="[882,939,252,275]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="54" unit="km" value="3.0">3 km</quantity>
41°NE Andranomay
</locationDeviation>
,
<locationDeviation id="628EFB071871FFEF47343012AA8AE878" box="[123,445,285,308]" location="Anjozorobe" pageId="53" pageNumber="54">
<quantity id="B47B4FC01871FFEF47343012ABE7E878" box="[123,208,285,308]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="53" pageNumber="54" unit="km" value="11.5">11.5 km</quantity>
147°SSE Anjozorobe
</locationDeviation>
,
<geoCoordinate id="16B784E21871FFEF46873012A975E878" box="[456,578,285,308]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="53" pageNumber="54" precision="15" seconds="24" value="-18.473333">18°28ʹ24ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21871FFEF45003012A9FBE878" box="[591,716,285,308]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="53" pageNumber="54" precision="15" seconds="36" value="47.96">47°57ʹ36ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01871FFEF44413012A86CE878" box="[782,859,285,308]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="54" unit="m" value="1300.0">
<elevation id="F8AE05161871FFEF44413012A86CE878" box="[782,859,285,308]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="54" unit="m" value="1300.0">1300m</elevation>
</quantity>
, general collecting –montane rainforest,
<date id="073DC4E51871FFEF47B53031AAC0E819" box="[250,503,318,341]" pageId="53" pageNumber="54" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="17793D0D1871FFEF47B53031AAC0E819" box="[250,503,318,341]" pageId="53" pageNumber="54" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5–13 December 2000</collectingDate>
</date>
,
<location id="765CB4FE1871FFEF45413031A8B1E819" LSID="urn:lsid:plazi:treatment:FB2A53331870FFEF473431D3A82BE8B4:765CB4FE1871FFEF45413031A8B1E819" box="[526,902,318,341]" country="Madagascar" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Fisher-Griswold Arthropod Team" pageId="53" pageNumber="54" stateProvince="Antananarivo">Fisher-Griswold Arthropod Team</location>
[BLF2543] (
<typeStatus id="AC385C871871FFEF43663031AFA5E819" box="[1065,1170,318,341]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1871FFEF47343051AA74E839" box="[123,323,350,373]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT9004079</specimenCode>
,
<specimenCount id="658529AC1871FFEF46013051AA47E839" box="[334,368,350,373]" count="1" pageId="53" pageNumber="54" type="female">1♀</specimenCount>
,
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)
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.
</paragraph>
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<emphasis id="41F73E371871FFEF47E43070AA69E8DA" bold="true" box="[171,350,383,406]" pageId="53" pageNumber="54">Natural history.</emphasis>
The
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female of the cribellate
<taxonomicName id="B48399A61871FFEF45BE3070A8B4E8DA" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[753,899,383,406]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E371871FFEF45BE3070A8B4E8DA" bold="true" box="[753,899,383,406]" italics="true" pageId="53" pageNumber="54">Uduba jayjay</emphasis>
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was collected in montane rainforest.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1871FFEF47E430CFA82BE8B4" pageId="53" pageNumber="54" type="distribution">
<paragraph id="733CE2251871FFEF47E430CFA82BE8B4" blockId="53.[123,1174,220,504]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E430CFAAEDE89B" bold="true" box="[171,474,448,471]" pageId="53" pageNumber="54">
Distribution.
<taxonomicName id="B48399A61871FFEF460930CFAAEDE89B" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[326,474,448,471]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jayjay">
<emphasis id="41F73E371871FFEF460930CFAAEDE89B" bold="true" box="[326,474,448,471]" italics="true" pageId="53" pageNumber="54">Uduba jayjay</emphasis>
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</emphasis>
are known only from the
<typeStatus id="AC385C871871FFEF45B130CFA81BE89B" box="[766,812,448,471]" pageId="53" pageNumber="54">type</typeStatus>
locality in montane rainforest in central east
<collectingRegion id="B1472CC71871FFEF47B530EEAAC2E8B4" box="[250,501,481,504]" country="Madagascar" name="Antananarivo" pageId="53" pageNumber="54">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51871FFEF454F30EEA9B6E8B4" box="[512,641,481,504]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
(
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, 20).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="12B48057633190D6B27665E8227D34FA">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="334EB368F6FE7AC2CBB56B9E2C84D06C">Ubick, Darrell</mods:namePart>
<mods:affiliation id="16A3043EA6C06F57B5A994EE4E62B49F">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<paragraph id="733CE2251871FFEF4734332EAAD6EB74" blockId="53.[123,707,545,599]" box="[123,481,545,568]" pageId="53" pageNumber="54">
<heading id="287455491871FFEF4734332EAAD6EB74" box="[123,481,545,568]" fontSize="10" level="2" pageId="53" pageNumber="54" reason="4">
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<taxonomicName id="B48399A61871FFEF4734332EAA79EB74" ID-CoL="C8KJL" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,334,545,568]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kavanaughi" status="sp. nov.">
<emphasis id="41F73E371871FFEF4734332EAA79EB74" bold="true" box="[123,334,545,568]" italics="true" pageId="53" pageNumber="54">Uduba kavanaughi</emphasis>
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,
<taxonomicNameLabel id="5AC4834C1871FFEF4613332EAAD6EB74" box="[348,481,545,568]" pageId="53" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="3B99B1AE1871FFEF4734334DA9EDEB33" pageId="53" pageNumber="54" type="description">
<paragraph id="733CE2251871FFEF4734334DA9F4EB1B" blockId="53.[123,707,545,599]" box="[123,707,578,599]" pageId="53" pageNumber="54">
<figureCitation id="EBB8FEA01871FFEF4734334DAA3EEB1B" box="[123,265,578,599]" captionStart="FIGURE" captionStartId="105.[155,167,1274,1293]" captionTargetBox="[120,1176,216,1257]" captionTargetId="figure-16@105.[120,1176,216,1257]" captionTargetPageId="105" captionText="FIGURE 1. Uduba spp., live individuals. A, B. U. kavanaughi, Holotype ♂ (CASENT9029870) from evergreen forest at 970m elevation at Talatakely, Parc National Ranomafana, January 2009. C, D. U. kavanaughi, Paratype ♀, CASENT9003517, Talatakely, 1.68 km SW Ranomafana, Parc National Ranomafana, January 2009. E. U. kavanaughi, Juvenile ♀, Parc National Ranomafana, April 1998. F. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. Photos A, B, F by Nikolaj Scharff, C, D by Charles Griswold, E by Dong Lin. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158556" httpUri="https://zenodo.org/record/13158556/files/figure.png" pageId="53" pageNumber="54">Figures 1 A–E</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF465C334DAA07EB1B" box="[275,304,578,599]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="53" pageNumber="54">7 I</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF4675334DAA57EB1B" box="[314,352,578,599]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="53" pageNumber="54">9 G</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF4624334DAAA0EB1B" box="[363,407,578,599]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="53" pageNumber="54">28 F</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF46EE334DAAD9EB1B" box="[417,494,578,599]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="53" pageNumber="54">47 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF46B6334DA971EB1B" box="[505,582,578,599]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="53" pageNumber="54">76 D, E</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF451F334DA9ADEB1B" box="[592,666,578,599]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="53" pageNumber="54">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF45EA334DA988EB1B" box="[677,703,578,599]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="53" pageNumber="54">13</figureCitation>
.
</paragraph>
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<taxonomicName id="B48399A61871FFEF47343365ABD6EB33" box="[123,225,618,639]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="41F73E371871FFEF47343365ABD6EB33" bold="true" box="[123,225,618,639]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47343365AB89EB33" bold="true" box="[123,190,618,639]" italics="true" pageId="53" pageNumber="54">Uduba</emphasis>
sp.
</emphasis>
</taxonomicName>
CG301
<bibRefCitation id="17129FD41871FFEF467C3365AACBEB33" author="WHEELER, W. C. &amp; J. A. CODDINGTON &amp; L. M. CROWLEY &amp; D. DIMITROV &amp; GOLOBO &amp; P. A &amp; C. E. GRISWOLD &amp; G. HORMIGA &amp; L. PRENDINI &amp; M. J. RAMIREZ &amp; P. SIERWALD &amp; L. M. ALMEIDA-SILVA &amp; F. ALVAREZ-PADILLA &amp; M. A. ARNEDO &amp; L. R. BENAVIDES &amp; S. P. BENJAMIN &amp; J. E. BOND &amp; C. J. GRISMADO &amp; E. HASAN &amp; M. HEDIN &amp; M. A. IZQUIERDO &amp; F. M. LABARQUE &amp; J. LEDFORD &amp; L. LOPARDO &amp; W. P. MADDISON &amp; J. A. MILLER &amp; L. N. PIACENTINI &amp; N. I. PLATNICK &amp; D. POLOTOW &amp; D. SILVA-DAVILA &amp; N. SCHARFF &amp; T. SZUTS &amp; D. UBICK &amp; C. VINK &amp; H. M. WOOD &amp; J. X. ZHANG" box="[307,508,618,639]" pageId="53" pageNumber="54" pagination="576 - 616" refId="ref110043" refString="WHEELER, W. C., J. A. CODDINGTON, L. M. CROWLEY, D. DIMITROV, GOLOBO ♀, P. A., C. E. GRISWOLD, G. HORMIGA, L. PRENDINI, M. J. RAMIREZ, P. SIERWALD, L. M. ALMEIDA-SILVA, F. ALVAREZ-PADILLA, M. A. ARNEDO, L. R. BENAVIDES, S. P. BENJAMIN, J. E. BOND, C. J. GRISMADO, E. HASAN, M. HEDIN, M. A. IZQUIERDO, F. M. LABARQUE, J. LEDFORD, L. LOPARDO, W. P. MADDISON, J. A. MILLER, L. N. PIACENTINI, N. I. PLATNICK, D. POLOTOW, D. SILVA-DAVILA, N. SCHARFF, T. SZUTS, D. UBICK, C. VINK, H. M. WOOD, AND J. X. ZHANG. (2017). The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. &lt;doi: 10.1111 / cla. 12182&gt;" type="journal article" year="2017">Wheeler et al. (2017</bibRefCitation>
: fig. 6), DNA voucher.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1871FFEF47E4339EA8FAEAC1" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="733CE2251871FFEF47E4339EA8FAEAC1" blockId="53.[123,1174,657,1725]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E4339EAA67EBE4" bold="true" box="[171,336,657,680]" pageId="53" pageNumber="54">Type material.</emphasis>
<materialsCitation id="C3EBE8781871FFEF4616339EA8FAEAC1" collectingDate="2001-01-02" collectingDateMax="2009-01-12" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="C. Griswold &amp; D. H. and K. M. Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; Talatakely &amp; A. Saucedo &amp; H. Wood" country="Madagascar" county="Ranomafana National Park" elevation="970" latitude="-21.264055" location="Parc National Ranomafana" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana" pageId="53" pageNumber="54" specimenCode="CASENT9029870, CGM01, CASENT9003516, CASENT9003517, CASENT9031127" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871871FFEF4616339EAA8BEBE4" box="[345,444,657,680]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1871FFEF4547339EA9E7EBE4" box="[520,720,657,680]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT9029870</specimenCode>
) from evergreen forest at
<quantity id="B47B4FC01871FFEF44A1339EAF1CEBE4" box="[1006,1067,657,680]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="53" pageNumber="54" unit="m" value="970.0">
<elevation id="F8AE05161871FFEF44A1339EAF1CEBE4" box="[1006,1067,657,680]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="53" pageNumber="54" unit="m" value="970.0">970m</elevation>
</quantity>
elevation in
<location id="765CB4FE1871FFEF47D933BDAA8FEB85" LSID="urn:lsid:plazi:treatment:FB2A53331871FFED4734332EA817EC05:765CB4FE1871FFEF47D933BDAA8FEB85" box="[150,440,690,713]" country="Madagascar" county="Ranomafana National Park" latitude="-21.264055" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana" name="Parc National Ranomafana" pageId="53" pageNumber="54" stateProvince="Fianarantsoa">
Parc National
<collectingMunicipality id="9358785F1871FFEF466133BDAA8FEB85" box="[302,440,690,713]" pageId="53" pageNumber="54">Ranomafana</collectingMunicipality>
</location>
,
<geoCoordinate id="16B784E21871FFEF468D33BDA979EB85" box="[450,590,690,713]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="53" pageNumber="54" precision="1" seconds="50.6" value="-21.264055">21º15ʹ50.6ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21871FFEF451533BDA9ECEB85" box="[602,731,690,713]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="53" pageNumber="54" precision="1" seconds="9.6" value="47.419334">47º25ʹ9.6ʺE</geoCoordinate>
,
<collectingRegion id="B1472CC71871FFEF45AA33BDA8E1EB85" box="[741,982,690,713]" country="Madagascar" name="Fianarantsoa" pageId="53" pageNumber="54">Fianarantsoa Province</collectingRegion>
,
<collectingCountry id="0B94A2B51871FFEF44AF33BDAF68EB85" box="[992,1119,690,713]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
, collected
<date id="073DC4E51871FFEF478933DDAA9DEBA5" box="[198,426,722,745]" pageId="53" pageNumber="54" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">
<collectingDate id="17793D0D1871FFEF478933DDAA9DEBA5" box="[198,426,722,745]" pageId="53" pageNumber="54" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">11–12 January 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31871FFEF46F533DDA976EBA5" box="[442,577,722,745]" pageId="53" pageNumber="54">C. Griswold</collectorName>
(
<specimenCode id="23254A5E1871FFEF451A33DDA986EBA5" box="[597,689,722,745]" collectionCode="CGM" pageId="53" pageNumber="54">CGM01</specimenCode>
), deposited in
<collectionCode id="15927AE01871FFEF441333DDA8A4EBA5" box="[860,915,722,745]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="53" pageNumber="54" type="Museum">CAS</collectionCode>
.
<specimenCount id="658529AC1871FFEF44EE33DDAFA2EBA5" box="[929,1173,722,745]" count="2" pageId="53" pageNumber="54" type="female" typeStatus="paratype">Two paratype females</specimenCount>
(
<specimenCode id="23254A5E1871FFEF47CD33FCAA7DEA46" box="[130,330,755,778]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT9003516</specimenCode>
) and (
<specimenCode id="23254A5E1871FFEF46C333FCA960EA46" box="[396,599,755,778]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT9003517</specimenCode>
), both from mixed tropical forest at
<quantity id="B47B4FC01871FFEF449A33FCAF25EA46" box="[981,1042,755,778]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="m" value="900.0">
<elevation id="F8AE05161871FFEF449A33FCAF25EA46" box="[981,1042,755,778]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="m" value="900.0">900m</elevation>
</quantity>
elevation in
<collectingCounty id="9A5D9AA91871FFEF4734321BAA95EA67" box="[123,418,788,811]" pageId="53" pageNumber="54">Ranomafana National Park</collectingCounty>
,
<geoCoordinate id="16B784E21871FFEF46E1321BA91AEA67" box="[430,557,788,811]" degrees="21.25041" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
,
<geoCoordinate id="16B784E21871FFEF4575321BA98BEA67" box="[570,700,788,811]" degrees="47.41945" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, collected
<date id="073DC4E51871FFEF447E321BA8C8EA67" box="[817,1023,788,811]" pageId="53" pageNumber="54" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="17793D0D1871FFEF447E321BA8C8EA67" box="[817,1023,788,811]" pageId="53" pageNumber="54" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2–22 January 2001</collectingDate>
</date>
by
<collectorName id="DE7687F31871FFEF4365321BAA0AEA07" pageId="53" pageNumber="54">D. H. and K. M. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31871FFEF4607323BAA8AEA07" box="[328,445,820,843]" pageId="53" pageNumber="54">R. L. Brett</collectorName>
,
<collectorName id="DE7687F31871FFEF4687323BA91DEA07" box="[456,554,820,843]" pageId="53" pageNumber="54">E. Elsom</collectorName>
and
<collectorName id="DE7687F31871FFEF4510323BA989EA07" box="[607,702,820,843]" pageId="53" pageNumber="54">F. Vargas</collectorName>
, and
<specimenCount id="658529AC1871FFEF45BA323BA88DEA07" box="[757,954,820,843]" count="1" pageId="53" pageNumber="54" type="male" typeStatus="paratype">one paratype male</specimenCount>
(
<specimenCode id="23254A5E1871FFEF4488323BAFB9EA07" box="[967,1166,820,843]" collectionCode="CASENT" pageId="53" pageNumber="54">CASENT9031127</specimenCode>
) from
<collectorName id="DE7687F31871FFEF47F8325AAA17EA20" box="[183,288,853,876]" pageId="53" pageNumber="54">Talatakely</collectorName>
,
<locationDeviation id="628EFB071871FFEF467E325AA97AEA20" box="[305,589,853,876]" location="Ranomafana" pageId="53" pageNumber="54">
<quantity id="B47B4FC01871FFEF467E325AAABEEA20" box="[305,393,853,876]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="53" pageNumber="54" unit="km" value="1.68">1.68 km</quantity>
SW Ranomafana
</locationDeviation>
, elev.
<quantity id="B47B4FC01871FFEF45DB325AA9E3EA20" box="[660,724,853,876]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="53" pageNumber="54" unit="m" value="970.0">
<elevation id="F8AE05161871FFEF45DB325AA9E3EA20" box="[660,724,853,876]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="53" pageNumber="54" unit="m" value="970.0">970m</elevation>
</quantity>
,
<geoCoordinate id="16B784E21871FFEF45AE325AA85AEA20" box="[737,877,853,876]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="53" pageNumber="54" precision="1" seconds="50.6" value="-21.264055">21º15ʹ50.6ʺS</geoCoordinate>
, 47º9.6ʺE, collected
<date id="073DC4E51871FFEF431A325AAA3CEAC1" pageId="53" pageNumber="54" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">
<collectingDate id="17793D0D1871FFEF431A325AAA3CEAC1" pageId="53" pageNumber="54" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">11–12 January 2009</collectingDate>
</date>
by
<collectorName id="DE7687F31871FFEF46793279AA8BEAC1" box="[310,444,886,909]" pageId="53" pageNumber="54">C. Griswold</collectorName>
,
<collectorName id="DE7687F31871FFEF468A3279A976EAC1" box="[453,577,886,909]" pageId="53" pageNumber="54">A. Saucedo</collectorName>
and
<collectorName id="DE7687F31871FFEF45373279A9EEEAC1" box="[632,729,886,909]" pageId="53" pageNumber="54">H. Wood</collectorName>
, all deposited in
<collectionCode id="15927AE01871FFEF44DD3279A8FEEAC1" box="[914,969,886,909]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="53" pageNumber="54" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1871FFEF47E43299AF76EDFE" pageId="53" pageNumber="54" type="etymology">
<paragraph id="733CE2251871FFEF47E43299A827EAA2" blockId="53.[123,1174,657,1725]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E43299AA1DEAE1" bold="true" box="[171,298,918,941]" pageId="53" pageNumber="54">Etymology.</emphasis>
The species epithet is a patronym in honor of Dr. David H. Kavanaugh,
<taxonomicName id="B48399A61871FFEF436F3299AFA2EAE1" authorityName="Linnaeus" authorityYear="1758" box="[1056,1173,918,941]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
curator and natural history explorer from the
<collectingRegion id="B1472CC71871FFEF453232B8A9DEEA82" box="[637,745,951,974]" country="United States of America" name="California" pageId="53" pageNumber="54">California</collectingRegion>
Academy of Sciences, who dauntlessly pursued and collected this species as well as other
<taxonomicName id="B48399A61871FFEF45F032D8A83EEAA2" authorityName="Simon" authorityYear="1880" box="[703,777,983,1006]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371871FFEF45F032D8A83EEAA2" bold="true" box="[703,777,983,1006]" italics="true" pageId="53" pageNumber="54">Uduba</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="733CE2251871FFEF47E432F7AF76EDFE" blockId="53.[123,1174,657,1725]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E432F7ABD7ED43" bold="true" box="[171,224,1016,1039]" pageId="53" pageNumber="54">Note</emphasis>
. In the phylogeny publication by
<bibRefCitation id="17129FD41871FFEF452A32F7A86FED43" author="WHEELER, W. C. &amp; J. A. CODDINGTON &amp; L. M. CROWLEY &amp; D. DIMITROV &amp; GOLOBO &amp; P. A &amp; C. E. GRISWOLD &amp; G. HORMIGA &amp; L. PRENDINI &amp; M. J. RAMIREZ &amp; P. SIERWALD &amp; L. M. ALMEIDA-SILVA &amp; F. ALVAREZ-PADILLA &amp; M. A. ARNEDO &amp; L. R. BENAVIDES &amp; S. P. BENJAMIN &amp; J. E. BOND &amp; C. J. GRISMADO &amp; E. HASAN &amp; M. HEDIN &amp; M. A. IZQUIERDO &amp; F. M. LABARQUE &amp; J. LEDFORD &amp; L. LOPARDO &amp; W. P. MADDISON &amp; J. A. MILLER &amp; L. N. PIACENTINI &amp; N. I. PLATNICK &amp; D. POLOTOW &amp; D. SILVA-DAVILA &amp; N. SCHARFF &amp; T. SZUTS &amp; D. UBICK &amp; C. VINK &amp; H. M. WOOD &amp; J. X. ZHANG" box="[613,856,1016,1039]" pageId="53" pageNumber="54" pagination="576 - 616" refId="ref110043" refString="WHEELER, W. C., J. A. CODDINGTON, L. M. CROWLEY, D. DIMITROV, GOLOBO ♀, P. A., C. E. GRISWOLD, G. HORMIGA, L. PRENDINI, M. J. RAMIREZ, P. SIERWALD, L. M. ALMEIDA-SILVA, F. ALVAREZ-PADILLA, M. A. ARNEDO, L. R. BENAVIDES, S. P. BENJAMIN, J. E. BOND, C. J. GRISMADO, E. HASAN, M. HEDIN, M. A. IZQUIERDO, F. M. LABARQUE, J. LEDFORD, L. LOPARDO, W. P. MADDISON, J. A. MILLER, L. N. PIACENTINI, N. I. PLATNICK, D. POLOTOW, D. SILVA-DAVILA, N. SCHARFF, T. SZUTS, D. UBICK, C. VINK, H. M. WOOD, AND J. X. ZHANG. (2017). The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. &lt;doi: 10.1111 / cla. 12182&gt;" type="journal article" year="2017">
Wheeler
<emphasis id="41F73E371871FFEF458432F7A831ED43" box="[715,774,1016,1039]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</bibRefCitation>
a voucher specimen of this species was included and listed as
<emphasis id="41F73E371871FFEF46A13516A98DED7C" bold="true" box="[494,698,1049,1072]" pageId="53" pageNumber="54">
<taxonomicName id="B48399A61871FFEF46A13516A955ED7C" box="[494,610,1049,1072]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="undetermined">Uduba sp.</taxonomicName>
CG301
</emphasis>
“big red juvenile” (
<bibRefCitation id="17129FD41871FFEF44C03516AF5AED7C" author="WHEELER, W. C. &amp; J. A. CODDINGTON &amp; L. M. CROWLEY &amp; D. DIMITROV &amp; GOLOBO &amp; P. A &amp; C. E. GRISWOLD &amp; G. HORMIGA &amp; L. PRENDINI &amp; M. J. RAMIREZ &amp; P. SIERWALD &amp; L. M. ALMEIDA-SILVA &amp; F. ALVAREZ-PADILLA &amp; M. A. ARNEDO &amp; L. R. BENAVIDES &amp; S. P. BENJAMIN &amp; J. E. BOND &amp; C. J. GRISMADO &amp; E. HASAN &amp; M. HEDIN &amp; M. A. IZQUIERDO &amp; F. M. LABARQUE &amp; J. LEDFORD &amp; L. LOPARDO &amp; W. P. MADDISON &amp; J. A. MILLER &amp; L. N. PIACENTINI &amp; N. I. PLATNICK &amp; D. POLOTOW &amp; D. SILVA-DAVILA &amp; N. SCHARFF &amp; T. SZUTS &amp; D. UBICK &amp; C. VINK &amp; H. M. WOOD &amp; J. X. ZHANG" box="[911,1133,1049,1072]" pageId="53" pageNumber="54" pagination="576 - 616" refId="ref110043" refString="WHEELER, W. C., J. A. CODDINGTON, L. M. CROWLEY, D. DIMITROV, GOLOBO ♀, P. A., C. E. GRISWOLD, G. HORMIGA, L. PRENDINI, M. J. RAMIREZ, P. SIERWALD, L. M. ALMEIDA-SILVA, F. ALVAREZ-PADILLA, M. A. ARNEDO, L. R. BENAVIDES, S. P. BENJAMIN, J. E. BOND, C. J. GRISMADO, E. HASAN, M. HEDIN, M. A. IZQUIERDO, F. M. LABARQUE, J. LEDFORD, L. LOPARDO, W. P. MADDISON, J. A. MILLER, L. N. PIACENTINI, N. I. PLATNICK, D. POLOTOW, D. SILVA-DAVILA, N. SCHARFF, T. SZUTS, D. UBICK, C. VINK, H. M. WOOD, AND J. X. ZHANG. (2017). The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. &lt;doi: 10.1111 / cla. 12182&gt;" type="journal article" year="2017">Wheeler et al., 2017</bibRefCitation>
:fig. 6), DNA voucher, with the following collection data: “Talatakely forest,
<quantity id="B47B4FC01871FFEF44E73536AF36ED1C" box="[936,1025,1081,1104]" metricMagnitude="4" metricUnit="m" metricValue="4.2299999999999995" pageId="53" pageNumber="54" unit="km" value="42.3">42.3 km</quantity>
58˚ NE
<collectingRegion id="B1472CC71871FFEF43253536ABD3ED3D" country="Madagascar" name="Fianarantsoa" pageId="53" pageNumber="54">Fianarantsoa</collectingRegion>
, S21˚15ʹ28.0ʺ, E47˚25ʹ21.8ʺ, elev.
<quantity id="B47B4FC01871FFEF45253555A980ED3D" box="[618,695,1114,1137]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="53" pageNumber="54" unit="m" value="1050.0">1050m</quantity>
, montane rainforest, general collecting,
<date id="073DC4E51871FFEF43343555AA12EDDE" pageId="53" pageNumber="54" value="2005-12-24">24 December 2005</date>
to
<date id="073DC4E51871FFEF46053574AAC8EDDE" box="[330,511,1147,1170]" pageId="53" pageNumber="54" value="2006-01-14">14 January 2006</date>
, H. Wood, J. Miller, J.J. Rafanomezantsoa, E. Rajeriarison, V. Andriamananona, [HW009] [DNA-MAD-111] 1j, (CASENT9024063,
<specimenCount id="658529AC1871FFEF44383594A8D2EDFE" box="[887,997,1179,1202]" count="1" pageId="53" pageNumber="54" type="juvenile">1 juvenile</specimenCount>
, CAS)”.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1871FFEF47E435B3A9F0EF30" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="733CE2251871FFEF47E435B3A9F0EF30" blockId="53.[123,1174,657,1725]" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E435B3AACFED9F" bold="true" box="[171,504,1212,1235]" pageId="53" pageNumber="54">
Diagnosis.
<taxonomicName id="B48399A61871FFEF466935B3AACFED9F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,504,1212,1235]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="41F73E371871FFEF466935B3AACFED9F" bold="true" box="[294,504,1212,1235]" italics="true" pageId="53" pageNumber="54">Uduba kavanaughi</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.a, the
<taxonomicName id="B48399A61871FFEF440935B3A8FEED9F" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[838,969,1212,1235]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA01871FFEF435335B3AF47ED9F" box="[1052,1136,1212,1235]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="53" pageNumber="54">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate and very large, with males greater than
<quantity id="B47B4FC01871FFEF473435F2ABF5EC58" box="[123,194,1277,1300]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="53" pageNumber="54" unit="mm" value="22.0">22mm</quantity>
, females greater than
<quantity id="B47B4FC01871FFEF46FA35F2AACEEC58" box="[437,505,1277,1300]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="54" unit="mm" value="30.0">30mm</quantity>
total length. Males can be distinguished from those of other
<taxonomicName id="B48399A61871FFEF47343411ABF2EC79" authorityName="Simon" authorityYear="1880" box="[123,197,1310,1333]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371871FFEF47343411ABF2EC79" bold="true" box="[123,197,1310,1333]" italics="true" pageId="53" pageNumber="54">Uduba</emphasis>
</taxonomicName>
with a screw shaped TA3 by having the TA2 that is hidden ventrally behind TA3 (
<figureCitation id="EBB8FEA01871FFEF43723411AB9AEC1A" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="53" pageNumber="54">Figs. 47 A–C</figureCitation>
<emphasis id="41F73E371871FFEF47E23430AB81EC1A" bold="true" box="[173,182,1343,1366]" pageId="53" pageNumber="54">)</emphasis>
and are distinguished from
<taxonomicName id="B48399A61871FFEF46BE3430A992EC1A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[497,677,1343,1366]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="41F73E371871FFEF46BE3430A992EC1A" bold="true" box="[497,677,1343,1366]" italics="true" pageId="53" pageNumber="54">Uduba platnicki</emphasis>
</taxonomicName>
by having the TA2 formed as a small, blunt blade with a median longitudinal ridge (
<figureCitation id="EBB8FEA01871FFEF457A3450A9ABEC3A" box="[565,668,1375,1398]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="53" pageNumber="54">Fig. 28 F</figureCitation>
<emphasis id="41F73E371871FFEF45D43450A993EC3A" bold="true" box="[667,676,1375,1398]" pageId="53" pageNumber="54">)</emphasis>
(in
<taxonomicName id="B48399A61871FFEF459B3450A8BFEC3A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[724,904,1375,1398]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="41F73E371871FFEF459B3450A8BFEC3A" bold="true" box="[724,904,1375,1398]" italics="true" pageId="53" pageNumber="54">Uduba platnicki</emphasis>
</taxonomicName>
the TA2 a small, simple mound, lacking such a ridge,
<figureCitation id="EBB8FEA01871FFEF46F9348FA92AECDB" box="[438,541,1408,1431]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="53" pageNumber="54">Fig. 28 E</figureCitation>
). Females can be distinguished from those of other ecribellate
<taxonomicName id="B48399A61871FFEF47E734AEABC5ECF4" authorityName="Simon" authorityYear="1880" box="[168,242,1441,1464]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371871FFEF47E734AEABC5ECF4" bold="true" box="[168,242,1441,1464]" italics="true" pageId="53" pageNumber="54">Uduba</emphasis>
</taxonomicName>
that have an epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01871FFEF437034AEAB98EC94" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="53" pageNumber="54">Figs. 76 D, E</figureCitation>
) by having the plate short or nearly absent posteriad of atrium, height less than 0.12 times epigynal length, lateral lobes meet behind atrium to form short posterior plate, equal to 0.08–0.12 times epigynum length, median lobe short and broad, width atrium less than 3 times ML width, ML length less than 1.50 times width, atrium a broad depression on each side of ML; CO (
<figureCitation id="EBB8FEA01871FFEF4352372CABBBEF17" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="53" pageNumber="54">Figs. 76 D, E</figureCitation>
) appear to be beneath the ML (as in
<taxonomicName id="B48399A61871FFEF455E374BA94CEF17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[529,635,1604,1627]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="halabe">
<emphasis id="41F73E371871FFEF455E374BA94CEF17" bold="true" box="[529,635,1604,1627]" italics="true" pageId="53" pageNumber="54">U. halabe</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01871FFEF45C6374BA822EF17" box="[649,789,1604,1627]" captionStart="FIGURE" captionStartId="162.[155,167,918,937]" captionTargetBox="[120,1176,216,898]" captionTargetId="figure-11@162.[120,1176,216,898]" captionTargetPageId="162" captionText="FIGURE 74. Uduba spp., female genitalia drawings. A, B. U. salegy, new species, paratype (CASENT9006045) Parc National Montagne d’Ambre. C, F. U. halabe, new species, paratype (CASENT9006015) Réserve Naturelle Intégrale d’Andohahela. D, E. U. balsama, new species, paratype (CASENT9013532) from Forêt de Kirindy. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations A–C, F by Jenny Speckels, D, E by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158739" httpUri="https://zenodo.org/record/13158739/files/figure.png" pageId="53" pageNumber="54">Figs. 74 C, F</figureCitation>
), but
<taxonomicName id="B48399A61871FFEF441E374BA88BEF17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[849,956,1604,1627]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="halabe">
<emphasis id="41F73E371871FFEF441E374BA88BEF17" bold="true" box="[849,956,1604,1627]" italics="true" pageId="53" pageNumber="54">U. halabe</emphasis>
</taxonomicName>
has an entire cribellum (
<figureCitation id="EBB8FEA01871FFEF47FB376AAA2BEF30" box="[180,284,1637,1660]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="53" pageNumber="54">Fig. 20 A</figureCitation>
) whereas
<taxonomicName id="B48399A61871FFEF46C5376AA919EF30" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[394,558,1637,1660]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="41F73E371871FFEF46C5376AA919EF30" bold="true" box="[394,558,1637,1660]" italics="true" pageId="53" pageNumber="54">U. kavanaughi</emphasis>
</taxonomicName>
is ecribellate.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1871FFED47E4378AA996EBF0" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
<paragraph id="733CE2251871FFEC47E4378AA9C1EFD1" blockId="53.[123,1174,657,1725]" lastBlockId="54.[123,1174,220,1725]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<emphasis id="41F73E371871FFEF47E4378AAA42EFD0" bold="true" box="[171,373,1669,1692]" pageId="53" pageNumber="54">Description. Male</emphasis>
(
<typeStatus id="AC385C871871FFEF46CC378AAADAEFD0" box="[387,493,1669,1692]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
): Total length 24.00. Markings as in
<figureCitation id="EBB8FEA01871FFEF443B378AA8C2EFD0" box="[884,1013,1669,1692]" captionStart="FIGURE" captionStartId="105.[155,167,1274,1293]" captionTargetBox="[120,1176,216,1257]" captionTargetId="figure-16@105.[120,1176,216,1257]" captionTargetPageId="105" captionText="FIGURE 1. Uduba spp., live individuals. A, B. U. kavanaughi, Holotype ♂ (CASENT9029870) from evergreen forest at 970m elevation at Talatakely, Parc National Ranomafana, January 2009. C, D. U. kavanaughi, Paratype ♀, CASENT9003517, Talatakely, 1.68 km SW Ranomafana, Parc National Ranomafana, January 2009. E. U. kavanaughi, Juvenile ♀, Parc National Ranomafana, April 1998. F. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. Photos A, B, F by Nikolaj Scharff, C, D by Charles Griswold, E by Dong Lin. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158556" httpUri="https://zenodo.org/record/13158556/files/figure.png" pageId="53" pageNumber="54">Figs. 1 A–B</figureCitation>
,
<figureCitation id="EBB8FEA01871FFEF44B0378AAF1EEFD0" box="[1023,1065,1669,1692]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="53" pageNumber="54">9 G</figureCitation>
, body and legs to patellae dark, tibiae to tarsi segments light. Carapace 13.80 long, 10.60 wide, 8.00 high; clypeus 3.60 high. Eye diameters: AME 0.50, ALE and PME 0.56, PLE 0.52. Chelicerae 18.00 long; sternum 14.20 long, 11.80 wide; labium 8.00 long; palpal coxae 11.50 long. With a colulus. Spination (
<typeStatus id="AC385C871872FFEC47BE3013AA66E87F" box="[241,337,284,307]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
, CASENT9031127): palpus–femur d0-0-1, p0-0-1, r0-0-1; leg I–femur d1-0- 1-0, p0-0-1, r0-0-1, tibia d0-0-0-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-0-0, v2-1-3, r1-0-0; leg II–femur d1-1-2, p0-1-1-0, r0-1-1-0, patella r1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-1, v1-1-2, r1-0-0; leg III–femur d1-1-0-1, p0-1-0-1-1, r0-1-0-1-1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-2-2, tarsus v0-1-0; leg IV–femur d1-1-1- 0-1, p0-1-0-1, r0-0-0-1, tibia d0-0-0-1, p0-1-1-0, v2-1-2-0, r0-1-1-0, metatarsus p0-1-2, v1-1-1-2, r0-1-1-1. Scopulae: cymbium, apicodorsal; beneath all tarsi, metatarsi I and II, and apices of metatarsi III and IV. Leg measurements [
<typeStatus id="AC385C871872FFEC456730F1A9B9EB59" box="[552,654,510,533]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
]: I: 15.00 + 6.00 + 13.00 + 15.00 + 7.00 = 56.00; II: 14.50 + 5.50 + 10.00 + 13.00 + 2.50 = 48.50; III: 10.10 + 4.50 + 7.00 + 11.00 + 4.00 = 36.50; IV: 14.00 + 4.50 + 11.00 + 17.20 + 6.50 = 53.20; palpus: 9.00 + 2.20 + 2.10 + NA + 8.30 = 21.60. Leg formula 1423.
<emphasis id="41F73E371872FFEC461F3351AAF2EB39" bold="true" box="[336,453,606,629]" pageId="54" pageNumber="55">Male palp</emphasis>
(
<typeStatus id="AC385C871872FFEC46993351A90FEB39" box="[470,568,606,629]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
,
<figureCitation id="EBB8FEA01872FFEC450A3351A9EBEB39" box="[581,732,606,629]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Figs. 47 A–C</figureCitation>
): palpal tibia 0.34 times cymbial length, RTA conical, length 0.16 tibia length, length 0.56 times width, with bluntly pointed apex (
<figureCitation id="EBB8FEA01872FFEC43083370ABB8EBFA" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Fig. 47 B</figureCitation>
); VTA conical, length 1.25 times width, length 0.31 tibia width; tibia lacking stout spines; tegulum convex, without ridge (
<figureCitation id="EBB8FEA01872FFEC46ED33B0A904EB9A" box="[418,563,703,726]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Figs. 47 B, C</figureCitation>
), tegulum length 1.02 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.07 times tegulum length; TA3 a large, sharp-point- ed screw, with proximal ridges (
<figureCitation id="EBB8FEA01872FFEC469C33F0A951EA5A" box="[467,614,767,790]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Figs. 47 B, C</figureCitation>
), TA3 extends apicad of TA2 by 1.15 times tegulum length, TA2 hidden behind apex of TA3, forming a small, blunt blade with a median longitudinal ridge (
<figureCitation id="EBB8FEA01872FFEC478D324FAA1DEA1B" box="[194,298,832,855]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="54" pageNumber="55">Fig. 28 F</figureCitation>
); MA simple, origin at 0.41 times tegulum length, tegulum apex at 0.34 tegulum length from MA apex, MA small (
<figureCitation id="EBB8FEA01872FFEC46BD326FA9BDEA3B" box="[498,650,864,887]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Figs. 47 B, C</figureCitation>
), length (including apical processes) 0.56 times width, width 0.37 tegulum width; conductor fan margin irregular (
<figureCitation id="EBB8FEA01872FFEC4402328FA88FEADB" box="[845,952,896,919]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="54" pageNumber="55">Fig. 47 C</figureCitation>
).
<emphasis id="41F73E371872FFEC4482328FAF17EADB" bold="true" box="[973,1056,896,919]" pageId="54" pageNumber="55">Female</emphasis>
(
<typeStatus id="AC385C871872FFEC437F328FAFA5EADB" box="[1072,1170,896,919]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
, CASENT9003516): Total length 31.00. Markings as in
<figureCitation id="EBB8FEA01872FFEC458032AEA867EAF4" box="[719,848,929,952]" captionStart="FIGURE" captionStartId="105.[155,167,1274,1293]" captionTargetBox="[120,1176,216,1257]" captionTargetId="figure-16@105.[120,1176,216,1257]" captionTargetPageId="105" captionText="FIGURE 1. Uduba spp., live individuals. A, B. U. kavanaughi, Holotype ♂ (CASENT9029870) from evergreen forest at 970m elevation at Talatakely, Parc National Ranomafana, January 2009. C, D. U. kavanaughi, Paratype ♀, CASENT9003517, Talatakely, 1.68 km SW Ranomafana, Parc National Ranomafana, January 2009. E. U. kavanaughi, Juvenile ♀, Parc National Ranomafana, April 1998. F. U. madagascariensis (Vinson, 1863), ♂ CASENT9006030, specimen from suburbs of Antananarivo. Photos A, B, F by Nikolaj Scharff, C, D by Charles Griswold, E by Dong Lin. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158556" httpUri="https://zenodo.org/record/13158556/files/figure.png" pageId="54" pageNumber="55">Figs. 1 C–E</figureCitation>
,
<figureCitation id="EBB8FEA01872FFEC441532AEA84EEAF4" box="[858,889,929,952]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="54" pageNumber="55">7 I</figureCitation>
, carapace and legs reddish in life. Carapace 16.00 long, 11.60 wide, 6.50 high; clypeus 1.40 high. Eye diameters: AME 0.40, ALE 0.48, PME 0.46, PLE 0.50. Chelicerae 7.80 long; sternum 6.50 long, 5.20 wide; labium 3.50 long; palpal coxae 5.40 long. With a colulus. Spination: Female (
<typeStatus id="AC385C871872FFEC441D350EA885ED54" box="[850,946,1025,1048]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
, CASENT9003517): palpus–femur d1-0-3, patella p1, tibia p2-0, tarsus p2-1, v2-0; leg I–femur d0-1-0-1, p0-0-0-2, patella p1, v1, tibia p0-1-1-0, v2-1-2-2, metatarsus v2-2-1; leg II–femur d1-0-0-1, p0-1-0-2, r0-1- 0-0, tibia p0-1-1-0, v2-2-2-2, metatarsus p0-1-1-0, v2-1-2-1; leg III–femur d1-1-0-1, p1-0-1-1, r1- 0-1-1, tibia d1-0-0-0, p0-1-1-0, v2-2-3, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-2-2, tarsus v0-1-0; leg IV–femur d0-1-0-1, p0-1-0-1, r0-0-0-1, tibia p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1, v2- 1-2-0, r2-2-1. Scopulae: strong ventral beneath all tarsi, beneath metatarsi I-III and apical half of metatarsus IV, and beneath apices of tibiae I and II. Leg measurements (
<typeStatus id="AC385C871872FFEC436835ECAFA6EDB6" box="[1063,1169,1251,1274]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
, CASENT9003516): I: 12.00 + 5.80 + 9.60 + 10.00 + 4.80 = 42.20; II: 10.30 + 5.30 + 8.00 + 8.50 + 4.50 = 36.60; III: 8.60 + 4.60 + 5.50 + 8.00 + 4.40 = 31.10; IV: 11.20 + 4.80 + 8.70 + 12.00 + 4.90 = 41.60; palpus: 6.00 + 3.10 + 3.50 + NA + 6.80 = 19.40. Leg formula 1423.
<emphasis id="41F73E371872FFEC4345344CAB82EC37" bold="true" pageId="54" pageNumber="55">Female genitalia</emphasis>
(
<typeStatus id="AC385C871872FFEC478E346BAA14EC37" box="[193,291,1380,1403]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
, CASENT9003517): epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA01872FFEC47E3348BAA22ECD7" box="[172,277,1412,1435]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="54" pageNumber="55">Fig. 76 D</figureCitation>
), plate width 1.50 times length; atrium sides evenly concave, atrium extends anteriad and posteriad of ML, width 1.425 times atrium length; epigynum length 1.75 times atrium length, epigynum width 1.84 times atrium width; atrium width at side of ML 0.46 times ML width; atrium ends just anteriad of epigastric groove, atrium origin at 0.09 of epigynal plate length, medi- an lobe broad, notched apically and widest at base, median lobe length 1.27 times width, ML ends at atrium apex, ML length 0.49 times epigynal length; lateral lobes broad, atrium width 2.85 times LL width; copulatory openings appear to open beneath ML of atrium (
<figureCitation id="EBB8FEA01872FFEC4426374AA8E5EF10" box="[873,978,1605,1628]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="54" pageNumber="55">Fig. 76 D</figureCitation>
). Vulva with spermathecal ducts making two large, transverse curves, vulva length 0.60 times width, vulva width 4.16 times distance between fertilization ducts (
<figureCitation id="EBB8FEA01872FFEC45CF3789A9DDEFD1" box="[640,746,1670,1693]" captionStart="FIGURE" captionStartId="164.[155,167,1141,1160]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-11@164.[120,1176,216,1117]" captionTargetPageId="164" captionText="FIGURE 76. Uduba spp., female genitalia drawings. A, B. U. barbarae, new species, paratype (CASENT9006064) from Lokobe Forest on the island of Nosy Be. C, F, I. U. milamina, new species. C. Holotype (CASENT9010797) from a malaise canopy trap at the Ranomafana-Radio Tower. F, I. Paratype (CASENT9006100) from Andringitra Reserve. D, E. U. kavanaughi, new species, paratype (CASENT9003517) from Talatakely, Parc National Ranomafana. G, H. U. platnicki, new species, paratype (CASENT9030253) from Mikira Forest. A, C, D. H, I. Epigynum, ventral. B, E, F, G. Vulva, dorsal. Illustrations A, B by Jenny Speckels, C–I by Charles Griswold. At, atrium of epigynum; AtSp, atrial side plate of epigynum; EF, epigastric furrow; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158743" httpUri="https://zenodo.org/record/13158743/files/figure.png" pageId="54" pageNumber="55">Fig. 76 E</figureCitation>
).
</paragraph>
<paragraph id="733CE2251872FFED47E437A9A996EBF0" blockId="54.[123,1174,220,1725]" lastBlockId="55.[122,1175,220,1353]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis id="41F73E371872FFEC47E437A9AA6BEFF1" bold="true" box="[171,348,1702,1725]" pageId="54" pageNumber="55">
Variation.
<emphasis id="41F73E371872FFEC466B37A9AA6BEFF1" bold="true" box="[292,348,1702,1725]" italics="true" pageId="54" pageNumber="55">Male</emphasis>
</emphasis>
(N = 2): Total length = 24.00–26.00, carapace length / width = 1.30–1.37, carapace height / width = 0.55–0.75, PER / carapace width = 0.35–0.35, PER / OAL = 2.82–2.86, PER / AER = 1.33–1.35, OAL / OQL = 1.16–1.35, OQP / OQA 1.09–1.21, cheliceral length / clypeus height = 5.00–15.00, femur I length / carapace width = 1.29–1.42, metatarsus I length / carapace width 1.33–1.43, femur IV length / carapace width 1.23–1.32, cymbium length / carapace width = 0.62–0.78, cymbium length / palpal patella length = 2.83–3.77, cymbium length / palpal tibia length = 2.50–3.95, cymbium length / palpal femur length = 0.92–1.00, palpal tibia length / palpal patella length = 0.95–1.13. Male genitalia as in
<figureCitation id="EBB8FEA01873FFED458630AFA805E8FB" box="[713,818,416,439]" captionStart="FIGURE" captionStartId="128.[155,167,1002,1021]" captionTargetBox="[120,1176,216,980]" captionTargetId="figure-11@128.[120,1176,216,980]" captionTargetPageId="128" captionText="FIGURE 28. Uduba spp., left male palpal bulbs, apical views. A. U. hiragasy, holotype (CASENT9065679) from Tsinjoarivo Forest. B. U. valiha, paratype (CASENT9082181) from Forêt de Vevembe. C. U. fisheri, paratype (CASENT9026388) from Anjozorobe. D. U. halabe, (CASENT9042520) from Parc National Ranomafana. E. U. platnicki, holotype (CASENT9006021) from Marojejy Reserve. F. U. kavanaughi, paratype (CASENT9031127) from Talatakely, Parc National Ranomafana. C, conductor; MA, median apophysis; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum, typically with apex extending to prolateral. Scale bar = 0.2 mm (A–C); 0.5 mm (D–F). Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.13158629" httpUri="https://zenodo.org/record/13158629/files/figure.png" pageId="55" pageNumber="56">Figs. 28 F</figureCitation>
,
<figureCitation id="EBB8FEA01873FFED440F30AFA8A2E8FB" box="[832,917,416,439]" captionStart="FIGURE" captionStartId="143.[155,167,912,931]" captionTargetBox="[135,1159,226,890]" captionTargetId="figure-16@143.[120,1176,216,899]" captionTargetPageId="143" captionText="FIGURE 47. Uduba kavanaughi, new species, Holotype (CASENT9029870) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158679" httpUri="https://zenodo.org/record/13158679/files/figure.png" pageId="55" pageNumber="56">47 A–C</figureCitation>
.
<emphasis id="41F73E371873FFED44EE30AFA8C3E8FB" bold="true" box="[929,1012,416,439]" pageId="55" pageNumber="56">Female</emphasis>
(N = 2): Total length = 31.00–37.00, carapace length / width = 1.38–1.52, carapace height / width = 0.56–0.57, PER / carapace width = 0.41–0.42, PER / OAL = 3.64–4.05, OAL / OQL = 1.00–1.02, OQP / OQA = 1.20–1.25, clypeus height / AME = 2.50–3.50, cheliceral length / clypeus height = 5.57–7.75, sternum length / width = 1.21–1.25, palpal coxa length / width = 2.35–2.39, femur I length / carapace width = 0.94–1.03, metatarsus I length / carapace width = 0.80–0.86, palpal tarsus length / carapace width = 0.49–0.59, palpal tarsus length / palpal patella length = 1.62–2.19, palpal tarsus length / palpal tibia length = 1.58–1.94, palpal tarsus length / palpal femur length = 1.00–1.13, palpal tibia length / palpal patella length = 1.03–1.13.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFED47E433C9A88BEC44" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="733CE2251873FFED47E433C9A9EAED28" blockId="55.[122,1175,220,1353]" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E433C9AABBEB91" bold="true" box="[171,396,710,733]" pageId="55" pageNumber="56">Material examined.</emphasis>
<materialsCitation id="C3EBE8781873FFED46DB33C9ABDBEA72" collectingDate="2009-01-11" collectingDateMax="2009-01-12" collectingDateMin="2009-01-11" collectionCode="CAS" collectorName="C. Griswold" country="Madagascar" elevation="970" latitude="-21.264055" location="Talatakely" longLatPrecision="1" longitude="47.419334" municipality="Parc National Ranomafana" pageId="55" pageNumber="56" specimenCode="CASENT9029870" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectingCountry id="0B94A2B51873FFED46DB33C9A961EB91" box="[404,598,710,733]" name="Madagascar" pageId="55" pageNumber="56">MADAGASCAR</collectingCountry>
:
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<collectingRegion id="B1472CC71873FFED452D33C9A86DEB91" box="[610,858,710,733]" country="Madagascar" name="Fianarantsoa" pageId="55" pageNumber="56">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1873FFED442433C9AFA5EB91" box="[875,1170,710,733]" pageId="55" pageNumber="56">Parc National Ranomafana</collectingMunicipality>
,
<location id="765CB4FE1873FFED473433E9ABD3EBB1" LSID="urn:lsid:plazi:treatment:FB2A53331871FFED4734332EA817EC05:765CB4FE1873FFED473433E9ABD3EBB1" box="[123,228,742,765]" country="Madagascar" latitude="-21.264055" longLatPrecision="1" longitude="47.419334" municipality="Parc National Ranomafana" name="Talatakely" pageId="55" pageNumber="56" stateProvince="Fianarantsoa">Talatakely</location>
,
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<quantity id="B47B4FC01873FFED47BA33E9AA7AEBB1" box="[245,333,742,765]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="55" pageNumber="56" unit="km" value="1.68">1.68 km</quantity>
SW Ranomafana
</locationDeviation>
,
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,
<geoCoordinate id="16B784E21873FFED45F333E9A809EBB1" box="[700,830,742,765]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="55" pageNumber="56" precision="1" seconds="9.6" value="47.419334">47º25ʹ9.6ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01873FFED44C933E9A8F2EBB1" box="[902,965,742,765]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="55" pageNumber="56" unit="m" value="970.0">
<elevation id="F8AE05161873FFED44C933E9A8F2EBB1" box="[902,965,742,765]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="55" pageNumber="56" unit="m" value="970.0">970m</elevation>
</quantity>
, general collecting day and night, evergreen forest,
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<collectingDate id="17793D0D1873FFED46953208A981EA52" box="[474,694,775,798]" pageId="55" pageNumber="56" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">11–12 January 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31873FFED458C3208A871EA52" box="[707,838,775,798]" pageId="55" pageNumber="56">C. Griswold</collectorName>
(
<typeStatus id="AC385C871873FFED44193208A88AEA52" box="[854,957,775,798]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1873FFED44853208AFA5EA52" box="[970,1170,775,798]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9029870</specimenCode>
,
<specimenCount id="658529AC1873FFED47343228ABABEA72" box="[123,156,807,830]" count="1" pageId="55" pageNumber="56" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01873FFED47E23228ABD3EA72" box="[173,228,807,830]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781873FFED47B63228A8A6EA13" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" pageId="55" pageNumber="56" specimenCode="CASENT9003516" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa Province" typeStatus="paratype">
<location id="765CB4FE1873FFED47B63228AAA8EA72" LSID="urn:lsid:plazi:treatment:FB2A53331871FFED4734332EA817EC05:765CB4FE1873FFED47B63228AAA8EA72" box="[249,415,807,830]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" name="Talatakely area" pageId="55" pageNumber="56" stateProvince="Fianarantsoa Province">Talatakely area</location>
, elev.
<quantity id="B47B4FC01873FFED46A83228A913EA72" box="[487,548,807,830]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="m" value="900.0">
<elevation id="F8AE05161873FFED46A83228A913EA72" box="[487,548,807,830]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="m" value="900.0">900m</elevation>
</quantity>
, mixed tropical forest,
<date id="073DC4E51873FFED44603228AF34EA72" box="[815,1027,807,830]" pageId="55" pageNumber="56" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="17793D0D1873FFED44603228AF34EA72" box="[815,1027,807,830]" pageId="55" pageNumber="56" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2–22 January 2001</collectingDate>
</date>
,
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,
<geoCoordinate id="16B784E21873FFED47343247ABCAEA13" box="[123,253,840,863]" degrees="47.41945" direction="east" orientation="longitude" pageId="55" pageNumber="56" precision="1" value="47.41945">47.41945°E</geoCoordinate>
,
<collectorName id="DE7687F31873FFED46453247AAFCEA13" box="[266,459,840,863]" pageId="55" pageNumber="56">D. H. Kavanaugh</collectorName>
(
<typeStatus id="AC385C871873FFED46933247A90BEA13" box="[476,572,840,863]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1873FFED45053247A825EA13" box="[586,786,840,863]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9003516</specimenCode>
,
<specimenCount id="658529AC1873FFED446F3247A876EA13" box="[800,833,840,863]" count="1" pageId="55" pageNumber="56" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01873FFED441D3247A8BEEA13" box="[850,905,840,863]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781873FFED44D13247AF2CEAEC" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="D. H. and K. M. Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" pageId="55" pageNumber="56" specimenCode="CASENT9003517" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa Province" typeStatus="paratype">
<location id="765CB4FE1873FFED44D13247AF73EA13" LSID="urn:lsid:plazi:treatment:FB2A53331871FFED4734332EA817EC05:765CB4FE1873FFED44D13247AF73EA13" box="[926,1092,840,863]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" name="Talatakely area" pageId="55" pageNumber="56" stateProvince="Fianarantsoa Province">Talatakely area</location>
,
<quantity id="B47B4FC01873FFED431D3247AFA6EA13" box="[1106,1169,840,863]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="m" value="900.0">
<elevation id="F8AE05161873FFED431D3247AFA6EA13" box="[1106,1169,840,863]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="m" value="900.0">900m</elevation>
</quantity>
, mixed tropical forest,
<date id="073DC4E51873FFED46263266A90FEACC" box="[361,568,873,896]" pageId="55" pageNumber="56" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="17793D0D1873FFED46263266A90FEACC" box="[361,568,873,896]" pageId="55" pageNumber="56" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2–22 January 2001</collectingDate>
</date>
,
<geoCoordinate id="16B784E21873FFED450B3266A9F3EACC" box="[580,708,873,896]" degrees="21.25041" direction="south" orientation="latitude" pageId="55" pageNumber="56" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
,
<geoCoordinate id="16B784E21873FFED459F3266A865EACC" box="[720,850,873,896]" degrees="47.41945" direction="east" orientation="longitude" pageId="55" pageNumber="56" precision="1" value="47.41945">47.41945°E</geoCoordinate>
,
<collectorName id="DE7687F31873FFED44123266AFA6EACC" box="[861,1169,873,896]" pageId="55" pageNumber="56">D. H. and K. M. Kavanaugh</collectorName>
,
<collectorName id="DE7687F31873FFED47343286ABC6EAEC" box="[123,241,905,928]" pageId="55" pageNumber="56">R. L. Brett</collectorName>
,
<collectorName id="DE7687F31873FFED47B23286AA54EAEC" box="[253,355,905,928]" pageId="55" pageNumber="56">E. Elsom</collectorName>
, and
<collectorName id="DE7687F31873FFED46D33286AACEEAEC" box="[412,505,905,928]" pageId="55" pageNumber="56">F. Vargas</collectorName>
(
<typeStatus id="AC385C871873FFED45473286A95FEAEC" box="[520,616,905,928]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1873FFED453B3286A80BEAEC" box="[628,828,905,928]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9003517</specimenCode>
,
<specimenCount id="658529AC1873FFED44083286A85EEAEC" box="[839,873,905,928]" count="1" pageId="55" pageNumber="56" type="female">1♀</specimenCount>
,
<specimenCount id="658529AC1873FFED44383286A8FAEAEC" box="[887,973,905,928]" count="1" pageId="55" pageNumber="56" type="male">1 juv. ♂</specimenCount>
,
<collectionCode id="15927AE01873FFED44943286AF25EAEC" box="[987,1042,905,928]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781873FFED436A3286A86EED4E" collectingDate="2009-01-11" collectingDateMax="2009-01-12" collectingDateMin="2009-01-11" collectionCode="CAS" collectorName="C. Griswold &amp; A. Saucedo &amp; H. Wood" country="Madagascar" elevation="970" latitude="-21.264055" location="Talatakely" pageId="55" pageNumber="56" specimenCode="CASENT9031127" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa Province" typeStatus="paratype">
<location id="765CB4FE1873FFED436A3286AFB9EAEC" LSID="urn:lsid:plazi:treatment:FB2A53331871FFED4734332EA817EC05:765CB4FE1873FFED436A3286AFB9EAEC" box="[1061,1166,905,928]" country="Madagascar" latitude="-21.264055" name="Talatakely" pageId="55" pageNumber="56" stateProvince="Fianarantsoa Province">Talatakely</location>
,
<locationDeviation id="628EFB071873FFED473432A5AAA2EA8D" box="[123,405,938,961]" location="Ranomafana" pageId="55" pageNumber="56">
<quantity id="B47B4FC01873FFED473432A5ABE5EA8D" box="[123,210,938,961]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="55" pageNumber="56" unit="km" value="1.68">1.68 km</quantity>
SW Ranomafana
</locationDeviation>
,
<geoCoordinate id="16B784E21873FFED46ED32A5A919EA8D" box="[418,558,938,961]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="55" pageNumber="56" precision="1" seconds="50.6" value="-21.264055">21º15ʹ50.6ʺS</geoCoordinate>
, 47º9.6ʺE, elev.
<quantity id="B47B4FC01873FFED45AC32A5A815EA8D" box="[739,802,938,961]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="55" pageNumber="56" unit="m" value="970.0">
<elevation id="F8AE05161873FFED45AC32A5A815EA8D" box="[739,802,938,961]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="55" pageNumber="56" unit="m" value="970.0">970m</elevation>
</quantity>
, general collecting day and night, evergreen forest,
<date id="073DC4E51873FFED467B32C4A939EAAE" box="[308,526,971,994]" pageId="55" pageNumber="56" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">
<collectingDate id="17793D0D1873FFED467B32C4A939EAAE" box="[308,526,971,994]" pageId="55" pageNumber="56" value="2009-01-11" valueMax="2009-01-12" valueMin="2009-01-11">11–12 January 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31873FFED455632C4A9A9EAAE" box="[537,670,971,994]" pageId="55" pageNumber="56">C. Griswold</collectorName>
,
<collectorName id="DE7687F31873FFED45E932C4A816EAAE" box="[678,801,971,994]" pageId="55" pageNumber="56">A. Saucedo</collectorName>
and
<collectorName id="DE7687F31873FFED441932C4A884EAAE" box="[854,947,971,994]" pageId="55" pageNumber="56">H. Wood</collectorName>
[CGM01] (
<typeStatus id="AC385C871873FFED437E32C4AFA6EAAE" box="[1073,1169,971,994]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1873FFED473432E4AA76ED4E" box="[123,321,1003,1026]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9031127</specimenCode>
,
<specimenCount id="658529AC1873FFED461D32E4AA43ED4E" box="[338,372,1003,1026]" count="1" pageId="55" pageNumber="56" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01873FFED46C732E4AA88ED4E" box="[392,447,1003,1026]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
), Talatakely forest,
<quantity id="B47B4FC01873FFED45EA32E4A833ED4E" box="[677,772,1003,1026]" metricMagnitude="4" metricUnit="m" metricValue="4.2299999999999995" pageId="55" pageNumber="56" unit="km" value="42.3">42.3 km</quantity>
58˚NE
</materialsCitation>
<materialsCitation id="C3EBE8781873FFED442832E4A9EEED28" collectingDate="2005-12-24" collectingDateMax="2006-01-14" collectingDateMin="2005-12-24" collectionCode="CAS" collectorName="H. Wood &amp; J. Miller &amp; J. J. Rafanomezantsoa &amp; Rajeriarison &amp; Andriamananona" country="Madagascar" elevation="1050" location="Fianarantsoa" pageId="55" pageNumber="56" specimenCode="CASENT9024063" specimenCount="1" specimenCount-juvenile="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingRegion id="B1472CC71873FFED442832E4A8C5ED4E" box="[871,1010,1003,1026]" country="Madagascar" name="Fianarantsoa" pageId="55" pageNumber="56">Fianarantsoa</collectingRegion>
, S21˚15ʹ28.0ʺ, E47˚25ʹ21.8ʺ, elev.
<quantity id="B47B4FC01873FFED46033503AAAEED6F" box="[332,409,1036,1059]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="55" pageNumber="56" unit="m" value="1050.0">
<elevation id="F8AE05161873FFED46033503AAAEED6F" box="[332,409,1036,1059]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="55" pageNumber="56" unit="m" value="1050.0">1050m</elevation>
</quantity>
, montane rainforest, general collecting,
<date id="073DC4E51873FFED44083503AF23ED6F" box="[839,1044,1036,1059]" pageId="55" pageNumber="56" value="2005-12-24">
<collectingDate id="17793D0D1873FFED44083503AF23ED6F" box="[839,1044,1036,1059]" pageId="55" pageNumber="56" value="2005-12-24">24 December 2005</collectingDate>
</date>
–
<date id="073DC4E51873FFED436C3503AB84ED08" pageId="55" pageNumber="56" value="2006-01-14">
<collectingDate id="17793D0D1873FFED436C3503AB84ED08" pageId="55" pageNumber="56" value="2006-01-14">14 January 2006</collectingDate>
</date>
,
<collectorName id="DE7687F31873FFED478F3522AA15ED08" box="[192,290,1069,1092]" pageId="55" pageNumber="56">H. Wood</collectorName>
,
<collectorName id="DE7687F31873FFED46603522AABCED08" box="[303,395,1069,1092]" pageId="55" pageNumber="56">J. Miller</collectorName>
,
<collectorName id="DE7687F31873FFED46D53522A9B8ED08" box="[410,655,1069,1092]" pageId="55" pageNumber="56">J. J. Rafanomezantsoa</collectorName>
, E.
<collectorName id="DE7687F31873FFED45F23522A875ED08" box="[701,834,1069,1092]" pageId="55" pageNumber="56">Rajeriarison</collectorName>
, V.
<collectorName id="DE7687F31873FFED44273522AF15ED08" box="[872,1058,1069,1092]" pageId="55" pageNumber="56">Andriamananona</collectorName>
, [HW009] [DNA-MAD-111] (
<specimenCode id="23254A5E1873FFED46053542A924ED28" box="[330,531,1101,1124]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9024063</specimenCode>
,
<specimenCount id="658529AC1873FFED45503542A9BAED28" box="[543,653,1101,1124]" count="1" pageId="55" pageNumber="56" type="juvenile">1 juvenile</specimenCount>
,
<collectionCode id="15927AE01873FFED45D63542A9E6ED28" box="[665,721,1101,1124]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251873FFED47E43561A88BEC44" blockId="55.[122,1175,220,1353]" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E43561AA68EDC9" bold="true" box="[171,351,1134,1157]" pageId="55" pageNumber="56">Natural history.</emphasis>
The species
<taxonomicName id="B48399A61873FFED46A33561A9F7EDC9" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[492,704,1134,1157]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="41F73E371873FFED46A33561A9F7EDC9" bold="true" box="[492,704,1134,1157]" italics="true" pageId="55" pageNumber="56">Uduba kavanaughi</emphasis>
</taxonomicName>
has been collected in rainforest. Males and females of this large, ecribellate species have been found wandering at night. David Kavanaugh (pers. Comm.) reports that several females were found in burrows from which they emerged for short distances at night, and into which they quickly retreated when disturbed. These burrows never had any extended turrets or even evident silk atop or in any of these burrows.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFED47E4341EA817EC05" pageId="55" pageNumber="56" type="distribution">
<paragraph id="733CE2251873FFED47E4341EA817EC05" blockId="55.[122,1175,220,1353]" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E4341EAA0AEC64" bold="true" box="[171,317,1297,1320]" pageId="55" pageNumber="56">Distribution.</emphasis>
Specimens of
<taxonomicName id="B48399A61873FFED46B9341EA9E4EC64" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[502,723,1297,1320]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="41F73E371873FFED46B9341EA9E4EC64" bold="true" box="[502,723,1297,1320]" italics="true" pageId="55" pageNumber="56">Uduba kavanaughi</emphasis>
</taxonomicName>
are known only from the vicinity of Ranomafana Forest, in southeastern
<collectingCountry id="0B94A2B51873FFED454A343DA9B1EC05" box="[517,646,1330,1353]" name="Madagascar" pageId="55" pageNumber="56">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA01873FFED45DA343DA9DEEC05" box="[661,745,1330,1353]" captionStart="MAP" captionStartId="180.[155,174,1080,1099]" captionTargetBox="[120,1176,216,1057]" captionTargetId="figure-12@180.[648,1176,216,1047]" captionTargetPageId="180" captionText="MAP 9. Distributions of Uduba fisheri, Uduba MAP 10. Distributions of Uduba hiragasy, Uduba rakotofrah, Uduba rajery, Uduba rakotozafy and Uduba valiha, Uduba taralily, Uduba danielae and Uduba sarotra. kavanaughi." figureDoi="http://doi.org/10.5281/zenodo.13158785" httpUri="https://zenodo.org/record/13158785/files/figure.png" pageId="55" pageNumber="56">Maps 9</figureCitation>
,
<figureCitation id="EBB8FEA01873FFED45B9343DA824EC05" box="[758,787,1330,1353]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="55" pageNumber="56">13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

291
data/FB/2A/53/FB2A53331873FFE247343475AA50EF16.xml Normal file
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@ -0,0 +1,291 @@
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<mods:title id="86573798B4799A63F3A53E4CF44BD1E5">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:name id="3ED59552553607C2DB98A2FA05FECC0F" type="personal">
<mods:role id="91F0310FF3A83CA390061CE5AD0A1402">
<mods:roleTerm id="6BA70B50153F85BC6077090CAF8B9EE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CECCC017FB4A22AB027C04BABD8CBE35">Griswold, Charles</mods:namePart>
<mods:affiliation id="0FAFA4EF0B16F880DB4342041E118D2E">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="6BF9693859D458456E5D6CF4924A9EB3" type="personal">
<mods:role id="6EC9AF699E9C2923FFEB18B335713D53">
<mods:roleTerm id="A06B8122F73543384C9E8B2FAE169FD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D92F13431E86AD3E563F451F1BD96E54">Ubick, Darrell</mods:namePart>
<mods:affiliation id="2F400A6DEDC5BC97A7B4D34ECACED4AA">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="DF81539C43DDCC631FAE4D98DC232915" type="personal">
<mods:role id="C1C8AFBEB8935DA54574670DC1A9DE1F">
<mods:roleTerm id="D10A4474D362F7B067851A08A05D10FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1049DF69D81E0D4E47F2CA4B6C2D864D">Ledford, Joel</mods:namePart>
<mods:affiliation id="9D4F1CE49DD3806DE6B72E68D56C50D2">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
</mods:name>
<mods:name id="12964D854E6A1B310E6D70FAEECEAE5B" type="personal">
<mods:role id="71E0CA4E60157AC3A424C17CC8DAE11C">
<mods:roleTerm id="B15C61719380753F5F38FBD01E26EC51">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD68A97B4449F9887A6B24B291C9249F">Polotow, Daniele</mods:namePart>
<mods:affiliation id="B13A53D3E34EC4F423AB38A9EFDC75BC">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
</mods:name>
<mods:typeOfResource id="010D0AB304B81018EC37324256001852">text</mods:typeOfResource>
<mods:relatedItem id="D09D020305EA81193F153B972827341B" type="host">
<mods:titleInfo id="D90625C3F1AD474EF014B6773C2D939E">
<mods:title id="515BBBEFF340412349B625ABB4FA202D">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="657E62FF5B3272809CD009349C3755EE">
<mods:date id="67199177F31CDF2038B96E7667CE4110">2022</mods:date>
<mods:detail id="DCBB1E2E1FAC11A2DF4E4CAD3EC4DE95" type="pubDate">
<mods:number id="4373E4321A98353F3C31C532FF1CEC2C">2022-12-15</mods:number>
</mods:detail>
<mods:detail id="CF804875605FAD6FA2F2064BBCDD4C0F" type="volume">
<mods:number id="B8DF42B247F983E55C23DEACB06922FA">67</mods:number>
</mods:detail>
<mods:extent id="A8C38E2B9CD8DAAD157D06D150B3BF88" unit="page">
<mods:start id="AD64D234FDDFCD599D87C4DCA40A2BA0">1</mods:start>
<mods:end id="B194E5AAB631D180EF7DD6A8A88E01C3">193</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D608CBCABEEBB404B8597D2D137BC159">journal article</mods:classification>
<mods:identifier id="3257AC0E9B48C5D7EA3B1B6B8CDB6651" type="DOI">10.5281/zenodo.13158554</mods:identifier>
<mods:identifier id="7408D1F6CAB6A844A7A21A81FB6279B9" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="8A3DB4ED9ECD6DE7CC25AB9D6A622A1B" type="Zenodo-Dep">13158554</mods:identifier>
<mods:identifier id="0CD223B55D4172D79C948FE692393BC2" type="ZooBank">74E0D22E-9890-4FF0-80AB-8FCBBA976B2F</mods:identifier>
</mods:mods>
<treatment id="FB2A53331873FFE247343475AA50EF16" LSID="urn:lsid:zoobank.org:act:A425BEC2-CFA9-4B1E-A380-508DC80A2E42" httpUri="http://treatment.plazi.org/id/FB2A53331873FFE247343475AA50EF16" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
<subSubSection id="3B99B1AE1873FFED47343475AA92ECDD" box="[123,421,1402,1425]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="733CE2251873FFED47343475AA92ECDD" blockId="55.[123,421,1402,1456]" box="[123,421,1402,1425]" pageId="55" pageNumber="56">
<heading id="287455491873FFED47343475AA92ECDD" box="[123,421,1402,1425]" fontSize="10" level="2" pageId="55" pageNumber="56" reason="4">
<emphasis id="41F73E371873FFED47343475AA92ECDD" bold="true" box="[123,421,1402,1425]" pageId="55" pageNumber="56">
<taxonomicName id="B48399A61873FFED47343475AA24ECDD" ID-CoL="C8KJM" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,275,1402,1425]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="lakroa" status="sp. nov.">
<emphasis id="41F73E371873FFED47343475AA24ECDD" bold="true" box="[123,275,1402,1425]" italics="true" pageId="55" pageNumber="56">Uduba lakroa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1873FFED466E3475AA92ECDD" box="[289,421,1402,1425]" pageId="55" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFED47343494AA97ECFC" box="[123,416,1435,1456]" pageId="55" pageNumber="56" type="description">
<paragraph id="733CE2251873FFED47343494AA97ECFC" blockId="55.[123,421,1402,1456]" box="[123,416,1435,1456]" pageId="55" pageNumber="56">
<figureCitation id="EBB8FEA01873FFED47343494AA21ECFC" box="[123,278,1435,1456]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="55" pageNumber="56">Figures 48 A–C</figureCitation>
, Maps 12, 21.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFED47E434CEA8DEEF17" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="733CE2251873FFED47E434CEA8DEEF17" blockId="55.[123,1174,1473,1725]" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E434CEAA79EC94" bold="true" box="[171,334,1473,1496]" pageId="55" pageNumber="56">Type material.</emphasis>
<materialsCitation id="C3EBE8781873FFED461B34CEA8DEEF17" collectingDate="2003-10-26" collectingDateMax="2003-10-31" collectingDateMin="2003-10-26" collectionCode="FMNH, CAS" collectorName="L. E. Olsen" country="Madagascar" elevation="875" location="Andrauorena River" municipality="Mt. Papango in Parc National de Midousy" pageId="55" pageNumber="56" specimenCode="CASENT9064782, CASENT9062274, CASENT9062280, CASENT9062275" specimenCount="3" specimenCount-male="3" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="AC385C871873FFED461B34CEAA80EC94" box="[340,439,1473,1496]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E1873FFED46B034CEA9F0EC94" box="[511,711,1473,1496]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9064782</specimenCode>
)
<collectingMethod id="AAC29A321873FFED459A34CEA8BEEC94" box="[725,905,1473,1496]" pageId="55" pageNumber="56">from pitfall traps</collectingMethod>
at
<quantity id="B47B4FC01873FFED44E634CEA8DBEC94" box="[937,1004,1473,1496]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="55" pageNumber="56" unit="m" value="875.0">
<elevation id="F8AE05161873FFED44E634CEA8DBEC94" box="[937,1004,1473,1496]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="55" pageNumber="56" unit="m" value="875.0">875 m</elevation>
</quantity>
elevation along
<location id="765CB4FE1873FFED473434EDAA78ECB5" LSID="urn:lsid:plazi:treatment:FB2A53331873FFE247343475AA50EF16:765CB4FE1873FFED473434EDAA78ECB5" box="[123,335,1506,1529]" country="Madagascar" municipality="Mt. Papango in Parc National de Midousy" name="Andrauorena River" pageId="55" pageNumber="56" stateProvince="Fianarantsoa">Andrauorena River</location>
on the NE slope
<collectingMunicipality id="9358785F1873FFED456534EDAF26ECB5" box="[554,1041,1506,1529]" pageId="55" pageNumber="56">Mt. Papango in Parc National de Midousy</collectingMunicipality>
, 23°50.1ʹS, 46°57.8ʹE,
<collectingRegion id="B1472CC71873FFED47B4370CAAFFEF56" box="[251,456,1539,1562]" country="Madagascar" name="Fianarantsoa" pageId="55" pageNumber="56">Fianarantsoa Prov.</collectingRegion>
,
<collectingCountry id="0B94A2B51873FFED4690370CA969EF56" box="[479,606,1539,1562]" name="Madagascar" pageId="55" pageNumber="56">Madagascar</collectingCountry>
, collected
<date id="073DC4E51873FFED45AC370CA8E7EF56" box="[739,976,1539,1562]" pageId="55" pageNumber="56" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">
<collectingDate id="17793D0D1873FFED45AC370CA8E7EF56" box="[739,976,1539,1562]" pageId="55" pageNumber="56" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">26–31 October 2003</collectingDate>
</date>
by
<collectorName id="DE7687F31873FFED4345370CAFA2EF56" box="[1034,1173,1539,1562]" pageId="55" pageNumber="56">L. E. Olsen</collectorName>
[FMNHLEO128], deposited in
<collectionCode id="15927AE01873FFED4683372CA92AEF76" box="[460,541,1571,1594]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="55" pageNumber="56" type="Museum">FMNH</collectionCode>
.
<specimenCount id="658529AC1873FFED456A372CA839EF76" box="[549,782,1571,1594]" count="3" pageId="55" pageNumber="56" type="male" typeStatus="paratype">Three paratype males</specimenCount>
, same data, two deposited in
<collectionCode id="15927AE01873FFED4307372CAFA2EF76" box="[1096,1173,1571,1594]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="55" pageNumber="56" type="Museum">FMNH</collectionCode>
(
<specimenCode id="23254A5E1873FFED47CD374BAA7FEF17" box="[130,328,1604,1627]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9062274</specimenCode>
and
<specimenCode id="23254A5E1873FFED4630374BA971EF17" box="[383,582,1604,1627]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9062280</specimenCode>
) and one in
<collectionCode id="15927AE01873FFED4580374BA834EF17" box="[719,771,1604,1627]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="55" pageNumber="56" type="Museum">CAS</collectionCode>
(
<specimenCode id="23254A5E1873FFED445D374BA8E9EF17" box="[786,990,1604,1627]" collectionCode="CASENT" pageId="55" pageNumber="56">CASENT9062275</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFED47E4376AAB9BEFD0" pageId="55" pageNumber="56" type="etymology">
<paragraph id="733CE2251873FFED47E4376AAB9BEFD0" blockId="55.[123,1174,1473,1725]" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E4376AAA1DEF30" bold="true" box="[171,298,1637,1660]" pageId="55" pageNumber="56">Etymology.</emphasis>
The species epithet is from a Malagasy word for cross,
<emphasis id="41F73E371873FFED44C7376AA8FBEF30" box="[904,972,1637,1660]" italics="true" pageId="55" pageNumber="56">lakroa</emphasis>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1873FFE247E437A9A80DE87F" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="733CE2251873FFE247E437A9A80DE87F" blockId="55.[123,1174,1473,1725]" lastBlockId="56.[123,1175,220,1626]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
<emphasis id="41F73E371873FFED47E437A9AA89EFF1" bold="true" box="[171,446,1702,1725]" pageId="55" pageNumber="56">
Diagnosis.
<taxonomicName id="B48399A61873FFED466937A9AA89EFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,446,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="lakroa">
<emphasis id="41F73E371873FFED466937A9AA89EFF1" bold="true" box="[294,446,1702,1725]" italics="true" pageId="55" pageNumber="56">Uduba lakroa</emphasis>
</taxonomicName>
</emphasis>
are one of the Unclassified species (
<figureCitation id="EBB8FEA01873FFED440837A9A8AAEFF1" box="[839,925,1702,1725]" captionStart="MAP" captionStartId="186.[421,440,1168,1187]" captionTargetBox="[388,916,216,1150]" captionTargetId="figure-11@186.[388,916,216,1150]" captionTargetPageId="186" captionText="MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863)." figureDoi="http://doi.org/10.5281/zenodo.13158799" httpUri="https://zenodo.org/record/13158799/files/figure.png" pageId="55" pageNumber="56">Map 21</figureCitation>
), which have a divided cribellum. Males of
<taxonomicName id="B48399A6187CFFE2461931D3AA88E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[342,447,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="lakroa">
<emphasis id="41F73E37187CFFE2461931D3AA88E9BF" bold="true" box="[342,447,220,243]" italics="true" pageId="56" pageNumber="57">U. lakroa</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="B48399A6187CFFE244CF31D3A8FDE9BF" authorityName="Simon" authorityYear="1880" box="[896,970,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37187CFFE244CF31D3A8FDE9BF" bold="true" box="[896,970,220,243]" italics="true" pageId="56" pageNumber="57">Uduba</emphasis>
</taxonomicName>
by having the palpus with a swollen, dark lobe near the base of TA2 and swollen, unsclerotized lobe near the base of TA3 and the conductor (
<figureCitation id="EBB8FEA0187CFFE246D13013A919E87F" box="[414,558,284,307]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Figs. 48 A–C</figureCitation>
<emphasis id="41F73E37187CFFE245613013A900E87F" bold="true" box="[558,567,284,307]" pageId="56" pageNumber="57">)</emphasis>
. Females are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187CFFE247E43033A8F6EF56" pageId="56" pageNumber="57" type="description">
<paragraph id="733CE225187CFFE247E43033AAC1ED7B" blockId="56.[123,1175,220,1626]" pageId="56" pageNumber="57">
<emphasis id="41F73E37187CFFE247E43033AA01E81F" bold="true" box="[171,310,316,339]" pageId="56" pageNumber="57">Description.</emphasis>
Male (
<typeStatus id="AC385C87187CFFE246CF3033AADDE81F" box="[384,490,316,339]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
): Total length 13.80. Markings typical of
<taxonomicName id="B48399A6187CFFE244E93033A8C7E81F" authorityName="Simon" authorityYear="1880" box="[934,1008,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37187CFFE244E93033A8C7E81F" bold="true" box="[934,1008,316,339]" italics="true" pageId="56" pageNumber="57">Uduba</emphasis>
</taxonomicName>
. Carapace 8.10 long, 5.60 wide, 3.40 high; clypeus 0.40 high. Eye diameters: AME 0.24, ALE 0.28, PME 0.26, PLE 0.28. Chelicerae 3.70 long; sternum 3.40 long, 2.60 wide; labium 1.70 long, 1.20 wide; palpal coxae 2.60 long, 1.00 wide. Cribellum divided. Spination (
<typeStatus id="AC385C87187CFFE2446D3092A8B5E8F8" box="[802,898,413,436]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
, CASENT9062275): palpus–femur 1-1-0-1, p0-0-2, patella p0-1-0, tibia p1 [elongate, stout spine]-0-0; leg I–femur d1-1-1, p0-1-2, r0-1-1-1, patella p1, r1, tibia d0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p01-1-1, v2- 2-2, r1-1-0; leg II–femur d1-1-1, p0-1-2, r1-1-1-1, patella p1, r1, tibia d1-1-1, p0-1-1-0, v2-1-2-2- 2, r0-1-1-0, metatarsus p1-1-1, v2-2-2, r1-1-1; leg III–femur d1-0-1-1, patella p1, r1, tibia d1-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2; leg IV–femur d0-1-1-1, patella p1, r1, tibia d0-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements (combination of males CASENT9062274 and CASENT9062275): I: 6.70 + 2.80 + 6.20 + 6.10 + 3.60 = 25.40; II: 6.00 + 2.70 + 5.40 + 5.60 + 3.20 = 22.90; III: 5.60 + 2.30 + 4.10 + 5.50 + 3.00 = 20.50; IV: 6.70 + 2.50 + 5.70 + 7.30 + 3.40 = 25.60; palpus: 3.00 + 1.10 + 1.10 + NA + 3.30 = 8.50. Leg formula 4123.
<emphasis id="41F73E37187CFFE2435033D1AFA2EBB9" bold="true" box="[1055,1173,734,757]" pageId="56" pageNumber="57">Male palp</emphasis>
(
<typeStatus id="AC385C87187CFFE247CD33F0ABD5EA5A" box="[130,226,767,790]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
, CASENT9062275): palpal tibia 0.30 cymbial length, RTA very broad (
<figureCitation id="EBB8FEA0187CFFE244A533F0AF63EA5A" box="[1002,1108,767,790]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Fig. 48 C</figureCitation>
), RTA length 0.27 tibia length, length 0.57 times width, with blunt, curved apex, VTA small, slender, curved (
<figureCitation id="EBB8FEA0187CFFE2479B3230AA08EA1A" box="[212,319,831,854]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Fig. 48 A</figureCitation>
), length 1.60 times width, length 0.33 tibia width; tibia with an elongate, stout prolateral spine; tegulum convex, without ridge (
<figureCitation id="EBB8FEA0187CFFE245C73250A82BEA3A" box="[648,796,863,886]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Figs. 48 B, C</figureCitation>
), tegulum length 0.92 times width; TA1 extends to tegulum apex and slightly past TA3 apex; TA3 and TA2 both thick, curved, TA2 crosses in front of TA3, TA3 extends apicad of TA2 by 1.05 times tegulum length (
<figureCitation id="EBB8FEA0187CFFE244BF3290AF61EAFA" box="[1008,1110,927,950]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Fig. 48 B</figureCitation>
), MA transverse, bilobed, origin at 0.47 times tegulum length, tegulum apex at 0.32 tegulum length from MA apex, MA length (including apical processes) 0.72 times width, MA large but narrowly transverse (
<figureCitation id="EBB8FEA0187CFFE2465B350FAA9BED5B" box="[276,428,1024,1047]" captionStart="FIGURE" captionStartId="143.[155,167,1623,1642]" captionTargetBox="[123,1166,998,1596]" captionTargetId="figure-52@143.[120,1176,990,1610]" captionTargetPageId="143" captionText="FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158681" httpUri="https://zenodo.org/record/13158681/files/figure.png" pageId="56" pageNumber="57">Figs. 48 B, C</figureCitation>
), length 0.28 tegulum length, width 0.35 tegulum width; conductor fan entire.
<emphasis id="41F73E37187CFFE2466C352FAA41ED7B" bold="true" box="[291,374,1056,1079]" pageId="56" pageNumber="57">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="733CE225187CFFE247E4354FAFA1EC75" blockId="56.[123,1175,220,1626]" pageId="56" pageNumber="57">
<emphasis id="41F73E37187CFFE247E4354FAA6CED1B" bold="true" box="[171,347,1088,1111]" pageId="56" pageNumber="57">Variation. Male</emphasis>
(N= 4): Total length 13.00 to 14.50; ratios of carapace length / width = 1.40– 1.48, carapace height / width = 0.49–0.61, PER / carapace width = 0.37–0.40, PER / OAL = 3.03– 3.48, PER / AER = 1.32–1.39, OQP / OQA = 1.03–1.07, clypeal height / diameter AME = 1.25– 1.67, cheliceral length /clypeal height = 9.25–12.33, sternum length / width = 1.31–1.46; length femur I / carapace width = 1.25–1.37, metatarsus I /carapace width = 1.07–1.27, femur IV length / carapace width = 1.26–1.35, cymbium length / carapace width = 0.58–0.63, palpal tibia length / palpal patella length = 0.83–1.00, cymbium length / palpal tibia length = 2.75–3.00, cymbium length / palpal femur length = 1.00–1.11. Female variation is unknown: only males are recognized.
</paragraph>
<paragraph id="733CE225187CFFE247E4344DA8A9EC96" blockId="56.[123,1175,220,1626]" pageId="56" pageNumber="57">
<materialsCitation id="C3EBE878187CFFE247E4344DA8ADEC96" collectingDate="2003-10-26" collectingDateMax="2003-10-31" collectingDateMin="2003-10-26" collectionCode="FMNH, CAS" collectorName="Parc National de Midousy, NE &amp; L. E. Olsen" country="Madagascar" county="Parc National de Midousy" elevation="875" location="Andrauorena River" municipality="Mt. Papango" pageId="56" pageNumber="57" specimenCode="LEO128, CASENT9064782, CASENT9062274, CASENT9062280, CASENT9062275" specimenCount="4" specimenCount-male="4" stateProvince="Fianarantsoa" typeStatus="holotype">
M
<emphasis id="41F73E37187CFFE2478D344DAABCEC15" bold="true" box="[194,395,1346,1369]" pageId="56" pageNumber="57">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5187CFFE246DA344DA960EC15" box="[405,599,1346,1369]" name="Madagascar" pageId="56" pageNumber="57">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37187CFFE2452B344DA869EC15" box="[612,862,1346,1369]" italics="true" pageId="56" pageNumber="57">
<collectingRegion id="B1472CC7187CFFE2452B344DA869EC15" box="[612,862,1346,1369]" country="Madagascar" name="Fianarantsoa" pageId="56" pageNumber="57">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectorName id="DE7687F3187CFFE2443F344DABA8EC35" pageId="56" pageNumber="57">
<collectingCounty id="9A5D9AA9187CFFE2443F344DAFB9EC15" box="[880,1166,1346,1369]" pageId="56" pageNumber="57">Parc National de Midousy</collectingCounty>
, NE
</collectorName>
slope
<collectingMunicipality id="9358785F187CFFE247A9346DAA44EC35" box="[230,371,1378,1401]" pageId="56" pageNumber="57">Mt. Papango</collectingMunicipality>
, along
<location id="765CB4FE187CFFE246F0346DA9BCEC35" LSID="urn:lsid:plazi:treatment:FB2A53331873FFE247343475AA50EF16:765CB4FE187CFFE246F0346DA9BCEC35" box="[447,651,1378,1401]" country="Madagascar" county="Parc National de Midousy" municipality="Mt. Papango" name="Andrauorena River" pageId="56" pageNumber="57" stateProvince="Fianarantsoa">Andrauorena River</location>
,
<locationDeviation id="628EFB07187CFFE245D5346DA85DEC35" box="[666,874,1378,1401]" location="Befotaka" pageId="56" pageNumber="57">
<quantity id="B47B4FC0187CFFE245D5346DA9E7EC35" box="[666,720,1378,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="km" value="2.0">2 km</quantity>
SW Befotaka
</locationDeviation>
, 23°50.1ʹS, 46°57.8ʹE, elev.
<quantity id="B47B4FC0187CFFE24734348DAB8DECD5" box="[123,186,1410,1433]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="56" pageNumber="57" unit="m" value="875.0">
<elevation id="F8AE0516187CFFE24734348DAB8DECD5" box="[123,186,1410,1433]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="56" pageNumber="57" unit="m" value="875.0">875m</elevation>
</quantity>
, camp #1,
<collectingMethod id="AAC29A32187CFFE2467F348DAA96ECD5" box="[304,417,1410,1433]" pageId="56" pageNumber="57">pitfall trap</collectingMethod>
line #3,
<date id="073DC4E5187CFFE2454E348DA9D6ECD5" box="[513,737,1410,1433]" pageId="56" pageNumber="57" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">
<collectingDate id="17793D0D187CFFE2454E348DA9D6ECD5" box="[513,737,1410,1433]" pageId="56" pageNumber="57" value="2003-10-26" valueMax="2003-10-31" valueMin="2003-10-26">26–31 October 2003</collectingDate>
</date>
,
<collectorName id="DE7687F3187CFFE245A2348DA85CECD5" box="[749,875,1410,1433]" pageId="56" pageNumber="57">L. E. Olsen</collectorName>
, [
<collectionCode id="15927AE0187CFFE24432348DA8FAECD5" box="[893,973,1410,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
#
<specimenCode id="23254A5E187CFFE244AC348DAF76ECD5" box="[995,1089,1410,1433]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">LEO128</specimenCode>
] (
<typeStatus id="AC385C87187CFFE24319348DAB9BECF5" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
,
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,
<specimenCount id="658529AC187CFFE246D834ADAA8FECF5" box="[407,440,1442,1465]" count="1" pageId="56" pageNumber="57" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187CFFE2468334ADA92AECF5" box="[460,541,1442,1465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
), (
<typeStatus id="AC385C87187CFFE2457134ADA9A9ECF5" box="[574,670,1442,1465]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187CFFE245FF34ADA84FECF5" box="[688,888,1442,1465]" collectionCode="CASENT" pageId="56" pageNumber="57">CASENT9062274</specimenCode>
,
<specimenCount id="658529AC187CFFE244C534ADA89CECF5" box="[906,939,1442,1465]" count="1" pageId="56" pageNumber="57" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187CFFE244F034ADAF27ECF5" box="[959,1040,1442,1465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
), (
<typeStatus id="AC385C87187CFFE2437E34ADAFA6ECF5" box="[1073,1169,1442,1465]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187CFFE2473434CCAA74EC96" box="[123,323,1475,1498]" collectionCode="CASENT" pageId="56" pageNumber="57">CASENT9062280</specimenCode>
,
<specimenCount id="658529AC187CFFE2460134CCAA47EC96" box="[334,368,1475,1498]" count="1" pageId="56" pageNumber="57" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187CFFE2463134CCAAF8EC96" box="[382,463,1475,1498]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
), (
<typeStatus id="AC385C87187CFFE246A534CCA97DEC96" box="[490,586,1475,1498]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187CFFE2451934CCA829EC96" box="[598,798,1475,1498]" collectionCode="CASENT" pageId="56" pageNumber="57">CASENT9062275</specimenCode>
,
<specimenCount id="658529AC187CFFE2446534CCA87CEC96" box="[810,843,1475,1498]" count="1" pageId="56" pageNumber="57" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187CFFE2441534CCA8A5EC96" box="[858,914,1475,1498]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="56" pageNumber="57" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225187CFFE247E434ECA8F6EF56" blockId="56.[123,1175,220,1626]" pageId="56" pageNumber="57">
<emphasis id="41F73E37187CFFE247E434ECAA69ECB6" bold="true" box="[171,350,1507,1530]" pageId="56" pageNumber="57">Natural history.</emphasis>
The
<typeStatus id="AC385C87187CFFE246DD34ECAAF7ECB6" box="[402,448,1507,1530]" pageId="56" pageNumber="57">type</typeStatus>
locality is in a region of rainforest. Collection data reveal that males of the cribellate
<taxonomicName id="B48399A6187CFFE24662370CAAF1EF56" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[301,454,1539,1562]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="lakroa">
<emphasis id="41F73E37187CFFE24662370CAAF1EF56" bold="true" box="[301,454,1539,1562]" italics="true" pageId="56" pageNumber="57">Uduba lakroa</emphasis>
</taxonomicName>
wander, as they were collected in pitfall traps.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187CFFE247E4372CAA50EF16" pageId="56" pageNumber="57" type="distribution">
<paragraph id="733CE225187CFFE247E4372CAA50EF16" blockId="56.[123,1175,220,1626]" pageId="56" pageNumber="57">
<emphasis id="41F73E37187CFFE247E4372CAAEBEF76" bold="true" box="[171,476,1571,1594]" pageId="56" pageNumber="57">
Distribution.
<taxonomicName id="B48399A6187CFFE2460B372CAAEBEF76" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[324,476,1571,1594]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="lakroa">
<emphasis id="41F73E37187CFFE2460B372CAAEBEF76" bold="true" box="[324,476,1571,1594]" italics="true" pageId="56" pageNumber="57">Uduba lakroa</emphasis>
</taxonomicName>
</emphasis>
are known only from the
<typeStatus id="AC385C87187CFFE245B9372CA813EF76" box="[758,804,1571,1594]" pageId="56" pageNumber="57">type</typeStatus>
locality in far southeastern
<collectingCountry id="0B94A2B5187CFFE2431F372CAB88EF16" name="Madagascar" pageId="56" pageNumber="57">Madagascar</collectingCountry>
(Maps 12, 21).
</paragraph>
</subSubSection>
</treatment>
</document>

249
data/FB/2A/53/FB2A53331874FFEB47343211AA63EAD9.xml Normal file
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@ -0,0 +1,249 @@
<document id="DB7DAC37012362F32B3C0743B23DAA31" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331874FFEB47343211AA63EAD9" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba ida Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="E47530E8-6E43-4ECA-B6E2-D23973506817" docUuidSource="ZooBank" docVersion="2" lastPageNumber="50" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="49" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="612385DD65A3E5FA11E8E15790E8CEFF">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="89EDEBDA0135117A631CD1B17BF11058" type="personal">
<mods:role id="9A3611B5F0156E65214A99D80CE9023C">
<mods:roleTerm id="406FE5112E033CFA468A39CEDC6BFEF4">Author</mods:roleTerm>
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<mods:namePart id="60982ADDBD1A98775F2220771DD4031D">Griswold, Charles</mods:namePart>
<mods:affiliation id="BEF38EF703CE68523893AC32AACC8C31">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="6DA7A9A7BC73F1E716DD82F337566352">Ubick, Darrell</mods:namePart>
<mods:affiliation id="31F221E42899DA3ACEC8EF8AB8C52922">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="D7E38E2F03727B873AFB386317E7F498" type="personal">
<mods:role id="A0BC1BE34EBB78E32C539F169AE850E9">
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<mods:namePart id="4583C57FCB0EC9E017726399DE7040D3">Ledford, Joel</mods:namePart>
<mods:affiliation id="E35E9ED67509693908F8A368E6B601DE">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
</mods:name>
<mods:name id="5D48685DCD4C764E926930745E1CA603" type="personal">
<mods:role id="233D537DA6F2BA2FA629D830EDC54E82">
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<mods:namePart id="87E3286D39F1371ADBB47011FF555689">Polotow, Daniele</mods:namePart>
<mods:affiliation id="8201DCEB58E9342168308FAD572E1E7A">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:title id="F77DE118DDE26C34DA0E711C11A1FE19">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="66BCDBF3C8504A44841C3FB9B9753357">
<mods:date id="EE1307E04D6498DAE917D97B94E925A6">2022</mods:date>
<mods:detail id="2165CB917CEE63F4DD29E675F5138A6F" type="pubDate">
<mods:number id="A40F67833D351E19C270A625576A5548">2022-12-15</mods:number>
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<mods:number id="48C0B8FEE5745EE0C26AEFD2D831F218">67</mods:number>
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<mods:start id="C29991451CB1B98C5BC34D42A1371924">1</mods:start>
<mods:end id="88AC7655C0000C189F8CF885CF51905C">193</mods:end>
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<mods:classification id="1D54E030F2830607B9838DF252E35351">journal article</mods:classification>
<mods:identifier id="18676364A8BA0C2FD71BEA2290468BDB" type="DOI">10.5281/zenodo.13158554</mods:identifier>
<mods:identifier id="0A5D4E6170160DF2D35DCF09CFAE509C" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="643F1A3181124E1E75567D47EBE8EBD4" type="Zenodo-Dep">13158554</mods:identifier>
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<treatment id="FB2A53331874FFEB47343211AA63EAD9" LSID="urn:lsid:zoobank.org:act:E47530E8-6E43-4ECA-B6E2-D23973506817" httpUri="http://treatment.plazi.org/id/FB2A53331874FFEB47343211AA63EAD9" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="3B99B1AE1874FFEA47343211AAB7EA79" box="[123,384,798,821]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="733CE2251874FFEA47343211AAB7EA79" blockId="48.[123,406,798,851]" box="[123,384,798,821]" pageId="48" pageNumber="49">
<heading id="287455491874FFEA47343211AAB7EA79" box="[123,384,798,821]" fontSize="10" level="2" pageId="48" pageNumber="49" reason="4">
<emphasis id="41F73E371874FFEA47343211AAB7EA79" bold="true" box="[123,384,798,821]" pageId="48" pageNumber="49">
<taxonomicName id="B48399A61874FFEA47343211ABD9EA79" ID-CoL="C8KJH" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,238,798,821]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ida" status="sp. nov.">
<emphasis id="41F73E371874FFEA47343211ABD9EA79" bold="true" box="[123,238,798,821]" italics="true" pageId="48" pageNumber="49">Uduba ida</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1874FFEA47B33211AAB7EA79" box="[252,384,798,821]" pageId="48" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEA47343231AAA1EA1F" box="[123,406,830,851]" pageId="48" pageNumber="49" type="description">
<paragraph id="733CE2251874FFEA47343231AAA1EA1F" blockId="48.[123,406,798,851]" box="[123,406,830,851]" pageId="48" pageNumber="49">
<figureCitation id="EBB8FEA01874FFEA47343231AA2FEA1F" box="[123,280,830,851]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="48" pageNumber="49">Figures. 73 C, F</figureCitation>
,
<figureCitation id="EBB8FEA01874FFEA466D3231AA5CEA1F" box="[290,363,830,851]" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="48" pageNumber="49">Maps 5</figureCitation>
,
<figureCitation id="EBB8FEA01874FFEA46383231AAA5EA1F" box="[375,402,830,851]" captionStart="MAP" captionStartId="186.[421,440,1168,1187]" captionTargetBox="[388,916,216,1150]" captionTargetId="figure-11@186.[388,916,216,1150]" captionTargetPageId="186" captionText="MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863)." figureDoi="http://doi.org/10.5281/zenodo.13158799" httpUri="https://zenodo.org/record/13158799/files/figure.png" pageId="48" pageNumber="49">21</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEA47E4326CAF43EAF6" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="733CE2251874FFEA47E4326CAF43EAF6" blockId="48.[123,1174,867,1725]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA47E4326CAA67EA36" bold="true" box="[171,336,867,890]" pageId="48" pageNumber="49">Type material.</emphasis>
<materialsCitation id="C3EBE8781874FFEA4617326CAF43EAF6" collectingDate="2007-12-11" collectingDateMax="2007-12-13" collectingDateMin="2007-12-11" collectionCode="CAS" collectorName="Brian L. Fisher" country="Madagascar" elevation="980" latitude="-18.384722" location="Station Forestiere Analamazaotra" longLatPrecision="21" longitude="48.41278" pageId="48" pageNumber="49" specimenCode="CASENT9042534" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C871874FFEA4617326CAA8CEA36" box="[344,443,867,890]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="23254A5E1874FFEA4553326CA9D3EA36" box="[540,740,867,890]" collectionCode="CASENT" pageId="48" pageNumber="49">CASENT9042534</specimenCode>
) from sifted litter in montane rainforest at
<quantity id="B47B4FC01874FFEA47D5328CABE0EAD6" box="[154,215,899,922]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="48" pageNumber="49" unit="m" value="980.0">
<elevation id="F8AE05161874FFEA47D5328CABE0EAD6" box="[154,215,899,922]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="48" pageNumber="49" unit="m" value="980.0">980m</elevation>
</quantity>
elevation in
<location id="765CB4FE1874FFEA463C328CA9D7EAD6" LSID="urn:lsid:plazi:treatment:FB2A53331874FFEB47343211AA63EAD9:765CB4FE1874FFEA463C328CA9D7EAD6" box="[371,736,899,922]" country="Madagascar" latitude="-18.384722" longLatPrecision="21" longitude="48.41278" name="Station Forestiere Analamazaotra" pageId="48" pageNumber="49" stateProvince="Toamasina">Station Forestière Analamazaotra</location>
(
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,
<geoCoordinate id="16B784E21874FFEA44CE328CAF3BEAD6" box="[897,1036,899,922]" degrees="048" direction="east" minutes="24" orientation="longitude" pageId="48" pageNumber="49" precision="15" seconds="46" value="48.41278">048°24ʹ46ʺE</geoCoordinate>
),
<collectingRegion id="B1472CC71874FFEA436B328CABEAEAF6" country="Madagascar" name="Toamasina" pageId="48" pageNumber="49">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B51874FFEA47A732ACAA50EAF6" box="[232,359,931,954]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
, collected
<date id="073DC4E51874FFEA469132ACA9E6EAF6" box="[478,721,931,954]" pageId="48" pageNumber="49" value="2007-12-11" valueMax="2007-12-13" valueMin="2007-12-11">
<collectingDate id="17793D0D1874FFEA469132ACA9E6EAF6" box="[478,721,931,954]" pageId="48" pageNumber="49" value="2007-12-11" valueMax="2007-12-13" valueMin="2007-12-11">11–13 December 2007</collectingDate>
</date>
by
<collectorName id="DE7687F31874FFEA45B332ACA8A8EAF6" box="[764,927,931,954]" pageId="48" pageNumber="49">Brian L. Fisher</collectorName>
, deposited in
<collectionCode id="15927AE01874FFEA437732ACAF58EAF6" box="[1080,1135,931,954]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="48" pageNumber="49" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEA47E432CCAA28ED57" pageId="48" pageNumber="49" type="etymology">
<paragraph id="733CE2251874FFEA47E432CCAA28ED57" blockId="48.[123,1174,867,1725]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA47E432CCAA1DEA96" bold="true" box="[171,298,963,986]" pageId="48" pageNumber="49">Etymology.</emphasis>
The species name honors Mrs.
<taxonomicName id="B48399A61874FFEA453632CCA874EA96" authority="George Meikle" authorityName="George Meikle" box="[633,835,963,986]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ida">Ida George Meikle</taxonomicName>
, mother-in-law of Charles Griswold. The name memorializes her for her encouragement and appreciation of all things natural and arachnological.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEA47E4352BA9B7EDB7" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="733CE2251874FFEA47E4352BA9B7EDB7" blockId="48.[123,1174,867,1725]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA47E4352BAAAFED77" bold="true" box="[171,408,1060,1083]" pageId="48" pageNumber="49">
Diagnosis.
<taxonomicName id="B48399A61874FFEA466A352BAAAFED77" box="[293,408,1060,1083]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ida">
<emphasis id="41F73E371874FFEA466A352BAAAFED77" bold="true" box="[293,408,1060,1083]" italics="true" pageId="48" pageNumber="49">Uduba ida</emphasis>
</taxonomicName>
</emphasis>
are an Unclassified species (
<figureCitation id="EBB8FEA01874FFEA4585352BA82AED77" box="[714,797,1060,1083]" captionStart="MAP" captionStartId="186.[421,440,1168,1187]" captionTargetBox="[388,916,216,1150]" captionTargetId="figure-11@186.[388,916,216,1150]" captionTargetPageId="186" captionText="MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863)." figureDoi="http://doi.org/10.5281/zenodo.13158799" httpUri="https://zenodo.org/record/13158799/files/figure.png" pageId="48" pageNumber="49">Map 21</figureCitation>
) with a divided cribellum. Females of
<taxonomicName id="B48399A61874FFEA47D7354BAA3CED17" box="[152,267,1092,1115]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ida">
<emphasis id="41F73E371874FFEA47D7354BAA3CED17" bold="true" box="[152,267,1092,1115]" italics="true" pageId="48" pageNumber="49">Uduba ida</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="B48399A61874FFEA4586354BA850ED17" box="[713,871,1092,1115]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA4586354BA824ED17" bold="true" box="[713,787,1092,1115]" italics="true" pageId="48" pageNumber="49">Uduba</emphasis>
species
</taxonomicName>
by their unique female genitalia: the epigynal plate (
<figureCitation id="EBB8FEA01874FFEA46C7356BAADBED37" box="[392,492,1124,1147]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="48" pageNumber="49">Fig. 73 C</figureCitation>
) is trapezoidal, narrowed anteriorly, with no distinction between ML and LL and with a strongly marked posterior margin at the epigastric furrow and copulatory openings exposed near the posterior margin of plate; vulva (
<figureCitation id="EBB8FEA01874FFEA445B35ABA84AEDF7" box="[788,893,1188,1211]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="48" pageNumber="49">Fig. 73 F</figureCitation>
) with spermathecal ducts forming five tight transverse loops in a longitudinal stack, duct returns posteriad within these loops, vulva length 1.30 times width. Males unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1874FFEB47E4340AAA84EA79" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="733CE2251874FFEA47E4340AAFA2EC50" blockId="48.[123,1174,867,1725]" box="[171,1173,1285,1308]" pageId="48" pageNumber="49">
<emphasis id="41F73E371874FFEA47E4340AAA4EEC50" bold="true" box="[171,377,1285,1308]" pageId="48" pageNumber="49">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E371874FFEA454A340AA96FEC50" bold="true" box="[517,600,1285,1308]" pageId="48" pageNumber="49">Female</emphasis>
(
<typeStatus id="AC385C871874FFEA4525340AA9E3EC50" box="[618,724,1285,1308]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
): Total length 7.60. Markings typical for
</paragraph>
<paragraph id="733CE2251874FFEB4734342AA837EB18" blockId="48.[123,1174,867,1725]" lastBlockId="49.[122,1174,220,917]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<taxonomicName id="B48399A61874FFEA4734342AABF2EC70" authorityName="Simon" authorityYear="1880" box="[123,197,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371874FFEA4734342AABF2EC70" bold="true" box="[123,197,1317,1340]" italics="true" pageId="48" pageNumber="49">Uduba</emphasis>
</taxonomicName>
. Carapace 3.40 long, 2.45 wide, 1.50 high; clypeus 0.15 high. Eye diameters: AME 0.12, ALE 0.15, PME 0.16, PLE 0.18. Chelicerae 1.80 long; sternum 1.60 long, 1.40 wide; labium 0.80 long; palpal coxae 1.25 long.
<emphasis id="41F73E371874FFEA468A346AA92CEC30" bold="true" box="[453,539,1381,1404]" pageId="48" pageNumber="49">Ratios–</emphasis>
carapace length / width = 1.39, carapace height / width = 0.61, PER / carapace width = 0.47, PER/OAL = 2.85, PER/AER = 1.36, OAL/OQL = 1.25, OQP/ OQA = 1.33, clypeus height / AME = 1.25, cheliceral length / clypeus height = 12.00, sternum length / width = 1.14, palpal coxa length / width = 2.50, femur I length / carapace width = 1.02, metatarsus I length / carapace width = 0.78, femur IV length / carapace width = 1.02, palpal tarsus length / carapace width = 0.41, palpal tibia length / palpal patella length = 1.67, palpal tarsus length / palpal tibia length = 1.43, palpal tarsus length / palpal femur length = 0.83, palpal tibia length / palpal patella length = 1.17. Cribellum divided. Spination: palpus–femur d0-1-2, tibia p2-1, tarsus p2-1; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2-2, metatarsus p0-0-1, v2-2-2; leg II–femur d1-0-0, p0-0-1, tibia p0-0-1, v1-1-1-2, metatarsus p0-0-1, v2-2-2; leg III–femur d1-0-0-1, p0-0-0-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0-0, p1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-0-0, tibia v2-2-2, metatarsus p0-1-2, v2-2-2, r1-1-2. Scopulae: weak ventral beneath tarsi I and II. Leg measurements: I: 2.50 + 1.10 + 1.90 + 1.90 + 1.30 = 8.70; II: 2.10 + 1.10 + 1.60 + 1.60 + 1.20 = 7.60; III: 2.30 + 0.80 + 1.10 + 1.20 + 1.00 = 6.40; IV: 2.50 + 1.00 + 1.90 + 2.30 + 1.30 = 9.00; palpus: 1.20 + 0.60 + 0.70 + NA + 1.00 = 3.50. Leg formula 4123.
<emphasis id="41F73E371875FFEB44C83033AF74E81F" bold="true" box="[903,1091,316,339]" pageId="49" pageNumber="50">Female genitalia</emphasis>
: epigynal plate (
<figureCitation id="EBB8FEA01875FFEB47A43053AA6FE83F" box="[235,344,348,371]" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="49" pageNumber="50">Fig. 73 C</figureCitation>
<emphasis id="41F73E371875FFEB46173053AA56E83F" bold="true" box="[344,353,348,371]" pageId="49" pageNumber="50">)</emphasis>
without depressed atrium, trapezoidal, narrowed anteriorly, no distinction between ML and LL, epigynal plate width 1.11 times length, with strongly marked posterior margin at epigastric furrow and copulatory openings exposed near posterior margin of plate; distance between copulatory openings 0.46 times epigynum width, CO origin at 0.15 of epigynum length, extant of plate laterad to copulatory openings equals 0.125 times plate width. Vulva unique (
<figureCitation id="EBB8FEA01875FFEB432530D2AB9BEB58" captionStart="FIGURE" captionStartId="161.[155,167,918,937]" captionTargetBox="[124,1164,219,893]" captionTargetId="figure-16@161.[120,1176,216,905]" captionTargetPageId="161" captionText="FIGURE 73. Uduba spp., female genitalia drawings. A, B. U. ibonia, new species, paratype (CASENT9006035), from Parc National Ranomafana. C, F. U. ida, new species, holotype (CASENT9042534) from Station Forestière Analamazaotra. D, E. U. schlingeri, new species (CASENT9006057) from Parc National Ranomafana. B–D. Epigynum, ventral. A, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158737" httpUri="https://zenodo.org/record/13158737/files/figure.png" pageId="49" pageNumber="50">Fig. 73 F</figureCitation>
) with spermathecal ducts forming five tight transverse loops in longitudinal stack, duct returns posteriad within these loops, vulva length 1.30 times width, fertilization ducts widely spaced, vulva width 3.64 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE2251875FFEB47E43352ABFCEBD8" blockId="49.[122,1174,220,917]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E43352AA2DEB38" bold="true" box="[171,282,605,628]" pageId="49" pageNumber="50">Variation.</emphasis>
Unknown: no males have been examined and only the unique
<typeStatus id="AC385C871875FFEB44823352AF1DEB38" box="[973,1066,605,628]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
female is known.
</paragraph>
<paragraph id="733CE2251875FFEB47E43392A83CEBB9" blockId="49.[122,1174,220,917]" pageId="49" pageNumber="50">
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M
<emphasis id="41F73E371875FFEB478C3392AABEEBF8" bold="true" box="[195,393,669,692]" pageId="49" pageNumber="50">aterial examined.</emphasis>
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:
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<collectingRegion id="B1472CC71875FFEB45123392A802EBF8" box="[605,821,669,692]" country="Madagascar" name="Toamasina" pageId="49" pageNumber="50">Toamasina Province</collectingRegion>
</emphasis>
:
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,
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,
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, elev.
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, sifted litter in montane rainforest,
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,
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(
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,
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,
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,
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.
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<paragraph id="733CE2251875FFEB47E433F1AA84EA79" blockId="49.[122,1174,220,917]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E433F1AA54EA59" bold="true" box="[171,355,766,789]" pageId="49" pageNumber="50">Natural history.</emphasis>
The unique specimen of the cribellate species
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<emphasis id="41F73E371875FFEB443133F1A8C1EA59" bold="true" box="[894,1014,766,789]" italics="true" pageId="49" pageNumber="50">Uduba ida</emphasis>
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was found in montane rainforest leaf litter.
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</subSubSection>
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<emphasis id="41F73E371875FFEB47E43231AA8BEA19" bold="true" box="[171,444,830,853]" pageId="49" pageNumber="50">
Distribution.
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<emphasis id="41F73E371875FFEB46093231AA8BEA19" bold="true" box="[326,444,830,853]" italics="true" pageId="49" pageNumber="50">Uduba ida</emphasis>
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are known only from the
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locality in east central
<collectingCountry id="0B94A2B51875FFEB435A3231AFA1EA19" box="[1045,1174,830,853]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA01875FFEB47CD3251ABE2EA39" box="[130,213,862,885]" captionStart="MAP" captionStartId="178.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1068]" captionTargetId="figure-12@178.[648,1176,216,1068]" captionTargetPageId="178" captionText="MAP 5. Distributions of Uduba schlingeri, Uduba MAP 6. Distributions of Uduba andriamihajai and volana and Uduba ida. Uduba goodmani." figureDoi="http://doi.org/10.5281/zenodo.13158779" httpUri="https://zenodo.org/record/13158779/files/figure.png" pageId="49" pageNumber="50">Maps 5</figureCitation>
,
<figureCitation id="EBB8FEA01875FFEB47AE3251ABCBEA39" box="[225,252,862,885]" captionStart="MAP" captionStartId="186.[421,440,1168,1187]" captionTargetBox="[388,916,216,1150]" captionTargetId="figure-11@186.[388,916,216,1150]" captionTargetPageId="186" captionText="MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863)." figureDoi="http://doi.org/10.5281/zenodo.13158799" httpUri="https://zenodo.org/record/13158799/files/figure.png" pageId="49" pageNumber="50">21</figureCitation>
<emphasis id="41F73E371875FFEB47B43251AA33EA39" bold="true" box="[251,260,862,885]" pageId="49" pageNumber="50">)</emphasis>
, near the town of Périnet, which is an area renowned in
<collectingCountry id="0B94A2B51875FFEB442E3251A8D5EA39" box="[865,994,862,885]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
for species richness and endemism.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8A9D556F2618C82A7DEF2BDE92B9BD7F">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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, 18.
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<emphasis id="41F73E371875FFEB47E4350CAA79ED56" bold="true" box="[171,334,1027,1050]" pageId="49" pageNumber="50">Type material.</emphasis>
<materialsCitation id="C3EBE8781875FFEB461C350CABD9ED36" collectingDate="2009-01-12" collectingDateMax="2009-01-22" collectingDateMin="2009-01-12" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="1700" location="Ankokoy Forest" pageId="49" pageNumber="50" specimenCode="CASENT9062802" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
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<collectingMethod id="AAC29A321875FFEB459D350CA8C5ED56" box="[722,1010,1027,1050]" pageId="49" pageNumber="50">collected from malaise trap</collectingMethod>
in
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elevation in the
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(20°04.05ʹS, 46°59.97ʹE),
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<quantity id="B47B4FC01875FFEB44DE352CA8F0ED76" box="[913,967,1059,1082]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="km" value="3.0">3 km</quantity>
E of Ibity
</locationDeviation>
,
<collectingRegion id="B1472CC71875FFEB4370352CAA10ED16" country="Madagascar" name="Antananarivo" pageId="49" pageNumber="50">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51875FFEB467F354CAA98ED16" box="[304,431,1091,1114]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
,
<date id="073DC4E51875FFEB46F4354CA9A2ED16" box="[443,661,1091,1114]" pageId="49" pageNumber="50" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">
<collectingDate id="17793D0D1875FFEB46F4354CA9A2ED16" box="[443,661,1091,1114]" pageId="49" pageNumber="50" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">12–22 January 2009</collectingDate>
</date>
, collectors
<collectorName id="DE7687F31875FFEB4443354CA85DED16" box="[780,874,1091,1114]" pageId="49" pageNumber="50">M. Irwin</collectorName>
and
<collectorName id="DE7687F31875FFEB44D2354CAF01ED16" box="[925,1078,1091,1114]" pageId="49" pageNumber="50">R. Harin’Hala</collectorName>
, deposit- ed in
<collectionCode id="15927AE01875FFEB47F8356CABD9ED36" box="[183,238,1123,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="49" pageNumber="50" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781875FFEB47B6356CAAF4ED97" collectingDate="2001-11-16" collectingDateMax="2001-11-20" collectingDateMin="2001-11-16" collectionCode="CAS" country="Madagascar" elevation="150" latitude="-19.709446" location="Parc National Tsingy de Bemaraha" longLatPrecision="21" longitude="44.718056" municipality="Fisher-Griswold Arthropod Team" pageId="49" pageNumber="50" specimenCode="CASENT9009383" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="AC385C871875FFEB47B6356CAA61ED36" box="[249,342,1123,1146]" pageId="49" pageNumber="50" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1875FFEB46FB356CA94AED36" box="[436,637,1123,1146]" collectionCode="CASENT" pageId="49" pageNumber="50">CASENT9009383</specimenCode>
) collected at
<quantity id="B47B4FC01875FFEB4442356CA87DED36" box="[781,842,1123,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="50" unit="m" value="150.0">
<elevation id="F8AE05161875FFEB4442356CA87DED36" box="[781,842,1123,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="50" unit="m" value="150.0">150m</elevation>
</quantity>
elevation in tropical dry forest on tsingy from
<location id="765CB4FE1875FFEB4661358BA980EDD7" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1875FFEB4661358BA980EDD7" box="[302,695,1156,1179]" country="Madagascar" latitude="-19.709446" longLatPrecision="21" longitude="44.718056" municipality="Fisher-Griswold Arthropod Team" name="Parc National Tsingy de Bemaraha" pageId="49" pageNumber="50" stateProvince="Antananarivo">Parc National Tsingy de Bemaraha</location>
,
<locationDeviation id="628EFB071875FFEB4589358BAF36EDD7" box="[710,1025,1156,1179]" location="Antsalova" pageId="49" pageNumber="50">
<quantity id="B47B4FC01875FFEB4589358BA829EDD7" box="[710,798,1156,1179]" metricMagnitude="4" metricUnit="m" metricValue="1.06" pageId="49" pageNumber="50" unit="km" value="10.6">10.6 km</quantity>
123°ESE Antsalova
</locationDeviation>
(
<geoCoordinate id="16B784E21875FFEB435A358BAFA7EDD7" box="[1045,1168,1156,1179]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="49" pageNumber="50" precision="15" seconds="34" value="-19.709446">19°42ʹ34ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21875FFEB473435ABABCCEDF7" box="[123,251,1188,1211]" degrees="44" direction="east" minutes="43" orientation="longitude" pageId="49" pageNumber="50" precision="15" seconds="05" value="44.718056">44°43ʹ05ʺE</geoCoordinate>
), collected
<date id="073DC4E51875FFEB46DF35ABA992EDF7" box="[400,677,1188,1211]" pageId="49" pageNumber="50" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">
<collectingDate id="17793D0D1875FFEB46DF35ABA992EDF7" box="[400,677,1188,1211]" pageId="49" pageNumber="50" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">16–20 November 2001</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F1875FFEB445B35ABAFA2EDF7" box="[788,1173,1188,1211]" pageId="49" pageNumber="50">Fisher-Griswold Arthropod Team</collectingMunicipality>
[BLF4503], deposited in
<collectionCode id="15927AE01875FFEB46C335CBAAF4ED97" box="[396,451,1220,1243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="49" pageNumber="50" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFEB47E435EBAF43ECF0" pageId="49" pageNumber="50" type="discussion">
<paragraph id="733CE2251875FFEB47E435EBAF43ECF0" blockId="49.[123,1174,1027,1725]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E435EBAA24EDB7" bold="true" box="[171,275,1252,1275]" pageId="49" pageNumber="50">Remarks</emphasis>
. Males of the similar species
<taxonomicName id="B48399A61875FFEB451B35EBA9D4EDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[596,739,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB451B35EBA9D4EDB7" bold="true" box="[596,739,1252,1275]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61875FFEB445735EBA89DEDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[792,938,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB445735EBA89DEDB7" bold="true" box="[792,938,1252,1275]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
may be distinguished by the TA2 form. Association of the males and females of
<taxonomicName id="B48399A61875FFEB45B4340BA8BDEC57" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[763,906,1284,1307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB45B4340BA8BDEC57" bold="true" box="[763,906,1284,1307]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
is unproblematic in that several were collected together at Parc National de Namoroka. Females associated with
<taxonomicName id="B48399A61875FFEB4304342BAB8FEC17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB4304342BAB8FEC17" bold="true" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
have the epigynal plate posterior of the ML smooth or grooved and the CO hidden beneath the ML hood (
<figureCitation id="EBB8FEA01875FFEB4659346AAABBEC30" box="[278,396,1381,1404]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01875FFEB46D7346AAAFAEC30" box="[408,461,1381,1404]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. A–C. Epigynum, ventral. D–F. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="49" pageNumber="50">81 B</figureCitation>
) whereas a female from Tsingy de Bemeraha (CASENT9009383) differs from these in that it has exposed copulatory openings just behind the ML (
<figureCitation id="EBB8FEA01875FFEB44A6348AAF64ECD0" box="[1001,1107,1413,1436]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 C</figureCitation>
). This female is considered a
<typeStatus id="AC385C871875FFEB463B34AAAAE7ECF0" box="[372,464,1445,1468]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of
<taxonomicName id="B48399A61875FFEB46B934AAA9BEECF0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[502,649,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB46B934AAA9BEECF0" bold="true" box="[502,649,1445,1468]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
, which is otherwise known only from males.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFEB47E434CAA92CEF50" pageId="49" pageNumber="50" type="etymology">
<paragraph id="733CE2251875FFEB47E434CAA92CEF50" blockId="49.[123,1174,1027,1725]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E434CAAA1DEC90" bold="true" box="[171,298,1477,1500]" pageId="49" pageNumber="50">Etymology.</emphasis>
The species epithet is a patronym on honor of Dr. Mike Irwin, entomologist and philanthropist and a principal architect of the malaise trap survey of
<collectingCountry id="0B94A2B51875FFEB440134EAA8F8ECB0" box="[846,975,1509,1532]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
arthropods, which revealed many new species of spiders.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFE847E4372AA9B5E81F" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="733CE2251875FFE847E4372AA9B5E81F" blockId="49.[123,1174,1027,1725]" lastBlockId="50.[122,1175,220,1725]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E4372AAA8BEF70" bold="true" box="[171,444,1573,1596]" pageId="49" pageNumber="50">
Diagnosis.
<taxonomicName id="B48399A61875FFEB4667372AAA8BEF70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,444,1573,1596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB4667372AAA8BEF70" bold="true" box="[296,444,1573,1596]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
</emphasis>
are members of Group IV, the
<taxonomicName id="B48399A61875FFEB445C372AA8ADEF70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[787,922,1573,1596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">Uduba rinha</taxonomicName>
group (Map 18). They have a divided cribellum. Males can be distinguished from those of other
<taxonomicName id="B48399A61875FFEB44E63749A8C4EF11" authorityName="Simon" authorityYear="1880" box="[937,1011,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371875FFEB44E63749A8C4EF11" bold="true" box="[937,1011,1606,1629]" italics="true" pageId="49" pageNumber="50">Uduba</emphasis>
</taxonomicName>
except
<taxonomicName id="B48399A61875FFEB43043749AB8FEF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB43043749AB8FEF31" bold="true" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
by the simple, trapezoidal MA (
<figureCitation id="EBB8FEA01875FFEB456B3769A984EF31" box="[548,691,1638,1661]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="49" pageNumber="50">Figs 46 A–C</figureCitation>
<emphasis id="41F73E371875FFEB45FC3769A98BEF31" bold="true" box="[691,700,1638,1661]" pageId="49" pageNumber="50">)</emphasis>
;
<taxonomicName id="B48399A61875FFEB45823769A854EF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[717,867,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB45823769A854EF31" bold="true" box="[717,867,1638,1661]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A61875FFEB47343789AA3EEFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,265,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB47343789AA3EEFD1" bold="true" box="[123,265,1670,1693]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
by having the TA2 an erect, rectangular spike (
<figureCitation id="EBB8FEA01875FFEB444B3789A85DEFD1" box="[772,874,1670,1693]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="49" pageNumber="50">Fig. 46 B</figureCitation>
) (
<taxonomicName id="B48399A61875FFEB44CD3789AF26EFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[898,1041,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB44CD3789AF26EFD1" bold="true" box="[898,1041,1670,1693]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
has the TA2 a flexible hook or curlicue,
<figureCitation id="EBB8FEA01875FFEB46E937A9A939EFF1" box="[422,526,1702,1725]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="49" pageNumber="50">Fig. 59 B</figureCitation>
). The females (
<figureCitation id="EBB8FEA01875FFEB45FA37A9A82BEFF1" box="[693,796,1702,1725]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 C</figureCitation>
<emphasis id="41F73E371875FFEB445437A9A813EFF1" bold="true" box="[795,804,1702,1725]" pageId="49" pageNumber="50">)</emphasis>
and
<taxonomicName id="B48399A61875FFEB441537A9A8DFEFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[858,1000,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB441537A9A8DFEFF1" bold="true" box="[858,1000,1702,1725]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01875FFEB44B737A9AF57EFF1" box="[1016,1120,1702,1725]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 G</figureCitation>
<emphasis id="41F73E371875FFEB432F37A9AF5EEFF1" bold="true" box="[1120,1129,1702,1725]" pageId="49" pageNumber="50">)</emphasis>
can be distinguished from those of other cribellate
<taxonomicName id="B48399A61876FFE8453C31D3A98AE9BF" authorityName="Simon" authorityYear="1880" box="[627,701,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE8453C31D3A98AE9BF" bold="true" box="[627,701,220,243]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
by having the epigynum lateral lobes fused medially; the lone female of
<taxonomicName id="B48399A61876FFE846FA31F3A97FE85F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[437,584,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371876FFE846FA31F3A97FE85F" bold="true" box="[437,584,252,275]" italics="true" pageId="50" pageNumber="51">Uduba irwini</emphasis>
</taxonomicName>
has the copulatory openings exposed just posteriad of the ML (
<figureCitation id="EBB8FEA01876FFE8479A3013AA0EE87F" box="[213,313,284,307]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Fig. 71 C</figureCitation>
<emphasis id="41F73E371876FFE846763013AA75E87F" bold="true" box="[313,322,284,307]" pageId="50" pageNumber="51">)</emphasis>
whereas the females of
<taxonomicName id="B48399A61876FFE8450F3013A9FAE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[576,717,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371876FFE8450F3013A9FAE87F" bold="true" box="[576,717,284,307]" italics="true" pageId="50" pageNumber="51">Uduba rinha</emphasis>
</taxonomicName>
have the copulatory openings that are each hidden beneath the sides of the ML (
<figureCitation id="EBB8FEA01876FFE845463033A941E81F" box="[521,630,316,339]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Fig. 71 G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1876FFE947E43053A933EF70" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="733CE2251876FFE947E43053AAEAE898" blockId="50.[122,1175,220,1725]" lastBlockId="51.[123,1175,220,1660]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="41F73E371876FFE847E43053AA40E83F" bold="true" box="[171,375,348,371]" pageId="50" pageNumber="51">Description. Male</emphasis>
(
<typeStatus id="AC385C871876FFE846C93053AAC7E83F" box="[390,496,348,371]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
): Total length 9.70. Markings typical of
<taxonomicName id="B48399A61876FFE844EC3053A8DAE83F" authorityName="Simon" authorityYear="1880" box="[931,1005,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE844EC3053A8DAE83F" bold="true" box="[931,1005,348,371]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
. Carapace 5.50 long, 3.60 wide, 2.10 high; clypeus 0.30 high. Eye diameters: AME 0.14, ALE 0.16, PME 0.18, PLE 0.20. Chelicerae 2.30 long; sternum 2.40 long, 1.80 wide; labium 1.10 long; palpal coxae 1.60 long. Cribellum divided. Spination. (ANJA reserve, CASENT9065405) palpus–femur d1-0-3; leg I–femur d1-1-1, p0-1-2, r0-1-1-0, tibia d0-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p0- 1-2, v2-2-1, r1-1-1; leg II–femur d1-1-1, p0-1-2, r0-1-0-0, tibia d0-1-1-0, p0-1-1-0, v2-2-2, r0-1-1- 0, metatarsus p1-1-2, v2-2-1, r1-1-2; leg III–femur d1-1-1, p0-1-0-2, r0-1-0-2, tibia d0-1-0, p0-1- 1-0, v2-1-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-1, p0-1- 1-0, r0-0-0-1, patella r1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2- 1-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements (
<typeStatus id="AC385C871876FFE843683371AFBCEBD9" box="[1063,1163,638,661]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
): I: 4.40 + 1.90 + 4.00 + 4.10 + 2.60 = 17.00; II: 3.90 + 1.80 + 3.20 + 3.40 + 2.20 = 14.50; III: 3.40 + 1.50 + 2.10 + 2.70 + 1.70 = 11.40; IV: 4.20 + 1.70 + 3.80 + 4.20 + 2.40 = 16.30; palpus: 2.00 + 0.70 + 0.80 + NA + 2.50 = 6.00. Leg formula 1423.
<emphasis id="41F73E371876FFE8440733D0A8FFEBBA" bold="true" box="[840,968,735,758]" pageId="50" pageNumber="51">Male palp</emphasis>
(ANJA Reserve, CASENT9065405) (
<figureCitation id="EBB8FEA01876FFE8461633F0AAC7EA5A" box="[345,496,767,790]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 A–C</figureCitation>
): palpal tibia 0.35 cymbial length, RTA long, straight (
<figureCitation id="EBB8FEA01876FFE8430933F0ABB8EA7B" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 C</figureCitation>
), RTA length 0.33 times tibia length, length 2.25 times width, with sharply-pointed apex, VTA short, length 1.50 times width, conical, blunt (
<figureCitation id="EBB8FEA01876FFE8453D324FA830EA1B" box="[626,775,832,855]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 A, C</figureCitation>
), length 0.52 times tibia width; tibia without stout spines; tegulum convex, without ridge, tegulum length 1.09 times width; TA1 slen- der, extends far distad of tegulum apex, TA1 extends past TA3 apex by 1.38 times tegulum length; TA3 a transverse, blunt hook (
<figureCitation id="EBB8FEA01876FFE846F632AFA929EAFB" box="[441,542,928,951]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 B</figureCitation>
<emphasis id="41F73E371876FFE8455232AFA911EAFB" bold="true" box="[541,550,928,951]" pageId="50" pageNumber="51">)</emphasis>
, TA3 extends apicad of TA2 by 1.21 times tegulum length, TA2 a small, erect trapezoid with concave apex, stiff, sclerotized, extends partially in front of TA3 (
<figureCitation id="EBB8FEA01876FFE847CC32EEABD0EAB4" box="[131,231,993,1016]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 B</figureCitation>
<emphasis id="41F73E371876FFE847A832EEABC7EAB4" bold="true" box="[231,240,993,1016]" pageId="50" pageNumber="51">)</emphasis>
; MA a broad, flat trapezoid, origin at 0.54 times tegulum length, extends nearly to tegulum apex, tegulum apex at 0.085 tegulum length from MA apex, MA length (including apical processes) 1.22 times width, MA large (but smaller than in
<taxonomicName id="B48399A61876FFE845B7352EA861ED74" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[760,854,1057,1080]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371876FFE845B7352EA861ED74" bold="true" box="[760,854,1057,1080]" italics="true" pageId="50" pageNumber="51">U. rinha</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01876FFE8442B352EA8FDED74" box="[868,970,1057,1080]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="50" pageNumber="51">Fig. 59 A</figureCitation>
), length 0.31 tegulum length, width 0.29 tegulum width (
<figureCitation id="EBB8FEA01876FFE84568354DA98DED15" box="[551,698,1090,1113]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 A–C</figureCitation>
); conductor fan elongate, entire (
<figureCitation id="EBB8FEA01876FFE84350354DAFB1ED15" box="[1055,1158,1090,1113]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 C</figureCitation>
<emphasis id="41F73E371876FFE843C9354DAFB8ED15" bold="true" box="[1158,1167,1090,1113]" pageId="50" pageNumber="51">)</emphasis>
.
<emphasis id="41F73E371876FFE84734356DABF9ED35" bold="true" box="[123,206,1122,1145]" pageId="50" pageNumber="51">Female</emphasis>
(
<typeStatus id="AC385C871876FFE84795356DAA09ED35" box="[218,318,1122,1145]" pageId="50" pageNumber="51" type="paratype">Paratype</typeStatus>
): Total length 11.56. Markings typical for
<taxonomicName id="B48399A61876FFE845B9356DA877ED35" authorityName="Simon" authorityYear="1880" box="[758,832,1122,1145]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE845B9356DA877ED35" bold="true" box="[758,832,1122,1145]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
. Carapace 5.60 long, 3.80 wide, 2.30 high; clypeus 0.30 high. Eye diameters: AME 0.16, ALE and PME 0.22, PLE 0.24. Chelicerae 2.70 long; sternum 2.40 long, 1.90 wide; labium 1.30 long; palpal coxae 1.90 long.
<emphasis id="41F73E371876FFE8434535ADAF57EDF5" bold="true" box="[1034,1120,1186,1209]" pageId="50" pageNumber="51">Ratios–</emphasis>
carapace length / width = 1.47, carapace height / width = 0.61, PER / carapace width = 0.42, PER / OAL = 3.20, PER / AER = 1.36, OAL / OQL = 1.09, OQP / OQA = 1.38, clypeus height / AME diameter width = 1.88, cheliceral length / clypeus height = 9.00, sternum length / width = 1.26, palpal coxa length / width = 2.38, femur I length / carapace width = 1.00, metatarsus I length / carapace width = 0.79, femur IV length / carapace width = 1.03, palpal tarsus length / carapace width = 0.45, palpal tarsus length / palpal femur length = 0.89. Cribellum divided. Spination: Palpus–femur d0-1-2, patella p1, tibia p2-0, tarsus p2-0, v0-0-2; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2, metatarsus v2-2-3; leg II–femur d1-0-0, p0-0-1, tibia p0-0-1, v1-1-2, metatarsus v2-2-3; leg III–femur p0-1-0-0, r0-1-0-0, tibia d1-0-1-0, p0-1-1-0, v1-0-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IV–femur d1-1-0-0-0, tibia v2-1-2, r0-1-1-0, metatarsus p0-1-2, v1-2-2, r0-2-2. Scopulae: strong ventral on all tarsi, most of metatarsi I and II, and apices of tarsi III and IV. Leg measurements: I: 3.80 + 1.90 + 3.30 + 3.00 + 1.90 = 13.90; II: 3.20 + 1.60 + 2.60 + 2.30 + 1.80 = 11.50; III: 2.60 + 1.50 + 1.30 + 2.10 + 1.40 = 8.90; IV: 3.90 + 1.80 + 2.90 + 3.40 + 1.70 = 13.70; palpus: 1.90 + 1.00 + 1.00 + NA + 1.70 = 5.69. Leg formula 1423.
<emphasis id="41F73E371876FFE847F23789AA4EEFD1" bold="true" box="[189,377,1670,1693]" pageId="50" pageNumber="51">Female genitalia</emphasis>
(
<figureCitation id="EBB8FEA01876FFE846C83789A92CEFD1" box="[391,539,1670,1693]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Figs. 71 C, D</figureCitation>
): epigynal plate without depressed atrium, plate width 1.35 times length; median lobe and lateral lobes differentiated only in anterior half of plate, distance from posterior margin of ML to epigastric furrow 0.35 times epigynum length; median lobe a flat plate with lateral margins extending around to posterior, broad, median lobe width 0.63 times epigynum width, 0.45 times ML length, ML width 2.64 times LL width at sides of ML (
<figureCitation id="EBB8FEA01877FFE943553013AFBFE87F" box="[1050,1160,284,307]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="51" pageNumber="52">Fig. 71 C</figureCitation>
); with copulatory openings exposed on epigynal plate beneath a transverse, crescentic groove on each side just behind ML, distance between copulatory openings 0.26 times epigynum width, CO origin at 0.35 of epigynum length. Vulva (
<figureCitation id="EBB8FEA01877FFE945003072A98FE8D8" box="[591,696,381,404]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="51" pageNumber="52">Fig. 71 D</figureCitation>
) with spermathecal ducts forming three longitudinal loops, vulva length 0.64 times width, fertilization ducts widely separated, vulva width 3.57 times distance between FD.
</paragraph>
<paragraph id="733CE2251877FFE947E430D2A9B4EA5A" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E430D2AA2DE8B8" bold="true" box="[171,282,477,500]" pageId="51" pageNumber="52">Variation.</emphasis>
Male (N= 7): Total length 9.70–11.50; carapace length / width = 1.33–1.53, carapace height / width = 0.40–0.59, PER / carapace width = 0.34–0.38, PER / OAL = 2.50–3.24, PER / AER = 1.37–1.42, OAL / OQL = 1.05–1.26, OQP / OQA = 1.18–1.25, clypeus height / AME 1.11– 2.14, cheliceral length / clypeus height = 7.67–11.00, sternum length / width = 1.25–1.39, palpal coxa length / width = 2.25–2.67, femur I length / carapace width =1.12–1.24, metatarsus I length / carapace width = 1.09–1.28, femur IV length / carapace width = 1.15–1.26, palpal tarsus length / carapace width = 0.64–0.74, palpal tarsus length / palpal patella length = 3.33–3.75, palpal tarsus length / palpal tibia length = 3.00–3.38, palpal tarsus length / palpal femur length = 1.17–1.33, palpal tibia length / palpal patella length = 1.00–1.25. Female variation is unknown: only
<specimenCount id="658529AC1877FFE9435633D0AFA6EBBA" box="[1049,1169,735,758]" count="1" pageId="51" pageNumber="52" type="female">one female</specimenCount>
, the
<typeStatus id="AC385C871877FFE947EC33F0ABC8EA5A" box="[163,255,767,790]" pageId="51" pageNumber="52" type="paratype">paratype</typeStatus>
(CASENT9009383), is recognized.
</paragraph>
<paragraph id="733CE2251877FFE947E4322FABEFEC37" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
M
<emphasis id="41F73E371877FFE9478D322FAAA5EA7B" bold="true" box="[194,402,800,823]" pageId="51" pageNumber="52">aterial examined.</emphasis>
<materialsCitation id="C3EBE8781877FFE946ED322FABDBEADB" collectingDate="2009-01-12" collectingDateMax="2009-01-22" collectingDateMin="2009-01-12" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="1700" location="Ankokoy Forest" pageId="51" pageNumber="52" specimenCode="CASENT9062802" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="0B94A2B51877FFE946ED322FA953EA7B" box="[418,612,800,823]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371877FFE94538322FA84EEA7B" box="[631,889,800,823]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE94538322FA84EEA7B" box="[631,889,800,823]" country="Madagascar" name="Antananarivo" pageId="51" pageNumber="52">Antananarivo Province</collectingRegion>
</emphasis>
:
<collectingRegion id="B1472CC71877FFE944C1322FAFA5EA7B" box="[910,1170,800,823]" country="Madagascar" name="Antananarivo" pageId="51" pageNumber="52">Antananarivo Province</collectingRegion>
,
<location id="765CB4FE1877FFE94734324FAA1BEA1B" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE94734324FAA1BEA1B" box="[123,300,832,855]" country="Madagascar" name="Ankokoy Forest" pageId="51" pageNumber="52" stateProvince="Antananarivo">Ankokoy Forest</location>
,
<locationDeviation id="628EFB071877FFE94676324FAAD7EA1B" box="[313,480,832,855]" location="Ibity" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE94676324FAA47EA1B" box="[313,368,832,855]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="51" pageNumber="52" unit="km" value="3.0">3 km</quantity>
E of Ibity
</locationDeviation>
, 20°04.05ʹS, 46°59.97ʹE, elev.
<quantity id="B47B4FC01877FFE9447B324FA8B4EA1B" box="[820,899,832,855]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="51" pageNumber="52" unit="m" value="1700.0">
<elevation id="F8AE05161877FFE9447B324FA8B4EA1B" box="[820,899,832,855]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="51" pageNumber="52" unit="m" value="1700.0">1700m</elevation>
</quantity>
, malaise, in
<taxonomicName id="B48399A61877FFE9435F324FAF53EA1B" box="[1040,1124,832,855]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="51" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E371877FFE9435F324FAF53EA1B" box="[1040,1124,832,855]" italics="true" pageId="51" pageNumber="52">Uapaca</emphasis>
</taxonomicName>
forest,
<date id="073DC4E51877FFE947E8326FAAB2EA3B" box="[167,389,864,887]" pageId="51" pageNumber="52" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">
<collectingDate id="17793D0D1877FFE947E8326FAAB2EA3B" box="[167,389,864,887]" pageId="51" pageNumber="52" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">12–22 January 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE946DE326FAAC5EA3B" box="[401,498,864,887]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and
<collectorName id="DE7687F31877FFE94567326FA9F6EA3B" box="[552,705,864,887]" pageId="51" pageNumber="52">R. Harin’Hala</collectorName>
[MG-56-08] (
<typeStatus id="AC385C871877FFE94413326FA889EA3B" box="[860,958,864,887]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E1877FFE94485326FAFA5EA3B" box="[970,1170,864,887]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9062802</specimenCode>
,
<specimenCount id="658529AC1877FFE94734328FABABEADB" box="[123,156,896,919]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE947E3328FABD3EADB" box="[172,228,896,919]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781877FFE947B5328FABE3ED54" collectingDate="1997-12-07" collectingDateMax="1997-12-12" collectingDateMin="1997-12-07" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1450" location="Reserve Speciale d'Ambohitantely" pageId="51" pageNumber="52" specimenCode="CASENT9016347, CASENT9016348, CASENT9006222, CASENT9016346, CASENT9016349" specimenCount="7" specimenCount-male="7" stateProvince="Antananarivo" typeStatus="holotype">
<location id="765CB4FE1877FFE947B5328FA947EADB" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE947B5328FA947EADB" box="[250,624,896,919]" country="Madagascar" name="Reserve Speciale d'Ambohitantely" pageId="51" pageNumber="52" stateProvince="Antananarivo">Réserve Spéciale d’Ambohitantely</location>
,
<locationDeviation id="628EFB071877FFE94530328FA851EADB" box="[639,870,896,919]" location="Ankazobe" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE94530328FA9F4EADB" box="[639,707,896,919]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="52" unit="km" value="24.0">24 km</quantity>
NE Ankazobe
</locationDeviation>
, 18°10.1ʹS, 47°16.6ʹE, elev.
<quantity id="B47B4FC01877FFE9473432AEABF0EAF4" box="[123,199,929,952]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">
<elevation id="F8AE05161877FFE9473432AEABF0EAF4" box="[123,199,929,952]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450m</elevation>
</quantity>
, disturbed transitional montane mossy forest, 30-hectare parcel,
<collectingMethod id="AAC29A321877FFE944DD32AEAF27EAF4" box="[914,1040,929,952]" pageId="51" pageNumber="52">pitfall traps</collectingMethod>
#1–3,
<date id="073DC4E51877FFE9432F32AEAA10EA94" pageId="51" pageNumber="52" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">
<collectingDate id="17793D0D1877FFE9432F32AEAA10EA94" pageId="51" pageNumber="52" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">7–12 December 1997</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE9467D32CEAA8FEA94" box="[306,440,961,984]" pageId="51" pageNumber="52">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1877FFE9468832CEA9A7EA94" box="[455,656,961,984]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016347</specimenCode>
,
<specimenCount id="658529AC1877FFE945D432CEA98BEA94" box="[667,700,961,984]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9458532CEA82CEA94" box="[714,795,961,984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE9447A32CEA8C9EA94" box="[821,1022,961,984]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016348</specimenCode>
,
<specimenCount id="658529AC1877FFE9434532CEAF1CEA94" box="[1034,1067,961,984]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9437632CEAFBFEA94" box="[1081,1160,961,984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE947CD32EEAA7BEAB4" box="[130,332,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9006222</specimenCode>
,
<specimenCount id="658529AC1877FFE9461032EEAAB7EAB4" box="[351,384,993,1016]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE946D932EEAAD0EAB4" box="[406,487,993,1016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE9454632EEA9E4EAB4" box="[521,723,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016346</specimenCode>
,
<specimenCount id="658529AC1877FFE945A932EEA830EAB4" box="[742,775,993,1016]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9445232EEA859EAB4" box="[797,878,993,1016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE944DE32EEAF6DEAB4" box="[913,1114,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016349</specimenCode>
,
<specimenCount id="658529AC1877FFE9432232EEAFB8EAB4" box="[1133,1167,993,1016]" count="3" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
,
<collectionCode id="15927AE01877FFE94734350EABFAED54" box="[123,205,1025,1048]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781877FFE947AE350EAF58ED15" collectingDate="2014-12-10" collectingDateMax="2014-12-19" collectingDateMin="2014-12-10" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" county="Haute Matsiatra Region" elevation="1097" location="Ambalavao" municipality="Reserve" pageId="51" pageNumber="52" specimenCode="CASENT9065491" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="41F73E371877FFE947AE350EAAEDED54" box="[225,474,1025,1048]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE947AE350EAAEDED54" box="[225,474,1025,1048]" country="Madagascar" name="Fianarantsoa" pageId="51" pageNumber="52">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectingCounty id="9A5D9AA91877FFE946A5350EA9C4ED54" box="[490,755,1025,1048]" pageId="51" pageNumber="52">Haute Matsiatra Region</collectingCounty>
, ANJA
<collectingMunicipality id="9358785F1877FFE94406350EA895ED54" box="[841,930,1025,1048]" pageId="51" pageNumber="52">Reserve</collectingMunicipality>
,
<quantity id="B47B4FC01877FFE944E0350EA8C3ED54" box="[943,1012,1025,1048]" metricMagnitude="4" metricUnit="m" metricValue="6.1" pageId="51" pageNumber="52" unit="km" value="61.0">61 km</quantity>
S
<location id="765CB4FE1877FFE9435B350EAFA5ED54" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE9435B350EAFA5ED54" box="[1044,1170,1025,1048]" country="Madagascar" county="Haute Matsiatra Region" municipality="Reserve" name="Ambalavao" pageId="51" pageNumber="52" stateProvince="Fianarantsoa">Ambalavao</location>
,
<locationDeviation id="628EFB071877FFE94734352EAABFED74" box="[123,392,1057,1080]" location="Fianarantsoa" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE94734352EAB88ED74" box="[123,191,1057,1080]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="51" pageNumber="52" unit="km" value="13.0">13 km</quantity>
S of
<collectingRegion id="B1472CC71877FFE947B2352EAABFED74" box="[253,392,1057,1080]" country="Madagascar" name="Fianarantsoa" pageId="51" pageNumber="52">Fianarantsoa</collectingRegion>
</locationDeviation>
, 22°43.23ʹS, 46°15.47ʹE, elev.
<quantity id="B47B4FC01877FFE94597352EA812ED74" box="[728,805,1057,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="51" pageNumber="52" unit="m" value="1097.0">
<elevation id="F8AE05161877FFE94597352EA812ED74" box="[728,805,1057,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="51" pageNumber="52" unit="m" value="1097.0">1097m</elevation>
</quantity>
, malaise, isleberg vegetation,
<date id="073DC4E51877FFE94322352EAA7EED15" pageId="51" pageNumber="52" value="2014-12-10" valueMax="2014-12-19" valueMin="2014-12-10">
<collectingDate id="17793D0D1877FFE94322352EAA7EED15" pageId="51" pageNumber="52" value="2014-12-10" valueMax="2014-12-19" valueMin="2014-12-10">10– 19 December 2014</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE9461A354DAA83ED15" box="[341,436,1090,1113]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and
<collectorName id="DE7687F31877FFE946A6354DA9B6ED15" box="[489,641,1090,1113]" pageId="51" pageNumber="52">R. Harin’Hala</collectorName>
[MG-64B-95] (
<specimenCode id="23254A5E1877FFE94463354DA8C2ED15" box="[812,1013,1090,1113]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9065491</specimenCode>
,
<specimenCount id="658529AC1877FFE9434E354DAF15ED15" box="[1025,1058,1090,1113]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9437F354DAF50ED15" box="[1072,1127,1090,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781877FFE94334354DAFA5ED35" collectingDate="2013-12-05" collectingDateMax="2013-12-19" collectingDateMin="2013-12-05" collectionCode="CAS" collectorName="M. Irwin &amp; Harin'Hala" country="Madagascar" pageId="51" pageNumber="52" specimenCode="CASENT9065405" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<date id="073DC4E51877FFE94334354DAA79ED35" pageId="51" pageNumber="52" value="2013-12-05" valueMax="2013-12-19" valueMin="2013-12-05">
<collectingDate id="17793D0D1877FFE94334354DAA79ED35" pageId="51" pageNumber="52" value="2013-12-05" valueMax="2013-12-19" valueMin="2013-12-05">5– 19 December 2013</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE94613356DAA89ED35" box="[348,446,1122,1145]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and R.
<collectorName id="DE7687F31877FFE94556356DA9A6ED35" box="[537,657,1122,1145]" pageId="51" pageNumber="52">Harin’Hala</collectorName>
{MG-64B-63} (
<specimenCode id="23254A5E1877FFE94406356DAF24ED35" box="[841,1043,1122,1145]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9065405</specimenCode>
,
<specimenCount id="658529AC1877FFE9436F356DAF75ED35" box="[1056,1090,1122,1145]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9431D356DAFBDED35" box="[1106,1162,1122,1145]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781877FFE94734358DAAF0EDB6" collectingDate="2001-11-16" collectingDateMax="2001-11-20" collectingDateMin="2001-11-16" collectionCode="CAS" collectorName="Fisher-Griswold Arthropod Team" country="Madagascar" elevation="150" latitude="-19.709446" location="Antsalova" longLatPrecision="21" longitude="44.718056" municipality="Parc National Tsingy de Bemaraha" pageId="51" pageNumber="52" specimenCode="CASENT9009383" specimenCount="1" specimenCount-female="1" stateProvince="Mahajanga" typeStatus="paratype">
<emphasis id="41F73E371877FFE94734358DAA52EDD5" box="[123,357,1154,1177]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE94734358DAA52EDD5" box="[123,357,1154,1177]" country="Madagascar" name="Mahajanga" pageId="51" pageNumber="52">Mahajanga Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1877FFE94632358DA82EEDD5" box="[381,793,1154,1177]" pageId="51" pageNumber="52">Parc National Tsingy de Bemaraha</collectingMunicipality>
,
<quantity id="B47B4FC01877FFE94461358DA8B8EDD5" box="[814,911,1154,1177]" metricMagnitude="4" metricUnit="m" metricValue="1.06" pageId="51" pageNumber="52" unit="km" value="10.6">10.6 km</quantity>
ESE 123°
<location id="765CB4FE1877FFE9436D358DAFA6EDD5" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE9436D358DAFA6EDD5" box="[1058,1169,1154,1177]" country="Madagascar" latitude="-19.709446" longLatPrecision="21" longitude="44.718056" municipality="Parc National Tsingy de Bemaraha" name="Antsalova" pageId="51" pageNumber="52" stateProvince="Mahajanga">Antsalova</location>
,
<geoCoordinate id="16B784E21877FFE9473435ADABC2EDF5" box="[123,245,1186,1209]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="34" value="-19.709446">19°42ʹ34ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21877FFE9464B35ADAA43EDF5" box="[260,372,1186,1209]" degrees="44" direction="east" minutes="43" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="5" value="44.718056">44°43ʹ5ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01877FFE946F335ADAACCEDF5" box="[444,507,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="m" value="150.0">
<elevation id="F8AE05161877FFE946F335ADAACCEDF5" box="[444,507,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="m" value="150.0">150m</elevation>
</quantity>
, tropical dry forest on tsingy, general collecting beating and puffing spiders,
<date id="073DC4E51877FFE9466135CCA91BED96" box="[302,556,1219,1242]" pageId="51" pageNumber="52" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">
<collectingDate id="17793D0D1877FFE9466135CCA91BED96" box="[302,556,1219,1242]" pageId="51" pageNumber="52" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">16–20 November 2001</collectingDate>
</date>
,
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[BLF4503] (
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,
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,
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,
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<materialsCitation id="C3EBE8781877FFE9469935ECABE3EC37" collectingDate="2000-11-02" collectingDateMax="2000-11-08" collectingDateMin="2000-11-02" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1250" location="Manasy River" municipality="Foret d'Analavelona" pageId="51" pageNumber="52" specimenCode="CASENT9062273, CASENT9062271, CASENT9062272" specimenCount="6" specimenCount-male="6" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="41F73E371877FFE9469935ECA9BCEDB6" box="[470,651,1251,1274]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE9469935ECA9BCEDB6" box="[470,651,1251,1274]" country="Madagascar" name="Toliara" pageId="51" pageNumber="52">Toliara Province</collectingRegion>
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:
<collectingMunicipality id="9358785F1877FFE945D035ECA8B4EDB6" box="[671,899,1251,1274]" pageId="51" pageNumber="52">Fôret d’Analavelona</collectingMunicipality>
, near source of
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,
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NW d’Andranoheza
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, 22°38.6ʹS, 044°10.3ʹE, elev.
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, undisturbed mid-elevation forest w/dry and humid elements,
<collectingMethod id="AAC29A321877FFE94559342CA96FEC76" box="[534,600,1315,1338]" pageId="51" pageNumber="52">pitfall</collectingMethod>
,
<date id="073DC4E51877FFE9452C342CA876EC76" box="[611,833,1315,1338]" pageId="51" pageNumber="52" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">
<collectingDate id="17793D0D1877FFE9452C342CA876EC76" box="[611,833,1315,1338]" pageId="51" pageNumber="52" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">2–8 November 2000</collectingDate>
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,
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, [FMHD#00-224] (
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,
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,
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,
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.
</paragraph>
<paragraph id="733CE2251877FFE947E4348BA933EF70" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E4348BAA54ECD7" bold="true" box="[171,355,1412,1435]" pageId="51" pageNumber="52">Natural history.</emphasis>
Except for a series of males collected in disturbed transitional montane mossy forest at Réserve Spéciale d’Ambohitantely, other collections for
<taxonomicName id="B48399A61877FFE9443634ABAF3CECF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[889,1035,1444,1467]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE9443634ABAF3CECF7" bold="true" box="[889,1035,1444,1467]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
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are from dry habitats including in
<taxonomicName id="B48399A61877FFE9462834CBAA8CEC97" box="[359,443,1476,1499]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E371877FFE9462834CBAA8CEC97" bold="true" box="[359,443,1476,1499]" italics="true" pageId="51" pageNumber="52">Uapaca</emphasis>
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forest, on inselbergs and in tsingy. Females of
<taxonomicName id="B48399A61877FFE9449B34CBAF5DEC97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[980,1130,1476,1499]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE9449B34CBAF5DEC97" bold="true" box="[980,1130,1476,1499]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
</taxonomicName>
are cribellate and may produce sticky silk, though the collecting data that state “general collecting…’ reveal nothing of burrows or retreats. As is typical of male
<taxonomicName id="B48399A61877FFE945A0370AA80EEF50" authorityName="Simon" authorityYear="1880" box="[751,825,1541,1564]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371877FFE945A0370AA80EEF50" bold="true" box="[751,825,1541,1564]" italics="true" pageId="51" pageNumber="52">Uduba</emphasis>
</taxonomicName>
, these were collected wandering on the ground and in the vegetation.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1877FFE947E4374AAFA2EF30" pageId="51" pageNumber="52" type="distribution">
<paragraph id="733CE2251877FFE947E4374AAFA2EF30" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E4374AAA0AEF10" bold="true" box="[171,317,1605,1628]" pageId="51" pageNumber="52">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61877FFE94685374AA96AEF10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[458,605,1605,1628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE94685374AA96AEF10" bold="true" box="[458,605,1605,1628]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
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is recorded on the west side of the great escarpment across the southern half of
<collectingCountry id="0B94A2B51877FFE946EE376AA915EF30" box="[417,546,1637,1660]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
on the high plateau and western dry forests (
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, 18).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C1E5BBEB30897E616F657AFEA7E8786D">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<emphasis id="41F73E37187DFFE3473431D3AA27E9BF" bold="true" box="[123,272,220,243]" italics="true" pageId="57" pageNumber="58">Uduba lamba</emphasis>
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,
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<figureCitation id="EBB8FEA0187DFFE3473431F3ABD8E85D" box="[123,239,252,273]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="57" pageNumber="58">Figures 9 H</figureCitation>
,
<figureCitation id="EBB8FEA0187DFFE347B531F3AA1DE85D" box="[250,298,252,273]" captionStart="FIGURE" captionStartId="127.[155,167,738,757]" captionTargetBox="[120,1176,216,721]" captionTargetId="figure-16@127.[120,1176,216,721]" captionTargetPageId="127" captionText="FIGURE 26. Spinning organs of Uduba spp. A. B. U. schlingeri ♀, (CASENT9006075), Vatoharanana, Parc Nacional Ranomafana. C. U. lamba, ♂, holotype, (CASENT9042542), R.N.I. Betampona. Cl, colulus, Cr, cribellum. Scale bars: A, B = 0.5 mm, C = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158625" httpUri="https://zenodo.org/record/13158625/files/figure.png" pageId="57" pageNumber="58">26 C</figureCitation>
,
<figureCitation id="EBB8FEA0187DFFE3467A31F3AAB5E85D" box="[309,386,252,273]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">49 A–C</figureCitation>
, Maps 12, 13.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187DFFE347E4302EA9E4E8D4" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="733CE225187DFFE347E4302EA9E4E8D4" blockId="57.[122,1175,289,1725]" pageId="57" pageNumber="58">
<emphasis id="41F73E37187DFFE347E4302EAA6FE874" bold="true" box="[171,344,289,312]" pageId="57" pageNumber="58">Type material.</emphasis>
<materialsCitation id="C3EBE878187DFFE34628302EA8D4E834" collectingDate="2008-05-25" collectingDateMax="2008-06-01" collectingDateMin="2008-05-25" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="525" latitude="-17.919722" location="Reserve Nationale Integrale Betampona" longLatPrecision="21" longitude="49.20028" pageId="57" pageNumber="58" specimenCode="CASENT9042542" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C87187DFFE34628302EAAFDE874" box="[359,458,289,312]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="23254A5E187DFFE3456A302EA9DAE874" box="[549,749,289,312]" collectionCode="CASENT" pageId="57" pageNumber="58">CASENT9042542</specimenCode>
) from rainforest at
<quantity id="B47B4FC0187DFFE344AE302EAF2AE874" box="[993,1053,289,312]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="57" pageNumber="58" unit="m" value="525.0">
<elevation id="F8AE0516187DFFE344AE302EAF2AE874" box="[993,1053,289,312]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="57" pageNumber="58" unit="m" value="525.0">525m</elevation>
</quantity>
elevation,
<location id="765CB4FE187DFFE34734304EA904E814" LSID="urn:lsid:plazi:treatment:FB2A5333187DFFE0473431D3AAC0EB98:765CB4FE187DFFE34734304EA904E814" box="[123,563,321,344]" country="Madagascar" latitude="-17.919722" longLatPrecision="21" longitude="49.20028" name="Reserve Nationale Integrale Betampona" pageId="57" pageNumber="58" stateProvince="Toamasina">Réserve Nationale Intégrale Betampona</location>
,
<geoCoordinate id="16B784E2187DFFE3450D304EA98CE814" box="[578,699,321,344]" degrees="17" direction="south" minutes="55" orientation="latitude" pageId="57" pageNumber="58" precision="15" seconds="11" value="-17.919722">17°55ʹ11ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E2187DFFE34584304EA861E814" box="[715,854,321,344]" degrees="049" direction="east" minutes="12" orientation="longitude" pageId="57" pageNumber="58" precision="15" seconds="01" value="49.20028">049°12ʹ01ʺE</geoCoordinate>
,
<collectingRegion id="B1472CC7187DFFE3442C304EAF76E814" box="[867,1089,321,344]" country="Madagascar" name="Toamasina" pageId="57" pageNumber="58">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B5187DFFE3431F304EAB8AE834" name="Madagascar" pageId="57" pageNumber="58">Madagascar</collectingCountry>
, collected
<date id="073DC4E5187DFFE3460E306EA90EE834" box="[321,569,353,376]" pageId="57" pageNumber="58" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">
<collectingDate id="17793D0D187DFFE3460E306EA90EE834" box="[321,569,353,376]" pageId="57" pageNumber="58" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">25 May–1 June 2008</collectingDate>
</date>
by
<collectorName id="DE7687F3187DFFE3453E306EA9C8E834" box="[625,767,353,376]" pageId="57" pageNumber="58">B. L. Fisher</collectorName>
, deposited in
<collectionCode id="15927AE0187DFFE344E3306EA8D4E834" box="[940,995,353,376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="57" pageNumber="58" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE878187DFFE344B9306EA9F8E8D4" collectingDate="2008-05-25" collectingDateMax="2008-06-01" collectingDateMin="2008-05-25" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="525" latitude="-17.919722" location="Reserve Nationale Integrale Betampona" longLatPrecision="21" longitude="49.20028" pageId="57" pageNumber="58" specimenCode="CASENT9042359" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="AC385C87187DFFE344B9306EAF64E834" box="[1014,1107,353,376]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
male (
<specimenCode id="23254A5E187DFFE347CD308EAA7AE8D4" box="[130,333,385,408]" collectionCode="CASENT" pageId="57" pageNumber="58">CASENT9042359</specimenCode>
), same data, also deposited in
<collectionCode id="15927AE0187DFFE345D7308EA9F8E8D4" box="[664,719,385,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="57" pageNumber="58" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187DFFE347E430ADA954E895" pageId="57" pageNumber="58" type="etymology">
<paragraph id="733CE225187DFFE347E430ADA954E895" blockId="57.[122,1175,289,1725]" pageId="57" pageNumber="58">
<emphasis id="41F73E37187DFFE347E430ADAA1DE8F5" bold="true" box="[171,298,418,441]" pageId="57" pageNumber="58">Etymology.</emphasis>
The species epithet,
<emphasis id="41F73E37187DFFE3455230ADA957E8F5" box="[541,608,418,441]" italics="true" pageId="57" pageNumber="58">lamba</emphasis>
, refers to a traditional garment worn by men and women in
<collectingCountry id="0B94A2B5187DFFE347A130CDAA45E895" box="[238,370,450,473]" name="Madagascar" pageId="57" pageNumber="58">Madagascar</collectingCountry>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187DFFE347E430EDAA0BEBB6" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="733CE225187DFFE347E430EDAA0BEBB6" blockId="57.[122,1175,289,1725]" pageId="57" pageNumber="58">
<emphasis id="41F73E37187DFFE347E430EDAA8EE8B5" bold="true" box="[171,441,482,505]" pageId="57" pageNumber="58">
Diagnosis.
<taxonomicName id="B48399A6187DFFE3466A30EDAA8EE8B5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,441,482,505]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="lamba">
<emphasis id="41F73E37187DFFE3466A30EDAA8EE8B5" bold="true" box="[293,441,482,505]" italics="true" pageId="57" pageNumber="58">Uduba lamba</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.a, the
<taxonomicName id="B48399A6187DFFE3444E30EDA8B5E8B5" authorityName="E. Simon" authorityYear="1903" baseAuthorityName="Simon" baseAuthorityYear="1903" box="[769,898,482,505]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="dahli">Uduba dahli</taxonomicName>
group (
<figureCitation id="EBB8FEA0187DFFE3449B30EDAF10E8B5" box="[980,1063,482,505]" captionStart="MAP" captionStartId="182.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-12@182.[648,1176,216,1153]" captionTargetPageId="182" captionText="MAP 13. Distribution of Group I.a, the Uduba dahli MAP 14. Distribution of Group I.b, the Uduba woodae group of Group I, the Epigynal atrium group: Uduba group of Group I, the Epigynal atrium group: Uduba lamba, U. fandroana, U. hainteny, U. kavanaughi, U. plat- andriamihajai, U. balsama, U. schlingeri, U. taralily, nicki, U. halabe, U. salegy, U. dahli Simon 1903 and U. lehibekokoa and U. woodae. U. ibonia." figureDoi="http://doi.org/10.5281/zenodo.13158789" httpUri="https://zenodo.org/record/13158789/files/figure.png" pageId="57" pageNumber="58">Map 13</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate, with a colulus (
<figureCitation id="EBB8FEA0187DFFE3440D330DA89BEB55" box="[834,940,514,537]" captionStart="FIGURE" captionStartId="127.[155,167,738,757]" captionTargetBox="[120,1176,216,721]" captionTargetId="figure-16@127.[120,1176,216,721]" captionTargetPageId="127" captionText="FIGURE 26. Spinning organs of Uduba spp. A. B. U. schlingeri ♀, (CASENT9006075), Vatoharanana, Parc Nacional Ranomafana. C. U. lamba, ♂, holotype, (CASENT9042542), R.N.I. Betampona. Cl, colulus, Cr, cribellum. Scale bars: A, B = 0.5 mm, C = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158625" httpUri="https://zenodo.org/record/13158625/files/figure.png" pageId="57" pageNumber="58">Fig. 26 C</figureCitation>
). Males have the TA3 with a small, screw-shaped apex, TA2 erect, truncate, apex a sharp point, with a large, complex MA, concave in center with strong marginal ridge with a transverse ventral lobe or with ventral point to prolateral (
<figureCitation id="EBB8FEA0187DFFE34605336DAAEBEB35" box="[330,476,610,633]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Figs. 49 A–C</figureCitation>
); distinguished from
<emphasis id="41F73E37187DFFE3458F336DAF0AEB35" box="[704,1085,610,633]" italics="true" pageId="57" pageNumber="58">
<taxonomicName id="B48399A6187DFFE3458F336DA8B2EB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[704,901,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E37187DFFE3458F336DA8B2EB35" bold="true" box="[704,901,610,633]" italics="true" pageId="57" pageNumber="58">Uduba fandroana</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A6187DFFE344F4336DAF0AEB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[955,1085,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E37187DFFE344F4336DAF0AEB35" bold="true" box="[955,1085,610,633]" italics="true" pageId="57" pageNumber="58">U. hainteny</emphasis>
</taxonomicName>
</emphasis>
by having the MA relatively large, with pronounced ventral projection (
<figureCitation id="EBB8FEA0187DFFE34407338DA8D7EBD5" box="[840,992,642,665]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Figs. 49 A–C</figureCitation>
): in
<taxonomicName id="B48399A6187DFFE3435C338DABFDEBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="fandroana">
<emphasis id="41F73E37187DFFE3435C338DABFDEBF5" bold="true" italics="true" pageId="57" pageNumber="58">Uduba fandroana</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA0187DFFE3479733ADAA50EBF5" box="[216,359,674,697]" captionStart="FIGURE" captionStartId="137.[155,167,832,851]" captionTargetBox="[125,1166,230,807]" captionTargetId="figure-16@137.[120,1176,216,819]" captionTargetPageId="137" captionText="FIGURE 39. Uduba fandroana, new species, Holotype (CASENT9002691) from Anjozorobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. C, conductor; Cm, cymbium; Eb, embolar base; MA, median apophysis; Pt, petiole of subtegulum; RTA, retrolateral tibial apophysis on male palp; ST, subtegulum; T, tegulum; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum, arising just prolaterad of distal notch; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp, which is located at proximoventral margin of tibia extending above patellar intersegmental membrane; VTA, ventral tibial apophysis on male palp, typically arising at midsegment. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158657" httpUri="https://zenodo.org/record/13158657/files/figure.png" pageId="57" pageNumber="58">Figs. 39 A–C</figureCitation>
) and
<taxonomicName id="B48399A6187DFFE346EC33ADA913EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[419,548,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hainteny">
<emphasis id="41F73E37187DFFE346EC33ADA913EBF5" bold="true" box="[419,548,674,697]" italics="true" pageId="57" pageNumber="58">U. hainteny</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA0187DFFE3457B33ADA9F4EBF5" box="[564,707,674,697]" captionStart="FIGURE" captionStartId="140.[155,167,888,907]" captionTargetBox="[120,1176,216,871]" captionTargetId="figure-11@140.[120,1176,216,872]" captionTargetPageId="140" captionText="FIGURE 42. Uduba hainteny, new species, Holotype (CASENT9032857) from Forêt Ivohibe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158663" httpUri="https://zenodo.org/record/13158663/files/figure.png" pageId="57" pageNumber="58">Figs. 42 A–C</figureCitation>
) the MA is relatively small, width less than 0.35 tegulum width, height less than 0.25 tegulum height and without prominent projections. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187DFFE047E4320CA9CAEBD8" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="733CE225187DFFE347E4320CAF4DEC90" blockId="57.[122,1175,289,1725]" pageId="57" pageNumber="58">
<emphasis id="41F73E37187DFFE347E4320CAA02EA56" bold="true" box="[171,309,771,794]" pageId="57" pageNumber="58">Description.</emphasis>
Male (
<typeStatus id="AC385C87187DFFE346C7320CAADBEA56" box="[392,492,771,794]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
): Total length 12.50. Markings as in
<figureCitation id="EBB8FEA0187DFFE344C6320CA8DCEA56" box="[905,1003,771,794]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="57" pageNumber="58">Fig. 9 H</figureCitation>
. Carapace 8.60 long, 5.70 wide, 2.80 high; clypeus 0.40 high. Eye diameters:AME and ALE 0.26, PME 0.28, PLE 0.36. Chelicerae 3.40 long; sternum 3.50 long, 2.80 wide; labium 2.20 long; palpal coxae 2.70 long. With a colulus (
<figureCitation id="EBB8FEA0187DFFE3466D326CAABDEA36" box="[290,394,867,890]" captionStart="FIGURE" captionStartId="127.[155,167,738,757]" captionTargetBox="[120,1176,216,721]" captionTargetId="figure-16@127.[120,1176,216,721]" captionTargetPageId="127" captionText="FIGURE 26. Spinning organs of Uduba spp. A. B. U. schlingeri ♀, (CASENT9006075), Vatoharanana, Parc Nacional Ranomafana. C. U. lamba, ♂, holotype, (CASENT9042542), R.N.I. Betampona. Cl, colulus, Cr, cribellum. Scale bars: A, B = 0.5 mm, C = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158625" httpUri="https://zenodo.org/record/13158625/files/figure.png" pageId="57" pageNumber="58">Fig. 26 C</figureCitation>
). Spination (
<typeStatus id="AC385C87187DFFE3455E326CA942EA36" box="[529,629,867,890]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
): palpus–femur d0-0-1, p0-0-1, r0-0-1; leg I–femur d1-1-1, p0-0-2, r0-0-1, tibia p0-1-1-0, v2-2-2-2, r0-1-0-0, metatarsus p1-1-0, v2-2-3, r1-1-0; leg II– femur d1-1-1, p0-1-1, r0-1-0-1, tibia d0-1-1-0, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1-0, v2- 2-3, r1-1-0; leg III–femur d1-1-1, p0-1-1-1, r0-1-1-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-1-2, v2-2-2, r1-1-2; leg IV–femur d1-1-0-1, p0-1-0-1, r0-0-0-1, tibia v2[segments missing from both sides]. Scopulae: cymbium, apicodorsal; weak beneath tarsi and apices of all metatarsi. Leg measurements (
<typeStatus id="AC385C87187DFFE3468F352BA917ED77" box="[448,544,1060,1083]" pageId="57" pageNumber="58" type="paratype">paratype</typeStatus>
, CASENT9042359): I: 8.50 + 3.30 + 8.20 + 9.00 + 4.60 = 33.60; II: 7.20 + 2.70 + 6.70 + 7.20 + 3.90 = 27.70; III: 6.00 + 3.40 + 4.00 + 6.30 + 3.20 = 22.90; IV [metatarsus and tarsus are missing, so their lengths and leg IV total length have been estimat- ed]: 8.40 + 3.60 + 7.10 + 9.14 + 4.03 = 32.27; palpus: 3.20 + 1.20 + 1.40 + NA + 4.00 = 9.80. Leg formula 1423.
<emphasis id="41F73E37187DFFE3465235ABAAA5EDF7" bold="true" box="[285,402,1188,1211]" pageId="57" pageNumber="58">Male palp</emphasis>
(
<typeStatus id="AC385C87187DFFE346ED35ABA933EDF7" box="[418,516,1188,1211]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
,
<figureCitation id="EBB8FEA0187DFFE3455E35ABA990EDF7" box="[529,679,1188,1211]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Figs. 49 A–C</figureCitation>
): palpal tibia 0.40 times cymbial length, RTA short, triangular, with blunt apex (
<figureCitation id="EBB8FEA0187DFFE346A535CBA964ED97" box="[490,595,1220,1243]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Fig. 49 C</figureCitation>
), RTA length 0.78 times width, RTA length 0.21 times tibia length; VTA slender, nearly cylindrical (
<figureCitation id="EBB8FEA0187DFFE3451635EBA9F7EDB7" box="[601,704,1252,1275]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Fig. 49 A</figureCitation>
), length 1.71 times width, length 0.40 times tibia width; tibia lacks stout spines; tegulum convex, without ridge but with retrolateral concavity (
<figureCitation id="EBB8FEA0187DFFE347CC342AABD9EC70" box="[131,238,1317,1340]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Fig. 49 B</figureCitation>
), tegulum length 1.08 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.36 times tegulum length; TA3 with a small, screw-shaped apex, TA3 even with TA2 apex, TA2 erect, truncate with apex a sharp point (
<figureCitation id="EBB8FEA0187DFFE34588346AA819EC30" box="[711,814,1381,1404]" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Fig. 49 B</figureCitation>
); MA large, complex, length 0.31 tegulum length, width 0.435 tegulum width, MA origin at 0.36 tegulum length, tegulum apex at 0.48 tegulum length from tegulum apex, length (including apical processes) 0.76 times width (
<figureCitation id="EBB8FEA0187DFFE3432F34AAABE7EC90" captionStart="FIGURE" captionStartId="144.[155,167,841,860]" captionTargetBox="[134,1158,233,814]" captionTargetId="figure-11@144.[120,1176,216,831]" captionTargetPageId="144" captionText="FIGURE 49. Uduba lamba, new species, Holotype (CASENT9052542) from Réserve National Intégrale Betampona, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Arrow to retrolateral notch in tegulum. Arrow to bump on tegulum. Illustrations by Rachel Diaz-Bastin. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158683" httpUri="https://zenodo.org/record/13158683/files/figure.png" pageId="57" pageNumber="58">Figs. 49 A–C</figureCitation>
), MA with transverse ventral lobe; conductor fan concave or entire.
<emphasis id="41F73E37187DFFE344FB34CAAF30EC90" bold="true" box="[948,1031,1477,1500]" pageId="57" pageNumber="58">Female</emphasis>
unknown.
</paragraph>
<paragraph id="733CE225187DFFE047E534EAA92BE85F" blockId="57.[122,1175,289,1725]" lastBlockId="58.[123,1174,220,724]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="41F73E37187DFFE347E534EAAA69ECB0" bold="true" box="[170,350,1509,1532]" pageId="57" pageNumber="58">Variation. Male</emphasis>
(N = 2) Total length = 12.50–14.29, carapace length / width = 1.40–1.51, carapace height / width = 0.41–0.49, PER / carapace width = 0.37–0.39, PER / OAL = 2.33–2.76, PER / AER = 1.33–1.37, OAL / OQL = 1.04–1.12, OQP / OQA = 1.05–1.08, clypeus height / AME = 1.25–1.54, cheliceral length / clypeus height = 8.50–9.00, palpal coxa length / width = 2.45–2.70, femur I length / carapace width = 1.37–1.47, metatarsus I length / carapace width = 1.44–1.55, femur IV length / carapace width = 1.45–1.46, cymbium length / carapace width = 0.63–0.69, cymbium length / palpal patella length = 3.27–3.33, cymbium length / palpal tibia length = 2.86–3.00, cymbium length / palpal femur length = 1.20–1.25, palpal tibia length / palpal patella length = 1.09–1.17. Female variation unknown.
</paragraph>
<paragraph id="733CE225187EFFE047E43013ABD8EB78" blockId="58.[123,1174,220,724]" pageId="58" pageNumber="59">
<materialsCitation id="C3EBE878187EFFE047E43013A979E83F" collectingDate="1994-10-19" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="875" location="Res. Anjanaharibe-Sud" pageId="58" pageNumber="59" specimenCode="CASENT9006020" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
M
<emphasis id="41F73E37187EFFE0478D3013AABEE87F" bold="true" box="[194,393,284,307]" pageId="58" pageNumber="59">aterial examined.</emphasis>
<collectingCountry id="0B94A2B5187EFFE046DF3013A965E87F" box="[400,594,284,307]" name="Madagascar" pageId="58" pageNumber="59">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37187EFFE045123013A870E87F" box="[605,839,284,307]" italics="true" pageId="58" pageNumber="59">
<collectingRegion id="B1472CC7187EFFE045123013A870E87F" box="[605,839,284,307]" country="Madagascar" name="Antsiranana" pageId="58" pageNumber="59">Antsiranana Province</collectingRegion>
</emphasis>
:
<locationDeviation id="628EFB07187EFFE044193013AF6CE87F" box="[854,1115,284,307]" location="Befingotra" pageId="58" pageNumber="59">
<quantity id="B47B4FC0187EFFE044193013A896E87F" box="[854,929,284,307]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="58" pageNumber="59" unit="km" value="6.5">6.5 km</quantity>
SSW Befingotra
</locationDeviation>
,
<location id="765CB4FE187EFFE043283013AA77E81F" LSID="urn:lsid:plazi:treatment:FB2A5333187DFFE0473431D3AAC0EB98:765CB4FE187EFFE043283013AA77E81F" country="Madagascar" name="Res. Anjanaharibe-Sud" pageId="58" pageNumber="59" stateProvince="Antsiranana">Rés. Anjanaharibe-Sud</location>
, 14º45ʹS; 49º30ʹE, elev.
<quantity id="B47B4FC0187EFFE0452E3033A997E81F" box="[609,672,316,339]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="58" pageNumber="59" unit="m" value="875.0">
<elevation id="F8AE0516187EFFE0452E3033A997E81F" box="[609,672,316,339]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="58" pageNumber="59" unit="m" value="875.0">875m</elevation>
</quantity>
, rainforest,
<date id="073DC4E5187EFFE044613033A8C3E81F" box="[814,1012,316,339]" pageId="58" pageNumber="59" value="1994-10-19">
<collectingDate id="17793D0D187EFFE044613033A8C3E81F" box="[814,1012,316,339]" pageId="58" pageNumber="59" value="1994-10-19">19 October 1994</collectingDate>
</date>
,
<collectorName id="DE7687F3187EFFE043493033AFA1E81F" box="[1030,1174,316,339]" pageId="58" pageNumber="59">B. L. Fisher</collectorName>
[BLF1076
<emphasis id="41F73E37187EFFE047A53053ABC4E83F" bold="true" box="[234,243,348,371]" pageId="58" pageNumber="59">]</emphasis>
(
<specimenCode id="23254A5E187EFFE0464B3053AAF9E83F" box="[260,462,348,371]" collectionCode="CASENT" pageId="58" pageNumber="59">CASENT9006020</specimenCode>
,
<specimenCount id="658529AC187EFFE046933053AACAE83F" box="[476,509,348,371]" count="1" pageId="58" pageNumber="59" type="male">1♂</specimenCount>
,
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)
</materialsCitation>
.
<materialsCitation id="C3EBE878187EFFE045143053AA33E89F" collectingDate="1997-03-05" collectingDateMax="1997-03-11" collectingDateMin="1997-03-05" collectionCode="FMNH" collectorName="S. E. Goodman" country="Madagascar" county="Reserve Nationale Integrale Andringitra" elevation="1990" location="Philippia" municipality="Anjavidilay" pageId="58" pageNumber="59" specimenCode="CASENT9064649" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa">
<emphasis id="41F73E37187EFFE045143053A862E83F" box="[603,853,348,371]" italics="true" pageId="58" pageNumber="59">
<collectingRegion id="B1472CC7187EFFE045143053A862E83F" box="[603,853,348,371]" country="Madagascar" name="Fianarantsoa" pageId="58" pageNumber="59">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectingCounty id="9A5D9AA9187EFFE0442A3053ABCDE8DF" pageId="58" pageNumber="59">Réserve Nationale Intégrale Andringitra</collectingCounty>
,
<collectingMunicipality id="9358785F187EFFE0464B3073AA49E8DF" box="[260,382,380,403]" pageId="58" pageNumber="59">Anjavidilay</collectingMunicipality>
,
<locationDeviation id="628EFB07187EFFE046C13073A944E8DF" box="[398,627,380,403]" location="Atanifotsy" pageId="58" pageNumber="59">
<quantity id="B47B4FC0187EFFE046C13073AAEFE8DF" box="[398,472,380,403]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="58" pageNumber="59" unit="km" value="8.5">8.5 km</quantity>
SE Atanifotsy
</locationDeviation>
, 22°09.5ʹS, 46°57.6ʹE, elev.
<quantity id="B47B4FC0187EFFE044E23073A8CEE8DF" box="[941,1017,380,403]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="58" pageNumber="59" unit="m" value="1990.0">
<elevation id="F8AE0516187EFFE044E23073A8CEE8DF" box="[941,1017,380,403]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="58" pageNumber="59" unit="m" value="1990.0">1990m</elevation>
</quantity>
,
<collectingMethod id="AAC29A32187EFFE043493073AF40E8DF" box="[1030,1143,380,403]" pageId="58" pageNumber="59">pitfall trap</collectingMethod>
in
<emphasis id="41F73E37187EFFE047343093ABE9E8FF" box="[123,222,412,435]" italics="true" pageId="58" pageNumber="59">
<location id="765CB4FE187EFFE047343093ABE9E8FF" LSID="urn:lsid:plazi:treatment:FB2A5333187DFFE0473431D3AAC0EB98:765CB4FE187EFFE047343093ABE9E8FF" box="[123,222,412,435]" country="Madagascar" county="Reserve Nationale Integrale Andringitra" municipality="Anjavidilay" name="Philippia" pageId="58" pageNumber="59" stateProvince="Fianarantsoa">Philippia</location>
</emphasis>
-dominated sclerophyllus forest,
<date id="073DC4E5187EFFE0450C3093A832E8FF" box="[579,773,412,435]" pageId="58" pageNumber="59" value="1997-03-05" valueMax="1997-03-11" valueMin="1997-03-05">
<collectingDate id="17793D0D187EFFE0450C3093A832E8FF" box="[579,773,412,435]" pageId="58" pageNumber="59" value="1997-03-05" valueMax="1997-03-11" valueMin="1997-03-05">5–11 March 1997</collectingDate>
</date>
,
<collectorName id="DE7687F3187EFFE0445D3093A88FE8FF" box="[786,952,412,435]" pageId="58" pageNumber="59">S. E. Goodman</collectorName>
(
<specimenCode id="23254A5E187EFFE044873093AFA6E8FF" box="[968,1169,412,435]" collectionCode="CASENT" pageId="58" pageNumber="59">CASENT9064649</specimenCode>
,
<specimenCount id="658529AC187EFFE0473430B3ABABE89F" box="[123,156,444,467]" count="2" pageId="58" pageNumber="59" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0187EFFE047E430B3ABCAE89F" box="[171,253,444,467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="58" pageNumber="59" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE878187EFFE0464030B3A879EB58" collectingDate="2008-05-25" collectingDateMax="2008-06-01" collectingDateMin="2008-05-25" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="525" latitude="-17.919722" location="Reserve Nationale Integrale Betampona" longLatPrecision="21" longitude="49.20028" pageId="58" pageNumber="59" specimenCode="CASENT9052542" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="41F73E37187EFFE0464030B3AAD0E89F" box="[271,487,444,467]" italics="true" pageId="58" pageNumber="59">
<collectingRegion id="B1472CC7187EFFE0464030B3AAD0E89F" box="[271,487,444,467]" country="Madagascar" name="Toamasina" pageId="58" pageNumber="59">Toamasina Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE187EFFE046B830B3A890E89F" LSID="urn:lsid:plazi:treatment:FB2A5333187DFFE0473431D3AAC0EB98:765CB4FE187EFFE046B830B3A890E89F" box="[503,935,444,467]" country="Madagascar" latitude="-17.919722" longLatPrecision="21" longitude="49.20028" name="Reserve Nationale Integrale Betampona" pageId="58" pageNumber="59" stateProvince="Toamasina">Réserve Nationale Intégrale Betampona</location>
,
<locationDeviation id="628EFB07187EFFE044FB30B3AB9FE8BF" location="Toamasina" pageId="58" pageNumber="59">
<quantity id="B47B4FC0187EFFE044FB30B3AF3CE89F" box="[948,1035,444,467]" metricMagnitude="4" metricUnit="m" metricValue="3.51" pageId="58" pageNumber="59" unit="km" value="35.1">35.1 km</quantity>
NW
<collectingRegion id="B1472CC7187EFFE0430930B3AB9FE8BF" country="Madagascar" name="Toamasina" pageId="58" pageNumber="59">Toamasina</collectingRegion>
</locationDeviation>
,
<geoCoordinate id="16B784E2187EFFE047FB30D3AA1BE8BF" box="[180,300,476,499]" degrees="17" direction="south" minutes="55" orientation="latitude" pageId="58" pageNumber="59" precision="15" seconds="11" value="-17.919722">17°55ʹ11ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E2187EFFE0467630D3AAF3E8BF" box="[313,452,476,499]" degrees="049" direction="east" minutes="12" orientation="longitude" pageId="58" pageNumber="59" precision="15" seconds="01" value="49.20028">049°12ʹ01ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC0187EFFE0454930D3A972E8BF" box="[518,581,476,499]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="58" pageNumber="59" unit="m" value="525.0">
<elevation id="F8AE0516187EFFE0454930D3A972E8BF" box="[518,581,476,499]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="58" pageNumber="59" unit="m" value="525.0">525m</elevation>
</quantity>
, rainforest,
<collectingMethod id="AAC29A32187EFFE0458A30D3A87DE8BF" box="[709,842,476,499]" pageId="58" pageNumber="59">malaise trap</collectingMethod>
,
<date id="073DC4E5187EFFE0441930D3AF0CE8BF" box="[854,1083,476,499]" pageId="58" pageNumber="59" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">
<collectingDate id="17793D0D187EFFE0441930D3AF0CE8BF" box="[854,1083,476,499]" pageId="58" pageNumber="59" value="2008-05-25" valueMax="2008-06-01" valueMin="2008-05-25">25 May–1 June 2008</collectingDate>
</date>
, coll.
<collectorName id="DE7687F3187EFFE0433230D3ABECEB58" pageId="58" pageNumber="59">B. L. Fisher</collectorName>
[BLF19594_23] (
<typeStatus id="AC385C87187EFFE046D330F2A934EB58" box="[412,515,509,532]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E187EFFE0454130F2A9E1EB58" box="[526,726,509,532]" collectionCode="CASENT" pageId="58" pageNumber="59">CASENT9052542</specimenCode>
,
<specimenCount id="658529AC187EFFE045AF30F2A836EB58" box="[736,769,509,532]" count="1" pageId="58" pageNumber="59" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187EFFE0444130F2A872EB58" box="[782,837,509,532]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="58" pageNumber="59" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE878187EFFE0441730F2ABDCEB78" collectingDate="2008-05-25" collectingDateMax="2008-06-01" collectingDateMin="2008-05-25" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="525" isTailEnumeration="true" latitude="-17.919722" location="Reserve Nationale Integrale Betampona" longLatPrecision="21" longitude="49.20028" pageId="58" pageNumber="59" specimenCode="CASENT9042359" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
(
<typeStatus id="AC385C87187EFFE0441030F2A888EB58" box="[863,959,509,532]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187EFFE0448530F2AFA5EB58" box="[970,1170,509,532]" collectionCode="CASENT" pageId="58" pageNumber="59">CASENT9042359</specimenCode>
,
<specimenCount id="658529AC187EFFE047343312ABABEB78" box="[123,156,541,564]" count="1" pageId="58" pageNumber="59" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0187EFFE047E43312ABD4EB78" box="[171,227,541,564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="58" pageNumber="59" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225187EFFE047E43332A9CAEBD8" blockId="58.[123,1174,220,724]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E43332AA56EB18" bold="true" box="[171,353,573,596]" pageId="58" pageNumber="59">Natural history.</emphasis>
Collection records document the occurrence of
<taxonomicName id="B48399A6187EFFE0443C3332AF3CEB18" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[883,1035,573,596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lamba">
<emphasis id="41F73E37187EFFE0443C3332AF3CEB18" bold="true" box="[883,1035,573,596]" italics="true" pageId="58" pageNumber="59">Uduba lamba</emphasis>
</taxonomicName>
in
<emphasis id="41F73E37187EFFE0437D3332AFA2EB18" box="[1074,1173,573,596]" italics="true" pageId="58" pageNumber="59">Philippia</emphasis>
dominated sclerophyllus forest in far southern
<collectingCountry id="0B94A2B5187EFFE0453C3352A9C3EB38" box="[627,756,605,628]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
and in rainforest at scattered localities in eastern
<collectingCountry id="0B94A2B5187EFFE047A43372AA5EEBD8" box="[235,361,637,660]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
. They make no use of cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187EFFE047E43392AAC0EB98" pageId="58" pageNumber="59" type="distribution">
<paragraph id="733CE225187EFFE047E43392AAC0EB98" blockId="58.[123,1174,220,724]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E43392AA0AEBF8" bold="true" box="[171,317,669,692]" pageId="58" pageNumber="59">Distribution.</emphasis>
The few records suggest that the species
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<emphasis id="41F73E37187EFFE044483392A8A9EBF8" bold="true" box="[775,926,669,692]" italics="true" pageId="58" pageNumber="59">Uduba lamba</emphasis>
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may be widespread in eastern
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(Maps 12, 13).
</paragraph>
</subSubSection>
</treatment>
</document>

313
data/FB/2A/53/FB2A5333187EFFE1473433F2A98FEC3B.xml Normal file
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<mods:title id="BAF4C148AED0780D42A536BDB470E189">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:affiliation id="8321CC303A4BCA77E996DFB9660B2E54">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="21A7DE9A20ED34F8C33E884EC50C6C10">Ubick, Darrell</mods:namePart>
<mods:affiliation id="2C9713465904D3FB9D23F4E4ED09B8C1">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:affiliation id="46C6B10CF7E6F7BFF072A1645709D84B">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="C84E2BB26220967EF0C92EC8473F99A5">Polotow, Daniele</mods:namePart>
<mods:affiliation id="8F396D641227271534207B3B5D3A280B">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<emphasis id="41F73E37187EFFE0473433F2AA92EA58" bold="true" box="[123,421,765,788]" pageId="58" pageNumber="59">
<taxonomicName id="B48399A6187EFFE0473433F2AA24EA58" ID-CoL="C8KJP" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,275,765,788]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lavitra" status="sp. nov.">
<emphasis id="41F73E37187EFFE0473433F2AA24EA58" bold="true" box="[123,275,765,788]" italics="true" pageId="58" pageNumber="59">Uduba lavitra</emphasis>
</taxonomicName>
,
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</emphasis>
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</paragraph>
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<paragraph id="733CE225187EFFE047343212AA9CEA7E" blockId="58.[123,427,765,818]" box="[123,427,797,818]" pageId="58" pageNumber="59">
<figureCitation id="EBB8FEA0187EFFE047343212AA19EA7E" box="[123,302,797,818]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="58" pageNumber="59">Figures 75 A–C, F</figureCitation>
, Maps 8, 15.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187EFFE047E4324CAFB6EAB6" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="733CE225187EFFE047E4324CAFB6EAB6" blockId="58.[122,1174,835,1725]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E4324CAA47EA16" bold="true" box="[171,368,835,858]" pageId="58" pageNumber="59">Type material.</emphasis>
<materialsCitation id="C3EBE878187EFFE046C2324CA989EAF6" collectingDate="2004-03-29" collectionCode="CAS" country="Madagascar" elevation="1070" latitude="-18.870834" location="Malagasy" longLatPrecision="21" longitude="48.3475" municipality="Torotorofotsy" pageId="58" pageNumber="59" specimenCode="CASENT9030898, CASENT9064776" specimenCount="2" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="AC385C87187EFFE046C2324CAACFEA16" box="[397,504,835,858]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
female (
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) and
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female (
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) collected
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<collectingDate id="17793D0D187EFFE046F0326CA956EA36" box="[447,609,867,890]" pageId="58" pageNumber="59" value="2004-03-29">29 March 2004</collectingDate>
</date>
by the
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ant team from a rotten log at marsh edge in montane rainforest at
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</quantity>
elevation at
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(
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,
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),
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,
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, deposited in the
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</materialsCitation>
.
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<typeStatus id="AC385C87187EFFE0458432ACA81FEAF6" box="[715,808,931,954]" pageId="58" pageNumber="59" type="paratype">Paratype</typeStatus>
female (
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), collected
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<collectingDate id="17793D0D187EFFE0478C32CCAA5DEA96" box="[195,362,963,986]" pageId="58" pageNumber="59" value="2004-03-23">23 March 2004</collectingDate>
</date>
by the
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ant team at
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elevation from montane rainforest at
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,
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,
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,
<collectingRegion id="B1472CC7187EFFE0455B32ECA9D8EAB6" box="[532,751,995,1018]" country="Madagascar" name="Toamasina" pageId="58" pageNumber="59">Toamasina Province</collectingRegion>
,
<collectingCountry id="0B94A2B5187EFFE045B532ECA84EEAB6" box="[762,889,995,1018]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
, also deposited in
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187EFFE047E4350BAA39ED17" pageId="58" pageNumber="59" type="etymology">
<paragraph id="733CE225187EFFE047E4350BAA39ED17" blockId="58.[122,1174,835,1725]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E4350BAA1DED57" bold="true" box="[171,298,1028,1051]" pageId="58" pageNumber="59">Etymology.</emphasis>
The species epithet is from a Malagasy word
<emphasis id="41F73E37187EFFE0445A350BA86BED57" box="[789,860,1028,1051]" italics="true" pageId="58" pageNumber="59">lavitra</emphasis>
, meaning far, referring to the great distance from the end of the epigynal medial lobe (ML) to the epigynal groove; an adjective in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187EFFE047E4356BA878EDB7" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="733CE225187EFFE047E4356BA878EDB7" blockId="58.[122,1174,835,1725]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E4356BAA89ED37" bold="true" box="[171,446,1124,1147]" pageId="58" pageNumber="59">
Diagnosis.
<taxonomicName id="B48399A6187EFFE04669356BAA89ED37" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[294,446,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lavitra">
<emphasis id="41F73E37187EFFE04669356BAA89ED37" bold="true" box="[294,446,1124,1147]" italics="true" pageId="58" pageNumber="59">Uduba lavitra</emphasis>
</taxonomicName>
</emphasis>
belong to Group I.c, the
<taxonomicName id="B48399A6187EFFE04585356BA85FED37" authorityName="Simon" authorityYear="1906" box="[714,872,1124,1147]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="funerea">Uduba funerea</taxonomicName>
group (
<figureCitation id="EBB8FEA0187EFFE044F4356BAF38ED37" box="[955,1039,1124,1147]" captionStart="MAP" captionStartId="183.[155,174,1173,1192]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-54@183.[648,1176,216,1157]" captionTargetPageId="183" captionText="MAP 15. Distribution of Group I.c, the Uduba funerea MAP 16. Distribution of Group II, the Epigynum lateral group of Group I, the Epigynal atrium group: Uduba projection group, or Uduba evanescens group: Uduba funerea Simon, 1906, U. barbarae, U. lavitra, U. orona and evanescens (Dahl, 1901), U. rakotofrah, U. volana, U. milamina. U. heliani, U. goodmani and Uduba pseudoevanescens." figureDoi="http://doi.org/10.5281/zenodo.13158793" httpUri="https://zenodo.org/record/13158793/files/figure.png" pageId="58" pageNumber="59">Map 15</figureCitation>
) of Group I, the Epigynal atrium group. They are ecribellate, with a colulus; female with epigynal plate with median lobe surrounded by depressed atrium, epigynal plate extensive posteriad of atrium, height greater than 0.15 times epigynal length, median lobe narrow, especially anteriorly, atrium width greater than three times ML width (
<figureCitation id="EBB8FEA0187EFFE046B335EBA9B9EDB7" box="[508,654,1252,1275]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="58" pageNumber="59">Figs. 75 A, C</figureCitation>
<emphasis id="41F73E37187EFFE045C235EBA9A1EDB7" bold="true" box="[653,662,1252,1275]" pageId="58" pageNumber="59">)</emphasis>
. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187EFFE147E4340BAA2AEAB9" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
<paragraph id="733CE225187EFFE047E4340BAFA2EC57" blockId="58.[122,1174,835,1725]" box="[171,1173,1284,1307]" pageId="58" pageNumber="59">
<emphasis id="41F73E37187EFFE047E4340BAA4DEC57" bold="true" box="[171,378,1284,1307]" pageId="58" pageNumber="59">Description. Male</emphasis>
unknown.
<emphasis id="41F73E37187EFFE046B5340BA97AEC57" bold="true" box="[506,589,1284,1307]" pageId="58" pageNumber="59">Female</emphasis>
(
<typeStatus id="AC385C87187EFFE0452F340BA9FDEC57" box="[608,714,1284,1307]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
): Total length 17.50. Markings typical of
</paragraph>
<paragraph id="733CE225187EFFE14734342AAA3DE8BF" blockId="58.[122,1174,835,1725]" lastBlockId="59.[122,1174,220,1399]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<taxonomicName id="B48399A6187EFFE04734342AABF2EC70" authorityName="Simon" authorityYear="1880" box="[123,197,1317,1340]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E37187EFFE04734342AABF2EC70" bold="true" box="[123,197,1317,1340]" italics="true" pageId="58" pageNumber="59">Uduba</emphasis>
</taxonomicName>
. Carapace 7.70 long, 5.50 wide, 3.20 high; clypeus 0.55 high. Eye diameters: AME 0.24, ALE 0.26, PME 0.28, PLE 0.34. Chelicerae 2.00 long; sternum 3.40 long, 2.70 wide; labium 1.90 long; palpal coxae 2.70 long. With a colulus. Spination: palpus–femur d0-1-2, patella p1-0, tibia d1-0, p2-0, tarsus p2-1, r0-1-0; leg I–femur d1-0-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II–femur d1-0-0, p0-0-1, tibia p0-1-0, v2-1-1-2, metatarsus v2-2-3; leg III–femur d1-0-0-1, p0-1- 1-0, r0-1-1-0, tibia p0-1-1-0, v1-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2; leg IV–femur d1- 0-0, r0-0-1, tibia v1-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-2-2. Scopulae: strong ventral beneath tarsi I–IV and metatarsus I, apical half of metatarsi I and II and apices of metatarsi III and IV. Leg measurements: I: 6.40 + 2.80 + 5.50 + 4.80 + 3.00 = 22.50; II: 5.50 + 2.60 + 5.50 + 4.30 + 2.60 = 20.50; III: 4.40 + 2.20 + 2.70 + 3.70 + 2.10 = 15.10; IV: 6.00 + 2.40 + 5.10 + 6.00 + 2.40 = 21.90; palpus: 2.60 + 1.35 + 1.70 + NA + 2.70 = 8.35. Leg formula 1423.
<emphasis id="41F73E37187EFFE043453769AB82EFD1" bold="true" pageId="58" pageNumber="59">Female genitalia</emphasis>
: epigynal plate with median lobe surrounded by depressed atrium (
<figureCitation id="EBB8FEA0187EFFE044CA3789A8DCEFD1" box="[901,1003,1670,1693]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="58" pageNumber="59">Fig. 75 A</figureCitation>
), epigynal plate width 1.25 times length; atrium sides convex, atrium width 1.82 times atrium length; epigynum length 2.36 times atrium length, atrium width 0.62 epigynum width; atrium width at side of ML 1.22 times ML width; epigynal plate extending far posteriad of atrium, atrium origin at 0.28 times epigynum length from epigastric groove; median lobe (
<figureCitation id="EBB8FEA0187FFFE1458C3013A810E87F" box="[707,807,284,307]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 A</figureCitation>
) very narrow, nearly straight sided and slightly wider at base, median lobe length 2.67 times width, length 1.47 times atrium length, 0.60 times epigynal length, extending far anteriad of atrium, ML extent anteriad of atrium 0.59 times epigynal length; lateral lobes narrow, atrium width 5.17 times LL width; copulatory openings beneath midpoint of lateral margin of atrium. Vulva (
<figureCitation id="EBB8FEA0187FFFE145F23093A811E8FF" box="[701,806,412,435]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 B</figureCitation>
) with spermathecal ducts making 2–3 broad curves, vulva length 0.68 times width, vulva width 3.88 times distance between fertilization ducts.
</paragraph>
<paragraph id="733CE225187FFFE147E430F2AA2AEAB9" blockId="59.[122,1174,220,1399]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E430F2AA2DEB58" bold="true" box="[171,282,509,532]" pageId="59" pageNumber="60">Variation.</emphasis>
(N=3). Total length 17.00–17.50; carapace length / width 1.40–1.54, carapace height/ width 0.56–0.61, PER/ carapace width 0.45–0.47, PER/OAL 3.24–3.34, PER/AER 1.29– 1.43, OAL/OQL 1.12–1.19, OQP/ OQA 1.09–1.18, clypeus height/ AME diameter width 1.54– 2.29, cheliceral length / clypeus height 7.65–9.25, sternum length / width 1.26–1.29, palpal coxa length / width 2.00–2.45, femur I length / carapace width =1.08–1.16, metatarsus I length / carapace width 0.87–0.90, femur IV length / carapace width 1.09–1.17, palpal tarsus length / carapace width 0.49–0.52, palpal tibia length / palpal patella length 1.86–2.00, palpal tarsus length / palpal tibia length 1.59–1.73, palpal tarsus length / palpal femur length 0.96–1.04, palpal tibia length / palpal patella length 1.07–1.26. Epigynal plate (
<figureCitation id="EBB8FEA0187FFFE145CB33F2A825EA58" box="[644,786,765,788]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Figs. 75 A, C</figureCitation>
<emphasis id="41F73E37187FFFE1445D33F2A82CEA58" bold="true" box="[786,795,765,788]" pageId="59" pageNumber="60">)</emphasis>
width 1.25–1.36 times length; atrium width 0.56–0.62 epigynum width; atrium origin at 0.18–0.275 times epigynum length from epigastric groove; median lobe length 2.67–2.83 times width, length 1.24–1.47 times atrium length, 0.52–0.60 times epigynal length, ML spreads narrowly (
<figureCitation id="EBB8FEA0187FFFE145973251A877EA39" box="[728,832,862,885]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 C</figureCitation>
) or broadly (
<figureCitation id="EBB8FEA0187FFFE144813251AF02EA39" box="[974,1077,862,885]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 A</figureCitation>
) at base; atrium width 4.17–5.17 times LL width. Vulva with spermathecal ducts making two (
<figureCitation id="EBB8FEA0187FFFE143463271AF58EAD9" box="[1033,1135,894,917]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 B</figureCitation>
) or three (
<figureCitation id="EBB8FEA0187FFFE1478F3291AA10EAF9" box="[192,295,926,949]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Fig. 75 F</figureCitation>
) broad curves, vulva length 0.68–0.83 times width, vulva width 3.88–7.25 times distance between fertilization ducts (
<figureCitation id="EBB8FEA0187FFFE1454432B1A9AAEA99" box="[523,669,958,981]" captionStart="FIGURE" captionStartId="163.[155,167,954,973]" captionTargetBox="[133,1157,216,925]" captionTargetId="figure-16@163.[120,1176,216,940]" captionTargetPageId="163" captionText="FIGURE 75. Uduba spp., female genitalia drawings. A, B. U. lavitra, new species, holotype (CASENT9030898) from Torotorofotsy. C, F. U. lavitra, new species, paratype (CASENT9030895) from Ambatovy. D, E. U. orona, new species, holotype (CASENT9006101) from Andringitra Reserve. A, C, D. Epigynum, ventral. B, E, F. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158741" httpUri="https://zenodo.org/record/13158741/files/figure.png" pageId="59" pageNumber="60">Figs. 75 B, F</figureCitation>
<emphasis id="41F73E37187FFFE145D232B1A991EA99" bold="true" box="[669,678,958,981]" pageId="59" pageNumber="60">)</emphasis>
. Male variation unknown, only females have been collected.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187FFFE147E532F1ABF0EDBA" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="733CE225187FFFE147E532F1A802EDDA" blockId="59.[122,1174,220,1399]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E532F1AAA2ED59" bold="true" box="[170,405,1022,1045]" pageId="59" pageNumber="60">Material examined.</emphasis>
<materialsCitation id="C3EBE878187FFFE146E832F1ABF6ED3A" collectingDate="2004-03-29" collectionCode="CAS" collectorName="Malagasy" country="Madagascar" elevation="1070" latitude="-18.870834" location="Torotorofotsy" longLatPrecision="21" longitude="48.3475" pageId="59" pageNumber="60" specimenCode="CASENT9030898, CASENT9064776" specimenCount="2" specimenCount-female="2" stateProvince="Toamasina" typeStatus="holotype">
<collectingCountry id="0B94A2B5187FFFE146E832F1A95EED59" box="[423,617,1022,1045]" name="Madagascar" pageId="59" pageNumber="60">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E37187FFFE1453632F1A86CED59" box="[633,859,1022,1045]" italics="true" pageId="59" pageNumber="60">
<collectingRegion id="B1472CC7187FFFE1453632F1A86CED59" box="[633,859,1022,1045]" country="Madagascar" name="Toamasina" pageId="59" pageNumber="60">Toamasina Province</collectingRegion>
</emphasis>
:
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,
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,
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, elev.
<quantity id="B47B4FC0187FFFE146073511AAA3ED79" box="[328,404,1054,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="59" pageNumber="60" unit="m" value="1070.0">
<elevation id="F8AE0516187FFFE146073511AAA3ED79" box="[328,404,1054,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="59" pageNumber="60" unit="m" value="1070.0">1070m</elevation>
</quantity>
, ex-rotten log, marsh edge, montane rainforest,
<date id="073DC4E5187FFFE144D53511AF76ED79" box="[922,1089,1054,1077]" pageId="59" pageNumber="60" value="2004-03-29">
<collectingDate id="17793D0D187FFFE144D53511AF76ED79" box="[922,1089,1054,1077]" pageId="59" pageNumber="60" value="2004-03-29">29 March 2004</collectingDate>
</date>
, collectors
<collectorName id="DE7687F3187FFFE147E63530AA26ED1A" box="[169,273,1087,1110]" pageId="59" pageNumber="60">Malagasy</collectorName>
ant team (
<typeStatus id="AC385C87187FFFE146CD3530AAD3ED1A" box="[386,484,1087,1110]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E187FFFE146A13530A981ED1A" box="[494,694,1087,1110]" collectionCode="CASENT" pageId="59" pageNumber="60">CASENT9030898</specimenCode>
,
<specimenCount id="658529AC187FFFE1458E3530A9D5ED1A" box="[705,738,1087,1110]" count="1" pageId="59" pageNumber="60" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0187FFFE145BF3530A811ED1A" box="[752,806,1087,1110]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="60" type="Museum">CAS</collectionCode>
,
<typeStatus id="AC385C87187FFFE1447E3530A8A7ED1A" box="[817,912,1087,1110]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187FFFE144D43530AF54ED1A" box="[923,1123,1087,1110]" collectionCode="CASENT" pageId="59" pageNumber="60">CASENT9064776</specimenCode>
,
<specimenCount id="658529AC187FFFE143223530AFB9ED1A" box="[1133,1166,1087,1110]" count="1" pageId="59" pageNumber="60" type="female">1♀</specimenCount>
,
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,)
</materialsCitation>
;
<materialsCitation id="C3EBE878187FFFE147873550A805EDDA" collectingDate="2004-03-23" collectionCode="CAS" country="Madagascar" elevation="1075" latitude="-18.850834" location="Ambatovy" longLatPrecision="21" longitude="48.32139" pageId="59" pageNumber="60" specimenCode="CASENT9030895" specimenCount="1" specimenCount-female="1" stateProvince="Toamasina Province" typeStatus="paratype">
<location id="765CB4FE187FFFE147873550AA02ED3A" LSID="urn:lsid:plazi:treatment:FB2A5333187EFFE1473433F2A98FEC3B:765CB4FE187FFFE147873550AA02ED3A" box="[200,309,1119,1142]" country="Madagascar" latitude="-18.850834" longLatPrecision="21" longitude="48.32139" name="Ambatovy" pageId="59" pageNumber="60" stateProvince="Toamasina Province">Ambatovy</location>
,
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,
<geoCoordinate id="16B784E2187FFFE146883550A966ED3A" box="[455,593,1119,1142]" degrees="048" direction="east" minutes="19" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="17" value="48.32139">048°19ʹ17ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC0187FFFE145DF3550A9EAED3A" box="[656,733,1119,1142]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="59" pageNumber="60" unit="m" value="1075.0">
<elevation id="F8AE0516187FFFE145DF3550A9EAED3A" box="[656,733,1119,1142]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="59" pageNumber="60" unit="m" value="1075.0">1075m</elevation>
</quantity>
, general collecting spiders, montane rainforest,
<date id="073DC4E5187FFFE147893570AA59EDDA" box="[198,366,1151,1174]" pageId="59" pageNumber="60" value="2004-03-23">
<collectingDate id="17793D0D187FFFE147893570AA59EDDA" box="[198,366,1151,1174]" pageId="59" pageNumber="60" value="2004-03-23">23 March 2004</collectingDate>
</date>
, (
<typeStatus id="AC385C87187FFFE146CD3570AAD5EDDA" box="[386,482,1151,1174]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E187FFFE146A13570A982EDDA" box="[494,693,1151,1174]" collectionCode="CASENT" pageId="59" pageNumber="60">CASENT9030895</specimenCode>
,
<specimenCount id="658529AC187FFFE1458E3570A9D4EDDA" box="[705,739,1151,1174]" count="1" pageId="59" pageNumber="60" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0187FFFE145BE3570A81EEDDA" box="[753,809,1151,1174]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="60" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225187FFFE147E43590ABF0EDBA" blockId="59.[122,1174,220,1399]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E43590AA54EDFA" bold="true" box="[171,355,1183,1206]" pageId="59" pageNumber="60">Natural history.</emphasis>
These ecribellate spiders have been collected in montane rainforest. The
<typeStatus id="AC385C87187FFFE1473435B0ABEFED9A" box="[123,216,1215,1238]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B48399A6187FFFE147B135B0AAA0ED9A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[254,407,1215,1238]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lavitra">
<emphasis id="41F73E37187FFFE147B135B0AAA0ED9A" bold="true" box="[254,407,1215,1238]" italics="true" pageId="59" pageNumber="60">Uduba lavitra</emphasis>
</taxonomicName>
was found hiding within a rotten log, but we know nothing of any use of silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187FFFE147E435F0A98FEC3B" pageId="59" pageNumber="60" type="distribution">
<paragraph id="733CE225187FFFE147E435F0A98FEC3B" blockId="59.[122,1174,220,1399]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E435F0AAECEC5A" bold="true" box="[171,475,1279,1302]" pageId="59" pageNumber="60">
Distribution.
<taxonomicName id="B48399A6187FFFE1460C35F0AAECEC5A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[323,475,1279,1302]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lavitra">
<emphasis id="41F73E37187FFFE1460C35F0AAECEC5A" bold="true" box="[323,475,1279,1302]" italics="true" pageId="59" pageNumber="60">Uduba lavitra</emphasis>
</taxonomicName>
</emphasis>
are known only from two nearby localities in montane rainforest in
<collectingRegion id="B1472CC7187FFFE147D93410AA59EC7A" box="[150,366,1311,1334]" country="Madagascar" name="Toamasina" pageId="59" pageNumber="60">Toamasina Province</collectingRegion>
in central
<collectingCountry id="0B94A2B5187FFFE146AE3410A955EC7A" box="[481,610,1311,1334]" name="Madagascar" pageId="59" pageNumber="60">Madagascar</collectingCountry>
(Maps 8, 15). Their forest habitat is threatened by a huge chrome-ore strip mine, which currently extends to less than a kilometer from the collection site of the
<typeStatus id="AC385C87187FFFE147A1346FAA7DEC3B" box="[238,330,1376,1399]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
(GoogleEarth,
<date id="073DC4E5187FFFE146BD346FA99AEC3B" box="[498,685,1376,1399]" pageId="59" pageNumber="60" value="2021-10-26">26 October 2021</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>

303
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@ -0,0 +1,303 @@
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<mods:title id="AEF868D1A76DCA5ED96B086431508E1E">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="FCFDE173C3D2D9265B66BB7B869C4F9F" type="personal">
<mods:role id="CD3F982DEC4922F18518C50C0203610B">
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<mods:namePart id="367550F970EF816E8CDD47CD52578225">Griswold, Charles</mods:namePart>
<mods:affiliation id="AA4C78DB9154AE8F92C7C342AC9B4FCA">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="E4B0344273B3860CE1C5EB5AC0674385">Ubick, Darrell</mods:namePart>
<mods:affiliation id="43F8E0240E334FB22050C0DF1151D60D">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="551B5FB02D7BC7629179D74281296749" type="personal">
<mods:role id="DF61595E6272F079C948477378705BB1">
<mods:roleTerm id="5D6D494E75E091F9380945A5053E2672">Author</mods:roleTerm>
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<mods:namePart id="8346A5986E93EA65645CD764012EA98E">Ledford, Joel</mods:namePart>
<mods:affiliation id="809FF8151A64519950223FBD643CFFD5">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
</mods:name>
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<mods:namePart id="EF006E2CF432918BB1C53DCCB357BDD1">Polotow, Daniele</mods:namePart>
<mods:affiliation id="4E6383DA433D80A847538E0D319B11D5">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:typeOfResource id="098BA810F72BB82EA8B08E8A8E4F42BB">text</mods:typeOfResource>
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<mods:date id="151D90D48D4C00108F6D9EAF0F3BEAA7">2022</mods:date>
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<subSubSection id="3B99B1AE187FFFE1473434AFAAD3ECFB" box="[123,484,1440,1463]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="733CE225187FFFE1473434AFAAD3ECFB" blockId="59.[123,484,1440,1493]" box="[123,484,1440,1463]" pageId="59" pageNumber="60">
<heading id="28745549187FFFE1473434AFAAD3ECFB" box="[123,484,1440,1463]" fontSize="10" level="2" pageId="59" pageNumber="60" reason="4">
<emphasis id="41F73E37187FFFE1473434AFAAD3ECFB" bold="true" box="[123,484,1440,1463]" pageId="59" pageNumber="60">
<taxonomicName id="B48399A6187FFFE1473434AFAA66ECFB" ID-CoL="C8KJQ" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,337,1440,1463]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lehibekokoa" status="sp. nov.">
<emphasis id="41F73E37187FFFE1473434AFAA66ECFB" bold="true" box="[123,337,1440,1463]" italics="true" pageId="59" pageNumber="60">Uduba lehibekokoa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C187FFFE1461034AFAAD3ECFB" box="[351,484,1440,1463]" pageId="59" pageNumber="60" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187FFFE1473434CFAAF4EC99" box="[123,451,1472,1493]" pageId="59" pageNumber="60" type="description">
<paragraph id="733CE225187FFFE1473434CFAAF4EC99" blockId="59.[123,484,1440,1493]" box="[123,451,1472,1493]" pageId="59" pageNumber="60">
<figureCitation id="EBB8FEA0187FFFE1473434CFABD9EC99" box="[123,238,1472,1493]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="59" pageNumber="60">Figures 9 C</figureCitation>
,
<figureCitation id="EBB8FEA0187FFFE147B734CFAA72EC99" box="[248,325,1472,1493]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="59" pageNumber="60">50 A–C</figureCitation>
,
<figureCitation id="EBB8FEA0187FFFE1461F34CFAAADEC99" box="[336,410,1472,1493]" captionStart="MAP" captionStartId="179.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1064]" captionTargetId="figure-17@179.[648,1176,216,1061]" captionTargetPageId="179" captionText="MAP 7. Distributions of Uduba evanescens (Dahl, 1901) MAP 8. Distributions of Uduba woodae, Uduba lavitra and Uduba lehibekokoa. and Uduba milamina." figureDoi="http://doi.org/10.5281/zenodo.13158781" httpUri="https://zenodo.org/record/13158781/files/figure.png" pageId="59" pageNumber="60">Maps 7</figureCitation>
, 14.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187FFFE147E434EAA9A6EF11" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="733CE225187FFFE147E434EAA9A6EF11" blockId="59.[123,1174,1509,1725]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E434EAAA66ECB0" bold="true" box="[171,337,1509,1532]" pageId="59" pageNumber="60">Type material.</emphasis>
<materialsCitation id="C3EBE878187FFFE1461534EAA9A6EF11" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CASENT" country="Madagascar" county="California Academy of Sciences." elevation="140" location="Parc National de Namoroka" municipality="Fisher-Griswold Arthropod Team" pageId="59" pageNumber="60" specimenCode="CASENT9017929, CASENT9017361" specimenCount="2" stateProvince="Mahajanga" typeStatus="holotype">
<typeStatus id="AC385C87187FFFE1461534EAAA8AECB0" box="[346,445,1509,1532]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
(
<specimenCode id="23254A5E187FFFE1468234EAA9A1ECB0" box="[461,662,1509,1532]" collectionCode="CASENT" pageId="59" pageNumber="60">CASENT9017929</specimenCode>
) and
<typeStatus id="AC385C87187FFFE1459A34EAA806ECB0" box="[725,817,1509,1532]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
(
<specimenCode id="23254A5E187FFFE1440E34EAAF3DECB0" box="[833,1034,1509,1532]" collectionCode="CASENT" pageId="59" pageNumber="60">CASENT9017361</specimenCode>
) males from tropical dry forest at
<quantity id="B47B4FC0187FFFE14613370AAAADEF50" box="[348,410,1541,1564]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="59" pageNumber="60" unit="m" value="140.0">
<elevation id="F8AE0516187FFFE14613370AAAADEF50" box="[348,410,1541,1564]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="59" pageNumber="60" unit="m" value="140.0">140m</elevation>
</quantity>
elevation (16°28ʹS, 45°21ʹE) in
<location id="765CB4FE187FFFE145B8370AAF13EF50" LSID="urn:lsid:plazi:treatment:FB2A5333187FFFE7473434AFAAC6E81F:765CB4FE187FFFE145B8370AAF13EF50" box="[759,1060,1541,1564]" country="Madagascar" county="California Academy of Sciences." municipality="Fisher-Griswold Arthropod Team" name="Parc National de Namoroka" pageId="59" pageNumber="60" stateProvince="Mahajanga">Parc National de Namoroka</location>
,
<collectingRegion id="B1472CC7187FFFE14360370AABC9EF71" country="Madagascar" name="Mahajanga" pageId="59" pageNumber="60">Mahajanga Province</collectingRegion>
,
<collectingCountry id="0B94A2B5187FFFE146463729AABFEF71" box="[265,392,1574,1597]" name="Madagascar" pageId="59" pageNumber="60">Madagascar</collectingCountry>
, collected
<date id="073DC4E5187FFFE146B13729A9ECEF71" box="[510,731,1574,1597]" pageId="59" pageNumber="60" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D187FFFE146B13729A9ECEF71" box="[510,731,1574,1597]" pageId="59" pageNumber="60" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">4–8 November 2002</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F187FFFE144623729AFA5EF71" box="[813,1170,1574,1597]" pageId="59" pageNumber="60">Fisher-Griswold Arthropod Team</collectingMunicipality>
, deposited in the
<collectingCounty id="9A5D9AA9187FFFE146613749A9A6EF11" box="[302,657,1606,1629]" pageId="59" pageNumber="60">California Academy of Sciences.</collectingCounty>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE187FFFE147E43769AFA1EFF1" pageId="59" pageNumber="60" type="etymology">
<paragraph id="733CE225187FFFE147E43769AFA1EFF1" blockId="59.[123,1174,1509,1725]" pageId="59" pageNumber="60">
<emphasis id="41F73E37187FFFE147E43769AA1DEF31" bold="true" box="[171,298,1638,1661]" pageId="59" pageNumber="60">Etymology.</emphasis>
The species name is from the Malagasy words
<emphasis id="41F73E37187FFFE1447F3769A846EF31" box="[816,881,1638,1661]" italics="true" pageId="59" pageNumber="60">lehibe</emphasis>
, meaning big or large, and
<emphasis id="41F73E37187FFFE147343789AB8CEFD1" box="[123,187,1670,1693]" italics="true" pageId="59" pageNumber="60">kokoa</emphasis>
, signifying bigger or larger. This refers to the large size of spiders of this species:
<taxonomicName id="B48399A6187FFFE143043789AA37EFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E37187FFFE143043789AA37EFF1" bold="true" italics="true" pageId="59" pageNumber="60">Uduba lehibekokoa</emphasis>
</taxonomicName>
are much larger than the similar
<taxonomicName id="B48399A6187FFFE1452E37A9A833EFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[609,772,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E37187FFFE1452E37A9A833EFF1" bold="true" box="[609,772,1702,1725]" italics="true" pageId="59" pageNumber="60">Uduba woodae</emphasis>
</taxonomicName>
. A compound adjective in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1878FFE647E431D3A899EB59" pageId="60" pageNumber="61" type="diagnosis">
<paragraph id="733CE2251878FFE647E431D3A899EB59" blockId="60.[122,1175,220,1725]" pageId="60" pageNumber="61">
<emphasis id="41F73E371878FFE647E431D3AACCE9BF" bold="true" box="[171,507,220,243]" pageId="60" pageNumber="61">
Diagnosis.
<taxonomicName id="B48399A61878FFE6466A31D3AACCE9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[293,507,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371878FFE6466A31D3AACCE9BF" bold="true" box="[293,507,220,243]" italics="true" pageId="60" pageNumber="61">Uduba lehibekokoa</emphasis>
</taxonomicName>
</emphasis>
are members of Group I.b, the
<taxonomicName id="B48399A61878FFE6440831D3A8D0E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[839,999,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="woodae">Uduba woodae</taxonomicName>
group (Map 14) of Group I, the Epigynal atrium group. They are cribellate, with divided cribellum or vestige. Males can be distinguished from those of other
<taxonomicName id="B48399A61878FFE645733013A9B1E87F" authorityName="Simon" authorityYear="1880" box="[572,646,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371878FFE645733013A9B1E87F" bold="true" box="[572,646,284,307]" italics="true" pageId="60" pageNumber="61">Uduba</emphasis>
</taxonomicName>
by having the palp with TA2 highly sclerotized, black, concave medially against TA3 (
<figureCitation id="EBB8FEA01878FFE645573033A9B5E81F" box="[536,642,316,339]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Fig. 50 A</figureCitation>
), distinguished from
<taxonomicName id="B48399A61878FFE644263033A8CEE81F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[873,1017,316,339]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371878FFE644263033A8CEE81F" bold="true" box="[873,1017,316,339]" italics="true" pageId="60" pageNumber="61">U. schlingeri</emphasis>
</taxonomicName>
by having the TA2 concave but not chelate (TA2 chelate in
<taxonomicName id="B48399A61878FFE6453B3053A831E83F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[628,774,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="41F73E371878FFE6453B3053A831E83F" bold="true" box="[628,774,348,371]" italics="true" pageId="60" pageNumber="61">U. schlingeri</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01878FFE644593053A8B9E83F" box="[790,910,348,371]" captionStart="FIGURE" captionStartId="152.[155,167,903,922]" captionTargetBox="[140,1164,228,880]" captionTargetId="figure-11@152.[120,1176,216,889]" captionTargetPageId="152" captionText="FIGURE 61. Uduba schlingeri, new species, (CASENT9006062) from Parc National Ranomafana, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158711" httpUri="https://zenodo.org/record/13158711/files/figure.png" pageId="60" pageNumber="61">Figs. 61 B</figureCitation>
,
<figureCitation id="EBB8FEA01878FFE644D33053A8C1E83F" box="[924,1014,348,371]" captionStart="FIGURE" captionStartId="153.[155,167,1176,1195]" captionTargetBox="[120,1176,216,1157]" captionTargetId="figure-16@153.[120,1176,216,1157]" captionTargetPageId="153" captionText="FIGURE 62. Uduba schlingeri, new species, CASENT9006059 from Andohahela, right male palpus, SEM. A. Tibia and tarsus, retrolateral. B. Tibia and tarsus, ventral. C. Tibia and tarsus, ventrolateral. D. Bulb, apical. E. Bulb, proventral. MA, median apophysis; RTA, retrolateral tibial apophysis on male palp; TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum; UTA, basal tibial apophysis on male palp; VTA, ventral tibial apophysis on male palp. Scale bars: A–C = 1500 µm, D = 600 µm, E = 1000 µm." figureDoi="http://doi.org/10.5281/zenodo.13158713" httpUri="https://zenodo.org/record/13158713/files/figure.png" pageId="60" pageNumber="61">62 C, E</figureCitation>
), and from
<taxonomicName id="B48399A61878FFE643333053ABFDE8D8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371878FFE643333053ABFDE8D8" bold="true" italics="true" pageId="60" pageNumber="61">U. taralily</emphasis>
</taxonomicName>
by having a simple MA (
<figureCitation id="EBB8FEA01878FFE646943072A975E8D8" box="[475,578,381,404]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Fig. 50 B</figureCitation>
) (
<taxonomicName id="B48399A61878FFE645143072A9FCE8D8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[603,715,381,404]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="taralily">
<emphasis id="41F73E371878FFE645143072A9FCE8D8" bold="true" box="[603,715,381,404]" italics="true" pageId="60" pageNumber="61">U. taralily</emphasis>
</taxonomicName>
with MA concave with lateral projections,
<figureCitation id="EBB8FEA01878FFE647343092ABD2E8F8" box="[123,229,413,436]" captionStart="FIGURE" captionStartId="154.[155,167,800,819]" captionTargetBox="[135,1162,230,786]" captionTargetId="figure-11@154.[120,1176,216,796]" captionTargetPageId="154" captionText="FIGURE 63. Uduba taralily, new species, Holotype (CASENT9042524) from Miandritsara Forest, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158715" httpUri="https://zenodo.org/record/13158715/files/figure.png" pageId="60" pageNumber="61">Fig. 63 B</figureCitation>
).
<taxonomicName id="B48399A61878FFE647B43092AAE4E8F8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[251,467,413,436]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371878FFE647B43092AAE4E8F8" bold="true" box="[251,467,413,436]" italics="true" pageId="60" pageNumber="61">Uduba lehibekokoa</emphasis>
</taxonomicName>
specimens are distinguished from
<taxonomicName id="B48399A61878FFE6441F3092A8F2E8F8" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[848,965,413,436]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371878FFE6441F3092A8F2E8F8" bold="true" box="[848,965,413,436]" italics="true" pageId="60" pageNumber="61">U. woodae</emphasis>
</taxonomicName>
by having the MA A–C, and specimens are larger size, total length 14.00–15.50 (
<taxonomicName id="B48399A61878FFE6445B30B2A8B0E898" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[788,903,445,468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="woodae">
<emphasis id="41F73E371878FFE6445B30B2A8B0E898" bold="true" box="[788,903,445,468]" italics="true" pageId="60" pageNumber="61">U. woodae</emphasis>
</taxonomicName>
males have a sharp point on the MA, the TA3 apex is blunt, the VTA is short and straight, length less than twice width,
<figureCitation id="EBB8FEA01878FFE6432E30D2ABE0EB59" captionStart="FIGURE" captionStartId="156.[155,167,847,866]" captionTargetBox="[129,1162,229,828]" captionTargetId="figure-11@156.[120,1176,216,837]" captionTargetPageId="156" captionText="FIGURE 66. Uduba woodae, new species, Holotype (CASENT9017834) from Lake Ranobe, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Giovanni Maki. TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158723" httpUri="https://zenodo.org/record/13158723/files/figure.png" pageId="60" pageNumber="61">Figs. 66 A–C</figureCitation>
, and specimens are less than 12.50 total length). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1878FFE747E43311A865E85F" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="description">
<paragraph id="733CE2251878FFE647E43311AFA1EDB6" blockId="60.[122,1175,220,1725]" pageId="60" pageNumber="61">
<emphasis id="41F73E371878FFE647E43311AA4EEB79" bold="true" box="[171,377,542,565]" pageId="60" pageNumber="61">Description. Male</emphasis>
(
<typeStatus id="AC385C871878FFE646C53311AAC3EB79" box="[394,500,542,565]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
): Total length 15.50. Markings as in
<figureCitation id="EBB8FEA01878FFE644C23311A8DAEB79" box="[909,1005,542,565]" captionStart="FIGURE" captionStartId="113.[155,167,1128,1147]" captionTargetBox="[120,1140,216,1104]" captionTargetId="figure-16@113.[120,1140,216,1104]" captionTargetPageId="113" captionText="FIGURE 9. Habitus photos of Uduba spp., males, dorsal views. A. U. schlingeri, (CASENT9006077) from R. S. de Ivohibe. B. U. woodae, CASENT9017831 from Lake Ranobe. C. U. lehibekokoa, Holotype (CASENT9017929) from tropical dry forest in Parc National de Namoroka. D. U. salegy, Holotype (CASENT9006044) from montane rainforest in Parc National Montagne d’Ambre. E. U. ibonia, CASENT9006034 from Parc National Ranomafana. F. U. fandroana, CASENT9006103 from Forêt d’Ankazomivady. G. U. kavanaughi, Holotype (CASENT9029870) from evergreen forest in Parc National Ranomafana. H. U. lamba, Holotype (CASENT9042542) from rainforest in Réserve Nationale Intégrale Betampona. Scale bars: A–F, H = 5 mm, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158580" httpUri="https://zenodo.org/record/13158580/files/figure.png" pageId="60" pageNumber="61">Fig. 9 C</figureCitation>
. Carapace 8.80 long, 6.70 wide, 3.80 high; clypeus 0.40 high. Eye diameters:AME 0.26, ALE 0.28, PME and PLE 0.34. Chelicerae 3.80 long; sternum 4.10 long, 3.00 wide; labium 2.10 long; palpal coxae 2.80 long. Cribellum divided. Spination: palpus–femur d1-1-3; leg I–femur d1-1-1, p0-1-1-1, r1-1-1-1, patella p1, r1, tibia d0-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-0-1-1, v2-2-1-2, r1-1-1-1; leg II–femur d1-1-1, p1-1-1-1, r0-1-1-1, patella p1, r1, tibia d0-0-1-0, p0-1-2-0, v2-2-2-2, r0-1-0-0, metatarsus p1-1-1, v2-2-1-2, r1-1-1; leg III–femur d1-1-1, p1-1-1-1, r1-0-1-1, patella r1, tibia d0- 1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-2-2, v2-2-1-2, r1-1-2; leg IV–femur d1-1-0, p1-1-1- 1, patella r1, tibia d0-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p0-1-1-2, v2-2-2-1-2, r0-1-1-2. Scopulae: cymbium, apicodorsal; tarsi and apices of metatarsi I–IV, ventral. Leg measurements: I: 9.00 + 3.90 + 9.30 + 9.60 + 4.70 = 36.50; II: 8.10 + 3.50 + 7.70 + 8.80 + 5.00 = 33.10; III: 7.10 + 3.10 + 5.00 + 7.30 + 3.80 = 26.30; IV: 9.20 + 3.20 + 7.60 + 11.50 + 5.00 = 36.50; palpus: 3.30 + 1.60 + 1.50 + NA + 4.30 = 10.70. Leg formula [1=4]23.
<emphasis id="41F73E371878FFE645A532AEA86AEAF4" bold="true" box="[746,861,929,952]" pageId="60" pageNumber="61">Male palp</emphasis>
: palpal tibia 0.36 times cymbial length, RTA broadly triangular, RTA length 0.29 tibia length, length 1.11 times width, with pointed apex (
<figureCitation id="EBB8FEA01878FFE6465D32EEAA4EEAB4" box="[274,377,993,1016]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Fig. 50 C</figureCitation>
), VTA tapering to a slender, hooked apex (
<figureCitation id="EBB8FEA01878FFE6447432EEA8FEEAB4" box="[827,969,993,1016]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Figs. 50 A, C</figureCitation>
), length 3.67 times width, VTA length 0.305 times tibia width; tibia lacking stout spines; tegulum convex, without ridge (
<figureCitation id="EBB8FEA01878FFE6478E352EAA62ED74" box="[193,341,1057,1080]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Figs. 50 B, C</figureCitation>
), tegulum length 0.98 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.32 times tegulum length; TA3 transverse, concave dorsally, tapering to a point that is recurved apically (
<figureCitation id="EBB8FEA01878FFE646AF356DA97DED35" box="[480,586,1122,1145]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Fig. 50 B</figureCitation>
), TA3 apex visible in front of TA2; TA2 highly sclerotized, black (
<figureCitation id="EBB8FEA01878FFE6464B358DAAA2EDD5" box="[260,405,1154,1177]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Figs. 50 B, C</figureCitation>
), concave medially against TA3, TA2 and TA3 apices at same level; MA transverse, simple, origin at 0.53 times tegulum length, tegulum apex at 0.22 tegulum length from MA apex, length (including apical processes) 0.44 times width, MA small, length 0.18 tegulum length, width 0.39 times tegulum width (
<figureCitation id="EBB8FEA01878FFE6457B35ECA9F0EDB6" box="[564,711,1251,1274]" captionStart="FIGURE" captionStartId="144.[155,167,1561,1580]" captionTargetBox="[130,1161,986,1550]" captionTargetId="figure-77@144.[120,1176,972,1551]" captionTargetPageId="144" captionText="FIGURE 50. Uduba lehibekokoa, new species, Holotype (CASENT9017929) from Parc National de Namoroka, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Daniele Polotow. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158685" httpUri="https://zenodo.org/record/13158685/files/figure.png" pageId="60" pageNumber="61">Figs. 50 A–C</figureCitation>
); conductor fan entire.
<emphasis id="41F73E371878FFE6448C35ECAF21EDB6" bold="true" box="[963,1046,1251,1274]" pageId="60" pageNumber="61">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="733CE2251878FFE647E5340CA94BEF50" blockId="60.[122,1175,220,1725]" pageId="60" pageNumber="61">
<emphasis id="41F73E371878FFE647E5340CAA24EC56" bold="true" box="[170,275,1283,1306]" pageId="60" pageNumber="61">Variation</emphasis>
(Males, N = 2). Total length 14.00–15.50, carapace length / width = 1.29–1.31, carapace height / width = 0.38–0.57, PER / carapace width = 0.31–0.34, PER / OAL = 2.64–2.82, PER / AER = 1.36–1.47, OAL / OQL = 1.00–1.05, OQP / OQA = 1.21–1.31, clypeus height / AME diameter width = 1.54–1.73, cheliceral length / clypeus height = 8.00–9.50, sternum length / width = 1.37–1.41, femur I length / carapace width =1.31–1.34, metatarsus I length / carapace width = 1.43–1.46, femur IV length / carapace width = 1.35–1.37, cymbium length / carapace width = 0.64– 0.65, cymbium length / palpal patella length = 2.69–4.20, cymbium length / palpal tibia length = 2.87–3.00, cymbium length / palpal femur length 1.24–1.30, palpal tibia length / palpal patella length = 0.94–1.40. Female variation unknown.
</paragraph>
<paragraph id="733CE2251878FFE647E4372AA890EFF1" blockId="60.[122,1175,220,1725]" pageId="60" pageNumber="61">
<materialsCitation id="C3EBE8781878FFE647E4372AAA8AEFD1" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" country="Madagascar" elevation="140" location="Fisher-Griswold Arthropod Team" municipality="Parc National de Namoroka" pageId="60" pageNumber="61" specimenCode="CASENT9017929" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="holotype">
M
<emphasis id="41F73E371878FFE6478D372AAABCEF70" bold="true" box="[194,395,1573,1596]" pageId="60" pageNumber="61">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51878FFE646DA372AA960EF70" box="[405,599,1573,1596]" name="Madagascar" pageId="60" pageNumber="61">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371878FFE6452A372AA87EEF70" box="[613,841,1573,1596]" italics="true" pageId="60" pageNumber="61">
<collectingRegion id="B1472CC71878FFE6452A372AA87EEF70" box="[613,841,1573,1596]" country="Madagascar" name="Mahajanga" pageId="60" pageNumber="61">Mahajanga Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1878FFE64415372AAFA6EF70" box="[858,1169,1573,1596]" pageId="60" pageNumber="61">Parc National de Namoroka</collectingMunicipality>
,
<locationDeviation id="628EFB071878FFE64735374AAAF3EF10" box="[122,452,1605,1628]" location="Vilanandro" pageId="60" pageNumber="61">
<quantity id="B47B4FC01878FFE64735374AABF2EF10" box="[122,197,1605,1628]" metricMagnitude="3" metricUnit="m" metricValue="9.8" pageId="60" pageNumber="61" unit="km" value="9.8">9.8 km</quantity>
300°WNW Vilanandro
</locationDeviation>
, 16°28ʹS, 45°21ʹE, elev.
<quantity id="B47B4FC01878FFE6459E374AA826EF10" box="[721,785,1605,1628]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="60" pageNumber="61" unit="m" value="140.0">
<elevation id="F8AE05161878FFE6459E374AA826EF10" box="[721,785,1605,1628]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="60" pageNumber="61" unit="m" value="140.0">140m</elevation>
</quantity>
, tropical dry forest, general collecting day spiders,
<date id="073DC4E51878FFE646773769A917EF31" box="[312,544,1638,1661]" pageId="60" pageNumber="61" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D1878FFE646773769A917EF31" box="[312,544,1638,1661]" pageId="60" pageNumber="61" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">4–8 November 2002</collectingDate>
</date>
,
<location id="765CB4FE1878FFE6457F3769A8AEEF31" LSID="urn:lsid:plazi:treatment:FB2A5333187FFFE7473434AFAAC6E81F:765CB4FE1878FFE6457F3769A8AEEF31" box="[560,921,1638,1661]" country="Madagascar" municipality="Parc National de Namoroka" name="Fisher-Griswold Arthropod Team" pageId="60" pageNumber="61" stateProvince="Mahajanga">Fisher-Griswold Arthropod Team</location>
[BLF6451] (
<typeStatus id="AC385C871878FFE6437F3769AFA5EF31" box="[1072,1170,1638,1661]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E1878FFE647343789AA74EFD1" box="[123,323,1670,1693]" collectionCode="CASENT" pageId="60" pageNumber="61">CASENT9017929</specimenCode>
,
<specimenCount id="658529AC1878FFE646013789AA58EFD1" box="[334,367,1670,1693]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01878FFE646313789AA82EFD1" box="[382,437,1670,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="61" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781878FFE646863789A894EFF1" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" country="Madagascar" location="Fisher-Griswold Arthropod Team" pageId="60" pageNumber="61" specimenCode="CASENT9017361" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="paratype">
general collecting night spiders,
<date id="073DC4E51878FFE644673789AF3FEFD1" box="[808,1032,1670,1693]" pageId="60" pageNumber="61" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D1878FFE644673789AF3FEFD1" box="[808,1032,1670,1693]" pageId="60" pageNumber="61" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">4–8 November 2002</collectingDate>
</date>
,
<location id="765CB4FE1878FFE6435B3789AA53EFF1" LSID="urn:lsid:plazi:treatment:FB2A5333187FFFE7473434AFAAC6E81F:765CB4FE1878FFE6435B3789AA53EFF1" country="Madagascar" name="Fisher-Griswold Arthropod Team" pageId="60" pageNumber="61" stateProvince="Mahajanga">Fisher-Griswold Arthropod Team</location>
[BLF6449] (
<typeStatus id="AC385C871878FFE646BC37A9A964EFF1" box="[499,595,1702,1725]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1878FFE6452F37A9A81FEFF1" box="[608,808,1702,1725]" collectionCode="CASENT" pageId="60" pageNumber="61">CASENT9017361</specimenCode>
,
<specimenCount id="658529AC1878FFE6447C37A9A862EFF1" box="[819,853,1702,1725]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01878FFE6442C37A9A8ACEFF1" box="[867,923,1702,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="61" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251879FFE747E431D3A865E85F" blockId="61.[123,1174,220,339]" pageId="61" pageNumber="62">
<emphasis id="41F73E371879FFE747E431D3AA69E9BF" bold="true" box="[171,350,220,243]" pageId="61" pageNumber="62">Natural history.</emphasis>
Members of the cribellate species
<taxonomicName id="B48399A61879FFE7458031D3A893E9BF" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[719,932,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371879FFE7458031D3A893E9BF" bold="true" box="[719,932,220,243]" italics="true" pageId="61" pageNumber="62">Uduba lehibekokoa</emphasis>
</taxonomicName>
occur at low elevation (
<quantity id="B47B4FC01879FFE747CD31F3ABF5E85F" box="[130,194,252,275]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="61" pageNumber="62" unit="m" value="140.0">140m</quantity>
) in tropical dry forest. We know nothing of their use of silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1879FFE747E43013AAC6E81F" pageId="61" pageNumber="62" type="distribution">
<paragraph id="733CE2251879FFE747E43013AAC6E81F" blockId="61.[123,1174,220,339]" pageId="61" pageNumber="62">
<emphasis id="41F73E371879FFE747E43013AA0AE87F" bold="true" box="[171,317,284,307]" pageId="61" pageNumber="62">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61879FFE7469D3013A99CE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[466,683,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="lehibekokoa">
<emphasis id="41F73E371879FFE7469D3013A99CE87F" bold="true" box="[466,683,284,307]" italics="true" pageId="61" pageNumber="62">Uduba lehibekokoa</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="AC385C871879FFE744883013A8C2E87F" box="[967,1013,284,307]" pageId="61" pageNumber="62">type</typeStatus>
locality in far western
<collectingCountry id="0B94A2B51879FFE747993033AA60E81F" box="[214,343,316,339]" name="Madagascar" pageId="61" pageNumber="62">Madagascar</collectingCountry>
(
<figureCitation id="EBB8FEA01879FFE746293033AA8DE81F" box="[358,442,316,339]" captionStart="MAP" captionStartId="179.[155,174,1076,1095]" captionTargetBox="[120,1176,216,1064]" captionTargetId="figure-17@179.[648,1176,216,1061]" captionTargetPageId="179" captionText="MAP 7. Distributions of Uduba evanescens (Dahl, 1901) MAP 8. Distributions of Uduba woodae, Uduba lavitra and Uduba lehibekokoa. and Uduba milamina." figureDoi="http://doi.org/10.5281/zenodo.13158781" httpUri="https://zenodo.org/record/13158781/files/figure.png" pageId="61" pageNumber="62">Maps 7</figureCitation>
, 14).
</paragraph>
</subSubSection>
</treatment>
</document>